450 lines
68 KiB
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450 lines
68 KiB
XML
<document id="D46331B05A597E6CCAC95252B0565729" ID-CLB-Dataset="110556" ID-DOI="10.5852/ejt.2022.832.1877" ID-GBIF-Dataset="a5effd4e-8a15-48fe-9e66-d8f46ff4694e" ID-ISSN="2118-9773" ID-Zenodo-Dep="6916850" ID-ZooBank="6AD25640-2B25-458D-942D-22ED24D177BD" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1658962493167" checkinUser="felipe" docAuthor="Steinpress, Igor Armiach, Cohen, Mira, Pétillon, Julien, Chipman, Ariel D. & Gavish-Regev, Efrat" docDate="2022" docId="1C43EA6E9B56FFF3FD255977FDAA5764" docLanguage="en" docName="ejt-832_1-54.pdf.imf" docOrigin="European Journal of Taxonomy 832 (1)" docSource="http://dx.doi.org/10.5852/ejt.2022.832.1877" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Lycosa gesserit Steinpress & Cohen & Pétillon & Chipman & Gavish-Regev 2022, sp. nov." docType="treatment" docUuid="A5A1F6AD-14B4-488F-BC74-A2B5E240A86B" docUuidSource="ZooBank" docVersion="7" lastPageNumber="17" masterDocId="E07A92169B5BFFE2FFAB5F34FF9D5622" masterDocTitle="Lycosa Latreille, 1804 (Araneae, Lycosidae) of Israel, with a note on Geolycosa Montgomery, 1904" masterLastPageNumber="54" masterPageNumber="1" pageNumber="14" updateTime="1699362406483" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="AFF18FD05A5F1C3EB5BA4C5377F76BF8">Lycosa Latreille, 1804 (Araneae, Lycosidae) of Israel, with a note on Geolycosa Montgomery, 1904</mods:title>
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<mods:namePart id="2E62948DE6A344CA651C006B96E479FC">Steinpress, Igor Armiach</mods:namePart>
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<mods:affiliation id="B3D0E9F6D0A011AD8A2BA9F5D287E068">The Arachnid National Natural History Collection, The Hebrew University of Jerusalem, Edmond J. Safra Campus, Giv’at Ram, Jerusalem, Israel. The Department of Ecology, Evolution and Behavior, The Hebrew University of Jerusalem, Edmond J. Safra Campus, Giv’at Ram, Jerusalem, Israel. University of Rennes, Rennes, France. Nelson Mandela University, Port Elizabeth, South Africa.</mods:affiliation>
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<mods:nameIdentifier id="4C64C4C0C45E2CEEC75158CBE3FCABC2" type="email">bomtombadil@gmail.com</mods:nameIdentifier>
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<mods:namePart id="901BE7A23323E4C3C5998E957D1FA808">Cohen, Mira</mods:namePart>
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<mods:nameIdentifier id="DF82C9AD6B5ADD91B18DCB3B9AB9CEB9" type="email">mira.cohen@mail.huji.ac.il</mods:nameIdentifier>
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<mods:namePart id="8AE47A8373272675718AFC75D21897C6">Pétillon, Julien</mods:namePart>
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<mods:nameIdentifier id="566BAC613FE08E83EF543252F1509EB3" type="email">julien.petillon@univ-rennes1.fr</mods:nameIdentifier>
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<mods:namePart id="AF0E639F77ED7D21F4CD59A54227BA52">Chipman, Ariel D.</mods:namePart>
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<mods:nameIdentifier id="BBB1714746B440B2D97A7CB367FB9F8D" type="email">ariel.chipman@huji.ac.il</mods:nameIdentifier>
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<mods:namePart id="6409A60E9AA273668366E45896EE1097">Gavish-Regev, Efrat</mods:namePart>
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<mods:affiliation id="6260A287CD64278128F391B89A1FDAEE">The Arachnid National Natural History Collection, The Hebrew University of Jerusalem, Edmond J. Safra Campus, Giv’at Ram, Jerusalem, Israel. The Department of Ecology, Evolution and Behavior, The Hebrew University of Jerusalem, Edmond J. Safra Campus, Giv’at Ram, Jerusalem, Israel. University of Rennes, Rennes, France. Nelson Mandela University, Port Elizabeth, South Africa.</mods:affiliation>
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<mods:nameIdentifier id="AC02DAAA4F3356FCABE14CAA4261F5F6" type="email">efrat.gavish-regev@mail.huji.ac.il</mods:nameIdentifier>
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<mods:title id="02865AAC364EE961EC2F027D2CE47BC6">European Journal of Taxonomy</mods:title>
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<mods:date id="4BB0ABAAB22250E9634D24553B5855EF">2022</mods:date>
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<treatment id="1C43EA6E9B56FFF3FD255977FDAA5764" ID-DOI="http://doi.org/10.5281/zenodo.6949786" ID-GBIF-Taxon="198139653" ID-Zenodo-Dep="6949786" LSID="urn:lsid:zoobank.org:act:A5A1F6AD-14B4-488F-BC74-A2B5E240A86B" httpUri="http://treatment.plazi.org/id/1C43EA6E9B56FFF3FD255977FDAA5764" lastPageId="17" lastPageNumber="17" pageId="13" pageNumber="14">
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<taxonomicName id="53EA20FB9B56FFEFFD255977FCD9507E" authority="Steinpress & Cohen & Pétillon & Chipman & Gavish-Regev, 2022" authorityName="Steinpress & Cohen & Pétillon & Chipman & Gavish-Regev" authorityYear="2022" box="[654,836,1602,1629]" class="Arachnida" family="Lycosidae" genus="Lycosa" kingdom="Animalia" order="Araneae" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="gesserit" status="sp. nov.">
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<emphasis id="A69E876A9B56FFEFFD255977FCD9507E" bold="true" box="[654,836,1602,1629]" italics="true" pageId="13" pageNumber="14">Lycosa gesserit</emphasis>
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<taxonomicNameLabel id="BDAD3A119B56FFEFFCE05977FC3B507F" box="[843,934,1603,1629]" pageId="13" pageNumber="14" rank="species">sp. nov.</taxonomicNameLabel>
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urn:lsid:zoobank.org:act:
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<uuid id="E04C61AD9B56FFEFFD235951FB5350A2" box="[648,1230,1637,1664]" pageId="13" pageNumber="14">A5A1F6AD-14B4-488F-BC74-A2B5E240A86B</uuid>
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<paragraph id="94555B789B56FFEFFE5459BDFBA95081" blockId="13.[358,1230,1602,1699]" box="[511,1076,1672,1699]" pageId="13" pageNumber="14">
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<figureCitation id="0CD147FD9B56FFEFFE5459BDFDDA5081" box="[511,583,1673,1699]" captionStart="Fig" captionStartId="5.[189,232,1882,1908]" captionTargetBox="[369,1219,265,1845]" captionTargetId="figure-11@5.[369,1219,265,1845]" captionTargetPageId="5" captionText="Fig. 1. Collecting localities for this study (small map points to a locality in southern Sinai). ■: Geolycosa vultuosa C.L. Koch, 1838.x: Lycosa gesseritsp. nov. o: Lycosasp. ★: Lycosa hyraculussp. nov. ●: Lycosa piochardi Simon, 1876. Isohyets represent mean annual precipitation in mm. Arrows point to localities represented in molecular phylogeny." figureDoi="http://doi.org/10.5281/zenodo.6916852" httpUri="https://zenodo.org/record/6916852/files/figure.png" pageId="13" pageNumber="14">Figs 1</figureCitation>
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<figureCitation id="0CD147FD9B56FFEFFDFD59BDFDE65081" box="[598,635,1673,1699]" captionStart="Fig" captionStartId="11.[189,232,1853,1879]" captionTargetBox="[202,1388,1047,1808]" captionTargetId="figure-169@11.[189,1398,1041,1814]" captionTargetPageId="11" captionText="Fig. 2. ♂♂ Pedipalps, ventral view. A. Geolycosa vultuosa C.L. Koch, 1838 (HUJ INV-AR20960). B. Lycosa gesserit sp. nov., holotype (HUJ INV-AR20631). C. Lycosa hyraculus sp. nov., holotype (HUJ INV-AR20818). D. Lycosa piochardi Simon, 1876 (HUJ INV-AR20948). E. Lycosa praegrandis C.L. Koch, 1836 (Veles, North Macedonia; 1 Jun. 2002; courtesy of Marjan Komnenov). Scale bars = 0.5 mm. Photos by I. Armiach Steinpress." figureDoi="http://doi.org/10.5281/zenodo.6916854" httpUri="https://zenodo.org/record/6916854/files/figure.png" pageId="13" pageNumber="14">2B</figureCitation>
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<figureCitation id="0CD147FD9B56FFEFFD2C59BDFD365081" box="[647,683,1673,1699]" captionStart="Fig" captionStartId="14.[189,232,1888,1914]" captionTargetBox="[206,1392,1257,1834]" captionTargetId="figure-249@14.[189,1398,1244,1849]" captionTargetPageId="14" captionText="Fig. 3. Male pedipalps, prolateral view. A. Geolycosa vultuosa C.L. Koch, 1838 (HUJ INV-AR20960). B. Lycosa gesserit sp. nov., holotype (HUJ INV-AR20631). C. Lycosa hyraculus sp. nov., holotype (HUJ INV-AR20818). D. Lycosa piochardi Simon, 1876 (HUJ INV-AR20948). Scale bars = 0.5 mm. Photos by I. Armiach Steinpress." figureDoi="http://doi.org/10.5281/zenodo.6916858" httpUri="https://zenodo.org/record/6916858/files/figure.png" pageId="13" pageNumber="14">3B</figureCitation>
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<figureCitation id="0CD147FD9B56FFEFFD1359BDFD415081" box="[696,732,1673,1699]" captionStart="Fig" captionStartId="15.[189,232,1885,1911]" captionTargetBox="[211,1382,1365,1839]" captionTargetId="figure-279@15.[189,1398,1357,1845]" captionTargetPageId="15" captionText="Fig. 4. Male pedipalps retrolateral view. A. Geolycosa vultuosa C.L. Koch, 1838 (HUJ INV-AR20960). B. Lycosa gesserit sp. nov., holotype (HUJ INV-AR20631). C. Lycosa hyraculus sp. nov., holotype (HUJ INV-AR20818). D. Lycosa piochardi Simon, 1876 (HUJ INV-AR20948). Scale bars = 0.5 mm. Photos by I. Armiach Steinpress." figureDoi="http://doi.org/10.5281/zenodo.6916860" httpUri="https://zenodo.org/record/6916860/files/figure.png" pageId="13" pageNumber="14">4B</figureCitation>
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<figureCitation id="0CD147FD9B56FFEFFD4359BCFC915081" box="[744,780,1672,1699]" captionStart="Fig" captionStartId="16.[190,233,1892,1918]" captionTargetBox="[195,1394,765,1850]" captionTargetId="figure-183@16.[189,1398,759,1856]" captionTargetPageId="16" captionText="Fig. 5. Male pedipalps, distal view. A. Geolycosa vultuosa C.L. Koch, 1838 (HUJ INV-AR20960). B. Lycosa gesserit sp. nov., holotype (HUJ INV-AR20631). C. Lycosa hyraculus sp. nov., holotype (HUJ INV-AR20818). D. Lycosa piochardi Simon, 1876 (HUJ INV-AR20948). Scale bars = 0.5 mm. Photos by I. Armiach Steinpress." figureDoi="http://doi.org/10.5281/zenodo.6916862" httpUri="https://zenodo.org/record/6916862/files/figure.png" pageId="13" pageNumber="14">5B</figureCitation>
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<figureCitation id="0CD147FD9B56FFEFFCB259BDFCA35081" box="[793,830,1673,1699]" captionStart="Fig" captionStartId="17.[190,233,1851,1877]" captionTargetBox="[209,1328,1004,1783]" captionTargetId="figure-192@17.[189,1398,984,1809]" captionTargetPageId="17" captionText="Fig. 6.Habitus, dorsal view. A. Geolycosa vultuosa C.L. Koch, 1838, ♂ (HUJ INV-AR20961). B. Lycosa gesserit sp. nov., holotype, ♂ (HUJ INV-AR20631). C. Lycosa hyraculus sp. nov., holotype, ♂ (HUJ INV-AR20818). D. Lycosa piochardi Simon, 1876, ♂ (HUJ INV-AR20597). E. Lycosa sp., ♀ (HUJ INV-AR20573). F. Lycosa hyraculus sp. nov., paratype, ♀ (HUJ INV-AR20817). G. Lycosa piochardi Simon, 1876, ♀ (SMFD11934). Scale bars = 10 mm. Photos by I. Armiach Steinpress." figureDoi="http://doi.org/10.5281/zenodo.6916866" httpUri="https://zenodo.org/record/6916866/files/figure.png" pageId="13" pageNumber="14">6B</figureCitation>
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<figureCitation id="0CD147FD9B56FFEFFCE159BDFCF25081" box="[842,879,1673,1699]" captionStart="Fig" captionStartId="18.[189,232,1851,1877]" captionTargetBox="[202,1335,1007,1798]" captionTargetId="figure-133@18.[189,1398,985,1812]" captionTargetPageId="18" captionText="Fig. 7. Habitus, ventral view. A. Geolycosa vultuosa C.L. Koch, 1838, ♂ (HUJ INV-AR20961). B. Lycosa gesserit sp. nov., holotype, ♂ (HUJ INV-AR20631). C. Lycosa hyraculus sp. nov., holotype, ♂ (HUJ INV-AR20818). D. Lycosa piochardi Simon, 1876, ♂ (HUJ INV-AR20597). E. Lycosa sp., ♀ (HUJ INV-AR20573). F. Lycosa hyraculus sp. nov., paratype, ♀ (HUJ INV-AR20817). G. Lycosa piochardi Simon, 1876, ♀ (SMFD11934). Scale bars = 10 mm. Photos by I. Armiach Steinpress." figureDoi="http://doi.org/10.5281/zenodo.6916868" httpUri="https://zenodo.org/record/6916868/files/figure.png" pageId="13" pageNumber="14">7B</figureCitation>
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<figureCitation id="0CD147FD9B56FFEFFCD059BDFC3D5081" box="[891,928,1673,1699]" captionStart="Fig" captionStartId="20.[189,232,1890,1916]" captionTargetBox="[189,1398,490,1850]" captionTargetId="figure-76@20.[189,1398,490,1850]" captionTargetPageId="20" captionText="Fig. 8. Live specimens. A. Geolycosa vultuosa C.L. Koch, 1838. B. Lycosa gesserit sp. nov., holotype, ♂ (HUJ INV-AR20631) C. Lycosa hyraculus sp. nov., juveniles. D. Lycosa hyraculus sp. nov., ♂ (specimen not collected). E. Hogna (cf.) graeca (specimen not collected). F. Hogna effera (O. Pickard- Cambridge, 1872). Photos by I. Armiach Steinpress." figureDoi="http://doi.org/10.5281/zenodo.6916870" httpUri="https://zenodo.org/record/6916870/files/figure.png" pageId="13" pageNumber="14">8B</figureCitation>
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<figureCitation id="0CD147FD9B56FFEFFC0759BDFC7D5081" box="[940,992,1673,1699]" captionStart="Fig" captionStartId="25.[188,231,1888,1914]" captionTargetBox="[189,1381,1297,1830]" captionTargetId="figure-278@25.[189,1398,1282,1850]" captionTargetPageId="25" captionText="Fig. 12. Lycosa ♂♂, conductor tip, photograph and outline. A. Lycosa praegrandis C.L. Koch, 1836. B. Lycosa gesserit sp. nov., holotype (HUJ INV-AR20631). C. Lycosa hyraculus sp. nov., holotype (HUJ INV-AR20818). D. Lycosa piochardi Simon, 1876 (HUJ INV-AR20855). Scale bar = 0.5 mm. Photos and drawings by I. Armiach Steinpress." figureDoi="http://doi.org/10.5281/zenodo.6916882" httpUri="https://zenodo.org/record/6916882/files/figure.png" pageId="13" pageNumber="14">12B</figureCitation>
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<figureCitation id="0CD147FD9B56FFEFFC4759BDFBA95081" box="[1004,1076,1673,1699]" captionStart-0="Fig" captionStart-1="Fig" captionStartId-0="43.[187,230,1916,1942]" captionStartId-1="44.[188,231,1952,1978]" captionTargetBox-0="[187,1396,685,1877]" captionTargetBox-1="[252,1343,812,1849]" captionTargetId-0="figure-155@43.[187,1396,685,1877]" captionTargetId-1="figure-145@44.[189,1398,729,1908]" captionTargetPageId-0="43" captionTargetPageId-1="44" captionText-0="Fig. 23. Bayesian phylogeny cladogram, showing the topology of Lycosa Latreille, 1804. Posterior probabilities of 1 marked by asterisk. Specimens used for this phylogeny detailed in Table 2. Photos by I. Armiach Steinpress." captionText-1="Fig. 24. Bayesian phylogeny cladogram, showing the topology of the oculata group. Posterior probabilities of 1 marked by asterisk. Specimens used for this phylogeny detailed in Table 2." figureDoi-0="http://doi.org/10.5281/zenodo.6916906" figureDoi-1="http://doi.org/10.5281/zenodo.6916908" httpUri-0="https://zenodo.org/record/6916906/files/figure.png" httpUri-1="https://zenodo.org/record/6916908/files/figure.png" pageId="13" pageNumber="14">23–24</figureCitation>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="DCF008F39B56FFEFFF1659FEFEA451C3" pageId="13" pageNumber="14" type="diagnosis">
|
||
<paragraph id="94555B789B56FFEFFF1659FEFEAB50C6" blockId="13.[189,310,1738,1764]" box="[189,310,1738,1764]" pageId="13" pageNumber="14">
|
||
<heading id="CF1DEC149B56FFEFFF1659FEFEAB50C6" bold="true" box="[189,310,1738,1764]" fontSize="11" level="3" pageId="13" pageNumber="14" reason="3">
|
||
<emphasis id="A69E876A9B56FFEFFF1659FEFEAB50C6" bold="true" box="[189,310,1738,1764]" pageId="13" pageNumber="14">Diagnosis</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="94555B789B56FFEFFF4159CDFEB75131" blockId="13.[187,1399,1785,1917]" box="[234,298,1785,1811]" pageId="13" pageNumber="14">
|
||
<emphasis id="A69E876A9B56FFEFFF4159CDFEB75131" bold="true" box="[234,298,1785,1811]" pageId="13" pageNumber="14">Male</emphasis>
|
||
</paragraph>
|
||
<paragraph id="94555B789B56FFEFFF165828FB6F515E" blockId="13.[187,1399,1785,1917]" pageId="13" pageNumber="14">
|
||
Recognized by pedipalp structure: TAT not bent posteriorly, directed retrolaterally. Contrasts with
|
||
<taxonomicName id="53EA20FB9B56FFEFFAB05829FF64517B" authority="Steinpress & Cohen & Pétillon & Chipman & Gavish-Regev, 2022" authorityName="Steinpress & Cohen & Pétillon & Chipman & Gavish-Regev" authorityYear="2022" class="Arachnida" family="Lycosidae" genus="Lycosa" kingdom="Animalia" order="Araneae" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="hyraculus" status="sp. nov.">
|
||
<emphasis id="A69E876A9B56FFEFFAB05829FF64517B" italics="true" pageId="13" pageNumber="14">L. hyraculus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="BDAD3A119B56FFEFFF555874FEC85178" box="[254,341,1856,1882]" pageId="13" pageNumber="14" rank="species">sp. nov.</taxonomicNameLabel>
|
||
by having TAT shorter than TAC (TAT as long as TAC in
|
||
<taxonomicName id="53EA20FB9B56FFEFFC7F5874FBFC517B" authority="Steinpress & Cohen & Pétillon & Chipman & Gavish-Regev, 2022" authorityName="Steinpress & Cohen & Pétillon & Chipman & Gavish-Regev" authorityYear="2022" box="[980,1121,1855,1881]" class="Arachnida" family="Lycosidae" genus="Lycosa" kingdom="Animalia" order="Araneae" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="hyraculus" status="sp. nov.">
|
||
<emphasis id="A69E876A9B56FFEFFC7F5874FBFC517B" box="[980,1121,1855,1881]" italics="true" pageId="13" pageNumber="14">L. hyraculus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="BDAD3A119B56FFEFFBCD5874FB5D5178" box="[1126,1216,1856,1882]" pageId="13" pageNumber="14" rank="species">sp. nov.</taxonomicNameLabel>
|
||
). Contrasts with all other known males of
|
||
<taxonomicName id="53EA20FB9B56FFEFFE775857FDB1515E" authorityName="Latreille" authorityYear="1804" box="[476,556,1891,1916]" class="Arachnida" family="Lycosidae" genus="Lycosa" kingdom="Animalia" order="Araneae" pageId="13" pageNumber="14" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="A69E876A9B56FFEFFE775857FDB1515E" box="[476,556,1891,1916]" italics="true" pageId="13" pageNumber="14">Lycosa</emphasis>
|
||
</taxonomicName>
|
||
, with TAT bent posteriorly (
|
||
<figureCitation id="0CD147FD9B56FFEFFCC75856FC54515E" box="[876,969,1890,1916]" captionStart="Fig" captionStartId="11.[189,232,1853,1879]" captionTargetBox="[202,1388,1047,1808]" captionTargetId="figure-169@11.[189,1398,1041,1814]" captionTargetPageId="11" captionText="Fig. 2. ♂♂ Pedipalps, ventral view. A. Geolycosa vultuosa C.L. Koch, 1838 (HUJ INV-AR20960). B. Lycosa gesserit sp. nov., holotype (HUJ INV-AR20631). C. Lycosa hyraculus sp. nov., holotype (HUJ INV-AR20818). D. Lycosa piochardi Simon, 1876 (HUJ INV-AR20948). E. Lycosa praegrandis C.L. Koch, 1836 (Veles, North Macedonia; 1 Jun. 2002; courtesy of Marjan Komnenov). Scale bars = 0.5 mm. Photos by I. Armiach Steinpress." figureDoi="http://doi.org/10.5281/zenodo.6916854" httpUri="https://zenodo.org/record/6916854/files/figure.png" pageId="13" pageNumber="14">Figs 2B</figureCitation>
|
||
,
|
||
<figureCitation id="0CD147FD9B56FFEFFC7F5856FC65515E" box="[980,1016,1890,1916]" captionStart="Fig" captionStartId="14.[189,232,1888,1914]" captionTargetBox="[206,1392,1257,1834]" captionTargetId="figure-249@14.[189,1398,1244,1849]" captionTargetPageId="14" captionText="Fig. 3. Male pedipalps, prolateral view. A. Geolycosa vultuosa C.L. Koch, 1838 (HUJ INV-AR20960). B. Lycosa gesserit sp. nov., holotype (HUJ INV-AR20631). C. Lycosa hyraculus sp. nov., holotype (HUJ INV-AR20818). D. Lycosa piochardi Simon, 1876 (HUJ INV-AR20948). Scale bars = 0.5 mm. Photos by I. Armiach Steinpress." figureDoi="http://doi.org/10.5281/zenodo.6916858" httpUri="https://zenodo.org/record/6916858/files/figure.png" pageId="13" pageNumber="14">3B</figureCitation>
|
||
,
|
||
<figureCitation id="0CD147FD9B56FFEFFBAF5856FBB5515E" box="[1028,1064,1890,1916]" captionStart="Fig" captionStartId="15.[189,232,1885,1911]" captionTargetBox="[211,1382,1365,1839]" captionTargetId="figure-279@15.[189,1398,1357,1845]" captionTargetPageId="15" captionText="Fig. 4. Male pedipalps retrolateral view. A. Geolycosa vultuosa C.L. Koch, 1838 (HUJ INV-AR20960). B. Lycosa gesserit sp. nov., holotype (HUJ INV-AR20631). C. Lycosa hyraculus sp. nov., holotype (HUJ INV-AR20818). D. Lycosa piochardi Simon, 1876 (HUJ INV-AR20948). Scale bars = 0.5 mm. Photos by I. Armiach Steinpress." figureDoi="http://doi.org/10.5281/zenodo.6916860" httpUri="https://zenodo.org/record/6916860/files/figure.png" pageId="13" pageNumber="14">4B</figureCitation>
|
||
,
|
||
<figureCitation id="0CD147FD9B56FFEFFB985856FBCB515F" box="[1075,1110,1890,1917]" captionStart="Fig" captionStartId="16.[190,233,1892,1918]" captionTargetBox="[195,1394,765,1850]" captionTargetId="figure-183@16.[189,1398,759,1856]" captionTargetPageId="16" captionText="Fig. 5. Male pedipalps, distal view. A. Geolycosa vultuosa C.L. Koch, 1838 (HUJ INV-AR20960). B. Lycosa gesserit sp. nov., holotype (HUJ INV-AR20631). C. Lycosa hyraculus sp. nov., holotype (HUJ INV-AR20818). D. Lycosa piochardi Simon, 1876 (HUJ INV-AR20948). Scale bars = 0.5 mm. Photos by I. Armiach Steinpress." figureDoi="http://doi.org/10.5281/zenodo.6916862" httpUri="https://zenodo.org/record/6916862/files/figure.png" pageId="13" pageNumber="14">5B</figureCitation>
|
||
,
|
||
<figureCitation id="0CD147FD9B56FFEFFBC95856FB18515E" box="[1122,1157,1890,1916]" captionStart="Fig" captionStartId="17.[190,233,1851,1877]" captionTargetBox="[209,1328,1004,1783]" captionTargetId="figure-192@17.[189,1398,984,1809]" captionTargetPageId="17" captionText="Fig. 6.Habitus, dorsal view. A. Geolycosa vultuosa C.L. Koch, 1838, ♂ (HUJ INV-AR20961). B. Lycosa gesserit sp. nov., holotype, ♂ (HUJ INV-AR20631). C. Lycosa hyraculus sp. nov., holotype, ♂ (HUJ INV-AR20818). D. Lycosa piochardi Simon, 1876, ♂ (HUJ INV-AR20597). E. Lycosa sp., ♀ (HUJ INV-AR20573). F. Lycosa hyraculus sp. nov., paratype, ♀ (HUJ INV-AR20817). G. Lycosa piochardi Simon, 1876, ♀ (SMFD11934). Scale bars = 10 mm. Photos by I. Armiach Steinpress." figureDoi="http://doi.org/10.5281/zenodo.6916866" httpUri="https://zenodo.org/record/6916866/files/figure.png" pageId="13" pageNumber="14">6B</figureCitation>
|
||
,
|
||
<figureCitation id="0CD147FD9B56FFEFFB3A5857FB29515F" box="[1169,1204,1891,1917]" captionStart="Fig" captionStartId="18.[189,232,1851,1877]" captionTargetBox="[202,1335,1007,1798]" captionTargetId="figure-133@18.[189,1398,985,1812]" captionTargetPageId="18" captionText="Fig. 7. Habitus, ventral view. A. Geolycosa vultuosa C.L. Koch, 1838, ♂ (HUJ INV-AR20961). B. Lycosa gesserit sp. nov., holotype, ♂ (HUJ INV-AR20631). C. Lycosa hyraculus sp. nov., holotype, ♂ (HUJ INV-AR20818). D. Lycosa piochardi Simon, 1876, ♂ (HUJ INV-AR20597). E. Lycosa sp., ♀ (HUJ INV-AR20573). F. Lycosa hyraculus sp. nov., paratype, ♀ (HUJ INV-AR20817). G. Lycosa piochardi Simon, 1876, ♀ (SMFD11934). Scale bars = 10 mm. Photos by I. Armiach Steinpress." figureDoi="http://doi.org/10.5281/zenodo.6916868" httpUri="https://zenodo.org/record/6916868/files/figure.png" pageId="13" pageNumber="14">7B</figureCitation>
|
||
,
|
||
<figureCitation id="0CD147FD9B56FFEFFB6B5856FB78515E" box="[1216,1253,1890,1916]" captionStart="Fig" captionStartId="20.[189,232,1890,1916]" captionTargetBox="[189,1398,490,1850]" captionTargetId="figure-76@20.[189,1398,490,1850]" captionTargetPageId="20" captionText="Fig. 8. Live specimens. A. Geolycosa vultuosa C.L. Koch, 1838. B. Lycosa gesserit sp. nov., holotype, ♂ (HUJ INV-AR20631) C. Lycosa hyraculus sp. nov., juveniles. D. Lycosa hyraculus sp. nov., ♂ (specimen not collected). E. Hogna (cf.) graeca (specimen not collected). F. Hogna effera (O. Pickard- Cambridge, 1872). Photos by I. Armiach Steinpress." figureDoi="http://doi.org/10.5281/zenodo.6916870" httpUri="https://zenodo.org/record/6916870/files/figure.png" pageId="13" pageNumber="14">8B</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="94555B789B56FFEFFF415890FED8519C" blockId="13.[189,325,1956,2017]" box="[234,325,1956,1982]" pageId="13" pageNumber="14">
|
||
<emphasis id="A69E876A9B56FFEFFF415890FED8519C" bold="true" box="[234,325,1956,1982]" pageId="13" pageNumber="14">Female</emphasis>
|
||
</paragraph>
|
||
<paragraph id="94555B789B56FFEFFF1658F3FEA451C3" blockId="13.[189,325,1956,2017]" box="[189,313,1991,2017]" pageId="13" pageNumber="14">Unknown.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="DCF008F39B55FFECFF165E3DFB515770" box="[189,1228,265,338]" pageId="14" pageNumber="15" type="etymology">
|
||
<paragraph id="94555B789B55FFECFF165E3DFED95701" blockId="14.[189,324,265,291]" box="[189,324,265,291]" pageId="14" pageNumber="15">
|
||
<heading id="CF1DEC149B55FFECFF165E3DFED95701" bold="true" box="[189,324,265,291]" fontSize="11" level="3" pageId="14" pageNumber="15" reason="3">
|
||
<emphasis id="A69E876A9B55FFECFF165E3DFED95701" bold="true" box="[189,324,265,291]" pageId="14" pageNumber="15">Etymology</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="94555B789B55FFECFF165E03FB515770" blockId="14.[189,1228,311,338]" box="[189,1228,311,338]" pageId="14" pageNumber="15">
|
||
Species named after the order of
|
||
<emphasis id="A69E876A9B55FFECFDEA5E0CFD785773" box="[577,741,311,337]" italics="true" pageId="14" pageNumber="15">Bene Gesserit</emphasis>
|
||
, from Frank Herbert’s 1965 novel,
|
||
<emphasis id="A69E876A9B55FFECFB2E5E0CFB585773" box="[1157,1221,312,337]" italics="true" pageId="14" pageNumber="15">Dune</emphasis>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="DCF008F39B55FFECFF165E4DFD145425" pageId="14" pageNumber="15" type="materials_examined">
|
||
<paragraph id="94555B789B55FFECFF165E4DFEF157B0" blockId="14.[189,364,376,403]" box="[189,364,376,403]" pageId="14" pageNumber="15">
|
||
<heading id="CF1DEC149B55FFECFF165E4DFEF157B0" bold="true" box="[189,364,376,403]" fontSize="11" level="3" pageId="14" pageNumber="15" reason="3">
|
||
<emphasis id="A69E876A9B55FFECFF165E4DFEF157B0" bold="true" box="[189,364,376,403]" pageId="14" pageNumber="15">Type material</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="94555B789B55FFECFF415E93FD145425" blockId="14.[189,1398,423,520]" pageId="14" pageNumber="15">
|
||
<materialsCitation id="248251259B55FFECFF415E93FD145425" ID-GBIF-Occurrence="3872858380" collectingDate="2017-09-07" collectionCode="HUJ" collectorName="E. Gavish-Regev" country="Israel" latitude="30.28" location="Har Karkom" longLatPrecision="733" longitude="34.74" municipality="Har Karkom" pageId="14" pageNumber="15" specimenCode="HUJ INV-AR20631" specimenCount="1" specimenCount-male="1" stateProvince="Negev" typeStatus="holotype">
|
||
<emphasis id="A69E876A9B55FFECFF415E93FEC157E3" bold="true" box="[234,348,423,449]" pageId="14" pageNumber="15">
|
||
<typeStatus id="4B51E5DA9B55FFECFF415E93FEC157E3" box="[234,348,423,449]" pageId="14" pageNumber="15" type="holotype">Holotype</typeStatus>
|
||
</emphasis>
|
||
<collectingCountry id="ECFD1BE89B55FFECFF165EFEFEBE57C6" box="[189,291,458,484]" name="Israel" pageId="14" pageNumber="15">ISRAEL</collectingCountry>
|
||
–
|
||
<emphasis id="A69E876A9B55FFECFEEC5EFEFE0957C6" bold="true" box="[327,404,458,484]" pageId="14" pageNumber="15">
|
||
<collectingRegion id="562E959A9B55FFECFEEC5EFEFE0957C6" box="[327,404,458,484]" pageId="14" pageNumber="15">Negev</collectingRegion>
|
||
</emphasis>
|
||
•
|
||
<specimenCount id="82EC90F19B55FFECFE045EFFFE5557C7" box="[431,456,459,485]" pageId="14" pageNumber="15" type="male">♂</specimenCount>
|
||
;
|
||
<collectingMunicipality id="7431C1029B55FFECFE7C5EFFFDED57C6" box="[471,624,458,485]" pageId="14" pageNumber="15">Har Karkom</collectingMunicipality>
|
||
; [
|
||
<geoCoordinate id="F1DE3DBF9B55FFECFD2E5EFEFD6557C7" box="[645,760,458,485]" degrees="30.28" direction="north" orientation="latitude" pageId="14" pageNumber="15" precision="555" value="30.28">30.28° N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="F1DE3DBF9B55FFECFCAD5EFEFCEE57C7" box="[774,883,458,485]" degrees="34.74" direction="east" orientation="longitude" pageId="14" pageNumber="15" precision="555" value="34.74">34.74° E</geoCoordinate>
|
||
];
|
||
<date id="E0547DB89B55FFECFC205EFFFB8357C6" box="[907,1054,458,485]" pageId="14" pageNumber="15" value="2017-09-07">
|
||
<collectingDate id="F01084509B55FFECFC205EFFFB8357C6" box="[907,1054,458,485]" pageId="14" pageNumber="15" value="2017-09-07">7 Sep. 2017</collectingDate>
|
||
</date>
|
||
;
|
||
<collectorName id="391F3EAE9B55FFECFB865EFFFB6157C6" box="[1069,1276,458,485]" pageId="14" pageNumber="15">E. Gavish-Regev</collectorName>
|
||
; col. sub.; collected at night;
|
||
<specimenCode id="C44CF3039B55FFECFE3C5EDAFD185425" box="[407,645,493,520]" pageId="14" pageNumber="15">HUJ INV-AR20631</specimenCode>
|
||
.
|
||
</materialsCitation>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="DCF008F39B55FFEDFF165D1BFE74541A" lastPageId="15" lastPageNumber="16" pageId="14" pageNumber="15" type="description">
|
||
<paragraph id="94555B789B55FFECFF165D1BFED3546B" blockId="14.[189,334,559,585]" box="[189,334,559,585]" pageId="14" pageNumber="15">
|
||
<heading id="CF1DEC149B55FFECFF165D1BFED3546B" bold="true" box="[189,334,559,585]" fontSize="11" level="3" pageId="14" pageNumber="15" reason="3">
|
||
<emphasis id="A69E876A9B55FFECFF165D1BFED3546B" bold="true" box="[189,334,559,585]" pageId="14" pageNumber="15">Description</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="94555B789B55FFECFF415D69FEB75455" blockId="14.[185,1399,605,773]" box="[234,298,605,631]" pageId="14" pageNumber="15">
|
||
<emphasis id="A69E876A9B55FFECFF415D69FEB75455" bold="true" box="[234,298,605,631]" pageId="14" pageNumber="15">Male</emphasis>
|
||
</paragraph>
|
||
<paragraph id="94555B789B55FFECFF165DB5FAEB54C0" blockId="14.[185,1399,605,773]" pageId="14" pageNumber="15">
|
||
<typeStatus id="4B51E5DA9B55FFECFF165DB5FEAD54B9" box="[189,304,641,667]" pageId="14" pageNumber="15" type="holotype">HOLOTYPE</typeStatus>
|
||
MEASUREMENTS.AME diameter:0.2;ALE diameter:0.18;PME diameter:0.8;PLE diameter:0.7; carapace length: 6.6; carapace width: 4.8; carapace maximal height: 2.1; abdomen length: 5.2; leg I (Fe, Pa, Ti, Mt, Tr): 5.5, 2.5, 4.9, 5.5, 2.8; leg II: 5.1, 2.3, 4.6, 5.3, 2.9; leg III: 4.9, 2.4, 4.1, 5.9, 2.3; leg IV:
|
||
</paragraph>
|
||
<paragraph id="94555B789B55FFECFF165DDEFE3C5526" blockId="14.[185,1399,605,773]" box="[189,417,746,773]" pageId="14" pageNumber="15">6, 2.6, 5.5, 8.2, 3.4.</paragraph>
|
||
<paragraph id="94555B789B55FFECFF165C18FD09554B" blockId="14.[189,1398,810,873]" pageId="14" pageNumber="15">CARAPACE. Ocular area>⅓ length of carapace, distinctly raised, making the carapace between PLE and fovea appear slightly concave in profile.</paragraph>
|
||
<paragraph id="94555B789B55FFECFF165CA4FCFA525C" blockId="14.[189,1400,912,1150]" pageId="14" pageNumber="15">
|
||
COLOR. Carapace cream colored, with dark brown median bands, darker on margins, connecting to dark ocular area. Perimeter light brown. Sternum grey. Chelicerae reddish-brown. Abdomen yellow infused with light brown dorsad, cardiac mark brown, darker on margins. Abdomen venter with black patch surrounding epigastric fold, connecting to two parallel black bars stretching towards spinnerets (Π shaped patch). Spinnerets brown. Legs cream colored, infused with brown dorsad, yellow-grey ventrad, to reddish-brown distad, with two black bands on tibia. Coxae dark grey, trochanters brown. Pedipalps yellow, to dark brown distad (
|
||
<figureCitation id="0CD147FD9B55FFECFD3D5B57FD6B525F" box="[662,758,1123,1149]" captionStart="Fig" captionStartId="17.[190,233,1851,1877]" captionTargetBox="[209,1328,1004,1783]" captionTargetId="figure-192@17.[189,1398,984,1809]" captionTargetPageId="17" captionText="Fig. 6.Habitus, dorsal view. A. Geolycosa vultuosa C.L. Koch, 1838, ♂ (HUJ INV-AR20961). B. Lycosa gesserit sp. nov., holotype, ♂ (HUJ INV-AR20631). C. Lycosa hyraculus sp. nov., holotype, ♂ (HUJ INV-AR20818). D. Lycosa piochardi Simon, 1876, ♂ (HUJ INV-AR20597). E. Lycosa sp., ♀ (HUJ INV-AR20573). F. Lycosa hyraculus sp. nov., paratype, ♀ (HUJ INV-AR20817). G. Lycosa piochardi Simon, 1876, ♀ (SMFD11934). Scale bars = 10 mm. Photos by I. Armiach Steinpress." figureDoi="http://doi.org/10.5281/zenodo.6916866" httpUri="https://zenodo.org/record/6916866/files/figure.png" pageId="14" pageNumber="15">Figs 6B</figureCitation>
|
||
,
|
||
<figureCitation id="0CD147FD9B55FFECFCA85B50FCBA525C" box="[771,807,1124,1150]" captionStart="Fig" captionStartId="18.[189,232,1851,1877]" captionTargetBox="[202,1335,1007,1798]" captionTargetId="figure-133@18.[189,1398,985,1812]" captionTargetPageId="18" captionText="Fig. 7. Habitus, ventral view. A. Geolycosa vultuosa C.L. Koch, 1838, ♂ (HUJ INV-AR20961). B. Lycosa gesserit sp. nov., holotype, ♂ (HUJ INV-AR20631). C. Lycosa hyraculus sp. nov., holotype, ♂ (HUJ INV-AR20818). D. Lycosa piochardi Simon, 1876, ♂ (HUJ INV-AR20597). E. Lycosa sp., ♀ (HUJ INV-AR20573). F. Lycosa hyraculus sp. nov., paratype, ♀ (HUJ INV-AR20817). G. Lycosa piochardi Simon, 1876, ♀ (SMFD11934). Scale bars = 10 mm. Photos by I. Armiach Steinpress." figureDoi="http://doi.org/10.5281/zenodo.6916868" httpUri="https://zenodo.org/record/6916868/files/figure.png" pageId="14" pageNumber="15">7B</figureCitation>
|
||
,
|
||
<figureCitation id="0CD147FD9B55FFECFC9F5B50FCC4525C" box="[820,857,1124,1150]" captionStart="Fig" captionStartId="20.[189,232,1890,1916]" captionTargetBox="[189,1398,490,1850]" captionTargetId="figure-76@20.[189,1398,490,1850]" captionTargetPageId="20" captionText="Fig. 8. Live specimens. A. Geolycosa vultuosa C.L. Koch, 1838. B. Lycosa gesserit sp. nov., holotype, ♂ (HUJ INV-AR20631) C. Lycosa hyraculus sp. nov., juveniles. D. Lycosa hyraculus sp. nov., ♂ (specimen not collected). E. Hogna (cf.) graeca (specimen not collected). F. Hogna effera (O. Pickard- Cambridge, 1872). Photos by I. Armiach Steinpress." figureDoi="http://doi.org/10.5281/zenodo.6916870" httpUri="https://zenodo.org/record/6916870/files/figure.png" pageId="14" pageNumber="15">8B</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<caption id="C0950BF09B55FFECFF165854FDA351C1" ID-DOI="http://doi.org/10.5281/zenodo.6916858" ID-Zenodo-Dep="6916858" httpUri="https://zenodo.org/record/6916858/files/figure.png" pageId="14" pageNumber="15" startId="14.[189,232,1888,1914]" targetBox="[206,1392,1257,1834]" targetPageId="14">
|
||
<paragraph id="94555B789B55FFECFF165854FDA351C1" blockId="14.[189,1401,1888,2020]" pageId="14" pageNumber="15">
|
||
<emphasis id="A69E876A9B55FFECFF165854FE965158" bold="true" box="[189,267,1888,1914]" pageId="14" pageNumber="15">Fig. 3.</emphasis>
|
||
Male pedipalps, prolateral view.
|
||
<emphasis id="A69E876A9B55FFECFD385854FD355158" bold="true" box="[659,680,1888,1914]" pageId="14" pageNumber="15">A</emphasis>
|
||
.
|
||
<taxonomicName id="53EA20FB9B55FFECFD1C5854FBF75158" authority="C. L. Koch, 1838" authorityName="C. L. Koch" authorityYear="1838" box="[695,1130,1888,1914]" class="Arachnida" family="Lycosidae" genus="Geolycosa" kingdom="Animalia" order="Araneae" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="vultuosa">
|
||
<emphasis id="A69E876A9B55FFECFD1C5854FC005158" box="[695,925,1888,1914]" italics="true" pageId="14" pageNumber="15">Geolycosa vultuosa</emphasis>
|
||
C.L.
|
||
<bibRefCitation id="F07B26899B55FFECFC4A5854FBF75158" author="Koch C. L." box="[993,1130,1888,1914]" pageId="14" pageNumber="15" refId="ref31028" refString="Koch C. L. 1838. Die Arachniden. C. H. Zeh'sche Buchhandlung, Nurnberg. https: // doi. org / 10.5962 / bhl. title. 43744" type="book" year="1838">Koch, 1838</bibRefCitation>
|
||
</taxonomicName>
|
||
(HUJ INV-AR20960).
|
||
<emphasis id="A69E876A9B55FFECFF1658B7FF4C51BF" bold="true" box="[189,209,1923,1949]" pageId="14" pageNumber="15">B</emphasis>
|
||
.
|
||
<taxonomicName id="53EA20FB9B55FFECFF4858B0FE0651BF" authority="Steinpress & Cohen & Pétillon & Chipman & Gavish-Regev, 2022" authorityName="Steinpress & Cohen & Pétillon & Chipman & Gavish-Regev" authorityYear="2022" box="[227,411,1924,1949]" class="Arachnida" family="Lycosidae" genus="Lycosa" kingdom="Animalia" order="Araneae" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="gesserit" status="sp. nov.">
|
||
<emphasis id="A69E876A9B55FFECFF4858B0FE0651BF" box="[227,411,1924,1949]" italics="true" pageId="14" pageNumber="15">Lycosa gesserit</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="BDAD3A119B55FFECFE0E58B0FD9851BF" box="[421,517,1924,1950]" pageId="14" pageNumber="15" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, holotype (HUJ INV-AR20631).
|
||
<emphasis id="A69E876A9B55FFECFC3758B7FC2C51BF" bold="true" box="[924,945,1923,1949]" pageId="14" pageNumber="15">C</emphasis>
|
||
.
|
||
<taxonomicName id="53EA20FB9B55FFECFC6858B0FB0951BF" authority="Steinpress & Cohen & Pétillon & Chipman & Gavish-Regev, 2022" authorityName="Steinpress & Cohen & Pétillon & Chipman & Gavish-Regev" authorityYear="2022" box="[963,1172,1923,1949]" class="Arachnida" family="Lycosidae" genus="Lycosa" kingdom="Animalia" order="Araneae" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="hyraculus" status="sp. nov.">
|
||
<emphasis id="A69E876A9B55FFECFC6858B0FB0951BF" box="[963,1172,1923,1949]" italics="true" pageId="14" pageNumber="15">Lycosa hyraculus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="BDAD3A119B55FFECFB3558B0FB6051BF" box="[1182,1277,1924,1950]" pageId="14" pageNumber="15" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, holotype (HUJ INV-AR20818).
|
||
<emphasis id="A69E876A9B55FFECFE645892FE7E51E2" bold="true" box="[463,483,1958,1984]" pageId="14" pageNumber="15">D</emphasis>
|
||
.
|
||
<taxonomicName id="53EA20FB9B55FFECFE595893FCFC51E2" authority="Simon, 1876" authorityName="Simon" authorityYear="1876" box="[498,865,1958,1984]" class="Arachnida" family="Lycosidae" genus="Lycosa" kingdom="Animalia" order="Araneae" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="piochardi">
|
||
<emphasis id="A69E876A9B55FFECFE595893FD2251E2" box="[498,703,1958,1984]" italics="true" pageId="14" pageNumber="15">Lycosa piochardi</emphasis>
|
||
<bibRefCitation id="F07B26899B55FFECFD625892FCFC51E2" author="Simon E." box="[713,865,1958,1984]" pageId="14" pageNumber="15" pagination="57 - 91" refId="ref32282" refString="Simon E. 1876. Etudes arachnologiques. 4 e memoire. VII. Revision des especes europeennes du groupe de la Lycosa tarentula Rossi. Annales de la Societe entomologique de France (5) 6: 57 - 91." type="journal article" year="1876">Simon, 1876</bibRefCitation>
|
||
</taxonomicName>
|
||
(HUJ INV-AR20948). Scale bars = 0.5 mm. Photos by I. Armiach Steinpress.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="94555B789B54FFEDFF165E3DFD5157F1" blockId="15.[188,1400,265,468]" pageId="15" pageNumber="16">
|
||
GENITALIA. Genital opening sclerotized, similar to subadult epigyne. Cymbium reddish-brown. Tegular apophysis (TA) lying flat on bulb, oriented retrolaterally. TAT highly sclerotized, dark brown, not bent posteriorly (unlike most other species of
|
||
<taxonomicName id="53EA20FB9B54FFEDFD045E64FC9C574B" authorityName="Latreille" authorityYear="1804" box="[687,769,336,361]" class="Arachnida" family="Lycosidae" genus="Lycosa" kingdom="Animalia" order="Araneae" pageId="15" pageNumber="16" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="A69E876A9B54FFEDFD045E64FC9C574B" box="[687,769,336,361]" italics="true" pageId="15" pageNumber="16">Lycosa</emphasis>
|
||
</taxonomicName>
|
||
), but slightly bent ventrad. Crest of TA (CTA) with smooth edge, and abrupt, rounded end at base of TAT. TAT shorter than width of CTA. Tip of conductor membranous, triangular, with smooth edges. Synembolus with large, slightly sclerotized lobe (SEL), on retrolateral edge (
|
||
<figureCitation id="0CD147FD9B54FFEDFE205E8DFE7157F1" box="[395,492,441,467]" captionStart="Fig" captionStartId="11.[189,232,1853,1879]" captionTargetBox="[202,1388,1047,1808]" captionTargetId="figure-169@11.[189,1398,1041,1814]" captionTargetPageId="11" captionText="Fig. 2. ♂♂ Pedipalps, ventral view. A. Geolycosa vultuosa C.L. Koch, 1838 (HUJ INV-AR20960). B. Lycosa gesserit sp. nov., holotype (HUJ INV-AR20631). C. Lycosa hyraculus sp. nov., holotype (HUJ INV-AR20818). D. Lycosa piochardi Simon, 1876 (HUJ INV-AR20948). E. Lycosa praegrandis C.L. Koch, 1836 (Veles, North Macedonia; 1 Jun. 2002; courtesy of Marjan Komnenov). Scale bars = 0.5 mm. Photos by I. Armiach Steinpress." figureDoi="http://doi.org/10.5281/zenodo.6916854" httpUri="https://zenodo.org/record/6916854/files/figure.png" pageId="15" pageNumber="16">Figs 2B</figureCitation>
|
||
,
|
||
<figureCitation id="0CD147FD9B54FFEDFE535E8DFD8157F1" box="[504,540,441,467]" captionStart="Fig" captionStartId="14.[189,232,1888,1914]" captionTargetBox="[206,1392,1257,1834]" captionTargetId="figure-249@14.[189,1398,1244,1849]" captionTargetPageId="14" captionText="Fig. 3. Male pedipalps, prolateral view. A. Geolycosa vultuosa C.L. Koch, 1838 (HUJ INV-AR20960). B. Lycosa gesserit sp. nov., holotype (HUJ INV-AR20631). C. Lycosa hyraculus sp. nov., holotype (HUJ INV-AR20818). D. Lycosa piochardi Simon, 1876 (HUJ INV-AR20948). Scale bars = 0.5 mm. Photos by I. Armiach Steinpress." figureDoi="http://doi.org/10.5281/zenodo.6916858" httpUri="https://zenodo.org/record/6916858/files/figure.png" pageId="15" pageNumber="16">3B</figureCitation>
|
||
,
|
||
<figureCitation id="0CD147FD9B54FFEDFD825E8DFDD057F1" box="[553,589,441,467]" captionStart="Fig" captionStartId="15.[189,232,1885,1911]" captionTargetBox="[211,1382,1365,1839]" captionTargetId="figure-279@15.[189,1398,1357,1845]" captionTargetPageId="15" captionText="Fig. 4. Male pedipalps retrolateral view. A. Geolycosa vultuosa C.L. Koch, 1838 (HUJ INV-AR20960). B. Lycosa gesserit sp. nov., holotype (HUJ INV-AR20631). C. Lycosa hyraculus sp. nov., holotype (HUJ INV-AR20818). D. Lycosa piochardi Simon, 1876 (HUJ INV-AR20948). Scale bars = 0.5 mm. Photos by I. Armiach Steinpress." figureDoi="http://doi.org/10.5281/zenodo.6916860" httpUri="https://zenodo.org/record/6916860/files/figure.png" pageId="15" pageNumber="16">4B</figureCitation>
|
||
,
|
||
<figureCitation id="0CD147FD9B54FFEDFDF25E8DFDE357F6" box="[601,638,441,468]" captionStart="Fig" captionStartId="16.[190,233,1892,1918]" captionTargetBox="[195,1394,765,1850]" captionTargetId="figure-183@16.[189,1398,759,1856]" captionTargetPageId="16" captionText="Fig. 5. Male pedipalps, distal view. A. Geolycosa vultuosa C.L. Koch, 1838 (HUJ INV-AR20960). B. Lycosa gesserit sp. nov., holotype (HUJ INV-AR20631). C. Lycosa hyraculus sp. nov., holotype (HUJ INV-AR20818). D. Lycosa piochardi Simon, 1876 (HUJ INV-AR20948). Scale bars = 0.5 mm. Photos by I. Armiach Steinpress." figureDoi="http://doi.org/10.5281/zenodo.6916862" httpUri="https://zenodo.org/record/6916862/files/figure.png" pageId="15" pageNumber="16">5B</figureCitation>
|
||
,
|
||
<figureCitation id="0CD147FD9B54FFEDFD215E8DFD2257F1" box="[650,703,441,467]" captionStart="Fig" captionStartId="25.[188,231,1888,1914]" captionTargetBox="[189,1381,1297,1830]" captionTargetId="figure-278@25.[189,1398,1282,1850]" captionTargetPageId="25" captionText="Fig. 12. Lycosa ♂♂, conductor tip, photograph and outline. A. Lycosa praegrandis C.L. Koch, 1836. B. Lycosa gesserit sp. nov., holotype (HUJ INV-AR20631). C. Lycosa hyraculus sp. nov., holotype (HUJ INV-AR20818). D. Lycosa piochardi Simon, 1876 (HUJ INV-AR20855). Scale bar = 0.5 mm. Photos and drawings by I. Armiach Steinpress." figureDoi="http://doi.org/10.5281/zenodo.6916882" httpUri="https://zenodo.org/record/6916882/files/figure.png" pageId="15" pageNumber="16">12B</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="94555B789B54FFEDFF165ECFFE74541A" blockId="15.[189,1398,506,568]" pageId="15" pageNumber="16">LEGS. Metatarsus I ventral spination: 2 pairs + apical triplet. Tibia I ventral spination: 2 pairs + apical pair. Tarsus with scopula.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="DCF008F39B54FFEDFF165D6BFDB05516" pageId="15" pageNumber="16" type="biology_ecology">
|
||
<paragraph id="94555B789B54FFEDFF165D6BFEE3545B" blockId="15.[189,382,607,633]" box="[189,382,607,633]" pageId="15" pageNumber="16">
|
||
<heading id="CF1DEC149B54FFEDFF165D6BFEE3545B" bold="true" box="[189,382,607,633]" fontSize="11" level="3" pageId="15" pageNumber="16" reason="3">
|
||
<emphasis id="A69E876A9B54FFEDFF165D6BFEE3545B" bold="true" box="[189,382,607,633]" pageId="15" pageNumber="16">Natural history</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="94555B789B54FFEDFF165DB9FDB05516" blockId="15.[187,1399,653,821]" pageId="15" pageNumber="16">
|
||
This species is nocturnal. A single juvenile male was collected at night, in September, at
|
||
<quantity id="5312F69D9B54FFEDFB115DB9FA9E548A" box="[1210,1283,653,680]" metricMagnitude="2" metricUnit="m" metricValue="6.5" pageId="15" pageNumber="16" unit="m" value="650.0">650 m</quantity>
|
||
a.s.l., in a stony plain near Har Karkom, a secluded tableland in the hyperarid desert (annual precipitation <
|
||
<quantity id="5312F69D9B54FFEDFAB75D85FAF254E9" box="[1308,1391,689,716]" metricMagnitude="-2" metricUnit="m" metricValue="7.5" pageId="15" pageNumber="16" unit="mm" value="75.0">75 mm</quantity>
|
||
) (
|
||
<figureCitation id="0CD147FD9B54FFEDFF6F5DE0FE9354CC" box="[196,270,724,750]" captionStart="Fig" captionStartId="5.[189,232,1882,1908]" captionTargetBox="[369,1219,265,1845]" captionTargetId="figure-11@5.[369,1219,265,1845]" captionTargetPageId="5" captionText="Fig. 1. Collecting localities for this study (small map points to a locality in southern Sinai). ■: Geolycosa vultuosa C.L. Koch, 1838.x: Lycosa gesseritsp. nov. o: Lycosasp. ★: Lycosa hyraculussp. nov. ●: Lycosa piochardi Simon, 1876. Isohyets represent mean annual precipitation in mm. Arrows point to localities represented in molecular phylogeny." figureDoi="http://doi.org/10.5281/zenodo.6916852" httpUri="https://zenodo.org/record/6916852/files/figure.png" pageId="15" pageNumber="16">Fig. 1</figureCitation>
|
||
,
|
||
<tableCitation id="D9686EC39B54FFEDFE8B5DE0FEE354CC" box="[288,382,724,750]" captionStart="Table 4" captionStartId="13.[189,259,265,291]" captionTargetId="graphics-301@13.[189,1397,365,775]" captionTargetPageId="13" captionText="Table 4. Documented activity of Lycosa Latreille, 1804 and Geolycosa Montgomery, 1904 in Israel (by month). Females with egg-sacs marked by ‘o’. Grey squares without a sex sign denote juveniles only." httpUri="http://table.plazi.org/id/C0950BF09B56FFEFFF165E3DFAF95764" pageId="15" pageNumber="16" tableUuid="C0950BF09B56FFEFFF165E3DFAF95764">Table 4</tableCitation>
|
||
). The specimen was mature after two molts in captivity, in
|
||
<date id="E0547DB89B54FFEDFBC95DE0FA9D54CC" box="[1122,1280,724,750]" pageId="15" pageNumber="16" value="2018-08">August 2018</date>
|
||
. Like the closely related
|
||
<taxonomicName id="53EA20FB9B54FFEDFEC05DCCFE605533" authority="Steinpress & Cohen & Pétillon & Chipman & Gavish-Regev, 2022" authorityName="Steinpress & Cohen & Pétillon & Chipman & Gavish-Regev" authorityYear="2022" box="[363,509,759,785]" class="Arachnida" family="Lycosidae" genus="Lycosa" kingdom="Animalia" order="Araneae" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="hyraculus" status="sp. nov.">
|
||
<emphasis id="A69E876A9B54FFEDFEC05DCCFE605533" box="[363,509,759,785]" italics="true" pageId="15" pageNumber="16">L. hyraculus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="BDAD3A119B54FFEDFDAF5DCCFDC25530" box="[516,607,760,786]" pageId="15" pageNumber="16" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, it makes a burrow with a trapdoor. Shares habitat with
|
||
<taxonomicName id="53EA20FB9B54FFEDFB745DCCFAF35533" authorityName="Simon" authorityYear="1876" box="[1247,1390,759,785]" class="Arachnida" family="Lycosidae" genus="Lycosa" kingdom="Animalia" order="Araneae" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="piochardi">
|
||
<emphasis id="A69E876A9B54FFEDFB745DCCFAF35533" box="[1247,1390,759,785]" italics="true" pageId="15" pageNumber="16">L. piochardi</emphasis>
|
||
</taxonomicName>
|
||
. Reproductive season unknown.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="DCF008F39B54FFEDFF165C68FA945587" box="[189,1289,860,933]" pageId="15" pageNumber="16" type="distribution">
|
||
<paragraph id="94555B789B54FFEDFF165C68FECB5554" blockId="15.[189,342,860,886]" box="[189,342,860,886]" pageId="15" pageNumber="16">
|
||
<heading id="CF1DEC149B54FFEDFF165C68FECB5554" bold="true" box="[189,342,860,886]" fontSize="11" level="3" pageId="15" pageNumber="16" reason="3">
|
||
<emphasis id="A69E876A9B54FFEDFF165C68FECB5554" bold="true" box="[189,342,860,886]" pageId="15" pageNumber="16">Distribution</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="94555B789B54FFEDFF165CBEFA945587" blockId="15.[189,1289,906,933]" box="[189,1289,906,933]" pageId="15" pageNumber="16">
|
||
<collectingCountry id="ECFD1BE89B54FFEDFF165CBEFEB55586" box="[189,296,906,932]" name="Israel" pageId="15" pageNumber="16">ISRAEL</collectingCountry>
|
||
. Possibly endemic to the Har Karkom tableland or found throughout hyperarid zone.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="DCF008F39B54FFF3FF165CFFFDAA5764" lastPageId="17" lastPageNumber="18" pageId="15" pageNumber="16" type="discussion">
|
||
<paragraph id="94555B789B54FFEDFF165CFFFEB955C7" blockId="15.[189,292,971,997]" box="[189,292,971,997]" pageId="15" pageNumber="16">
|
||
<heading id="CF1DEC149B54FFEDFF165CFFFEB955C7" bold="true" box="[189,292,971,997]" fontSize="11" level="3" pageId="15" pageNumber="16" reason="3">
|
||
<emphasis id="A69E876A9B54FFEDFF165CFFFEB955C7" bold="true" box="[189,292,971,997]" pageId="15" pageNumber="16">Records</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="94555B789B54FFEDFF165CCCFD095230" blockId="15.[189,660,1016,1043]" box="[189,660,1016,1043]" pageId="15" pageNumber="16">
|
||
<collectingCountry id="ECFD1BE89B54FFEDFF165CCCFEB55230" box="[189,296,1016,1042]" name="Israel" pageId="15" pageNumber="16">ISRAEL</collectingCountry>
|
||
: Negev (Har Karkom) (
|
||
<figureCitation id="0CD147FD9B54FFEDFD955CCCFD1E5230" box="[574,643,1016,1042]" captionStart="Fig" captionStartId="5.[189,232,1882,1908]" captionTargetBox="[369,1219,265,1845]" captionTargetId="figure-11@5.[369,1219,265,1845]" captionTargetPageId="5" captionText="Fig. 1. Collecting localities for this study (small map points to a locality in southern Sinai). ■: Geolycosa vultuosa C.L. Koch, 1838.x: Lycosa gesseritsp. nov. o: Lycosasp. ★: Lycosa hyraculussp. nov. ●: Lycosa piochardi Simon, 1876. Isohyets represent mean annual precipitation in mm. Arrows point to localities represented in molecular phylogeny." figureDoi="http://doi.org/10.5281/zenodo.6916852" httpUri="https://zenodo.org/record/6916852/files/figure.png" pageId="15" pageNumber="16">Fig. 1</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="94555B789B54FFEDFF165B03FEFA5273" blockId="15.[189,359,1079,1105]" box="[189,359,1079,1105]" pageId="15" pageNumber="16">
|
||
<heading id="CF1DEC149B54FFEDFF165B03FEFA5273" bold="true" box="[189,359,1079,1105]" fontSize="11" level="3" pageId="15" pageNumber="16" reason="3">
|
||
<emphasis id="A69E876A9B54FFEDFF165B03FEFA5273" bold="true" box="[189,359,1079,1105]" pageId="15" pageNumber="16">Relationships</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="94555B789B54FFEDFF165B51FA8E52E1" blockId="15.[186,1399,1125,1219]" pageId="15" pageNumber="16">
|
||
According to our molecular phylogeny, this species is the sister taxon of
|
||
<taxonomicName id="53EA20FB9B54FFEDFBB85B52FB41525D" authority="Steinpress & Cohen & Pétillon & Chipman & Gavish-Regev, 2022" authorityName="Steinpress & Cohen & Pétillon & Chipman & Gavish-Regev" authorityYear="2022" box="[1043,1244,1125,1151]" class="Arachnida" family="Lycosidae" genus="Lycosa" kingdom="Animalia" order="Araneae" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="hyraculus" status="sp. nov.">
|
||
<emphasis id="A69E876A9B54FFEDFBB85B52FB41525D" box="[1043,1244,1125,1151]" italics="true" pageId="15" pageNumber="16">Lycosa hyraculus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="BDAD3A119B54FFEDFB485B51FADD525D" box="[1251,1344,1125,1151]" pageId="15" pageNumber="16" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, and closely related to
|
||
<taxonomicName id="53EA20FB9B54FFEDFED05BBCFDB95283" authorityName="CRBA-LC" authorityYear="1019" baseAuthorityName="Simon" baseAuthorityYear="1876" box="[379,548,1159,1185]" class="Arachnida" family="Lycosidae" genus="Lycosa" kingdom="Animalia" order="Araneae" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="oculata">
|
||
<emphasis id="A69E876A9B54FFEDFED05BBCFDB95283" box="[379,548,1159,1185]" italics="true" pageId="15" pageNumber="16">Lycosa oculata</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="53EA20FB9B54FFEDFD995BBCFC985283" authorityName="CRBA-LC" authorityYear="1019" baseAuthorityName="Simon" baseAuthorityYear="1876" box="[562,773,1159,1185]" class="Arachnida" family="Lycosidae" genus="Lycosa" isUncertain="true" kingdom="Animalia" order="Araneae" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="oculata">
|
||
<emphasis id="A69E876A9B54FFEDFD995BBCFD1F5283" box="[562,642,1160,1185]" italics="true" pageId="15" pageNumber="16">Lycosa</emphasis>
|
||
aff.
|
||
<emphasis id="A69E876A9B54FFEDFD1B5BB3FC985283" box="[688,773,1159,1185]" italics="true" pageId="15" pageNumber="16">oculata</emphasis>
|
||
</taxonomicName>
|
||
1 and
|
||
<taxonomicName id="53EA20FB9B54FFEDFCEC5BBCFB805283" authorityName="CRBA-LC" authorityYear="1019" baseAuthorityName="Simon" baseAuthorityYear="1876" box="[839,1053,1159,1185]" class="Arachnida" family="Lycosidae" genus="Lycosa" isUncertain="true" kingdom="Animalia" order="Araneae" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="oculata">
|
||
<emphasis id="A69E876A9B54FFEDFCEC5BBCFC0A5283" box="[839,919,1160,1185]" italics="true" pageId="15" pageNumber="16">Lycosa</emphasis>
|
||
aff.
|
||
<emphasis id="A69E876A9B54FFEDFC635BB3FB805283" box="[968,1053,1159,1185]" italics="true" pageId="15" pageNumber="16">oculata</emphasis>
|
||
</taxonomicName>
|
||
2 (
|
||
<figureCitation id="0CD147FD9B54FFEDFB915BB3FB215283" box="[1082,1212,1159,1185]" captionStart-0="Fig" captionStart-1="Fig" captionStart-2="Fig" captionStart-3="Fig" captionStartId-0="40.[189,231,1958,1984]" captionStartId-1="41.[189,232,1886,1912]" captionStartId-2="43.[187,230,1916,1942]" captionStartId-3="44.[188,231,1952,1978]" captionTargetBox-0="[189,1398,1015,1916]" captionTargetBox-1="[189,1398,938,1845]" captionTargetBox-2="[187,1396,685,1877]" captionTargetBox-3="[252,1343,812,1849]" captionTargetId-0="figure-221@40.[189,1398,1012,1916]" captionTargetId-1="figure-185@41.[189,1398,938,1845]" captionTargetId-2="figure-155@43.[187,1396,685,1877]" captionTargetId-3="figure-145@44.[189,1398,729,1908]" captionTargetPageId-0="40" captionTargetPageId-1="41" captionTargetPageId-2="43" captionTargetPageId-3="44" captionText-0="Fig. 21. Typical habitats. A. Mediterranean grassland, Karmiel. B. Desert loess plain, Yeruham. C. Desert rocky slope, Yeruham. D. Mediterranean dwarf scrub (batha), Yodfat. Photos by I. Armiach Steinpress." captionText-1="Fig. 22. Retreats. A. Geolycosa vultuosa C.L. Koch, 1838, burrow with turret made of soil. B. Lycosa hyraculus sp. nov., burrow with door, Midreshet Ben-Gurion. C–E. Lycosa piochardi Simon, 1876. C. Burrow with turret made of plant material, Midreshet Ben-Gurion. D. Burrow without turret, Tel Kofer. E. Retreat under stone, Ariel Sharon camp. Photos by I. Armiach Steinpress." captionText-2="Fig. 23. Bayesian phylogeny cladogram, showing the topology of Lycosa Latreille, 1804. Posterior probabilities of 1 marked by asterisk. Specimens used for this phylogeny detailed in Table 2. Photos by I. Armiach Steinpress." captionText-3="Fig. 24. Bayesian phylogeny cladogram, showing the topology of the oculata group. Posterior probabilities of 1 marked by asterisk. Specimens used for this phylogeny detailed in Table 2." figureDoi-0="http://doi.org/10.5281/zenodo.6916902" figureDoi-1="http://doi.org/10.5281/zenodo.6916904" figureDoi-2="http://doi.org/10.5281/zenodo.6916906" figureDoi-3="http://doi.org/10.5281/zenodo.6916908" httpUri-0="https://zenodo.org/record/6916902/files/figure.png" httpUri-1="https://zenodo.org/record/6916904/files/figure.png" httpUri-2="https://zenodo.org/record/6916906/files/figure.png" httpUri-3="https://zenodo.org/record/6916908/files/figure.png" pageId="15" pageNumber="16">Figs21–24</figureCitation>
|
||
). Cephalothorax is similar to
|
||
<taxonomicName id="53EA20FB9B54FFEDFEE35B9DFCE052E1" authority="Nadolny & Zamani, 2020" authorityName="Nadolny & Zamani" authorityYear="2020" box="[328,893,1193,1219]" class="Arachnida" family="Lycosidae" genus="Lycosa" kingdom="Animalia" order="Araneae" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="macrophthalma">
|
||
<emphasis id="A69E876A9B54FFEDFEE35B9DFDCF52E1" box="[328,594,1193,1219]" italics="true" pageId="15" pageNumber="16">Lycosa macrophthalma</emphasis>
|
||
<bibRefCitation id="F07B26899B54FFEDFDF25B9DFCE052E1" author="Nadolny A. A. & Zamani A." box="[601,893,1193,1219]" pageId="15" pageNumber="16" pagination="205 - 212" refId="ref31572" refString="Nadolny A. A. & Zamani A. 2020. A new species of wolf spiders of the genus Lycosa (Aranei: Lycosidae) from Iran. Zoosystematica Rossica 29 (2): 205 - 212. https: // doi. org / 10.31610 / zsr / 2020.29.2.205" type="journal article" year="2020">Nadolny & Zamani, 2020</bibRefCitation>
|
||
</taxonomicName>
|
||
and it might be related to it as well.
|
||
</paragraph>
|
||
<caption id="C0950BF09B54FFEDFF165869FDA351C2" ID-DOI="http://doi.org/10.5281/zenodo.6916860" ID-Zenodo-Dep="6916860" httpUri="https://zenodo.org/record/6916860/files/figure.png" pageId="15" pageNumber="16" startId="15.[189,232,1885,1911]" targetBox="[211,1382,1365,1839]" targetPageId="15">
|
||
<paragraph id="94555B789B54FFEDFF165869FDA351C2" blockId="15.[189,1401,1885,2017]" pageId="15" pageNumber="16">
|
||
<emphasis id="A69E876A9B54FFEDFF165869FE965155" bold="true" box="[189,267,1885,1911]" pageId="15" pageNumber="16">Fig. 4.</emphasis>
|
||
Male pedipalps retrolateral view.
|
||
<emphasis id="A69E876A9B54FFEDFD3C5869FD315155" bold="true" box="[663,684,1885,1911]" pageId="15" pageNumber="16">A</emphasis>
|
||
.
|
||
<taxonomicName id="53EA20FB9B54FFEDFD105869FBF65155" authority="C. L. Koch, 1838" authorityName="C. L. Koch" authorityYear="1838" box="[699,1131,1885,1911]" class="Arachnida" family="Lycosidae" genus="Geolycosa" kingdom="Animalia" order="Araneae" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="vultuosa">
|
||
<emphasis id="A69E876A9B54FFEDFD105869FC3C5155" box="[699,929,1885,1911]" italics="true" pageId="15" pageNumber="16">Geolycosa vultuosa</emphasis>
|
||
C.L.
|
||
<bibRefCitation id="F07B26899B54FFEDFC485869FBF65155" author="Koch C. L." box="[995,1131,1885,1911]" pageId="15" pageNumber="16" refId="ref31028" refString="Koch C. L. 1838. Die Arachniden. C. H. Zeh'sche Buchhandlung, Nurnberg. https: // doi. org / 10.5962 / bhl. title. 43744" type="book" year="1838">Koch, 1838</bibRefCitation>
|
||
</taxonomicName>
|
||
(HUJ INV-AR20960).
|
||
<emphasis id="A69E876A9B54FFEDFF1658B4FF4C51B8" bold="true" box="[189,209,1920,1946]" pageId="15" pageNumber="16">B</emphasis>
|
||
.
|
||
<taxonomicName id="53EA20FB9B54FFEDFF4858B5FE0651B8" authority="Steinpress & Cohen & Pétillon & Chipman & Gavish-Regev, 2022" authorityName="Steinpress & Cohen & Pétillon & Chipman & Gavish-Regev" authorityYear="2022" box="[227,411,1921,1946]" class="Arachnida" family="Lycosidae" genus="Lycosa" kingdom="Animalia" order="Araneae" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="gesserit" status="sp. nov.">
|
||
<emphasis id="A69E876A9B54FFEDFF4858B5FE0651B8" box="[227,411,1921,1946]" italics="true" pageId="15" pageNumber="16">Lycosa gesserit</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="BDAD3A119B54FFEDFE0D58B5FD9851B8" box="[422,517,1921,1947]" pageId="15" pageNumber="16" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, holotype (HUJ INV-AR20631).
|
||
<emphasis id="A69E876A9B54FFEDFC3658B4FC2F51B8" bold="true" box="[925,946,1920,1946]" pageId="15" pageNumber="16">C</emphasis>
|
||
.
|
||
<taxonomicName id="53EA20FB9B54FFEDFC6F58B5FB0851B8" authority="Steinpress & Cohen & Pétillon & Chipman & Gavish-Regev, 2022" authorityName="Steinpress & Cohen & Pétillon & Chipman & Gavish-Regev" authorityYear="2022" box="[964,1173,1920,1946]" class="Arachnida" family="Lycosidae" genus="Lycosa" kingdom="Animalia" order="Araneae" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="hyraculus" status="sp. nov.">
|
||
<emphasis id="A69E876A9B54FFEDFC6F58B5FB0851B8" box="[964,1173,1920,1946]" italics="true" pageId="15" pageNumber="16">Lycosa hyraculus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="BDAD3A119B54FFEDFB0B58B5FA9D51B9" box="[1184,1280,1921,1947]" pageId="15" pageNumber="16" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, holotype (HUJ INV-AR20818).
|
||
<emphasis id="A69E876A9B54FFEDFE665897FE7F519F" bold="true" box="[461,482,1955,1981]" pageId="15" pageNumber="16">D</emphasis>
|
||
.
|
||
<taxonomicName id="53EA20FB9B54FFEDFE595890FCFC519F" authority="Simon, 1876" authorityName="Simon" authorityYear="1876" box="[498,865,1955,1981]" class="Arachnida" family="Lycosidae" genus="Lycosa" kingdom="Animalia" order="Araneae" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="piochardi">
|
||
<emphasis id="A69E876A9B54FFEDFE595890FD22519F" box="[498,703,1955,1981]" italics="true" pageId="15" pageNumber="16">Lycosa piochardi</emphasis>
|
||
<bibRefCitation id="F07B26899B54FFEDFD625897FCFC519F" author="Simon E." box="[713,865,1955,1981]" pageId="15" pageNumber="16" pagination="57 - 91" refId="ref32282" refString="Simon E. 1876. Etudes arachnologiques. 4 e memoire. VII. Revision des especes europeennes du groupe de la Lycosa tarentula Rossi. Annales de la Societe entomologique de France (5) 6: 57 - 91." type="journal article" year="1876">Simon, 1876</bibRefCitation>
|
||
</taxonomicName>
|
||
(HUJ INV-AR20948). Scale bars = 0.5 mm. Photos by I. Armiach Steinpress.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="94555B789B4BFFF2FF165E3DFD7D578F" blockId="16.[188,1399,265,430]" pageId="16" pageNumber="17">
|
||
The single known male specimen was raised in the laboratory for two molts. This has probably affected the size of the adult spider. Even though we have observed an aberrant morphology in the genitalia of females grown in lab conditions, we have not observed the same in males, either of
|
||
<taxonomicName id="53EA20FB9B4BFFF2FB0E5E64FAAB574B" authorityName="Simon" authorityYear="1876" box="[1189,1334,335,361]" class="Arachnida" family="Lycosidae" genus="Lycosa" kingdom="Animalia" order="Araneae" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="piochardi">
|
||
<emphasis id="A69E876A9B4BFFF2FB0E5E64FAAB574B" box="[1189,1334,335,361]" italics="true" pageId="16" pageNumber="17">L. piochardi</emphasis>
|
||
</taxonomicName>
|
||
or of
|
||
<taxonomicName id="53EA20FB9B4BFFF2FF165E46FED257A9" authority="Steinpress & Cohen & Pétillon & Chipman & Gavish-Regev, 2022" authorityName="Steinpress & Cohen & Pétillon & Chipman & Gavish-Regev" authorityYear="2022" box="[189,335,369,395]" class="Arachnida" family="Lycosidae" genus="Lycosa" kingdom="Animalia" order="Araneae" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="hyraculus" status="sp. nov.">
|
||
<emphasis id="A69E876A9B4BFFF2FF165E46FED257A9" box="[189,335,369,395]" italics="true" pageId="16" pageNumber="17">L. hyraculus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="BDAD3A119B4BFFF2FEFE5E46FE3357AE" box="[341,430,370,396]" pageId="16" pageNumber="17" rank="species">sp. nov.</taxonomicNameLabel>
|
||
We consider it highly probable that the diagnostic characters in the male’s pedipalp were not affected by the laboratory conditions.
|
||
</paragraph>
|
||
<paragraph id="94555B789B4BFFF2FF165EE0FAEF5417" blockId="16.[189,1399,468,565]" pageId="16" pageNumber="17">
|
||
Although genitalia of other males of
|
||
<taxonomicName id="53EA20FB9B4BFFF2FD2C5EE1FD4457CC" authorityName="Latreille" authorityYear="1804" box="[647,729,469,494]" class="Arachnida" family="Lycosidae" genus="Lycosa" kingdom="Animalia" order="Araneae" pageId="16" pageNumber="17" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="A69E876A9B4BFFF2FD2C5EE1FD4457CC" box="[647,729,469,494]" italics="true" pageId="16" pageNumber="17">Lycosa</emphasis>
|
||
</taxonomicName>
|
||
grown in our laboratory did not show distortions (see discussion about distortions of female genitalia), we cannot dismiss the possibility that the pedipalps of specimens in the wild are somewhat different to the
|
||
<typeStatus id="4B51E5DA9B4BFFF2FC895D2FFCC85417" box="[802,853,539,565]" pageId="16" pageNumber="17">type</typeStatus>
|
||
specimen, which was raised in the laboratory.
|
||
</paragraph>
|
||
<paragraph id="94555B789B4BFFF2FF165D68FE9E5454" blockId="16.[189,259,604,630]" box="[189,259,604,630]" pageId="16" pageNumber="17">
|
||
<heading id="CF1DEC149B4BFFF2FF165D68FE9E5454" bold="true" box="[189,259,604,630]" fontSize="11" level="3" pageId="16" pageNumber="17" reason="3">
|
||
<emphasis id="A69E876A9B4BFFF2FF165D68FE9E5454" bold="true" box="[189,259,604,630]" pageId="16" pageNumber="17">Notes</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="94555B789B4BFFF3FF165DBFFDAA5764" blockId="16.[189,1398,651,713]" lastBlockId="17.[189,1399,265,326]" lastPageId="17" lastPageNumber="18" pageId="16" pageNumber="17">
|
||
The locality where the
|
||
<typeStatus id="4B51E5DA9B4BFFF2FE675DBFFE625487" box="[460,511,651,677]" pageId="16" pageNumber="17">type</typeStatus>
|
||
was found is a remote and not easily reachable nature reserve, encircled by army training areas, that are closed to the public most of the year. There are only few occasions every year that access to the public is possible. As of the writing of this paper, we have not succeeded in collecting additional specimens.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |