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<mods:title id="08A3B63A67D1718CC08FB302A289A2F9">Bucardium grateloupianum n. sp. from the Lower Miocene of Aquitaine with taxonomic comments on some fossil species from Europe (Bivalvia, Cardiidae)</mods:title>
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<treatment id="1C20AE74FF9C5B41F9E1B925BB5E3495" ID-DOI="http://doi.org/10.5281/zenodo.5673932" ID-GBIF-Taxon="124788224" ID-Zenodo-Dep="5673932" LSID="urn:lsid:plazi:treatment:1C20AE74FF9C5B41F9E1B925BB5E3495" httpUri="http://treatment.plazi.org/id/1C20AE74FF9C5B41F9E1B925BB5E3495" lastPageId="9" lastPageNumber="577" pageId="2" pageNumber="570">
<subSubSection id="DC934CE9FF9C5B4AF9E1B925B82732A1" pageId="2" pageNumber="570" type="nomenclature">
<paragraph id="94361F62FF9C5B4AF9E1B925BB653280" blockId="2.[151,562,1905,1965]" box="[151,562,1905,1932]" pageId="2" pageNumber="570">
<heading id="CF7EA80EFF9C5B4AF9E1B925BB653280" bold="true" box="[151,562,1905,1932]" fontSize="11" level="1" pageId="2" pageNumber="570" reason="1">
<emphasis id="A6FDC370FF9C5B4AF9E1B925BB653280" bold="true" box="[151,562,1905,1932]" pageId="2" pageNumber="570">
<taxonomicName id="538964E1FF9C5B4AF9E1B925B8BD3287" box="[151,490,1905,1931]" class="Bivalvia" family="Cardiidae" genus="Bucardium" kingdom="Animalia" order="Veneroida" pageId="2" pageNumber="570" phylum="Mollusca" rank="species" species="grateloupianum" status="sp. nov.">
<emphasis id="A6FDC370FF9C5B4AF9E1B925B8BD3287" bold="true" box="[151,490,1905,1931]" italics="true" pageId="2" pageNumber="570">Bucardium grateloupianum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="BDCE7E0BFF9C5B4AF886B926BB653280" box="[496,562,1906,1932]" pageId="2" pageNumber="570" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="94361F62FF9C5B4AF9E1B9C0B82732A1" blockId="2.[151,562,1905,1965]" box="[151,368,1940,1965]" pageId="2" pageNumber="570">
(
<figureCitation id="0CB203E7FF9C5B4AF9E9B9C0B9B232A1" box="[159,229,1940,1965]" captionStart="FIGURE 2" captionStartId="4.[151,250,1938,1960]" captionTargetBox="[151,1436,337,1915]" captionTargetId="figure@4.[151,1436,337,1917]" captionTargetPageId="4" captionText="FIGURE 2. Bucardium grateloupianum n. sp. A C. Holotype, St-Paul-lès-Dax, RV (MNHN Sciences de la Terre A 13845, d'Orbigny coll. 10986), photo J. J. ter Poorten, with permission. D F, K. Paratype 1, Mérignac, LF (MZB 60206). G J. Paratype 2, Mérignac, RV (MZB 60207). L. Mérignac, RV (ter Poorten coll. TP 1310), photo J. J. ter Poorten, with permission." httpUri="https://zenodo.org/record/272521/files/figure.png" pageId="2" pageNumber="570">Figs 2</figureCitation>
AL, 3AF)
</paragraph>
</subSubSection>
<subSubSection id="DC934CE9FF9C5B4BF9E1B988BB8835A2" lastPageId="3" lastPageNumber="571" pageId="2" pageNumber="570" type="reference_group">
<paragraph id="94361F62FF9C5B4AF9E1B988BDE132FE" blockId="2.[151,1206,2012,2034]" box="[151,1206,2012,2034]" pageId="2" pageNumber="570">
1912
<taxonomicName id="538964E1FF9C5B4AF9A6B988BB9432FE" authority="Mayer, 1866" authorityName="Mayer" authorityYear="1866" box="[208,707,2012,2034]" class="Bivalvia" family="Cardiidae" genus="Cardium" kingdom="Animalia" order="Veneroida" pageId="2" pageNumber="570" phylum="Mollusca" rank="species" species="grateloupi" subGenus="Ringicardium">
<emphasis id="A6FDC370FF9C5B4AF9A6B988BB6D32FE" box="[208,570,2012,2034]" italics="true" pageId="2" pageNumber="570">Cardium (Ringicardium) grateloupi</emphasis>
<bibRefCitation id="F0186293FF9C5B4AFB36B989BB9432FE" author="Mayer" box="[576,707,2012,2034]" pageId="2" pageNumber="570" refString="Mayer, M. C. (1866) Description de coquilles fossiles des terrains tertiaires superieurs (suite). Journal de Conchyliologie, 14, 67 - 76." type="journal article" year="1866">Mayer, 1866</bibRefCitation>
</taxonomicName>
,
<emphasis id="A6FDC370FF9C5B4AFBBBB989BA5232FE" box="[717,773,2013,2034]" italics="true" pageId="2" pageNumber="570">sensu</emphasis>
Cossmann &amp; Peyrot, 115, pl. 22, figs 36.
</paragraph>
<paragraph id="94361F62FF9D5B4BF9E1BECCBB8835A2" blockId="3.[151,1323,152,450]" box="[151,735,152,174]" pageId="3" pageNumber="571">
1950
<taxonomicName id="538964E1FF9D5B4BF9A6BECCBB5335A2" authority="Magne" authorityName="Magne" box="[208,516,152,174]" class="Bivalvia" family="Cardiidae" genus="Ringicardium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Veneroida" pageId="3" pageNumber="571" phylum="Mollusca" rank="species" species="aquense">
<emphasis id="A6FDC370FF9D5B4BF9A6BECCB8E435A2" box="[208,435,152,174]" italics="true" pageId="3" pageNumber="571">Ringicardium aquense</emphasis>
Magne
</taxonomicName>
: 122 (
<emphasis id="A6FDC370FF9D5B4BFB34BECDBB8635A2" box="[578,721,152,174]" italics="true" pageId="3" pageNumber="571">nomen nudum</emphasis>
).
</paragraph>
</subSubSection>
<subSubSection id="DC934CE9FF9D5B4BF9E1BEE2B9EA35E7" pageId="3" pageNumber="571" type="description">
<paragraph id="94361F62FF9D5B4BF9E1BEE2BA7535C0" blockId="3.[151,1323,152,450]" box="[151,802,182,204]" pageId="3" pageNumber="571">
2005
<taxonomicName id="538964E1FF9D5B4BF9A6BEE2B8CE35C0" box="[208,409,182,204]" class="Bivalvia" family="Cardiidae" genus="Cardium" kingdom="Animalia" order="Veneroida" pageId="3" pageNumber="571" phylum="Mollusca" rank="species" species="grateloupi">
<emphasis id="A6FDC370FF9D5B4BF9A6BEE2B8CE35C0" box="[208,409,182,204]" italics="true" pageId="3" pageNumber="571">Cardium grateloupi</emphasis>
</taxonomicName>
—Lesport &amp; Cahuzac, pl. 6, figs 9, 19.
</paragraph>
<paragraph id="94361F62FF9D5B4BF9E1BE82B9EA35E7" blockId="3.[151,1323,152,450]" box="[151,189,214,235]" pageId="3" pageNumber="571">non</paragraph>
</subSubSection>
<subSubSection id="DC934CE9FF9D5B4BF9E1BEA0BAF134CE" pageId="3" pageNumber="571" type="reference_group">
<paragraph id="94361F62FF9D5B4BF9E1BEA0BADE3488" blockId="3.[151,1323,152,450]" pageId="3" pageNumber="571">
1839
<taxonomicName id="538964E1FF9D5B4BF9A6BEA0BB1F3406" authority="Michelotti, 1839" authorityName="Michelotti" authorityYear="1839" box="[208,584,244,266]" class="Bivalvia" family="Cardiidae" genus="Cardium" kingdom="Animalia" order="Veneroida" pageId="3" pageNumber="571" phylum="Mollusca" rank="species" species="grateloupi">
<emphasis id="A6FDC370FF9D5B4BF9A6BEA0B8CE3406" box="[208,409,244,266]" italics="true" pageId="3" pageNumber="571">Cardium grateloupi</emphasis>
<bibRefCitation id="F0186293FF9D5B4BF8E9BEA0BB1F3406" author="Michelotti" box="[415,584,244,266]" pageId="3" pageNumber="571" refString="Michelotti, G. (1839). Brevi cenni di alcuni resti delle classi Brachiopodi ed Acefali, trovati fossili in Italia. Annali delle Scienze del Regno Lombardo-Veneto, 9, 119 - 174." type="journal article" year="1839">Michelotti, 1839</bibRefCitation>
</taxonomicName>
=
<taxonomicName id="538964E1FF9D5B4BFB15BEA1BA2D3406" authority="Brocchi, 1814" authorityName="Brocchi" authorityYear="1814" box="[611,890,244,266]" class="Bivalvia" family="Veneridae" genus="Venus" kingdom="Animalia" order="Veneroida" pageId="3" pageNumber="571" phylum="Mollusca" rank="species" species="cypria">
<emphasis id="A6FDC370FF9D5B4BFB15BEA1BBB23406" box="[611,741,245,266]" italics="true" pageId="3" pageNumber="571">Venus cypria</emphasis>
<bibRefCitation id="F0186293FF9D5B4BFB9DBEA0BA2D3406" author="Brocchi" box="[747,890,244,266]" pageId="3" pageNumber="571" refString="Brocchi, G. (1814) Conchiologia fossile subappennina con osservazioni geologiche sugli Appennini e sul suolo adiacente. Vol. 1 &amp; 2. Stamperia Reale, Milano, 472 pp. http: // dx. doi. org / 10.5962 / bhl. title. 11569" type="book" year="1814">Brocchi, 1814</bibRefCitation>
</taxonomicName>
=
<taxonomicName id="538964E1FF9D5B4BFAE3BEA0BC713406" authority="Brocchi, 1814" authorityName="Brocchi" authorityYear="1814" box="[917,1318,244,266]" class="Bivalvia" family="Cardiidae" genus="Nemocardium" kingdom="Animalia" order="Veneroida" pageId="3" pageNumber="571" phylum="Mollusca" rank="species" species="cyprium">
<emphasis id="A6FDC370FF9D5B4BFAE3BEA0BD2A3406" box="[917,1149,244,266]" italics="true" pageId="3" pageNumber="571">Nemocardium cyprium</emphasis>
(
<bibRefCitation id="F0186293FF9D5B4BFDFCBEA0BC773406" author="Brocchi" box="[1162,1312,244,266]" pageId="3" pageNumber="571" refString="Brocchi, G. (1814) Conchiologia fossile subappennina con osservazioni geologiche sugli Appennini e sul suolo adiacente. Vol. 1 &amp; 2. Stamperia Reale, Milano, 472 pp. http: // dx. doi. org / 10.5962 / bhl. title. 11569" type="book" year="1814">Brocchi, 1814</bibRefCitation>
)
</taxonomicName>
. 1862
<taxonomicName id="538964E1FF9D5B4BF9A6BF46BB843424" authority="Grateloup in Hornes, 1862" authorityName="Grateloup in Hornes" authorityYear="1862" box="[208,723,274,296]" class="Bivalvia" family="Cardiidae" genus="Cardium" kingdom="Animalia" order="Veneroida" pageId="3" pageNumber="571" phylum="Mollusca" rank="species" species="hoernesianum">
<emphasis id="A6FDC370FF9D5B4BF9A6BF46B8E93424" box="[208,446,274,296]" italics="true" pageId="3" pageNumber="571">Cardium hoernesianum</emphasis>
Grateloup in
<bibRefCitation id="F0186293FF9D5B4BFB3DBF47BB843424" author="Hornes" box="[587,723,274,296]" pageId="3" pageNumber="571" refString="Hornes, M. (1862) Die fossilen Mollusken des Tertiarbeckens von Wien. Band II: Bivalvia. Abhandlungen der Kaiserlich- Koniglichen Geologischen Reichsanstalt, 4, 117 - 214." type="journal article" year="1862">Hörnes, 1862</bibRefCitation>
</taxonomicName>
=
<taxonomicName id="538964E1FF9D5B4BFB98BF46BD383424" authority="Hornes, 1862" authorityName="Hornes" authorityYear="1862" box="[750,1135,274,296]" class="Bivalvia" family="Cardiidae" genus="Cardium" kingdom="Animalia" order="Veneroida" pageId="3" pageNumber="571" phylum="Mollusca" rank="species" species="hoernesianum">
<emphasis id="A6FDC370FF9D5B4BFB98BF46BA8B3424" box="[750,988,274,296]" italics="true" pageId="3" pageNumber="571">Cardium hoernesianum</emphasis>
<bibRefCitation id="F0186293FF9D5B4BFA95BF47BD383424" author="Hornes" box="[995,1135,274,296]" pageId="3" pageNumber="571" refString="Hornes, M. (1862) Die fossilen Mollusken des Tertiarbeckens von Wien. Band II: Bivalvia. Abhandlungen der Kaiserlich- Koniglichen Geologischen Reichsanstalt, 4, 117 - 214." type="journal article" year="1862">Hörnes, 1862</bibRefCitation>
</taxonomicName>
. 1866
<taxonomicName id="538964E1FF9D5B4BF9A6BF65BB48344B" authority="Mayer, 1866" authorityName="Mayer" authorityYear="1866" box="[208,543,305,327]" class="Bivalvia" family="Cardiidae" genus="Cardium" kingdom="Animalia" order="Veneroida" pageId="3" pageNumber="571" phylum="Mollusca" rank="species" species="grateloupi">
<emphasis id="A6FDC370FF9D5B4BF9A6BF65B8CE344B" box="[208,409,305,327]" italics="true" pageId="3" pageNumber="571">Cardium grateloupi</emphasis>
<bibRefCitation id="F0186293FF9D5B4BF8E9BF66BB48344B" author="Mayer" box="[415,543,305,327]" pageId="3" pageNumber="571" refString="Mayer, M. C. (1866) Description de coquilles fossiles des terrains tertiaires superieurs (suite). Journal de Conchyliologie, 14, 67 - 76." type="journal article" year="1866">Mayer, 1866</bibRefCitation>
</taxonomicName>
=
<taxonomicName id="538964E1FF9D5B4BFB4CBF65BAEC344B" authority="Hornes, 1862" authorityName="Hornes" authorityYear="1862" box="[570,955,305,327]" class="Bivalvia" family="Cardiidae" genus="Cardium" kingdom="Animalia" order="Veneroida" pageId="3" pageNumber="571" phylum="Mollusca" rank="species" species="hoernesianum">
<emphasis id="A6FDC370FF9D5B4BFB4CBF65BA7E344B" box="[570,809,305,327]" italics="true" pageId="3" pageNumber="571">Cardium hoernesianum</emphasis>
<bibRefCitation id="F0186293FF9D5B4BFA59BF66BAEC344B" author="Hornes" box="[815,955,305,327]" pageId="3" pageNumber="571" refString="Hornes, M. (1862) Die fossilen Mollusken des Tertiarbeckens von Wien. Band II: Bivalvia. Abhandlungen der Kaiserlich- Koniglichen Geologischen Reichsanstalt, 4, 117 - 214." type="journal article" year="1862">Hörnes, 1862</bibRefCitation>
</taxonomicName>
. 1910
<taxonomicName id="538964E1FF9D5B4BF9A6BF04BBD6346A" authority="Hornes, 1862" authorityName="Hornes" authorityYear="1862" box="[208,641,336,358]" class="Bivalvia" family="Cardiidae" genus="Ringicardium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Veneroida" pageId="3" pageNumber="571" phylum="Mollusca" rank="species" species="hoernesianum">
<emphasis id="A6FDC370FF9D5B4BF9A6BF04B8B8346A" box="[208,495,336,358]" italics="true" pageId="3" pageNumber="571">Ringicardium hoernesianum</emphasis>
<bibRefCitation id="F0186293FF9D5B4BF880BF05BBD6346A" author="Hornes" box="[502,641,336,358]" pageId="3" pageNumber="571" refString="Hornes, M. (1862) Die fossilen Mollusken des Tertiarbeckens von Wien. Band II: Bivalvia. Abhandlungen der Kaiserlich- Koniglichen Geologischen Reichsanstalt, 4, 117 - 214." type="journal article" year="1862">Hörnes, 1862</bibRefCitation>
</taxonomicName>
,
<emphasis id="A6FDC370FF9D5B4BFBFDBF05BB94346A" box="[651,707,337,358]" italics="true" pageId="3" pageNumber="571">sensu</emphasis>
<bibRefCitation id="F0186293FF9D5B4BFBBCBF04BA36346A" author="Schaffer" box="[714,865,336,358]" pageId="3" pageNumber="571" refString="Schaffer, F. X. (1910) Die Bivalven der Miocanbildungen von Eggenburg. In: Das Miocan von Eggenburg. Die Fauna der erstern Mediterranstufe des Wiener Beckens und die geologischen Verhaltnisse der Umgebung des Manhartsberges in Niederosterreich. Abhandlungen der Kaiserlich-Koniglichen Geologischen Reichsanstalt, 22, 1 - 112." type="journal article" year="1910">Schaffer, 1910</bibRefCitation>
: 66, pl. 32, figs 1, 2. 1910
<taxonomicName id="538964E1FF9D5B4BF9A6BF3ABB8F3488" authority="Schaffer" authorityName="Schaffer" box="[208,728,366,388]" class="Bivalvia" family="Cardiidae" genus="Ringicardium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Veneroida" pageId="3" pageNumber="571" phylum="Mollusca" rank="variety" species="hoernesianum" variety="elongata">
<emphasis id="A6FDC370FF9D5B4BF9A6BF3AB8B83488" box="[208,495,366,388]" italics="true" pageId="3" pageNumber="571">Ringicardium hoernesianum</emphasis>
var.
<emphasis id="A6FDC370FF9D5B4BFB54BF3ABB2C3488" box="[546,635,366,388]" italics="true" pageId="3" pageNumber="571">elongata</emphasis>
Schaffer
</taxonomicName>
, 67, pl. 32, fig. 3.
</paragraph>
<paragraph id="94361F62FF9D5B4BF9E1BFD9BAF134CE" blockId="3.[151,1323,152,450]" pageId="3" pageNumber="571">
<treatmentCitationGroup id="B499384CFF9D5B4BF9E1BFD9BAF134CE" pageId="3" pageNumber="571">
1936
<taxonomicName id="538964E1FF9D5B4BF9A6BFD9BACB34AF" authority="Noszky, 1936: 88" authorityName="Noszky" authorityPageNumber="88" authorityYear="1936" box="[208,924,397,419]" class="Bivalvia" family="Cardiidae" genus="Cardium" kingdom="Animalia" order="Veneroida" pageId="3" pageNumber="571" phylum="Mollusca" rank="variety" species="grateloupi" subGenus="Ringicardium" variety="oligocenica">
<emphasis id="A6FDC370FF9D5B4BF9A6BFD9BB6D34AF" box="[208,570,397,419]" italics="true" pageId="3" pageNumber="571">Cardium (Ringicardium) grateloupi</emphasis>
var.
<emphasis id="A6FDC370FF9D5B4BFB1ABFD9BBB534AF" box="[620,738,397,419]" italics="true" pageId="3" pageNumber="571">oligocenica</emphasis>
<treatmentCitation id="15283973FF9D5B4BFB9EBFD9BACB34AF" author="Noszky" box="[744,924,397,419]" page="88" pageId="3" pageNumber="571" year="1936">
<bibRefCitation id="F0186293FF9D5B4BFB9EBFD9BA2034AF" author="Noszky" box="[744,887,397,419]" pageId="3" pageNumber="571" refString="Noszky, J. (1936) Az egri felso cattien molluszkafaunaja. Die Molluskenfauna des oberen Cattiens von Eger, in Ungarn. Annales Musei Nationalis Hungarici. Pars Mineralogica, Geologica, Paleontologica, 30, 53 - 115." type="journal article" year="1936">Noszky, 1936</bibRefCitation>
: 88
</treatmentCitation>
</taxonomicName>
, 111, pl. 6, fig. 7. 2003
<taxonomicName id="538964E1FF9D5B4BF9A6BFF8BB0734CE" authority="Hornes, 1862" authorityName="Hornes" authorityYear="1862" box="[208,592,428,450]" class="Bivalvia" family="Cardiidae" genus="Cardium" kingdom="Animalia" order="Veneroida" pageId="3" pageNumber="571" phylum="Mollusca" rank="species" species="hoernesianum">
<emphasis id="A6FDC370FF9D5B4BF9A6BFF8B8E934CE" box="[208,446,428,450]" italics="true" pageId="3" pageNumber="571">Cardium hoernesianum</emphasis>
<bibRefCitation id="F0186293FF9D5B4BF8B3BFF9BB0734CE" author="Hornes" box="[453,592,428,450]" pageId="3" pageNumber="571" refString="Hornes, M. (1862) Die fossilen Mollusken des Tertiarbeckens von Wien. Band II: Bivalvia. Abhandlungen der Kaiserlich- Koniglichen Geologischen Reichsanstalt, 4, 117 - 214." type="journal article" year="1862">Hörnes, 1862</bibRefCitation>
</taxonomicName>
,
<emphasis id="A6FDC370FF9D5B4BFB2CBFF9BBC534CE" box="[602,658,429,450]" italics="true" pageId="3" pageNumber="571">sensu</emphasis>
Schultz: 517, pl. 75, fig. 1.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="DC934CE9FF9D5B4BF9E1BFBABC273727" pageId="3" pageNumber="571" type="materials_examined">
<paragraph id="94361F62FF9D5B4BF9E1BFBABC273727" blockId="3.[151,1436,494,2031]" pageId="3" pageNumber="571">
<emphasis id="A6FDC370FF9D5B4BF9E1BFBAB810370B" bold="true" box="[151,327,494,519]" pageId="3" pageNumber="571">Type material.</emphasis>
Holotype, RV, Burdigalian of St-Paul-lès-Dax (MNHN Sciences de la Terre A13845, d'Orbigny coll. 10986); Paratype 1, RV, Burdigalian of Mérignac (MZB 60206); Paratype 2, LV, Mérignac (MZB 60207).
</paragraph>
</subSubSection>
<subSubSection id="DC934CE9FF9D5B4BF9B1BC62BC503012" pageId="3" pageNumber="571" type="description">
<paragraph id="94361F62FF9D5B4BF9B1BC62B9B8360F" blockId="3.[151,1436,494,2031]" pageId="3" pageNumber="571">
<emphasis id="A6FDC370FF9D5B4BF9B1BC62B80E3743" bold="true" box="[199,345,566,591]" pageId="3" pageNumber="571">Description.</emphasis>
Shell moderately large, robust and convex; poorly elongate, inequilateral, slightly gaping. Umbo median, moderately large, barely prosogyrate. Antero-dorsal margin long, straight to slightly convex, barely sloping. Posterior margin moderately convex, ventral margin strongly convex, posterior margin consisting of some, usually six, rather strong, well developed, sharply ending digitations, dorsalward increasing in strength, formed by prolongations of posterior ribs; postero-dorsal margin short. Radial sculpture moderately robust, covering all shell surface.
</paragraph>
<paragraph id="94361F62FF9D5B4BF9B1BD5AB8A836BB" blockId="3.[151,1436,494,2031]" pageId="3" pageNumber="571">Sculpture on antero-dorsal area consisting of seven or eight fine ribs, symmetrically triangular in cross-section, with interspaces of similar width. Following sculpture much stronger, of about 13 wide, convex ribs separated by narrow and flat interspaces; rib cross-section slightly asymmetrical, with wider, convex anterior side and narrower, steeper posterior side, then becoming almost symmetrical and flatter medially, and again markedly aymmetrical up to the postero-ventral transition.</paragraph>
<paragraph id="94361F62FF9D5B4BF9B1BD96BCD8312F" blockId="3.[151,1436,494,2031]" pageId="3" pageNumber="571">Six posterior ribs flattish, strongly asymmetrical in cross-section, each rib with a thin, continuous laminar ridge on its posterior side; ridges finely and irregularly wavy; interspaces very narrow, deep. Commarginal sculpture consisting of fairly distinct growth striae, crossing ribs and interspaces, slightly crescent-shaped on posterior ribs.</paragraph>
<paragraph id="94361F62FF9D5B4BF9B1BA7ABCDA31F7" blockId="3.[151,1436,494,2031]" pageId="3" pageNumber="571">Internal margin crenulate and serrate, finely antero-dorsally, in accordance with external ribbing. Adductor muscle scars ovate, of similar size; pallial line entire, poorly distinct. Hinge moderately strong, rather straight, with well developed lunular flap. Right valve with a deeply bifid cardinal tooth, two anterior triangular, elongate lateral teeth, dorsal one much weaker; one strong, posterior lateral tooth. Left valve with a cardinal, deeply bifid tooth, two anterior triangular, lateral teeth, dorsal one stronger; two posterior triangular, elongate posterior teeth, dorsal one much stronger. Lunule lanceolate, moderately deep, smooth; escutcheon small, mostly occupied by ligament.</paragraph>
<paragraph id="94361F62FF9D5B4BF9B1BB52BC503012" blockId="3.[151,1436,494,2031]" box="[199,1287,1286,1311]" pageId="3" pageNumber="571">
Holotype: L
<quantity id="5371B287FF9D5B4BF820BB52B8F03012" box="[342,423,1286,1311]" metricMagnitude="-2" metricUnit="m" metricValue="6.1" pageId="3" pageNumber="571" unit="mm" value="61.0">61 mm</quantity>
, H
<quantity id="5371B287FF9D5B4BF8B8BB52BB773012" box="[462,544,1286,1311]" metricMagnitude="-2" metricUnit="m" metricValue="5.3" pageId="3" pageNumber="571" unit="mm" value="53.0">53 mm</quantity>
; paratype 1: L
<quantity id="5371B287FF9D5B4BFBBEBB52BA4F3012" box="[712,792,1286,1311]" metricMagnitude="-2" metricUnit="m" metricValue="6.5" pageId="3" pageNumber="571" unit="mm" value="65.0">65 mm</quantity>
, H
<quantity id="5371B287FF9D5B4BFA49BB52BAC63012" box="[831,913,1286,1311]" metricMagnitude="-2" metricUnit="m" metricValue="5.6" pageId="3" pageNumber="571" unit="mm" value="56.0">56 mm</quantity>
; paratype 2: L
<quantity id="5371B287FF9D5B4BFD4FBB53BDDD3012" box="[1081,1162,1287,1311]" metricMagnitude="-2" metricUnit="m" metricValue="4.3" pageId="3" pageNumber="571" unit="mm" value="43.0">43 mm</quantity>
, H
<quantity id="5371B287FF9D5B4BFDC7BB52BC543012" box="[1201,1283,1286,1311]" metricMagnitude="-2" metricUnit="m" metricValue="3.6" pageId="3" pageNumber="571" unit="mm" value="36.0">36 mm</quantity>
.
</paragraph>
</subSubSection>
<subSubSection id="DC934CE9FF9D5B4BF9B1BB7EBD2B3333" pageId="3" pageNumber="571" type="materials_examined">
<paragraph id="94361F62FF9D5B4BF9B1BB7EBBA330DF" blockId="3.[151,1436,494,2031]" pageId="3" pageNumber="571">
<materialsCitation id="24E1153FFF9D5B4BF9B1BB7EBBA330DF" ID-GBIF-Occurrence="1328377998" collectionCode="MNHN" collectorName="Landes, SW &amp; d'Orbigny" country="France" location="France" pageId="3" pageNumber="571" specimenCount="3" typeStatus="holotype">
<emphasis id="A6FDC370FF9D5B4BF9B1BB7EB83D304F" bold="true" box="[199,362,1322,1347]" pageId="3" pageNumber="571">
<typeStatus id="4B32A1C0FF9D5B4BF9B1BB7EB855304F" box="[199,258,1322,1347]" pageId="3" pageNumber="571">Type</typeStatus>
locality.
</emphasis>
Saint-Paul-lès-Dax,
<collectorName id="397C7AB4FF9D5B4BFB28BB7EBBBB304F" box="[606,748,1322,1347]" pageId="3" pageNumber="571">Landes, SW</collectorName>
<collectingCountry id="EC9E5FF2FF9D5B4BFB81BB7FBA10304E" box="[759,839,1323,1346]" name="France" pageId="3" pageNumber="571">France</collectingCountry>
. The
<typeStatus id="4B32A1C0FF9D5B4BFAF8BB7EBAA7304F" box="[910,1008,1322,1347]" pageId="3" pageNumber="571" type="holotype">holotype</typeStatus>
is from a lot in the
<collectorName id="397C7AB4FF9D5B4BFD99BB7EBC08304F" box="[1263,1375,1322,1347]" pageId="3" pageNumber="571">d'Orbigny</collectorName>
coll. (
<collectionCode id="F29887A7FF9D5B4BF9E8BB1BB9A0306B" box="[158,247,1359,1383]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="3" pageNumber="571">MNHN</collectionCode>
) from Saint-Paul-lès-Dax, labelled as &quot;
<taxonomicName id="538964E1FF9D5B4BFBC4BB1ABDFB306B" authority="Lamarck, Falunien B" authorityName="Lamarck, Falunien B" box="[690,1196,1358,1383]" class="Bivalvia" family="Cardiidae" genus="Cardium" kingdom="Animalia" order="Veneroida" pageId="3" pageNumber="571" phylum="Mollusca" rank="species" species="burdigalinum">
<emphasis id="A6FDC370FF9D5B4BFBC4BB1ABAE4306A" box="[690,947,1358,1382]" italics="true" pageId="3" pageNumber="571">Cardium burdigalinum</emphasis>
Lamarck, Falunien B
</taxonomicName>
&quot;. The outcrops in the area of Saint-Paul-lès-Dax (SE of Bordeaux) are shallow-water, coarse-grained deposits (&quot;
<emphasis id="A6FDC370FF9D5B4BFDF8BB27BD9F3087" box="[1166,1224,1395,1419]" italics="true" pageId="3" pageNumber="571">falun</emphasis>
&quot;) of Aquitanian to Burdigalian age (
<bibRefCitation id="F0186293FF9D5B4BF82FBBC2BB5530A3" author="Poignant" box="[345,514,1430,1455]" pageId="3" pageNumber="571" refString="Poignant, A. (1966). Le Neogene de St-Paul-les-Dax. In: Drooger, C. W, Reiss, Z., Rutsch, R. F. &amp; Marks, P. (Eds.), Proceedings of the Third Session in Berne (8 - 13 June 1964). International Union of Geological Sciences, Commission on Stratigraphy, Committee on Mediterranean Neogene Stratigraphy. E. J. Brill, Leiden, 225 - 230." type="journal article" year="1966">Poignant 1966</bibRefCitation>
). The
<typeStatus id="4B32A1C0FF9D5B4BFB3FBBC3BBE330A3" box="[585,692,1431,1455]" pageId="3" pageNumber="571" type="paratype">paratypes</typeStatus>
are from Mérignac, a well known Burdigalian locality in the area of Bordeaux (
<bibRefCitation id="F0186293FF9D5B4BF847BBEEB89730DF" author="Folliot" box="[305,448,1466,1491]" pageId="3" pageNumber="571" refString="Folliot M., 1989. Les depots fossiliferes du Miocene du Bordelais. Mineraux et Fossiles, 163, 17 - 28." type="journal article" year="1989">Folliot 1989</bibRefCitation>
;
<bibRefCitation id="F0186293FF9D5B4BF8BDBBEEBBBE30DF" author="Cahuzac" box="[459,745,1466,1491]" pageId="3" pageNumber="571" refString="Cahuzac, B. &amp; Janssen, A. W. (2010) Eocene to Miocene holoplanktonic Mollusca (Gastropoda) of the Aquitaine Basin, southwest France. Scripta Geologica, 141, 1 - 192." type="journal article" year="2010">Cahuzac &amp; Janssen 2010</bibRefCitation>
).
</materialsCitation>
</paragraph>
<paragraph id="94361F62FF9D5B4BF9B1BB8ABD2B3333" blockId="3.[151,1436,494,2031]" pageId="3" pageNumber="571">
<emphasis id="A6FDC370FF9D5B4BF9B1BB8AB8D130FB" bold="true" box="[199,390,1502,1527]" pageId="3" pageNumber="571">Other material.</emphasis>
Mérignac, 4 vs (La Perna coll.); Mérignac,
<date id="E03739A2FF9D5B4BFAE2BB8BBAEB30FA" box="[916,956,1503,1527]" pageId="3" pageNumber="571">1 v</date>
(ter Poorten coll. 1310); Mérignac, 2 vs (NHMW 1850.XXV.99); Léognan,
<date id="E03739A2FF9D5B4BFB37B857BB3A3316" box="[577,621,1539,1563]" pageId="3" pageNumber="571">1 v</date>
(NHMW 1851.XVII.1324); Salles, 2 vs (NHMW 1851.XXIX.56); Léognan, 2 vs (NHMW 1860.XXIX.58); Saucats, 2 vs (NHMW no registration number).
</paragraph>
</subSubSection>
<subSubSection id="DC934CE9FF9D5B4DF9B1B81EBD043253" lastPageId="5" lastPageNumber="573" pageId="3" pageNumber="571" type="etymology">
<paragraph id="94361F62FF9D5B4BF9B1B81EBA0A338A" blockId="3.[151,1436,494,2031]" pageId="3" pageNumber="571">
<emphasis id="A6FDC370FF9D5B4BF9B1B81EB819336F" bold="true" box="[199,334,1610,1635]" pageId="3" pageNumber="571">Etymology.</emphasis>
The species is named after the French naturalist Jean-Pierre Sylvestre de Grateloup (
<date id="E03739A2FF9D5B4BFC7AB81EBCD8336F" box="[1292,1423,1610,1635]" pageId="3" pageNumber="571" value="1782" valueMax="1861">17821861</date>
), who was the first to recognise and name this species
<emphasis id="A6FDC370FF9D5B4BFB91B83BBA01338A" box="[743,854,1646,1670]" italics="true" pageId="3" pageNumber="571">in schedis</emphasis>
.
</paragraph>
<paragraph id="94361F62FF9D5B4BF9B1B8C6BC6C3237" blockId="3.[151,1436,494,2031]" pageId="3" pageNumber="571">
<emphasis id="A6FDC370FF9D5B4BF9B1B8C6B83533A7" bold="true" box="[199,354,1682,1707]" pageId="3" pageNumber="571">Comparison.</emphasis>
The only species which
<taxonomicName id="538964E1FF9D5B4BFB0DB8C7BAE633A7" box="[635,945,1683,1707]" class="Bivalvia" family="Cardiidae" genus="Bucardium" kingdom="Animalia" order="Veneroida" pageId="3" pageNumber="571" phylum="Mollusca" rank="species" species="grateloupianum" status="sp. nov.">
<emphasis id="A6FDC370FF9D5B4BFB0DB8C7BAE633A7" box="[635,945,1683,1707]" italics="true" pageId="3" pageNumber="571">Bucardium grateloupianum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="BDCE7E0BFF9D5B4BFACFB8C6BAAE33A7" box="[953,1017,1682,1707]" pageId="3" pageNumber="571" rank="species">
<emphasis id="A6FDC370FF9D5B4BFACFB8C6BAAE33A7" bold="true" box="[953,1017,1682,1707]" pageId="3" pageNumber="571">n. sp.</emphasis>
</taxonomicNameLabel>
surely deserves to be compared to is
<taxonomicName id="538964E1FF9D5B4BF9E1B8E3B85D33C2" box="[151,266,1719,1742]" class="Bivalvia" family="Cardiidae" genus="Cardium" kingdom="Animalia" order="Veneroida" pageId="3" pageNumber="571" phylum="Mollusca" rank="species" species="ringens">
<emphasis id="A6FDC370FF9D5B4BF9E1B8E3B85D33C2" box="[151,266,1719,1742]" italics="true" pageId="3" pageNumber="571">B. ringens</emphasis>
</taxonomicName>
(
<figureCitation id="0CB203E7FF9D5B4BF86CB8E2B80B33C3" box="[282,348,1718,1743]" captionStart="FIGURE 1" captionStartId="2.[151,250,1416,1438]" captionTargetBox="[151,1436,193,1394]" captionTargetId="figure@2.[151,1436,193,1395]" captionTargetPageId="2" captionText="FIGURE 1. Bucardium ringens (Bruguière, 1789). A F. Casamance, Senegal, depth 10 20 m (ter Poorten coll., TP 634), photo J. J. ter Poorten, with permission (A C, RV; D, LV). G, H. Hann Bay, Senegal, RV (La Perna coll.)." httpUri="https://zenodo.org/record/272520/files/figure.png" pageId="3" pageNumber="571">Fig. 1</figureCitation>
). In
<taxonomicName id="538964E1FF9D5B4BF8E2B8E3BB3133C2" box="[404,614,1718,1742]" class="Bivalvia" family="Cardiidae" genus="Bucardium" kingdom="Animalia" order="Veneroida" pageId="3" pageNumber="571" phylum="Mollusca" rank="species" species="grateloupianum" status="sp. nov.">
<emphasis id="A6FDC370FF9D5B4BF8E2B8E3BB3133C2" box="[404,614,1718,1742]" italics="true" pageId="3" pageNumber="571">B. grateloupianum</emphasis>
</taxonomicName>
<emphasis id="A6FDC370FF9D5B4BFB1BB8E2BBFA33C3" bold="true" box="[621,685,1718,1743]" pageId="3" pageNumber="571">
<taxonomicNameLabel id="BDCE7E0BFF9D5B4BFB1BB8E2BBFA33C3" box="[621,685,1718,1743]" pageId="3" pageNumber="571" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, the antero-dorsal margin is long, almost straight and only slightly sloping whereas it is shorter and distinctly sloping in
<taxonomicName id="538964E1FF9D5B4BFB80B88FBA3D33FE" box="[758,874,1755,1778]" class="Bivalvia" family="Cardiidae" genus="Cardium" kingdom="Animalia" order="Veneroida" pageId="3" pageNumber="571" phylum="Mollusca" rank="species" species="ringens">
<emphasis id="A6FDC370FF9D5B4BFB80B88FBA3D33FE" box="[758,874,1755,1778]" italics="true" pageId="3" pageNumber="571">B. ringens</emphasis>
</taxonomicName>
. The antero-dorsal area with fine radial ribs (
<figureCitation id="0CB203E7FF9D5B4BFC18B88EB9F2321B" captionStart="FIGURE 2" captionStartId="4.[151,250,1938,1960]" captionTargetBox="[151,1436,337,1915]" captionTargetId="figure@4.[151,1436,337,1917]" captionTargetPageId="4" captionText="FIGURE 2. Bucardium grateloupianum n. sp. A C. Holotype, St-Paul-lès-Dax, RV (MNHN Sciences de la Terre A 13845, d'Orbigny coll. 10986), photo J. J. ter Poorten, with permission. D F, K. Paratype 1, Mérignac, LF (MZB 60206). G J. Paratype 2, Mérignac, RV (MZB 60207). L. Mérignac, RV (ter Poorten coll. TP 1310), photo J. J. ter Poorten, with permission." httpUri="https://zenodo.org/record/272521/files/figure.png" pageId="3" pageNumber="571">Fig. 2</figureCitation>
J) is much better defined in the new species, and the posterior digitations are coarser and less numerous with the dorsal one pointing upwards and extending above the umbo, in the large valves at least (
<figureCitation id="0CB203E7FF9D5B4BFD05B976BDE23237" box="[1139,1205,1826,1851]" captionStart="FIGURE 2" captionStartId="4.[151,250,1938,1960]" captionTargetBox="[151,1436,337,1915]" captionTargetId="figure@4.[151,1436,337,1917]" captionTargetPageId="4" captionText="FIGURE 2. Bucardium grateloupianum n. sp. A C. Holotype, St-Paul-lès-Dax, RV (MNHN Sciences de la Terre A 13845, d'Orbigny coll. 10986), photo J. J. ter Poorten, with permission. D F, K. Paratype 1, Mérignac, LF (MZB 60206). G J. Paratype 2, Mérignac, RV (MZB 60207). L. Mérignac, RV (ter Poorten coll. TP 1310), photo J. J. ter Poorten, with permission." httpUri="https://zenodo.org/record/272521/files/figure.png" pageId="3" pageNumber="571">Fig. 2</figureCitation>
AC, G, H).
</paragraph>
<paragraph id="94361F62FF9D5B4BF9B1B912BC0B32E2" blockId="3.[151,1436,494,2031]" pageId="3" pageNumber="571">
The radial sculpture follows the same pattern as in
<taxonomicName id="538964E1FF9D5B4BFB88B913BA273252" box="[766,880,1863,1886]" class="Bivalvia" family="Cardiidae" genus="Cardium" kingdom="Animalia" order="Veneroida" pageId="3" pageNumber="571" phylum="Mollusca" rank="species" species="ringens">
<emphasis id="A6FDC370FF9D5B4BFB88B913BA273252" box="[766,880,1863,1886]" italics="true" pageId="3" pageNumber="571">B. ringens</emphasis>
</taxonomicName>
, but in the new species the medial ribs are mainly symmetrically convex and only becoming asymmetrical and then ridged near the posterior margin. The crescentshaped growth striae on the posterior ribs are well distinct in
<taxonomicName id="538964E1FF9D5B4BFA48B9DBBAE532AA" box="[830,946,1935,1958]" class="Bivalvia" family="Cardiidae" genus="Cardium" kingdom="Animalia" order="Veneroida" pageId="3" pageNumber="571" phylum="Mollusca" rank="species" species="ringens">
<emphasis id="A6FDC370FF9D5B4BFA48B9DBBAE532AA" box="[830,946,1935,1958]" italics="true" pageId="3" pageNumber="571">B. ringens</emphasis>
</taxonomicName>
but ill-defined in the new species (
<figureCitation id="0CB203E7FF9D5B4BFC41B9DABC2D32AB" box="[1335,1402,1934,1959]" captionStart="FIGURE 2" captionStartId="4.[151,250,1938,1960]" captionTargetBox="[151,1436,337,1915]" captionTargetId="figure@4.[151,1436,337,1917]" captionTargetPageId="4" captionText="FIGURE 2. Bucardium grateloupianum n. sp. A C. Holotype, St-Paul-lès-Dax, RV (MNHN Sciences de la Terre A 13845, d'Orbigny coll. 10986), photo J. J. ter Poorten, with permission. D F, K. Paratype 1, Mérignac, LF (MZB 60206). G J. Paratype 2, Mérignac, RV (MZB 60207). L. Mérignac, RV (ter Poorten coll. TP 1310), photo J. J. ter Poorten, with permission." httpUri="https://zenodo.org/record/272521/files/figure.png" pageId="3" pageNumber="571">Fig. 2</figureCitation>
C).
<taxonomicName id="538964E1FF9D5B4BF9E1B9E7B89932C7" box="[151,462,1971,1995]" class="Bivalvia" family="Cardiidae" genus="Bucardium" kingdom="Animalia" order="Veneroida" pageId="3" pageNumber="571" phylum="Mollusca" rank="species" species="grateloupianum" status="sp. nov.">
<emphasis id="A6FDC370FF9D5B4BF9E1B9E7B89932C7" box="[151,462,1971,1995]" italics="true" pageId="3" pageNumber="571">Bucardium grateloupianum</emphasis>
</taxonomicName>
<emphasis id="A6FDC370FF9D5B4BF8A3B9E6BB4132C7" bold="true" box="[469,534,1970,1995]" pageId="3" pageNumber="571">
<taxonomicNameLabel id="BDCE7E0BFF9D5B4BF8A3B9E6BB4132C7" box="[469,534,1970,1995]" pageId="3" pageNumber="571" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
is slightly elongate antero-posteriorly, particular at larger size, instead of being almost equidimensional as
<emphasis id="A6FDC370FF9D5B4BF8B1B983BB1632E2" box="[455,577,2007,2030]" italics="true" pageId="3" pageNumber="571">
<taxonomicName id="538964E1FF9D5B4BF8B1B983BB6A32E2" box="[455,573,2007,2030]" class="Bivalvia" family="Cardiidae" genus="Cardium" kingdom="Animalia" order="Veneroida" pageId="3" pageNumber="571" phylum="Mollusca" rank="species" species="ringens">B. ringens</taxonomicName>
,
</emphasis>
and with a larger maximum size (slightly exceeding
<quantity id="5371B287FF9D5B4BFDE5B982BDB432E2" box="[1171,1251,2006,2031]" metricMagnitude="-2" metricUnit="m" metricValue="6.0" pageId="3" pageNumber="571" unit="mm" value="60.0">60 mm</quantity>
in length).
</paragraph>
<paragraph id="94361F62FF9A5B4CF9B1BECCB9AE3411" blockId="4.[151,1436,151,285]" pageId="4" pageNumber="572">
Some differences can be also seen in the hinge, which is straighter in the new species, with the posterior lateral tooth in the right valve further away from the cardinal teeth than in
<taxonomicName id="538964E1FF9A5B4CFAC2BEE9BD7B35D8" box="[948,1068,189,212]" class="Bivalvia" family="Cardiidae" genus="Cardium" kingdom="Animalia" order="Veneroida" pageId="4" pageNumber="572" phylum="Mollusca" rank="species" species="ringens">
<emphasis id="A6FDC370FF9A5B4CFAC2BEE9BD7B35D8" box="[948,1068,189,212]" italics="true" pageId="4" pageNumber="572">B. ringens</emphasis>
</taxonomicName>
(
<figureCitation id="0CB203E7FF9A5B4CFD4BBEE8BDD435D8" box="[1085,1155,188,213]" captionStart="FIGURE 2" captionStartId="4.[151,250,1938,1960]" captionTargetBox="[151,1436,337,1915]" captionTargetId="figure@4.[151,1436,337,1917]" captionTargetPageId="4" captionText="FIGURE 2. Bucardium grateloupianum n. sp. A C. Holotype, St-Paul-lès-Dax, RV (MNHN Sciences de la Terre A 13845, d'Orbigny coll. 10986), photo J. J. ter Poorten, with permission. D F, K. Paratype 1, Mérignac, LF (MZB 60206). G J. Paratype 2, Mérignac, RV (MZB 60207). L. Mérignac, RV (ter Poorten coll. TP 1310), photo J. J. ter Poorten, with permission." httpUri="https://zenodo.org/record/272521/files/figure.png" pageId="4" pageNumber="572">Fig. 2</figureCitation>
F, I). Crenulation on the lunular flap is generally coarser and better developed in the new species (
<figureCitation id="0CB203E7FF9A5B4CFAA3BE8BBD4E35F4" box="[981,1049,223,248]" captionStart="FIGURE 2" captionStartId="4.[151,250,1938,1960]" captionTargetBox="[151,1436,337,1915]" captionTargetId="figure@4.[151,1436,337,1917]" captionTargetPageId="4" captionText="FIGURE 2. Bucardium grateloupianum n. sp. A C. Holotype, St-Paul-lès-Dax, RV (MNHN Sciences de la Terre A 13845, d'Orbigny coll. 10986), photo J. J. ter Poorten, with permission. D F, K. Paratype 1, Mérignac, LF (MZB 60206). G J. Paratype 2, Mérignac, RV (MZB 60207). L. Mérignac, RV (ter Poorten coll. TP 1310), photo J. J. ter Poorten, with permission." httpUri="https://zenodo.org/record/272521/files/figure.png" pageId="4" pageNumber="572">Fig. 2</figureCitation>
K), but this character seems rather variable.
</paragraph>
<caption id="C0F64FEAFF9A5B4CF9E1B9C6BCCD32E9" httpUri="https://zenodo.org/record/272521/files/figure.png" pageId="4" pageNumber="572" targetBox="[151,1436,337,1915]" targetPageId="4">
<paragraph id="94361F62FF9A5B4CF9E1B9C6BCCD32E9" blockId="4.[151,1436,1938,2021]" pageId="4" pageNumber="572">
<emphasis id="A6FDC370FF9A5B4CF9E1B9C6B85832AB" bold="true" box="[151,271,1938,1960]" pageId="4" pageNumber="572">FIGURE 2</emphasis>
.
<taxonomicName id="538964E1FF9A5B4CF86BB9C6BB6F32A4" box="[285,568,1938,1960]" class="Bivalvia" family="Cardiidae" genus="Bucardium" kingdom="Animalia" order="Veneroida" pageId="4" pageNumber="572" phylum="Mollusca" rank="species" species="grateloupianum" status="sp. nov.">
<emphasis id="A6FDC370FF9A5B4CF86BB9C6BB6F32A4" box="[285,568,1938,1960]" italics="true" pageId="4" pageNumber="572">Bucardium grateloupianum</emphasis>
</taxonomicName>
<emphasis id="A6FDC370FF9A5B4CFB36B9C6BBE232A4" bold="true" box="[576,693,1938,1960]" pageId="4" pageNumber="572">
<taxonomicNameLabel id="BDCE7E0BFF9A5B4CFB36B9C6BB2B32AB" box="[576,636,1938,1959]" pageId="4" pageNumber="572" rank="species">n. sp.</taxonomicNameLabel>
AC
</emphasis>
. Holotype, St-Paul-lès-Dax, RV (MNHN Sciences de la Terre A13845, d'Orbigny coll. 10986), photo J.J. ter Poorten, with permission.
<emphasis id="A6FDC370FF9A5B4CFA30B9E5BAC232CA" bold="true" box="[838,917,1969,1991]" pageId="4" pageNumber="572">DF, K</emphasis>
. Paratype 1, Mérignac, LF (MZB 60206).
<emphasis id="A6FDC370FF9A5B4CFC1FB9E5BCC232CA" bold="true" box="[1385,1429,1969,1991]" pageId="4" pageNumber="572">GJ</emphasis>
. Paratype 2, Mérignac, RV (MZB 60207).
<emphasis id="A6FDC370FF9A5B4CFB48B984BB1832E9" bold="true" box="[574,591,2000,2021]" pageId="4" pageNumber="572">L</emphasis>
. Mérignac, RV (ter Poorten coll. TP 1310), photo J.J. ter Poorten, with permission.
</paragraph>
</caption>
<caption id="C0F64FEAFF9B5B4DF9E1BDBEBBA63131" httpUri="https://zenodo.org/record/272522/files/figure.png" pageId="5" pageNumber="573" targetBox="[151,1436,193,980]" targetPageId="5">
<paragraph id="94361F62FF9B5B4DF9E1BDBEBBA63131" blockId="5.[151,1436,1002,1085]" pageId="5" pageNumber="573">
<emphasis id="A6FDC370FF9B5B4DF9E1BDBEB85A36F3" bold="true" box="[151,269,1002,1024]" pageId="5" pageNumber="573">FIGURE 3</emphasis>
.
<taxonomicName id="538964E1FF9B5B4DF86CBDBEBB65310C" box="[282,562,1002,1024]" class="Bivalvia" family="Cardiidae" genus="Bucardium" kingdom="Animalia" order="Veneroida" pageId="5" pageNumber="573" phylum="Mollusca" rank="species" species="grateloupianum" status="sp. nov.">
<emphasis id="A6FDC370FF9B5B4DF86CBDBEBB65310C" box="[282,562,1002,1024]" italics="true" pageId="5" pageNumber="573">Bucardium grateloupianum</emphasis>
</taxonomicName>
<emphasis id="A6FDC370FF9B5B4DFB4EBDBEBBDD36F3" bold="true" box="[568,650,1002,1023]" pageId="5" pageNumber="573">
<taxonomicNameLabel id="BDCE7E0BFF9B5B4DFB4EBDBEBB2536F3" box="[568,626,1002,1023]" pageId="5" pageNumber="573" rank="species">n. sp.</taxonomicNameLabel>
A
</emphasis>
. Salles, RV (NHMW 1851.XIX.56).
<emphasis id="A6FDC370FF9B5B4DFD70BDBEBD4036F3" bold="true" box="[1030,1047,1002,1023]" pageId="5" pageNumber="573">B</emphasis>
. Saucats, RV (NHMW no registration number).
<emphasis id="A6FDC370FF9B5B4DF98FBA5DB85C3113" bold="true" box="[249,267,1033,1055]" pageId="5" pageNumber="573">C</emphasis>
. Mérignac, LV (NHMW 1850.XXV.99).
<emphasis id="A6FDC370FF9B5B4DFBDBBA5DBBE83112" bold="true" box="[685,703,1033,1054]" pageId="5" pageNumber="573">D</emphasis>
. Salles, LV (NHMW no registration number).
<emphasis id="A6FDC370FF9B5B4DFDE1BA5DBDFF3112" bold="true" box="[1175,1192,1033,1054]" pageId="5" pageNumber="573">E</emphasis>
, Léognan, RV (NHMW 1860.XXIX.58).
<emphasis id="A6FDC370FF9B5B4DF832BA7CB8063131" bold="true" box="[324,337,1064,1085]" pageId="5" pageNumber="573">F</emphasis>
. Léognan, LV (NHMW 1860.XXIX.58).
</paragraph>
</caption>
<caption id="C0F64FEAFF9B5B4DF9E1B8CCBC3333A2" box="[151,1380,1688,1710]" httpUri="https://zenodo.org/record/272523/files/figure.png" pageId="5" pageNumber="573" targetBox="[151,1436,1135,1666]" targetPageId="5">
<paragraph id="94361F62FF9B5B4DF9E1B8CCBC3333A2" blockId="5.[151,1380,1688,1710]" box="[151,1380,1688,1710]" pageId="5" pageNumber="573">
<emphasis id="A6FDC370FF9B5B4DF9E1B8CCB84533A1" bold="true" box="[151,274,1688,1710]" pageId="5" pageNumber="573">FIGURE 4.</emphasis>
<taxonomicName id="538964E1FF9B5B4DF86FB8CCBB3B33A2" authority="Lamarck, 1819" authorityName="Lamarck" authorityYear="1819" box="[281,620,1688,1710]" class="Bivalvia" family="Cardiidae" genus="Cardium" kingdom="Animalia" order="Veneroida" pageId="5" pageNumber="573" phylum="Mollusca" rank="species" species="indicum">
<emphasis id="A6FDC370FF9B5B4DF86FB8CCB89D33A2" box="[281,458,1688,1710]" italics="true" pageId="5" pageNumber="573">Cardium indicum</emphasis>
Lamarck, 1819
</taxonomicName>
.
<emphasis id="A6FDC370FF9B5B4DFB00B8CCBBF033A2" bold="true" box="[630,679,1688,1710]" pageId="5" pageNumber="573">AC</emphasis>
. Gravina in Puglia, Early Pleistocene, LV, sculpture details (MPUB).
</paragraph>
</caption>
<paragraph id="94361F62FF9B5B4DF9B1B888BD043253" blockId="5.[151,1437,1755,2032]" pageId="5" pageNumber="573">
In
<taxonomicName id="538964E1FF9B5B4DF993B888B80E33FF" box="[229,345,1756,1779]" class="Bivalvia" family="Cardiidae" genus="Cardium" kingdom="Animalia" order="Veneroida" pageId="5" pageNumber="573" phylum="Mollusca" rank="species" species="ringens">
<emphasis id="A6FDC370FF9B5B4DF993B888B80E33FF" box="[229,345,1756,1779]" italics="true" pageId="5" pageNumber="573">B. ringens</emphasis>
</taxonomicName>
, right and left digitations touch each other at least in part (
<figureCitation id="0CB203E7FF9B5B4DFA98B88FBD7A33FF" box="[1006,1069,1755,1780]" captionStart="FIGURE 1" captionStartId="2.[151,250,1416,1438]" captionTargetBox="[151,1436,193,1394]" captionTargetId="figure@2.[151,1436,193,1395]" captionTargetPageId="2" captionText="FIGURE 1. Bucardium ringens (Bruguière, 1789). A F. Casamance, Senegal, depth 10 20 m (ter Poorten coll., TP 634), photo J. J. ter Poorten, with permission (A C, RV; D, LV). G, H. Hann Bay, Senegal, RV (La Perna coll.)." httpUri="https://zenodo.org/record/272520/files/figure.png" pageId="5" pageNumber="573">Fig. 1</figureCitation>
E) whereas the posterior gape in
<taxonomicName id="538964E1FF9B5B4DF9E1B954B83F321B" box="[151,360,1791,1815]" class="Bivalvia" family="Cardiidae" genus="Bucardium" kingdom="Animalia" order="Veneroida" pageId="5" pageNumber="573" phylum="Mollusca" rank="species" species="grateloupianum" status="sp. nov.">
<emphasis id="A6FDC370FF9B5B4DF9E1B954B83F321B" box="[151,360,1791,1815]" italics="true" pageId="5" pageNumber="573">B. grateloupianum</emphasis>
</taxonomicName>
<emphasis id="A6FDC370FF9B5B4DF819B8AAB8FA321B" bold="true" box="[367,429,1790,1815]" pageId="5" pageNumber="573">
<taxonomicNameLabel id="BDCE7E0BFF9B5B4DF819B8AAB8FA321B" box="[367,429,1790,1815]" pageId="5" pageNumber="573" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
seems wider, with digitations not touching each other. Variations mainly regard the length and number of posterior digitations: they are typically well developed, up to six, but specimens with few, short digitations (34) (
<figureCitation id="0CB203E7FF9B5B4DF816B912B8F73253" box="[352,416,1862,1887]" captionStart="FIGURE 3" captionStartId="5.[151,250,1002,1024]" captionTargetBox="[151,1436,193,980]" captionTargetId="figure@5.[151,1436,193,981]" captionTargetPageId="5" captionText="FIGURE 3. Bucardium grateloupianum n. sp. A. Salles, RV (NHMW 1851. XIX. 56). B. Saucats, RV (NHMW no registration number). C. Mérignac, LV (NHMW 1850. XXV. 99). D. Salles, LV (NHMW no registration number). E, Léognan, RV (NHMW 1860. XXIX. 58). F. Léognan, LV (NHMW 1860. XXIX. 58)." httpUri="https://zenodo.org/record/272522/files/figure.png" pageId="5" pageNumber="573">Fig. 3</figureCitation>
AC) and even almost mutic specimens are known (
<figureCitation id="0CB203E7FF9B5B4DFA9AB912BD793253" box="[1004,1070,1862,1887]" captionStart="FIGURE 3" captionStartId="5.[151,250,1002,1024]" captionTargetBox="[151,1436,193,980]" captionTargetId="figure@5.[151,1436,193,981]" captionTargetPageId="5" captionText="FIGURE 3. Bucardium grateloupianum n. sp. A. Salles, RV (NHMW 1851. XIX. 56). B. Saucats, RV (NHMW no registration number). C. Mérignac, LV (NHMW 1850. XXV. 99). D. Salles, LV (NHMW no registration number). E, Léognan, RV (NHMW 1860. XXIX. 58). F. Léognan, LV (NHMW 1860. XXIX. 58)." httpUri="https://zenodo.org/record/272522/files/figure.png" pageId="5" pageNumber="573">Fig. 3</figureCitation>
D).
</paragraph>
</subSubSection>
<subSubSection id="DC934CE9FF9B5B4EF9B1B93FBD9B363B" lastPageId="6" lastPageNumber="574" pageId="5" pageNumber="573" type="description">
<paragraph id="94361F62FF9B5B4EF9B1B93FBC123410" blockId="5.[151,1437,1755,2032]" lastBlockId="6.[151,1437,151,1977]" lastPageId="6" lastPageNumber="574" pageId="5" pageNumber="573">
<emphasis id="A6FDC370FF9B5B4DF9B1B93FB8B63288" bold="true" box="[199,481,1899,1924]" pageId="5" pageNumber="573">Nomenclatorial history.</emphasis>
<taxonomicName id="538964E1FF9B5B4DF891B93FBA4A328F" box="[487,797,1899,1923]" class="Bivalvia" family="Cardiidae" genus="Bucardium" kingdom="Animalia" order="Veneroida" pageId="5" pageNumber="573" phylum="Mollusca" rank="species" species="grateloupianum" status="sp. nov.">
<emphasis id="A6FDC370FF9B5B4DF891B93FBA4A328F" box="[487,797,1899,1923]" italics="true" pageId="5" pageNumber="573">Bucardium grateloupianum</emphasis>
</taxonomicName>
<emphasis id="A6FDC370FF9B5B4DFA52B93FBA343288" bold="true" box="[804,867,1899,1924]" pageId="5" pageNumber="573">
<taxonomicNameLabel id="BDCE7E0BFF9B5B4DFA52B93FBA343288" box="[804,867,1899,1924]" pageId="5" pageNumber="573" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
was first illustrated by
<bibRefCitation id="F0186293FF9B5B4DFD1CB93FB8DD32AB" author="Cossmann" pageId="5" pageNumber="573" refString="Cossmann, M. &amp; Peyrot, A. (1912) Conchologie neogenique de l'Aquitaine. Actes de la Societe Linneenne de Bordeaux, 65, 99 - 178." type="journal article" year="1912">Cossmann &amp; Peyrot (1912: 115, pl. 22, figs 36)</bibRefCitation>
as
<taxonomicName id="538964E1FF9B5B4DF8C2B9DBBAB032AB" authority="Mayer, 1866" authorityName="Mayer" authorityYear="1866" box="[436,999,1934,1960]" class="Bivalvia" family="Cardiidae" genus="Cardium" kingdom="Animalia" order="Veneroida" pageId="5" pageNumber="573" phylum="Mollusca" rank="species" species="grateloupi" subGenus="Ringicardium">
<emphasis id="A6FDC370FF9B5B4DF8C2B9DBBA1D32AB" box="[436,842,1935,1959]" italics="true" pageId="5" pageNumber="573">Cardium (Ringicardium) grateloupi</emphasis>
<bibRefCitation id="F0186293FF9B5B4DFA22B9C4BAB032AB" author="Mayer" box="[852,999,1934,1960]" pageId="5" pageNumber="573" refString="Mayer, M. C. (1866) Description de coquilles fossiles des terrains tertiaires superieurs (suite). Journal de Conchyliologie, 14, 67 - 76." type="journal article" year="1866">Mayer, 1866</bibRefCitation>
</taxonomicName>
. They tried, though unsuccessfully, to clarify the species identity, already confused at that time (“
<emphasis id="A6FDC370FF9B5B4DFA45B9E7BC0432C7" box="[819,1363,1971,1995]" italics="true" pageId="5" pageNumber="573">Cette espèce a donné lieu a plusieurs confusions</emphasis>
”). As observed by
<bibRefCitation id="F0186293FF9B5B4DF80AB98CBB4A32E3" author="Magne" box="[380,541,2006,2032]" pageId="5" pageNumber="573" refString="Magne, A. (1950) Les Ringicardium neogenes du Bassin d'Aquitaine. Journal de Conchyliologie, 90, 122 - 124." type="journal article" year="1950">Magne (1950)</bibRefCitation>
,
<taxonomicName id="538964E1FF9B5B4DFB5FB983BA5F32E3" box="[553,776,2007,2031]" class="Bivalvia" family="Cardiidae" genus="Cardium" kingdom="Animalia" order="Veneroida" pageId="5" pageNumber="573" phylum="Mollusca" rank="species" species="grateloupi">
<emphasis id="A6FDC370FF9B5B4DFB5FB983BA5F32E3" box="[553,776,2007,2031]" italics="true" pageId="5" pageNumber="573">Cardium grateloupi</emphasis>
</taxonomicName>
was proposed by
<bibRefCitation id="F0186293FF9B5B4DFAA7B98CBDC132E3" author="Mayer" box="[977,1174,2006,2032]" pageId="5" pageNumber="573" refString="Mayer, M. C. (1866) Description de coquilles fossiles des terrains tertiaires superieurs (suite). Journal de Conchyliologie, 14, 67 - 76." type="journal article" year="1866">Mayer (1866: 71)</bibRefCitation>
as a replacement name for “
<taxonomicName id="538964E1FF985B4EF9BABECCBB0535BC" authority="Grateloup" authorityName="Grateloup" box="[204,594,151,176]" class="Bivalvia" family="Cardiidae" genus="Cardium" kingdom="Animalia" order="Veneroida" pageId="6" pageNumber="574" phylum="Mollusca" rank="species" species="hoernesianum">
<emphasis id="A6FDC370FF985B4EF9BABECCB88035BC" box="[204,471,152,176]" italics="true" pageId="6" pageNumber="574">Cardium hoernesianum</emphasis>
Grateloup
</taxonomicName>
” of
<bibRefCitation id="F0186293FF985B4EFBF2BECCBD6735BC" author="Hornes" box="[644,1072,151,176]" pageId="6" pageNumber="574" refString="Hornes, M. (1862) Die fossilen Mollusken des Tertiarbeckens von Wien. Band II: Bivalvia. Abhandlungen der Kaiserlich- Koniglichen Geologischen Reichsanstalt, 4, 117 - 214." type="journal article" year="1862">Hörnes (1862: 183, pl. 27, figs. 1a-c)</bibRefCitation>
, from the Eggenburgian (Upper Miocene) of
<collectingCountry id="EC9E5FF2FF985B4EF85DBEE8B8D735D9" box="[299,384,188,213]" name="Austria" pageId="6" pageNumber="574">Austria</collectingCountry>
, thought to be preoccupied by
<taxonomicName id="538964E1FF985B4EFB95BEE8BD7335D9" authority="Deshayes, 1858" authorityName="Deshayes" authorityYear="1858" box="[739,1060,188,213]" class="Bivalvia" family="Cardiidae" genus="Cardium" kingdom="Animalia" order="Veneroida" pageId="6" pageNumber="574" phylum="Mollusca" rank="species" species="hoernesi">
<emphasis id="A6FDC370FF985B4EFB95BEE8BA3235D8" box="[739,869,188,212]" italics="true" pageId="6" pageNumber="574">C. hoernesi</emphasis>
Deshayes, 1858
</taxonomicName>
. This is very clear from Mayers comments: “…
<emphasis id="A6FDC370FF985B4EF831BEB4B8793410" italics="true" pageId="6" pageNumber="574">
je ferai remarquer que le C. Hörnesianum Grat. (Hörn., Foss. Mollusk. Von
<collectingRegion id="564DD180FF985B4EFDE1BEB4BD8535FB" box="[1175,1234,224,247]" country="Austria" name="Wien" pageId="6" pageNumber="574">Wien</collectingRegion>
, vol. II, p. 183, pl. XXVII, fig. 1)
</emphasis>
[…]
<emphasis id="A6FDC370FF985B4EF81CBF50BC653410" box="[362,1330,260,284]" italics="true" pageId="6" pageNumber="574">doit encore changer de nom, puisque le C. Hörnesi Desh. à la priorité de près dun an</emphasis>
”.
</paragraph>
<paragraph id="94361F62FF985B4EF9B1BF7CBA3734DC" blockId="6.[151,1437,151,1977]" pageId="6" pageNumber="574">
<bibRefCitation id="F0186293FF985B4EF9B1BF7CB821344C" author="Hornes" box="[199,374,295,320]" pageId="6" pageNumber="574" refString="Hornes, M. (1862) Die fossilen Mollusken des Tertiarbeckens von Wien. Band II: Bivalvia. Abhandlungen der Kaiserlich- Koniglichen Geologischen Reichsanstalt, 4, 117 - 214." type="journal article" year="1862">Hörnes (1862)</bibRefCitation>
and Schaffer (1919: 66) explained that Grateloup sent
<taxonomicName id="538964E1FF985B4EFD27BF7CBDE3344C" box="[1105,1204,296,320]" class="Bivalvia" family="Cardiidae" genus="Cardium" kingdom="Animalia" order="Veneroida" pageId="6" pageNumber="574" phylum="Mollusca" rank="genus">
<emphasis id="A6FDC370FF985B4EFD27BF7CBDE3344C" box="[1105,1204,296,320]" italics="true" pageId="6" pageNumber="574">Cardium</emphasis>
</taxonomicName>
specimens to the
<emphasis id="A6FDC370FF985B4EF9E1BF18B8C23468" box="[151,405,332,356]" italics="true" pageId="6" pageNumber="574">Hofmineralienkabinett</emphasis>
in
<collectingRegion id="564DD180FF985B4EF8C1BF18BB513469" box="[439,518,332,357]" country="Austria" name="Wien" pageId="6" pageNumber="574">Vienna</collectingRegion>
(the modern Naturhistorisches Museum), one year before his death. This material, labelled as
<taxonomicName id="538964E1FF985B4EF861BF24BBCF3484" authority="Grateloup" authorityName="Grateloup" box="[279,664,367,392]" class="Bivalvia" family="Cardiidae" genus="Cardium" kingdom="Animalia" order="Veneroida" pageId="6" pageNumber="574" phylum="Mollusca" rank="species" species="hoernesianum" status="sp. nov.">
<emphasis id="A6FDC370FF985B4EF861BF24BB773484" box="[279,544,368,392]" italics="true" pageId="6" pageNumber="574">Cardium hoernesianum</emphasis>
Grateloup
</taxonomicName>
<taxonomicNameLabel id="BDCE7E0BFF985B4EFBD6BF24BBAD348B" box="[672,762,368,391]" pageId="6" pageNumber="574" rank="species">sp. nov.</taxonomicNameLabel>
, from the Burdigalian yellowish
<emphasis id="A6FDC370FF985B4EFD07BF24BDFC3484" box="[1137,1195,368,392]" italics="true" pageId="6" pageNumber="574">falun</emphasis>
at Léognan, south of Bordeaux, is deemed to be conspecific with
<taxonomicName id="538964E1FF985B4EFBF5BFC0BA0134A0" box="[643,854,404,428]" class="Bivalvia" family="Cardiidae" genus="Bucardium" kingdom="Animalia" order="Veneroida" pageId="6" pageNumber="574" phylum="Mollusca" rank="species" species="grateloupianum" status="sp. nov.">
<emphasis id="A6FDC370FF985B4EFBF5BFC0BA0134A0" box="[643,854,404,428]" italics="true" pageId="6" pageNumber="574">C. grateloupianum</emphasis>
</taxonomicName>
<emphasis id="A6FDC370FF985B4EFA2ABFC0BACC34A1" bold="true" box="[860,923,404,429]" pageId="6" pageNumber="574">
<taxonomicNameLabel id="BDCE7E0BFF985B4EFA2ABFC0BACC34A1" box="[860,923,404,429]" pageId="6" pageNumber="574" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
Grateloup also annotated that his new species had been confused with
<taxonomicName id="538964E1FF985B4EF8D1BFECBA0C34DC" authority="Lamarck, 1819" authorityName="Lamarck" authorityYear="1819" box="[423,859,439,464]" class="Bivalvia" family="Cardiidae" genus="Cardium" kingdom="Animalia" order="Veneroida" pageId="6" pageNumber="574" phylum="Mollusca" rank="species" species="burdigalinum">
<emphasis id="A6FDC370FF985B4EF8D1BFECBBF034DC" box="[423,679,440,464]" italics="true" pageId="6" pageNumber="574">Cardium burdigalinum</emphasis>
<bibRefCitation id="F0186293FF985B4EFBD9BFE3BA0C34DC" author="Lamarck" box="[687,859,439,464]" pageId="6" pageNumber="574" refString="Lamarck, J. - B. (1819) Histoire naturelle des animaux sans vertebre s. Tome 6, partie 1 ere, 343 pp." type="book" year="1819">Lamarck, 1819</bibRefCitation>
</taxonomicName>
.
</paragraph>
<paragraph id="94361F62FF985B4EF9B1BF88BD9B363B" blockId="6.[151,1437,151,1977]" pageId="6" pageNumber="574">
Actually, some material labelled as &quot;
<taxonomicName id="538964E1FF985B4EFB03BF88BD5634F9" authority="Grateloup" authorityName="Grateloup" box="[629,1025,476,501]" class="Bivalvia" family="Cardiidae" genus="Cardium" kingdom="Animalia" order="Veneroida" pageId="6" pageNumber="574" phylum="Mollusca" rank="species" species="hoernesianum">
<emphasis id="A6FDC370FF985B4EFB03BF88BAD534F8" box="[629,898,476,500]" italics="true" pageId="6" pageNumber="574">Cardium hoernesianum</emphasis>
Grateloup
</taxonomicName>
&quot;, all from the area of Bordeaux, is present at NHMW (
<figureCitation id="0CB203E7FF985B4EF80ABFABB8973714" box="[380,448,511,536]" captionStart="FIGURE 3" captionStartId="5.[151,250,1002,1024]" captionTargetBox="[151,1436,193,980]" captionTargetId="figure@5.[151,1436,193,981]" captionTargetPageId="5" captionText="FIGURE 3. Bucardium grateloupianum n. sp. A. Salles, RV (NHMW 1851. XIX. 56). B. Saucats, RV (NHMW no registration number). C. Mérignac, LV (NHMW 1850. XXV. 99). D. Salles, LV (NHMW no registration number). E, Léognan, RV (NHMW 1860. XXIX. 58). F. Léognan, LV (NHMW 1860. XXIX. 58)." httpUri="https://zenodo.org/record/272522/files/figure.png" pageId="6" pageNumber="574">Fig. 3</figureCitation>
AF). One lot, consisting of two worn valves, almost lacking the posterior digitations (
<figureCitation id="0CB203E7FF985B4EF9E9BC70B9B63730" box="[159,225,548,573]" captionStart="FIGURE 3" captionStartId="5.[151,250,1002,1024]" captionTargetBox="[151,1436,193,980]" captionTargetId="figure@5.[151,1436,193,981]" captionTargetPageId="5" captionText="FIGURE 3. Bucardium grateloupianum n. sp. A. Salles, RV (NHMW 1851. XIX. 56). B. Saucats, RV (NHMW no registration number). C. Mérignac, LV (NHMW 1850. XXV. 99). D. Salles, LV (NHMW no registration number). E, Léognan, RV (NHMW 1860. XXIX. 58). F. Léognan, LV (NHMW 1860. XXIX. 58)." httpUri="https://zenodo.org/record/272522/files/figure.png" pageId="6" pageNumber="574">Fig. 3</figureCitation>
E, F), is from Léognan and was acquired in 1860 (as can be inferred from the registration number), i.e. one year before Grateloups death in 1861 (
<bibRefCitation id="F0186293FF985B4EFB2EBC1CBA6F376C" author="Lesport" box="[600,824,584,608]" pageId="6" pageNumber="574" refString="Lesport, J. - F., Cluzaud, A. &amp; Verhecken, A. (2012) Les publications du Docteur Jean-Pierre Sylvestre de Grateloup sur les mollusques fossiles du Bassin d'Aquitaine (S. - O. France): dates de parutions et commentaires. Bulletin de la Societe Linneenne de Bordeaux, New Series, 147, 40, 417 - 485." type="journal article" year="2012">
Lesport
<emphasis id="A6FDC370FF985B4EFBCEBC1DBBBC376C" box="[696,747,584,608]" italics="true" pageId="6" pageNumber="574">et al</emphasis>
. 2012
</bibRefCitation>
). It could represent Grateloups original material, and their worn condition could explain why
<bibRefCitation id="F0186293FF985B4EFB2ABC39BBA93789" author="Hornes" box="[604,766,620,645]" pageId="6" pageNumber="574" refString="Hornes, M. (1862) Die fossilen Mollusken des Tertiarbeckens von Wien. Band II: Bivalvia. Abhandlungen der Kaiserlich- Koniglichen Geologischen Reichsanstalt, 4, 117 - 214." type="journal article" year="1862">Hörnes (1862)</bibRefCitation>
applied Grateloups manuscript name to a different species (further discussed below), remarking that “
<emphasis id="A6FDC370FF985B4EFB0BBCC4BCC737C0" italics="true" pageId="6" pageNumber="574">Die Wiener Exemplare gleichen vollkommen denen aus der Umgebung von Bordeaux, nur sind sie wie alle hiesigen Vorkommnisse verhältnissmässig grösser und dicker in der Schale</emphasis>
” (&quot;The specimens from
<collectingRegion id="564DD180FF985B4EF8D6BC83B8B837FC" box="[416,495,727,752]" country="Austria" name="Wien" pageId="6" pageNumber="574">Vienna</collectingRegion>
are completely identical to those from the Bordeaux area, but […] the shells are comparatively larger and thicker&quot;). The correct authorship of
<taxonomicName id="538964E1FF985B4EFA20BCA8BD403618" box="[854,1047,764,788]" class="Bivalvia" family="Cardiidae" genus="Cardium" kingdom="Animalia" order="Veneroida" pageId="6" pageNumber="574" phylum="Mollusca" rank="species" species="hoernesianum">
<emphasis id="A6FDC370FF985B4EFA20BCA8BD403618" box="[854,1047,764,788]" italics="true" pageId="6" pageNumber="574">C. hoernesianum</emphasis>
</taxonomicName>
should be then
<bibRefCitation id="F0186293FF985B4EFDA6BCA9BC3C3619" author="Hornes" box="[1232,1387,764,789]" pageId="6" pageNumber="574" refString="Hornes, M. (1862) Die fossilen Mollusken des Tertiarbeckens von Wien. Band II: Bivalvia. Abhandlungen der Kaiserlich- Koniglichen Geologischen Reichsanstalt, 4, 117 - 214." type="journal article" year="1862">Hörnes, 1862</bibRefCitation>
, not Grateloup in Hörnes, given that Hörnes did not describe Grateloup's species, but a different one.
</paragraph>
</subSubSection>
<subSubSection id="DC934CE9FF985B4EF9B1BD11BB033304" pageId="6" pageNumber="574" type="reference_group">
<paragraph id="94361F62FF985B4EF9B1BD11BAA53170" blockId="6.[151,1437,151,1977]" pageId="6" pageNumber="574">
For
<taxonomicName id="538964E1FF985B4EF982BD10BA453650" authority="Cossmann &amp; Peyrot (1912)" authorityName="Cossmann &amp; Peyrot" authorityYear="1912" box="[244,786,836,861]" class="Bivalvia" family="Cardiidae" genus="Cardium" kingdom="Animalia" order="Veneroida" pageId="6" pageNumber="574" phylum="Mollusca" rank="species" species="grateloupi">
<emphasis id="A6FDC370FF985B4EF982BD10B8863650" box="[244,465,836,860]" italics="true" pageId="6" pageNumber="574">Cardium grateloupi</emphasis>
,
<bibRefCitation id="F0186293FF985B4EF896BD10BA453650" author="Cossmann" box="[480,786,836,861]" pageId="6" pageNumber="574" refString="Cossmann, M. &amp; Peyrot, A. (1912) Conchologie neogenique de l'Aquitaine. Actes de la Societe Linneenne de Bordeaux, 65, 99 - 178." type="journal article" year="1912">Cossmann &amp; Peyrot (1912)</bibRefCitation>
</taxonomicName>
reported the following synonymy:
<taxonomicName id="538964E1FF985B4EFDEDBD10B817368C" authority="Basterot, 1825" authorityName="Basterot" authorityYear="1825" class="Bivalvia" family="Cardiidae" genus="Cardium" kingdom="Animalia" order="Veneroida" pageId="6" pageNumber="574" phylum="Mollusca" rank="species" species="burdigalinum">
<emphasis id="A6FDC370FF985B4EFDEDBD10BCCC3650" box="[1179,1435,836,860]" italics="true" pageId="6" pageNumber="574">Cardium burdigalinum</emphasis>
<bibRefCitation id="F0186293FF985B4EF9E1BD3CB817368C" author="Basterot" box="[151,320,871,896]" pageId="6" pageNumber="574" refString="Basterot, B. de (1825) Description geologique du bassin tertiaire du sud-ouest de la France. Memoires de la Societe d'Histoire naturelle de Paris, 2, 1 - 100." type="journal article" year="1825">Basterot, 1825</bibRefCitation>
</taxonomicName>
(
<emphasis id="A6FDC370FF985B4EF823BD3DB828368C" box="[341,383,873,896]" italics="true" pageId="6" pageNumber="574">non</emphasis>
Lamarck),
<taxonomicName id="538964E1FF985B4EFB7ABD3CBACE368C" authority="Grateloup, 1838" authorityName="Grateloup" authorityYear="1838" box="[524,921,871,896]" class="Bivalvia" family="Cardiidae" genus="Cardium" kingdom="Animalia" order="Veneroida" pageId="6" pageNumber="574" phylum="Mollusca" rank="species" species="hoernesianum">
<emphasis id="A6FDC370FF985B4EFB7ABD3CBB85368C" box="[524,722,872,896]" italics="true" pageId="6" pageNumber="574">C. hoernesianum</emphasis>
Grateloup, 1838
</taxonomicName>
(
<emphasis id="A6FDC370FF985B4EFAD9BD3DBA8E368C" box="[943,985,873,896]" italics="true" pageId="6" pageNumber="574">non</emphasis>
<taxonomicName id="538964E1FF985B4EFA90BD3CBDBE368C" authority="Deshayes" authorityName="Deshayes" box="[998,1257,871,896]" class="Bivalvia" family="Cardiidae" genus="Cardium" kingdom="Animalia" order="Veneroida" pageId="6" pageNumber="574" phylum="Mollusca" rank="species" species="hoernesi">
<emphasis id="A6FDC370FF985B4EFA90BD3CBD3B368C" box="[998,1132,872,896]" italics="true" pageId="6" pageNumber="574">C. hoernesi</emphasis>
Deshayes
</taxonomicName>
),
<taxonomicName id="538964E1FF985B4EFC77BD3CB87B36A9" authority="Mayer, 1866" authorityName="Mayer" authorityYear="1866" class="Bivalvia" family="Cardiidae" genus="Cardium" kingdom="Animalia" order="Veneroida" pageId="6" pageNumber="574" phylum="Mollusca" rank="species" species="grateloupi">
<emphasis id="A6FDC370FF985B4EFC77BD3CBCCB368C" box="[1281,1436,872,896]" italics="true" pageId="6" pageNumber="574">C. grateloupi</emphasis>
<bibRefCitation id="F0186293FF985B4EF9E1BDD9B87B36A9" author="Mayer" box="[151,300,908,933]" pageId="6" pageNumber="574" refString="Mayer, M. C. (1866) Description de coquilles fossiles des terrains tertiaires superieurs (suite). Journal de Conchyliologie, 14, 67 - 76." type="journal article" year="1866">Mayer, 1866</bibRefCitation>
</taxonomicName>
, and
<taxonomicName id="538964E1FF985B4EF804BDD8BBEA36A8" authority="Benoist, 1873" authorityName="Benoist" authorityYear="1873" box="[370,701,908,933]" class="Bivalvia" family="Cardiidae" genus="Cardium" kingdom="Animalia" order="Veneroida" pageId="6" pageNumber="574" phylum="Mollusca" rank="species" species="grateloupi">
<emphasis id="A6FDC370FF985B4EF804BDD8BB5A36A8" box="[370,525,908,932]" italics="true" pageId="6" pageNumber="574">C. grateloupi</emphasis>
<bibRefCitation id="F0186293FF985B4EFB6FBDD8BBEA36A8" author="Benoist" box="[537,701,908,933]" pageId="6" pageNumber="574" refString="Benoist, E. A. (1873) Catalogue synonymique er raisonne des Testaces fossiles recueillis dans les faluns miocenes des communes de La Brede et de Saucats. Actes de la Societe Linneenne de Bordeaux, New Series, 9, 5 - 77." type="journal article" year="1873">Benoist, 1873</bibRefCitation>
</taxonomicName>
.
<bibRefCitation id="F0186293FF985B4EFBBBBDD9BD0836A8" author="Basterot" box="[717,1119,908,933]" pageId="6" pageNumber="574" refString="Basterot, B. de (1825) Description geologique du bassin tertiaire du sud-ouest de la France. Memoires de la Societe d'Histoire naturelle de Paris, 2, 1 - 100." type="journal article" year="1825">Basterot (1825: 82, pl. 6, fig. 12)</bibRefCitation>
actually reported
<taxonomicName id="538964E1FF985B4EFC4FBDD8B87936C4" class="Bivalvia" family="Cardiidae" genus="Cardium" kingdom="Animalia" order="Veneroida" pageId="6" pageNumber="574" phylum="Mollusca" rank="species" species="burdigalinum">
<emphasis id="A6FDC370FF985B4EFC4FBDD8B87936C4" italics="true" pageId="6" pageNumber="574">Cardium burdigalinum</emphasis>
</taxonomicName>
from the area of Bordeaux and it really is Lamarcks
<emphasis id="A6FDC370FF985B4EFAE7BDE4BD7836C4" box="[913,1071,944,968]" italics="true" pageId="6" pageNumber="574">
<taxonomicName id="538964E1FF985B4EFAE7BDE4BD7C36C4" box="[913,1067,944,968]" class="Bivalvia" family="Cardiidae" genus="Cardium" kingdom="Animalia" order="Veneroida" pageId="6" pageNumber="574" phylum="Mollusca" rank="species" species="burdigalinum">burdigalinum</taxonomicName>
,
</emphasis>
as discussed below. The second synonymous entry, &quot;
<taxonomicName id="538964E1FF985B4EF80DBD80BA1736E0" authority="Grateloup, 1838" authorityName="Grateloup" authorityYear="1838" box="[379,832,980,1005]" class="Bivalvia" family="Cardiidae" genus="Cardium" kingdom="Animalia" order="Veneroida" pageId="6" pageNumber="574" phylum="Mollusca" rank="species" species="hoernesianum">
<emphasis id="A6FDC370FF985B4EF80DBD80BBD536E0" box="[379,642,980,1004]" italics="true" pageId="6" pageNumber="574">Cardium hoernesianum</emphasis>
Grateloup, 1838
</taxonomicName>
&quot; does not exist. According to the citation (&quot;
<emphasis id="A6FDC370FF985B4EFC50BD80BCDF36E7" box="[1318,1416,980,1003]" italics="true" pageId="6" pageNumber="574">Cat. Gir.</emphasis>
&quot;) by Cossmann &amp; Peyrot, it was published in the catalogue of fossils from Gironde (Grateloup &quot;1838&quot;; published in 1839:
<bibRefCitation id="F0186293FF985B4EF9A9BA49B8ED3138" author="Lesport" box="[223,442,1052,1076]" pageId="6" pageNumber="574" refString="Lesport, J. - F., Cluzaud, A. &amp; Verhecken, A. (2012) Les publications du Docteur Jean-Pierre Sylvestre de Grateloup sur les mollusques fossiles du Bassin d'Aquitaine (S. - O. France): dates de parutions et commentaires. Bulletin de la Societe Linneenne de Bordeaux, New Series, 147, 40, 417 - 485." type="journal article" year="2012">
Lesport
<emphasis id="A6FDC370FF985B4EF84BBA49B8393138" box="[317,366,1052,1076]" italics="true" pageId="6" pageNumber="574">et al</emphasis>
. 2012
</bibRefCitation>
), where only
<taxonomicName id="538964E1FF985B4EFB23BA48BA583138" box="[597,783,1052,1076]" class="Bivalvia" family="Cardiidae" genus="Cardium" kingdom="Animalia" order="Veneroida" pageId="6" pageNumber="574" phylum="Mollusca" rank="species" species="burdigalinum">
<emphasis id="A6FDC370FF985B4EFB23BA48BB3F313F" box="[597,616,1052,1075]" italics="true" pageId="6" pageNumber="574">C</emphasis>
.
<emphasis id="A6FDC370FF985B4EFB0EBA48BA583138" box="[632,783,1052,1076]" italics="true" pageId="6" pageNumber="574">burdigalinum</emphasis>
</taxonomicName>
was listed. The third entry evidently refers to the species from
<collectingCountry id="EC9E5FF2FF985B4EF9A2BA6BB8713154" box="[212,294,1087,1112]" name="Austria" pageId="6" pageNumber="574">Austria</collectingCountry>
renamed by
<bibRefCitation id="F0186293FF985B4EF8C0BA14BB043154" author="Mayer" box="[438,595,1087,1112]" pageId="6" pageNumber="574" refString="Mayer, M. C. (1866) Description de coquilles fossiles des terrains tertiaires superieurs (suite). Journal de Conchyliologie, 14, 67 - 76." type="journal article" year="1866">Mayer (1866)</bibRefCitation>
, while the last one is the record of
<taxonomicName id="538964E1FF985B4EFAA0BA14BD3C3154" box="[982,1131,1088,1112]" class="Bivalvia" family="Cardiidae" genus="Cardium" kingdom="Animalia" order="Veneroida" pageId="6" pageNumber="574" phylum="Mollusca" rank="species" species="grateloupi">
<emphasis id="A6FDC370FF985B4EFAA0BA14BABE315B" box="[982,1001,1088,1111]" italics="true" pageId="6" pageNumber="574">C</emphasis>
.
<emphasis id="A6FDC370FF985B4EFA80BA14BD3C3154" box="[1014,1131,1088,1112]" italics="true" pageId="6" pageNumber="574">grateloupi</emphasis>
</taxonomicName>
from the
<emphasis id="A6FDC370FF985B4EFDA1BA14BC463154" box="[1239,1297,1088,1112]" italics="true" pageId="6" pageNumber="574">falun</emphasis>
of La Bréde and Saucats by
<bibRefCitation id="F0186293FF985B4EF830BA30BB4D3171" author="Benoist" box="[326,538,1124,1149]" pageId="6" pageNumber="574" refString="Benoist, E. A. (1873) Catalogue synonymique er raisonne des Testaces fossiles recueillis dans les faluns miocenes des communes de La Brede et de Saucats. Actes de la Societe Linneenne de Bordeaux, New Series, 9, 5 - 77." type="journal article" year="1873">Benoist (1873: 45)</bibRefCitation>
, who evidently misapplied Mayer's name.
</paragraph>
<paragraph id="94361F62FF985B4EF9B1BADCBA003059" blockId="6.[151,1437,151,1977]" pageId="6" pageNumber="574">
<taxonomicName id="538964E1FF985B4EF9B1BADCB8B031AC" box="[199,487,1160,1184]" class="Bivalvia" family="Cardiidae" genus="Cardium" kingdom="Animalia" order="Veneroida" pageId="6" pageNumber="574" phylum="Mollusca" rank="species" species="grateloupianum" status="sp. nov.">
<emphasis id="A6FDC370FF985B4EF9B1BADCB8B031AC" box="[199,487,1160,1184]" italics="true" pageId="6" pageNumber="574">Cardium grateloupianum</emphasis>
</taxonomicName>
<emphasis id="A6FDC370FF985B4EF884BAD3BB6231AC" bold="true" box="[498,565,1159,1184]" pageId="6" pageNumber="574">
<taxonomicNameLabel id="BDCE7E0BFF985B4EF884BAD3BB6231AC" box="[498,565,1159,1184]" pageId="6" pageNumber="574" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
was dealt with by
<bibRefCitation id="F0186293FF985B4EFA6DBADDBA9631AC" author="Magne" box="[795,961,1159,1185]" pageId="6" pageNumber="574" refString="Magne, A. (1950) Les Ringicardium neogenes du Bassin d'Aquitaine. Journal de Conchyliologie, 90, 122 - 124." type="journal article" year="1950">Magne (1950)</bibRefCitation>
, as
<taxonomicName id="538964E1FF985B4EFA85BADCBCCE31AC" authority="Magne, 1944" authorityName="Magne" authorityYear="1944" box="[1011,1433,1160,1185]" class="Bivalvia" family="Cardiidae" genus="Ringicardium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Veneroida" pageId="6" pageNumber="574" phylum="Mollusca" rank="species" species="aquense">
<emphasis id="A6FDC370FF985B4EFA85BADCBDA431AC" box="[1011,1267,1160,1184]" italics="true" pageId="6" pageNumber="574">Ringicardium aquense</emphasis>
Magne, 1944
</taxonomicName>
. While the older citation refers to Magnes unpublished graduation thesis at the University of Bordeaux (1944), the same author failed to provide a nomenclatorially valid description in his later work;
<taxonomicName id="538964E1FF985B4EFD12BA85BCD731E4" authority="Magne, 1950" authorityName="Magne" authorityYear="1950" box="[1124,1408,1231,1256]" class="Bivalvia" family="Cardiidae" genus="Ringicardium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Veneroida" pageId="6" pageNumber="574" phylum="Mollusca" rank="species" species="aquense">
<emphasis id="A6FDC370FF985B4EFD12BA85BD2231E4" box="[1124,1141,1233,1256]" italics="true" pageId="6" pageNumber="574">R</emphasis>
.
<emphasis id="A6FDC370FF985B4EFDF3BA85BDB631E4" box="[1157,1249,1233,1256]" italics="true" pageId="6" pageNumber="574">aquense</emphasis>
<bibRefCitation id="F0186293FF985B4EFD9CBA84BCD731E4" author="Magne" box="[1258,1408,1231,1256]" pageId="6" pageNumber="574" refString="Magne, A. (1950) Les Ringicardium neogenes du Bassin d'Aquitaine. Journal de Conchyliologie, 90, 122 - 124." type="journal article" year="1950">Magne, 1950</bibRefCitation>
</taxonomicName>
is considered a
<emphasis id="A6FDC370FF985B4EF85CBAA1B89F3000" box="[298,456,1268,1292]" italics="true" pageId="6" pageNumber="574">nomen nudum</emphasis>
, therefore. However, Magne has the merit of having realized that the species illustrated by
<bibRefCitation id="F0186293FF985B4EF9CABB4CB8A4303C" author="Cossmann" box="[188,499,1304,1328]" pageId="6" pageNumber="574" refString="Cossmann, M. &amp; Peyrot, A. (1912) Conchologie neogenique de l'Aquitaine. Actes de la Societe Linneenne de Bordeaux, 65, 99 - 178." type="journal article" year="1912">Cossmann &amp; Peyrot (1912)</bibRefCitation>
as
<taxonomicName id="538964E1FF985B4EFB6ABB4CBBAB303C" box="[540,764,1304,1328]" class="Bivalvia" family="Cardiidae" genus="Cardium" kingdom="Animalia" order="Veneroida" pageId="6" pageNumber="574" phylum="Mollusca" rank="species" species="grateloupi">
<emphasis id="A6FDC370FF985B4EFB6ABB4CBBAB303C" box="[540,764,1304,1328]" italics="true" pageId="6" pageNumber="574">Cardium grateloupi</emphasis>
</taxonomicName>
had never been described and that
<taxonomicName id="538964E1FF985B4EFDE4BB4CBC04303C" box="[1170,1363,1304,1328]" class="Bivalvia" family="Cardiidae" genus="Cardium" kingdom="Animalia" order="Veneroida" pageId="6" pageNumber="574" phylum="Mollusca" rank="species" species="hoernesianum">
<emphasis id="A6FDC370FF985B4EFDE4BB4CBC04303C" box="[1170,1363,1304,1328]" italics="true" pageId="6" pageNumber="574">C. hoernesianum</emphasis>
</taxonomicName>
was a replacement name for a distinct Miocene species from
<collectingCountry id="EC9E5FF2FF985B4EFB8BBB68BA043059" box="[765,851,1340,1365]" name="Austria" pageId="6" pageNumber="574">Austria</collectingCountry>
.
</paragraph>
<paragraph id="94361F62FF985B4EF9B1BB0BBB033304" blockId="6.[151,1437,151,1977]" pageId="6" pageNumber="574">
<materialsCitation id="24E1153FFF985B4EF9B1BB0BBA4E3091" ID-GBIF-Occurrence="1328377999" country="Hungary" location="Upper Oligocene" pageId="6" pageNumber="574" specimenCount="1" stateProvince="Eger">
<bibRefCitation id="F0186293FF985B4EF9B1BB0BBB333074" author="Noszky" box="[199,612,1375,1400]" pageId="6" pageNumber="574" refString="Noszky, J. (1936) Az egri felso cattien molluszkafaunaja. Die Molluskenfauna des oberen Cattiens von Eger, in Ungarn. Annales Musei Nationalis Hungarici. Pars Mineralogica, Geologica, Paleontologica, 30, 53 - 115." type="journal article" year="1936">Noszky (1936: 88, 111, pl. 6, fig. 7)</bibRefCitation>
described
<taxonomicName id="538964E1FF985B4EFB96BB34BC653074" box="[736,1330,1376,1400]" class="Bivalvia" family="Cardiidae" genus="Cardium" kingdom="Animalia" order="Veneroida" pageId="6" pageNumber="574" phylum="Mollusca" rank="variety" species="grateloupi" subGenus="Ringicardium" variety="oligocenica">
<emphasis id="A6FDC370FF985B4EFB96BB34BD223074" box="[736,1141,1376,1400]" italics="true" pageId="6" pageNumber="574">Cardium (Ringicardium) grateloupi</emphasis>
var.
<emphasis id="A6FDC370FF985B4EFDD9BB34BC653074" box="[1199,1330,1376,1400]" italics="true" pageId="6" pageNumber="574">oligocenica</emphasis>
</taxonomicName>
from the Egerian (Chattian,
<location id="915649B9FF985B4EF802BBD1BB6D3091" LSID="urn:lsid:plazi:treatment:1C20AE74FF9C5B41F9E1B925BB5E3495:915649B9FF985B4EF802BBD1BB6D3091" box="[372,570,1412,1437]" country="Hungary" name="Upper Oligocene" pageId="6" pageNumber="574" stateProvince="Eger">Upper Oligocene</location>
) of
<collectingRegion id="564DD180FF985B4EFB19BBD1BBF13091" box="[623,678,1413,1437]" country="Hungary" name="Eger" pageId="6" pageNumber="574">Eger</collectingRegion>
,
<collectingCountry id="EC9E5FF2FF985B4EFBC3BBD1BA4E3091" box="[693,793,1413,1437]" name="Hungary" pageId="6" pageNumber="574">Hungary</collectingCountry>
</materialsCitation>
. The new variety was compared with
<taxonomicName id="538964E1FF985B4EFDA9BBD0BC2E3090" box="[1247,1401,1412,1436]" class="Bivalvia" family="Cardiidae" genus="Cardium" kingdom="Animalia" order="Veneroida" pageId="6" pageNumber="574" phylum="Mollusca" rank="species" species="grateloupi">
<emphasis id="A6FDC370FF985B4EFDA9BBD0BC2E3090" box="[1247,1401,1412,1436]" italics="true" pageId="6" pageNumber="574">C. grateloupi</emphasis>
</taxonomicName>
of
<bibRefCitation id="F0186293FF985B4EF9E1BBFCB89930CC" author="Cossmann" box="[151,462,1448,1472]" pageId="6" pageNumber="574" refString="Cossmann, M. &amp; Peyrot, A. (1912) Conchologie neogenique de l'Aquitaine. Actes de la Societe Linneenne de Bordeaux, 65, 99 - 178." type="journal article" year="1912">Cossmann &amp; Peyrot (1912)</bibRefCitation>
but the original material, consisting of a badly preserved internal mould, is definitely different from
<taxonomicName id="538964E1FF985B4EF832BB98BB2930E8" box="[324,638,1484,1508]" class="Bivalvia" family="Cardiidae" genus="Bucardium" kingdom="Animalia" order="Veneroida" pageId="6" pageNumber="574" phylum="Mollusca" rank="species" species="grateloupianum" status="sp. nov.">
<emphasis id="A6FDC370FF985B4EF832BB98BB2930E8" box="[324,638,1484,1508]" italics="true" pageId="6" pageNumber="574">Bucardium grateloupianum</emphasis>
</taxonomicName>
<emphasis id="A6FDC370FF985B4EFBFCBB98BB9A30E9" bold="true" box="[650,717,1484,1509]" pageId="6" pageNumber="574">
<taxonomicNameLabel id="BDCE7E0BFF985B4EFBFCBB98BB9A30E9" box="[650,717,1484,1509]" pageId="6" pageNumber="574" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
Further comments on this taxon and other cardiids from the Egerian of Hungary are reported below.
</paragraph>
</subSubSection>
<subSubSection id="DC934CE9FF985B4EF9B1B840BBBF3378" pageId="6" pageNumber="574" type="description">
<paragraph id="94361F62FF985B4EF9B1B840BBBF3378" blockId="6.[151,1437,151,1977]" pageId="6" pageNumber="574">
Finally,
<taxonomicName id="538964E1FF985B4EF852B840BBCA3321" authority="Mayer, 1866" authorityName="Mayer" authorityYear="1866" box="[292,669,1556,1581]" class="Bivalvia" family="Cardiidae" genus="Cardium" kingdom="Animalia" order="Veneroida" pageId="6" pageNumber="574" phylum="Mollusca" rank="species" species="grateloupi">
<emphasis id="A6FDC370FF985B4EF852B840BB543320" box="[292,515,1556,1580]" italics="true" pageId="6" pageNumber="574">Cardium grateloupi</emphasis>
<bibRefCitation id="F0186293FF985B4EFB7DB841BBCA3321" author="Mayer" box="[523,669,1556,1581]" pageId="6" pageNumber="574" refString="Mayer, M. C. (1866) Description de coquilles fossiles des terrains tertiaires superieurs (suite). Journal de Conchyliologie, 14, 67 - 76." type="journal article" year="1866">Mayer, 1866</bibRefCitation>
</taxonomicName>
, is a primary homonym of
<taxonomicName id="538964E1FF985B4EFAA5B840BC673320" authority="Michelotti, 1839" authorityName="Michelotti" authorityYear="1839" box="[979,1328,1556,1581]" class="Bivalvia" family="Cardiidae" genus="Cardium" kingdom="Animalia" order="Veneroida" pageId="6" pageNumber="574" phylum="Mollusca" rank="species" species="grateloupi">
<emphasis id="A6FDC370FF985B4EFAA5B840BD3D3320" box="[979,1130,1556,1580]" italics="true" pageId="6" pageNumber="574">C. grateloupi</emphasis>
<bibRefCitation id="F0186293FF985B4EFD04B840BC673320" author="Michelotti" box="[1138,1328,1556,1581]" pageId="6" pageNumber="574" refString="Michelotti, G. (1839). Brevi cenni di alcuni resti delle classi Brachiopodi ed Acefali, trovati fossili in Italia. Annali delle Scienze del Regno Lombardo-Veneto, 9, 119 - 174." type="journal article" year="1839">Michelotti, 1839</bibRefCitation>
</taxonomicName>
, which is considered a junior synonym of
<taxonomicName id="538964E1FF985B4EFB79B86DBA06335C" authority="Brocchi, 1814" authorityName="Brocchi" authorityYear="1814" box="[527,849,1591,1616]" class="Bivalvia" family="Veneridae" genus="Venus" kingdom="Animalia" order="Veneroida" pageId="6" pageNumber="574" phylum="Mollusca" rank="species" species="cypria">
<emphasis id="A6FDC370FF985B4EFB79B86DBBF73343" box="[527,672,1592,1616]" italics="true" pageId="6" pageNumber="574">Venus cypria</emphasis>
<bibRefCitation id="F0186293FF985B4EFBDAB863BA06335C" author="Brocchi" box="[684,849,1591,1616]" pageId="6" pageNumber="574" refString="Brocchi, G. (1814) Conchiologia fossile subappennina con osservazioni geologiche sugli Appennini e sul suolo adiacente. Vol. 1 &amp; 2. Stamperia Reale, Milano, 472 pp. http: // dx. doi. org / 10.5962 / bhl. title. 11569" type="book" year="1814">Brocchi, 1814</bibRefCitation>
</taxonomicName>
, now
<taxonomicName id="538964E1FF985B4EFAEFB86CBDCB3343" box="[921,1180,1592,1616]" class="Bivalvia" family="Cardiidae" genus="Nemocardium" kingdom="Animalia" order="Veneroida" pageId="6" pageNumber="574" phylum="Mollusca" rank="species" species="cyprium">
<emphasis id="A6FDC370FF985B4EFAEFB86CBDCB3343" box="[921,1180,1592,1616]" italics="true" pageId="6" pageNumber="574">Nemocardium cyprium</emphasis>
</taxonomicName>
, from the Pliocene of
<collectingCountry id="EC9E5FF2FF985B4EF9E1B808B99C3379" box="[151,203,1628,1653]" name="Italy" pageId="6" pageNumber="574">Italy</collectingCountry>
(
<bibRefCitation id="F0186293FF985B4EF9AEB808B8DB3378" author="Hylleberg" box="[216,396,1628,1653]" pageId="6" pageNumber="574" refString="Hylleberg, J. (2004) Lexical approach to Cardiacea. Vol. 2. Records. Annotated and illustrated, AM. Phuket Marine Biological Center Special Publication, 30, 352645." type="journal article" year="2004">Hylleberg 2004</bibRefCitation>
; La Perna &amp; DAbramo 2011).
</paragraph>
</subSubSection>
<subSubSection id="DC934CE9FF985B41F9B1B82BBB5E3495" lastPageId="9" lastPageNumber="577" pageId="6" pageNumber="574" type="distribution">
<paragraph id="94361F62FF985B4EF9B1B82BB9B532B5" blockId="6.[151,1437,151,1977]" pageId="6" pageNumber="574">
<emphasis id="A6FDC370FF985B4EF9B1B82BB8373394" bold="true" box="[199,352,1663,1688]" pageId="6" pageNumber="574">Distribution.</emphasis>
In addition to Saint-Paul-lès-Dax and Mérignac, from where the
<typeStatus id="4B32A1C0FF985B4EFD32B8D4BD233394" box="[1092,1140,1664,1688]" pageId="6" pageNumber="574">type</typeStatus>
material comes, the study material is from Lèognan, Saucats and Salles, all well known fossiliferous localities in the
<collectingRegion id="564DD180FF985B4EFDE6B8F0BDA933B1" box="[1168,1278,1700,1725]" country="France" name="Aquitaine" pageId="6" pageNumber="574">Aquitaine</collectingRegion>
basin (
<bibRefCitation id="F0186293FF985B4EFC26B8F0B98433EC" author="Folliot" pageId="6" pageNumber="574" refString="Folliot M., 1989. Les depots fossiliferes du Miocene du Bordelais. Mineraux et Fossiles, 163, 17 - 28." type="journal article" year="1989">Folliot 1989</bibRefCitation>
). The outcrops in the areas of Lèognan and Saucats are of Burdigalian age, though some of the Saucats outcrops include Late Aquitanian beds, while the outcrops in the area of Salles are Serravallian (
<bibRefCitation id="F0186293FF985B4EFD85B8B8B9853224" author="Poignant" pageId="6" pageNumber="574" refString="Poignant, A., Pujol, C., Ringeade, M. &amp; Londeix, L. (1997) The Burdigalian historical stratotype. In: Montanari, A., Odin G. S. &amp; Coccioni, R. (Eds.), Miocene stratigraphy: an integrated approach. Elsevier, Amsterdam, pp. 17 - 24." type="book chapter" year="1997">
Poignant
<emphasis id="A6FDC370FF985B4EFC17B8B9BCC23208" box="[1377,1429,1772,1796]" italics="true" pageId="6" pageNumber="574">et al</emphasis>
. 1997
</bibRefCitation>
;
<bibRefCitation id="F0186293FF985B4EF9A8B95BBB533224" author="Cahuzac" box="[222,516,1807,1832]" pageId="6" pageNumber="574" refString="Cahuzac, B. &amp; Cluzaud, A. (1999) Bilan scientifique et synthese des donnees bibliographiques sur le Miocene moyen (Serravallien) de Saucats (Gironde, Bassin d'Aquitaine). Bulletin de la Societe Linneenne de Bordeaux, 27, 95 - 111." type="journal article" year="1999">Cahuzac &amp; Cluzaud 1999</bibRefCitation>
;
<bibRefCitation id="F0186293FF985B4EFB66B95BBA643224" author="Cahuzac" box="[528,819,1807,1832]" pageId="6" pageNumber="574" refString="Cahuzac, B. &amp; Janssen, A. W. (2010) Eocene to Miocene holoplanktonic Mollusca (Gastropoda) of the Aquitaine Basin, southwest France. Scripta Geologica, 141, 1 - 192." type="journal article" year="2010">Cahuzac &amp; Jannsen 2010</bibRefCitation>
). According to
<bibRefCitation id="F0186293FF985B4EFA97B944BC403224" author="Cossmann" box="[993,1303,1808,1832]" pageId="6" pageNumber="574" refString="Cossmann, M. &amp; Peyrot, A. (1912) Conchologie neogenique de l'Aquitaine. Actes de la Societe Linneenne de Bordeaux, 65, 99 - 178." type="journal article" year="1912">Cossmann &amp; Peyrot (1912)</bibRefCitation>
and
<bibRefCitation id="F0186293FF985B4EFC38B944B9B43241" author="Magne" pageId="6" pageNumber="574" refString="Magne, A. (1950) Les Ringicardium neogenes du Bassin d'Aquitaine. Journal de Conchyliologie, 90, 122 - 124." type="journal article" year="1950">Magne (1950)</bibRefCitation>
, this species is only known from Aquitanian and Burdigalian deposits. Most probably, the &quot;Serravallian&quot; material (NHMW) was mislabelled or its locality inaccurately recorded, considering that in some Miocene localities in SW
<collectingCountry id="EC9E5FF2FF985B4EF822B929B8F43298" box="[340,419,1917,1940]" name="France" pageId="6" pageNumber="574">France</collectingCountry>
, the Burdigalian deposits are directly overlain by Serravallian beds (
<bibRefCitation id="F0186293FF985B4EFDDBB928BCCF3298" author="Poignant" box="[1197,1432,1916,1941]" pageId="6" pageNumber="574" refString="Poignant, A., Pujol, C., Ringeade, M. &amp; Londeix, L. (1997) The Burdigalian historical stratotype. In: Montanari, A., Odin G. S. &amp; Coccioni, R. (Eds.), Miocene stratigraphy: an integrated approach. Elsevier, Amsterdam, pp. 17 - 24." type="book chapter" year="1997">
Poignant
<emphasis id="A6FDC370FF985B4EFC6DB929BC1A3298" box="[1307,1357,1916,1940]" italics="true" pageId="6" pageNumber="574">et al</emphasis>
. 1997
</bibRefCitation>
: fig. 5).
</paragraph>
<caption id="C0F64FEAFF965B40F9E1BE81BBF834CE" pageId="8" pageNumber="576">
<paragraph id="94361F62FF965B40F9E1BE81BBF834CE" blockId="8.[151,1437,213,450]" pageId="8" pageNumber="576">
<emphasis id="A6FDC370FF965B40F9E1BE81B84735E6" bold="true" box="[151,272,213,235]" pageId="8" pageNumber="576">FIGURE 5</emphasis>
.
<taxonomicName id="538964E1FF965B40F869BE81B82F35E7" box="[287,376,213,235]" class="Bivalvia" family="Cardiidae" genus="Cardium" kingdom="Animalia" order="Veneroida" pageId="8" pageNumber="576" phylum="Mollusca" rank="genus">
<emphasis id="A6FDC370FF965B40F869BE81B82F35E7" box="[287,376,213,235]" italics="true" pageId="8" pageNumber="576">Cardium</emphasis>
</taxonomicName>
spp. from the Eggenburgian of Austria.
<emphasis id="A6FDC370FF965B40FA50BE81BA0E35E6" bold="true" box="[806,857,213,234]" pageId="8" pageNumber="576">A, B</emphasis>
. Syntype of
<taxonomicName id="538964E1FF965B40FA95BE81BC3B35E7" authority="Hornes, 1862" authorityName="Hornes" authorityYear="1862" box="[995,1388,213,235]" class="Bivalvia" family="Cardiidae" genus="Cardium" kingdom="Animalia" order="Veneroida" pageId="8" pageNumber="576" phylum="Mollusca" rank="species" species="hoernesianum">
<emphasis id="A6FDC370FF965B40FA95BE81BD8235E7" box="[995,1237,213,235]" italics="true" pageId="8" pageNumber="576">Cardium hoernesianum</emphasis>
Hörnes, 1862
</taxonomicName>
, LV, Gaudendorf (NHMW 1851.VI.41) (Hörnes 1862: pl. 27, figs 1a, c).
<emphasis id="A6FDC370FF965B40FA17BEA0BA243406" bold="true" box="[865,883,244,266]" pageId="8" pageNumber="576">C</emphasis>
. Syntype of
<taxonomicName id="538964E1FF965B40FA8ABEA0B8623424" authority="Schaffer, 1910" authorityName="Schaffer" authorityYear="1910" class="Bivalvia" family="Cardiidae" genus="Ringicardium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Veneroida" pageId="8" pageNumber="576" phylum="Mollusca" rank="variety" species="burdigalinum" variety="grandis">
<emphasis id="A6FDC370FF965B40FA8ABEA0BC4F3406" box="[1020,1304,244,266]" italics="true" pageId="8" pageNumber="576">Ringicardium burdigalinum</emphasis>
var.
<emphasis id="A6FDC370FF965B40FC38BEA0BCCC3406" box="[1358,1435,244,266]" italics="true" pageId="8" pageNumber="576">grandis</emphasis>
Schaffer, 1910
</taxonomicName>
, LV, Loibersdorf (NHMW 1866.XL.20) (Schaffer 1910: pl. 30, fig. 4).
<emphasis id="A6FDC370FF965B40FD12BF47BD213424" bold="true" box="[1124,1142,275,296]" pageId="8" pageNumber="576">D</emphasis>
. Syntype of
<taxonomicName id="538964E1FF965B40FC64BF46BB6A344B" authority="Schaffer, 1910" authorityName="Schaffer" authorityYear="1910" class="Bivalvia" family="Cardiidae" genus="Ringicardium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Veneroida" pageId="8" pageNumber="576" phylum="Mollusca" rank="variety" species="burdigalinum" variety="grandis">
<emphasis id="A6FDC370FF965B40FC64BF46B877344B" italics="true" pageId="8" pageNumber="576">Ringicardium burdigalinum</emphasis>
var.
<emphasis id="A6FDC370FF965B40F824BF65B8C8344B" box="[338,415,305,327]" italics="true" pageId="8" pageNumber="576">grandis</emphasis>
Schaffer, 1910
</taxonomicName>
, RV, Loibersdorf (Hörnes 1862: pl. 27, figs 2a, b; Schaffer 1910: pl. 30, fig. 2, 3).
<emphasis id="A6FDC370FF965B40FCF2BF65BCC2344A" bold="true" box="[1412,1429,305,326]" pageId="8" pageNumber="576">E</emphasis>
.
<taxonomicName id="538964E1FF965B40F9E1BF04B9A7346A" box="[151,240,336,358]" class="Bivalvia" family="Cardiidae" genus="Cardium" kingdom="Animalia" order="Veneroida" pageId="8" pageNumber="576" phylum="Mollusca" rank="genus">
<emphasis id="A6FDC370FF965B40F9E1BF04B9A7346A" box="[151,240,336,358]" italics="true" pageId="8" pageNumber="576">Cardium</emphasis>
</taxonomicName>
sp., RV Loibersdorf (no registration number).
<emphasis id="A6FDC370FF965B40FBA1BF04BBB33469" bold="true" box="[727,740,336,357]" pageId="8" pageNumber="576">F</emphasis>
. Eggenburg (no registration number).
<emphasis id="A6FDC370FF965B40FD02BF04BDD2346A" bold="true" box="[1140,1157,336,358]" pageId="8" pageNumber="576">G</emphasis>
. Anterior sculpture of Fig. 5D.
<emphasis id="A6FDC370FF965B40F9B5BF3BB9803488" bold="true" box="[195,215,367,388]" pageId="8" pageNumber="576">H</emphasis>
. Anterior sculpure of Fig. 5C.
<emphasis id="A6FDC370FF965B40FB62BF3BBB493488" bold="true" box="[532,542,367,388]" pageId="8" pageNumber="576">I</emphasis>
. Posterior sculpture of Fig. 5A.
<emphasis id="A6FDC370FF965B40FA1FBF3BBA223488" bold="true" box="[873,885,367,388]" pageId="8" pageNumber="576">J</emphasis>
. Posterior sculpture of Fig. 5D.
<emphasis id="A6FDC370FF965B40FDB7BF3BBDAD3488" bold="true" box="[1217,1274,367,388]" pageId="8" pageNumber="576">KM</emphasis>
. Illustrations of
<taxonomicName id="538964E1FF965B40F9E1BFD9BB4934AF" authority="Hornes, 1862" authorityName="Hornes" authorityYear="1862" box="[151,542,397,419]" class="Bivalvia" family="Cardiidae" genus="Cardium" kingdom="Animalia" order="Veneroida" pageId="8" pageNumber="576" phylum="Mollusca" rank="species" species="hoernesianum">
<emphasis id="A6FDC370FF965B40F9E1BFD9B8DF34AF" box="[151,392,397,419]" italics="true" pageId="8" pageNumber="576">Cardium hoernesianum</emphasis>
Hörnes, 1862
</taxonomicName>
(after Hoernes 1862: pl. 27, figs 1ac).
<emphasis id="A6FDC370FF965B40FABDBFD9BAA834AF" bold="true" box="[971,1023,397,419]" pageId="8" pageNumber="576">N, O</emphasis>
. Illustrations of
<taxonomicName id="538964E1FF965B40FDC7BFD9B86734CE" authority="Lamarck, 1819" authorityName="Lamarck" authorityYear="1819" class="Bivalvia" family="Cardiidae" genus="Cardium" kingdom="Animalia" order="Veneroida" pageId="8" pageNumber="576" phylum="Mollusca" rank="species" species="burdigalinum">
<emphasis id="A6FDC370FF965B40FDC7BFD9BCCB34AF" box="[1201,1436,397,419]" italics="true" pageId="8" pageNumber="576">Cardium burdigalinum</emphasis>
Lamarck, 1819
</taxonomicName>
(after Hörnes 1862: pl. 27, figs 2a,b).
</paragraph>
</caption>
<caption id="C0F64FEAFF965B40F9E1B978B80D3292" httpUri="https://zenodo.org/record/272524/files/figure.png" pageId="8" pageNumber="576" targetBox="[151,1436,499,1814]" targetPageId="8">
<paragraph id="94361F62FF965B40F9E1B978B80D3292" blockId="8.[151,1436,1836,1950]" pageId="8" pageNumber="576">
<emphasis id="A6FDC370FF965B40F9E1B978B844324D" bold="true" box="[151,275,1836,1858]" pageId="8" pageNumber="576">FIGURE 6</emphasis>
.
<taxonomicName id="538964E1FF965B40F853B978BB94324E" authority="Lamarck, 1819" authorityName="Lamarck" authorityYear="1819" box="[293,707,1836,1858]" class="Bivalvia" family="Cardiidae" genus="Cardium" kingdom="Animalia" order="Veneroida" pageId="8" pageNumber="576" phylum="Mollusca" rank="species" species="burdigalinum">
<emphasis id="A6FDC370FF965B40F853B978BB43324E" box="[293,532,1836,1858]" italics="true" pageId="8" pageNumber="576">Cardium burdigalinum</emphasis>
Lamarck, 1819
</taxonomicName>
.
<emphasis id="A6FDC370FF965B40FBA4B978BA5F324E" bold="true" box="[722,776,1836,1858]" pageId="8" pageNumber="576">A, B</emphasis>
. Syntype, surroundings of Bordeaux, RV (MNHN.F.A50146, Lamarck coll. 200814), photo E. Porez (http://coldb.mnhn.fr/catalognumber/mnhn/f/a50146).
<emphasis id="A6FDC370FF965B40FD2EB91FBDDF326C" bold="true" box="[1112,1160,1867,1889]" pageId="8" pageNumber="576">C, D</emphasis>
. Léognan, RV (MPUB).
<emphasis id="A6FDC370FF965B40FCF2B91FB9F33273" bold="true" pageId="8" pageNumber="576">E, F</emphasis>
. Léognan, LV (MPUB).
<emphasis id="A6FDC370FF965B40F8C5B93EB893328C" bold="true" box="[435,452,1898,1920]" pageId="8" pageNumber="576">G</emphasis>
. Léognan, RV, detail of posterior sculpture (MPUB).
<emphasis id="A6FDC370FF965B40FD71B93EBD613273" bold="true" box="[1031,1078,1898,1919]" pageId="8" pageNumber="576">H, I</emphasis>
. Léognan, RV, details of anterior sculpture (MPUB).
</paragraph>
</caption>
<paragraph id="94361F62FF975B41F9B1BEC3B8CF3419" blockId="9.[151,1436,151,410]" pageId="9" pageNumber="577">
The species was recently described as
<taxonomicName id="538964E1FF975B41FB04BECCBA1835BC" box="[626,847,152,176]" class="Bivalvia" family="Cardiidae" genus="Cardium" kingdom="Animalia" order="Veneroida" pageId="9" pageNumber="577" phylum="Mollusca" rank="species" species="grateloupi">
<emphasis id="A6FDC370FF975B41FB04BECCBA1835BC" box="[626,847,152,176]" italics="true" pageId="9" pageNumber="577">Cardium grateloupi</emphasis>
</taxonomicName>
by
<bibRefCitation id="F0186293FF975B41FA0EBECCBDF635BC" author="Lesport" box="[888,1185,151,176]" pageId="9" pageNumber="577" refString="Lesport, J. - F. &amp; Cahuzac, B. (2005) Decouverte des sables a Mactres de la base du Burdigalien a Martillac (Nord Aquitaine; SW France); donnees paleontologiques. Annales de Paleontologie, 91, 73 - 116. http: // dx. doi. org / 10.1016 / j. annpal. 2004.11.004" type="journal article" year="2005">Lesport &amp; Cahuzac (2005)</bibRefCitation>
from a rich molluscan assemblage of Early Burdigalian age at Martillac, south of Bordeaux. It is a shallow water assemblage of sandy substrate, comparable to the modern
<emphasis id="A6FDC370FF975B41FB35BE8FBA2735FF" box="[579,880,219,243]" italics="true" pageId="9" pageNumber="577">Sables Fins Bien Calibrés</emphasis>
community, as defined in the Mediterranean by
<bibRefCitation id="F0186293FF975B41F9E1BEA8B8C43419" author="Peres" box="[151,403,252,277]" pageId="9" pageNumber="577" refString="Peres, J. M. &amp; Picard, J. (1964) Nouveau manuel de bionomie benthique. Recueil des Travaux de la Station marine d'Endoume, 31, 5 - 137." type="journal article" year="1964">Pérès &amp; Picard (1964)</bibRefCitation>
.
</paragraph>
<paragraph id="94361F62FF975B41F9B1BF49BB5E3495" blockId="9.[151,1436,151,410]" pageId="9" pageNumber="577">
The living
<taxonomicName id="538964E1FF975B41F830BF4AB8E63439" box="[326,433,286,309]" class="Bivalvia" family="Cardiidae" genus="Bucardium" kingdom="Animalia" order="Veneroida" pageId="9" pageNumber="577" phylum="Mollusca" rank="species" species="ringen">
<emphasis id="A6FDC370FF975B41F830BF4AB8E63439" box="[326,433,286,309]" italics="true" pageId="9" pageNumber="577">B. ringen</emphasis>
</taxonomicName>
s, a tropical species as indicated by its West African distribution, is probably the direct descent of
<taxonomicName id="538964E1FF975B41F86CBF6BB8A6345A" box="[282,497,318,342]" class="Bivalvia" family="Cardiidae" genus="Bucardium" kingdom="Animalia" order="Veneroida" pageId="9" pageNumber="577" phylum="Mollusca" rank="species" species="grateloupianum" status="sp. nov.">
<emphasis id="A6FDC370FF975B41F86CBF6BB8A6345A" box="[282,497,318,342]" italics="true" pageId="9" pageNumber="577">B. grateloupianum</emphasis>
</taxonomicName>
<emphasis id="A6FDC370FF975B41F88BBF6ABB17345B" bold="true" box="[509,576,318,343]" pageId="9" pageNumber="577">
<taxonomicNameLabel id="BDCE7E0BFF975B41F88BBF6ABB17345B" box="[509,576,318,343]" pageId="9" pageNumber="577" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, which evidently also had warm water preferences, as inferred from the palaeoclimatic and palaeogeographic characteristics of the Lower Miocene fauna of the
<collectingRegion id="564DD180FF975B41FD09BF0BBDBA3474" box="[1151,1261,351,376]" country="France" name="Aquitaine" pageId="9" pageNumber="577">Aquitaine</collectingRegion>
basin (Lauriat- Rage
<emphasis id="A6FDC370FF975B41F9A1BFD6B8473495" box="[215,272,385,409]" italics="true" pageId="9" pageNumber="577">et al.</emphasis>
1993;
<bibRefCitation id="F0186293FF975B41F82BBFD6B8AA3495" author="Lozouet" box="[349,509,385,409]" pageId="9" pageNumber="577" refString="Lozouet, P. (2014) Temporal and latitudinal trends in the biodiversity of European Atlantic Cenozoic gastropod (Mollusca) faunas. A base for the history of biogeographic provinces. Carnets de Geologie (Notebooks on Geology), 14 (14), 273 - 314. http: // dx. doi. org / 10.4267 / 2042 / 54150" type="journal article" year="2014">Lozouet 2014</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>