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<document id="1B60F9351C08683AD80E41708B2E2875" ID-DOI="10.11646/zootaxa.3946.1.4" ID-GBIF-Dataset="d846b7cd-fdad-4608-b416-0abfe4444abc" ID-ISSN="1175-5326" ID-Zenodo-Dep="239416" ID-ZooBank="172B1204-0AFC-4593-ACD6-2FF94A8CF933" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461032209056" checkinUser="plazi" docAuthor="Rizzo, V. &amp; Comas, J." docDate="2015" docId="03B2D61C7E2C6D2DFF42B337FC4DCACB" docLanguage="en" docName="zt03946p112.pdf" docOrigin="Zootaxa 3946 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Troglocharinus pallisei Rizzo &amp; Comas, sp. n." docType="treatment" docVersion="9" lastPageNumber="109" masterDocId="FF8BAE647E2F6D28FFD5B028FFF8CB47" masterDocTitle="A new species of Troglocharinus Reitter, 1908 (Coleoptera, Leiodidae, Cholevinae, Leptodirini) from southern Catalonia, with a molecular phylogeny of the related species group" masterLastPageNumber="112" masterPageNumber="104" pageNumber="107" updateTime="1698573624724" updateUser="plazi">
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<mods:title id="1155DEA9B02C8BD96AF992D7884E18E3">A new species of Troglocharinus Reitter, 1908 (Coleoptera, Leiodidae, Cholevinae, Leptodirini) from southern Catalonia, with a molecular phylogeny of the related species group</mods:title>
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<mods:namePart id="3A679F6EB0617303BE562B836E958FA2">Rizzo, V.</mods:namePart>
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<mods:namePart id="B02EA137446F3061ADFF09D7189A0817">Comas, J.</mods:namePart>
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<treatment id="03B2D61C7E2C6D2DFF42B337FC4DCACB" ID-DOI="http://doi.org/10.5281/zenodo.5679725" ID-GBIF-Taxon="119608355" ID-Zenodo-Dep="5679725" LSID="urn:lsid:plazi:treatment:03B2D61C7E2C6D2DFF42B337FC4DCACB" httpUri="http://treatment.plazi.org/id/03B2D61C7E2C6D2DFF42B337FC4DCACB" lastPageId="5" lastPageNumber="109" pageId="3" pageNumber="107">
<subSubSection id="C30134817E2C6D2BFF42B337FEF3C81B" pageId="3" pageNumber="107" type="nomenclature">
<paragraph id="8BA4670A7E2C6D2BFF42B337FD4BC87F" blockId="3.[151,691,799,861]" box="[151,691,799,824]" pageId="3" pageNumber="107">
<heading id="D0ECD0667E2C6D2BFF42B337FD4BC87F" bold="true" box="[151,691,799,824]" fontSize="10" level="2" pageId="3" pageNumber="107" reason="2">
<emphasis id="B96FBB187E2C6D2BFF42B337FD4BC87F" bold="true" box="[151,691,799,824]" pageId="3" pageNumber="107">
<taxonomicName id="4C1B1C897E2C6D2BFF42B337FD9DC87F" ID-CoL="598PX" authority="Rizzo &amp; Comas" authorityName="Rizzo &amp; Comas" box="[151,613,799,824]" class="Insecta" family="Leiodidae" genus="Troglocharinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="107" phylum="Arthropoda" rank="species" species="pallisei" status="sp. nov.">
<emphasis id="B96FBB187E2C6D2BFF42B337FE5BC87F" bold="true" box="[151,419,799,824]" italics="true" pageId="3" pageNumber="107">Troglocharinus pallisei</emphasis>
Rizzo &amp; Comas
</taxonomicName>
,
<taxonomicNameLabel id="A25C06637E2C6D2BFDA6B337FD4BC87F" box="[627,691,799,824]" pageId="3" pageNumber="107" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BA4670A7E2C6D2BFF42B36CFEF3C81B" blockId="3.[151,691,799,861]" box="[151,267,836,861]" pageId="3" pageNumber="107">
(
<figureCitation id="13207B8F7E2C6D2BFF4AB36CFEFAC81B" box="[159,258,836,861]" captionStart-0="FIGURE 2" captionStart-1="FIGURE 3" captionStart-2="FIGURE 4, 5" captionStart-3="FIGURE 6, 7" captionStart-4="FIGURE 8, 9" captionStartId-0="4.[151,250,1370,1392]" captionStartId-1="4.[151,250,1989,2011]" captionStartId-2="5.[151,250,1529,1551]" captionStartId-3="6.[151,250,1121,1143]" captionStartId-4="6.[151,250,1564,1586]" captionTargetBox-0="[491,1095,176,1347]" captionTargetBox-2="[154,1386,484,1476]" captionTargetBox-3="[546,1036,302,1083]" captionTargetBox-4="[213,1354,1193,1542]" captionTargetId-0="figure@4.[491,1096,161,1348]" captionTargetId-2="figure@5.[151,1436,448,1507]" captionTargetId-3="figure@6.[542,1046,297,1098]" captionTargetId-4="figure@6.[213,1374,1193,1542]" captionTargetPageId-0="4" captionTargetPageId-2="5" captionTargetPageId-3="6" captionTargetPageId-4="6" captionText-0="FIGURE 2. Habitus of Troglocharinus pallisei sp. n. (holotype, male). Scale bar: 0.5 mm." captionText-1="FIGURE 3. Troglocharinus pallisei sp. n., process of the mesoventrite." captionText-2="FIGURE 4, 5. Aedeagus of Troglocharinus pallisei sp. n.; 4) lateral view; 5) ventral view. Scale bar: 0.5 mm." captionText-3="FIGURE 6, 7. Aedeagus of Troglocharinus pallisei sp. n.; 6) apex of the paramere; 7) internal sac (endophallus)." captionText-4="FIGURE 8, 9. Troglocharinus pallisei sp. n. 8) spermatheca; 9) visible female abdominal sternite." httpUri-0="https://zenodo.org/record/239418/files/figure.png" httpUri-2="https://zenodo.org/record/239419/files/figure.png" httpUri-3="https://zenodo.org/record/239420/files/figure.png" httpUri-4="https://zenodo.org/record/239421/files/figure.png" pageId="3" pageNumber="107">Figs 29</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C30134817E2C6D2BFF42B3A4FECECF3B" pageId="3" pageNumber="107" type="materials_examined">
<paragraph id="8BA4670A7E2C6D2BFF42B3A4FECECF3B" blockId="3.[151,1437,908,1941]" pageId="3" pageNumber="107">
<emphasis id="B96FBB187E2C6D2BFF42B3A4FDAEC8E2" bold="true" box="[151,598,908,933]" pageId="3" pageNumber="107">
<typeStatus id="54A0D9A87E2C6D2BFF42B3A4FF2AC8E2" box="[151,210,908,933]" pageId="3" pageNumber="107">Type</typeStatus>
material.
<typeStatus id="54A0D9A87E2C6D2BFE9BB3A4FE47C8E2" box="[334,447,908,933]" pageId="3" pageNumber="107" type="holotype">Holotype</typeStatus>
: COUNTRY
</emphasis>
:
<collectingCountry id="F30C279A7E2C6D2BFDB3B3A4FD52C8E2" box="[614,682,908,933]" name="Spain" pageId="3" pageNumber="107">Spain</collectingCountry>
:
<emphasis id="B96FBB187E2C6D2BFD63B3A4FCE7C8E2" bold="true" box="[694,799,908,933]" pageId="3" pageNumber="107">Province</emphasis>
: Tarragona: ♂, labelled “Cv. de la Riba o / del Talc -La Riba- / (Alt Camp)
<date id="FFA541CA7E2C6D2BFEA8B398FE2AC88F" box="[381,466,944,968]" pageId="3" pageNumber="107" value="2012-02">II-2012</date>
/ R. Sendra leg.; X 34895.7 /
<collectingCountry id="F30C279A7E2C6D2BFCEDB398FCB4C880" box="[824,844,944,967]" name="Italy" pageId="3" pageNumber="107">Y</collectingCountry>
4575816.3 / Z 240 /
<collectionCode id="ED0AFFCF7E2C6D2BFB9CB398FB68C88F" box="[1097,1168,944,968]" pageId="3" pageNumber="107">ETRS</collectionCode>
89” (
<collectionCode id="ED0AFFCF7E2C6D2BFB1DB398FAFDC88F" box="[1224,1285,944,968]" httpUri="http://grbio.org/cool/e3tf-9780" name="Colecao de Culturas de Basidiomicetos" pageId="3" pageNumber="107">CCB</collectionCode>
).
<emphasis id="B96FBB187E2C6D2BFACEB387FED9C8AA" bold="true" pageId="3" pageNumber="107">
<typeStatus id="54A0D9A87E2C6D2BFACEB387FA6FC88F" box="[1307,1431,943,968]" pageId="3" pageNumber="107" type="paratype">Paratypes</typeStatus>
: COUNTRY
</emphasis>
:
<collectingCountry id="F30C279A7E2C6D2BFEFAB3FCFE8BC8AA" box="[303,371,980,1005]" name="Spain" pageId="3" pageNumber="107">Spain</collectingCountry>
:
<emphasis id="B96FBB187E2C6D2BFEA8B3FCFE1EC8AA" bold="true" box="[381,486,980,1005]" pageId="3" pageNumber="107">Province</emphasis>
: Tarragona: all with same locality data as
<typeStatus id="54A0D9A87E2C6D2BFC6BB3FCFBDDC8AA" box="[958,1061,980,1005]" pageId="3" pageNumber="107" type="holotype">holotype</typeStatus>
:
<specimenCount id="9D1DAC837E2C6D2BFBFBB3FCFBA2C8AB" box="[1070,1114,980,1004]" pageId="3" pageNumber="107" type="generic">1 ♂</specimenCount>
, 1 ♀ same date and collector as
<typeStatus id="54A0D9A87E2C6D2BFF62B3DFFEE5CF57" box="[183,285,1015,1040]" pageId="3" pageNumber="107" type="holotype">holotype</typeStatus>
;
<specimenCount id="9D1DAC837E2C6D2BFEFCB3D0FEAFCF48" box="[297,343,1016,1040]" pageId="3" pageNumber="107" type="generic">3 ♂</specimenCount>
, 2 ♀,
<date id="FFA541CA7E2C6D2BFE75B3D0FDF8CF57" box="[416,512,1016,1040]" pageId="3" pageNumber="107" value="2012-03">III-2012</date>
, R. Sendra leg.;
<specimenCount id="9D1DAC837E2C6D2BFD6AB3D0FD15CF48" box="[703,749,1016,1040]" pageId="3" pageNumber="107" type="generic">4 ♂</specimenCount>
, 13 ♀,
<date id="FFA541CA7E2C6D2BFC91B3D0FC2CCF57" box="[836,980,1016,1040]" pageId="3" pageNumber="107" value="2013-07-28">28-VII-2013</date>
, J. Pallisé leg. (1 ♀,
<specimenCount id="9D1DAC837E2C6D2BFB10B3D0FB09CF48" box="[1221,1265,1016,1040]" pageId="3" pageNumber="107" type="generic">1 ♂</specimenCount>
used for
<collectionCode id="ED0AFFCF7E2C6D2BFA8AB3D0FA64CF57" LSID="urn:lsid:biocol.org:col:15536" box="[1375,1436,1016,1040]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15536" name="Department of Natural Resources, Environment, The Arts and Sport" pageId="3" pageNumber="107">DNA</collectionCode>
extraction, voucher numbers IBE-RA1190 and IBE-RA1191 respectively);
<specimenCount id="9D1DAC837E2C6D2BFC3FB434FBEFCF73" box="[1002,1047,1052,1076]" pageId="3" pageNumber="107" type="generic">3 ♂</specimenCount>
, 1 ♀ (
<specimenCount id="9D1DAC837E2C6D2BFBB7B434FB73CF73" box="[1122,1163,1052,1076]" pageId="3" pageNumber="107" type="generic">1 ♂</specimenCount>
damaged),
<date id="FFA541CA7E2C6D2BFAC4B434FA7ACF73" box="[1297,1410,1052,1076]" pageId="3" pageNumber="107" value="1971-07-17">17-VII-71</date>
J. Pallisé leg. (
<collectionCode id="ED0AFFCF7E2C6D2BFECAB468FE8BCF1F" box="[287,371,1088,1112]" httpUri="http://grbio.org/cool/fs2t-231f" name="Museu de Ciències Naturals de Barcelona" pageId="3" pageNumber="107">MCNB</collectionCode>
Ref. Nos 74-3360
<collectionCode id="ED0AFFCF7E2C6D2BFD92B468FD7ECF10" box="[583,646,1088,1111]" httpUri="http://grbio.org/cool/b9gz-izvg" name="Museum Zoologicum Bogoriense" pageId="3" pageNumber="107">MZB</collectionCode>
, 74-3361
<collectionCode id="ED0AFFCF7E2C6D2BFD23B468FCCDCF10" box="[758,821,1088,1111]" httpUri="http://grbio.org/cool/b9gz-izvg" name="Museum Zoologicum Bogoriense" pageId="3" pageNumber="107">MZB</collectionCode>
, 74-3362
<collectionCode id="ED0AFFCF7E2C6D2BFC76B468FC18CF10" box="[931,992,1088,1111]" httpUri="http://grbio.org/cool/b9gz-izvg" name="Museum Zoologicum Bogoriense" pageId="3" pageNumber="107">MZB</collectionCode>
and 74-3363
<collectionCode id="ED0AFFCF7E2C6D2BFBACB468FB4DCF10" box="[1145,1205,1088,1111]" httpUri="http://grbio.org/cool/b9gz-izvg" name="Museum Zoologicum Bogoriense" pageId="3" pageNumber="107">MZB</collectionCode>
respectively) (
<collectionCode id="ED0AFFCF7E2C6D2BFA89B468FA60CF1F" box="[1372,1432,1088,1112]" httpUri="http://grbio.org/cool/e3tf-9780" name="Colecao de Culturas de Basidiomicetos" pageId="3" pageNumber="107">CCB</collectionCode>
,
<collectionCode id="ED0AFFCF7E2C6D2BFF42B44DFF3FCF3B" box="[151,199,1125,1148]" httpUri="http://grbio.org/cool/qyi3-sxew" name="Institut de Biologia Evolutiva, (CSIC-UPF)" pageId="3" pageNumber="107">IBE</collectionCode>
,
<collectionCode id="ED0AFFCF7E2C6D2BFF07B44CFED2CF3B" box="[210,298,1124,1148]" httpUri="http://grbio.org/cool/fs2t-231f" name="Museu de Ciències Naturals de Barcelona" pageId="3" pageNumber="107">MCNB</collectionCode>
).
</paragraph>
</subSubSection>
<subSubSection id="C30134817E2C6D2BFF12B4AFFC36CE77" pageId="3" pageNumber="107" type="diagnosis">
<paragraph id="8BA4670A7E2C6D2BFF12B4AFFC36CE77" blockId="3.[151,1437,908,1941]" pageId="3" pageNumber="107">
<emphasis id="B96FBB187E2C6D2BFF12B4AFFEB9CFE7" bold="true" box="[199,321,1159,1184]" pageId="3" pageNumber="107">Diagnosis.</emphasis>
The new species can be separated from the geographically closest species of
<taxonomicName id="4C1B1C897E2C6D2BFB71B4AFFAB7CFE7" box="[1188,1359,1159,1184]" class="Insecta" family="Leiodidae" genus="Troglocharinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="107" phylum="Arthropoda" rank="genus">
<emphasis id="B96FBB187E2C6D2BFB71B4AFFAB7CFE7" box="[1188,1359,1159,1184]" italics="true" pageId="3" pageNumber="107">Troglocharinus</emphasis>
</taxonomicName>
by the presence of a brush of setae in the apex of the parameres of the adeagus (a penicillus, see
<bibRefCitation id="EF8A1AFB7E2C6D2BFB7CB484FA6CCF83" author="Fresneda" box="[1193,1428,1196,1221]" pageId="3" pageNumber="107" refString="Fresneda, J., Salgado, J. &amp; Ribera, I. (2007) Phylogeny of western Mediterranean Leptodirini, with an emphasis on genital characters (Coleoptera: Leiodidae: Cholevinae). Systematic Entomology, 32 (2), 332 - 358. http: // dx. doi. org / 10.1111 / j. 1365 - 3113.2006.00366. x" type="journal article" year="2007">
Fresneda
<emphasis id="B96FBB187E2C6D2BFAC2B485FAB2CF82" box="[1303,1354,1196,1221]" italics="true" pageId="3" pageNumber="107">et al</emphasis>
. 2007
</bibRefCitation>
) (absent in
<taxonomicName id="4C1B1C897E2C6D2BFED8B4F9FE84CFAF" box="[269,380,1233,1256]" class="Insecta" family="Leiodidae" genus="Troglocharinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="107" phylum="Arthropoda" rank="species" species="orcinus">
<emphasis id="B96FBB187E2C6D2BFED8B4F9FE84CFAF" box="[269,380,1233,1256]" italics="true" pageId="3" pageNumber="107">T. orcinus</emphasis>
</taxonomicName>
), the slightly sinuated sides or the pronotum (regularly curved in
<taxonomicName id="4C1B1C897E2C6D2BFBB7B4F9FB29CFAF" box="[1122,1233,1233,1256]" class="Insecta" family="Leiodidae" genus="Troglocharinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="107" phylum="Arthropoda" rank="species" species="orcinus">
<emphasis id="B96FBB187E2C6D2BFBB7B4F9FB29CFAF" box="[1122,1233,1233,1256]" italics="true" pageId="3" pageNumber="107">T. orcinus</emphasis>
</taxonomicName>
), the shape of the process of the mesoventrite (differently conformed in
<taxonomicName id="4C1B1C897E2C6D2BFD25B4DDFBF6CE4B" authority="Jeannel, 1930" authorityName="Jeannel" authorityYear="1930" box="[752,1038,1268,1293]" class="Insecta" family="Leiodidae" genus="Troglocharinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="107" phylum="Arthropoda" rank="species" species="espanoli">
<emphasis id="B96FBB187E2C6D2BFD25B4DDFC92CE4A" box="[752,874,1268,1293]" italics="true" pageId="3" pageNumber="107">T. espanoli</emphasis>
<bibRefCitation id="EF8A1AFB7E2C6D2BFCA5B4DCFBF6CE4B" author="Jeannel" box="[880,1038,1268,1293]" pageId="3" pageNumber="107" refString="Jeannel, R. (1930) Bathysciinae nouveaux d'Espagne. Butlleti Institucio Catatalana d'Historia Natural, 30 (6), 90 - 91." type="journal article" year="1930">Jeannel, 1930</bibRefCitation>
</taxonomicName>
), and the presence of long and erect pubescence (finer and more adpressed in both
<taxonomicName id="4C1B1C897E2C6D2BFD4AB531FCF5CE77" box="[671,781,1305,1328]" class="Insecta" family="Leiodidae" genus="Troglocharinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="107" phylum="Arthropoda" rank="species" species="orcinus">
<emphasis id="B96FBB187E2C6D2BFD4AB531FCF5CE77" box="[671,781,1305,1328]" italics="true" pageId="3" pageNumber="107">T. orcinus</emphasis>
</taxonomicName>
and
<taxonomicName id="4C1B1C897E2C6D2BFC91B531FC46CE77" box="[836,958,1303,1328]" class="Insecta" family="Leiodidae" genus="Troglocharinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="107" phylum="Arthropoda" rank="species" species="espanoli">
<emphasis id="B96FBB187E2C6D2BFC91B531FC46CE77" box="[836,958,1303,1328]" italics="true" pageId="3" pageNumber="107">T. espanoli</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
<subSubSection id="C30134817E2C6D2DFF12B514FF0ACA67" lastPageId="5" lastPageNumber="109" pageId="3" pageNumber="107" type="description">
<paragraph id="8BA4670A7E2C6D2BFF12B514FE06CE3F" blockId="3.[151,1437,908,1941]" pageId="3" pageNumber="107">
<emphasis id="B96FBB187E2C6D2BFF12B514FDEFCE12" bold="true" box="[199,535,1340,1365]" pageId="3" pageNumber="107">
Description of the
<typeStatus id="54A0D9A87E2C6D2BFE7FB514FDEBCE12" box="[426,531,1340,1365]" pageId="3" pageNumber="107" type="holotype">holotype</typeStatus>
.
</emphasis>
Total length (head excluded)
<quantity id="4CE3CAEF7E2C6D2BFCA4B514FC34CE13" box="[881,972,1340,1364]" metricMagnitude="-3" metricUnit="m" metricValue="3.2" pageId="3" pageNumber="107" unit="mm" value="3.2">3.2 mm</quantity>
; anophthalmous; body colour yellowish, with long and erect pubescence.
</paragraph>
<paragraph id="8BA4670A7E2C6D2BFF12B5ACFE5DCE87" blockId="3.[151,1437,908,1941]" pageId="3" pageNumber="107">Head surface finely punctured, with long and erect pubescence. Antennae long, longer than apex of elytra by ca. 1/5th of their length.</paragraph>
<paragraph id="8BA4670A7E2C6D2BFF12B5E5FD90CD08" blockId="3.[151,1437,908,1941]" pageId="3" pageNumber="107">
Pronotum length
<quantity id="4CE3CAEF7E2C6D2BFE58B5E4FE0DCEA3" box="[397,501,1484,1508]" metricMagnitude="-4" metricUnit="m" metricValue="9.0" pageId="3" pageNumber="107" unit="mm" value="0.9">0.90 mm</quantity>
, trapezoidal, maximum width (
<quantity id="4CE3CAEF7E2C6D2BFC8EB5E4FC3CCEA3" box="[859,964,1484,1509]" metricMagnitude="-3" metricUnit="m" metricValue="1.06" pageId="3" pageNumber="107" unit="mm" value="1.06">1.06 mm</quantity>
) in anterior part; anterior margin slightly concave, angles straight and slightly rounded; posterior margin regularly curved; anterior part of lateral margins curved, middle part straight and slightly sinuated before posterior angles; posterior angles obtuse. Surface of pronotum with well impressed punctures.
</paragraph>
<paragraph id="8BA4670A7E2C6D2BFF12B674FA90CDA7" blockId="3.[151,1437,908,1941]" pageId="3" pageNumber="107">
Elytra oval, elongated, maximum width in anterior third to middle, narrowing posteriorly; apex round, with elytra slightly opened at apex; surface with strongly impressed striolae associated to equally impressed punctures; striolae effacing towards apex; elytral pubescence long, erected (
<figureCitation id="13207B8F7E2C6D2BFC7CB68CFC0ECDFB" box="[937,1014,1700,1725]" captionStart="FIGURE 2" captionStartId="4.[151,250,1370,1392]" captionTargetBox="[491,1095,176,1347]" captionTargetId="figure@4.[491,1096,161,1348]" captionTargetPageId="4" captionText="FIGURE 2. Habitus of Troglocharinus pallisei sp. n. (holotype, male). Scale bar: 0.5 mm." httpUri="https://zenodo.org/record/239418/files/figure.png" pageId="3" pageNumber="107">Fig. 2</figureCitation>
). Process of the mesoventrite well developed, with a variable posterior acute extension (
<figureCitation id="13207B8F7E2C6D2BFD38B6EFFCC9CDA7" box="[749,817,1735,1760]" captionStart="FIGURE 3" captionStartId="4.[151,250,1989,2011]" captionText="FIGURE 3. Troglocharinus pallisei sp. n., process of the mesoventrite." pageId="3" pageNumber="107">Fig. 3</figureCitation>
); visible abdominal sternites with glandular pores.
</paragraph>
<paragraph id="8BA4670A7E2C6D2BFF12B6C4FDBFCCD3" blockId="3.[151,1437,908,1941]" pageId="3" pageNumber="107">
Aedeagus as in
<figureCitation id="13207B8F7E2C6D2BFE56B6C4FE13CC43" box="[387,491,1772,1797]" captionStart-0="FIGURE 4, 5" captionStart-1="FIGURE 6, 7" captionStartId-0="5.[151,250,1529,1551]" captionStartId-1="6.[151,250,1121,1143]" captionTargetBox-0="[154,1386,484,1476]" captionTargetBox-1="[546,1036,302,1083]" captionTargetId-0="figure@5.[151,1436,448,1507]" captionTargetId-1="figure@6.[542,1046,297,1098]" captionTargetPageId-0="5" captionTargetPageId-1="6" captionText-0="FIGURE 4, 5. Aedeagus of Troglocharinus pallisei sp. n.; 4) lateral view; 5) ventral view. Scale bar: 0.5 mm." captionText-1="FIGURE 6, 7. Aedeagus of Troglocharinus pallisei sp. n.; 6) apex of the paramere; 7) internal sac (endophallus)." httpUri-0="https://zenodo.org/record/239419/files/figure.png" httpUri-1="https://zenodo.org/record/239420/files/figure.png" pageId="3" pageNumber="107">Figs 47</figureCitation>
, curved in lateral view, forming an obtuse angle; apex slightly bended ventrally; parameres with three apical thick setae, with a penicillus (
<figureCitation id="13207B8F7E2C6D2BFCE5B727FC8ECC6F" box="[816,886,1807,1832]" captionStart="FIGURE 6, 7" captionStartId="6.[151,250,1121,1143]" captionTargetBox="[546,1036,302,1083]" captionTargetId="figure@6.[542,1046,297,1098]" captionTargetPageId="6" captionText="FIGURE 6, 7. Aedeagus of Troglocharinus pallisei sp. n.; 6) apex of the paramere; 7) internal sac (endophallus)." httpUri="https://zenodo.org/record/239420/files/figure.png" pageId="3" pageNumber="107">Fig. 6</figureCitation>
); internal sac (endophallus) typical of the genus
<taxonomicName id="4C1B1C897E2C6D2BFF42B71CFEBBCC0A" box="[151,323,1844,1869]" class="Insecta" family="Leiodidae" genus="Troglocharinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="107" phylum="Arthropoda" rank="genus">
<emphasis id="B96FBB187E2C6D2BFF42B71CFEBBCC0A" box="[151,323,1844,1869]" italics="true" pageId="3" pageNumber="107">Troglocharinus</emphasis>
</taxonomicName>
(
<figureCitation id="13207B8F7E2C6D2BFE82B71CFE59CC0B" box="[343,417,1844,1869]" captionStart="FIGURE 6, 7" captionStartId="6.[151,250,1121,1143]" captionTargetBox="[546,1036,302,1083]" captionTargetId="figure@6.[542,1046,297,1098]" captionTargetPageId="6" captionText="FIGURE 6, 7. Aedeagus of Troglocharinus pallisei sp. n.; 6) apex of the paramere; 7) internal sac (endophallus)." httpUri="https://zenodo.org/record/239420/files/figure.png" pageId="3" pageNumber="107">Fig. 7</figureCitation>
), with a sclerotized basal &quot;Y-piece&quot; with divergent arms, a complex medial part with feather-like structures and strongly sclerotized nodules at the base of the reinforcing bands of the internal sac (
<bibRefCitation id="EF8A1AFB7E2C6D2BFF4BB754FEBFCCD3" author="Fresneda" box="[158,327,1916,1941]" pageId="3" pageNumber="107" refString="Fresneda, J. (1998) Revision de los generos de Leptodirinae de la seccion Speonomus del Sur de los Pirineos: Generos Antrocharidius Jeannel 1910, Perriniella Jeannel 1910, Speonomus Jeannel 1908, Troglocharinus Reitter 1908 y Troglophyes Abeille, 1894 (Coleoptera, Cholevidae). Memoires de Biospeologie, 25, 53 - 85" type="journal article" year="1998">Fresneda 1998</bibRefCitation>
;
<bibRefCitation id="EF8A1AFB7E2C6D2BFE86B754FDC3CCD3" author="Fresneda" box="[339,571,1916,1941]" pageId="3" pageNumber="107" refString="Fresneda, J., Salgado, J. &amp; Ribera, I. (2007) Phylogeny of western Mediterranean Leptodirini, with an emphasis on genital characters (Coleoptera: Leiodidae: Cholevinae). Systematic Entomology, 32 (2), 332 - 358. http: // dx. doi. org / 10.1111 / j. 1365 - 3113.2006.00366. x" type="journal article" year="2007">
Fresneda
<emphasis id="B96FBB187E2C6D2BFE6AB755FE08CCD2" box="[447,496,1916,1941]" italics="true" pageId="3" pageNumber="107">et al</emphasis>
. 2007
</bibRefCitation>
).
</paragraph>
<caption id="DF6437827E2B6D2CFF42B572FBCCCE37" box="[151,1076,1370,1392]" httpUri="https://zenodo.org/record/239418/files/figure.png" pageId="4" pageNumber="108" targetBox="[491,1095,176,1347]" targetPageId="4">
<paragraph id="8BA4670A7E2B6D2CFF42B572FBCCCE37" blockId="4.[151,1076,1370,1392]" box="[151,1076,1370,1392]" pageId="4" pageNumber="108">
<emphasis id="B96FBB187E2B6D2CFF42B572FEF5CE28" bold="true" box="[151,269,1370,1392]" pageId="4" pageNumber="108">FIGURE 2</emphasis>
. Habitus of
<taxonomicName id="4C1B1C897E2B6D2CFE5CB572FD8ECE37" box="[393,630,1370,1392]" class="Insecta" family="Leiodidae" genus="Troglocharinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="108" phylum="Arthropoda" rank="species" species="pallisei" status="sp. nov.">
<emphasis id="B96FBB187E2B6D2CFE5CB572FD8ECE37" box="[393,630,1370,1392]" italics="true" pageId="4" pageNumber="108">Troglocharinus pallisei</emphasis>
</taxonomicName>
<emphasis id="B96FBB187E2B6D2CFDA9B572FD4CCE28" bold="true" box="[636,692,1370,1391]" pageId="4" pageNumber="108">
<taxonomicNameLabel id="A25C06637E2B6D2CFDA9B572FD4CCE28" box="[636,692,1370,1391]" pageId="4" pageNumber="108" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
(holotype, male). Scale bar: 0.5 mm.
</paragraph>
</caption>
<caption id="DF6437827E2B6D2CFF42B7EDFC96CC9D" box="[151,878,1988,2011]" pageId="4" pageNumber="108">
<paragraph id="8BA4670A7E2B6D2CFF42B7EDFC96CC9D" blockId="4.[151,878,1988,2011]" box="[151,878,1988,2011]" pageId="4" pageNumber="108">
<emphasis id="B96FBB187E2B6D2CFF42B7EDFEF5CC9D" bold="true" box="[151,269,1989,2011]" pageId="4" pageNumber="108">FIGURE 3</emphasis>
.
<taxonomicName id="4C1B1C897E2B6D2CFECCB7ECFDFDCC9D" box="[281,517,1988,2010]" class="Insecta" family="Leiodidae" genus="Troglocharinus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="108" phylum="Arthropoda" rank="species" species="pallisei" status="sp. nov.">
<emphasis id="B96FBB187E2B6D2CFECCB7ECFDFDCC9D" box="[281,517,1988,2010]" italics="true" pageId="4" pageNumber="108">Troglocharinus pallisei</emphasis>
</taxonomicName>
<emphasis id="B96FBB187E2B6D2CFDDEB7EDFDBCCC9D" bold="true" box="[523,580,1989,2010]" pageId="4" pageNumber="108">
<taxonomicNameLabel id="A25C06637E2B6D2CFDDEB7EDFDBCCC9D" box="[523,580,1989,2010]" pageId="4" pageNumber="108" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
, process of the mesoventrite.
</paragraph>
</caption>
<paragraph id="8BA4670A7E2A6D2DFF20B0B3FF0ACA67" blockId="5.[151,1436,155,396]" pageId="5" pageNumber="109">
<emphasis id="B96FBB187E2A6D2DFF20B0B3FE83CBF3" bold="true" box="[245,379,155,180]" pageId="5" pageNumber="109">Variability.</emphasis>
Total length
<specimenCount id="9D1DAC837E2A6D2DFDD8B0B4FDC6CBF4" box="[525,574,156,180]" pageId="5" pageNumber="109" type="generic">2 ♂</specimenCount>
<quantity id="4CE3CAEF7E2A6D2DFD90B0B3FD11CBF4" box="[581,745,155,180]" metricMagnitude="-3" metricUnit="m" metricValue="3.21" metricValueMax="3.27" metricValueMin="3.15" pageId="5" pageNumber="109" unit="mm" value="3.21" valueMax="3.27" valueMin="3.15">3.153.27 mm</quantity>
,
<quantity id="4CE3CAEF7E2A6D2DFD20B0B4FCDDCBF3" box="[757,805,156,180]" metricMagnitude="-3" metricUnit="kg" metricValue="2.0" pageId="5" pageNumber="109" unit="g" value="2.0">2 g</quantity>
<quantity id="4CE3CAEF7E2A6D2DFCF9B0B3FC29CBF4" box="[812,977,155,180]" metricMagnitude="-3" metricUnit="m" metricValue="3.2249999999999996" metricValueMax="3.4" metricValueMin="3.05" pageId="5" pageNumber="109" unit="mm" value="3.2249999999999996" valueMax="3.4" valueMin="3.05">3.053.40 mm</quantity>
. Females with external morphology as in males except for non-dilated protarsi; spermatheca long and slender (
<figureCitation id="13207B8F7E2A6D2DFC6BB097FBFDCB9F" box="[958,1029,191,216]" captionStart="FIGURE 8, 9" captionStartId="6.[151,250,1564,1586]" captionTargetBox="[213,1354,1193,1542]" captionTargetId="figure@6.[213,1374,1193,1542]" captionTargetPageId="6" captionText="FIGURE 8, 9. Troglocharinus pallisei sp. n. 8) spermatheca; 9) visible female abdominal sternite." httpUri="https://zenodo.org/record/239421/files/figure.png" pageId="5" pageNumber="109">Fig. 8</figureCitation>
); basal and apical lobes forming a straight angle; accessory gland next to insertion of basal duct, visible abdominal sternites without glandular pores (
<figureCitation id="13207B8F7E2A6D2DFF4AB12FFF1CCA67" box="[159,228,263,288]" captionStart="FIGURE 8, 9" captionStartId="6.[151,250,1564,1586]" captionTargetBox="[213,1354,1193,1542]" captionTargetId="figure@6.[213,1374,1193,1542]" captionTargetPageId="6" captionText="FIGURE 8, 9. Troglocharinus pallisei sp. n. 8) spermatheca; 9) visible female abdominal sternite." httpUri="https://zenodo.org/record/239421/files/figure.png" pageId="5" pageNumber="109">Fig. 9</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C30134817E2A6D2DFF12B103FC4DCACB" pageId="5" pageNumber="109" type="etymology">
<paragraph id="8BA4670A7E2A6D2DFF12B103FC4DCACB" blockId="5.[151,1436,155,396]" pageId="5" pageNumber="109">
<emphasis id="B96FBB187E2A6D2DFF12B103FEB6CA03" bold="true" box="[199,334,299,324]" pageId="5" pageNumber="109">Etymology.</emphasis>
Dedicated to the speleologist Joan Pallisè, for his long and active explorations in this area.
<emphasis id="B96FBB187E2A6D2DFF12B167FE00CA2F" bold="true" box="[199,504,335,360]" pageId="5" pageNumber="109">Distribution and habitat.</emphasis>
Only known from the
<typeStatus id="54A0D9A87E2A6D2DFCDEB178FCC3CA2F" box="[779,827,336,360]" pageId="5" pageNumber="109">type</typeStatus>
locality. The new species can be considered a truly troglobiont both in morphology and habitat (in the sense of
<bibRefCitation id="EF8A1AFB7E2A6D2DFCE1B15BFC52CACB" author="Sket" box="[820,938,371,396]" pageId="5" pageNumber="109" refString="Sket, B. (2008) Can we agree on an ecological classification of subterranean animals? Journal of Natural History, 42, 1549 - 1563." type="journal article" year="2008">Sket 2008</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>