183 lines
22 KiB
XML
183 lines
22 KiB
XML
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<mods:title id="BFA69168D5944AD21952A85ECE2782B7">Into the Chamber of Horrors: A proposal for the resolution of nomenclatural chaos in the Scaphiophryne calcarata complex (Anura: Microhylidae), with a new species-level phylogenetic hypothesis for Scaphiophryninae</mods:title>
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<mods:namePart id="BE62FEF5739787901662986B47990743">Scherz, Mark D.</mods:namePart>
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<mods:affiliation id="BC4FBA0DE1ABD67E68815FB214F86FC9">Department of Evolutionary Biology, University of Konstanz, Universitätsstrasse 10, 78464 Konstanz, Germany mark. scherz @ gmail. com; https: // orcid. org / 0000 - 0002 - 4613 - 7761</mods:affiliation>
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<mods:namePart id="B813CF4FD53EDC907097B5E9C2B772EB">Schmidt, Luca</mods:namePart>
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<mods:affiliation id="E2A2640AC3ECD67DDC2E093213FDFBA3">Braunschweig University of Technology, Zoological Institute, Mendelssohnstrasse 4, 38106 Braunschweig, Germany m. vences @ tu-braunschweig. de; https: // orcid. org / 0000 - 0003 - 0747 - 0817</mods:affiliation>
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<mods:namePart id="0A76C502BCF825692E298D3DE561D8C2">Crottini, Angelica</mods:namePart>
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<mods:namePart id="1C5CECFCF78B4DD19B85FADCD5A6DCF8">Miralles, Aurélien</mods:namePart>
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<mods:affiliation id="6BAF14CB60773F1CB80D863F9CA5C56A">Institut de Systématique, Évolution, Biodiversité (ISYEB), Muséum national d'Histoire naturelle, CNRS, Sorbonne Université, EPHE, Université des Antilles, 57 rue Cuvier, CP 50, 75005 Paris, France</mods:affiliation>
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<mods:namePart id="A429A81E3489CC46F965484AD85ADE0D">Rakotoarison, Andolalao</mods:namePart>
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<mods:affiliation id="4083EBAAF622CA0EA0AFC6A06EB7621F">Zoologie et Biodiversité Animale, Université d’Antananarivo, BP 906, Antananarivo, 101 Madagascar</mods:affiliation>
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<mods:namePart id="8FFC84FC305B65A33426A2BD295930F5">Raselimanana, Achille P.</mods:namePart>
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<mods:affiliation id="B95924065FCC3038F70EC248073322AF">Zoologie et Biodiversité Animale, Université d’Antananarivo, BP 906, Antananarivo, 101 Madagascar & Association Vahatra, lot VA 38 LB Ter A, Ambohidempona- Tsiadana, Antananarivo 101, Madagascar raselimananaachille @ gmail. com; https: // orcid. org / 0000 - 0003 - 1610 - 7307</mods:affiliation>
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<mods:namePart id="6981C6A369F63F351D63FDDF07C174C9">Köhler, Jörn</mods:namePart>
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<mods:affiliation id="01CD41DBE4FA4716D5EB8BFE84D23DA4">Hessisches Landesmuseum Darmstadt, Friedensplatz 1, 64283 Darmstadt, Germany</mods:affiliation>
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<mods:namePart id="A6236B4B1DF17EAE68C91A0970325A48">Glaw, Frank</mods:namePart>
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<mods:affiliation id="085ADEB978CF6A9A3F4A83B45A0D7365">Zoologische Staatssammlung München (ZSM-SNSB), Münchhausenstrasse 21, 81247 München, Germany</mods:affiliation>
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<mods:namePart id="D052BA323816E66B9BDA9114E31E69FD">Vences, Miguel</mods:namePart>
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<mods:affiliation id="F6A656AA082564E7A5228AF2D6100800">Braunschweig University of Technology, Zoological Institute, Mendelssohnstrasse 4, 38106 Braunschweig, Germany m. vences @ tu-braunschweig. de; https: // orcid. org / 0000 - 0003 - 0747 - 0817</mods:affiliation>
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<mods:date id="56928A86A0F1E7E487D9B454836DEFDD">2021</mods:date>
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<treatment id="1C7C8218FF88FFEBFF017681FCF1FD28" ID-DOI="http://doi.org/10.5281/zenodo.4605112" ID-GBIF-Taxon="180568741" ID-Zenodo-Dep="4605112" LSID="urn:lsid:plazi:treatment:1C7C8218FF88FFEBFF017681FCF1FD28" httpUri="http://treatment.plazi.org/id/1C7C8218FF88FFEBFF017681FCF1FD28" lastPageNumber="409" pageId="17" pageNumber="409">
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<paragraph id="946A330EFF88FFEBFF017681FDB8FF33" blockId="17.[151,594,151,178]" box="[151,594,151,178]" pageId="17" pageNumber="409">
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<taxonomicName id="53D5488DFF88FFEBFF017681FDB8FF33" ID-CoL="68VGT" authority="Boulenger, 1896" authorityName="Boulenger" authorityYear="1896" box="[151,594,151,178]" class="Amphibia" family="Microhylidae" genus="Calophrynus" higherTaxonomySource="CoL" kingdom="Animalia" order="Anura" pageId="17" pageNumber="409" phylum="Chordata" rank="species" species="brevis">
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<emphasis id="A6A1EF1CFF88FFEBFF017681FDB8FF33" bold="true" box="[151,594,151,178]" pageId="17" pageNumber="409">
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<emphasis id="A6A1EF1CFF88FFEBFF017681FE69FF30" bold="true" box="[151,387,151,178]" italics="true" pageId="17" pageNumber="409">Calophrynus brevis</emphasis>
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<bibRefCitation id="F0444EFFFF88FFEBFE1C7681FDB8FF33" author="Boulenger, G. A." box="[394,594,151,177]" pageId="17" pageNumber="409" pagination="401 - 406" refId="ref15999" refString="Boulenger, G. A. (1896) Descriptions of new batrachians in the British Museum. Annals and Magazine of Natural History, Series 6, 17, 401 - 406. https: // doi. org / 10.1080 / 00222939608680389" type="journal article" year="1896">Boulenger, 1896</bibRefCitation>
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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<paragraph id="946A330EFF88FFEBFF0176C9FA73FF7B" blockId="17.[151,1437,223,682]" box="[151,1433,223,250]" pageId="17" pageNumber="409">
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<materialsCitation id="24BD3953FF88FFEBFF0176C9FA73FF7B" ID-GBIF-Occurrence="3053023305" box="[151,1433,223,250]" collectionCode="BMNH" country="Madagascar" location="Madagascar" pageId="17" pageNumber="409" specimenCode="BMNH 1947.2" specimenCount="1" typeStatus="holotype">
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Name-bearing type:
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<typeStatus id="4B6E8DACFF88FFEBFEEB76C9FE0FFF7B" box="[381,485,223,249]" pageId="17" pageNumber="409" type="holotype">Holotype</typeStatus>
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<specimenCode id="C4739B75FF88FFEBFE7B76F6FD72FF7B" box="[493,664,223,250]" collectionCode="BMNH" pageId="17" pageNumber="409">BMNH 1947.2</specimenCode>
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.7.42, from “S.W.
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<collectingCountry id="ECC2739EFF88FFEBFCF776C9FC01FF7B" box="[865,1003,223,249]" name="Madagascar" pageId="17" pageNumber="409">Madagascar</collectingCountry>
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” according to the original description.
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</paragraph>
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<subSubSection id="DCCF6085FF88FFEBFF517715FCF1FD28" pageId="17" pageNumber="409" type="description">
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<paragraph id="946A330EFF88FFEBFF517715FCF1FD28" blockId="17.[151,1437,223,682]" pageId="17" pageNumber="409">
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The
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<typeStatus id="4B6E8DACFF88FFEBFF6C7715FEB6FE9F" box="[250,348,259,285]" pageId="17" pageNumber="409" type="holotype">holotype</typeStatus>
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of this nomen is in good state of preservation, allowing the examination of most or all characters relevant for taxonomic assignment. Although it is smaller than most other adult specimens typically assigned to
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<taxonomicName id="53D5488DFF88FFEBFF01775DFF12FEE7" authorityName="Boulenger" authorityYear="1896" box="[151,248,331,357]" class="Amphibia" family="Microhylidae" genus="Calophrynus" higherTaxonomySource="CoL" kingdom="Animalia" order="Anura" pageId="17" pageNumber="409" phylum="Chordata" rank="species" species="brevis">
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<emphasis id="A6A1EF1CFF88FFEBFF01775DFF12FEE7" box="[151,248,331,357]" italics="true" pageId="17" pageNumber="409">S. brevis</emphasis>
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</taxonomicName>
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and therefore may be a subadult, its size is similar to one small-sized adult male collected by us (ZSM 577/2000;
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<tableCitation id="D95706B5FF88FFEBFE867779FE89FE0B" box="[272,355,367,393]" captionStart="TABLE 1" captionStartId="20.[151,239,461,487]" captionText="TABLE 1. Morphometric measurements (all in mm) of name bearing types and other specimens of Scaphiophryne brevis, S. calcarata and S. obscura. For abbreviations of morphometric measurements and collection acronyms, see Materials and Methods. RHL (relative hindlimb length) indicates the point reached by the tibiotarsal articulation when hindlimb is adpressed along body: 1, forelimb; 2, between forelimb and tympanum; 3, tympanum; 4, eye. Additional abbreviations: nm, not measured; na, not applicable; M, male; F, female; J, juvenile; LT, lectotype; PLT, paralectotype; ST, syntype; HT, holotype. Nomina are abbreviated as follows: Cb, Calophrynus brevis; Cc, Calophrynus calcaratus; Ho, Hemisus obscurus; Pg, Pseudohemisus granulosus; Pl, Pseudohemisus longimanus; Plm; Pseudohemisus longimanus var. melanopleura; Pv, Pseudohemisus verrucosus." pageId="17" pageNumber="409">Table 1</tableCitation>
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). The general body shape and absence of dorsolateral folds or a sharp dorsolateral color border (i.e., a clearly delimited boundary between the colors of the dorsum and flanks) differentiate the specimen from those of the
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<taxonomicName id="53D5488DFF88FFEBFEB677A1FE4DFE53" baseAuthorityName="Mocquard" baseAuthorityYear="1895" box="[288,423,439,465]" class="Amphibia" family="Microhylidae" genus="Scaphiophryne" kingdom="Animalia" order="Anura" pageId="17" pageNumber="409" phylum="Chordata" rank="species" species="calcarata">
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<emphasis id="A6A1EF1CFF88FFEBFEB677A1FE4DFE53" box="[288,423,439,465]" italics="true" pageId="17" pageNumber="409">S. calcarata</emphasis>
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</taxonomicName>
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complex, and confirm that
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<taxonomicName id="53D5488DFF88FFEBFD4D77A1FB4AFE50" authority="(Boulenger, 1896)" baseAuthorityName="Boulenger" baseAuthorityYear="1896" box="[731,1184,439,466]" class="Amphibia" family="Microhylidae" genus="Scaphiophryne" kingdom="Animalia" order="Anura" pageId="17" pageNumber="409" phylum="Chordata" rank="species" species="brevis">
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<emphasis id="A6A1EF1CFF88FFEBFD4D77A1FC27FE53" box="[731,973,439,465]" italics="true" pageId="17" pageNumber="409">Scaphiophryne brevis</emphasis>
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(
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<bibRefCitation id="F0444EFFFF88FFEBFC4A77A1FB72FE50" author="Boulenger, G. A." box="[988,1176,439,466]" pageId="17" pageNumber="409" pagination="401 - 406" refId="ref15999" refString="Boulenger, G. A. (1896) Descriptions of new batrachians in the British Museum. Annals and Magazine of Natural History, Series 6, 17, 401 - 406. https: // doi. org / 10.1080 / 00222939608680389" type="journal article" year="1896">Boulenger, 1896</bibRefCitation>
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)
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</taxonomicName>
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||
has been correctly applied by previous researchers (e.g., Blommers-Schl̂sser & Blanc 1991;
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<bibRefCitation id="F0444EFFFF88FFEBFC5577CDFB5EFE77" author="Glaw, F. & Vences, M." box="[963,1204,475,502]" pageId="17" pageNumber="409" refId="ref16536" refString="Glaw, F. & Vences, M. (2007) A Field Guide to the Amphibians and Reptiles of Madagascar. Third Edition. K ˆ ln, Vences & Glaw, 496 pp." type="book" year="2007">Glaw & Vences 2007</bibRefCitation>
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;
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||
<bibRefCitation id="F0444EFFFF88FFEBFB5677CDFA7BFE77" author="Vieites, D. R. & Wollenberg, K. C. & Andreone, F. & Glaw, F. & Vences, M." box="[1216,1425,475,502]" pageId="17" pageNumber="409" pagination="8267 - 8272" refId="ref19847" refString="Vieites, D. R., Wollenberg, K. C., Andreone, F., K ˆ hler, J., Glaw, F. & Vences, M. (2009) Vast underestimation of Madagascar's biodiversity evidenced by an integrative amphibian inventory. Proceedings of the National Academy of Sciences of the United States of America, 106, 8267 - 8272. https: // doi. org / 10.1073 / pnas. 0810821106" type="journal article" year="2009">
|
||
Vieites
|
||
<emphasis id="A6A1EF1CFF88FFEBFA8377CAFAACFE77" box="[1301,1350,475,501]" italics="true" pageId="17" pageNumber="409">et al</emphasis>
|
||
. 2009
|
||
</bibRefCitation>
|
||
).
|
||
<bibRefCitation id="F0444EFFFF88FFEBFF0177E9FE6DFD9B" author="D'Cruze, N. & Olsonn, A. & Henson, D. & Kumar, S. & Emmett, D." box="[151,391,511,538]" pageId="17" pageNumber="409" pagination="62 - 79" refId="ref16300" refString="D'Cruze, N., Olsonn, A., Henson, D., Kumar, S. & Emmett, D. (2009) Amphibians and reptiles of the Onilahy River valley. Herpetological Conservation and Biology, 4, 62 - 79." type="journal article" year="2009">
|
||
D’Cruze
|
||
<emphasis id="A6A1EF1CFF88FFEBFF687416FEC5FD9B" box="[254,303,511,537]" italics="true" pageId="17" pageNumber="409">et al</emphasis>
|
||
. (2009)
|
||
</bibRefCitation>
|
||
reported this species from the Onilahy River valley.
|
||
<taxonomicName id="53D5488DFF88FFEBFC4277E9FB2FFD9B" baseAuthorityName="Boulenger" baseAuthorityYear="1896" box="[980,1221,511,537]" class="Amphibia" family="Microhylidae" genus="Scaphiophryne" kingdom="Animalia" order="Anura" pageId="17" pageNumber="409" phylum="Chordata" rank="species" species="brevis">
|
||
<emphasis id="A6A1EF1CFF88FFEBFC4277E9FB2FFD9B" box="[980,1221,511,537]" italics="true" pageId="17" pageNumber="409">Scaphiophryne brevis</emphasis>
|
||
</taxonomicName>
|
||
seems to exhibit at least two pronounced color morphs, one with a diffuse dorsolateral color border and typically a light midvertebral stripe (first row of
|
||
<figureCitation id="0CEE2F8BFF88FFEBFEFA7451FE44FDE3" box="[364,430,583,609]" captionStart="FIGURE 3" captionStartId="8.[151,250,1854,1878]" captionTargetBox="[174,1405,181,1831]" captionTargetId="figure-17@8.[174,1413,181,1831]" captionTargetPageId="8" captionText="FIGURE 3. Individuals in life of the three species of Scaphiophryne in the subgenus Pseudohemisus according to the classification proposed in the present paper: Scaphiophryne brevis from Ifaty (A–E), Kirindy (F–G) and Isalo (H); S. calcarata from Pic St. Louis near Tolagnaro (I–K), Ambalavao (L), Isalo (Ilakaka) (M–N), Sakaviro (O) and near Anja (P); S. obscura from Isalo (Q), Kirindy (R–V), Ankarafantsika (W), and Bongolava Plateau (X). Photos by F. Glaw, J. Lobon-Rovira, A. Miralles, G. M. Rosa, and M. Vences." figureDoi="http://doi.org/10.5281/zenodo.4574877" httpUri="https://zenodo.org/record/4574877/files/figure.png" pageId="17" pageNumber="409">Fig. 3</figureCitation>
|
||
), and one that is more marbled (second row of
|
||
<figureCitation id="0CEE2F8BFF88FFEBFC557451FBECFDE3" box="[963,1030,583,609]" captionStart="FIGURE 3" captionStartId="8.[151,250,1854,1878]" captionTargetBox="[174,1405,181,1831]" captionTargetId="figure-17@8.[174,1413,181,1831]" captionTargetPageId="8" captionText="FIGURE 3. Individuals in life of the three species of Scaphiophryne in the subgenus Pseudohemisus according to the classification proposed in the present paper: Scaphiophryne brevis from Ifaty (A–E), Kirindy (F–G) and Isalo (H); S. calcarata from Pic St. Louis near Tolagnaro (I–K), Ambalavao (L), Isalo (Ilakaka) (M–N), Sakaviro (O) and near Anja (P); S. obscura from Isalo (Q), Kirindy (R–V), Ankarafantsika (W), and Bongolava Plateau (X). Photos by F. Glaw, J. Lobon-Rovira, A. Miralles, G. M. Rosa, and M. Vences." figureDoi="http://doi.org/10.5281/zenodo.4574877" httpUri="https://zenodo.org/record/4574877/files/figure.png" pageId="17" pageNumber="409">Fig. 3</figureCitation>
|
||
). A potential third color morph with green elements is figured in
|
||
<bibRefCitation id="F0444EFFFF88FFEBFE4E747DFD23FD07" author="Glaw, F. & Vences, M." box="[472,713,619,646]" pageId="17" pageNumber="409" refId="ref16536" refString="Glaw, F. & Vences, M. (2007) A Field Guide to the Amphibians and Reptiles of Madagascar. Third Edition. K ˆ ln, Vences & Glaw, 496 pp." type="book" year="2007">Glaw & Vences 2007</bibRefCitation>
|
||
(page 448), and discussed in the context of
|
||
<taxonomicName id="53D5488DFF88FFEBFB2D747DFF04FD2B" authorityName="Angel" authorityYear="1930" class="Amphibia" family="Microhylidae" genus="Pseudohemisus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="17" pageNumber="409" phylum="Chordata" rank="species" species="verrucosus">
|
||
<emphasis id="A6A1EF1CFF88FFEBFB2D747DFF04FD2B" italics="true" pageId="17" pageNumber="409">Pseudohemisus verrucosus</emphasis>
|
||
</taxonomicName>
|
||
below. These color morphs are not related to sex.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |