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<document ID-DOI="10.15407/zoo2022.04.291" ID-ISSN="2707-7268" ID-Zenodo-Dep="7175521" approvalRequired="10" approvalRequired_for_textStreams="10" checkinTime="1665293873903" checkinUser="felipe" docAuthor="Zamani, A. &amp; Marusik, Y. M." docDate="2022" docId="D76C87CB113CFFABF8BD9991C5C7FD16" docLanguage="en" docName="Zoodiversity.56.4.291–306.pdf" docOrigin="Zoodiversity 56 (4)" docSource="http://dx.doi.org/10.15407/zoo2022.04.291" docStyle="DocumentStyle:65E6F982F72656C606C1639D3996B172.1:Zoodiversity.2020-.journal_article" docStyleId="65E6F982F72656C606C1639D3996B172" docStyleName="Zoodiversity.2020-.journal_article" docStyleVersion="1" docTitle="Zelotes jakesi Zamani &amp; Marusik 2022, sp. n." docType="treatment" docVersion="3" lastPageNumber="295" masterDocId="2B55FFB3113FFFAFF813985EC526FFBC" masterDocTitle="On A Small Collection Of Spiders (Arachnida, Araneae) From Iraq, With New Species And Records" masterLastPageNumber="306" masterPageNumber="291" pageNumber="294" updateTime="1665311832057" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-NC-ND-4.0">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>On A Small Collection Of Spiders (Arachnida, Araneae) From Iraq, With New Species And Records</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Zamani, A.</mods:namePart>
<mods:affiliation>Zoological Museum, Biodiversity Unit, University of Turku, FI- 20014, Finland</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Marusik, Y. M.</mods:namePart>
<mods:affiliation>Department of Zoology &amp; Entomology, University of the Free State, Bloemfontein 9300, South Africa</mods:affiliation>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>Zoodiversity</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2022</mods:date>
<mods:detail type="volume">
<mods:number>56</mods:number>
</mods:detail>
<mods:detail type="issue">
<mods:number>4</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>291</mods:start>
<mods:end>306</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location>
<mods:url>http://dx.doi.org/10.15407/zoo2022.04.291</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">10.15407/zoo2022.04.291</mods:identifier>
<mods:identifier type="ISSN">2707-7268</mods:identifier>
<mods:identifier type="Zenodo-Dep">7175521</mods:identifier>
</mods:mods>
<treatment ID-DOI="http://doi.org/10.5281/zenodo.7171810" ID-Zenodo-Dep="7171810" LSID="urn:lsid:plazi:treatment:D76C87CB113CFFABF8BD9991C5C7FD16" httpUri="http://treatment.plazi.org/id/D76C87CB113CFFABF8BD9991C5C7FD16" lastPageId="4" lastPageNumber="295" pageId="3" pageNumber="294">
<subSubSection pageId="3" pageNumber="294" type="nomenclature">
<paragraph blockId="3.[174,559,463,490]" box="[174,559,463,490]" pageId="3" pageNumber="294">
<heading box="[174,559,463,490]" fontSize="36" level="2" pageId="3" pageNumber="294" reason="3">
<taxonomicName authority="Zamani &amp; Marusik, 2022" authorityName="Zamani &amp; Marusik" authorityYear="2022" box="[174,334,463,490]" class="Arachnida" family="Gnaphosidae" genus="Zelotes" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="294" phylum="Arthropoda" rank="species" species="jakesi" status="sp. n.">Zelotes jakesi</taxonomicName>
<taxonomicNameLabel box="[341,405,463,490]" pageId="3" pageNumber="294" rank="species">
<emphasis bold="true" box="[341,399,463,489]" pageId="3" pageNumber="294">sp. n</emphasis>
.
</taxonomicNameLabel>
(
<figureCitation box="[419,480,463,489]" captionStart="Fig" captionStartId="3.[174,207,1774,1796]" captionTargetBox="[231,1176,991,1758]" captionTargetId="figure-338@3.[231,1176,982,1764]" captionTargetPageId="3" captionText="Fig. 2. Female of Zelotes jakesi sp. n.: A — habitus, dorsal view; B — epigyne, ventral view; C — vulva, dorsal view; D — same, dorsolateral view. Abbreviations: Ah — anterior hood, Ag — accessorial gland, Re — receptacle. Scale bars = 0.2 mm, unless stated otherwise." figureDoi="http://doi.org/10.5281/zenodo.7175525" httpUri="https://zenodo.org/record/7175525/files/figure.png" pageId="3" pageNumber="294">fig. 2</figureCitation>
, A–D)
</heading>
</paragraph>
<paragraph blockId="3.[174,850,509,531]" box="[174,850,509,531]" pageId="3" pageNumber="294">
<uri box="[174,850,509,531]" pageId="3" pageNumber="294">
urn:lsid:zoobank.org:act:
<uuid box="[402,850,509,531]" pageId="3" pageNumber="294">E3C7D40C-129A-48BE-9927-0572A2696B50</uuid>
</uri>
</paragraph>
</subSubSection>
<subSubSection lastPageId="4" lastPageNumber="295" pageId="3" pageNumber="294" type="materials_examined">
<paragraph blockId="3.[174,1233,551,601]" pageId="3" pageNumber="294">
<materialsCitation collectingDate="1978-11-07" collectionCode="MMB" collectorName="O. Jakes" country="Iraq" elevation="260" latitude="30.8" location="Ash Shabakah" longLatPrecision="1220" longitude="43.65" pageId="3" pageNumber="294" specimenCount="1" stateProvince="An Najaf" typeStatus="holotype">
Material. Type.
<emphasis bold="true" box="[405,499,552,573]" pageId="3" pageNumber="294">
<typeStatus box="[405,499,552,573]" pageId="3" pageNumber="294">Holotype</typeStatus>
</emphasis>
}:
<collectingCountry box="[536,578,553,574]" name="Iraq" pageId="3" pageNumber="294">Iraq</collectingCountry>
:
<collectingRegion box="[589,737,552,574]" country="Iraq" name="An Najaf" pageId="3" pageNumber="294">Najaf Province</collectingRegion>
:
<location LSID="urn:lsid:plazi:treatment:D76C87CB113CFFABF8BD9991C5C7FD16:5A1A6006113CFFACFAFE9A76C65EFD82" box="[749,888,552,574]" country="Iraq" latitude="30.8" longLatPrecision="1220" longitude="43.65" name="Ash Shabakah" pageId="3" pageNumber="294" stateProvince="An Najaf">Ash Shabakah</location>
camp,
<quantity box="[962,1037,552,574]" metricMagnitude="5" metricUnit="m" metricValue="1.5" pageId="3" pageNumber="294" unit="km" value="150.0">150 km</quantity>
SW of
<collectingRegion box="[1115,1169,552,574]" country="Iraq" name="An Najaf" pageId="3" pageNumber="294">Najaf</collectingRegion>
, stone desert, [
<geoCoordinate box="[249,334,579,600]" degrees="30" direction="north" minutes="48" orientation="latitude" pageId="3" pageNumber="294" precision="925" value="30.8">30°48' N</geoCoordinate>
,
<geoCoordinate box="[344,425,579,600]" degrees="43" direction="east" minutes="39" orientation="longitude" pageId="3" pageNumber="294" precision="925" value="43.65">43°39' E</geoCoordinate>
],
<quantity box="[440,501,579,601]" metricMagnitude="2" metricUnit="m" metricValue="2.6" pageId="3" pageNumber="294" unit="m" value="260.0">
<elevation box="[440,501,579,601]" metricMagnitude="2" metricUnit="m" metricValue="2.6" pageId="3" pageNumber="294" unit="m" value="260.0">260 m</elevation>
</quantity>
,
<date box="[511,614,579,600]" pageId="3" pageNumber="294" value="1978-11-07">
<collectingDate box="[511,614,579,600]" pageId="3" pageNumber="294" value="1978-11-07">11.07.1978</collectingDate>
</date>
(
<collectorName box="[627,708,579,601]" pageId="3" pageNumber="294">O. Jakeš</collectorName>
) (
<collectionCode box="[727,789,579,600]" country="Czech Republic" httpUri="http://grbio.org/cool/ry0g-1y6k" name="Moravske Muzeum" pageId="3" pageNumber="294">MMB</collectionCode>
).
</materialsCitation>
</paragraph>
<paragraph blockId="3.[174,1234,623,970]" box="[227,1206,623,650]" pageId="3" pageNumber="294">
Etymology. This species is named after Oldřich Jakeš, who collected the
<typeStatus box="[1096,1202,623,649]" pageId="3" pageNumber="294">holotype</typeStatus>
.
</paragraph>
<paragraph blockId="3.[174,1234,623,970]" pageId="3" pageNumber="294">
Diagnosis. The new species is most similar to
<taxonomicName authority="Fomichev &amp; Marusik, 2021" authorityName="Fomichev &amp; Marusik" authorityYear="2021" class="Arachnida" family="Gnaphosidae" genus="Zelotes" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="294" phylum="Arthropoda" rank="species" species="khatlonicus">
<emphasis box="[800,966,655,681]" italics="true" pageId="3" pageNumber="294">Z. khatlonicus</emphasis>
Fomichev &amp; Marusik, 2021
</taxonomicName>
from
<collectingCountry box="[308,428,687,713]" name="Tajikistan" pageId="3" pageNumber="294">Tajikistan</collectingCountry>
, but differs by the accessorial glands directed mesally (
<emphasis box="[1083,1106,688,713]" italics="true" pageId="3" pageNumber="294">vs</emphasis>
. laterally), shape of the copulatory ducts (cf.
<figureCitation box="[577,636,719,745]" captionStart="Fig" captionStartId="3.[174,207,1774,1796]" captionTargetBox="[231,1176,991,1758]" captionTargetId="figure-338@3.[231,1176,982,1764]" captionTargetPageId="3" captionText="Fig. 2. Female of Zelotes jakesi sp. n.: A — habitus, dorsal view; B — epigyne, ventral view; C — vulva, dorsal view; D — same, dorsolateral view. Abbreviations: Ah — anterior hood, Ag — accessorial gland, Re — receptacle. Scale bars = 0.2 mm, unless stated otherwise." figureDoi="http://doi.org/10.5281/zenodo.7175525" httpUri="https://zenodo.org/record/7175525/files/figure.png" pageId="3" pageNumber="294">fig. 2</figureCitation>
, B–D and Fomichev &amp; Marusik, 2021: figs 47–49), and wider receptacles (receptacle as wide as anterior hood
<emphasis box="[860,883,752,777]" italics="true" pageId="3" pageNumber="294">vs</emphasis>
. hood wider than receptacle).
</paragraph>
<paragraph blockId="3.[174,1234,623,970]" pageId="3" pageNumber="294">
Description. Female. Habitus as in
<figureCitation box="[674,735,783,809]" captionStart="Fig" captionStartId="3.[174,207,1774,1796]" captionTargetBox="[231,1176,991,1758]" captionTargetId="figure-338@3.[231,1176,982,1764]" captionTargetPageId="3" captionText="Fig. 2. Female of Zelotes jakesi sp. n.: A — habitus, dorsal view; B — epigyne, ventral view; C — vulva, dorsal view; D — same, dorsolateral view. Abbreviations: Ah — anterior hood, Ag — accessorial gland, Re — receptacle. Scale bars = 0.2 mm, unless stated otherwise." figureDoi="http://doi.org/10.5281/zenodo.7175525" httpUri="https://zenodo.org/record/7175525/files/figure.png" pageId="3" pageNumber="294">fig. 2</figureCitation>
, A. Total length 7.05. Carapace 2.53 long, 1.94 wide. Eye sizes: AME: 0.09, ALE: 0.11, PME: 0.08, PLE: 0.09. Carapace, sternum, chelicerae, labium and maxillae dark reddish brown. Legs slightly lighter than carapace, without annulations. Abdomen beige. Spinnerets dark greyish brown, lighter distally. Measurements of legs: I: 6.16 (1.82, 1.11, 1.34, 1.13, 0.76), II: 5.53 (1.63, 0.91, 1.16, 1.06, 0.77), III: missing, IV: 6.56 + missing Ta (1.91, 1.03, 1.61, 2.01, missing).
</paragraph>
<caption ID-DOI="http://doi.org/10.5281/zenodo.7175525" ID-Zenodo-Dep="7175525" httpUri="https://zenodo.org/record/7175525/files/figure.png" pageId="3" pageNumber="294" startId="3.[174,207,1774,1796]" targetBox="[231,1176,991,1758]" targetPageId="3">
<paragraph blockId="3.[174,1232,1774,1849]" pageId="3" pageNumber="294">
Fig. 2. Female of
<taxonomicName authority="Zamani &amp; Marusik, 2022" authorityName="Zamani &amp; Marusik" authorityYear="2022" box="[342,468,1774,1795]" class="Arachnida" family="Gnaphosidae" genus="Zelotes" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="294" phylum="Arthropoda" rank="species" species="jakesi" status="sp. n.">
<emphasis box="[342,468,1774,1795]" italics="true" pageId="3" pageNumber="294">Zelotes jakesi</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[474,527,1774,1795]" pageId="3" pageNumber="294" rank="species">sp. n.</taxonomicNameLabel>
: A — habitus, dorsal view; B — epigyne, ventral view; C — vulva, dorsal view; D — same, dorsolateral view. Abbreviations:
<emphasis box="[718,746,1800,1821]" italics="true" pageId="3" pageNumber="294">Ah</emphasis>
— anterior hood,
<emphasis box="[935,961,1801,1822]" italics="true" pageId="3" pageNumber="294">Ag</emphasis>
— accessorial gland,
<emphasis box="[1178,1202,1801,1822]" italics="true" pageId="3" pageNumber="294">Re</emphasis>
— receptacle. Scale bars = 0.2 mm, unless stated otherwise.
</paragraph>
</caption>
<paragraph blockId="4.[166,1225,175,362]" pageId="4" pageNumber="295">
Epigyne as in
<figureCitation box="[384,444,175,201]" captionStart="Fig" captionStartId="3.[174,207,1774,1796]" captionTargetBox="[231,1176,991,1758]" captionTargetId="figure-338@3.[231,1176,982,1764]" captionTargetPageId="3" captionText="Fig. 2. Female of Zelotes jakesi sp. n.: A — habitus, dorsal view; B — epigyne, ventral view; C — vulva, dorsal view; D — same, dorsolateral view. Abbreviations: Ah — anterior hood, Ag — accessorial gland, Re — receptacle. Scale bars = 0.2 mm, unless stated otherwise." figureDoi="http://doi.org/10.5281/zenodo.7175525" httpUri="https://zenodo.org/record/7175525/files/figure.png" pageId="4" pageNumber="295">fig. 2</figureCitation>
, B–D; epigynal plate about 1.4 times longer than wide; fovea long, 3 times longer than wide; anterior hood (
<emphasis box="[656,690,207,233]" italics="true" pageId="4" pageNumber="295">Ah</emphasis>
) short, as wide as receptacle (
<emphasis box="[1045,1074,208,233]" italics="true" pageId="4" pageNumber="295">Re</emphasis>
); accessorial glands (
<emphasis box="[259,291,240,266]" italics="true" pageId="4" pageNumber="295">Ag</emphasis>
) directed antero-mesally.
</paragraph>
<paragraph blockId="4.[166,1225,175,362]" box="[219,418,271,297]" pageId="4" pageNumber="295">Male. Unknown.</paragraph>
<paragraph blockId="4.[166,1225,175,362]" pageId="4" pageNumber="295">
Distribution. Currently known only from the
<typeStatus box="[833,883,304,330]" pageId="4" pageNumber="295">type</typeStatus>
locality in
<collectingRegion box="[1035,1220,303,329]" country="Iraq" name="An Najaf" pageId="4" pageNumber="295">Najaf Province</collectingRegion>
, southern
<collectingCountry box="[278,330,336,362]" name="Iraq" pageId="4" pageNumber="295">Iraq</collectingCountry>
.
</paragraph>
<paragraph blockId="4.[166,550,399,458]" box="[166,550,399,425]" pageId="4" pageNumber="295">
Family
<taxonomicName authority="Thorell, 1870" authorityName="Thorell" authorityYear="1870" box="[254,550,399,425]" class="Arachnida" family="Hersiliidae" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="295" phylum="Arthropoda" rank="family">
<emphasis bold="true" box="[254,390,399,425]" pageId="4" pageNumber="295">Hersiliidae</emphasis>
Thorell, 1870
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection pageId="4" pageNumber="295" type="description">
<paragraph blockId="4.[166,550,399,458]" box="[166,529,431,458]" pageId="4" pageNumber="295">
Genus
<taxonomicName authority="Thorell, 1870" authorityName="Thorell" authorityYear="1870" box="[249,529,431,458]" class="Arachnida" family="Hersiliidae" genus="Hersiliola" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="295" phylum="Arthropoda" rank="genus">
<emphasis bold="true" box="[249,369,431,458]" italics="true" pageId="4" pageNumber="295">Hersiliola</emphasis>
Thorell, 1870
</taxonomicName>
</paragraph>
<paragraph blockId="4.[166,1226,495,682]" pageId="4" pageNumber="295">
Comments. The genus is rather well studied in the Middle East thanks to several regional revisions and smaller taxonomic contributions (e. g. Foord &amp; DippenaarSchoeman, 2005;
<bibRefCitation author="Danisman, T. &amp; Sancak, Z. &amp; Erdek, M. &amp; Cosar, I." box="[385,658,559,586]" pageId="4" pageNumber="295" pagination="85 - 88" refId="ref7569" refString="Danisman, T., Sancak, Z., Erdek, M., Cosar, I. 2012. A new species of the genus Hersiliola Thorell, 1870 from Turkey (Araneae: Hersiliidae). Zoology in the Middle East, 55, 85 - 88. https: // doi. org / 10.1080 / 09397140. 2012.10648922" type="journal article" year="2012">Danışman et al., 2012</bibRefCitation>
;
<bibRefCitation author="Marusik, Y. M. &amp; Fet, V." box="[671,929,559,586]" pageId="4" pageNumber="295" pagination="75 - 114" refId="ref8195" refString="Marusik, Y. M., Fet, V. 2009. A survey of east Palearctic Hersiliola Thorell, 1870 (Araneae, Hersiliidae), with a description of three new genera. ZooKeys, 16, 75 - 114. https: // doi. org / 10.3897 / zookeys. 16.229" type="journal article" year="2009">Marusik &amp; Fet, 2009</bibRefCitation>
;
<bibRefCitation author="Mirshamsi, O. &amp; Zamani, A. &amp; Marusik, Y. M." box="[942,1220,559,586]" pageId="4" pageNumber="295" pagination="1447 - 1461" refId="ref8251" refString="Mirshamsi, O., Zamani, A., Marusik, Y. M. 2016. A survey of Hersiliidae (Arachnida: Araneae) of Iran with description of one new genus and two new species. Journal of Natural History, 50 (23 - 24), 1447 - 1461. https: // doi. org / 10.1080 / 00222933.2015.1113318" type="journal article" year="2016">Mirshamsi et al., 2016</bibRefCitation>
;
<bibRefCitation author="Zamani, A. &amp; Mirshamsi, O. &amp; Marusik, Y. M." box="[168,420,591,618]" pageId="4" pageNumber="295" pagination="624 - 629" refId="ref8950" refString="Zamani, A., Mirshamsi, O., Marusik, Y. M. 2017. Description of a new species of Hersiliola and the male of Duninia rheimsae Marusik &amp; Fet, 2009 from Iran (Araneae: Hersiliidae). Turkish Journal of Zoology, 41 (4), 624 - 629. https: // doi. org / 10.3906 / zoo- 1609 - 53" type="journal article" year="2017">Zamani et al., 2017</bibRefCitation>
). Six species are currently known from the Middle East (
<bibRefCitation author="WSC" pageId="4" pageNumber="295" refId="ref8714" refString="WSC. 2022. World Spider Catalog. Version 23.0. Natural History Museum Bern, online at http: // wsc. nmbe. ch (accessed on 18.05.2022). https: // doi. org / 10.24436 / 2" type="url" year="2022">WSC, 2022</bibRefCitation>
). The new species described herein represents the first record of
<taxonomicName authorityName="Thorell" authorityYear="1870" box="[1069,1189,623,649]" class="Arachnida" family="Hersiliidae" genus="Hersiliola" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="295" phylum="Arthropoda" rank="genus">
<emphasis box="[1069,1189,623,649]" italics="true" pageId="4" pageNumber="295">Hersiliola</emphasis>
</taxonomicName>
in
<collectingCountry box="[166,221,656,682]" name="Iraq" pageId="4" pageNumber="295">Iraq</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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