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<document id="5BE17EEC7B8E6C8E766A6090055EF1F4" ID-DOI="10.15298/rusentj.28.2.04" ID-ISSN="0132-8069" ID-Zenodo-Dep="13163815" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1722579156484" checkinUser="felipe" docAuthor="Gildenkov, M. Yu." docDate="2019" docId="1C2087FB83231F176A58FE71FAE0FE6D" docLanguage="en" docName="RussEntomolJ.28.2.138-145.pdf" docOrigin="Russian Entomological Journal 28 (2)" docSource="http://dx.doi.org/10.15298/rusentj.28.2.04" docStyle="DocumentStyle:0D5BAC5AA0C130B3809AF578A601D837.2:RussEntomolJ.2019-.journal_article" docStyleId="0D5BAC5AA0C130B3809AF578A601D837" docStyleName="RussEntomolJ.2019-.journal_article" docStyleVersion="2" docTitle="Carpelimus (Trogophloeus) keraliensis Gildenkov, 2019, sp.n." docType="treatment" docVersion="2" lastPageNumber="142" masterDocId="E019FF8383201F136B55FF9BFF97FFEA" masterDocTitle="Seven new species of the genus Carpelimus Leach, 1819 from the “ taprobanae ” group (Coleoptera: Staphylinidae: Oxytelinae)" masterLastPageNumber="145" masterPageNumber="138" pageNumber="141" updateTime="1722580280181" updateUser="ExternalLinkService">
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<mods:title id="9EF63C2906452F01B3B901FEFF3EBDA6">Seven new species of the genus Carpelimus Leach, 1819 from the “ taprobanae ” group (Coleoptera: Staphylinidae: Oxytelinae)</mods:title>
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<mods:namePart id="1D4D3AA10CE4B0C35D8DC03BF0D0FB57">Gildenkov, M. Yu.</mods:namePart>
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<mods:title id="720EFF7EBBA73F4F1803D69F853FD93E">Russian Entomological Journal</mods:title>
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<mods:part id="060562B989341A1C2AB87947F831D851">
<mods:date id="C372783103FD3B7AA1B35F783847D3A6">2019</mods:date>
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<mods:number id="A0A392C810B007E74C890BBE65490389">2019-09-30</mods:number>
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<mods:classification id="CBC7F92367E732B13B02E1C0904039C8">journal article</mods:classification>
<mods:identifier id="CAEFABFAA0936EBC3341E883A8932BDE" type="DOI">10.15298/rusentj.28.2.04</mods:identifier>
<mods:identifier id="92623A5AAA8DD2C3E78AEA3097F456C0" type="ISSN">0132-8069</mods:identifier>
<mods:identifier id="28C545559651AE067C4BA66EEC9E60B4" type="Zenodo-Dep">13163815</mods:identifier>
</mods:mods>
<treatment id="1C2087FB83231F176A58FE71FAE0FE6D" LSID="urn:lsid:plazi:treatment:1C2087FB83231F176A58FE71FAE0FE6D" httpUri="http://treatment.plazi.org/id/1C2087FB83231F176A58FE71FAE0FE6D" lastPageId="4" lastPageNumber="142" pageId="3" pageNumber="141">
<subSubSection id="DC93656683231F106A58FE71FDACFDCA" pageId="3" pageNumber="141" type="nomenclature">
<paragraph id="943636ED83231F106A58FE71FD21FDE8" blockId="3.[189,775,490,835]" box="[269,694,490,515]" pageId="3" pageNumber="141">
<heading id="CF7E818183231F106A58FE71FD21FDE8" box="[269,694,490,515]" centered="true" fontSize="10" level="2" pageId="3" pageNumber="141" reason="8">
<taxonomicName id="53894D6E83231F106A58FE71FD21FDE8" ID-CoL="8XWNF" authorityName="Gildenkov" authorityYear="2019" box="[269,694,490,515]" class="Insecta" family="Staphylinidae" genus="Carpelimus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="141" phylum="Arthropoda" rank="species" species="keraliensis" status="sp. n." subGenus="Trogophloeus">
<emphasis id="A6FDEAFF83231F106A58FE71FE1DFDE8" box="[269,394,490,514]" italics="true" pageId="3" pageNumber="141">Carpelimus</emphasis>
(
<emphasis id="A6FDEAFF83231F106AC9FE71FDA7FDE8" box="[412,560,490,514]" italics="true" pageId="3" pageNumber="141">Trogophloeus</emphasis>
)
<emphasis id="A6FDEAFF83231F106917FE71FD21FDE8" box="[578,694,490,514]" italics="true" pageId="3" pageNumber="141">keraliensis</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="943636ED83231F106AD2FD93FDACFDCA" blockId="3.[189,775,490,835]" box="[391,571,520,544]" pageId="3" pageNumber="141">
Gildenkov,
<emphasis id="A6FDEAFF83231F106953FD93FDACFDCA" bold="true" box="[518,571,520,544]" pageId="3" pageNumber="141">
<taxonomicNameLabel id="BDCE578483231F106953FD93FDACFDCA" box="[518,571,520,544]" pageId="3" pageNumber="141" rank="species">sp.n.</taxonomicNameLabel>
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="DC93656683231F106AC1FDBEFDB8FDD7" box="[404,559,549,573]" pageId="3" pageNumber="141" type="description">
<paragraph id="943636ED83231F106AC1FDBEFDB8FDD7" blockId="3.[189,775,490,835]" box="[404,559,549,573]" pageId="3" pageNumber="141">
<figureCitation id="0CB22A6883231F106AC1FDBEFE4EFDD7" box="[404,473,549,573]" captionStart="Figs 3–4" captionStartId="3.[226,263,1966,1985]" captionTargetBox="[189,1372,859,2064]" captionTargetId="figure-327@3.[189,1399,858,2068]" captionTargetPageId="3" captionText="Figs 3–4. Carpelimus (Trogophloeus) spp., holotypes, males, dorsal view: 3 — C. hoaensis, sp.n.; 4 — C. keraliensis, sp.n. Рис. 34. Carpelimus (Trogophloeus) spp., глотипы, самцы, сверху: 3 — C. hoaensis, sp.n.; 4 — C.keraliensis, sp.n." figureDoi="http://doi.org/10.5281/zenodo.13163821" httpUri="https://zenodo.org/record/13163821/files/figure.png" pageId="3" pageNumber="141">Figs 4</figureCitation>
,
<figureCitation id="0CB22A6883231F106AB0FDBEFDBCFDD7" box="[485,555,549,573]" captionStart="Figs 7–22" captionStartId="5.[226,262,1918,1937]" captionTargetBox="[202,1367,323,1859]" captionTargetId="figure-184@5.[198,1367,323,1863]" captionTargetPageId="5" captionText="Figs 7–22. Genitalia of Carpelimus (Trogophloeus) spp.: 7–8 — C. cachariensis sp.n.; 9–11— C. finelypunctus sp.n.; 12–14 — C. hoaensis sp.n.; 15–17 — C. keraliensis sp.n.; 18–19 — C. khasiensis sp.n.; 20–22 — C. madrasiensis sp.n.; 7, 9, 12, 15, 18, 20 — aedeagus, ventral view (holotypes); 8, 10, 13, 16, 19, 21 — aedeagus, lateral view (holotypes); 11, 14, 17, 22 — spermatheca (paratypes). Scale bar: 0.25 mm. Рис. 7–22. Гениталии Carpelimus (Trogophloeus) spp.: 7–8 — C. cachariensis sp.n.; 9–11— C. finelypunctus sp.n.; 12–14 — C. hoaensis sp.n.; 15–17 — C. keraliensis sp.n.; 18–19 — C. khasiensis sp.n.; 20–22 — C. madrasiensis sp.n.; 7, 9, 12, 15, 18, 20 — Эдеагус, сниЗу (голотипы); 8, 10, 13, 16, 19, 21 — Эдеагус, сбоку (голотипы); 11, 14, 17, 22 — сперматека (паратипы). МасШтаб: 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.13163825" httpUri="https://zenodo.org/record/13163825/files/figure.png" pageId="3" pageNumber="141">15–17</figureCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="DC93656683231F106BB7FDD1FD64FD57" pageId="3" pageNumber="141" type="materials_examined">
<paragraph id="943636ED83231F106BB7FDD1FD64FD57" blockId="3.[189,775,490,835]" pageId="3" pageNumber="141">
<materialsCitation id="24E13CB083231F106BB7FDD1FD78FD57" collectingDate="1993-12-24" collectionCode="NHMW" collectorName="Boukai" elevation="1000" latitude="9.516666" location="Cardamom Hills" longLatPrecision="1299" longitude="77.11667" pageId="3" pageNumber="141" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Kerala" typeStatus="holotype">
MATERIAL.
<typeStatus id="4B32884F83231F106A0FFDD1FE3CFDB7" box="[346,427,586,605]" pageId="3" pageNumber="141" type="holotype">Holotype</typeStatus>
,
<specimenCount id="828FFD6483231F106AE6FDDCFE50FDB7" box="[435,455,583,605]" count="1" pageId="3" pageNumber="141" type="male">♂</specimenCount>
“S-INDIEN:
<collectingRegion id="564DF80F83231F106968FDD1FDE2FDB7" box="[573,629,586,605]" country="India" name="Kerala" pageId="3" pageNumber="141">Kerala</collectingRegion>
<location id="9156603683231F10692FFDD1FC91FDB7" LSID="urn:lsid:plazi:treatment:1C2087FB83231F176A58FE71FAE0FE6D:9156603683231F10692FFDD1FC91FDB7" box="[634,774,586,605]" latitude="9.516666" longLatPrecision="1299" longitude="77.11667" name="Cardamom Hills" pageId="3" pageNumber="141" stateProvince="Kerala">Cardamom Hills</location>
<locationDeviation id="85842FCF83231F106BE8FDF9FEF9FD9F" box="[189,366,610,629]" location="Kumily" pageId="3" pageNumber="141">
<quantity id="53719B0883231F106BE8FDF9FF78FD9F" box="[189,239,610,629]" metricMagnitude="4" metricUnit="m" metricValue="1.0" pageId="3" pageNumber="141" unit="km" value="10.0">10km</quantity>
SW Kumily
</locationDeviation>
<geoCoordinate id="F1BD502A83231F106A2FFDF9FE51FD9F" box="[378,454,610,629]" degrees="77" direction="east" minutes="07" orientation="longitude" pageId="3" pageNumber="141" precision="925" value="77.11667">77°07´E</geoCoordinate>
<geoCoordinate id="F1BD502A83231F106A86FDF9FDB4FD9F" box="[467,547,610,629]" degrees="09" direction="north" minutes="31" orientation="latitude" pageId="3" pageNumber="141" precision="925" value="9.516666">09°31´N</geoCoordinate>
” “Vallakadavu.
<quantity id="53719B0883231F10699DFDF9FC91FD9F" box="[712,774,610,629]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="3" pageNumber="141" unit="m" value="1000.0">
<elevation id="1FA4D1DE83231F10699DFDF9FC91FD9F" box="[712,774,610,629]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="3" pageNumber="141" unit="m" value="1000.0">1000m</elevation>
</quantity>
<date id="E037102D83231F106BE8FDE1FE88FD67" box="[189,287,634,653]" pageId="3" pageNumber="141" value="1993-12-24">
<collectingDate id="F073E9C583231F106BE8FDE1FE88FD67" box="[189,287,634,653]" pageId="3" pageNumber="141" value="1993-12-24">24.12.1993</collectingDate>
</date>
, leg.
<collectorName id="397C533B83231F106A02FDE1FE02FD67" box="[343,405,634,653]" pageId="3" pageNumber="141">Boukai</collectorName>
&amp; Kejval” (
<collectionCode id="F298AE2883231F10695BFDE1FDCEFD67" box="[526,601,634,653]" country="Austria" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" pageId="3" pageNumber="141" type="Museum">NHMW</collectionCode>
).
<typeStatus id="4B32884F83231F106938FDE1FD2CFD67" box="[621,699,634,653]" pageId="3" pageNumber="141" type="paratype">Paratype</typeStatus>
:
<specimenCount id="828FFD6483231F10699DFDE1FD48FD67" box="[712,735,631,653]" count="1" pageId="3" pageNumber="141" type="female">1♀</specimenCount>
“S-INDIEN:
<collectingRegion id="564DF80F83231F106A58FD09FED2FD4F" box="[269,325,658,677]" country="India" name="Kerala" pageId="3" pageNumber="141">Kerala</collectingRegion>
Cardamom Hills
<locationDeviation id="85842FCF83231F106A81FD09FDFEFD4F" box="[468,617,658,677]" location="Kumily" pageId="3" pageNumber="141">
<quantity id="53719B0883231F106A81FD09FD93FD4F" box="[468,516,658,677]" metricMagnitude="4" metricUnit="m" metricValue="1.0" pageId="3" pageNumber="141" unit="km" value="10.0">10km</quantity>
SW Kumily
</locationDeviation>
<geoCoordinate id="F1BD502A83231F106939FD09FD25FD4F" box="[620,690,658,677]" degrees="77" direction="east" minutes="07" orientation="longitude" pageId="3" pageNumber="141" precision="925" value="77.11667">77°07´E</geoCoordinate>
<geoCoordinate id="F1BD502A83231F1069E0FD09FD69FD4F" box="[693,766,658,677]" degrees="09" direction="north" minutes="31" orientation="latitude" pageId="3" pageNumber="141" precision="925" value="9.516666">09°31´N</geoCoordinate>
” “Vallakadavu.
<quantity id="53719B0883231F106A68FD31FEE0FD57" box="[317,375,682,701]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="3" pageNumber="141" unit="m" value="1000.0">
<elevation id="1FA4D1DE83231F106A68FD31FEE0FD57" box="[317,375,682,701]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="3" pageNumber="141" unit="m" value="1000.0">1000m</elevation>
</quantity>
<date id="E037102D83231F106A2BFD31FE48FD57" box="[382,479,682,701]" pageId="3" pageNumber="141" value="1993-12-24">
<collectingDate id="F073E9C583231F106A2BFD31FE48FD57" box="[382,479,682,701]" pageId="3" pageNumber="141" value="1993-12-24">24.12.1993</collectingDate>
</date>
, leg.
<collectorName id="397C533B83231F10695AFD31FDDBFD57" box="[527,588,682,701]" pageId="3" pageNumber="141">Boukai</collectorName>
&amp; Kejval” (cMG)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="DC93656683231F176BB7FD59FB15FEBB" lastPageId="4" lastPageNumber="142" pageId="3" pageNumber="141" type="description">
<paragraph id="943636ED83231F106BB7FD59FD22FCA9" blockId="3.[189,775,490,835]" pageId="3" pageNumber="141">
DESCRIPTION (
<typeStatus id="4B32884F83231F106ADFFD59FE77FD32" box="[394,480,706,728]" pageId="3" pageNumber="141" type="holotype">holotype</typeStatus>
). Length 2.2 mm. Colouration brown. Head, pronotum and abdomen dark brown, elytra slightly lighter, dark brown with reddish tint; legs and antennae yellow brown, antennal segments 911 lighter than others. Integument slightly shining, body with short, light-coloured hairs.
</paragraph>
<paragraph id="943636ED83231F106807FEA0FAE0FD7A" blockId="3.[812,1399,315,844]" pageId="3" pageNumber="141">
Head transverse, with wide base, ratio of its length (from posterior margin of head to anterior margin of clypeus) to maximum width about 18:29. Neck constriction prominent. Eyes large, convex. Temples well-developed, round, eye diameter in dorsal view more than twice as long as temple length. Head widest across eyes (
<figureCitation id="0CB22A6883231F106F7AFE5AFBFAFE3C" box="[1071,1133,449,471]" captionStart="Figs 3–4" captionStartId="3.[226,263,1966,1985]" captionTargetBox="[189,1372,859,2064]" captionTargetId="figure-327@3.[189,1399,858,2068]" captionTargetPageId="3" captionText="Figs 3–4. Carpelimus (Trogophloeus) spp., holotypes, males, dorsal view: 3 — C. hoaensis, sp.n.; 4 — C. keraliensis, sp.n. Рис. 34. Carpelimus (Trogophloeus) spp., глотипы, самцы, сверху: 3 — C. hoaensis, sp.n.; 4 — C.keraliensis, sp.n." figureDoi="http://doi.org/10.5281/zenodo.13163821" httpUri="https://zenodo.org/record/13163821/files/figure.png" pageId="3" pageNumber="141">Fig. 4</figureCitation>
). Head surface with rather distinct, fine and dense punctation. Puncture diameter about twice as small as eye facet. Distances between punctures significantly smaller than their diameter, interspaces smooth, slightly shining. Antennae rather long, antennal segments 1–6 elongate; segments 78 about as wide as long; segments 9–10 slightly transverse; segment 11 elongate, conical. Last 3 segments more massive than others and form loose club (
<figureCitation id="0CB22A6883231F106E65FDE0FAFCFD7A" box="[1328,1387,635,657]" captionStart="Figs 3–4" captionStartId="3.[226,263,1966,1985]" captionTargetBox="[189,1372,859,2064]" captionTargetId="figure-327@3.[189,1399,858,2068]" captionTargetPageId="3" captionText="Figs 3–4. Carpelimus (Trogophloeus) spp., holotypes, males, dorsal view: 3 — C. hoaensis, sp.n.; 4 — C. keraliensis, sp.n. Рис. 34. Carpelimus (Trogophloeus) spp., глотипы, самцы, сверху: 3 — C. hoaensis, sp.n.; 4 — C.keraliensis, sp.n." figureDoi="http://doi.org/10.5281/zenodo.13163821" httpUri="https://zenodo.org/record/13163821/files/figure.png" pageId="3" pageNumber="141">Fig. 4</figureCitation>
).
</paragraph>
<paragraph id="943636ED83231F176807FD0DFE4FFE81" blockId="3.[812,1399,315,844]" lastBlockId="4.[189,776,315,871]" lastPageId="4" lastPageNumber="142" pageId="3" pageNumber="141">
Pronotum widest about 2/3 its length from base, then narrowed. Lateral margins broadly rounded (
<figureCitation id="0CB22A6883231F106F89FD2AFA80FD2C" box="[1244,1303,689,711]" captionStart="Figs 3–4" captionStartId="3.[226,263,1966,1985]" captionTargetBox="[189,1372,859,2064]" captionTargetId="figure-327@3.[189,1399,858,2068]" captionTargetPageId="3" captionText="Figs 3–4. Carpelimus (Trogophloeus) spp., holotypes, males, dorsal view: 3 — C. hoaensis, sp.n.; 4 — C. keraliensis, sp.n. Рис. 34. Carpelimus (Trogophloeus) spp., глотипы, самцы, сверху: 3 — C. hoaensis, sp.n.; 4 — C.keraliensis, sp.n." figureDoi="http://doi.org/10.5281/zenodo.13163821" httpUri="https://zenodo.org/record/13163821/files/figure.png" pageId="3" pageNumber="141">Fig. 4</figureCitation>
). Ratio of pronotum length to its maximum width about 25:34. Surface of pronotum with distinct, fine and dense punctation; punctation similar to that on head. Puncture diameter about twice as small as eye facet. Distances between punctures significantly smaller than their diameter, interspaces smooth, slightly shining. Depressions on pronotal disc absent, except for small flattening at its base (
<figureCitation id="0CB22A6883241F176ADAFECDFE5BFE81" box="[399,460,342,364]" captionStart="Figs 3–4" captionStartId="3.[226,263,1966,1985]" captionTargetBox="[189,1372,859,2064]" captionTargetId="figure-327@3.[189,1399,858,2068]" captionTargetPageId="3" captionText="Figs 3–4. Carpelimus (Trogophloeus) spp., holotypes, males, dorsal view: 3 — C. hoaensis, sp.n.; 4 — C. keraliensis, sp.n. Рис. 34. Carpelimus (Trogophloeus) spp., глотипы, самцы, сверху: 3 — C. hoaensis, sp.n.; 4 — C.keraliensis, sp.n." figureDoi="http://doi.org/10.5281/zenodo.13163821" httpUri="https://zenodo.org/record/13163821/files/figure.png" pageId="4" pageNumber="142">Fig. 4</figureCitation>
).
</paragraph>
<caption id="C0F6666583231F106BB7F835FDA8F7E6" ID-DOI="http://doi.org/10.5281/zenodo.13163821" ID-Zenodo-Dep="13163821" httpUri="https://zenodo.org/record/13163821/files/figure.png" pageId="3" pageNumber="141" startId="3.[226,263,1966,1985]" targetBox="[189,1372,859,2064]" targetPageId="3" targetType="figure">
<paragraph id="943636ED83231F106BB7F835FCF1F82B" blockId="3.[189,870,1966,2061]" box="[226,870,1966,1985]" pageId="3" pageNumber="141">
Figs 3–4.
<taxonomicName id="53894D6E83231F106A6CF835FDBEF82B" authorityName="Mannerheim" authorityYear="1830" box="[313,553,1966,1985]" class="Insecta" family="Staphylinidae" genus="Carpelimus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="141" phylum="Arthropoda" rank="subGenus" subGenus="Trogophloeus">
<emphasis id="A6FDEAFF83231F106A6CF835FE0AF82B" box="[313,413,1966,1985]" italics="true" pageId="3" pageNumber="141">Carpelimus</emphasis>
(
<emphasis id="A6FDEAFF83231F106AFFF835FDB6F82B" box="[426,545,1966,1985]" italics="true" pageId="3" pageNumber="141">Trogophloeus</emphasis>
)
</taxonomicName>
spp., holotypes, males, dorsal view:
</paragraph>
<paragraph id="943636ED83231F106BE8F85DFDE6F833" blockId="3.[189,870,1966,2061]" box="[189,625,1990,2009]" pageId="3" pageNumber="141">
3 —
<taxonomicName id="53894D6E83231F106BBFF85DFEC7F833" authorityName="Gildenkov" authorityYear="2019" box="[234,336,1990,2009]" class="Insecta" family="Staphylinidae" genus="Carpelimus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="141" phylum="Arthropoda" rank="species" species="hoaensis" status="sp.n.">
<emphasis id="A6FDEAFF83231F106BBFF85DFEC7F833" box="[234,336,1990,2009]" italics="true" pageId="3" pageNumber="141">C. hoaensis</emphasis>
</taxonomicName>
,
<emphasis id="A6FDEAFF83231F106A09F85DFE10F833" bold="true" box="[348,391,1990,2009]" pageId="3" pageNumber="141">
<taxonomicNameLabel id="BDCE578483231F106A09F85DFE10F833" box="[348,391,1990,2009]" pageId="3" pageNumber="141" rank="species">sp.n.</taxonomicNameLabel>
</emphasis>
; 4 —
<taxonomicName id="53894D6E83231F106A97F85DFDADF833" authorityName="Gildenkov" authorityYear="2019" box="[450,570,1990,2009]" class="Insecta" family="Staphylinidae" genus="Carpelimus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="141" phylum="Arthropoda" rank="species" species="keraliensis" status="sp.n.">
<emphasis id="A6FDEAFF83231F106A97F85DFDADF833" box="[450,570,1990,2009]" italics="true" pageId="3" pageNumber="141">C. keraliensis</emphasis>
</taxonomicName>
,
<emphasis id="A6FDEAFF83231F106913F85DFDE6F833" bold="true" box="[582,625,1990,2009]" pageId="3" pageNumber="141">
<taxonomicNameLabel id="BDCE578483231F106913F85DFDE6F833" box="[582,625,1990,2009]" pageId="3" pageNumber="141" rank="species">sp.n.</taxonomicNameLabel>
</emphasis>
</paragraph>
<paragraph id="943636ED83231F106BB7F879FCF2F81F" blockId="3.[189,870,1966,2061]" box="[226,869,2017,2037]" pageId="3" pageNumber="141">
Рис. 34.
<taxonomicName id="53894D6E83231F106A65F87AFDB7F81E" authorityName="Mannerheim" authorityYear="1830" box="[304,544,2017,2036]" class="Insecta" family="Staphylinidae" genus="Carpelimus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="141" phylum="Arthropoda" rank="subGenus" subGenus="Trogophloeus">
<emphasis id="A6FDEAFF83231F106A65F87AFE03F81E" box="[304,404,2017,2036]" italics="true" pageId="3" pageNumber="141">Carpelimus</emphasis>
(
<emphasis id="A6FDEAFF83231F106AF7F87AFD8EF81E" box="[418,537,2017,2036]" italics="true" pageId="3" pageNumber="141">Trogophloeus</emphasis>
)
</taxonomicName>
spp., глотипы, самцы, сверху: 3 —
</paragraph>
<paragraph id="943636ED83231F106BE8F861FDA8F7E6" blockId="3.[189,870,1966,2061]" box="[189,575,2041,2061]" pageId="3" pageNumber="141">
<taxonomicName id="53894D6E83231F106BE8F861FEB5F7E6" authorityName="Gildenkov" authorityYear="2019" box="[189,290,2041,2061]" class="Insecta" family="Staphylinidae" genus="Carpelimus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="141" phylum="Arthropoda" rank="species" species="hoaensis" status="sp.n.">
<emphasis id="A6FDEAFF83231F106BE8F861FEB5F7E6" box="[189,290,2041,2061]" italics="true" pageId="3" pageNumber="141">C. hoaensis</emphasis>
</taxonomicName>
,
<emphasis id="A6FDEAFF83231F106A7AF862FECDF7E6" bold="true" box="[303,346,2041,2060]" pageId="3" pageNumber="141">
<taxonomicNameLabel id="BDCE578483231F106A7AF862FECDF7E6" box="[303,346,2041,2060]" pageId="3" pageNumber="141" rank="species">sp.n.</taxonomicNameLabel>
</emphasis>
; 4 —
<taxonomicName id="53894D6E83231F106AC3F862FD9FF7E6" authorityName="Gildenkov" authorityYear="2019" box="[406,520,2041,2060]" class="Insecta" family="Staphylinidae" genus="Carpelimus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="141" phylum="Arthropoda" rank="species" species="keraliensis" status="sp.n.">
<emphasis id="A6FDEAFF83231F106AC3F862FD9FF7E6" box="[406,520,2041,2060]" italics="true" pageId="3" pageNumber="141">C.keraliensis</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="BDCE578483231F106941F862FDA8F7E6" box="[532,575,2041,2060]" pageId="3" pageNumber="141" rank="species">sp.n.</taxonomicNameLabel>
</paragraph>
</caption>
<paragraph id="943636ED83241F176BB7FEEAFDE6FDE6" blockId="4.[189,776,315,871]" pageId="4" pageNumber="142">
Ratio of length of elytra to their combined width about 35:40. Scutellum with shallow, round depressions (
<figureCitation id="0CB22A6883241F1769E9FE10FD6CFE4A" box="[700,763,395,417]" captionStart="Figs 3–4" captionStartId="3.[226,263,1966,1985]" captionTargetBox="[189,1372,859,2064]" captionTargetId="figure-327@3.[189,1399,858,2068]" captionTargetPageId="3" captionText="Figs 3–4. Carpelimus (Trogophloeus) spp., holotypes, males, dorsal view: 3 — C. hoaensis, sp.n.; 4 — C. keraliensis, sp.n. Рис. 34. Carpelimus (Trogophloeus) spp., глотипы, самцы, сверху: 3 — C. hoaensis, sp.n.; 4 — C.keraliensis, sp.n." figureDoi="http://doi.org/10.5281/zenodo.13163821" httpUri="https://zenodo.org/record/13163821/files/figure.png" pageId="4" pageNumber="142">Fig. 4</figureCitation>
). Surface of elytra with rather distinct, fine and dense punctation. Puncture diameter about equal to diameter of eye facet. Distances between punctures slightly smaller than their diameter, interspaces smooth, slightly shining.
</paragraph>
<paragraph id="943636ED83241F176BB7FD8AFDB5FDCD" blockId="4.[189,776,315,871]" box="[226,546,529,551]" pageId="4" pageNumber="142">Abdomen delicately shagreened.</paragraph>
<paragraph id="943636ED83241F176BB7FDB0FD5AFDAB" blockId="4.[189,776,315,871]" box="[226,717,555,577]" pageId="4" pageNumber="142">
Aedeagus of characteristic structure (
<figureCitation id="0CB22A6883241F17691AFDB0FD56FDAB" box="[591,705,555,577]" captionStart="Figs 7–22" captionStartId="5.[226,262,1918,1937]" captionTargetBox="[202,1367,323,1859]" captionTargetId="figure-184@5.[198,1367,323,1863]" captionTargetPageId="5" captionText="Figs 7–22. Genitalia of Carpelimus (Trogophloeus) spp.: 7–8 — C. cachariensis sp.n.; 9–11— C. finelypunctus sp.n.; 12–14 — C. hoaensis sp.n.; 15–17 — C. keraliensis sp.n.; 18–19 — C. khasiensis sp.n.; 20–22 — C. madrasiensis sp.n.; 7, 9, 12, 15, 18, 20 — aedeagus, ventral view (holotypes); 8, 10, 13, 16, 19, 21 — aedeagus, lateral view (holotypes); 11, 14, 17, 22 — spermatheca (paratypes). Scale bar: 0.25 mm. Рис. 7–22. Гениталии Carpelimus (Trogophloeus) spp.: 7–8 — C. cachariensis sp.n.; 9–11— C. finelypunctus sp.n.; 12–14 — C. hoaensis sp.n.; 15–17 — C. keraliensis sp.n.; 18–19 — C. khasiensis sp.n.; 20–22 — C. madrasiensis sp.n.; 7, 9, 12, 15, 18, 20 — Эдеагус, сниЗу (голотипы); 8, 10, 13, 16, 19, 21 — Эдеагус, сбоку (голотипы); 11, 14, 17, 22 — сперматека (паратипы). МасШтаб: 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.13163825" httpUri="https://zenodo.org/record/13163825/files/figure.png" pageId="4" pageNumber="142">Figs 15–16</figureCitation>
).
</paragraph>
<paragraph id="943636ED83241F176BB7FDDDFC90FD9C" blockId="4.[189,776,315,871]" pageId="4" pageNumber="142">
Female. Sexual dimorphism absent, female morphologically similar to male. Spermatheca of characteristic structure (
<figureCitation id="0CB22A6883241F1769ECFDFAFD6CFD9C" box="[697,763,609,631]" captionStart="Figs 7–22" captionStartId="5.[226,262,1918,1937]" captionTargetBox="[202,1367,323,1859]" captionTargetId="figure-184@5.[198,1367,323,1863]" captionTargetPageId="5" captionText="Figs 7–22. Genitalia of Carpelimus (Trogophloeus) spp.: 7–8 — C. cachariensis sp.n.; 9–11— C. finelypunctus sp.n.; 12–14 — C. hoaensis sp.n.; 15–17 — C. keraliensis sp.n.; 18–19 — C. khasiensis sp.n.; 20–22 — C. madrasiensis sp.n.; 7, 9, 12, 15, 18, 20 — aedeagus, ventral view (holotypes); 8, 10, 13, 16, 19, 21 — aedeagus, lateral view (holotypes); 11, 14, 17, 22 — spermatheca (paratypes). Scale bar: 0.25 mm. Рис. 7–22. Гениталии Carpelimus (Trogophloeus) spp.: 7–8 — C. cachariensis sp.n.; 9–11— C. finelypunctus sp.n.; 12–14 — C. hoaensis sp.n.; 15–17 — C. keraliensis sp.n.; 18–19 — C. khasiensis sp.n.; 20–22 — C. madrasiensis sp.n.; 7, 9, 12, 15, 18, 20 — Эдеагус, сниЗу (голотипы); 8, 10, 13, 16, 19, 21 — Эдеагус, сбоку (голотипы); 11, 14, 17, 22 — сперматека (паратипы). МасШтаб: 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.13163825" httpUri="https://zenodo.org/record/13163825/files/figure.png" pageId="4" pageNumber="142">Fig. 17</figureCitation>
).
</paragraph>
<paragraph id="943636ED83241F176BB7FDE0FB15FEBB" blockId="4.[189,776,315,871]" lastBlockId="4.[812,1399,315,391]" pageId="4" pageNumber="142">
COMPARATIVE REMARKS. The new species is most similar in size, the structure of the pronotum, colouration and punctation patterns to closely related species,
<taxonomicName id="53894D6E83241F17692CFD2AFC90FD2C" authorityName="Gildenkov" authorityYear="2014" box="[633,775,689,710]" class="Insecta" family="Staphylinidae" genus="Carpelimus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="142" phylum="Arthropoda" rank="species" species="magnicollis">
<emphasis id="A6FDEAFF83241F17692CFD2AFC90FD2C" box="[633,775,689,710]" italics="true" pageId="4" pageNumber="142">C. magnicollis</emphasis>
</taxonomicName>
and
<taxonomicName id="53894D6E83241F176BBDFD57FE16FD0A" authorityName="Gildenkov" authorityYear="2019" box="[232,385,715,737]" class="Insecta" family="Staphylinidae" genus="Carpelimus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="142" phylum="Arthropoda" rank="species" species="cachariensis" status="sp.n.">
<emphasis id="A6FDEAFF83241F176BBDFD57FF6FFD0B" box="[232,248,716,737]" italics="true" pageId="4" pageNumber="142">C</emphasis>
.
<emphasis id="A6FDEAFF83241F176A52FD50FE16FD0A" box="[263,385,715,736]" italics="true" pageId="4" pageNumber="142">cachariensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="BDCE578483241F176ADFFD50FE2CFD0B" box="[394,443,715,737]" pageId="4" pageNumber="142" rank="species">
<emphasis id="A6FDEAFF83241F176ADFFD50FE23FD0A" bold="true" box="[394,436,715,736]" pageId="4" pageNumber="142">sp.n</emphasis>
.
</taxonomicNameLabel>
The new species differs from
<taxonomicName id="53894D6E83241F1769A5FD57FEB9FD11" authorityName="Gildenkov" authorityYear="2014" class="Insecta" family="Staphylinidae" genus="Carpelimus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="142" phylum="Arthropoda" rank="species" species="magnicollis">
<emphasis id="A6FDEAFF83241F1769A5FD57FC97FD0B" box="[752,768,716,737]" italics="true" pageId="4" pageNumber="142">C</emphasis>
.
<emphasis id="A6FDEAFF83241F176BE8FD7DFEB9FD11" box="[189,302,742,763]" italics="true" pageId="4" pageNumber="142">magnicollis</emphasis>
</taxonomicName>
in having smaller size and is clearly distinguished by the structure of the aedeagus [
<bibRefCitation id="F0184B1C83241F176910FC9AFD7BFCFC" author="Gildenkov M. Yu." box="[581,748,768,791]" pageId="4" pageNumber="142" refId="ref5809" refString="Gildenkov M. Yu. 2015. [Fauna of Carpelimus of the Old World (Coleoptera: Staphylinidae)]. Smolensk: SmolSU. 414 pp. [In Russian, with English summary]" type="book" year="2015">Gildenkov, 2015</bibRefCitation>
, p. 385,
<figureCitation id="0CB22A6883241F176BA5FC80FEA8FCDA" box="[240,319,795,817]" captionStart="Figs 7–22" captionStartId="5.[226,262,1918,1937]" captionTargetBox="[202,1367,323,1859]" captionTargetId="figure-184@5.[198,1367,323,1863]" captionTargetPageId="5" captionText="Figs 7–22. Genitalia of Carpelimus (Trogophloeus) spp.: 7–8 — C. cachariensis sp.n.; 9–11— C. finelypunctus sp.n.; 12–14 — C. hoaensis sp.n.; 15–17 — C. keraliensis sp.n.; 18–19 — C. khasiensis sp.n.; 20–22 — C. madrasiensis sp.n.; 7, 9, 12, 15, 18, 20 — aedeagus, ventral view (holotypes); 8, 10, 13, 16, 19, 21 — aedeagus, lateral view (holotypes); 11, 14, 17, 22 — spermatheca (paratypes). Scale bar: 0.25 mm. Рис. 7–22. Гениталии Carpelimus (Trogophloeus) spp.: 7–8 — C. cachariensis sp.n.; 9–11— C. finelypunctus sp.n.; 12–14 — C. hoaensis sp.n.; 15–17 — C. keraliensis sp.n.; 18–19 — C. khasiensis sp.n.; 20–22 — C. madrasiensis sp.n.; 7, 9, 12, 15, 18, 20 — Эдеагус, сниЗу (голотипы); 8, 10, 13, 16, 19, 21 — Эдеагус, сбоку (голотипы); 11, 14, 17, 22 — сперматека (паратипы). МасШтаб: 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.13163825" httpUri="https://zenodo.org/record/13163825/files/figure.png" pageId="4" pageNumber="142">Figs 19</figureCitation>
: 45]. It differs from
<taxonomicName id="53894D6E83241F176943FC87FD27FCDA" authorityName="Gildenkov" authorityYear="2019" box="[534,688,795,817]" class="Insecta" family="Staphylinidae" genus="Carpelimus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="142" phylum="Arthropoda" rank="species" species="cachariensis" status="sp.n.">
<emphasis id="A6FDEAFF83241F176943FC87FDB1FCDB" box="[534,550,796,817]" italics="true" pageId="4" pageNumber="142">C</emphasis>
.
<emphasis id="A6FDEAFF83241F176963FC80FD27FCDA" box="[566,688,795,816]" italics="true" pageId="4" pageNumber="142">cachariensis</emphasis>
</taxonomicName>
<emphasis id="A6FDEAFF83241F1769ECFC80FD7DFCDA" bold="true" box="[697,746,795,816]" pageId="4" pageNumber="142">
<taxonomicNameLabel id="BDCE578483241F1769ECFC80FD7DFCDA" box="[697,746,795,816]" pageId="4" pageNumber="142" rank="species">sp.n.</taxonomicNameLabel>
</emphasis>
in having slightly more distinct punctation on the surface of the pronotum and is clearly differentiated from it by the structure of the aedeagus (
<figureCitation id="0CB22A6883241F176886FEA0FBE1FEBB" box="[979,1142,315,337]" captionStart="Figs 7–22" captionStartId="5.[226,262,1918,1937]" captionTargetBox="[202,1367,323,1859]" captionTargetId="figure-184@5.[198,1367,323,1863]" captionTargetPageId="5" captionText="Figs 7–22. Genitalia of Carpelimus (Trogophloeus) spp.: 7–8 — C. cachariensis sp.n.; 9–11— C. finelypunctus sp.n.; 12–14 — C. hoaensis sp.n.; 15–17 — C. keraliensis sp.n.; 18–19 — C. khasiensis sp.n.; 20–22 — C. madrasiensis sp.n.; 7, 9, 12, 15, 18, 20 — aedeagus, ventral view (holotypes); 8, 10, 13, 16, 19, 21 — aedeagus, lateral view (holotypes); 11, 14, 17, 22 — spermatheca (paratypes). Scale bar: 0.25 mm. Рис. 7–22. Гениталии Carpelimus (Trogophloeus) spp.: 7–8 — C. cachariensis sp.n.; 9–11— C. finelypunctus sp.n.; 12–14 — C. hoaensis sp.n.; 15–17 — C. keraliensis sp.n.; 18–19 — C. khasiensis sp.n.; 20–22 — C. madrasiensis sp.n.; 7, 9, 12, 15, 18, 20 — Эдеагус, сниЗу (голотипы); 8, 10, 13, 16, 19, 21 — Эдеагус, сбоку (голотипы); 11, 14, 17, 22 — сперматека (паратипы). МасШтаб: 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.13163825" httpUri="https://zenodo.org/record/13163825/files/figure.png" pageId="4" pageNumber="142">Figs 7–8, 15–16</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="DC93656683241F176807FECDFBD5FE86" box="[850,1090,342,364]" pageId="4" pageNumber="142" type="distribution">
<paragraph id="943636ED83241F176807FECDFBD5FE86" blockId="4.[812,1399,315,391]" box="[850,1090,342,364]" pageId="4" pageNumber="142">
DISTRIBUTION.
<collectingCountry id="EC9E767D83241F176F5CFECDFBA9FE86" box="[1033,1086,342,364]" name="India" pageId="4" pageNumber="142">India</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="DC93656683241F176807FEEAFAE0FE6D" box="[850,1399,369,391]" pageId="4" pageNumber="142" type="etymology">
<paragraph id="943636ED83241F176807FEEAFAE0FE6D" blockId="4.[812,1399,315,391]" box="[850,1399,369,391]" pageId="4" pageNumber="142">ETYMOLOGY. Named for its geographical distribution.</paragraph>
</subSubSection>
</treatment>
</document>
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