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<document id="588890068AE1FD69A29D1FDED406600B" ID-CLB-Dataset="305638" ID-DOI="10.3897/zookeys.1219.137409" ID-GBIF-Dataset="74ccf9d0-0a5e-4c57-9e89-203eeab9cd69" ID-ZooBank="BD49B17B-1B71-4FB2-ABD3-84AEDD7ADB96" ID-publisher-id="137409" URI-arpha="BE76C7F3-C1C2-5150-937D-587FABC5B8C4" URI-zoobank="http://zoobank.org/BD49B17B-1B71-4FB2-ABD3-84AEDD7ADB96" XM.bibliography_approvedBy="admin" XM.materialsCitations_approvedBy="admin" XM.treatments_approvedBy="admin" article-type="research-article" checkinTime="1732785290019" checkinUser="pensoft" docAuthor="Cumming, Royce T. &amp; Mlynarek, Julia J." docDate="2024" docId="B0B5C9C1779A54B3AD1EF7E6FEA73551" docLanguage="en" docName="ZooKeys 1219: 123-133" docOrigin="ZooKeys 1219" docSource="https://zookeys.pensoft.net/article/137409/download/xml/" docStyle="DocumentStyle:PensoftTaxPub.0000.journal_article.generic" docStyleName="PensoftTaxPub.0000.journal_article.generic" docTitle="Sauronaradus meganae Cumming &amp; Mlynarek, 2024, sp. nov." docType="treatment" docVersion="5" dtd-version="3.0" lastPageNumber="133" masterDocId="BE76C7F3C1C25150937D587FABC5B8C4" masterDocTitle="An additional ✝ Archearadinae flat-bug species from Cretaceous Burmese Amber (Hemiptera, Aradidae)" masterLastPageNumber="133" masterPageNumber="123" pageNumber="123" updateTime="1732867382473" updateUser="ExternalLinkService">
<mods:mods id="D697FA1BEA9CE09A5252E3E5EC858305" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="3777FAA2DFAC2F52F1E236B6CE8D5185">
<mods:title id="C590EEF5DA1A67E2D9FC39428D1896F9">An additional ✝ Archearadinae flat-bug species from Cretaceous Burmese Amber (Hemiptera, Aradidae)</mods:title>
</mods:titleInfo>
<mods:name id="0A48BDC7555ED016018D25FF0FF2A183" type="personal">
<mods:role id="90B94544B667F7EBA2B6186B73AC4241">
<mods:roleTerm id="1B14D30645E5E56B56FE159D3F175704">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="FE8FAEF3E8B8AF699113AD1513D2EE7B">Cumming, Royce T.</mods:namePart>
<mods:nameIdentifier id="48D72035D9A2753DFDDB8B21F48F83F3" type="ORCID">0000-0001-7930-1292</mods:nameIdentifier>
<mods:affiliation id="E94E39D3308791CC21B2607E191B8428">Montreal Insectarium, 4581 rue Sherbrooke est, Montréal, H 1 X 2 B 2, Québec, Canada &amp; Richard Gilder Graduate School, American Museum of Natural History, New York, NY 10024, USA &amp; Biology, Graduate Center, City University of New York, New York, USA</mods:affiliation>
</mods:name>
<mods:name id="57B62AB474E6A06C7A00951B8D5D48D8" type="personal">
<mods:role id="18EEC307D25C6A7C8695561485BEFD04">
<mods:roleTerm id="11F25F51C7481CC1616FB15668C66814">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="D04C6FA54EF233D3F6DF9B7F0AD3725B">Mlynarek, Julia J.</mods:namePart>
<mods:nameIdentifier id="CE8F5FD299CD67358327EEEC7E375193" type="ORCID">0000-0002-1569-9403</mods:nameIdentifier>
<mods:affiliation id="4296BB6E5FA77AE4B60DAD0A2727B066">Montreal Insectarium, 4581 rue Sherbrooke est, Montréal, H 1 X 2 B 2, Québec, Canada</mods:affiliation>
</mods:name>
<mods:typeOfResource id="32B00724B8E84772BFFA0D4C7841D00F">text</mods:typeOfResource>
<mods:relatedItem id="65D8EEAA353E78FF97A32F78D99C97AB" type="host">
<mods:titleInfo id="42AB152D97447D9EFA519917C1A1C0CE">
<mods:title id="D7CB0C353604DBFF0447003AF421D560">ZooKeys</mods:title>
</mods:titleInfo>
<mods:part id="17C8B3428672B366C1E247EC31687EEA">
<mods:date id="C9D288A0B8357651C55AFD025C41BBCF">2024</mods:date>
<mods:detail id="CFBE423FA259094CA0621410432EEB18" type="pubDate">
<mods:number id="292D5B7F75C34FD11BFA23C6C64043CA">2024-11-28</mods:number>
</mods:detail>
<mods:detail id="8ADB2348CC88D9432C27D8FD085B38FA" type="volume">
<mods:number id="785F3BDAB66FD058E881EC20B7CE8011">1219</mods:number>
</mods:detail>
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<mods:start id="0EC818C16D30FD1BE772FB10DF520BCC">123</mods:start>
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</mods:extent>
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<mods:classification id="B3741F02C5F201C648DD92280CC33D00">journal article</mods:classification>
<mods:identifier id="E28987C4B1A193E71D8016219EE1C299" type="CLB-Dataset">305638</mods:identifier>
<mods:identifier id="C81CDD08E0AD7E65273A5B1A5919240C" type="DOI">10.3897/zookeys.1219.137409</mods:identifier>
<mods:identifier id="FCC7DBCCAB71C12C3765D6A9D18DD050" type="GBIF-Dataset">74ccf9d0-0a5e-4c57-9e89-203eeab9cd69</mods:identifier>
<mods:identifier id="795D55D9B6C65303A2AB70A34952A126" type="ZooBank">BD49B17B-1B71-4FB2-ABD3-84AEDD7ADB96</mods:identifier>
</mods:mods>
<treatment id="B0B5C9C1779A54B3AD1EF7E6FEA73551" ID-DOI="http://doi.org/10.5281/zenodo.14238469" ID-GBIF-Taxon="247966320" ID-Zenodo-Dep="14238469" ID-arpha="B0B5C9C1-779A-54B3-AD1E-F7E6FEA73551" ID-zoobank="https://zoobank.org/CC59C92B-2178-4C3C-B599-2114D420D433" LSID="urn:lsid:plazi:treatment:B0B5C9C1779A54B3AD1EF7E6FEA73551" LSID-ZBK="urn:lsid:zoobank.org:act:CC59C92B-2178-4C3C-B599-2114D420D433" httpUri="http://treatment.plazi.org/id/B0B5C9C1779A54B3AD1EF7E6FEA73551" scope_family="Aradidae" scope_infraOrder="Pentatomomorpha" scope_order="Hemiptera" scope_subOrder="Heteroptera">
<subSubSection id="C121AEA28CC486CF88BD250C4D326757" type="nomenclature">
<paragraph id="9454D928A19BCCA5C3C3B93D03050CF6">
<taxonomicName id="CB10ABB09DE048FF97838A4B32213941" authorityName="Cumming &amp; Mlynarek" authorityYear="2024" class="Insecta" family="Aradidae" genus="Sauronaradus" kingdom="Animalia" order="Hemiptera" phylum="Arthropoda" rank="species" species="meganae" status="sp. nov.">
<emphasis id="1429DABE85F5A3CB0B127FE087CC617B" italics="true">Sauronaradus meganae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="2E2C75DA2A3A7FF06421E707DCD49AA3" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="D15548A8D6B83B751E3CC9A7AA81CA9B" type="description">
<paragraph id="D0DDE877AB36D8AC7136123A6156266E">
<figureCitation id="9F9754A340B469BA5A9F7F774C81BAB0" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Sauronaradus meganae gen. et sp. nov. holotype A dorsal habitus B ventral habitus C abdomen, dorsal D abdomen, ventral E right antenna, dorsal." figureDoi="10.3897/zookeys.1219.137409.figure1" httpUri="https://binary.pensoft.net/fig/1184492">Figs 1</figureCitation>
,
<figureCitation id="F541E10C6A50541989920BAF9AA4A4B8" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Sauronaradus meganae gen. et sp. nov. holotype A anterior lit to highlight the spines of the head, dorsal B head through thorax, dorsal C pronotum, lit to highlight carinae, dorsal D full amber piece with holotype inclusion and Cretopiesma suukyiae syninclusion to the lower right of Sauronaradus meganae gen. et sp. nov. E head through thorax, ventral F pro- and meso- legs, mesotibia and tarsus visible, dorsal G right metatibia, dorsal." figureDoi="10.3897/zookeys.1219.137409.figure2" httpUri="https://binary.pensoft.net/fig/1184493">2</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="SECID0EAPAE" type="type material">
<paragraph id="9B4ED440F8AF93F86708B35F404C6057">
<heading id="6221C1C87ACF02A265017A72ED404176" reason="title">Type material.</heading>
</paragraph>
<paragraph id="A77F88BA3D1B4F365A1D048AA349E2CE">
<materialsCitation id="8A0C13573A1014B19D7D316FBEEFE1D5" ID-GBIF-Occurrence="4986844301" collectionCode="IMQC" country="Myanmar" location="Hukawng Valley" municipality="Montreal Insectarium" specimenCode="IMQC-AMB-ara 0001" specimenCount="1" typeStatus="holotype">
<emphasis id="FD5D0A0C2DB8271D2B2947D7347710F7" bold="true" italics="true">
<typeStatus id="7C1123607E64E4FA706802681DF22EAD" type="holotype">Holotype</typeStatus>
</emphasis>
: specimen number
<specimenCode id="99CB36694CB15266C829A74C5A964D6B">
<abbrev id="ABBRID0ELPAE" xlink_title="Montreal Insectarium">IMQC</abbrev>
-AMB-ara 0001
</specimenCode>
;
<location id="5409866EEED3DB78ADEF3BB69580DECF" LSID="urn:lsid:plazi:treatment:B0B5C9C1779A54B3AD1EF7E6FEA73551:5409866EEED3DB78ADEF3BB69580DECF" country="Myanmar" county="Hukawng Valley" municipality="Montreal Insectarium" name="Hukawng Valley" stateProvince="Quebec">Hukawng Valley</location>
,
<collectingCountry id="BA38AA69F7E7D29D07DA21988F9D292D" name="Myanmar">Myanmar</collectingCountry>
, accession c. 2010; male; specimen deposited within the Montreal Insectarium, Montreal, Quebec, Canada (
<abbrev id="ABBRID0EPPAE" xlink_title="Montreal Insectarium">
<collectionCode id="63052CDF666B829257540ADE5C078C94">IMQC</collectionCode>
</abbrev>
).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="SECID0ETPAE" type="Taphonomy">
<paragraph id="5589D6316F3A7AFD9729D1A2E9E246B5">
<heading id="198B6B9B8848AEE0E5D9E9A7559DF484" reason="title">Taphonomy.</heading>
</paragraph>
<paragraph id="0FE418C8CE1F0018EC74F32879A53095">
Ovular piece of clear amber (~ 15.5 ×
<quantity id="77E110A36D1D9BE73837A2F04D2AC2DE" metricMagnitude="-2" metricUnit="m" metricValue="1.09" unit="mm" value="10.9">10.9 mm</quantity>
), with minimal debris obscuring the
<typeStatus id="690BF3C06EE33A40F916453CA7D4BA7D" type="holotype">holotype</typeStatus>
(Fig.
<figureCitation id="B2703869D324C180FBA06E79B5B567F2" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Sauronaradus meganae gen. et sp. nov. holotype A anterior lit to highlight the spines of the head, dorsal B head through thorax, dorsal C pronotum, lit to highlight carinae, dorsal D full amber piece with holotype inclusion and Cretopiesma suukyiae syninclusion to the lower right of Sauronaradus meganae gen. et sp. nov. E head through thorax, ventral F pro- and meso- legs, mesotibia and tarsus visible, dorsal G right metatibia, dorsal." figureDoi="10.3897/zookeys.1219.137409.figure2" httpUri="https://binary.pensoft.net/fig/1184493">2 D</figureCitation>
). The
<typeStatus id="C5E5B3BA7F8037040243D4F8DA1A26D7" type="holotype">holotype</typeStatus>
is wholly intact and appears to have little warping. Due to the dark coloration the wings are difficult to discern as they are tucked closely to the abdomen surface. One syninclusion is present, a slightly damaged
<taxonomicName id="70BF573223E088916E0A3ED718B65FA0" authority="Grimaldi &amp; Engle, 2008" authorityName="Grimaldi &amp; Engle" authorityYear="2008" class="Insecta" family="Piesmatidae" genus="Cretopiesma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" phylum="Arthropoda" rank="species" species="suukyiae">
<emphasis id="0435EDEE6543178C64CB04073633B890" italics="true">Cretopiesma suukyiae</emphasis>
Grimaldi &amp; Engle, 2008
</taxonomicName>
.
</paragraph>
<caption id="9FB009E441732ACE1F9456D484AE76B8" ID-DOI="10.3897/zookeys.1219.137409.figure2" ID-arpha="DD023B40-C861-54EA-8707-B8ECA32E48D3" httpUri="https://binary.pensoft.net/fig/1184493" startId="F2">
<paragraph id="04211CE23C349293D948B44602425804">
<label id="282A4C95EDB1EC9765BAD5A5341CDD05">Figure 2.</label>
</paragraph>
<paragraph id="0A4C46E693091E4E0EAC183B069542B0">
<taxonomicName id="8592E8F23A1BEF2DF8CDDC056E40E615" authorityName="Cumming &amp; Mlynarek" authorityYear="2024" class="Insecta" family="Aradidae" genus="Sauronaradus" kingdom="Animalia" order="Hemiptera" phylum="Arthropoda" rank="species" species="meganae" status="gen. et sp. nov.">
<emphasis id="B64D4CE3C104C81CDC428E1F93EB3E59" italics="true">Sauronaradus meganae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="242FBAF914E10B4F6D5A6FF0A027CE5C" rank="species">gen. et sp. nov.</taxonomicNameLabel>
holotype
<emphasis id="AED39A4445E38DAC1BBAC8FC542AE62F" bold="true">A</emphasis>
anterior lit to highlight the spines of the head, dorsal
<emphasis id="B1FCA92BA182F98D71E481859811F49C" bold="true">B</emphasis>
head through thorax, dorsal
<emphasis id="809076F80BCD6B9A5A66C7D2F88E2D8A" bold="true">C</emphasis>
pronotum, lit to highlight carinae, dorsal
<emphasis id="6E3F430AA07A393A21D068B2069922D8" bold="true">D</emphasis>
full amber piece with holotype inclusion and
<taxonomicName id="0F1AAF1F93BA06CBE42586B6939C5F24" authorityName="Grimaldi &amp; Engle" authorityYear="2008" class="Insecta" family="Piesmatidae" genus="Cretopiesma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" phylum="Arthropoda" rank="species" species="suukyiae">
<emphasis id="200E8728F836CD6E7971EEBE45E8C7E1" italics="true">Cretopiesma suukyiae</emphasis>
</taxonomicName>
syninclusion to the lower right of
<taxonomicName id="7D03B6D2DC33B3ED36DC691DE826CB6D" authorityName="Cumming &amp; Mlynarek" authorityYear="2024" class="Insecta" family="Aradidae" genus="Sauronaradus" kingdom="Animalia" order="Hemiptera" phylum="Arthropoda" rank="species" species="meganae" status="gen. et sp. nov.">
<emphasis id="C080EFBAB80BC13EFADBA11BAD3AF7D9" italics="true">Sauronaradus meganae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="6870E1C64B24D94BCD564E8D3B73E604" rank="species">gen. et sp. nov.</taxonomicNameLabel>
<emphasis id="ED3C2418994C15C043A15B723934E8C8" bold="true">E</emphasis>
head through thorax, ventral
<emphasis id="F4A20B7DB9BA69476D2D8DAA6FEB4BC1" bold="true">F</emphasis>
pro- and meso- legs, mesotibia and tarsus visible, dorsal
<emphasis id="F1E4F47973ABE502675B533735518FE6" bold="true">G</emphasis>
right metatibia, dorsal.
</paragraph>
</caption>
</subSubSection>
<subSubSection id="SECID0EESAE" type="type locality">
<paragraph id="960D91907990317D6F2BDFDE44657E24">
<heading id="43549FEEDB210B16A5551E8AB4048B72" reason="title">Type locality and horizon.</heading>
</paragraph>
<paragraph id="CED1D31741D85DEB7CBFF4F870868BD7">
Latest Albian to lowermost Cenomanian (mid-Cretaceous), Hukawng Valley, northern
<collectingCountry id="F9D722038D0A1B3EFCF3B0B3FB71C6C6" name="Myanmar">Myanmar</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EJSAE" type="etymology">
<paragraph id="B6366702884BBA3B19E9F428263131B4">
<heading id="9B54EF7888BAFA4A082AD466428FE840" reason="title">Etymology.</heading>
</paragraph>
<paragraph id="E13090B2F042A473E21FA602703649ED">Patronym, named to honor Megan Solan, environmental toxicologist and entomologist. The first author wishes to thank Megan for her years of friendship and passion for entomology. Her enthusiasm for research is infectious and a positive driving force in the sciences.</paragraph>
</subSubSection>
<subSubSection id="SECID0EOSAE" type="diagnosis">
<paragraph id="3BF7E035AE1D4DDCCF44FDC0C66DF24F">
<heading id="055D38D0D1129CF1173BCBB226899C10" reason="title">Diagnosis.</heading>
</paragraph>
<paragraph id="C8DD701D5454FA646B3528B101D33893">Currently the genus is monotypic. See the above diagnosis of the genus for differentiation of this species from other aradids.</paragraph>
</subSubSection>
<subSubSection id="SECID0ETSAE" type="description">
<paragraph id="96100FB5B407674AB3661EC3346538A6">
<heading id="430F9536C0F231712F578EDF9F2103C0" reason="title">Description</heading>
</paragraph>
<paragraph id="B14787E265575E917D5E3ED2EC93B896">
(All measurements are in mm and based on the
<typeStatus id="4C21A44E67031E41C84EF1BC68D83548" type="holotype">holotype</typeStatus>
). Macropterous male; large, 5.05 long, 1.88 wide; body flattened (Figs
<figureCitation id="34C0E27475F6C671940F84F9D193D11C" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Sauronaradus meganae gen. et sp. nov. holotype A dorsal habitus B ventral habitus C abdomen, dorsal D abdomen, ventral E right antenna, dorsal." figureDoi="10.3897/zookeys.1219.137409.figure1" httpUri="https://binary.pensoft.net/fig/1184492">1</figureCitation>
,
<figureCitation id="F25E2D14BEADF1CB5C3D4471CCF2055F" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Sauronaradus meganae gen. et sp. nov. holotype A anterior lit to highlight the spines of the head, dorsal B head through thorax, dorsal C pronotum, lit to highlight carinae, dorsal D full amber piece with holotype inclusion and Cretopiesma suukyiae syninclusion to the lower right of Sauronaradus meganae gen. et sp. nov. E head through thorax, ventral F pro- and meso- legs, mesotibia and tarsus visible, dorsal G right metatibia, dorsal." figureDoi="10.3897/zookeys.1219.137409.figure2" httpUri="https://binary.pensoft.net/fig/1184493">2</figureCitation>
).
</paragraph>
<paragraph id="2EF4E80863BC24BA92FE3319F49C3398">
<emphasis id="8D7364883DCBD056BB70D64E948C2C88" bold="true" italics="true">Head</emphasis>
1.16 long, 0.89 wide (across the compound eyes); dorsally and ventrally rough textured. Compound eyes strongly protruding (Fig.
<figureCitation id="FCA49476F5D63CF8E3E51369B9BE73C1" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Sauronaradus meganae gen. et sp. nov. holotype A anterior lit to highlight the spines of the head, dorsal B head through thorax, dorsal C pronotum, lit to highlight carinae, dorsal D full amber piece with holotype inclusion and Cretopiesma suukyiae syninclusion to the lower right of Sauronaradus meganae gen. et sp. nov. E head through thorax, ventral F pro- and meso- legs, mesotibia and tarsus visible, dorsal G right metatibia, dorsal." figureDoi="10.3897/zookeys.1219.137409.figure2" httpUri="https://binary.pensoft.net/fig/1184493">2 A</figureCitation>
). Ocelli absent. Clypeus long, protruding, and boxy, with two clearly defined widths, broad for the posterior half (loral lobe), then half as wide on the apical half (maxillary lobe; Fig.
<figureCitation id="87A275C7C5A00A11FAF04FEC7440B71A" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Sauronaradus meganae gen. et sp. nov. holotype A anterior lit to highlight the spines of the head, dorsal B head through thorax, dorsal C pronotum, lit to highlight carinae, dorsal D full amber piece with holotype inclusion and Cretopiesma suukyiae syninclusion to the lower right of Sauronaradus meganae gen. et sp. nov. E head through thorax, ventral F pro- and meso- legs, mesotibia and tarsus visible, dorsal G right metatibia, dorsal." figureDoi="10.3897/zookeys.1219.137409.figure2" httpUri="https://binary.pensoft.net/fig/1184493">2 B</figureCitation>
). Dorsally there is a prominent anteclypeus on the posterior half (Fig.
<figureCitation id="CD85647A7BE9FE424C525B003B44FB8C" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Sauronaradus meganae gen. et sp. nov. holotype A anterior lit to highlight the spines of the head, dorsal B head through thorax, dorsal C pronotum, lit to highlight carinae, dorsal D full amber piece with holotype inclusion and Cretopiesma suukyiae syninclusion to the lower right of Sauronaradus meganae gen. et sp. nov. E head through thorax, ventral F pro- and meso- legs, mesotibia and tarsus visible, dorsal G right metatibia, dorsal." figureDoi="10.3897/zookeys.1219.137409.figure2" httpUri="https://binary.pensoft.net/fig/1184493">2 A</figureCitation>
). Clypeus loral lobe with two prominent apically pointing tubercles on each side; clypeus maxillary lobe with nodes on the dorsal and lateral surfaces (Fig.
<figureCitation id="655BBDD49700733A6413BDE1DE9184E3" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Sauronaradus meganae gen. et sp. nov. holotype A anterior lit to highlight the spines of the head, dorsal B head through thorax, dorsal C pronotum, lit to highlight carinae, dorsal D full amber piece with holotype inclusion and Cretopiesma suukyiae syninclusion to the lower right of Sauronaradus meganae gen. et sp. nov. E head through thorax, ventral F pro- and meso- legs, mesotibia and tarsus visible, dorsal G right metatibia, dorsal." figureDoi="10.3897/zookeys.1219.137409.figure2" httpUri="https://binary.pensoft.net/fig/1184493">2 B</figureCitation>
). Ventral surface of head with nodes similar to the dorsal surface nodes, and a narrow median depression of gula (Fig.
<figureCitation id="22464B802142CE3AA757746CCCE9BBF1" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Sauronaradus meganae gen. et sp. nov. holotype A anterior lit to highlight the spines of the head, dorsal B head through thorax, dorsal C pronotum, lit to highlight carinae, dorsal D full amber piece with holotype inclusion and Cretopiesma suukyiae syninclusion to the lower right of Sauronaradus meganae gen. et sp. nov. E head through thorax, ventral F pro- and meso- legs, mesotibia and tarsus visible, dorsal G right metatibia, dorsal." figureDoi="10.3897/zookeys.1219.137409.figure2" httpUri="https://binary.pensoft.net/fig/1184493">2 E</figureCitation>
). Bucculae not prominent. Open rostral atrium arising between the compound eyes, not at the apex of clypeus. Four, prominent, conical tubercles are present on the dorsal surface of the head. One pair are anteorbital tubercles present near the base of the compound eyes, and the second pair are situated evenly between the anteorbital tubercles (Fig.
<figureCitation id="F9CDD905DAE66878749B9F7069FB7036" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Sauronaradus meganae gen. et sp. nov. holotype A anterior lit to highlight the spines of the head, dorsal B head through thorax, dorsal C pronotum, lit to highlight carinae, dorsal D full amber piece with holotype inclusion and Cretopiesma suukyiae syninclusion to the lower right of Sauronaradus meganae gen. et sp. nov. E head through thorax, ventral F pro- and meso- legs, mesotibia and tarsus visible, dorsal G right metatibia, dorsal." figureDoi="10.3897/zookeys.1219.137409.figure2" httpUri="https://binary.pensoft.net/fig/1184493">2 A</figureCitation>
). These four tubercles are of similar size and shapes. Head lacking prominent postocular lobes, instead immediately behind the compound eyes is the notably constricted postocciput, which is slightly narrower than the anterior of the pronotum.
</paragraph>
<paragraph id="F0F1C315F6602F430E2A55DB5B992D71">
<emphasis id="4333BBBB58024FACDC80CDEF04A045B3" bold="true" italics="true">Antenna</emphasis>
exceptionally long and thin with antennomeres II, III, and IV with lengths c. 20–22 × their widths. Antenna with four antennomeres; basal antennomere (scape) the shortest, antennomere lengths: I: 0.27, II: 0.62, III: 0.70, IV: 0.74; antennomeres II, III, and IV tubiform; antennomeres II and III with surface granulation, and antennomere IV densely marked with setae throughout the surface with the seta length slightly less than the antennomere width, with the seta strongly angled apically (Fig.
<figureCitation id="7B6AE6C2DDD6BBB406A589A9EEA21CEE" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Sauronaradus meganae gen. et sp. nov. holotype A dorsal habitus B ventral habitus C abdomen, dorsal D abdomen, ventral E right antenna, dorsal." figureDoi="10.3897/zookeys.1219.137409.figure1" httpUri="https://binary.pensoft.net/fig/1184492">1 E</figureCitation>
). Antenniferous lobe prominent with a granular surface; stout, not projecting apically beyond the antennal insertion. Antennal insertion one-third of the way between the compound eyes and the apex of the clypeus (Fig.
<figureCitation id="893B78C1942A077757BD252BCC5C2D77" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Sauronaradus meganae gen. et sp. nov. holotype A anterior lit to highlight the spines of the head, dorsal B head through thorax, dorsal C pronotum, lit to highlight carinae, dorsal D full amber piece with holotype inclusion and Cretopiesma suukyiae syninclusion to the lower right of Sauronaradus meganae gen. et sp. nov. E head through thorax, ventral F pro- and meso- legs, mesotibia and tarsus visible, dorsal G right metatibia, dorsal." figureDoi="10.3897/zookeys.1219.137409.figure2" httpUri="https://binary.pensoft.net/fig/1184493">2 B</figureCitation>
). Rostrum with four segments extending to the posterior margin on the procoxal cavity (~
<quantity id="36E207A0B1E3F812E729B83BB922C338" metricMagnitude="-3" metricUnit="m" metricValue="1.73" unit="mm" value="1.73">1.73 mm</quantity>
long; Fig.
<figureCitation id="4C9815E3C1700F42C60BCDB6B13FF255" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Sauronaradus meganae gen. et sp. nov. holotype A anterior lit to highlight the spines of the head, dorsal B head through thorax, dorsal C pronotum, lit to highlight carinae, dorsal D full amber piece with holotype inclusion and Cretopiesma suukyiae syninclusion to the lower right of Sauronaradus meganae gen. et sp. nov. E head through thorax, ventral F pro- and meso- legs, mesotibia and tarsus visible, dorsal G right metatibia, dorsal." figureDoi="10.3897/zookeys.1219.137409.figure2" httpUri="https://binary.pensoft.net/fig/1184493">2 E</figureCitation>
).
</paragraph>
<paragraph id="DC43271DE61C0BA2F3EA71F24524AA73">
<emphasis id="C1780CD5556C65337B78B17A3E1EC19D" bold="true" italics="true">Thorax</emphasis>
pronotum (
<quantity id="A7731C5F640DCF0330557C7EABFFD87F" metricMagnitude="-4" metricUnit="m" metricValue="9.9" unit="mm" value="0.99">0.99 mm</quantity>
long, greatest width 1.44); roughly trapezoidal in shape (increasing in width caudally) but with somewhat undulating margins (approximate widths from the anterior to the posterior:
<quantity id="01D28A396B84484B43B254C6F3C0DE5B" metricMagnitude="-4" metricUnit="m" metricValue="4.2" unit="mm" value="0.42">0.42 mm</quantity>
,
<quantity id="E08238F43197A60ABEF7744B5D71000A" metricMagnitude="-4" metricUnit="m" metricValue="9.9" unit="mm" value="0.99">0.99 mm</quantity>
,
<quantity id="64330EE487F08851E4EF13F5D8E601DC" metricMagnitude="-3" metricUnit="m" metricValue="1.44" unit="mm" value="1.44">1.44 mm</quantity>
,
<quantity id="B1C8EDD2CAA9259711774EDF61DA2EF1" metricMagnitude="-3" metricUnit="m" metricValue="1.24" unit="mm" value="1.24">1.24 mm</quantity>
; Fig.
<figureCitation id="F017AD3E99C8F1492F3DEF3D40748FAC" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Sauronaradus meganae gen. et sp. nov. holotype A anterior lit to highlight the spines of the head, dorsal B head through thorax, dorsal C pronotum, lit to highlight carinae, dorsal D full amber piece with holotype inclusion and Cretopiesma suukyiae syninclusion to the lower right of Sauronaradus meganae gen. et sp. nov. E head through thorax, ventral F pro- and meso- legs, mesotibia and tarsus visible, dorsal G right metatibia, dorsal." figureDoi="10.3897/zookeys.1219.137409.figure2" httpUri="https://binary.pensoft.net/fig/1184493">2 C</figureCitation>
). Pronotum lateral margin armed on the anterior two-thirds with three prominent spiniform tubercles (anterior most
<quantity id="65C127C34FB05AF56D5940C32E1E7B1F" metricMagnitude="-4" metricUnit="m" metricValue="1.1" unit="mm" value="0.11">0.11 mm</quantity>
, middle
<quantity id="F6D1CB5257F12C4810DDF5050F1BF31F" metricMagnitude="-4" metricUnit="m" metricValue="1.0" unit="mm" value="0.1">0.10 mm</quantity>
, posterior-most
<quantity id="A2289FB0F6EEF431C50BBBE4CF2474AC" metricMagnitude="-4" metricUnit="m" metricValue="1.5" unit="mm" value="0.15">0.15 mm</quantity>
long; Fig.
<figureCitation id="10DDD676B2D53FCCD11BD2F69AD57A89" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Sauronaradus meganae gen. et sp. nov. holotype A anterior lit to highlight the spines of the head, dorsal B head through thorax, dorsal C pronotum, lit to highlight carinae, dorsal D full amber piece with holotype inclusion and Cretopiesma suukyiae syninclusion to the lower right of Sauronaradus meganae gen. et sp. nov. E head through thorax, ventral F pro- and meso- legs, mesotibia and tarsus visible, dorsal G right metatibia, dorsal." figureDoi="10.3897/zookeys.1219.137409.figure2" httpUri="https://binary.pensoft.net/fig/1184493">2 B</figureCitation>
). The posterior third of the pronotum bulges out to a width of
<quantity id="23D9063CCAB794BE8234B33552B11DC2" metricMagnitude="-3" metricUnit="m" metricValue="1.44" unit="mm" value="1.44">1.44 mm</quantity>
, and this bulge is armed with several smaller tubercles (Fig.
<figureCitation id="15B281BA13437A29F90855D1D227B448" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Sauronaradus meganae gen. et sp. nov. holotype A anterior lit to highlight the spines of the head, dorsal B head through thorax, dorsal C pronotum, lit to highlight carinae, dorsal D full amber piece with holotype inclusion and Cretopiesma suukyiae syninclusion to the lower right of Sauronaradus meganae gen. et sp. nov. E head through thorax, ventral F pro- and meso- legs, mesotibia and tarsus visible, dorsal G right metatibia, dorsal." figureDoi="10.3897/zookeys.1219.137409.figure2" httpUri="https://binary.pensoft.net/fig/1184493">2 B</figureCitation>
). Pronotum surface punctate throughout, with two paramedial carinae on each side of the sagittal plane, the central pair of carinae extend the full length of the pronotum and diverge in a similar fashion to the pronotum width, while the exterior carinae are less pronounced and are more prominent on the posterior half (Fig.
<figureCitation id="7E2FFA28C74D7C3B7644B1F7454AA950" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Sauronaradus meganae gen. et sp. nov. holotype A anterior lit to highlight the spines of the head, dorsal B head through thorax, dorsal C pronotum, lit to highlight carinae, dorsal D full amber piece with holotype inclusion and Cretopiesma suukyiae syninclusion to the lower right of Sauronaradus meganae gen. et sp. nov. E head through thorax, ventral F pro- and meso- legs, mesotibia and tarsus visible, dorsal G right metatibia, dorsal." figureDoi="10.3897/zookeys.1219.137409.figure2" httpUri="https://binary.pensoft.net/fig/1184493">2 C</figureCitation>
). Carinae jagged in form, not uniform in intensity throughout the lengths, somewhat rising and falling in rough textures (Fig.
<figureCitation id="8DF9A76DA6BB28D7A476F4886578D5C7" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Sauronaradus meganae gen. et sp. nov. holotype A anterior lit to highlight the spines of the head, dorsal B head through thorax, dorsal C pronotum, lit to highlight carinae, dorsal D full amber piece with holotype inclusion and Cretopiesma suukyiae syninclusion to the lower right of Sauronaradus meganae gen. et sp. nov. E head through thorax, ventral F pro- and meso- legs, mesotibia and tarsus visible, dorsal G right metatibia, dorsal." figureDoi="10.3897/zookeys.1219.137409.figure2" httpUri="https://binary.pensoft.net/fig/1184493">2 C</figureCitation>
). Scutellum long and prominent,
<quantity id="5441ED6560616F8F3FF2063AB022F869" metricMagnitude="-3" metricUnit="m" metricValue="1.24" unit="mm" value="1.24">1.24 mm</quantity>
wide,
<quantity id="710A76DDA1C3E7B8FE3746812A4902D0" metricMagnitude="-3" metricUnit="m" metricValue="1.16" unit="mm" value="1.16">1.16 mm</quantity>
long; dorsal surface with the anterior third slightly raised above the posterior two-thirds; surface flat but punctate throughout; apex broadly rounded. Prosternum;
<quantity id="644CA3A951F16A88B22E9E9C2743FFA1" metricMagnitude="-4" metricUnit="m" metricValue="4.9" unit="mm" value="0.49">0.49 mm</quantity>
long, surface punctate and marked with a moderate rostral groove which continues onto the mesosternum as a broader depression along the sagittal plane (Fig.
<figureCitation id="BE87E718097BB59F5FF5B68D20145997" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Sauronaradus meganae gen. et sp. nov. holotype A anterior lit to highlight the spines of the head, dorsal B head through thorax, dorsal C pronotum, lit to highlight carinae, dorsal D full amber piece with holotype inclusion and Cretopiesma suukyiae syninclusion to the lower right of Sauronaradus meganae gen. et sp. nov. E head through thorax, ventral F pro- and meso- legs, mesotibia and tarsus visible, dorsal G right metatibia, dorsal." figureDoi="10.3897/zookeys.1219.137409.figure2" httpUri="https://binary.pensoft.net/fig/1184493">2 E</figureCitation>
). Mesosternum maximum width
<quantity id="39B60DD89B0C158AC9789ED84E26386F" metricMagnitude="-3" metricUnit="m" metricValue="1.44" unit="mm" value="1.44">1.44 mm</quantity>
, length
<quantity id="F4E02F1B35691ECC1402387ED31F6CD1" metricMagnitude="-4" metricUnit="m" metricValue="5.1" unit="mm" value="0.51">0.51 mm</quantity>
; ventral surface punctate (Fig.
<figureCitation id="D82888365BC54BA505CD2AA350D68301" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Sauronaradus meganae gen. et sp. nov. holotype A anterior lit to highlight the spines of the head, dorsal B head through thorax, dorsal C pronotum, lit to highlight carinae, dorsal D full amber piece with holotype inclusion and Cretopiesma suukyiae syninclusion to the lower right of Sauronaradus meganae gen. et sp. nov. E head through thorax, ventral F pro- and meso- legs, mesotibia and tarsus visible, dorsal G right metatibia, dorsal." figureDoi="10.3897/zookeys.1219.137409.figure2" httpUri="https://binary.pensoft.net/fig/1184493">2 E</figureCitation>
). Metasternum
<quantity id="17D155C901E082D9B8B004B50B37B65D" metricMagnitude="-4" metricUnit="m" metricValue="5.1" unit="mm" value="0.51">0.51 mm</quantity>
long,
<quantity id="080A726D89DBC2D064DF5738DF90FB85" metricMagnitude="-3" metricUnit="m" metricValue="1.59" unit="mm" value="1.59">1.59 mm</quantity>
wide.
</paragraph>
<paragraph id="4428C6CC1899770C77F7D9382AC11C0B">
<emphasis id="0D14AA904E633639746C8FDEB3EDD2C9" bold="true" italics="true">Legs</emphasis>
long and thin, all with slightly granular surfaces. Profemora
<quantity id="C90DF58C077398E1B324D4961B4BEB19" metricMagnitude="-4" metricUnit="m" metricValue="9.0" unit="mm" value="0.9">0.90 mm</quantity>
long, thinner on the proximal third. Leg lengths: protibiae
<quantity id="8BBF83AEAA7169CC64DBBCC112DA1CED" metricMagnitude="-4" metricUnit="m" metricValue="7.5" unit="mm" value="0.75">0.75 mm</quantity>
, mesofemora
<quantity id="23D335030E76706F94C6CB35C40BEE00" metricMagnitude="-4" metricUnit="m" metricValue="9.2" unit="mm" value="0.92">0.92 mm</quantity>
, mesotibiae
<quantity id="B29AD35EB65917A1B8A7F1754B63DEFD" metricMagnitude="-4" metricUnit="m" metricValue="6.0" unit="mm" value="0.6">0.60 mm</quantity>
, metafemora
<quantity id="7C6DA7B70D041609C25E8787932DD72A" metricMagnitude="-3" metricUnit="m" metricValue="1.16" unit="mm" value="1.16">1.16 mm</quantity>
, metatibiae
<quantity id="F79A9279DD6274E8E7263E22B5CCF75C" metricMagnitude="-4" metricUnit="m" metricValue="9.3" unit="mm" value="0.93">0.93 mm</quantity>
. Tibiae with sparse setae throughout, with longer and thicker setae on the distal ends (Fig.
<figureCitation id="55FDCD1900FF697FD4176358402BB2A2" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Sauronaradus meganae gen. et sp. nov. holotype A anterior lit to highlight the spines of the head, dorsal B head through thorax, dorsal C pronotum, lit to highlight carinae, dorsal D full amber piece with holotype inclusion and Cretopiesma suukyiae syninclusion to the lower right of Sauronaradus meganae gen. et sp. nov. E head through thorax, ventral F pro- and meso- legs, mesotibia and tarsus visible, dorsal G right metatibia, dorsal." figureDoi="10.3897/zookeys.1219.137409.figure2" httpUri="https://binary.pensoft.net/fig/1184493">2 F</figureCitation>
). Tarsi with two tarsomeres; apical tarsomere is 1.5–2.0 times longer than the previous tarsomere; apical tarsomere thin proximally, widening notably for the proximal half, then a uniform broad width on the distal half (Fig.
<figureCitation id="883A4E105AE35E8B011FA34951E1049D" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Sauronaradus meganae gen. et sp. nov. holotype A anterior lit to highlight the spines of the head, dorsal B head through thorax, dorsal C pronotum, lit to highlight carinae, dorsal D full amber piece with holotype inclusion and Cretopiesma suukyiae syninclusion to the lower right of Sauronaradus meganae gen. et sp. nov. E head through thorax, ventral F pro- and meso- legs, mesotibia and tarsus visible, dorsal G right metatibia, dorsal." figureDoi="10.3897/zookeys.1219.137409.figure2" httpUri="https://binary.pensoft.net/fig/1184493">2 F, G</figureCitation>
). Apical tarsomere ventrally with few thin setae, previous tarsomere with thicker but sparse setae on the distal end (Fig.
<figureCitation id="7C21838EB80B40A41792717C3342E1E1" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Sauronaradus meganae gen. et sp. nov. holotype A anterior lit to highlight the spines of the head, dorsal B head through thorax, dorsal C pronotum, lit to highlight carinae, dorsal D full amber piece with holotype inclusion and Cretopiesma suukyiae syninclusion to the lower right of Sauronaradus meganae gen. et sp. nov. E head through thorax, ventral F pro- and meso- legs, mesotibia and tarsus visible, dorsal G right metatibia, dorsal." figureDoi="10.3897/zookeys.1219.137409.figure2" httpUri="https://binary.pensoft.net/fig/1184493">2 F</figureCitation>
). Tarsal claws long and simple; pulvilli approximately lemniscate, slightly shorter than tarsal claws (Fig.
<figureCitation id="1998C0048C43AC42657E13861F9F0FD3" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Sauronaradus meganae gen. et sp. nov. holotype A anterior lit to highlight the spines of the head, dorsal B head through thorax, dorsal C pronotum, lit to highlight carinae, dorsal D full amber piece with holotype inclusion and Cretopiesma suukyiae syninclusion to the lower right of Sauronaradus meganae gen. et sp. nov. E head through thorax, ventral F pro- and meso- legs, mesotibia and tarsus visible, dorsal G right metatibia, dorsal." figureDoi="10.3897/zookeys.1219.137409.figure2" httpUri="https://binary.pensoft.net/fig/1184493">2 F, G</figureCitation>
).
</paragraph>
<paragraph id="EC5CA52F4F619713EEC2F6A58A2C3492">
<emphasis id="73B11E9D3FCFB1A48242CA1D4E9335B7" bold="true" italics="true">Wings</emphasis>
fully developed, but details are indiscernible due to taphonomy of the
<typeStatus id="35F89C8D14A405E99916EA523772F263" type="holotype">holotype</typeStatus>
.
</paragraph>
<paragraph id="5DB27823959F3E5806610B075DF462E1">
<emphasis id="1F4E1D6062CE12047A41F6DF5CCF4195" bold="true" italics="true">Abdomen</emphasis>
broad and flat, all surfaces punctate; length 2.41, greatest width
<quantity id="15538E919565EA1B617B20D29F4E5C94" metricMagnitude="-3" metricUnit="m" metricValue="1.88" unit="mm" value="1.88">1.88 mm</quantity>
. Abdominal segment lengths: III =
<quantity id="69B0732A93ACB70CF47FF98B36C1873A" metricMagnitude="-4" metricUnit="m" metricValue="5.699999999999999" unit="mm" value="0.57">0.57 mm</quantity>
, IV =
<quantity id="E953A8E2F5B654BE754494A460AC0CD0" metricMagnitude="-4" metricUnit="m" metricValue="3.4" unit="mm" value="0.34">0.34 mm</quantity>
, V =
<quantity id="8E629437852983A9FDFB022582114561" metricMagnitude="-4" metricUnit="m" metricValue="3.2" unit="mm" value="0.32">0.32 mm</quantity>
, VI =
<quantity id="7DF31BCB9D79B4C63FF4A348B652D99D" metricMagnitude="-4" metricUnit="m" metricValue="3.7" unit="mm" value="0.37">0.37 mm</quantity>
, VII =
<quantity id="04B7155585C4F2285772879BB42F58CD" metricMagnitude="-4" metricUnit="m" metricValue="4.3" unit="mm" value="0.43">0.43 mm</quantity>
, VIII =
<quantity id="8A2F5501BE64E74EB2A0FC10290674BE" metricMagnitude="-4" metricUnit="m" metricValue="4.0" unit="mm" value="0.4">0.40 mm</quantity>
. Each abdominal segment has margins which gently undulate, with four or five humps, the posterior-most of which is on the posterior margin and larger than the others (Fig.
<figureCitation id="D00D73A45899F0DD0583542752DED087" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Sauronaradus meganae gen. et sp. nov. holotype A dorsal habitus B ventral habitus C abdomen, dorsal D abdomen, ventral E right antenna, dorsal." figureDoi="10.3897/zookeys.1219.137409.figure1" httpUri="https://binary.pensoft.net/fig/1184492">1 C, D</figureCitation>
). Male terminalia broad and roughly bell shaped, surfaces rugose, a distinctly raised median ridge protrudes on the dorsal surface (Fig.
<figureCitation id="34DD4B8886CB9516D5F4B070BB899A24" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Sauronaradus meganae gen. et sp. nov. holotype A dorsal habitus B ventral habitus C abdomen, dorsal D abdomen, ventral E right antenna, dorsal." figureDoi="10.3897/zookeys.1219.137409.figure1" httpUri="https://binary.pensoft.net/fig/1184492">1 C, D</figureCitation>
). Paratergite VIII rhomboid, angulated posteriorly, with the lateral margin slightly undulating (Fig.
<figureCitation id="2FCF060E1BEE558C3E6BEA2EEB8560EA" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Sauronaradus meganae gen. et sp. nov. holotype A dorsal habitus B ventral habitus C abdomen, dorsal D abdomen, ventral E right antenna, dorsal." figureDoi="10.3897/zookeys.1219.137409.figure1" httpUri="https://binary.pensoft.net/fig/1184492">1 C, D</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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