415 lines
56 KiB
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415 lines
56 KiB
XML
<document id="AA975771D36EF01108A9D7F3898C5BC5" ID-CLB-Dataset="299696" ID-DOI="10.5281/zenodo.11512952" ID-GBIF-Dataset="2b56e3b0-2274-4963-b243-9c991b07072c" ID-ISSN="0068-547X" ID-Zenodo-Dep="11512952" IM.bibliography_approvedBy="julia" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="julia" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="julia" IM.treatmentCitations_approvedBy="julia" IM.treatments_approvedBy="julia" checkinTime="1717721037103" checkinUser="felipe" docAuthor="Kavanaugh, David H. & Cueva-Dabkoski, Mollie" docDate="2023" docId="1C1187FCEB70ED5FFF9EFB4E675882CF" docLanguage="en" docName="ProcCalifAcadSci.67.21.493-537.pdf" docOrigin="Proceedings of the California Academy of Sciences 67 (21)" docStyle="DocumentStyle:E411DA009AF5E7F9A7E023431E2B7D5B.2:ProcCalifAcadSci.2015-.journal_article" docStyleId="E411DA009AF5E7F9A7E023431E2B7D5B" docStyleName="ProcCalifAcadSci.2015-.journal_article" docStyleVersion="2" docTitle="Cyclosomus sumatrensis Bouchard" docType="treatment" docVersion="5" lastPageNumber="519" masterDocId="E028FF84EB68ED45FFE5FFAF6327867B" masterDocTitle="A Review of Genus Cyclosomus Latreille (Coleoptera: Carabidae: Lebiinae: Cyclosomini) in Asia." masterLastPageNumber="537" masterPageNumber="493" pageNumber="517" updateTime="1720721490288" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:titleInfo id="FD6C7291D7DF33D8BE5190D95B7B2640">
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<mods:title id="C2288CC926DE218FE9D116976BA6BB2A">A Review of Genus Cyclosomus Latreille (Coleoptera: Carabidae: Lebiinae: Cyclosomini) in Asia.</mods:title>
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<mods:name id="2485DBF3C2B372D391C569E4006CBAF8" type="personal">
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<mods:namePart id="5F105567EE00B7E00D80AF62378DD2EA">Kavanaugh, David H.</mods:namePart>
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<mods:affiliation id="7A4AFAB418552E8C7B842B6018FB11B0">Department of Entomology, California Academy of Sciences, 55 Music Concourse Drive, Golden Gate Park, San Francisco, CA 94118.</mods:affiliation>
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<mods:name id="2724DF02D813291CCEBD6BFB123CE16F" type="personal">
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<mods:roleTerm id="568EE2BF1C45F5C0C2C2FE2BBFD133F6">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="BF980668DCA4410AA799FF60CDB6F37C">Cueva-Dabkoski, Mollie</mods:namePart>
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<mods:affiliation id="5C7E0F5BF3457011ADA9AE0B92089691">Student Science Fellow, California Academy of Sciences, San Francisco, and & 2355 Pearl Street, Santa Monica, CA 90405.</mods:affiliation>
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<mods:typeOfResource id="2352D16BF69E3CC6F6903C70EADCC326">text</mods:typeOfResource>
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<mods:title id="C5C3953D960DD7B71F55DBAC6866DB3F">Proceedings of the California Academy of Sciences</mods:title>
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<mods:part id="4CB320E372326CC96E461D1B8342EB3B">
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<mods:date id="DBCDD477B0F5BDF1ADB62B7723F6D661">2023</mods:date>
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<mods:number id="11074127C3186F8C33E93BAAED31E3DD">2023-02-15</mods:number>
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<mods:number id="0CBBEE9A7F6698D724A0E41E82646327">67</mods:number>
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<mods:number id="152BBFBDD0CB74C5F9B49127F3BFD274">21</mods:number>
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<mods:start id="86D08F258A59793827041B6C28E2150B">493</mods:start>
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<mods:identifier id="2AC5B9E1DB3521887E61E157C98EFDF5" type="DOI">10.5281/zenodo.11512952</mods:identifier>
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<treatment id="1C1187FCEB70ED5FFF9EFB4E675882CF" ID-DOI="http://doi.org/10.5281/zenodo.12726729" ID-GBIF-Taxon="235039991" ID-Zenodo-Dep="12726729" LSID="urn:lsid:plazi:treatment:1C1187FCEB70ED5FFF9EFB4E675882CF" httpUri="http://treatment.plazi.org/id/1C1187FCEB70ED5FFF9EFB4E675882CF" lastPageId="26" lastPageNumber="519" pageId="24" pageNumber="517">
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<subSubSection id="DCA26561EB70ED5DFF9EFB4E61258283" box="[123,514,1249,1273]" pageId="24" pageNumber="517" type="nomenclature">
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<paragraph id="940736EAEB70ED5DFF9EFB4E61258283" blockId="24.[123,1174,1249,1726]" box="[123,514,1249,1273]" pageId="24" pageNumber="517">
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<heading id="CF4F8186EB70ED5DFF9EFB4E61258283" box="[123,514,1249,1273]" fontSize="10" level="2" pageId="24" pageNumber="517" reason="4">
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<taxonomicName id="53B84D69EB70ED5DFF9EFB4E61258283" ID-CoL="6C69G" authority="Bouchard" authorityName="Bouchard" box="[123,514,1249,1273]" class="Insecta" family="Carabidae" genus="Cyclosomus" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="517" phylum="Arthropoda" rank="species" species="sumatrensis">
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<emphasis id="A6CCEAF8EB70ED5DFF9EFB4E61258283" bold="true" box="[123,514,1249,1273]" pageId="24" pageNumber="517">
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<emphasis id="A6CCEAF8EB70ED5DFF9EFB4E62AD8282" bold="true" box="[123,394,1249,1273]" italics="true" pageId="24" pageNumber="517">Cyclosomus sumatrensis</emphasis>
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Bouchard
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="DCA26561EB70ED5DFF9EFAAE62F7836C" box="[123,464,1281,1303]" pageId="24" pageNumber="517" type="description">
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<paragraph id="940736EAEB70ED5DFF9EFAAE62F7836C" blockId="24.[123,1174,1249,1726]" box="[123,464,1281,1303]" pageId="24" pageNumber="517">
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<figureCitation id="0C832A6FEB70ED5DFF9EFAAE63C5836C" box="[123,226,1281,1303]" captionStart="FIGURE" captionStartId="25.[171,183,1138,1157]" captionTargetBox="[120,1176,216,1118]" captionTargetId="figure-226@25.[120,1176,216,1118]" captionTargetPageId="25" captionText="FIGURE 11. Cyclosomus sumatrensis Bouchard, dorsal habitus. A. Dark male (Kras, Kediri, East Java, Java, Indonesia); B. Pale male (Taman Negara, Pahang, Malaysia). Scale lines = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.11512980" httpUri="https://zenodo.org/record/11512980/files/figure.png" pageId="24" pageNumber="517">Figures 11</figureCitation>
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,
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<figureCitation id="0C832A6FEB70ED5DFF08FAAE623F836C" box="[237,280,1281,1303]" captionStart="FIGURE" captionStartId="33.[171,183,1573,1592]" captionTargetBox="[139,1158,216,1560]" captionTargetId="figure-14@33.[135,1162,216,1560]" captionTargetPageId="33" captionText="FIGURE 14. Pronota, Cyclosomus spp., dorsal aspect. A. C. acutangulus Kavanaugh and Cueva-Dabkoski (Bhalukpong, Arunachal Pradesh, India); B. C. flexuosus (Fabricius) (Hetauda, Bagmati Province, Nepal); C. C. inustus Andrewes, holotype (Hong Kong, China); D. C. marginatus Motschulsky (India); E. C. pallidus Kavanaugh and Cueva- Dabkoski sp. nov. (Karachi, Sindh Province, Pakistan); F. C. philippinus Heller (Pili, Zambales Mountains, Luzon, Philippines); G. C. sumatrensis Bouchard (Taman Negra National Park, Pahang, Malaysia); H. C. suturalis (Wiedemann), lectotype (“Bengala”, India); I. C. vespertilio Cueva-Dabkoski and Kavanaugh sp. nov. Scale lines = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.11512988" httpUri="https://zenodo.org/record/11512988/files/figure.png" pageId="24" pageNumber="517">14G</figureCitation>
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,
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<figureCitation id="0C832A6FEB70ED5DFEC6FAAE626A836C" box="[291,333,1281,1303]" captionStart="FIGURE" captionStartId="34.[171,183,869,888]" captionTargetBox="[140,1176,216,846]" captionTargetId="figure-9@34.[120,1176,216,846]" captionTargetPageId="34" captionText="FIGURE 15. Prosternal intercoxal process, Cyclosomus spp., left lateral aspect. A. C. acutangulus Kavanaugh and Cueva-Dabkoski (Longchuan Jiang at Longjiang Bridge, Wuhe Township, Tengchong County, Yunnan, China); B. C. flexuosus Dejean (Cinkona, Anamalia Hills, Tamil Nadu, India); C. C. sumatrensis Bouchard (Taman Negara, Pahang, Malaysia); D. C. verspertilio Cueva-Dabkoski and Kavanaugh sp. nov. (New Delhi, Delhi, India). Scale lines = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.11512990" httpUri="https://zenodo.org/record/11512990/files/figure.png" pageId="24" pageNumber="517">15C</figureCitation>
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,
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<figureCitation id="0C832A6FEB70ED5DFEBDFAAE625A836C" box="[344,381,1281,1303]" captionStart="FIGURE" captionStartId="35.[171,183,1494,1513]" captionTargetBox="[120,1163,216,1464]" captionTargetId="figure-14@35.[120,1176,216,1479]" captionTargetPageId="35" captionText="FIGURE 16. Male genitalia, Cyclosomus spp., left lateral aspect. A. C. acutangulus Kavanaugh and Cueva-Dabkoski (Longchuan Jiang at Longjiang Bridge, Wuhe Township, Tengchong County, Yunnan, China); B. C. flexuosus Dejean (Karaikal, Puducherry, India); C. C. inustus Andrewes (“China”); D. C. marginatus Motschulsky (“Bengal”, India); E. C. philippinus Heller (Pili, Zambales Mountains, Luzon, Philippines); F. C. sumatrensis Bouchard (Taman Negara, Pahang, Malaysia); G. C. suturalis (Wiedemann), lectotype (“Bengala”, India); H. C. verspertilio Cueva-Dabkoski and Kavanaugh sp. nov. (New Delhi, Delhi, India). Scale lines = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.11512992" httpUri="https://zenodo.org/record/11512992/files/figure.png" pageId="24" pageNumber="517">16F</figureCitation>
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,
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<figureCitation id="0C832A6FEB70ED5DFE6DFAAE6289836C" box="[392,430,1281,1303]" captionStart="FIGURE" captionStartId="36.[171,183,1563,1582]" captionTargetBox="[120,1164,216,1526]" captionTargetId="figure-9@36.[120,1176,216,1548]" captionTargetPageId="36" captionText="FIGURE 17. Male genitalia, Cyclosomus spp., dorsal aspect. A. C. acutangulus Kavanaugh and Cueva-Dabkoski (Longchuan Jiang at Longjiang Bridge, Wuhe Township, Tengchong County, Yunnan, China); B. C. flexuosus Dejean (Karaikal, Puducherry, India); C. C. inustus Andrewes (“China”); D. C. marginatus Motschulsky (“Bengal”, India); E. C. philippinus Heller (Pili, Zambales Mountains, Luzon, Philippines); F. C. sumatrensis Bouchard (Taman Negara, Pahang, Malaysia); G. C. suturalis (Wiedemann), lectotype (“Bengala”, India); H. C. verspertilio Cueva-Dabkoski and Kavanaugh sp. nov. (New Delhi, Delhi, India). Scale lines = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.11512994" httpUri="https://zenodo.org/record/11512994/files/figure.png" pageId="24" pageNumber="517">17F</figureCitation>
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,
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<figureCitation id="0C832A6FEB70ED5DFE5DFAAE62F7836C" box="[440,464,1281,1303]" captionStart="FIGURE" captionStartId="40.[171,183,837,856]" captionTargetBox="[168,1128,216,826]" captionTargetId="figure-339@40.[168,1128,216,826]" captionTargetPageId="40" captionText="FIGURE 22. Map showing distribution of locality records for Cyclosomus specimens examined. White triangles = C. sumatrensis Bouchard; yellow highlighted area with black question mark = type area for C. suturalis (Wiedemann); white circles = other localities for C. suturalis; white squares = C. vespertilio Cueva-Dabkoski and Kavanaugh sp. nov. Modified from Wikimedia Commons, World Atlas of the World, at URL: <http://upload.wikimedia.org/wikipedia/ commons/8/8f/Whole_world_-_land_and_oceans_12000.jpg>. Scale line = 500 km." figureDoi="http://doi.org/10.5281/zenodo.11513004" httpUri="https://zenodo.org/record/11513004/files/figure.png" pageId="24" pageNumber="517">22</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="DCA26561EB70ED5CFF9EFA8A6255829C" lastPageId="25" lastPageNumber="518" pageId="24" pageNumber="517" type="reference_group">
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<paragraph id="940736EAEB70ED5DFF9EFA8A6233830E" blockId="24.[123,1174,1249,1726]" pageId="24" pageNumber="517">
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<treatmentCitationGroup id="B4A811C4EB70ED5DFF9EFA8A6233830E" pageId="24" pageNumber="517">
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<taxonomicName id="53B84D69EB70ED5DFF9EFA8A61638340" ID-CoL="6C69G" authority="Bouchard, 1903 a: 174" authorityName="Bouchard" authorityPageNumber="174" authorityYear="1903" box="[123,580,1317,1339]" class="Insecta" family="Carabidae" genus="Cyclosomus" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="517" phylum="Arthropoda" rank="species" species="sumatrensis">
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<emphasis id="A6CCEAF8EB70ED5DFF9EFA8A624E8340" box="[123,361,1317,1339]" italics="true" pageId="24" pageNumber="517">Cyclosomus sumatrensis</emphasis>
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<bibRefCitation id="F0294B1BEB70ED5DFE94FA8A61288340" author="BOUCHARD, J." box="[369,527,1317,1339]" pageId="24" pageNumber="517" pagination="169 - 176" refId="ref23110" refString="BOUCHARD, J. 1903. Insectes recueillis par M. le Professeur Dr Forster a Borneo, Java et Sumatra (Palembang). Annales de la Societe Entomologique de France 72: 169 - 176." type="journal article" year="1903">Bouchard, 1903</bibRefCitation>
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a:174
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</taxonomicName>
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.
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<typeStatus id="4B038848EB70ED5DFDB5FA8A61F68340" box="[592,721,1317,1339]" pageId="24" pageNumber="517" type="holotype">HOLOTYPE</typeStatus>
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, sex and present location unknown (see notes below). Type locality:
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<collectingCountry id="ECAF767AEB70ED5DFE64FAED62C68323" box="[385,481,1346,1368]" name="Indonesia" pageId="24" pageNumber="517">Indonesia</collectingCountry>
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,
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<collectingRegion id="567CF808EB70ED5DFE0FFAED611B8323" box="[490,572,1346,1368]" country="Indonesia" name="Sumatera Utara" pageId="24" pageNumber="517">Sumatra</collectingRegion>
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, Palembang.
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<treatmentCitation id="151910FBEB70ED5DFD5FFAED60A98323" author="ANDREWES, H. E." box="[698,910,1346,1368]" page="364" pageId="24" pageNumber="517" year="1930">
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<bibRefCitation id="F0294B1BEB70ED5DFD5FFAED60A98323" author="ANDREWES, H. E." box="[698,910,1346,1368]" pageId="24" pageNumber="517" pagination="363 - 365" refId="ref23077" refString="ANDREWES, H. E. 1930. Papers on the Oriental Carabidae. XXIII. Annals and Magazine of Natural History (Ser. 10) 6: 363 - 365." type="journal article" year="1930">Andrewes (1930:364)</bibRefCitation>
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</treatmentCitation>
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;
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<treatmentCitation id="151910FBEB70ED5DFC72FAED676F8323" author="CSIKI, E." box="[919,1096,1346,1368]" page="1295" pageId="24" pageNumber="517" year="1932">
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<bibRefCitation id="F0294B1BEB70ED5DFC72FAED676F8323" author="CSIKI, E." box="[919,1096,1346,1368]" pageId="24" pageNumber="517" pagination="1279 - 1598" refId="ref23258" refString="CSIKI, E. 1932. Coleopterorum catalogus. Pars 124. Carabidae: Harpalinae VII. W. Junk, Berlin, pp. 1279 - 1598." type="book chapter" year="1932">Csiki (1932:1295)</bibRefCitation>
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</treatmentCitation>
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;
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<treatmentCitation id="151910FBEB70ED5DFBB7FAEC622D830E" author="LORENZ, W." page="452" pageId="24" pageNumber="517" year="2005">
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<bibRefCitation id="F0294B1BEB70ED5DFBB7FAEC622D830E" author="LORENZ, W." pageId="24" pageNumber="517" refId="ref23840" refString="LORENZ, W. 2005. Systematic List of Extant Ground Beetles of the World (Insecta Coleoptera " Geadephaga ": Trachypachidae and Carabidae incl. Paussinae, Cicindelinae, Rhysodinae). Second edition. W. Lorenz, Tutzing, 530 pp." type="book" year="2005">Lorenz 2005:452</bibRefCitation>
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</treatmentCitation>
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).
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</treatmentCitationGroup>
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</paragraph>
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<paragraph id="940736EAEB70ED5CFF4EFA2B6255829C" blockId="24.[123,1174,1249,1726]" lastBlockId="25.[123,1174,1231,1703]" lastPageId="25" lastPageNumber="518" pageId="24" pageNumber="517">
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<emphasis id="A6CCEAF8EB70ED5DFF4EFA2B610B83E0" bold="true" box="[171,556,1412,1435]" pageId="24" pageNumber="517">Notes on types and nomenclature.</emphasis>
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<bibRefCitation id="F0294B1BEB70ED5DFDD5FA2C603D83E0" author="ANDREWES, H. E." box="[560,794,1411,1435]" pageId="24" pageNumber="517" pagination="363 - 365" refId="ref23077" refString="ANDREWES, H. E. 1930. Papers on the Oriental Carabidae. XXIII. Annals and Magazine of Natural History (Ser. 10) 6: 363 - 365." type="journal article" year="1930">Andrewes (1930:364)</bibRefCitation>
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reported on his study of the unique type of
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<taxonomicName id="53B84D69EB70ED5DFF2BFA0B624A83C7" ID-CoL="6C69G" authority="Bouchard, 1903 a: 174" authorityName="Bouchard" authorityPageNumber="174" authorityYear="1903" box="[206,365,1444,1468]" class="Insecta" family="Carabidae" genus="Cyclosomus" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="517" phylum="Arthropoda" rank="species" species="sumatrensis">
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<emphasis id="A6CCEAF8EB70ED5DFF2BFA0B624A83C7" box="[206,365,1444,1468]" italics="true" pageId="24" pageNumber="517">C. sumatrensis</emphasis>
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</taxonomicName>
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in the Förster collection, which was housed at that time in the Museum of the Lycée at Mulhouse, Strasbourg (
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<collectingCountry id="ECAF767AEB70ED5DFDE7FA6B617683A7" box="[514,593,1476,1500]" name="France" pageId="24" pageNumber="517">France</collectingCountry>
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). Our efforts to borrow and study the Bouchard
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<typeStatus id="4B038848EB70ED5DFBB8FA6B638E8386" pageId="24" pageNumber="517" type="holotype">holotype</typeStatus>
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were facilitated by Thierry Deuve, who contacted colleagues in several institutions in Strasbourg on our behalf. According to Mary Meister (Zoological Museum of Strasbourg (ZMSC)), the insects from the Lycée at Mulhouse collection were at one time offered to the ZMSC but a transfer was never completed. Efforts to learn whether or not the Mulhouse collection still exists have failed thus far. It seems likely that the collection, including the
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<typeStatus id="4B038848EB70ED5DFCF7F9CA60488006" box="[786,879,1637,1661]" pageId="24" pageNumber="517" type="holotype">holotype</typeStatus>
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of
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<taxonomicName id="53B84D69EB70ED5DFC74F9C9670A8006" ID-CoL="6C69G" authority="Bouchard, 1903 a: 174" authorityName="Bouchard" authorityPageNumber="174" authorityYear="1903" box="[913,1069,1638,1661]" class="Insecta" family="Carabidae" genus="Cyclosomus" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="517" phylum="Arthropoda" rank="species" species="sumatrensis">
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<emphasis id="A6CCEAF8EB70ED5DFC74F9C9670A8006" box="[913,1069,1638,1661]" italics="true" pageId="24" pageNumber="517">C. sumatrensis</emphasis>
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</taxonomicName>
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, has been lost or destroyed. However, this species is geographically and morphologically distinct enough from other
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<taxonomicName id="53B84D69EB70ED5DFF14F90A62E280C6" ID-CoL="6C69G" box="[241,453,1701,1725]" class="Insecta" family="Carabidae" genus="Cyclosomus" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="517" phylum="Arthropoda" rank="genus">
|
|
<emphasis id="A6CCEAF8EB70ED5DFF14F90A625580C6" box="[241,370,1701,1725]" italics="true" pageId="24" pageNumber="517">Cyclosomus</emphasis>
|
|
species
|
|
</taxonomicName>
|
|
that we see no need to designate a
|
|
<typeStatus id="4B038848EB70ED5DFCD1F90960AF80C5" box="[820,904,1702,1726]" pageId="24" pageNumber="517" type="neotype">neotype</typeStatus>
|
|
until the loss of the
|
|
<typeStatus id="4B038848EB70ED5CFBB8F90A638E8293" lastPageId="25" lastPageNumber="518" pageId="24" pageNumber="517" type="holotype">holotype</typeStatus>
|
|
can be confirmed.
|
|
</paragraph>
|
|
</subSubSection>
|
|
<caption id="C0C76662EB71ED5CFF4EFBDD602882E4" ID-DOI="http://doi.org/10.5281/zenodo.11512980" ID-Zenodo-Dep="11512980" httpUri="https://zenodo.org/record/11512980/files/figure.png" pageId="25" pageNumber="518" startId="25.[171,183,1138,1157]" targetBox="[120,1176,216,1118]" targetPageId="25" targetType="figure">
|
|
<paragraph id="940736EAEB71ED5CFF4EFBDD602882E4" blockId="25.[123,1173,1138,1183]" pageId="25" pageNumber="518">
|
|
<smallCapsWord id="92E1A036EB71ED5CFF4EFBDD63CF82FF" baselines="1152,1152" box="[171,232,1138,1157]" lowerCaseFontSize="6" mainFontSize="8" normCase="title" normString="Figure" pageId="25" pageNumber="518">FIGURE</smallCapsWord>
|
|
11.
|
|
<taxonomicName id="53B84D69EB71ED5CFEEBFBDD611A82FE" authority="Bouchard" authorityName="Bouchard" box="[270,573,1138,1157]" class="Insecta" family="Carabidae" genus="Cyclosomus" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="518" phylum="Arthropoda" rank="species" species="sumatrensis">
|
|
<emphasis id="A6CCEAF8EB71ED5CFEEBFBDD62C682FE" box="[270,481,1138,1157]" italics="true" pageId="25" pageNumber="518">Cyclosomus sumatrensis</emphasis>
|
|
Bouchard
|
|
</taxonomicName>
|
|
, dorsal habitus. A. Dark male (Kras, Kediri, East Java, Java, Indonesia); B. Pale male (Taman Negara, Pahang, Malaysia). Scale lines = 1.0 mm.
|
|
</paragraph>
|
|
</caption>
|
|
<subSubSection id="DCA26561EB71ED5FFF4EFB5F675882CF" lastPageId="26" lastPageNumber="519" pageId="25" pageNumber="518" type="diagnosis">
|
|
<paragraph id="940736EAEB71ED5CFF4EFB5F60B280DC" blockId="25.[123,1174,1231,1703]" pageId="25" pageNumber="518">
|
|
<emphasis id="A6CCEAF8EB71ED5CFF4EFB5F6238837C" bold="true" box="[171,287,1264,1287]" pageId="25" pageNumber="518">Diagnosis.</emphasis>
|
|
Adults of
|
|
<taxonomicName id="53B84D69EB71ED5CFE7FFB5F611C837C" box="[410,571,1264,1287]" class="Insecta" family="Carabidae" genus="Cyclosomus" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="518" phylum="Arthropoda" rank="species" species="sumatrensis">
|
|
<emphasis id="A6CCEAF8EB71ED5CFE7FFB5F611C837C" box="[410,571,1264,1287]" italics="true" pageId="25" pageNumber="518">C. sumatrensis</emphasis>
|
|
</taxonomicName>
|
|
can be distinguished from those of other
|
|
<taxonomicName id="53B84D69EB71ED5CFBF1FB4063EF835C" pageId="25" pageNumber="518">
|
|
<emphasis id="A6CCEAF8EB71ED5CFBF1FB4067B2837C" box="[1044,1173,1263,1287]" italics="true" pageId="25" pageNumber="518">Cyclosomus</emphasis>
|
|
species
|
|
</taxonomicName>
|
|
in Asia by the following combination of character states: Body size medium for genus, BL males =
|
|
<quantity id="53409B0FEB71ED5CFF3EFA9F625B833C" box="[219,380,1327,1351]" metricMagnitude="-3" metricUnit="m" metricValue="7.9" metricValueMax="8.1" metricValueMin="7.7" pageId="25" pageNumber="518" unit="mm" value="7.9" valueMax="8.1" valueMin="7.7">7.7 to 8.1 mm</quantity>
|
|
, females =
|
|
<quantity id="53409B0FEB71ED5CFE1AFA806187833C" box="[511,672,1327,1351]" metricMagnitude="-3" metricUnit="m" metricValue="7.25" metricValueMax="7.4" metricValueMin="7.1" pageId="25" pageNumber="518" unit="mm" value="7.25" valueMax="7.4" valueMin="7.1">7.1 to 7.4 mm</quantity>
|
|
; body form (
|
|
<figureCitation id="0C832A6FEB71ED5CFCD7FA8060A6833C" box="[818,897,1327,1351]" captionStart="FIGURE" captionStartId="25.[171,183,1138,1157]" captionTargetBox="[120,1176,216,1118]" captionTargetId="figure-226@25.[120,1176,216,1118]" captionTargetPageId="25" captionText="FIGURE 11. Cyclosomus sumatrensis Bouchard, dorsal habitus. A. Dark male (Kras, Kediri, East Java, Java, Indonesia); B. Pale male (Taman Negara, Pahang, Malaysia). Scale lines = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.11512980" httpUri="https://zenodo.org/record/11512980/files/figure.png" pageId="25" pageNumber="518">Fig. 11</figureCitation>
|
|
) slightly elongate ovoid; pronotum (
|
|
<figureCitation id="0C832A6FEB71ED5CFF17FAE06273831C" box="[242,340,1359,1383]" captionStart="FIGURE" captionStartId="33.[171,183,1573,1592]" captionTargetBox="[139,1158,216,1560]" captionTargetId="figure-14@33.[135,1162,216,1560]" captionTargetPageId="33" captionText="FIGURE 14. Pronota, Cyclosomus spp., dorsal aspect. A. C. acutangulus Kavanaugh and Cueva-Dabkoski (Bhalukpong, Arunachal Pradesh, India); B. C. flexuosus (Fabricius) (Hetauda, Bagmati Province, Nepal); C. C. inustus Andrewes, holotype (Hong Kong, China); D. C. marginatus Motschulsky (India); E. C. pallidus Kavanaugh and Cueva- Dabkoski sp. nov. (Karachi, Sindh Province, Pakistan); F. C. philippinus Heller (Pili, Zambales Mountains, Luzon, Philippines); G. C. sumatrensis Bouchard (Taman Negra National Park, Pahang, Malaysia); H. C. suturalis (Wiedemann), lectotype (“Bengala”, India); I. C. vespertilio Cueva-Dabkoski and Kavanaugh sp. nov. Scale lines = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.11512988" httpUri="https://zenodo.org/record/11512988/files/figure.png" pageId="25" pageNumber="518">Fig. 14G</figureCitation>
|
|
) relatively narrow (ratio PWM/PL = 2.25 to 2.43), disc piceous to black with lateral pale bands slightly narrower than average for genus, distinctly defined, and slightly narrowed basally, apical angles narrower and less broadly rounded, lateral margins smoothly arcuate, not sinuate near anterior angles; elytra (
|
|
<figureCitation id="0C832A6FEB71ED5CFDD6FA0061A583BC" box="[563,642,1455,1479]" captionStart="FIGURE" captionStartId="25.[171,183,1138,1157]" captionTargetBox="[120,1176,216,1118]" captionTargetId="figure-226@25.[120,1176,216,1118]" captionTargetPageId="25" captionText="FIGURE 11. Cyclosomus sumatrensis Bouchard, dorsal habitus. A. Dark male (Kras, Kediri, East Java, Java, Indonesia); B. Pale male (Taman Negara, Pahang, Malaysia). Scale lines = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.11512980" httpUri="https://zenodo.org/record/11512980/files/figure.png" pageId="25" pageNumber="518">Fig. 11</figureCitation>
|
|
) with pattern of dark maculae more extensively developed and less varied than average for genus, with middle transverse dark band thick and extended from midline at least across interval 8, also onto interval
|
|
<quantity id="53409B0FEB71ED5CFCA2FA406057807C" box="[839,880,1519,1543]" metricMagnitude="-1" metricUnit="m" metricValue="2.286" pageId="25" pageNumber="518" unit="in" value="9.0">9 in</quantity>
|
|
some specimens, preapical dark spot present; elytral striae deeply impressed and intervals slightly to moderately convex; elytral epipleura with long setae only in humeral and subhumeral areas, setae in apical two-thirds distinctly shorter; male median lobe with apical lamella extended in line with ventral curvature of shaft in lateral aspect (
|
|
<figureCitation id="0C832A6FEB71ED5CFE93F9C062F080FC" box="[374,471,1647,1671]" captionStart="FIGURE" captionStartId="35.[171,183,1494,1513]" captionTargetBox="[120,1163,216,1464]" captionTargetId="figure-14@35.[120,1176,216,1479]" captionTargetPageId="35" captionText="FIGURE 16. Male genitalia, Cyclosomus spp., left lateral aspect. A. C. acutangulus Kavanaugh and Cueva-Dabkoski (Longchuan Jiang at Longjiang Bridge, Wuhe Township, Tengchong County, Yunnan, China); B. C. flexuosus Dejean (Karaikal, Puducherry, India); C. C. inustus Andrewes (“China”); D. C. marginatus Motschulsky (“Bengal”, India); E. C. philippinus Heller (Pili, Zambales Mountains, Luzon, Philippines); F. C. sumatrensis Bouchard (Taman Negara, Pahang, Malaysia); G. C. suturalis (Wiedemann), lectotype (“Bengala”, India); H. C. verspertilio Cueva-Dabkoski and Kavanaugh sp. nov. (New Delhi, Delhi, India). Scale lines = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.11512992" httpUri="https://zenodo.org/record/11512992/files/figure.png" pageId="25" pageNumber="518">Fig. 16F</figureCitation>
|
|
), apical lamella rounded and of medium length in dorsal aspect (
|
|
<figureCitation id="0C832A6FEB71ED5CFF66F92063C580DC" box="[131,226,1679,1703]" captionStart="FIGURE" captionStartId="36.[171,183,1563,1582]" captionTargetBox="[120,1164,216,1526]" captionTargetId="figure-9@36.[120,1176,216,1548]" captionTargetPageId="36" captionText="FIGURE 17. Male genitalia, Cyclosomus spp., dorsal aspect. A. C. acutangulus Kavanaugh and Cueva-Dabkoski (Longchuan Jiang at Longjiang Bridge, Wuhe Township, Tengchong County, Yunnan, China); B. C. flexuosus Dejean (Karaikal, Puducherry, India); C. C. inustus Andrewes (“China”); D. C. marginatus Motschulsky (“Bengal”, India); E. C. philippinus Heller (Pili, Zambales Mountains, Luzon, Philippines); F. C. sumatrensis Bouchard (Taman Negara, Pahang, Malaysia); G. C. suturalis (Wiedemann), lectotype (“Bengala”, India); H. C. verspertilio Cueva-Dabkoski and Kavanaugh sp. nov. (New Delhi, Delhi, India). Scale lines = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.11512994" httpUri="https://zenodo.org/record/11512994/files/figure.png" pageId="25" pageNumber="518">Fig. 17F</figureCitation>
|
|
); specimen from West
|
|
<collectingCountry id="ECAF767AEB71ED5CFE3CF920611C80DC" box="[473,571,1679,1703]" name="Malaysia" pageId="25" pageNumber="518">Malaysia</collectingCountry>
|
|
or northern
|
|
<collectingCountry id="ECAF767AEB71ED5CFD24F920600E80DC" box="[705,809,1679,1703]" name="Indonesia" pageId="25" pageNumber="518">Indonesia</collectingCountry>
|
|
(
|
|
<figureCitation id="0C832A6FEB71ED5CFCDDF92060AE80DC" box="[824,905,1679,1703]" captionStart="FIGURE" captionStartId="40.[171,183,837,856]" captionTargetBox="[168,1128,216,826]" captionTargetId="figure-339@40.[168,1128,216,826]" captionTargetPageId="40" captionText="FIGURE 22. Map showing distribution of locality records for Cyclosomus specimens examined. White triangles = C. sumatrensis Bouchard; yellow highlighted area with black question mark = type area for C. suturalis (Wiedemann); white circles = other localities for C. suturalis; white squares = C. vespertilio Cueva-Dabkoski and Kavanaugh sp. nov. Modified from Wikimedia Commons, World Atlas of the World, at URL: <http://upload.wikimedia.org/wikipedia/ commons/8/8f/Whole_world_-_land_and_oceans_12000.jpg>. Scale line = 500 km." figureDoi="http://doi.org/10.5281/zenodo.11513004" httpUri="https://zenodo.org/record/11513004/files/figure.png" pageId="25" pageNumber="518">Fig. 22</figureCitation>
|
|
).
|
|
</paragraph>
|
|
<paragraph id="940736EAEB72ED5FFF4EFF7461C58448" blockId="26.[123,1174,219,1205]" pageId="26" pageNumber="519">
|
|
<bibRefCitation id="F0294B1BEB72ED5FFF4EFF7462418688" author="ANDREWES, H. E." box="[171,358,219,243]" pageId="26" pageNumber="519" pagination="363 - 365" refId="ref23077" refString="ANDREWES, H. E. 1930. Papers on the Oriental Carabidae. XXIII. Annals and Magazine of Natural History (Ser. 10) 6: 363 - 365." type="journal article" year="1930">Andrewes (1930)</bibRefCitation>
|
|
distinguished members of
|
|
<taxonomicName id="53B84D69EB72ED5FFD6CFF7460008688" box="[649,807,219,243]" class="Insecta" family="Carabidae" genus="Cyclosomus" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="519" phylum="Arthropoda" rank="species" species="sumatrensis">
|
|
<emphasis id="A6CCEAF8EB72ED5FFD6CFF7460008688" box="[649,807,219,243]" italics="true" pageId="26" pageNumber="519">C. sumatrensis</emphasis>
|
|
</taxonomicName>
|
|
from those of
|
|
<taxonomicName id="53B84D69EB72ED5FFC2DFF74676C8688" authorityName=", Kavanaugh" authorityYear="2015" box="[968,1099,219,243]" class="Insecta" family="Carabidae" genus="Cyclosomus" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="519" phylum="Arthropoda" rank="species" species="flexuosus">
|
|
<emphasis id="A6CCEAF8EB72ED5FFC2DFF74676C8688" box="[968,1099,219,243]" italics="true" pageId="26" pageNumber="519">C. flexuosus</emphasis>
|
|
</taxonomicName>
|
|
on the basis of their smaller size, narrower form, and deeper elytral striae, as well as minor differences in elytral color pattern. He concluded that
|
|
<taxonomicName id="53B84D69EB72ED5FFDF8FEB3619E8748" box="[541,697,284,307]" class="Insecta" family="Carabidae" genus="Cyclosomus" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="519" phylum="Arthropoda" rank="species" species="sumatrensis">
|
|
<emphasis id="A6CCEAF8EB72ED5FFDF8FEB3619E8748" box="[541,697,284,307]" italics="true" pageId="26" pageNumber="519">C. sumatrensis</emphasis>
|
|
</taxonomicName>
|
|
was probably a distinct species closely related to
|
|
<taxonomicName id="53B84D69EB72ED5FFF5CFE93621A8728" authorityName=", Kavanaugh" authorityYear="2015" box="[185,317,315,339]" class="Insecta" family="Carabidae" genus="Cyclosomus" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="519" phylum="Arthropoda" rank="species" species="flexuosus">
|
|
<emphasis id="A6CCEAF8EB72ED5FFF5CFE93621A8728" box="[185,317,315,339]" italics="true" pageId="26" pageNumber="519">C. flexuosus</emphasis>
|
|
</taxonomicName>
|
|
, but suggested that additional material might show it to be only a “variety” of the latter. Specimens of
|
|
<taxonomicName id="53B84D69EB72ED5FFE98FEF3613D8708" box="[381,538,348,371]" class="Insecta" family="Carabidae" genus="Cyclosomus" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="519" phylum="Arthropoda" rank="species" species="sumatrensis">
|
|
<emphasis id="A6CCEAF8EB72ED5FFE98FEF3613D8708" box="[381,538,348,371]" italics="true" pageId="26" pageNumber="519">C. sumatrensis</emphasis>
|
|
</taxonomicName>
|
|
can be confused only with those of
|
|
<taxonomicName id="53B84D69EB72ED5FFC7BFEF367078708" authorityName=", Kavanaugh" authorityYear="2015" box="[926,1056,347,371]" class="Insecta" family="Carabidae" genus="Cyclosomus" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="519" phylum="Arthropoda" rank="species" species="flexuosus">
|
|
<emphasis id="A6CCEAF8EB72ED5FFC7BFEF367078708" box="[926,1056,347,371]" italics="true" pageId="26" pageNumber="519">C. flexuosus</emphasis>
|
|
</taxonomicName>
|
|
, but members of these species are easily distinguished by the setae of the epipleura, which are long only in the humeral and subhumeral regions and progressively and distinctly shorter toward the apex in
|
|
<taxonomicName id="53B84D69EB72ED5FFF9EFE13623D87A8" box="[123,282,444,467]" class="Insecta" family="Carabidae" genus="Cyclosomus" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="519" phylum="Arthropoda" rank="species" species="sumatrensis">
|
|
<emphasis id="A6CCEAF8EB72ED5FFF9EFE13623D87A8" box="[123,282,444,467]" italics="true" pageId="26" pageNumber="519">C. sumatrensis</emphasis>
|
|
</taxonomicName>
|
|
specimens but longer throughout the epipleural length in
|
|
<taxonomicName id="53B84D69EB72ED5FFC70FE13670287A8" baseAuthorityName="Fabricius" baseAuthorityYear="1775" box="[917,1061,443,467]" class="Insecta" family="Carabidae" genus="Cyclosomus" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="519" phylum="Arthropoda" rank="species" species="flexusosus">
|
|
<emphasis id="A6CCEAF8EB72ED5FFC70FE13670287A8" box="[917,1061,443,467]" italics="true" pageId="26" pageNumber="519">C. flexusosus</emphasis>
|
|
</taxonomicName>
|
|
members. Specimens of
|
|
<taxonomicName id="53B84D69EB72ED5FFEF4FE7362898788" box="[273,430,476,499]" class="Insecta" family="Carabidae" genus="Cyclosomus" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="519" phylum="Arthropoda" rank="species" species="sumatrensis">
|
|
<emphasis id="A6CCEAF8EB72ED5FFEF4FE7362898788" box="[273,430,476,499]" italics="true" pageId="26" pageNumber="519">C. sumatrensis</emphasis>
|
|
</taxonomicName>
|
|
are also slightly smaller (
|
|
<quantity id="53409B0FEB72ED5FFD25FE7460728788" box="[704,853,475,499]" metricMagnitude="-3" metricUnit="m" metricValue="7.6" metricValueMax="8.1" metricValueMin="7.1" pageId="26" pageNumber="519" unit="mm" value="7.6" valueMax="8.1" valueMin="7.1">7.1 to 8.1 mm</quantity>
|
|
compared with
|
|
<quantity id="53409B0FEB72ED5FFBE4FE7467B28788" box="[1025,1173,475,499]" metricMagnitude="-3" metricUnit="m" metricValue="9.0" metricValueMax="9.8" metricValueMin="8.2" pageId="26" pageNumber="519" unit="mm" value="9.0" valueMax="9.8" valueMin="8.2">8.2 to 9.8 mm</quantity>
|
|
for
|
|
<taxonomicName id="53B84D69EB72ED5FFF44FE5362038468" authorityName=", Kavanaugh" authorityYear="2015" box="[161,292,507,531]" class="Insecta" family="Carabidae" genus="Cyclosomus" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="519" phylum="Arthropoda" rank="species" species="flexuosus">
|
|
<emphasis id="A6CCEAF8EB72ED5FFF44FE5362038468" box="[161,292,507,531]" italics="true" pageId="26" pageNumber="519">C. flexuosus</emphasis>
|
|
</taxonomicName>
|
|
specimens), and males have the apical lamella of the median lobe longer and narrower in dorsal aspect (compare
|
|
<figureCitation id="0C832A6FEB72ED5FFE3EFDB461118448" box="[475,566,539,563]" captionStart="FIGURE" captionStartId="36.[171,183,1563,1582]" captionTargetBox="[120,1164,216,1526]" captionTargetId="figure-9@36.[120,1176,216,1548]" captionTargetPageId="36" captionText="FIGURE 17. Male genitalia, Cyclosomus spp., dorsal aspect. A. C. acutangulus Kavanaugh and Cueva-Dabkoski (Longchuan Jiang at Longjiang Bridge, Wuhe Township, Tengchong County, Yunnan, China); B. C. flexuosus Dejean (Karaikal, Puducherry, India); C. C. inustus Andrewes (“China”); D. C. marginatus Motschulsky (“Bengal”, India); E. C. philippinus Heller (Pili, Zambales Mountains, Luzon, Philippines); F. C. sumatrensis Bouchard (Taman Negara, Pahang, Malaysia); G. C. suturalis (Wiedemann), lectotype (“Bengala”, India); H. C. verspertilio Cueva-Dabkoski and Kavanaugh sp. nov. (New Delhi, Delhi, India). Scale lines = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.11512994" httpUri="https://zenodo.org/record/11512994/files/figure.png" pageId="26" pageNumber="519">Fig. 17F</figureCitation>
|
|
with
|
|
<figureCitation id="0C832A6FEB72ED5FFD91FDB461F28448" box="[628,725,539,563]" captionStart="FIGURE" captionStartId="36.[171,183,1563,1582]" captionTargetBox="[120,1164,216,1526]" captionTargetId="figure-9@36.[120,1176,216,1548]" captionTargetPageId="36" captionText="FIGURE 17. Male genitalia, Cyclosomus spp., dorsal aspect. A. C. acutangulus Kavanaugh and Cueva-Dabkoski (Longchuan Jiang at Longjiang Bridge, Wuhe Township, Tengchong County, Yunnan, China); B. C. flexuosus Dejean (Karaikal, Puducherry, India); C. C. inustus Andrewes (“China”); D. C. marginatus Motschulsky (“Bengal”, India); E. C. philippinus Heller (Pili, Zambales Mountains, Luzon, Philippines); F. C. sumatrensis Bouchard (Taman Negara, Pahang, Malaysia); G. C. suturalis (Wiedemann), lectotype (“Bengala”, India); H. C. verspertilio Cueva-Dabkoski and Kavanaugh sp. nov. (New Delhi, Delhi, India). Scale lines = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.11512994" httpUri="https://zenodo.org/record/11512994/files/figure.png" pageId="26" pageNumber="519">Fig. 17B</figureCitation>
|
|
).
|
|
</paragraph>
|
|
<paragraph id="940736EAEB72ED5FFF4EFD9360F8840F" blockId="26.[123,1174,219,1205]" pageId="26" pageNumber="519">
|
|
<emphasis id="A6CCEAF8EB72ED5FFF4EFD9362B08428" bold="true" box="[171,407,572,595]" pageId="26" pageNumber="519">Habitat distribution.</emphasis>
|
|
Unknown, but presumed to be restricted to the sandy shores of medium to large rivers at low elevation, like members of most other species of the genus.
|
|
</paragraph>
|
|
<paragraph id="940736EAEB72ED5FFF4EFDD360F5854F" blockId="26.[123,1174,219,1205]" pageId="26" pageNumber="519">
|
|
<emphasis id="A6CCEAF8EB72ED5FFF4EFDD362FC84E8" bold="true" box="[171,475,636,659]" pageId="26" pageNumber="519">Geographical distribution.</emphasis>
|
|
<location id="91676031EB72ED5FFE07FDD3612284E8" LSID="urn:lsid:plazi:treatment:1C1187FCEB70ED5FFF9EFB4E675882CF:91676031EB72ED5FFE07FDD3612284E8" box="[482,517,636,659]" name="We" pageId="26" pageNumber="519">We</location>
|
|
have examined a total of
|
|
<specimenCount id="82BEFD63EB72ED5FFCC4FDD4609484E8" box="[801,947,635,659]" count="14" pageId="26" pageNumber="519" type="generic">14 specimens</specimenCount>
|
|
(
|
|
<specimenCount id="82BEFD63EB72ED5FFC26FDD4670E84E8" box="[963,1065,635,659]" count="6" pageId="26" pageNumber="519" type="male">six males</specimenCount>
|
|
and
|
|
<specimenCount id="82BEFD63EB72ED5FFB85FDD463F784C8" count="8" pageId="26" pageNumber="519" type="female">eight females</specimenCount>
|
|
) from the following localities:
|
|
<materialsCitation id="24D03CB7EB72ED5FFDC1FD3363FE84A8" ID-GBIF-Occurrence="4905152338" collectionCode="IRSNB, MFNB, NHMUK, NHRS" country="Indonesia" location="Kras" municipality="Kediri" pageId="26" pageNumber="519" specimenCount="11" specimenCount-female="6" specimenCount-male="5" stateProvince="Jawa Timur">
|
|
<emphasis id="A6CCEAF8EB72ED5FFDC1FD33619C84C8" bold="true" box="[548,699,668,691]" pageId="26" pageNumber="519">
|
|
<collectingCountry id="ECAF767AEB72ED5FFDC1FD33619C84C8" box="[548,699,668,691]" name="Indonesia" pageId="26" pageNumber="519">INDONESIA</collectingCountry>
|
|
</emphasis>
|
|
:
|
|
<collectingRegion id="567CF808EB72ED5FFD2FFD3361DA84C8" box="[714,765,668,691]" country="Indonesia" name="Jawa Timur" pageId="26" pageNumber="519">Java</collectingRegion>
|
|
:
|
|
<collectingRegion id="567CF808EB72ED5FFCECFD33604884C8" box="[777,879,668,691]" country="Indonesia" name="Jawa Timur" pageId="26" pageNumber="519">East Java</collectingRegion>
|
|
(
|
|
<collectingMunicipality id="7463AC90EB72ED5FFC9AFD3460E084C8" box="[895,967,667,691]" pageId="26" pageNumber="519">Kediri</collectingMunicipality>
|
|
,
|
|
<location id="91676031EB72ED5FFC36FD33672284C8" LSID="urn:lsid:plazi:treatment:1C1187FCEB70ED5FFF9EFB4E675882CF:91676031EB72ED5FFC36FD33672284C8" box="[979,1029,668,691]" country="Indonesia" municipality="Kediri" name="Kras" pageId="26" pageNumber="519" stateProvince="Jawa Timur">Kras</location>
|
|
[
|
|
<specimenCount id="82BEFD63EB72ED5FFBF1FD3367B684C8" box="[1044,1169,667,691]" count="1" pageId="26" pageNumber="519" type="female">one female</specimenCount>
|
|
;
|
|
<collectionCode id="F2A9AE2FEB72ED5FFF9EFD1463E884A8" box="[123,207,699,723]" country="Belgium" httpUri="http://grbio.org/cool/c7r2-61q8" name="Institut Royal des Sciences Naturelles de Belgique" pageId="26" pageNumber="519">IRSNB</collectionCode>
|
|
],
|
|
</materialsCitation>
|
|
<materialsCitation id="24D03CB7EB72ED5FFF04FD1362EA84A8" ID-GBIF-Occurrence="4905152341" box="[225,461,699,723]" collectionCode="IRSNB, MFNB, NHMUK, NHRS" country="Indonesia" location="Kras" municipality="Kediri" pageId="26" pageNumber="519" specimenCount="11" specimenCount-female="6" specimenCount-male="5" stateProvince="Jawa Timur">
|
|
[
|
|
<specimenCount id="82BEFD63EB72ED5FFF0DFD13624284A8" box="[232,357,699,723]" count="1" pageId="26" pageNumber="519" type="female">one female</specimenCount>
|
|
;
|
|
<collectionCode id="F2A9AE2FEB72ED5FFE97FD1362E484A8" box="[370,451,700,723]" country="Italy" httpUri="http://grbio.org/cool/irpn-c0nq" name="Museo Friulano di Storia Naturale" pageId="26" pageNumber="519">MFNB</collectionCode>
|
|
],
|
|
</materialsCitation>
|
|
<materialsCitation id="24D03CB7EB72ED5FFE33FD1460BB84A8" ID-GBIF-Occurrence="4905152323" box="[470,924,699,723]" collectionCode="IRSNB, MFNB, NHMUK, NHRS" country="Indonesia" location="Kras" municipality="Kediri" pageId="26" pageNumber="519" specimenCount="11" specimenCount-female="6" specimenCount-male="5" stateProvince="Jawa Timur">
|
|
[
|
|
<specimenCount id="82BEFD63EB72ED5FFE39FD14616884A8" box="[476,591,699,723]" count="5" pageId="26" pageNumber="519" type="male">five males</specimenCount>
|
|
and
|
|
<specimenCount id="82BEFD63EB72ED5FFD62FD14603C84A8" box="[647,795,699,723]" count="3" pageId="26" pageNumber="519" type="female">three females</specimenCount>
|
|
;
|
|
<collectionCode id="F2A9AE2FEB72ED5FFCCDFD1360B584A8" box="[808,914,700,723]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="26" pageNumber="519" type="Museum">NHMUK</collectionCode>
|
|
],
|
|
</materialsCitation>
|
|
<materialsCitation id="24D03CB7EB72ED5FFC40FD1367B684A8" ID-GBIF-Occurrence="4905152305" box="[933,1169,699,723]" collectionCode="IRSNB, MFNB, NHMUK, NHRS" country="Indonesia" location="Kras" municipality="Kediri" pageId="26" pageNumber="519" specimenCount="11" specimenCount-female="6" specimenCount-male="5" stateProvince="Jawa Timur">
|
|
[
|
|
<specimenCount id="82BEFD63EB72ED5FFC4EFD13670E84A8" box="[939,1065,699,723]" count="1" pageId="26" pageNumber="519" type="female">one female</specimenCount>
|
|
;
|
|
<collectionCode id="F2A9AE2FEB72ED5FFBD0FD1467A484A8" box="[1077,1155,699,723]" country="Sweden" httpUri="http://grbio.org/cool/rsbi-vwmq" name="Swedish Museum of Natural History, Entomology Collections" pageId="26" pageNumber="519">NHRS</collectionCode>
|
|
])
|
|
</materialsCitation>
|
|
;
|
|
<materialsCitation id="24D03CB7EB72ED5FFF9EFD736066848F" ID-GBIF-Occurrence="4905152352" box="[123,833,732,756]" collectionCode="NHMUK" collectorName="M. Knappert" country="Indonesia" location="Manna" municipality="Mana" pageId="26" pageNumber="519" specimenCount="1" specimenCount-female="1" stateProvince="Sumatera Utara">
|
|
<collectingRegion id="567CF808EB72ED5FFF9EFD7363FF848F" box="[123,216,732,756]" country="Indonesia" name="Sumatera Utara" pageId="26" pageNumber="519">Sumatra</collectingRegion>
|
|
:
|
|
<collectingRegion id="567CF808EB72ED5FFF04FD73626E848F" box="[225,329,732,756]" country="Indonesia" name="Bengkulu" pageId="26" pageNumber="519">Bengkulu</collectingRegion>
|
|
(
|
|
<collectingMunicipality id="7463AC90EB72ED5FFEB3FD7362B3848F" box="[342,404,732,756]" pageId="26" pageNumber="519">Mana</collectingMunicipality>
|
|
[“
|
|
<location id="91676031EB72ED5FFE4EFD7362DA848F" LSID="urn:lsid:plazi:treatment:1C1187FCEB70ED5FFF9EFB4E675882CF:91676031EB72ED5FFE4EFD7362DA848F" box="[427,509,732,756]" country="Indonesia" municipality="Mana" name="Manna" pageId="26" pageNumber="519" stateProvince="Sumatera Utara">Manna</location>
|
|
”] 1902,
|
|
<collectorName id="394D533CEB72ED5FFDB3FD7361F98488" box="[598,734,732,755]" pageId="26" pageNumber="519">M. Knappert</collectorName>
|
|
collector
|
|
</materialsCitation>
|
|
<materialsCitation id="24D03CB7EB72ED5FFCADFD73676A848F" ID-GBIF-Occurrence="4905152339" box="[840,1101,732,756]" collectionCode="NHMUK" collectorName="M. Knappert" country="Indonesia" location="Manna" municipality="Mana" pageId="26" pageNumber="519" specimenCount="1" specimenCount-female="1" stateProvince="Sumatera Utara">
|
|
[
|
|
<specimenCount id="82BEFD63EB72ED5FFCABFD7360EE848F" box="[846,969,732,756]" count="1" pageId="26" pageNumber="519" type="female">one female</specimenCount>
|
|
;
|
|
<collectionCode id="F2A9AE2FEB72ED5FFC36FD736718848F" box="[979,1087,732,756]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="26" pageNumber="519" type="Museum">NHMUK</collectionCode>
|
|
])
|
|
</materialsCitation>
|
|
;
|
|
<materialsCitation id="24D03CB7EB72ED5FFBB2FD7361DD856F" ID-GBIF-Occurrence="4905152331" collectionCode="MFNB" collectorName="W. Reisch" country="Indonesia" location="Deli" municipality="Deli Serdang" pageId="26" pageNumber="519" specimenCount="1" specimenCount-female="1" stateProvince="Sumatera Utara">
|
|
<collectingRegion id="567CF808EB72ED5FFBB2FD7363F3856F" country="Indonesia" name="Sumatera Utara" pageId="26" pageNumber="519">North Sumatra</collectingRegion>
|
|
(
|
|
<collectingMunicipality id="7463AC90EB72ED5FFF06FD536255856F" box="[227,370,764,788]" pageId="26" pageNumber="519">Deli Serdang</collectingMunicipality>
|
|
[“
|
|
<location id="91676031EB72ED5FFE68FD5362E4856F" LSID="urn:lsid:plazi:treatment:1C1187FCEB70ED5FFF9EFB4E675882CF:91676031EB72ED5FFE68FD5362E4856F" box="[397,451,764,788]" country="Indonesia" municipality="Deli Serdang" name="Deli" pageId="26" pageNumber="519" stateProvince="Sumatera Utara">Deli</location>
|
|
”], 1894,
|
|
<collectorName id="394D533CEB72ED5FFDC3FD5361B2856F" box="[550,661,764,788]" pageId="26" pageNumber="519">W. Reisch</collectorName>
|
|
collector
|
|
</materialsCitation>
|
|
<materialsCitation id="24D03CB7EB72ED5FFCE7FD5360D3856F" ID-GBIF-Occurrence="4905152312" box="[770,1012,764,788]" collectionCode="MFNB" collectorName="W. Reisch" country="Indonesia" location="Deli" municipality="Deli Serdang" pageId="26" pageNumber="519" specimenCount="1" specimenCount-female="1" stateProvince="Sumatera Utara">
|
|
[
|
|
<specimenCount id="82BEFD63EB72ED5FFCECFD5360A1856F" box="[777,902,764,788]" count="1" pageId="26" pageNumber="519" type="female">one female</specimenCount>
|
|
;
|
|
<collectionCode id="F2A9AE2FEB72ED5FFC76FD5360C1856F" box="[915,998,764,788]" country="Italy" httpUri="http://grbio.org/cool/irpn-c0nq" name="Museo Friulano di Storia Naturale" pageId="26" pageNumber="519">MFNB</collectionCode>
|
|
])
|
|
</materialsCitation>
|
|
.
|
|
<materialsCitation id="24D03CB7EB72ED5FFC1AFD5360E9854F" ID-GBIF-Occurrence="4905152344" collectingDate="1992-01-05" collectionCode="CAS" collectorName="J. Nielsen" country="Malaysia" location="Taman Negara" pageId="26" pageNumber="519" specimenCount="1" specimenCount-male="1" stateProvince="Pahang">
|
|
<emphasis id="A6CCEAF8EB72ED5FFC1AFD5367A98568" bold="true" box="[1023,1166,764,787]" pageId="26" pageNumber="519">
|
|
<collectingCountry id="ECAF767AEB72ED5FFC1AFD5367A98568" box="[1023,1166,764,787]" name="Malaysia" pageId="26" pageNumber="519">MALAYSIA</collectingCountry>
|
|
</emphasis>
|
|
:
|
|
<collectingRegion id="567CF808EB72ED5FFF9EFCB363E9854F" box="[123,206,796,820]" country="Malaysia" name="Pahang" pageId="26" pageNumber="519">Pahang</collectingRegion>
|
|
:
|
|
<location id="91676031EB72ED5FFF32FCB36253854F" LSID="urn:lsid:plazi:treatment:1C1187FCEB70ED5FFF9EFB4E675882CF:91676031EB72ED5FFF32FCB36253854F" box="[215,372,796,820]" country="Malaysia" name="Taman Negara" pageId="26" pageNumber="519" stateProvince="Pahang">Taman Negara</location>
|
|
(
|
|
<date id="E006102AEB72ED5FFE61FCB3610C854F" box="[388,555,796,820]" pageId="26" pageNumber="519" value="1992-01-05">
|
|
<collectingDate id="F042E9C2EB72ED5FFE61FCB3610C854F" box="[388,555,796,820]" pageId="26" pageNumber="519" value="1992-01-05">5 January 1992</collectingDate>
|
|
</date>
|
|
,
|
|
<collectorName id="394D533CEB72ED5FFDD3FCB36187854F" box="[566,672,796,820]" pageId="26" pageNumber="519">J. Nielsen</collectorName>
|
|
collector [
|
|
<specimenCount id="82BEFD63EB72ED5FFCF6FCB3605D854F" box="[787,890,796,820]" count="1" pageId="26" pageNumber="519" type="male">one male</specimenCount>
|
|
;
|
|
<collectionCode id="F2A9AE2FEB72ED5FFC63FCB360E7854F" box="[902,960,796,820]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="26" pageNumber="519" type="Museum">CAS</collectionCode>
|
|
])
|
|
</materialsCitation>
|
|
.
|
|
</paragraph>
|
|
<paragraph id="940736EAEB72ED5FFF4EFC93625485CF" blockId="26.[123,1174,219,1205]" pageId="26" pageNumber="519">
|
|
The known geographical range of
|
|
<taxonomicName id="53B84D69EB72ED5FFDC7FC9361E6852F" box="[546,705,828,852]" class="Insecta" family="Carabidae" genus="Cyclosomus" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="519" phylum="Arthropoda" rank="species" species="sumatrensis">
|
|
<emphasis id="A6CCEAF8EB72ED5FFDC7FC9361E6852F" box="[546,705,828,852]" italics="true" pageId="26" pageNumber="519">C. sumatrensis</emphasis>
|
|
</taxonomicName>
|
|
(
|
|
<figureCitation id="0C832A6FEB72ED5FFD37FC936005852F" box="[722,802,828,852]" captionStart="FIGURE" captionStartId="40.[171,183,837,856]" captionTargetBox="[168,1128,216,826]" captionTargetId="figure-339@40.[168,1128,216,826]" captionTargetPageId="40" captionText="FIGURE 22. Map showing distribution of locality records for Cyclosomus specimens examined. White triangles = C. sumatrensis Bouchard; yellow highlighted area with black question mark = type area for C. suturalis (Wiedemann); white circles = other localities for C. suturalis; white squares = C. vespertilio Cueva-Dabkoski and Kavanaugh sp. nov. Modified from Wikimedia Commons, World Atlas of the World, at URL: <http://upload.wikimedia.org/wikipedia/ commons/8/8f/Whole_world_-_land_and_oceans_12000.jpg>. Scale line = 500 km." figureDoi="http://doi.org/10.5281/zenodo.11513004" httpUri="https://zenodo.org/record/11513004/files/figure.png" pageId="26" pageNumber="519">Fig. 22</figureCitation>
|
|
) extends from the Malaysian portion of the Malay Peninsula south to
|
|
<collectingRegion id="567CF808EB72ED5FFDE3FCF36178850F" box="[518,607,860,884]" country="Indonesia" name="Sumatera Utara" pageId="26" pageNumber="519">Sumatra</collectingRegion>
|
|
and
|
|
<collectingRegion id="567CF808EB72ED5FFD76FCF261E1850F" box="[659,710,861,884]" country="Indonesia" name="Jawa Timur" pageId="26" pageNumber="519">Java</collectingRegion>
|
|
. It was previously known only from
|
|
<collectingRegion id="567CF808EB72ED5FFBB5FCF363BD85EF" country="Indonesia" name="Sumatera Utara" pageId="26" pageNumber="519">Sumatra</collectingRegion>
|
|
,
|
|
<collectingCountry id="ECAF767AEB72ED5FFF4CFCD3623685EF" box="[169,273,892,916]" name="Indonesia" pageId="26" pageNumber="519">Indonesia</collectingCountry>
|
|
and specimens reported here from the Malay Peninsula and
|
|
<collectingRegion id="567CF808EB72ED5FFC27FCD260D685EF" box="[962,1009,893,916]" country="Indonesia" name="Jawa Timur" pageId="26" pageNumber="519">Java</collectingRegion>
|
|
represent new records for these areas.
|
|
</paragraph>
|
|
<paragraph id="940736EAEB72ED5FFF4EFC1261D5824F" blockId="26.[123,1174,219,1205]" pageId="26" pageNumber="519">
|
|
<emphasis id="A6CCEAF8EB72ED5FFF4EFC12629C85AF" bold="true" box="[171,443,957,980]" pageId="26" pageNumber="519">Geographical variation.</emphasis>
|
|
We found slight individual variation (
|
|
<figureCitation id="0C832A6FEB72ED5FFCB3FC1360C785AF" box="[854,992,956,980]" captionStart="FIGURE" captionStartId="25.[171,183,1138,1157]" captionTargetBox="[120,1176,216,1118]" captionTargetId="figure-226@25.[120,1176,216,1118]" captionTargetPageId="25" captionText="FIGURE 11. Cyclosomus sumatrensis Bouchard, dorsal habitus. A. Dark male (Kras, Kediri, East Java, Java, Indonesia); B. Pale male (Taman Negara, Pahang, Malaysia). Scale lines = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.11512980" httpUri="https://zenodo.org/record/11512980/files/figure.png" pageId="26" pageNumber="519">Figs. 11A, B</figureCitation>
|
|
) in development of the dark color pattern of the elytra within the largest sample of this species (from Kras,
|
|
<collectingRegion id="567CF808EB72ED5FFB8DFC726388826F" country="Indonesia" name="Jawa Timur" pageId="26" pageNumber="519">East Java</collectingRegion>
|
|
). Specimens from all other localities were similar to at least some specimens from that sample, so we recognized no particular geographical variation.
|
|
</paragraph>
|
|
<paragraph id="940736EAEB72ED5FFF4EFB92675882CF" blockId="26.[123,1174,219,1205]" pageId="26" pageNumber="519">
|
|
<emphasis id="A6CCEAF8EB72ED5FFF4EFB926062822F" bold="true" box="[171,837,1085,1109]" pageId="26" pageNumber="519">
|
|
Geographical relationships with other
|
|
<taxonomicName id="53B84D69EB72ED5FFD81FB926062822F" box="[612,837,1085,1109]" class="Insecta" family="Carabidae" genus="Cyclosomus" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="519" phylum="Arthropoda" rank="species" species="undetermined">
|
|
<emphasis id="A6CCEAF8EB72ED5FFD81FB9261CF822E" bold="true" box="[612,744,1085,1109]" italics="true" pageId="26" pageNumber="519">Cyclosomus</emphasis>
|
|
species.
|
|
</taxonomicName>
|
|
</emphasis>
|
|
No other congeneric species is known to occur in
|
|
<collectingCountry id="ECAF767AEB72ED5FFEACFBF3628C820F" box="[329,427,1116,1140]" name="Malaysia" pageId="26" pageNumber="519">Malaysia</collectingCountry>
|
|
or
|
|
<collectingCountry id="ECAF767AEB72ED5FFE34FBF3611B820F" box="[465,572,1116,1140]" name="Indonesia" pageId="26" pageNumber="519">Indonesia</collectingCountry>
|
|
; and the nearest
|
|
<taxonomicName id="53B84D69EB72ED5FFD16FBF36053820F" authorityName="Latreille" authorityYear="1829" box="[755,884,1116,1140]" class="Insecta" family="Carabidae" genus="Cyclosomus" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="519" phylum="Arthropoda" rank="genus">
|
|
<emphasis id="A6CCEAF8EB72ED5FFD16FBF36053820F" box="[755,884,1116,1140]" italics="true" pageId="26" pageNumber="519">Cyclosomus</emphasis>
|
|
</taxonomicName>
|
|
records are for
|
|
<taxonomicName id="53B84D69EB72ED5FFBC6FBF267A8820F" box="[1059,1167,1117,1140]" class="Insecta" family="Carabidae" genus="Cyclosomus" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="519" phylum="Arthropoda" rank="species" species="inustus">
|
|
<emphasis id="A6CCEAF8EB72ED5FFBC6FBF267A8820F" box="[1059,1167,1117,1140]" italics="true" pageId="26" pageNumber="519">C. inustus</emphasis>
|
|
</taxonomicName>
|
|
, in
|
|
<collectingCountry id="ECAF767AEB72ED5FFF7FFBD3622D82EF" box="[154,266,1148,1172]" name="Cambodia" pageId="26" pageNumber="519">Cambodia</collectingCountry>
|
|
, across the Gulf of
|
|
<collectingCountry id="ECAF767AEB72ED5FFE08FBD3616B82EF" box="[493,588,1148,1172]" name="Thailand" pageId="26" pageNumber="519">Thailand</collectingCountry>
|
|
and more than
|
|
<quantity id="53409B0FEB72ED5FFCE4FBD3607282EF" box="[769,853,1148,1172]" metricMagnitude="5" metricUnit="m" metricValue="8.0" pageId="26" pageNumber="519" unit="km" value="800.0">800 km</quantity>
|
|
distant to the northeast, and
|
|
<taxonomicName id="53B84D69EB72ED5FFF9EFB3263DA82CF" authorityName=", Kavanaugh" authorityYear="2015" box="[123,253,1180,1204]" class="Insecta" family="Carabidae" genus="Cyclosomus" kingdom="Animalia" order="Coleoptera" pageId="26" pageNumber="519" phylum="Arthropoda" rank="species" species="flexuosus">
|
|
<emphasis id="A6CCEAF8EB72ED5FFF9EFB3263DA82CF" box="[123,253,1180,1204]" italics="true" pageId="26" pageNumber="519">C. flexuosus</emphasis>
|
|
</taxonomicName>
|
|
in
|
|
<collectingCountry id="ECAF767AEB72ED5FFEC4FB3362AA82CF" box="[289,397,1180,1204]" name="Sri Lanka" pageId="26" pageNumber="519">Sri Lanka</collectingCountry>
|
|
, across the Bay of Bengal and more than
|
|
<quantity id="53409B0FEB72ED5FFCABFB33608A82CF" box="[846,941,1180,1204]" metricMagnitude="6" metricUnit="m" metricValue="1.6" pageId="26" pageNumber="519" unit="km" value="1600.0">1600 km</quantity>
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distant to the west.
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|
</paragraph>
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|
</subSubSection>
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|
</treatment>
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|
</document> |