763 lines
100 KiB
XML
763 lines
100 KiB
XML
<document id="5F27F99AF2FAB93BB6A9174F91528C62" ID-DOI="10.5281/zenodo.214789" ID-GBIF-Dataset="4c5af5bf-c291-4031-86c9-f10683309f4c" ID-ISSN="1175-5326" ID-Zenodo-Dep="214789" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460517754413" checkinUser="plazi" docAuthor="Coad, Brian W. & Bogutskaya, Nina G." docDate="2012" docId="03A11D7EFFCCFFBDFF1AFE40E1B3E75F" docLanguage="en" docName="zt03453p055.pdf" docOrigin="Zootaxa 3453" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Alburnoides holciki Coad & Bogutskaya, 2012, new species" docType="treatment" docVersion="9" lastPageNumber="54" masterDocId="FF986506FFCDFFB6FF8DFFD4E027E563" masterDocTitle="A new species of riffle minnow, Alburnoides holciki, from the Hari River basin in Afghanistan and Iran (Actinopterygii: Cyprinidae)" masterLastPageNumber="55" masterPageNumber="43" pageNumber="44" updateTime="1698314252039" updateUser="plazi">
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<mods:titleInfo id="4DF2E29C8953B77FD7CA8B00BB87F0E9">
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<mods:title id="BC6760F5D8848AF11F23C4C0F2FE24C7">A new species of riffle minnow, Alburnoides holciki, from the Hari River basin in Afghanistan and Iran (Actinopterygii: Cyprinidae)</mods:title>
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</mods:titleInfo>
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<mods:name id="CF50A14C9305FC8BE8D1A54CE476E3BD" type="personal">
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<mods:roleTerm id="B7153A4CAAFFF166B20B8B5EDA74B7D2">Author</mods:roleTerm>
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<mods:namePart id="8FC97D7B106257466773772F836BCC02">Coad, Brian W.</mods:namePart>
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</mods:name>
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<mods:name id="425099B35F0592E515D1C7B7B1083257" type="personal">
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<mods:roleTerm id="4DD4762DE5EDDFA234D234C35BADC353">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="44507BEB6A0EC58CC37556601DE1696A">Bogutskaya, Nina G.</mods:namePart>
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</mods:name>
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<mods:typeOfResource id="80967D611FCFFADC6F6D00FDBDCA5D62">text</mods:typeOfResource>
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<mods:titleInfo id="06DBA295452B053BB11E358AE4DCFD9B">
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<mods:title id="BCA10A1C22A8163AB8DA59B2BA0558BF">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="BB41A8A0EA02EC75B2FF686361D43499">
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<mods:date id="CC4E16F9CDF80817246C9083AAE882FF">2012</mods:date>
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<mods:detail id="DA93401939E4B2DC226282FCF6A741A4" type="volume">
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<mods:number id="DA9C0606F55301D61CA90C96CE02F785">3453</mods:number>
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<mods:start id="C82253AFD89CD3CFF3F3EE9F7F263657">43</mods:start>
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<mods:end id="62FB5567DDD089FA8FFC7646ABB0AE72">55</mods:end>
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<mods:classification id="3E7537D180E1BE6269FD97C524CC5E86">journal article</mods:classification>
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<mods:identifier id="1ECDB8915C94ECB3C637D27ADDEFFCB6" type="DOI">10.5281/zenodo.214789</mods:identifier>
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<mods:identifier id="3A46E3B89B1FD108070AFEFFF1BD9AB7" type="GBIF-Dataset">4c5af5bf-c291-4031-86c9-f10683309f4c</mods:identifier>
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<mods:identifier id="6E83D211B29D38F0B3B04A66C4AAC35B" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="9AD27CE7CA795DB7FDBC4B6DA16E0452" type="Zenodo-Dep">214789</mods:identifier>
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</mods:mods>
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<treatment id="03A11D7EFFCCFFBDFF1AFE40E1B3E75F" ID-DOI="http://doi.org/10.5281/zenodo.5681569" ID-GBIF-Taxon="119524478" ID-Zenodo-Dep="5681569" LSID="urn:lsid:plazi:treatment:03A11D7EFFCCFFBDFF1AFE40E1B3E75F" httpUri="http://treatment.plazi.org/id/03A11D7EFFCCFFBDFF1AFE40E1B3E75F" lastPageId="11" lastPageNumber="54" pageId="1" pageNumber="44">
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<subSubSection id="C312FFE3FFCCFFB7FF1AFE40E135E4AD" pageId="1" pageNumber="44" type="nomenclature">
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<paragraph id="8BB7AC68FFCCFFB7FF1AFE40E203E4CE" blockId="1.[151,548,403,462]" box="[151,548,403,430]" pageId="1" pageNumber="44">
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<heading id="D0FF1B04FFCCFFB7FF1AFE40E203E4CE" bold="true" box="[151,548,403,430]" fontSize="11" level="1" pageId="1" pageNumber="44" reason="1">
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<emphasis id="B97C707AFFCCFFB7FF1AFE40E203E4CE" bold="true" box="[151,548,403,430]" pageId="1" pageNumber="44">
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<taxonomicName id="4C08D7EBFFCCFFB7FF1AFE40E1A2E4CD" ID-CoL="BGV6" box="[151,389,404,430]" class="Actinopterygii" family="Cyprinidae" genus="Alburnoides" kingdom="Animalia" order="Cypriniformes" pageId="1" pageNumber="44" phylum="Chordata" rank="species" species="holciki" status="sp. nov.">
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<emphasis id="B97C707AFFCCFFB7FF1AFE40E1A2E4CD" bold="true" box="[151,389,404,430]" italics="true" pageId="1" pageNumber="44">Alburnoides holciki</emphasis>
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</taxonomicName>
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,
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<taxonomicNameLabel id="A24FCD01FFCCFFB7FE19FE47E203E4CE" box="[404,548,403,429]" pageId="1" pageNumber="44" rank="species">new species</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8BB7AC68FFCCFFB7FF1AFE62E135E4AD" blockId="1.[151,548,403,462]" box="[151,274,438,462]" pageId="1" pageNumber="44">
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(
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<figureCitation id="1333B0EDFFCCFFB7FF12FE62E0CBE4AD" box="[159,236,438,462]" captionStart="FIGURE 1" captionStartId="1.[151,250,1404,1427]" captionTargetBox="[151,1435,1058,1380]" captionTargetId="figure@1.[151,1436,1058,1381]" captionTargetPageId="1" captionText="FIGURE 1. Alburnoides holciki sp. n., female holotype, 80.0 mm SL, Hari River at Herat (Afghanistan)." httpUri="https://zenodo.org/record/214790/files/figure.png" pageId="1" pageNumber="44">Figs. 1</figureCitation>
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,
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<figureCitation id="1333B0EDFFCCFFB7FF77FE62E12EE4AD" box="[250,265,438,462]" captionStart="FIGURE 2" captionStartId="1.[151,250,1939,1962]" captionTargetBox="[168,1435,1507,1920]" captionTargetId="figure@1.[168,1453,1507,1920]" captionTargetPageId="1" captionText="FIGURE 2. Alburnoides holciki sp. n., radiograph of a non-type specimen, BMNH 1914.1.1: 30 - 31, 82.4 mm SL, Kashaf River at Mashhad (Iran). Shows an upturned mouth, a straightened upper profile of the snout, 13 predorsal vertebrae, 20 abdominal vertebrae, and 21 caudal vertebrae." httpUri="https://zenodo.org/record/214791/files/figure.png" pageId="1" pageNumber="44">2</figureCitation>
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)
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||
</paragraph>
|
||
</subSubSection>
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||
<subSubSection id="C312FFE3FFCCFFB7FF1AFE2BE17FE78D" pageId="1" pageNumber="44" type="materials_examined">
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<paragraph id="8BB7AC68FFCCFFB7FF1AFE2BE324E758" blockId="1.[151,1436,510,1002]" pageId="1" pageNumber="44">
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<typeStatus id="54B312CAFFCCFFB7FF1AFE2BE123E774" box="[151,260,511,535]" pageId="1" pageNumber="44" type="holotype">
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||
<emphasis id="B97C707AFFCCFFB7FF1AFE2BE123E774" bold="true" box="[151,260,511,535]" pageId="1" pageNumber="44">Holotype</emphasis>
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||
</typeStatus>
|
||
(
|
||
<figureCitation id="1333B0EDFFCCFFB7FE9EFE2AE173E775" box="[275,340,510,534]" captionStart="FIGURE 1" captionStartId="1.[151,250,1404,1427]" captionTargetBox="[151,1435,1058,1380]" captionTargetId="figure@1.[151,1436,1058,1381]" captionTargetPageId="1" captionText="FIGURE 1. Alburnoides holciki sp. n., female holotype, 80.0 mm SL, Hari River at Herat (Afghanistan)." httpUri="https://zenodo.org/record/214790/files/figure.png" pageId="1" pageNumber="44">Fig. 1</figureCitation>
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).
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||
<collectionCode id="ED1934ADFFCCFFB7FEE6FE2AE180E775" box="[363,423,510,534]" httpUri="http://grbio.org/cool/n1s9-2rrz" name="Slovak National Museum" pageId="1" pageNumber="44">SNM</collectionCode>
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6788, female,
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||
<quantity id="4CF0018DFFCCFFB7FDC0FE2AE299E775" box="[589,702,510,534]" metricMagnitude="-1" metricUnit="m" metricValue="1.018" pageId="1" pageNumber="44" unit="mm" value="101.8">101.8 mm</quantity>
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TL, 80.0 mm SL,
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<collectingCountry id="F31FECF8FFCCFFB7FC01FE2AE473E775" box="[908,1108,510,534]" name="Afghanistan" pageId="1" pageNumber="44">AFGHANISTAN</collectingCountry>
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||
, Hari River at Herat (Herat is at
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<geoCoordinate id="EE3CCAAFFFCCFFB7FF3FFDF6E132E759" box="[178,277,546,570]" direction="north" orientation="latitude" pageId="1" pageNumber="44" precision="925" value="34.333332">34°20’N</geoCoordinate>
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||
,
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||
<geoCoordinate id="EE3CCAAFFFCCFFB7FEACFDF6E1A6E759" box="[289,385,546,570]" direction="east" orientation="longitude" pageId="1" pageNumber="44" precision="925" value="62.2">62°12’E</geoCoordinate>
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),
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<date id="FFB68AA8FFCCFFB7FE1BFDF6E219E759" box="[406,574,546,571]" pageId="1" pageNumber="44" value="1974-08-09">9 August 1974</date>
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, coll. L. Jedlička.
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||
</paragraph>
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||
<paragraph id="8BB7AC68FFCCFFB7FF44FD93E1D1E7E1" blockId="1.[151,1436,510,1002]" pageId="1" pageNumber="44">
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||
<typeStatus id="54B312CAFFCCFFB7FF44FD93E166E73C" box="[201,321,583,607]" pageId="1" pageNumber="44" type="paratype">
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||
<emphasis id="B97C707AFFCCFFB7FF44FD93E166E73C" bold="true" box="[201,321,583,607]" pageId="1" pageNumber="44">Paratypes</emphasis>
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||
</typeStatus>
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||
.
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<collectionCode id="ED1934ADFFCCFFB7FEC3FD92E1ADE73D" box="[334,394,582,606]" httpUri="http://grbio.org/cool/n1s9-2rrz" name="Slovak National Museum" pageId="1" pageNumber="44">SNM</collectionCode>
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||
6788,
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||
<specimenCount id="9D0E67E1FFCCFFB7FE5BFD92E256E73D" box="[470,625,582,606]" pageId="1" pageNumber="44" type="generic">18 specimens</specimenCount>
|
||
,
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||
<quantity id="4CF0018DFFCCFFB7FDF6FD92E33AE73D" box="[635,797,582,606]" metricMagnitude="-2" metricUnit="m" metricValue="7.109999999999999" metricValueMax="9.26" metricValueMin="4.96" pageId="1" pageNumber="44" unit="mm" value="71.1" valueMax="92.6" valueMin="49.6">49.6–92.6 mm</quantity>
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SL,
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||
<specimenCount id="9D0E67E1FFCCFFB7FCDCFD92E381E73D" box="[849,934,582,606]" pageId="1" pageNumber="44" type="male">8 males</specimenCount>
|
||
<quantity id="4CF0018DFFCCFFB7FC20FD92E477E73D" box="[941,1104,582,606]" metricMagnitude="-2" metricUnit="m" metricValue="5.99" metricValueMax="7.02" metricValueMin="4.96" pageId="1" pageNumber="44" unit="mm" value="59.9" valueMax="70.2" valueMin="49.6">49.6–70.2 mm</quantity>
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||
SL and
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||
<specimenCount id="9D0E67E1FFCCFFB7FB26FD92E502E73D" box="[1195,1317,582,606]" pageId="1" pageNumber="44" type="female">10 females</specimenCount>
|
||
<quantity id="4CF0018DFFCCFFB7FAA1FD92E0E4E7E1" metricMagnitude="-2" metricUnit="m" metricValue="7.45" metricValueMax="9.26" metricValueMin="5.64" pageId="1" pageNumber="44" unit="mm" value="74.5" valueMax="92.6" valueMin="56.4">56.4–92.6 mm</quantity>
|
||
SL, same data as
|
||
<typeStatus id="54B312CAFFCCFFB7FE00FDBEE1D4E7E1" box="[397,499,618,642]" pageId="1" pageNumber="44" type="holotype">holotype</typeStatus>
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||
.
|
||
</paragraph>
|
||
<paragraph id="8BB7AC68FFCCFFB7FF44FD5BE17FE78D" blockId="1.[151,1436,510,1002]" pageId="1" pageNumber="44">
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||
<emphasis id="B97C707AFFCCFFB7FF44FD5BE180E7C4" bold="true" box="[201,423,654,679]" pageId="1" pageNumber="44">Non-type material</emphasis>
|
||
(
|
||
<figureCitation id="1333B0EDFFCCFFB7FE37FD5AE225E7C5" box="[442,514,654,678]" captionStart="FIGURE 2" captionStartId="1.[151,250,1939,1962]" captionTargetBox="[168,1435,1507,1920]" captionTargetId="figure@1.[168,1453,1507,1920]" captionTargetPageId="1" captionText="FIGURE 2. Alburnoides holciki sp. n., radiograph of a non-type specimen, BMNH 1914.1.1: 30 - 31, 82.4 mm SL, Kashaf River at Mashhad (Iran). Shows an upturned mouth, a straightened upper profile of the snout, 13 predorsal vertebrae, 20 abdominal vertebrae, and 21 caudal vertebrae." httpUri="https://zenodo.org/record/214791/files/figure.png" pageId="1" pageNumber="44">Fig. 2</figureCitation>
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||
).
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||
<collectionCode id="ED1934ADFFCCFFB7FD97FD5BE249E7C5" box="[538,622,655,678]" pageId="1" pageNumber="44">BMNH</collectionCode>
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||
1914.1.1:30-31, 2,
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||
<quantity id="4CF0018DFFCCFFB7FCD9FD5AE3DCE7C5" box="[852,1019,654,678]" metricMagnitude="-2" metricUnit="m" metricValue="8.755" metricValueMax="9.27" metricValueMin="8.24" pageId="1" pageNumber="44" unit="mm" value="87.55" valueMax="92.7" valueMin="82.4">82.4–92.7 mm</quantity>
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||
SL,
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||
<collectingCountry id="F31FECF8FFCCFFB7FBBAFD5BE458E7C4" box="[1079,1151,655,679]" name="Iran" pageId="1" pageNumber="44">IRAN</collectingCountry>
|
||
, Kashaf River, Mashhad (Mashhad is at
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||
<geoCoordinate id="EE3CCAAFFFCCFFB7FECAFD66E18DE7A9" box="[327,426,690,714]" direction="north" orientation="latitude" pageId="1" pageNumber="44" precision="925" value="36.3">36°18’N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE3CCAAFFFCCFFB7FE3AFD66E233E7A9" box="[439,532,690,714]" direction="east" orientation="longitude" pageId="1" pageNumber="44" precision="925" value="59.6">59°36’E</geoCoordinate>
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||
), coll. P. Sykes (material dried at some point so not measurable in comparison with
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<typeStatus id="54B312CAFFCCFFB7FF5DFD03E127E78C" box="[208,256,727,751]" pageId="1" pageNumber="44">type</typeStatus>
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||
series).
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C312FFE3FFCCFFB7FF44FD2EE2D6E689" pageId="1" pageNumber="44" type="diagnosis">
|
||
<paragraph id="8BB7AC68FFCCFFB7FF44FD2EE2D6E689" blockId="1.[151,1436,510,1002]" pageId="1" pageNumber="44">
|
||
<emphasis id="B97C707AFFCCFFB7FF44FD2EE164E670" bold="true" box="[201,323,762,787]" pageId="1" pageNumber="44">Diagnosis.</emphasis>
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The species is distinguished by a combination of characters which includes a well-defined, sharp, scaleless or only slightly scaled, ventral keel; a short, slightly pointed snout; a terminal mouth with the tip of the mouth cleft on a level with the upper half of the pupil; a large eye (orbit width about equal to interorbital width), (46)47–51(55) lateral-line scales to posterior margin of hypurals (47–57 total lateral-line scales); 2.5–4.2 pharyngeal teeth; usually 8½ branched dorsal-fin rays; usually 13–16½ branched anal-fin rays; 40–42, usually 41, total vertebrae; caudal vertebral region longer than abdominal region (most frequent vertebral formulae 20+21, 20+22 and 19+21); and usually 13 or 14 predorsal vertebrae.
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||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="DF77FCE0FFCCFFB7FF1AFAA8E4E3E0F0" box="[151,1220,1404,1427]" httpUri="https://zenodo.org/record/214790/files/figure.png" pageId="1" pageNumber="44" targetBox="[151,1435,1058,1380]" targetPageId="1">
|
||
<paragraph id="8BB7AC68FFCCFFB7FF1AFAA8E4E3E0F0" blockId="1.[151,1220,1404,1427]" box="[151,1220,1404,1427]" pageId="1" pageNumber="44">
|
||
<emphasis id="B97C707AFFCCFFB7FF1AFAA8E134E0F1" bold="true" box="[151,275,1404,1427]" pageId="1" pageNumber="44">FIGURE 1.</emphasis>
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||
<taxonomicName id="4C08D7EBFFCCFFB7FE94FAA9E1C7E0F0" box="[281,480,1405,1427]" class="Actinopterygii" family="Cyprinidae" genus="Alburnoides" kingdom="Animalia" order="Cypriniformes" pageId="1" pageNumber="44" phylum="Chordata" rank="species" species="holciki" status="sp. nov.">
|
||
<emphasis id="B97C707AFFCCFFB7FE94FAA9E1C7E0F0" box="[281,480,1405,1427]" italics="true" pageId="1" pageNumber="44">Alburnoides holciki</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A24FCD01FFCCFFB7FE6BFAA9E238E0F1" box="[486,543,1405,1426]" pageId="1" pageNumber="44" rank="species">
|
||
<emphasis id="B97C707AFFCCFFB7FE6BFAA9E238E0F1" bold="true" box="[486,543,1405,1426]" pageId="1" pageNumber="44">sp. n.</emphasis>
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||
</taxonomicNameLabel>
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||
, female holotype, 80.0 mm SL, Hari River at Herat (Afghanistan).
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF77FCE0FFCCFFB7FF1AF847E1DDE284" httpUri="https://zenodo.org/record/214791/files/figure.png" pageId="1" pageNumber="44" targetBox="[168,1435,1507,1920]" targetPageId="1">
|
||
<paragraph id="8BB7AC68FFCCFFB7FF1AF847E1DDE284" blockId="1.[151,1436,1939,2023]" pageId="1" pageNumber="44">
|
||
<emphasis id="B97C707AFFCCFFB7FF1AF847E135E2CA" bold="true" box="[151,274,1939,1962]" pageId="1" pageNumber="44">FIGURE 2.</emphasis>
|
||
<taxonomicName id="4C08D7EBFFCCFFB7FE95F840E1F8E2C9" box="[280,479,1940,1962]" class="Actinopterygii" family="Cyprinidae" genus="Alburnoides" kingdom="Animalia" order="Cypriniformes" pageId="1" pageNumber="44" phylum="Chordata" rank="species" species="holciki" status="sp. nov.">
|
||
<emphasis id="B97C707AFFCCFFB7FE95F840E1F8E2C9" box="[280,479,1940,1962]" italics="true" pageId="1" pageNumber="44">Alburnoides holciki</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A24FCD01FFCCFFB7FE69F840E23BE2CA" box="[484,540,1940,1961]" pageId="1" pageNumber="44" rank="species">
|
||
<emphasis id="B97C707AFFCCFFB7FE69F840E23BE2CA" bold="true" box="[484,540,1940,1961]" pageId="1" pageNumber="44">sp. n.</emphasis>
|
||
</taxonomicNameLabel>
|
||
, radiograph of a non-type specimen, BMNH 1914.1.1:30-31, 82.4 mm SL, Kashaf River at Mashhad (Iran). Shows an upturned mouth, a straightened upper profile of the snout, 13 predorsal vertebrae, 20 abdominal vertebrae, and 21 caudal vertebrae.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C312FFE3FFCFFFB2FF4AFF43E2D3E68F" lastPageId="4" lastPageNumber="47" pageId="2" pageNumber="45" type="description">
|
||
<paragraph id="8BB7AC68FFCFFFB4FF4AFF43E1C0E407" blockId="2.[151,1436,151,1984]" pageId="2" pageNumber="45">
|
||
<emphasis id="B97C707AFFCFFFB4FF4AFF43E17EE5D3" bold="true" box="[199,345,151,176]" pageId="2" pageNumber="45">Description.</emphasis>
|
||
<typeStatus id="54B312CAFFCFFFB4FEE9FF43E1F7E5CC" box="[356,464,151,175]" pageId="2" pageNumber="45" type="holotype">Holotype</typeStatus>
|
||
. A ventral keel between the pelvics and the anal-fin is well-developed, sharp and scaleless. There is a pelvic axillary scale and scales extend over the proximal bases of the anal-fin. The lateral line is decurved and only the last few scales are elevated and on the mid-caudal peduncle. Lateral-line scales to posterior margin of hypurals number 47 (49 total), scales above lateral line to dorsal-fin origin are 11, scales below lateral line to anal-fin origin are 4, scales below lateral line to pelvic-fin origin are 5, predorsal scales are 23, and circumpeduncle scales are 17.
|
||
</paragraph>
|
||
<paragraph id="8BB7AC68FFCFFFB4FF44FEBBE1B7E706" blockId="2.[151,1436,151,1984]" pageId="2" pageNumber="45">Dorsal-fin rays are 3 unbranched and 8½ branched, anal-fin rays are 3 unbranched and 13½ branched, branched pectoral-fin rays are 13, pelvic-fin branched rays are 7. The anal-fin origin is under the 7th branched ray of the dorsal-fin. There are 6 gill rakers on the entire first gill arch and the longest raker reaches the raker below when appressed. Pharyngeal teeth are –4.2 on the right 5th ceratobranchial. Total vertebrae are 41 (including 4 Weberian vertebrae and last complex centrum), abdominal vertebrae (including intermediate ones; precaudal vertebrae auctorum) are 20, caudal vertebrae are 21, and predorsal abdominal vertebrae (anterior to the first dorsal-fin pterygiophore) are 14.</paragraph>
|
||
<paragraph id="8BB7AC68FFCFFFB4FF44FDA5E35FE796" blockId="2.[151,1436,151,1984]" pageId="2" pageNumber="45">
|
||
General topography of cephalic sensory canals and numbers of pores are typical of most
|
||
<taxonomicName id="4C08D7EBFFCFFFB4FB64FDA5E555E7EA" box="[1257,1394,625,649]" class="Actinopterygii" family="Cyprinidae" genus="Alburnoides" kingdom="Animalia" order="Cypriniformes" pageId="2" pageNumber="45" phylum="Chordata" rank="genus">
|
||
<emphasis id="B97C707AFFCFFFB4FB64FDA5E555E7EA" box="[1257,1394,625,649]" italics="true" pageId="2" pageNumber="45">Alburnoides</emphasis>
|
||
</taxonomicName>
|
||
, as described by
|
||
<bibRefCitation id="EF99D199FFCFFFB4FEA1FD41E221E7CE" author="Bogutskaya" box="[300,518,661,685]" pageId="2" pageNumber="45" refString="Bogutskaya, N. G. (1988) Canal topography of the seismosensory system of cyprinids of the subfamilies Leuciscinae, Xenocyprininae and Cultrinae. Voprosy Ikhtiologii, 28, 367 - 382 (in Russian; translated in Journal of Ichthyology, 28 (4), 91 - 107)." type="journal article" year="1988">Bogutskaya (1988)</bibRefCitation>
|
||
. The supraorbital canal is not lengthened in its posterior section and has 10 (left) / 9 (right) pores, the infraorbital canal has 16/13 pores; the preopercular-mandibular canal is complete, with 15/15 pores, and the supratemporal canal is also complete, with 5 pores.
|
||
</paragraph>
|
||
<paragraph id="8BB7AC68FFCFFFB4FF44FCD5E3D9E6E6" blockId="2.[151,1436,151,1984]" pageId="2" pageNumber="45">
|
||
The upper body profile is rounded while the lower profile is slightly more convex. The snout is short, slightly pointed, not stout. The mouth is upturned and terminal, the mouth cleft is straight, and its tip is on a level with the upper half of the pupil. Neither lower jaw nor upper jaw protrudes relative to each other. The caudal-fin lobes are pointed, the fin is clearly forked. Morphometrics are summarised in
|
||
<tableCitation id="C68A99D3FFCFFFB4FC29FCB9E3DFE6E6" box="[932,1016,877,901]" captionStart="TABLE 1" captionStartId="3.[151,235,152,173]" captionTargetBox="[151,1436,262,1732]" captionTargetPageId="3" captionText="TABLE 1. Morphometric characters for Alburnoides holciki. Significant differences (p <0.05) between males and females marked by an asterisk (*); for preanal length (marked by #) means and ranges are given to two decimal places as rounding gives identical means." httpUri="http://table.plazi.org/id/DF77FCE0FFCEFFB5FF1AFF4CE11BE588" pageId="2" pageNumber="45" tableUuid="DF77FCE0FFCEFFB5FF1AFF4CE11BE588">Table 1</tableCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BB7AC68FFCFFFB4FF44FC45E28CE1AA" blockId="2.[151,1436,151,1984]" pageId="2" pageNumber="45">
|
||
The
|
||
<typeStatus id="54B312CAFFCFFFB4FF71FC45E179E6CA" box="[252,350,913,937]" pageId="2" pageNumber="45" type="holotype">holotype</typeStatus>
|
||
in preservative is decoloured and discernible pigment is a light brown. Pigmentation consists of a darker back fading to a cream belly, pre- and post-dorsal-fin stripes on the back, small spots on the upper operculum and adjacently behind the eye that look like the remains of a dark lateral stripe, three rows of spots above the lateral line anteriorly before the pelvic-fin level, a dark line delimiting the hypaxial and epaxial muscle masses overlain by a diffuse stripe from the head to the tail, expanded on the tail base, a lateral line demarcated by pigment above and below it (the typical "stitched" pattern in many
|
||
<taxonomicName id="4C08D7EBFFCFFFB4FCA7FB92E394E13D" box="[810,947,1094,1118]" class="Actinopterygii" family="Cyprinidae" genus="Alburnoides" kingdom="Animalia" order="Cypriniformes" pageId="2" pageNumber="45" phylum="Chordata" rank="genus">
|
||
<emphasis id="B97C707AFFCFFFB4FCA7FB92E394E13D" box="[810,947,1094,1118]" italics="true" pageId="2" pageNumber="45">Alburnoides</emphasis>
|
||
</taxonomicName>
|
||
species), fins are mostly hyaline, faint pigment along the anterior dorsal-fin rays, a line of pigment along the proximal edge of the unbranched pectoral-fin ray and weak pigmentation on the first branched ray, and faint pigmentation on the caudal-fin centre branching distally to follow the inner margins of the fin fork.
|
||
</paragraph>
|
||
<paragraph id="8BB7AC68FFCFFFB4FF44FB02E29CE056" blockId="2.[151,1436,151,1984]" pageId="2" pageNumber="45">
|
||
<typeStatus id="54B312CAFFCFFFB4FF44FB02E11DE18D" box="[201,314,1238,1262]" pageId="2" pageNumber="45" type="paratype">Paratypes</typeStatus>
|
||
. The ventral keel between the pelvics and anal-fin is a well-developed, sharp and protruding scaleless keel in 15 fish, with one scale behind the pelvic-fins (scaled along 1/4 of its length) in two fish, and with three scales (scaled along 1/3 of its length) in one fish.
|
||
</paragraph>
|
||
<paragraph id="8BB7AC68FFCFFFB4FF44FA96E26FE08A" blockId="2.[151,1436,151,1984]" pageId="2" pageNumber="45">A pelvic axillary scale is present and the anal-fin base is proximally overlain by flank scales. Scales below the dorsal-fin on the upper flank are a vertical oval with rounded posterior, dorsal and ventral margins, and a wavy anterior margin. Scale radii are restricted to the posterior field encroaching laterally, circuli are fine and eccentric and the focus is anteriorly located. The lateral line is complete; it is decurved and only the last few scales are elevated and on the mid-caudal peduncle.</paragraph>
|
||
<paragraph id="8BB7AC68FFCFFFB4FF44FA21E3F3E386" blockId="2.[151,1436,151,1984]" pageId="2" pageNumber="45">
|
||
Meristic characters showed no significant differences (p>0.5) between males and females and the data were combined with the exception of the predorsal scale count. Lateral-line scales to posterior margin of hypurals 46(1), 47(4), 48(6), 49(2), 50(1), 51(3) or 55(1) (48.8, 2.16); total lateral-line scales 47–53 (
|
||
<quantity id="4CF0018DFFCFFFB4FBE4F9E9E482E336" box="[1129,1189,1597,1621]" metricMagnitude="0" metricUnit="m" metricValue="1.4478" pageId="2" pageNumber="45" unit="in" value="57.0">57 in</quantity>
|
||
one
|
||
<typeStatus id="54B312CAFFCFFFB4FB52F9E9E563E336" box="[1247,1348,1597,1621]" pageId="2" pageNumber="45" type="paratype">paratype</typeStatus>
|
||
); scales above lateral line to dorsal-fin origin 9(2), 10(14) or 11(2) (10.0, 0.49), scales below lateral line to pelvic-fin origin 4(3), 5(14) or 6(1) (4.9, 0.47), scales below lateral line to anal-fin origin 4(7) or 5(11) (4.6, 0.50), predorsal scales 20(1), 21(5) and 22(2) for males (21.1,0.64) and 20(1), 21(1), 22(2), 23(3), 24(1), 25(1) or 26(1) for females (22.9, 1.79 (p = 0.02), and circumpeduncular scales 16(2) or 17(16) (16.9, 0.32).
|
||
</paragraph>
|
||
<paragraph id="8BB7AC68FFCFFFB4FF44F925E12CE2A3" blockId="2.[151,1436,151,1984]" pageId="2" pageNumber="45">
|
||
Dorsal-fin unbranched rays 3 (in 12
|
||
<typeStatus id="54B312CAFFCFFFB4FDD2F925E2EBE26A" box="[607,716,1777,1801]" pageId="2" pageNumber="45" type="paratype">paratypes</typeStatus>
|
||
) or 4 (in 6
|
||
<typeStatus id="54B312CAFFCFFFB4FCC5F925E39FE26A" box="[840,952,1777,1801]" pageId="2" pageNumber="45" type="paratype">paratypes</typeStatus>
|
||
), dorsal-fin branched rays 8½(15) or 9½(3) (mean 8.2, standard deviation 0.38), anal-fin unbranched rays 3, anal-fin branched rays 10½(1), 14½(6), 15½(8) or 16½(3) (14.6, 1.34) (14.8 and 0.73 if the abnormal number, 10½, is excluded), branched pectoral-fin rays 13(8), 14(9) or 15(1) (13.6, 0.61), and pelvic-fin branched rays 7(18). The dorsal-fin outer margin is truncate to slightly rounded and the anal-fin outer margin is slightly concave. The anal-fin origin is under the 5–7th branched ray of the dorsal-fin.
|
||
</paragraph>
|
||
<caption id="DF77FCE0FFCEFFB5FF1AFF4CE11BE588" ID-Table-UUID="DF77FCE0FFCEFFB5FF1AFF4CE11BE588" httpUri="http://table.plazi.org/id/DF77FCE0FFCEFFB5FF1AFF4CE11BE588" pageId="3" pageNumber="46" targetBox="[151,1436,262,1732]" targetIsTable="true" targetPageId="3">
|
||
<paragraph id="8BB7AC68FFCEFFB5FF1AFF4CE11BE588" blockId="3.[151,1436,152,235]" pageId="3" pageNumber="46">
|
||
<emphasis id="B97C707AFFCEFFB5FF1AFF4CE12FE5CE" bold="true" box="[151,264,152,173]" pageId="3" pageNumber="46">TABLE 1.</emphasis>
|
||
Morphometric characters for
|
||
<taxonomicName id="4C08D7EBFFCEFFB5FDC5FF4CE334E5CD" box="[584,787,152,174]" class="Actinopterygii" family="Cyprinidae" genus="Alburnoides" kingdom="Animalia" order="Cypriniformes" pageId="3" pageNumber="46" phylum="Chordata" rank="species" species="holciki">
|
||
<emphasis id="B97C707AFFCEFFB5FDC5FF4CE334E5CD" box="[584,787,152,174]" italics="true" pageId="3" pageNumber="46">Alburnoides holciki</emphasis>
|
||
</taxonomicName>
|
||
. Significant differences (p<0.05) between males and females marked by an asterisk (*); for preanal length (marked by #) means and ranges are given to two decimal places as rounding gives identical means.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BB7AC68FFCEFFB5FF12FED2E560E3A3" pageId="3" pageNumber="46">
|
||
<table id="F9085EC8FFCE0049FF1AFED2E5BBE3A7" box="[151,1436,262,1732]" gridcols="5" gridrows="24" pageId="3" pageNumber="46">
|
||
<tr id="3538AE2AFFCE0049FF1AFED2E5BBE458" box="[151,1436,262,315]" gridrow="0" pageId="3" pageNumber="46">
|
||
<th id="76E9C756FFCE0049FF1AFED2E1F8E458" box="[151,479,262,315]" gridcol="0" gridrow="0" pageId="3" pageNumber="46">Character</th>
|
||
<th id="76E9C756FFCE0049FDA2FED2E2AAE458" box="[559,653,262,315]" gridcol="1" gridrow="0" pageId="3" pageNumber="46">Holotype</th>
|
||
<th id="76E9C756FFCE0049FD4FFED2E383E458" box="[706,932,262,315]" gridcol="2" gridrow="0" pageId="3" pageNumber="46">Male paratypes range (mean, S.D.)</th>
|
||
<th id="76E9C756FFCE0049FC6DFED2E4F7E458" box="[992,1232,262,315]" gridcol="3" gridrow="0" pageId="3" pageNumber="46">Female paratypes range (mean, S.D.)</th>
|
||
<th id="76E9C756FFCE0049FA96FED2E5BBE458" box="[1307,1436,262,315]" gridcol="4" gridrow="0" pageId="3" pageNumber="46">p</th>
|
||
</tr>
|
||
<tr id="3538AE2AFFCE0049FF1AFE9AE5BBE406" box="[151,1436,334,357]" gridrow="1" pageId="3" pageNumber="46" rowspan-4="1">
|
||
<th id="76E9C756FFCE0049FF1AFE9AE1F8E406" box="[151,479,334,357]" gridcol="0" gridrow="1" pageId="3" pageNumber="46">Size (mm SL)</th>
|
||
<td id="76E9C756FFCE0049FDA2FE9AE2AAE406" box="[559,653,334,357]" gridcol="1" gridrow="1" pageId="3" pageNumber="46">80.0</td>
|
||
<td id="76E9C756FFCE0049FD4FFE9AE383E406" box="[706,932,334,357]" gridcol="2" gridrow="1" pageId="3" pageNumber="46">49.6–70.2</td>
|
||
<td id="76E9C756FFCE0049FC6DFE9AE4F7E406" box="[992,1232,334,357]" gridcol="3" gridrow="1" pageId="3" pageNumber="46">56.4–92.6</td>
|
||
</tr>
|
||
<tr id="3538AE2AFFCE0049FF1AFEACE5BBE4ED" box="[151,1436,376,398]" gridrow="2" pageId="3" pageNumber="46">
|
||
<th id="76E9C756FFCE0049FF1AFEACE1F8E4ED" box="[151,479,376,398]" gridcol="0" gridrow="2" pageId="3" pageNumber="46">SL/Head length (HL)</th>
|
||
<td id="76E9C756FFCE0049FDA2FEACE2AAE4ED" box="[559,653,376,398]" gridcol="1" gridrow="2" pageId="3" pageNumber="46">3.7</td>
|
||
<td id="76E9C756FFCE0049FD4FFEACE383E4ED" box="[706,932,376,398]" gridcol="2" gridrow="2" pageId="3" pageNumber="46">3.8–4.0 (3.9, 0.06)</td>
|
||
<td id="76E9C756FFCE0049FC6DFEACE4F7E4ED" box="[992,1232,376,398]" gridcol="3" gridrow="2" pageId="3" pageNumber="46">3.6–4.1 (3.9, 0.14)</td>
|
||
<td id="76E9C756FFCE0049FA96FEACE5BBE4ED" box="[1307,1436,376,398]" gridcol="4" gridrow="2" pageId="3" pageNumber="46">0.90</td>
|
||
</tr>
|
||
<tr id="3538AE2AFFCE0049FF1AFE75E5BBE769" box="[151,1436,417,522]" gridrow="3" pageId="3" pageNumber="46">
|
||
<th id="76E9C756FFCE0049FF1AFE75E1F8E769" box="[151,479,417,522]" gridcol="0" gridrow="3" pageId="3" pageNumber="46">SL/Head depth SL/Head width SL/Body depth</th>
|
||
<td id="76E9C756FFCE0049FDA2FE75E2AAE769" box="[559,653,417,522]" gridcol="1" gridrow="3" pageId="3" pageNumber="46">5.2 7.1 3.5</td>
|
||
<td id="76E9C756FFCE0049FD4FFE75E383E769" box="[706,932,417,522]" gridcol="2" gridrow="3" pageId="3" pageNumber="46">4.8–5.1 (4.9, 0.12) 7.0–7.4 (7.2, 0.14) 3.2–3.7 (3.5, 0.16)</td>
|
||
<td id="76E9C756FFCE0049FC6DFE75E4F7E769" box="[992,1232,417,522]" gridcol="3" gridrow="3" pageId="3" pageNumber="46">4.7–5.1 (4.9, 0.14) 6.8–7.8 (7.2, 0.29) 3.5–37 (3.6, 0.08)</td>
|
||
<td id="76E9C756FFCE0049FA96FE75E5BBE769" box="[1307,1436,417,522]" gridcol="4" gridrow="3" pageId="3" pageNumber="46">0.83 0.94 0.60</td>
|
||
</tr>
|
||
<tr id="3538AE2AFFCE0049FF1AFDC9E5BBE7E8" box="[151,1436,541,651]" gridrow="4" pageId="3" pageNumber="46">
|
||
<th id="76E9C756FFCE0049FF1AFDC9E1F8E7E8" box="[151,479,541,651]" gridcol="0" gridrow="4" pageId="3" pageNumber="46">SL/Pectoral-fin length (P1) SL/Pelvic-fin length (P2) SL/Longest dorsal-fin ray</th>
|
||
<td id="76E9C756FFCE0049FDA2FDC9E2AAE7E8" box="[559,653,541,651]" gridcol="1" gridrow="4" pageId="3" pageNumber="46">4.6 5.6 4.1</td>
|
||
<td id="76E9C756FFCE0049FD4FFDC9E383E7E8" box="[706,932,541,651]" gridcol="2" gridrow="4" pageId="3" pageNumber="46">4.2–4.5 (4.4, 0.09) 5.1–6.4 (5.5, 0.42) 3.9–4.3 (4.1, 0.14)</td>
|
||
<td id="76E9C756FFCE0049FC6DFDC9E4F7E7E8" box="[992,1232,541,651]" gridcol="3" gridrow="4" pageId="3" pageNumber="46">4.2–4.8 (4.6, 0.22) 5.3–6.0 (5.6, 0.22) 4.1–4.6 (4.3, 0.16)</td>
|
||
<td id="76E9C756FFCE0049FA96FDC9E5BBE7E8" box="[1307,1436,541,651]" gridcol="4" gridrow="4" pageId="3" pageNumber="46">0.03* 0.52 0.02*</td>
|
||
</tr>
|
||
<tr id="3538AE2AFFCE0049FF1AFD4AE5BBE7BD" box="[151,1436,670,734]" gridrow="5" pageId="3" pageNumber="46">
|
||
<th id="76E9C756FFCE0049FF1AFD4AE1F8E7BD" box="[151,479,670,734]" gridcol="0" gridrow="5" pageId="3" pageNumber="46">SL/Longest anal-fin ray SL/Dorsal-fin base</th>
|
||
<td id="76E9C756FFCE0049FDA2FD4AE2AAE7BD" box="[559,653,670,734]" gridcol="1" gridrow="5" pageId="3" pageNumber="46">6.4 6.9</td>
|
||
<td id="76E9C756FFCE0049FD4FFD4AE383E7BD" box="[706,932,670,734]" gridcol="2" gridrow="5" pageId="3" pageNumber="46">5.7–6.4 (6.1, 0.25) 6.6–7.5 (7.1, 0.34)</td>
|
||
<td id="76E9C756FFCE0049FC6DFD4AE4F7E7BD" box="[992,1232,670,734]" gridcol="3" gridrow="5" pageId="3" pageNumber="46">5.7–6.8 (6.1, 0.40) 6.4–7.9 (7.2, 0.47)</td>
|
||
<td id="76E9C756FFCE0049FA96FD4AE5BBE7BD" box="[1307,1436,670,734]" gridcol="4" gridrow="5" pageId="3" pageNumber="46">0.79 0.61</td>
|
||
</tr>
|
||
<tr id="3538AE2AFFCE0049FF1AFD25E5BBE664" box="[151,1436,753,775]" gridrow="6" pageId="3" pageNumber="46">
|
||
<th id="76E9C756FFCE0049FF1AFD25E1F8E664" box="[151,479,753,775]" gridcol="0" gridrow="6" pageId="3" pageNumber="46">SL/Anal-fin base</th>
|
||
<td id="76E9C756FFCE0049FDA2FD25E2AAE664" box="[559,653,753,775]" gridcol="1" gridrow="6" pageId="3" pageNumber="46">5.6</td>
|
||
<td id="76E9C756FFCE0049FD4FFD25E383E664" box="[706,932,753,775]" gridcol="2" gridrow="6" pageId="3" pageNumber="46">4.6–6.2 (5.2, 0.47)</td>
|
||
<td id="76E9C756FFCE0049FC6DFD25E4F7E664" box="[992,1232,753,775]" gridcol="3" gridrow="6" pageId="3" pageNumber="46">4.7–5.6 (5.1, 0.32)</td>
|
||
<td id="76E9C756FFCE0049FA96FD25E5BBE664" box="[1307,1436,753,775]" gridcol="4" gridrow="6" pageId="3" pageNumber="46">0.41</td>
|
||
</tr>
|
||
<tr id="3538AE2AFFCE0049FF1AFCCEE5BBE639" box="[151,1436,794,858]" gridrow="7" pageId="3" pageNumber="46">
|
||
<th id="76E9C756FFCE0049FF1AFCCEE1F8E639" box="[151,479,794,858]" gridcol="0" gridrow="7" pageId="3" pageNumber="46">SL/Predorsal length SL/Prepelvic length</th>
|
||
<td id="76E9C756FFCE0049FDA2FCCEE2AAE639" box="[559,653,794,858]" gridcol="1" gridrow="7" pageId="3" pageNumber="46">1.8 2.0</td>
|
||
<td id="76E9C756FFCE0049FD4FFCCEE383E639" box="[706,932,794,858]" gridcol="2" gridrow="7" pageId="3" pageNumber="46">1.8–1.9 (1.9, 0.03) 2.1–2.2 (2.1, 0.04)</td>
|
||
<td id="76E9C756FFCE0049FC6DFCCEE4F7E639" box="[992,1232,794,858]" gridcol="3" gridrow="7" pageId="3" pageNumber="46">1.8–1.9 (1.9, 0.04) 1.8–2.2 (2.1, 0.11)</td>
|
||
<td id="76E9C756FFCE0049FA96FCCEE5BBE639" box="[1307,1436,794,858]" gridcol="4" gridrow="7" pageId="3" pageNumber="46">0.76 0.25</td>
|
||
</tr>
|
||
<tr id="3538AE2AFFCE0049FF1AFCB9E5BBE6E0" box="[151,1436,877,899]" gridrow="8" pageId="3" pageNumber="46">
|
||
<th id="76E9C756FFCE0049FF1AFCB9E1F8E6E0" box="[151,479,877,899]" gridcol="0" gridrow="8" pageId="3" pageNumber="46">SL/Preanal length#</th>
|
||
<td id="76E9C756FFCE0049FDA2FCB9E2AAE6E0" box="[559,653,877,899]" gridcol="1" gridrow="8" pageId="3" pageNumber="46">1.5</td>
|
||
<td id="76E9C756FFCE0049FD4FFCB9E383E6E0" box="[706,932,877,899]" gridcol="2" gridrow="8" pageId="3" pageNumber="46">1.55–1.65 (1.61, 0.03)</td>
|
||
<td id="76E9C756FFCE0049FC6DFCB9E4F7E6E0" box="[992,1232,877,899]" gridcol="3" gridrow="8" pageId="3" pageNumber="46">1.51–1.59 (1.56, 0.02)</td>
|
||
<td id="76E9C756FFCE0049FA96FCB9E5BBE6E0" box="[1307,1436,877,899]" gridcol="4" gridrow="8" pageId="3" pageNumber="46">0.001*</td>
|
||
</tr>
|
||
<tr id="3538AE2AFFCE0049FF1AFC42E5BBE6B5" box="[151,1436,918,982]" gridrow="9" pageId="3" pageNumber="46">
|
||
<th id="76E9C756FFCE0049FF1AFC42E1F8E6B5" box="[151,479,918,982]" gridcol="0" gridrow="9" pageId="3" pageNumber="46">SL/Caudal peduncle length SL/Caudal peduncle depth</th>
|
||
<td id="76E9C756FFCE0049FDA2FC42E2AAE6B5" box="[559,653,918,982]" gridcol="1" gridrow="9" pageId="3" pageNumber="46">5.2 9.4</td>
|
||
<td id="76E9C756FFCE0049FD4FFC42E383E6B5" box="[706,932,918,982]" gridcol="2" gridrow="9" pageId="3" pageNumber="46">4.3–5.2 (4.9, 0.28) 8.9–10.1 (9.5, 0.39)</td>
|
||
<td id="76E9C756FFCE0049FC6DFC42E4F7E6B5" box="[992,1232,918,982]" gridcol="3" gridrow="9" pageId="3" pageNumber="46">4.8–5.5 (5.1, 0.22) 9.1–10.6 (9.8, 0.57)</td>
|
||
<td id="76E9C756FFCE0049FA96FC42E5BBE6B5" box="[1307,1436,918,982]" gridcol="4" gridrow="9" pageId="3" pageNumber="46">0.10 0.19</td>
|
||
</tr>
|
||
<tr id="3538AE2AFFCE0049FF1AFC3DE5BBE69C" box="[151,1436,1001,1023]" gridrow="10" pageId="3" pageNumber="46">
|
||
<th id="76E9C756FFCE0049FF1AFC3DE1F8E69C" box="[151,479,1001,1023]" gridcol="0" gridrow="10" pageId="3" pageNumber="46">SL/Pectoral to pelvic-fin length</th>
|
||
<td id="76E9C756FFCE0049FDA2FC3DE2AAE69C" box="[559,653,1001,1023]" gridcol="1" gridrow="10" pageId="3" pageNumber="46">4.3</td>
|
||
<td id="76E9C756FFCE0049FD4FFC3DE383E69C" box="[706,932,1001,1023]" gridcol="2" gridrow="10" pageId="3" pageNumber="46">4.2–4.8 (4.6, 0.22)</td>
|
||
<td id="76E9C756FFCE0049FC6DFC3DE4F7E69C" box="[992,1232,1001,1023]" gridcol="3" gridrow="10" pageId="3" pageNumber="46">4.2–4.9 (4.5, 0.20)</td>
|
||
<td id="76E9C756FFCE0049FA96FC3DE5BBE69C" box="[1307,1436,1001,1023]" gridcol="4" gridrow="10" pageId="3" pageNumber="46">0.35</td>
|
||
</tr>
|
||
<tr id="3538AE2AFFCE0049FF1AFBC6E5BBE14B" box="[151,1436,1042,1064]" gridrow="11" pageId="3" pageNumber="46">
|
||
<th id="76E9C756FFCE0049FF1AFBC6E1F8E14B" box="[151,479,1042,1064]" gridcol="0" gridrow="11" pageId="3" pageNumber="46">SL/Pelvic to anal-fin length</th>
|
||
<td id="76E9C756FFCE0049FDA2FBC6E2AAE14B" box="[559,653,1042,1064]" gridcol="1" gridrow="11" pageId="3" pageNumber="46">5.3</td>
|
||
<td id="76E9C756FFCE0049FD4FFBC6E383E14B" box="[706,932,1042,1064]" gridcol="2" gridrow="11" pageId="3" pageNumber="46">5.5–6.3 (5.9, 0.25)</td>
|
||
<td id="76E9C756FFCE0049FC6DFBC6E4F7E14B" box="[992,1232,1042,1064]" gridcol="3" gridrow="11" pageId="3" pageNumber="46">5.3–6.2 (5.6, 0.25)</td>
|
||
<td id="76E9C756FFCE0049FA96FBC6E5BBE14B" box="[1307,1436,1042,1064]" gridcol="4" gridrow="11" pageId="3" pageNumber="46">0.02*</td>
|
||
</tr>
|
||
<tr id="3538AE2AFFCE0049FF1AFBEFE5BBE131" box="[151,1436,1083,1106]" gridrow="12" pageId="3" pageNumber="46">
|
||
<th id="76E9C756FFCE0049FF1AFBEFE1F8E131" box="[151,479,1083,1106]" gridcol="0" gridrow="12" pageId="3" pageNumber="46">HL/Mouth width</th>
|
||
<td id="76E9C756FFCE0049FDA2FBEFE2AAE131" box="[559,653,1083,1106]" gridcol="1" gridrow="12" pageId="3" pageNumber="46">4.6</td>
|
||
<td id="76E9C756FFCE0049FD4FFBEFE383E131" box="[706,932,1083,1106]" gridcol="2" gridrow="12" pageId="3" pageNumber="46">4.1–4.7 (4.5, 0.19)</td>
|
||
<td id="76E9C756FFCE0049FC6DFBEFE4F7E131" box="[992,1232,1083,1106]" gridcol="3" gridrow="12" pageId="3" pageNumber="46">3.6–4.9 (4.4, 0.48)</td>
|
||
<td id="76E9C756FFCE0049FA96FBEFE5BBE131" box="[1307,1436,1083,1106]" gridcol="4" gridrow="12" pageId="3" pageNumber="46">0.34</td>
|
||
</tr>
|
||
<tr id="3538AE2AFFCE0049FF1AFBB1E5BBE118" box="[151,1436,1125,1147]" gridrow="13" pageId="3" pageNumber="46">
|
||
<th id="76E9C756FFCE0049FF1AFBB1E1F8E118" box="[151,479,1125,1147]" gridcol="0" gridrow="13" pageId="3" pageNumber="46">HL/Head depth</th>
|
||
<td id="76E9C756FFCE0049FDA2FBB1E2AAE118" box="[559,653,1125,1147]" gridcol="1" gridrow="13" pageId="3" pageNumber="46">1.4</td>
|
||
<td id="76E9C756FFCE0049FD4FFBB1E383E118" box="[706,932,1125,1147]" gridcol="2" gridrow="13" pageId="3" pageNumber="46">1.2–1.3 (1.3, 0.04)</td>
|
||
<td id="76E9C756FFCE0049FC6DFBB1E4F7E118" box="[992,1232,1125,1147]" gridcol="3" gridrow="13" pageId="3" pageNumber="46">1.2–1.3 (1.3, 0.03)</td>
|
||
<td id="76E9C756FFCE0049FA96FBB1E5BBE118" box="[1307,1436,1125,1147]" gridcol="4" gridrow="13" pageId="3" pageNumber="46">0.95</td>
|
||
</tr>
|
||
<tr id="3538AE2AFFCE0049FF1AFB5AE5BBE1C7" box="[151,1436,1166,1188]" gridrow="14" pageId="3" pageNumber="46">
|
||
<th id="76E9C756FFCE0049FF1AFB5AE1F8E1C7" box="[151,479,1166,1188]" gridcol="0" gridrow="14" pageId="3" pageNumber="46">HL/Head width</th>
|
||
<td id="76E9C756FFCE0049FDA2FB5AE2AAE1C7" box="[559,653,1166,1188]" gridcol="1" gridrow="14" pageId="3" pageNumber="46">1.9</td>
|
||
<td id="76E9C756FFCE0049FD4FFB5AE383E1C7" box="[706,932,1166,1188]" gridcol="2" gridrow="14" pageId="3" pageNumber="46">1.8–1.9 (1.9, 0.05)</td>
|
||
<td id="76E9C756FFCE0049FC6DFB5AE4F7E1C7" box="[992,1232,1166,1188]" gridcol="3" gridrow="14" pageId="3" pageNumber="46">1.8–2.0 (1.9, 0.06)</td>
|
||
<td id="76E9C756FFCE0049FA96FB5AE5BBE1C7" box="[1307,1436,1166,1188]" gridcol="4" gridrow="14" pageId="3" pageNumber="46">0.83</td>
|
||
</tr>
|
||
<tr id="3538AE2AFFCE0049FF1AFB63E5BBE1AD" box="[151,1436,1207,1230]" gridrow="15" pageId="3" pageNumber="46">
|
||
<th id="76E9C756FFCE0049FF1AFB63E1F8E1AD" box="[151,479,1207,1230]" gridcol="0" gridrow="15" pageId="3" pageNumber="46">HL/Snout length</th>
|
||
<td id="76E9C756FFCE0049FDA2FB63E2AAE1AD" box="[559,653,1207,1230]" gridcol="1" gridrow="15" pageId="3" pageNumber="46">3.5</td>
|
||
<td id="76E9C756FFCE0049FD4FFB63E383E1AD" box="[706,932,1207,1230]" gridcol="2" gridrow="15" pageId="3" pageNumber="46">3.6–3.9 (3.7, 0.13)</td>
|
||
<td id="76E9C756FFCE0049FC6DFB63E4F7E1AD" box="[992,1232,1207,1230]" gridcol="3" gridrow="15" pageId="3" pageNumber="46">3.6–3.9 (3.7, 0.11)</td>
|
||
<td id="76E9C756FFCE0049FA96FB63E5BBE1AD" box="[1307,1436,1207,1230]" gridcol="4" gridrow="15" pageId="3" pageNumber="46">0.59</td>
|
||
</tr>
|
||
<tr id="3538AE2AFFCE0049FF1AFB35E5BBE194" box="[151,1436,1249,1271]" gridrow="16" pageId="3" pageNumber="46">
|
||
<th id="76E9C756FFCE0049FF1AFB35E1F8E194" box="[151,479,1249,1271]" gridcol="0" gridrow="16" pageId="3" pageNumber="46">HL/Orbit width</th>
|
||
<td id="76E9C756FFCE0049FDA2FB35E2AAE194" box="[559,653,1249,1271]" gridcol="1" gridrow="16" pageId="3" pageNumber="46">3.3</td>
|
||
<td id="76E9C756FFCE0049FD4FFB35E383E194" box="[706,932,1249,1271]" gridcol="2" gridrow="16" pageId="3" pageNumber="46">2.9–3.2 (3.0, 0.09)</td>
|
||
<td id="76E9C756FFCE0049FC6DFB35E4F7E194" box="[992,1232,1249,1271]" gridcol="3" gridrow="16" pageId="3" pageNumber="46">3.0–3.2 (3.1, 0.08)</td>
|
||
<td id="76E9C756FFCE0049FA96FB35E5BBE194" box="[1307,1436,1249,1271]" gridcol="4" gridrow="16" pageId="3" pageNumber="46">0.05</td>
|
||
</tr>
|
||
<tr id="3538AE2AFFCE0049FF1AFADEE5BBE029" box="[151,1436,1290,1354]" gridrow="17" pageId="3" pageNumber="46">
|
||
<th id="76E9C756FFCE0049FF1AFADEE1F8E029" box="[151,479,1290,1354]" gridcol="0" gridrow="17" pageId="3" pageNumber="46">HL/Interorbital width HL/Postorbital length</th>
|
||
<td id="76E9C756FFCE0049FDA2FADEE2AAE029" box="[559,653,1290,1354]" gridcol="1" gridrow="17" pageId="3" pageNumber="46">3.1 2.2</td>
|
||
<td id="76E9C756FFCE0049FD4FFADEE383E029" box="[706,932,1290,1354]" gridcol="2" gridrow="17" pageId="3" pageNumber="46">2.8–3.1 (3.0, 0.12) 2.2–2.3 (2.3, 0.03)</td>
|
||
<td id="76E9C756FFCE0049FC6DFADEE4F7E029" box="[992,1232,1290,1354]" gridcol="3" gridrow="17" pageId="3" pageNumber="46">3.0–3.2 (3.1, 0.06) 2.2–2.4 (2.3, 0.09)</td>
|
||
<td id="76E9C756FFCE0049FA96FADEE5BBE029" box="[1307,1436,1290,1354]" gridcol="4" gridrow="17" pageId="3" pageNumber="46">0.16 0.18</td>
|
||
</tr>
|
||
<tr id="3538AE2AFFCE0049FF1AFA89E5BBE010" box="[151,1436,1373,1395]" gridrow="18" pageId="3" pageNumber="46">
|
||
<th id="76E9C756FFCE0049FF1AFA89E1F8E010" box="[151,479,1373,1395]" gridcol="0" gridrow="18" pageId="3" pageNumber="46">HL/Longest dorsal-fin ray</th>
|
||
<td id="76E9C756FFCE0049FDA2FA89E2AAE010" box="[559,653,1373,1395]" gridcol="1" gridrow="18" pageId="3" pageNumber="46">1.1</td>
|
||
<td id="76E9C756FFCE0049FD4FFA89E383E010" box="[706,932,1373,1395]" gridcol="2" gridrow="18" pageId="3" pageNumber="46">1.0–1.1 (1.0, 0.03)</td>
|
||
<td id="76E9C756FFCE0049FC6DFA89E4F7E010" box="[992,1232,1373,1395]" gridcol="3" gridrow="18" pageId="3" pageNumber="46">1.1–1.2 (1.1, 0.04)</td>
|
||
<td id="76E9C756FFCE0049FA96FA89E5BBE010" box="[1307,1436,1373,1395]" gridcol="4" gridrow="18" pageId="3" pageNumber="46">0.01*</td>
|
||
</tr>
|
||
<tr id="3538AE2AFFCE0049FF1AFA52E5BBE0A5" box="[151,1436,1414,1478]" gridrow="19" pageId="3" pageNumber="46">
|
||
<th id="76E9C756FFCE0049FF1AFA52E1F8E0A5" box="[151,479,1414,1478]" gridcol="0" gridrow="19" pageId="3" pageNumber="46">HL/Longest anal-fin ray HL/Pectoral-fin length</th>
|
||
<td id="76E9C756FFCE0049FDA2FA52E2AAE0A5" box="[559,653,1414,1478]" gridcol="1" gridrow="19" pageId="3" pageNumber="46">1.7 1.2</td>
|
||
<td id="76E9C756FFCE0049FD4FFA52E383E0A5" box="[706,932,1414,1478]" gridcol="2" gridrow="19" pageId="3" pageNumber="46">1.5–1.6 (1.6, 0.06) 1.1–1.2 (1.1, 0.03)</td>
|
||
<td id="76E9C756FFCE0049FC6DFA52E4F7E0A5" box="[992,1232,1414,1478]" gridcol="3" gridrow="19" pageId="3" pageNumber="46">1.5–1.7 (1.6, 0.10) 1.1–1.2 (1.2, 0.03)</td>
|
||
<td id="76E9C756FFCE0049FA96FA52E5BBE0A5" box="[1307,1436,1414,1478]" gridcol="4" gridrow="19" pageId="3" pageNumber="46">0.71 0.001*</td>
|
||
</tr>
|
||
<tr id="3538AE2AFFCE0049FF1AFA0DE5BBE08C" box="[151,1436,1497,1519]" gridrow="20" pageId="3" pageNumber="46">
|
||
<th id="76E9C756FFCE0049FF1AFA0DE1F8E08C" box="[151,479,1497,1519]" gridcol="0" gridrow="20" pageId="3" pageNumber="46">HL/Pelvic-fin length</th>
|
||
<td id="76E9C756FFCE0049FDA2FA0DE2AAE08C" box="[559,653,1497,1519]" gridcol="1" gridrow="20" pageId="3" pageNumber="46">1.5</td>
|
||
<td id="76E9C756FFCE0049FD4FFA0DE383E08C" box="[706,932,1497,1519]" gridcol="2" gridrow="20" pageId="3" pageNumber="46">1.3–1.7 (1.4, 0.12)</td>
|
||
<td id="76E9C756FFCE0049FC6DFA0DE4F7E08C" box="[992,1232,1497,1519]" gridcol="3" gridrow="20" pageId="3" pageNumber="46">1.4–1.6 (1.5, 0.07)</td>
|
||
<td id="76E9C756FFCE0049FA96FA0DE5BBE08C" box="[1307,1436,1497,1519]" gridcol="4" gridrow="20" pageId="3" pageNumber="46">0.53</td>
|
||
</tr>
|
||
<tr id="3538AE2AFFCE0049FF1AF9D6E5BBE321" box="[151,1436,1538,1602]" gridrow="21" pageId="3" pageNumber="46">
|
||
<th id="76E9C756FFCE0049FF1AF9D6E1F8E321" box="[151,479,1538,1602]" gridcol="0" gridrow="21" pageId="3" pageNumber="46">HL/Dorsal-fin base HL/Anal-fin base</th>
|
||
<td id="76E9C756FFCE0049FDA2F9D6E2AAE321" box="[559,653,1538,1602]" gridcol="1" gridrow="21" pageId="3" pageNumber="46">1.8 1.5</td>
|
||
<td id="76E9C756FFCE0049FD4FF9D6E383E321" box="[706,932,1538,1602]" gridcol="2" gridrow="21" pageId="3" pageNumber="46">1.7–2.0 (1.8, 0.10) 1.2–1.6 (1.3, 0.13)</td>
|
||
<td id="76E9C756FFCE0049FC6DF9D6E4F7E321" box="[992,1232,1538,1602]" gridcol="3" gridrow="21" pageId="3" pageNumber="46">1.6–2.1 (1.8, 0.14) 1.2–1.5 (1.3, 0.10)</td>
|
||
<td id="76E9C756FFCE0049FA96F9D6E5BBE321" box="[1307,1436,1538,1602]" gridcol="4" gridrow="21" pageId="3" pageNumber="46">0.60 0.49</td>
|
||
</tr>
|
||
<tr id="3538AE2AFFCE0049FF1AF981E5BBE308" box="[151,1436,1621,1643]" gridrow="22" pageId="3" pageNumber="46">
|
||
<th id="76E9C756FFCE0049FF1AF981E1F8E308" box="[151,479,1621,1643]" gridcol="0" gridrow="22" pageId="3" pageNumber="46">Caudal peduncle length/depth</th>
|
||
<td id="76E9C756FFCE0049FDA2F981E2AAE308" box="[559,653,1621,1643]" gridcol="1" gridrow="22" pageId="3" pageNumber="46">1.8</td>
|
||
<td id="76E9C756FFCE0049FD4FF981E383E308" box="[706,932,1621,1643]" gridcol="2" gridrow="22" pageId="3" pageNumber="46">1.8–2.4 (1.9, 0.19)</td>
|
||
<td id="76E9C756FFCE0049FC6DF981E4F7E308" box="[992,1232,1621,1643]" gridcol="3" gridrow="22" pageId="3" pageNumber="46">1.8–2.1 (1.9, 0.09)</td>
|
||
<td id="76E9C756FFCE0049FA96F981E5BBE308" box="[1307,1436,1621,1643]" gridcol="4" gridrow="22" pageId="3" pageNumber="46">0.75</td>
|
||
</tr>
|
||
<tr id="3538AE2AFFCE0049FF1AF9AAE5BBE3A7" box="[151,1436,1662,1732]" gridrow="23" pageId="3" pageNumber="46">
|
||
<th id="76E9C756FFCE0049FF1AF9AAE1F8E3A7" box="[151,479,1662,1732]" gridcol="0" gridrow="23" pageId="3" pageNumber="46">P1 origin to P2 origin/P1 P2 origin to anal-fin origin/P2</th>
|
||
<td id="76E9C756FFCE0049FDA2F9AAE2AAE3A7" box="[559,653,1662,1732]" gridcol="1" gridrow="23" pageId="3" pageNumber="46">1.1 1.1</td>
|
||
<td id="76E9C756FFCE0049FD4FF9AAE383E3A7" box="[706,932,1662,1732]" gridcol="2" gridrow="23" pageId="3" pageNumber="46">0.9–1.1 (1.0, 0.06) 0.8–1.2 (0.9, 0.09)</td>
|
||
<td id="76E9C756FFCE0049FC6DF9AAE4F7E3A7" box="[992,1232,1662,1732]" gridcol="3" gridrow="23" pageId="3" pageNumber="46">0.9–1.1 (1.0, 0.08) 0.9–1.1 (1.0, 0.06)</td>
|
||
<td id="76E9C756FFCE0049FA96F9AAE5BBE3A7" box="[1307,1436,1662,1732]" gridcol="4" gridrow="23" pageId="3" pageNumber="46">0.07 0.06</td>
|
||
</tr>
|
||
</table>
|
||
</paragraph>
|
||
<paragraph id="8BB7AC68FFCEFFB5FF44F8D1E4E6E2CE" blockId="3.[151,1436,1794,2037]" pageId="3" pageNumber="46">
|
||
Pharyngeal tooth counts are
|
||
<quantity id="4CF0018DFFCEFFB5FD97F8D1E2A8E27E" box="[538,655,1797,1821]" metricMagnitude="-2" metricUnit="m" metricValue="8.509" metricValueMax="10.668" metricValueMin="6.35" pageId="3" pageNumber="46" unit="in" value="3.35" valueMax="4.2" valueMin="2.5">2.5–4.2 in</quantity>
|
||
6 fish, and in other 13 fish only the right 5th ceratobranchial was examined where –4.2 were found in
|
||
<specimenCount id="9D0E67E1FFCEFFB5FDB2F8FCE2FEE223" box="[575,729,1832,1856]" pageId="3" pageNumber="46" type="generic">12 specimens</specimenCount>
|
||
and –
|
||
<quantity id="4CF0018DFFCEFFB5FCAEF8FCE343E223" box="[803,868,1832,1856]" metricMagnitude="-1" metricUnit="m" metricValue="1.3208" pageId="3" pageNumber="46" unit="in" value="5.2">5.2 in</quantity>
|
||
one. Teeth are hooked at the tip and not serrated below it. The gut shape is a simple “S” with larger fish having a slight loop to the left anteriorly. Total gill rakers in the outer row on first left arch number 5(1), 6(4), 7(5), 8(7) or 9(1) (7.2, 1.04); gill rakers are not long and are widely spaced, touching the adjacent raker when appressed, often small and weakly developed.
|
||
</paragraph>
|
||
<paragraph id="8BB7AC68FFCEFFB2FF44F86CE3BCE47F" blockId="3.[151,1436,1794,2037]" lastBlockId="4.[151,1437,151,1978]" lastPageId="4" lastPageNumber="47" pageId="3" pageNumber="46">The total vertebrae are 40 (2), 41 (13) or 42 (3). Abdominal vertebrae are 19 (3), 20 (14) or 21 (1); predorsal abdominal vertebrae are 12 (1), 13 (13) or 14 (4); intermediate vertebrae are 2 (3), 3 (14) or 4 (1). Caudal vertebrae are 20 (1), 21 (13) or 22 (4). The vertebral formulae are 20+21 (11), 20+22 (3), 19+21 (3), 19+22 (1) or 21+20 (1). Thus, the caudal vertebral region most commonly (in all specimens but one) is longer then the abdominal region, the difference between abdominal and caudal counts being –1 (11), –2 (6), –3 (1) or 1 (1). A postcleithrum in the pectoral-fin skeleton is small and maybe absent on one or both sides.</paragraph>
|
||
<paragraph id="8BB7AC68FFC9FFB2FF44FEF3E18FE4CF" blockId="4.[151,1437,151,1978]" pageId="4" pageNumber="47">
|
||
Canal pores counts were examined in 13
|
||
<typeStatus id="54B312CAFFC9FFB2FD10FEFCE32FE423" box="[669,776,296,320]" pageId="4" pageNumber="47" type="paratype">paratypes</typeStatus>
|
||
on both sides. The supraorbital canal has 8 (11), 9 (13) or 10 (2) pores, the infraorbital canal has 12 (1), 13 (2), 14 (9), 15 (9), 16 (1), 17 (2), 18 (1) or 20 (1) pores, the preopercular-mandibular canal has 14 (11), 15 (12), 16 (1) or 17 (2) pores, and the supratemporal canal (unpaired) has 5 (3), 6 (5), or 7 (5) pores.
|
||
</paragraph>
|
||
<paragraph id="8BB7AC68FFC9FFB2FF44FE63E3CAE7C4" blockId="4.[151,1437,151,1978]" pageId="4" pageNumber="47">
|
||
The body is markedly compressed. The upper body profile is moderately rounded while the lower profile is more convex or both profiles are about equally convex. The eye is large, orbit width about equal to interorbital width. The snout is short, slightly pointed, not stout. The mouth is terminal, with the tip of the mouth cleft on a level with the upper half of the pupil. Neither lower jaw nor upper jaw protrudes relative to each other, so no ‘chin’ is formed by the symphysis of the lower jaws. The mouth cleft is always turned upward, never horizontal, and the junction of the lower jaw and the quadrate is on about a vertical through the anterior eye margin. In those
|
||
<typeStatus id="54B312CAFFC9FFB2FABCFDB9E5BBE7E6" box="[1329,1436,621,645]" pageId="4" pageNumber="47" type="paratype">paratypes</typeStatus>
|
||
with an unbroken caudal-fin its lobes are pointed, the fin is clearly forked.
|
||
</paragraph>
|
||
<paragraph id="8BB7AC68FFC9FFB2FF44FD60E1DFE654" blockId="4.[151,1437,151,1978]" pageId="4" pageNumber="47">
|
||
Morphometrics are summarised in
|
||
<tableCitation id="C68A99D3FFC9FFB2FDECFD60E29EE7AF" box="[609,697,692,716]" captionStart="TABLE 1" captionStartId="3.[151,235,152,173]" captionTargetBox="[151,1436,262,1732]" captionTargetPageId="3" captionText="TABLE 1. Morphometric characters for Alburnoides holciki. Significant differences (p <0.05) between males and females marked by an asterisk (*); for preanal length (marked by #) means and ranges are given to two decimal places as rounding gives identical means." httpUri="http://table.plazi.org/id/DF77FCE0FFCEFFB5FF1AFF4CE11BE588" pageId="4" pageNumber="47" tableUuid="DF77FCE0FFCEFFB5FF1AFF4CE11BE588">Table 1</tableCitation>
|
||
. The majority of morphometric characters are not significantly different between males and females as shown by values in
|
||
<tableCitation id="C68A99D3FFC9FFB2FCD8FD03E38BE78C" box="[853,940,727,751]" captionStart="TABLE 1" captionStartId="3.[151,235,152,173]" captionTargetBox="[151,1436,262,1732]" captionTargetPageId="3" captionText="TABLE 1. Morphometric characters for Alburnoides holciki. Significant differences (p <0.05) between males and females marked by an asterisk (*); for preanal length (marked by #) means and ranges are given to two decimal places as rounding gives identical means." httpUri="http://table.plazi.org/id/DF77FCE0FFCEFFB5FF1AFF4CE11BE588" pageId="4" pageNumber="47" tableUuid="DF77FCE0FFCEFFB5FF1AFF4CE11BE588">Table 1</tableCitation>
|
||
. Exceptions are pectoral-fin and dorsal-fin lengths are longer in males (pectoral-fin often reaching the pelvic-fin origin), and preanal and pelvic-fin to anal-fin distances are longer in females.
|
||
</paragraph>
|
||
<paragraph id="8BB7AC68FFC9FFB2FF44FC90E2D3E68F" blockId="4.[151,1437,151,1978]" pageId="4" pageNumber="47">
|
||
Colouration. Pigmentation in preserved fish is as described for the
|
||
<typeStatus id="54B312CAFFC9FFB2FC46FC90E40AE63F" box="[971,1069,836,860]" pageId="4" pageNumber="47" type="holotype">holotype</typeStatus>
|
||
but there are 3–4 rows of spots anteriorly on the flank and these rows may extend to the end of the pelvic-fin level or just beyond it. Additionally, there may be 1–2 rows of spots below the anterior lateral line in front of the pelvic-fin level and above the pectoralfin. The peritoneum is white-grey to light brown overall, the latter from distinct but crowded pale brown spots. Some fish have a few, very small, black pigment dots.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C312FFE3FFC9FFB2FF44FC23E366E11C" pageId="4" pageNumber="47" type="materials_examined">
|
||
<paragraph id="8BB7AC68FFC9FFB2FF44FC23E366E11C" blockId="4.[151,1437,151,1978]" pageId="4" pageNumber="47">
|
||
<emphasis id="B97C707AFFC9FFB2FF44FC23E27FE173" bold="true" box="[201,600,1015,1040]" pageId="4" pageNumber="47">Description of non-type material.</emphasis>
|
||
Two specimens from River Kashaf share with
|
||
<typeStatus id="54B312CAFFC9FFB2FBE0FC23E4E8E16C" box="[1133,1231,1015,1039]" pageId="4" pageNumber="47" type="holotype">holotype</typeStatus>
|
||
and the
|
||
<typeStatus id="54B312CAFFC9FFB2FABCFC2DE5BBE172" box="[1329,1436,1017,1041]" pageId="4" pageNumber="47" type="paratype">paratypes</typeStatus>
|
||
most diagnostic features. Thus, they have a completely scaleless sharp ventral keel, –4.2 teeth on the right 5th ceratobranchial, an upturned mouth, 8½ branched dorsal-fin rays, 13½ and 14½ anal-fin branched rays, 41 (20+21) total vertebrae and 13 predorsal abdominal vertebrae (
|
||
<figureCitation id="1333B0EDFFC9FFB2FD78FBB3E31FE11C" box="[757,824,1127,1151]" captionStart="FIGURE 2" captionStartId="1.[151,250,1939,1962]" captionTargetBox="[168,1435,1507,1920]" captionTargetId="figure@1.[168,1453,1507,1920]" captionTargetPageId="1" captionText="FIGURE 2. Alburnoides holciki sp. n., radiograph of a non-type specimen, BMNH 1914.1.1: 30 - 31, 82.4 mm SL, Kashaf River at Mashhad (Iran). Shows an upturned mouth, a straightened upper profile of the snout, 13 predorsal vertebrae, 20 abdominal vertebrae, and 21 caudal vertebrae." httpUri="https://zenodo.org/record/214791/files/figure.png" pageId="4" pageNumber="47">Fig. 2</figureCitation>
|
||
)
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C312FFE3FFC9FFB2FF44FB5FE31EE1A4" pageId="4" pageNumber="47" type="etymology">
|
||
<paragraph id="8BB7AC68FFC9FFB2FF44FB5FE31EE1A4" blockId="4.[151,1437,151,1978]" pageId="4" pageNumber="47">
|
||
<emphasis id="B97C707AFFC9FFB2FF44FB5FE177E1C0" bold="true" box="[201,336,1163,1187]" pageId="4" pageNumber="47">Etymology.</emphasis>
|
||
The new species is named for the late Juraj Holcík, Bratislava, colleague and friend, in recognition of his many contributions to the science of ichthyology.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C312FFE3FFC9FFB3FF44FB06E487E352" lastPageId="5" lastPageNumber="48" pageId="4" pageNumber="47" type="distribution">
|
||
<paragraph id="8BB7AC68FFC9FFB2FF44FB06E4A6E084" blockId="4.[151,1437,151,1978]" pageId="4" pageNumber="47">
|
||
<emphasis id="B97C707AFFC9FFB2FF44FB06E1D3E188" bold="true" box="[201,500,1234,1259]" pageId="4" pageNumber="47">Distribution and habitat.</emphasis>
|
||
The new species is only known from historical collections in the Hari (= Tedzhen) River of
|
||
<collectingCountry id="F31FECF8FFC9FFB2FF70FB23E1A2E06C" box="[253,389,1271,1295]" name="Afghanistan" pageId="4" pageNumber="47">Afghanistan</collectingCountry>
|
||
and its western tributary from
|
||
<collectingCountry id="F31FECF8FFC9FFB2FD67FB2CE33DE06C" box="[746,794,1272,1295]" name="Iran" pageId="4" pageNumber="47">Iran</collectingCountry>
|
||
, the Kashaf River (
|
||
<figureCitation id="1333B0EDFFC9FFB2FC75FB23E41AE06C" box="[1016,1085,1271,1295]" captionStart="FIGURE 3" captionStartId="5.[151,250,1083,1106]" captionTargetBox="[185,1401,193,1062]" captionTargetId="figure@5.[185,1402,193,1062]" captionTargetPageId="5" captionText="FIGURE 3. Map showing the type locality of Alburnoides holciki sp. n.: Hari River system, 34 ° 20 ’ N, 62 ° 12 ’ E (empty star) and a locality of non-type material from Kashaf River (solid star) and sampling sites of comparative material: Alburnoides cf. eichwaldii west of Safid River (empty circles), Alburnoides cf. e ichwaldii Safid River drainage (solid circles), A. cf. eichwaldii Tajan River (empty triangle), Alburnoides sp., Atrek River drainage (solid triangles), A. varentsovi rivers of northern slope of Kopetdag (solid quadrates), Alburnoides sp. Amu Darya drainage (empty quadrates)." httpUri="https://zenodo.org/record/214792/files/figure.png" pageId="4" pageNumber="47">Fig. 3</figureCitation>
|
||
). The Hari River or Harirud is known as the Tedzhen River in
|
||
<collectingCountry id="F31FECF8FFC9FFB2FE70FACEE2BFE051" box="[509,664,1306,1330]" name="Turkmenistan" pageId="4" pageNumber="47">Turkmenistan</collectingCountry>
|
||
where it is lost in the sands of the Kara Kum desert. The Hari River in
|
||
<collectingCountry id="F31FECF8FFC9FFB2FF38FAEBE119E034" box="[181,318,1343,1367]" name="Afghanistan" pageId="4" pageNumber="47">Afghanistan</collectingCountry>
|
||
dried up completely in 2000 (Payvand News at www.payvand.com/news/
|
||
<date id="FFB68AA8FFC9FFB2FB04FAEBE528E034" box="[1161,1295,1343,1367]" pageId="4" pageNumber="47" value="1011-01-01">01/jan/1011</date>
|
||
.html) and is usually dry from August to November as water is diverted for irrigation and precipitation is low. The city of Mashhad on the Kashaf River in
|
||
<collectingCountry id="F31FECF8FFC9FFB2FE5AFA5CE223E0FC" box="[471,516,1416,1439]" name="Iran" pageId="4" pageNumber="47">Iran</collectingCountry>
|
||
has a population of about 2.5 million people with a concomitant draw on water resources. Water quality in the Kashaf River at Mashhad is degraded (
|
||
<bibRefCitation id="EF99D199FFC9FFB2FC28FA7EE485E0A1" author="Najafpoor" box="[933,1186,1450,1475]" pageId="4" pageNumber="47" refString="Najafpoor, A. A., Vojoudi, Z., Dehgani, M. H., Changani, F. & Alidadai, H. (2007) Quality assessment of the Kashaf River in North East of Iran in 1996 - 2005. Journal of Applied Sciences, 7 (2), 253 - 257." type="journal article" year="2007">
|
||
Najafpoor
|
||
<emphasis id="B97C707AFFC9FFB2FB92FA78E477E0A0" box="[1055,1104,1451,1475]" italics="true" pageId="4" pageNumber="47">et al</emphasis>
|
||
., 2007
|
||
</bibRefCitation>
|
||
). These shallow rivers with little riparian vegetation in a desert climate have habitats that are stressful for fishes.
|
||
</paragraph>
|
||
<paragraph id="8BB7AC68FFC9FFB2FF44FA26E43FE2D9" blockId="4.[151,1437,151,1978]" pageId="4" pageNumber="47">
|
||
<emphasis id="B97C707AFFC9FFB2FF44FA26E1C3E368" bold="true" box="[201,484,1522,1547]" pageId="4" pageNumber="47">Comparative remarks.</emphasis>
|
||
<taxonomicName id="4C08D7EBFFC9FFB2FE7AFA27E2F9E368" box="[503,734,1523,1547]" class="Actinopterygii" family="Cyprinidae" genus="Alburnoides" kingdom="Animalia" order="Cypriniformes" pageId="4" pageNumber="47" phylum="Chordata" rank="species" species="holciki">
|
||
<emphasis id="B97C707AFFC9FFB2FE7AFA27E2F9E368" box="[503,734,1523,1547]" italics="true" pageId="4" pageNumber="47">Alburnoides holciki</emphasis>
|
||
</taxonomicName>
|
||
clearly differs from Iranian species
|
||
<taxonomicName id="4C08D7EBFFC9FFB2FB36FA27E1EFE34C" authority="Bogutskaya & Coad, 2009" authorityName="Bogutskaya & Coad" authorityYear="2009" class="Actinopterygii" family="Cyprinidae" genus="Alburnoides" kingdom="Animalia" order="Cypriniformes" pageId="4" pageNumber="47" phylum="Chordata" rank="species" species="petrubanarescui">
|
||
<emphasis id="B97C707AFFC9FFB2FB36FA27E5BCE368" box="[1211,1435,1523,1547]" italics="true" pageId="4" pageNumber="47">A. petrubanarescui</emphasis>
|
||
Bogutskaya & Coad, 2009
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C08D7EBFFC9FFB2FE59F9C3E3A7E34C" authority="Bogutskaya & Coad, 2009" authorityName="Bogutskaya & Coad" authorityYear="2009" box="[468,896,1559,1583]" class="Actinopterygii" family="Cyprinidae" genus="Alburnoides" kingdom="Animalia" order="Cypriniformes" pageId="4" pageNumber="47" phylum="Chordata" rank="species" species="namaki">
|
||
<emphasis id="B97C707AFFC9FFB2FE59F9C3E260E34C" box="[468,583,1559,1583]" italics="true" pageId="4" pageNumber="47">A. namaki</emphasis>
|
||
Bogutskaya & Coad, 2009
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C08D7EBFFC9FFB2FC00F9C3E56DE34C" authority="Bogutskaya & Coad, 2009" authorityName="Bogutskaya & Coad" authorityYear="2009" box="[909,1354,1559,1583]" class="Actinopterygii" family="Cyprinidae" genus="Alburnoides" kingdom="Animalia" order="Cypriniformes" pageId="4" pageNumber="47" phylum="Chordata" rank="species" species="nicolausi">
|
||
<emphasis id="B97C707AFFC9FFB2FC00F9C3E433E34C" box="[909,1044,1559,1583]" italics="true" pageId="4" pageNumber="47">A. nicolausi</emphasis>
|
||
Bogutskaya & Coad, 2009
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C08D7EBFFC9FFB2FA09F9C3E219E331" authority="Bogutskaya & Coad, 2009" authorityName="Bogutskaya & Coad" authorityYear="2009" class="Actinopterygii" family="Cyprinidae" genus="Alburnoides" kingdom="Animalia" order="Cypriniformes" pageId="4" pageNumber="47" phylum="Chordata" rank="species" species="idignensis">
|
||
<emphasis id="B97C707AFFC9FFB2FA09F9C3E12EE330" italics="true" pageId="4" pageNumber="47">A. idignensis</emphasis>
|
||
Bogutskaya & Coad, 2009
|
||
</taxonomicName>
|
||
distributed in the south of the Caspian Sea basin by having a short, slightly pointed snout (vs. stout and rounded), a terminal mouth with the tip of the mouth cleft on a level with the upper half of the pupil (vs. almost or completely subterminal with a tip of the mouth on a level with the lower margin of the pupil to below the lower margin of the eye), a clearly forked caudal-fin with pointed lobes (vs. shallowly forked with rounded lobes), and 13–16½, usually 14–15½, branched anal-fin rays (vs. 8–13½, usually 9–12½), and orbit width about equal to interorbital width (vs. smaller). The new species is further easily distinguishable from
|
||
<taxonomicName id="4C08D7EBFFC9FFB2FA09F93BE16AE248" class="Actinopterygii" family="Cyprinidae" genus="Alburnoides" kingdom="Animalia" order="Cypriniformes" pageId="4" pageNumber="47" phylum="Chordata" rank="species" species="petrubanarescui">
|
||
<emphasis id="B97C707AFFC9FFB2FA09F93BE16AE248" italics="true" pageId="4" pageNumber="47">A. petrubanarescui</emphasis>
|
||
</taxonomicName>
|
||
in having a sharp, usually scaleless ventral keel (vs. smooth and completely scaled) and a caudal vertebral region longer than an abdominal region: most frequent vertebral formulae 20+21, 20+22 and 19+21 (vs. shorter, usually 21+19 and 21+20), from
|
||
<taxonomicName id="4C08D7EBFFC9FFB2FDFDF88FE2DFE210" box="[624,760,1883,1907]" class="Actinopterygii" family="Cyprinidae" genus="Alburnoides" kingdom="Animalia" order="Cypriniformes" pageId="4" pageNumber="47" phylum="Chordata" rank="species" species="nicolausi">
|
||
<emphasis id="B97C707AFFC9FFB2FDFDF88FE2DFE210" box="[624,760,1883,1907]" italics="true" pageId="4" pageNumber="47">A. nicolausi</emphasis>
|
||
</taxonomicName>
|
||
– in having usually 8½ branched anal-fin rays (vs. usually 7½), and from both
|
||
<taxonomicName id="4C08D7EBFFC9FFB2FE09F8ABE22BE2F4" box="[388,524,1919,1943]" class="Actinopterygii" family="Cyprinidae" genus="Alburnoides" kingdom="Animalia" order="Cypriniformes" pageId="4" pageNumber="47" phylum="Chordata" rank="species" species="nicolausi">
|
||
<emphasis id="B97C707AFFC9FFB2FE09F8ABE22BE2F4" box="[388,524,1919,1943]" italics="true" pageId="4" pageNumber="47">A. nicolausi</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<emphasis id="B97C707AFFC9FFB2FDC7F8ABE2D1E2F4" box="[586,758,1919,1943]" italics="true" pageId="4" pageNumber="47">
|
||
<taxonomicName id="4C08D7EBFFC9FFB2FDC7F8ABE2FAE2F4" box="[586,733,1919,1943]" class="Actinopterygii" family="Cyprinidae" genus="Alburnoides" kingdom="Animalia" order="Cypriniformes" pageId="4" pageNumber="47" phylum="Chordata" rank="species" species="idignensis">A. idignensis</taxonomicName>
|
||
–
|
||
</emphasis>
|
||
in having 40–42, usually 41, total vertebrae and 13 or 14 predorsal vertebrae (vs. 37–40, usually 39, and 11–13, usually 12, respectively).
|
||
</paragraph>
|
||
<caption id="DF77FCE0FFC8FFB3FF1AFBEFE3D3E1AF" httpUri="https://zenodo.org/record/214792/files/figure.png" pageId="5" pageNumber="48" targetBox="[185,1401,193,1062]" targetPageId="5">
|
||
<paragraph id="8BB7AC68FFC8FFB3FF1AFBEFE3D3E1AF" blockId="5.[151,1437,1083,1228]" pageId="5" pageNumber="48">
|
||
<emphasis id="B97C707AFFC8FFB3FF1AFBEFE133E132" bold="true" box="[151,276,1083,1106]" pageId="5" pageNumber="48">FIGURE 3.</emphasis>
|
||
Map showing the type locality of
|
||
<taxonomicName id="4C08D7EBFFC8FFB3FDF7FBE8E364E131" box="[634,835,1084,1106]" class="Actinopterygii" family="Cyprinidae" genus="Alburnoides" kingdom="Animalia" order="Cypriniformes" pageId="5" pageNumber="48" phylum="Chordata" rank="species" species="holciki" status="sp. nov.">
|
||
<emphasis id="B97C707AFFC8FFB3FDF7FBE8E364E131" box="[634,835,1084,1106]" italics="true" pageId="5" pageNumber="48">Alburnoides holciki</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A24FCD01FFC8FFB3FCC6FBE8E3A2E132" box="[843,901,1084,1105]" pageId="5" pageNumber="48" rank="species">
|
||
<emphasis id="B97C707AFFC8FFB3FCC6FBE8E3A2E132" bold="true" box="[843,901,1084,1105]" pageId="5" pageNumber="48">sp. n.</emphasis>
|
||
</taxonomicNameLabel>
|
||
: Hari River system, 34°20’N, 62°12’E (empty star) and a locality of non-type material from Kashaf River (solid star) and sampling sites of comparative material:
|
||
<taxonomicName id="4C08D7EBFFC8FFB3FB72FB8EE126E1EC" class="Actinopterygii" family="Cyprinidae" genus="Alburnoides" kingdom="Animalia" order="Cypriniformes" pageId="5" pageNumber="48" phylum="Chordata" rank="species" species="eichwaldii">
|
||
<emphasis id="B97C707AFFC8FFB3FB72FB8EE55CE113" box="[1279,1403,1114,1136]" italics="true" pageId="5" pageNumber="48">Alburnoides</emphasis>
|
||
cf.
|
||
<emphasis id="B97C707AFFC8FFB3FF1AFBADE126E1EC" box="[151,257,1145,1167]" italics="true" pageId="5" pageNumber="48">eichwaldii</emphasis>
|
||
</taxonomicName>
|
||
west of Safid River (empty circles),
|
||
<taxonomicName id="4C08D7EBFFC8FFB3FDFBFBADE2D5E1EC" box="[630,754,1145,1167]" class="Actinopterygii" family="Cyprinidae" genus="Alburnoides" kingdom="Animalia" order="Cypriniformes" pageId="5" pageNumber="48" phylum="Chordata" rank="genus">
|
||
<emphasis id="B97C707AFFC8FFB3FDFBFBADE2D5E1EC" box="[630,754,1145,1167]" italics="true" pageId="5" pageNumber="48">Alburnoides</emphasis>
|
||
</taxonomicName>
|
||
cf. e
|
||
<emphasis id="B97C707AFFC8FFB3FCAEFBADE3A6E1EC" box="[803,897,1145,1167]" italics="true" pageId="5" pageNumber="48">ichwaldii</emphasis>
|
||
Safid River drainage (solid circles),
|
||
<taxonomicName id="4C08D7EBFFC8FFB3FB7BFBAEE129E1CE" authority="Tajan River" authorityName="Tajan River" class="Actinopterygii" family="Cyprinidae" genus="Alburnoides" kingdom="Animalia" order="Cypriniformes" pageId="5" pageNumber="48" phylum="Chordata" rank="species" species="eichwaldii">
|
||
<emphasis id="B97C707AFFC8FFB3FB7BFBAEE52AE1EC" box="[1270,1293,1146,1167]" italics="true" pageId="5" pageNumber="48">A.</emphasis>
|
||
cf.
|
||
<emphasis id="B97C707AFFC8FFB3FABFFBADE5BCE1EC" box="[1330,1435,1145,1167]" italics="true" pageId="5" pageNumber="48">eichwaldii</emphasis>
|
||
Tajan River
|
||
</taxonomicName>
|
||
(empty triangle),
|
||
<taxonomicName id="4C08D7EBFFC8FFB3FE45FB43E263E1CE" box="[456,580,1175,1197]" class="Actinopterygii" family="Cyprinidae" genus="Alburnoides" kingdom="Animalia" order="Cypriniformes" pageId="5" pageNumber="48" phylum="Chordata" rank="genus">
|
||
<emphasis id="B97C707AFFC8FFB3FE45FB43E263E1CE" box="[456,580,1175,1197]" italics="true" pageId="5" pageNumber="48">Alburnoides</emphasis>
|
||
</taxonomicName>
|
||
sp., Atrek River drainage (solid triangles),
|
||
<taxonomicName id="4C08D7EBFFC8FFB3FB8EFB4CE4AEE1CD" box="[1027,1161,1176,1198]" class="Actinopterygii" family="Cyprinidae" genus="Alburnoides" kingdom="Animalia" order="Cypriniformes" pageId="5" pageNumber="48" phylum="Chordata" rank="species" species="varentsovi">
|
||
<emphasis id="B97C707AFFC8FFB3FB8EFB4CE4AEE1CD" box="[1027,1161,1176,1198]" italics="true" pageId="5" pageNumber="48">A. varentsovi</emphasis>
|
||
</taxonomicName>
|
||
rivers of northern slope of Kopetdag (solid quadrates),
|
||
<taxonomicName id="4C08D7EBFFC8FFB3FE38FB62E216E1AF" box="[437,561,1206,1228]" class="Actinopterygii" family="Cyprinidae" genus="Alburnoides" kingdom="Animalia" order="Cypriniformes" pageId="5" pageNumber="48" phylum="Chordata" rank="genus">
|
||
<emphasis id="B97C707AFFC8FFB3FE38FB62E216E1AF" box="[437,561,1206,1228]" italics="true" pageId="5" pageNumber="48">Alburnoides</emphasis>
|
||
</taxonomicName>
|
||
sp. Amu Darya drainage (empty quadrates).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BB7AC68FFC8FFB3FF44FB2DE487E352" blockId="5.[151,1436,1273,2017]" pageId="5" pageNumber="48">
|
||
<taxonomicName id="4C08D7EBFFC8FFB3FF44FB2DE182E072" box="[201,421,1273,1297]" class="Actinopterygii" family="Cyprinidae" genus="Alburnoides" kingdom="Animalia" order="Cypriniformes" pageId="5" pageNumber="48" phylum="Chordata" rank="species" species="holciki">
|
||
<emphasis id="B97C707AFFC8FFB3FF44FB2DE182E072" box="[201,421,1273,1297]" italics="true" pageId="5" pageNumber="48">Alburnoides holciki</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C08D7EBFFC8FFB3FE50FB2DE277E072" box="[477,592,1273,1297]" class="Actinopterygii" family="Cyprinidae" genus="Alburnoides" kingdom="Animalia" order="Cypriniformes" pageId="5" pageNumber="48" phylum="Chordata" rank="species" species="namaki">
|
||
<emphasis id="B97C707AFFC8FFB3FE50FB2DE277E072" box="[477,592,1273,1297]" italics="true" pageId="5" pageNumber="48">A. namaki</emphasis>
|
||
</taxonomicName>
|
||
are similar in having a sharp scaleless ventral keel but
|
||
<taxonomicName id="4C08D7EBFFC8FFB3FB4FFB2DE50AE072" box="[1218,1325,1273,1297]" class="Actinopterygii" family="Cyprinidae" genus="Alburnoides" kingdom="Animalia" order="Cypriniformes" pageId="5" pageNumber="48" phylum="Chordata" rank="species" species="holciki">
|
||
<emphasis id="B97C707AFFC8FFB3FB4FFB2DE50AE072" box="[1218,1325,1273,1297]" italics="true" pageId="5" pageNumber="48">A. holciki</emphasis>
|
||
</taxonomicName>
|
||
is clearly different from the latter species in having a pointed snout and a terminal mouth with the tip of the mouth cleft on a level with the upper half of the pupil (vs. a stout markedly rounded snout, a subterminal mouth with the tip of the mouth cleft on a level below the lower margin of the eye), 40–42, usually 41, total vertebrae (vs. 39–41, usually 40), 13 or 14 predorsal vertebrae (vs. usually 12), and 13–16½, usually 14–15½, branched anal-fin rays (vs. 10–13½, usually 11–12½). The new species resembles
|
||
<taxonomicName id="4C08D7EBFFC8FFB3FC85FA79E532E0A6" authority="Coad & Bogutskaya, 2009" authorityName="Coad & Bogutskaya" authorityYear="2009" box="[776,1301,1453,1477]" class="Actinopterygii" family="Cyprinidae" genus="Alburnoides" kingdom="Animalia" order="Cypriniformes" pageId="5" pageNumber="48" phylum="Chordata" rank="species" species="qanati">
|
||
<emphasis id="B97C707AFFC8FFB3FC85FA79E3C7E0A6" box="[776,992,1453,1477]" italics="true" pageId="5" pageNumber="48">Alburnoides qanati</emphasis>
|
||
Coad & Bogutskaya,
|
||
<quantity id="4CF0018DFFC8FFB3FB50FA79E532E0A6" box="[1245,1301,1453,1477]" metricMagnitude="1" metricUnit="m" metricValue="5.10286" pageId="5" pageNumber="48" unit="in" value="2009.0">2009</quantity>
|
||
</taxonomicName>
|
||
in having a terminal upturned mouth but in the latter species the mouth cleft is even more vertical and the tip of the mouth cleft is on about a level with the upper margin of the pupil.
|
||
<taxonomicName id="4C08D7EBFFC8FFB3FD76FA21E3F0E36E" box="[763,983,1525,1549]" class="Actinopterygii" family="Cyprinidae" genus="Alburnoides" kingdom="Animalia" order="Cypriniformes" pageId="5" pageNumber="48" phylum="Chordata" rank="species" species="holciki">
|
||
<emphasis id="B97C707AFFC8FFB3FD76FA21E3F0E36E" box="[763,983,1525,1549]" italics="true" pageId="5" pageNumber="48">Alburnoides holciki</emphasis>
|
||
</taxonomicName>
|
||
differs from
|
||
<taxonomicName id="4C08D7EBFFC8FFB3FBE1FA21E4F3E36E" box="[1132,1236,1525,1549]" class="Actinopterygii" family="Cyprinidae" genus="Alburnoides" kingdom="Animalia" order="Cypriniformes" pageId="5" pageNumber="48" phylum="Chordata" rank="species" species="qanati">
|
||
<emphasis id="B97C707AFFC8FFB3FBE1FA21E4F3E36E" box="[1132,1236,1525,1549]" italics="true" pageId="5" pageNumber="48">A. qanati</emphasis>
|
||
</taxonomicName>
|
||
in having a sharp scaleless ventral keel (vs. smooth and scaled), 13–16½ branched anal-fin rays (vs. 10–12½).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C312FFE3FFC8FFBDFF44F9E9E1B3E75F" lastPageId="11" lastPageNumber="54" pageId="5" pageNumber="48" type="reference_group">
|
||
<paragraph id="8BB7AC68FFC8FFB0FF44F9E9E31EE45C" blockId="5.[151,1436,1273,2017]" lastBlockId="6.[151,1436,151,320]" lastPageId="6" lastPageNumber="49" pageId="5" pageNumber="48">
|
||
<taxonomicName id="4C08D7EBFFC8FFB3FF44F9E9E175E336" ID-CoL="RK5" box="[201,338,1597,1621]" class="Actinopterygii" family="Cyprinidae" genus="Alburnoides" kingdom="Animalia" order="Cypriniformes" pageId="5" pageNumber="48" phylum="Chordata" rank="genus">
|
||
<emphasis id="B97C707AFFC8FFB3FF44F9E9E175E336" box="[201,338,1597,1621]" italics="true" pageId="5" pageNumber="48">Alburnoides</emphasis>
|
||
</taxonomicName>
|
||
from the Caspian and Aral basins (including presently endorheic drainages such as Murghab, Hari and rivers of the Kopetdag Mountains) contain at least two species described up to now (the third one is being described in this paper),
|
||
<taxonomicName id="4C08D7EBFFC8FFB3FE27F951E329E3FE" ID-CoL="66875" authority="De Filippi, 1863" authorityName="De Filippi" authorityYear="1863" box="[426,782,1669,1693]" class="Actinopterygii" family="Cyprinidae" genus="Alburnoides" kingdom="Animalia" order="Cypriniformes" pageId="5" pageNumber="48" phylum="Chordata" rank="species" species="eichwaldii">
|
||
<emphasis id="B97C707AFFC8FFB3FE27F951E21AE3FE" box="[426,573,1669,1693]" italics="true" pageId="5" pageNumber="48">A. eichwaldii</emphasis>
|
||
(
|
||
<bibRefCitation id="EF99D199FFC8FFB3FDC1F951E321E3FE" author="De" box="[588,774,1669,1693]" pageId="5" pageNumber="48" refString="De Filippi, F. (1863) Nuove o poco note specie di animali vertebrati raccolte in un Viaggio in Persia. Archivio per la Zoologia, Anatomia e la Fisiologia, Modena, 2 (2), 377 - 394." type="journal article" year="1863">
|
||
<collectingCountry id="F31FECF8FFC8FFB3FDC1F951E249E3FE" box="[588,622,1669,1693]" name="South Georgia and the South Sandwich Islands" pageId="5" pageNumber="48">De</collectingCountry>
|
||
Filippi, 1863
|
||
</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C08D7EBFFC8FFB3FCC9F951E52FE3FE" ID-CoL="BGVV" authority="Bogutskaya & Coad, 2009" authorityName="Bogutskaya & Coad" authorityYear="2009" box="[836,1288,1669,1693]" class="Actinopterygii" family="Cyprinidae" genus="Alburnoides" kingdom="Animalia" order="Cypriniformes" pageId="5" pageNumber="48" phylum="Chordata" rank="species" species="varentsovi">
|
||
<emphasis id="B97C707AFFC8FFB3FCC9F951E3F0E3FE" box="[836,983,1669,1693]" italics="true" pageId="5" pageNumber="48">A. varentsovi</emphasis>
|
||
Bogutskaya & Coad, 2009
|
||
</taxonomicName>
|
||
.
|
||
<taxonomicName id="4C08D7EBFFC8FFB3FA9EF951E12BE3A2" ID-CoL="66875" class="Actinopterygii" family="Cyprinidae" genus="Alburnoides" kingdom="Animalia" order="Cypriniformes" pageId="5" pageNumber="48" phylum="Chordata" rank="species" species="eichwaldii">
|
||
<emphasis id="B97C707AFFC8FFB3FA9EF951E12BE3A2" italics="true" pageId="5" pageNumber="48">Alburnoides eichwaldii</emphasis>
|
||
</taxonomicName>
|
||
was originally described from the "Kur presso Tiflis" (= Kura River near Tbilisi, now
|
||
<collectingCountry id="F31FECF8FFC8FFB3FB60F97DE56CE3A2" box="[1261,1355,1705,1729]" name="Georgia" pageId="5" pageNumber="48">Georgia</collectingCountry>
|
||
), so an
|
||
<taxonomicName id="4C08D7EBFFC8FFB3FF1AF919E107E386" ID-CoL="RK5" box="[151,288,1741,1765]" class="Actinopterygii" family="Cyprinidae" genus="Alburnoides" kingdom="Animalia" order="Cypriniformes" pageId="5" pageNumber="48" phylum="Chordata" rank="genus">
|
||
<emphasis id="B97C707AFFC8FFB3FF1AF919E107E386" box="[151,288,1741,1765]" italics="true" pageId="5" pageNumber="48">Alburnoides</emphasis>
|
||
</taxonomicName>
|
||
from Kura River drainage is a bearer of the name; there is another nominal subspecies in the Aras River (a tributary of the Kura) system of
|
||
<collectingCountry id="F31FECF8FFC8FFB3FDEEF925E2EFE26A" box="[611,712,1777,1801]" name="Armenia" pageId="5" pageNumber="48">Armenia</collectingCountry>
|
||
,
|
||
<taxonomicName id="4C08D7EBFFC8FFB3FD5EF925E516E26A" ID-CoL="7J9CD" authority="Dadikyan, 1972" authorityName="Dadikyan" authorityYear="1972" box="[723,1329,1777,1801]" class="Actinopterygii" family="Cyprinidae" genus="Alburnoides" kingdom="Animalia" order="Cypriniformes" pageId="5" pageNumber="48" phylum="Chordata" rank="subSpecies" species="bipunctatus" subSpecies="armeniensis">
|
||
<emphasis id="B97C707AFFC8FFB3FD5EF925E452E26A" box="[723,1141,1777,1801]" italics="true" pageId="5" pageNumber="48">Alburnoides bipunctatus armeniensis</emphasis>
|
||
<bibRefCitation id="EF99D199FFC8FFB3FBF6F925E516E26A" author="Dadikyan" box="[1147,1329,1777,1801]" pageId="5" pageNumber="48" refString="Dadikyan, M. G. (1972) A new subspecies of European riffle minnow Alburnoides bipunctatus armeniensis subsp. nov. Voprosy Ikhtiologii, 12 (3), 566 - 569 (in Russian; translated in Journal of Ichthyology, 12 (3), 519 - 522)." type="journal article" year="1972">Dadikyan, 1972</bibRefCitation>
|
||
</taxonomicName>
|
||
, from the rivers Arpa, Vorotan, Vedi, Marmarik, Kasakh and their tributaries, now regarded as a synonym of
|
||
<taxonomicName id="4C08D7EBFFC8FFB3FAAAF8C1E1F3E232" ID-CoL="66875" authority="Bogutskaya & Coad 2009" authorityName="Bogutskaya & Coad" authorityYear="2009" class="Actinopterygii" family="Cyprinidae" genus="Alburnoides" kingdom="Animalia" order="Cypriniformes" pageId="5" pageNumber="48" phylum="Chordata" rank="species" species="eichwaldii">
|
||
<emphasis id="B97C707AFFC8FFB3FAAAF8C1E5BBE24E" box="[1319,1436,1813,1837]" italics="true" pageId="5" pageNumber="48">eichwaldii</emphasis>
|
||
(Bogutskaya & Coad 2009)
|
||
</taxonomicName>
|
||
. Bogutskaya & Coad (2009) made an assumption that riffle minnows from the Safid River drainage and the Amu Darya drainage may represent two distinct species. These issues are outside the goal of this paper; for comparisons that confirm the species status of
|
||
<taxonomicName id="4C08D7EBFFC8FFB3FCFDF855E3F8E2FA" ID-CoL="BGV6" box="[880,991,1921,1945]" class="Actinopterygii" family="Cyprinidae" genus="Alburnoides" kingdom="Animalia" order="Cypriniformes" pageId="5" pageNumber="48" phylum="Chordata" rank="species" species="holciki">
|
||
<emphasis id="B97C707AFFC8FFB3FCFDF855E3F8E2FA" box="[880,991,1921,1945]" italics="true" pageId="5" pageNumber="48">A. holciki</emphasis>
|
||
</taxonomicName>
|
||
we subdivided the examined samples besides those of
|
||
<taxonomicName id="4C08D7EBFFC8FFB3FEDDF871E19DE2DE" ID-CoL="BGV6" box="[336,442,1957,1981]" class="Actinopterygii" family="Cyprinidae" genus="Alburnoides" kingdom="Animalia" order="Cypriniformes" pageId="5" pageNumber="48" phylum="Chordata" rank="species" species="holciki">
|
||
<emphasis id="B97C707AFFC8FFB3FEDDF871E19DE2DE" box="[336,442,1957,1981]" italics="true" pageId="5" pageNumber="48">A. holciki</emphasis>
|
||
</taxonomicName>
|
||
into seven groups: 1.
|
||
<taxonomicName id="4C08D7EBFFC8FFB3FD3AF871E36CE2DE" ID-CoL="66875" box="[695,843,1957,1981]" class="Actinopterygii" family="Cyprinidae" genus="Alburnoides" kingdom="Animalia" order="Cypriniformes" pageId="5" pageNumber="48" phylum="Chordata" rank="species" species="eichwaldii">
|
||
<emphasis id="B97C707AFFC8FFB3FD3AF871E36CE2DE" box="[695,843,1957,1981]" italics="true" pageId="5" pageNumber="48">A. eichwaldii</emphasis>
|
||
</taxonomicName>
|
||
from Kura-Aras and Lenkoran (in the north from the Talysh Mountains), 2.
|
||
<taxonomicName id="4C08D7EBFFC8FFB3FE1CF81DE29EE282" ID-CoL="66875" box="[401,697,1993,2017]" class="Actinopterygii" family="Cyprinidae" genus="Alburnoides" kingdom="Animalia" order="Cypriniformes" pageId="5" pageNumber="48" phylum="Chordata" rank="species" species="eichwaldii">
|
||
<emphasis id="B97C707AFFC8FFB3FE1CF81DE23DE282" box="[401,538,1993,2017]" italics="true" pageId="5" pageNumber="48">Alburnoides</emphasis>
|
||
cf.
|
||
<emphasis id="B97C707AFFC8FFB3FDC9F81DE29EE282" box="[580,697,1993,2017]" italics="true" pageId="5" pageNumber="48">eichwaldii</emphasis>
|
||
</taxonomicName>
|
||
from rivers in the south from the Talysh Mountains and west from the Safid, 3.
|
||
<taxonomicName id="4C08D7EBFFCBFFB0FEABFF4CE277E5D3" ID-CoL="66875" box="[294,592,151,176]" class="Actinopterygii" family="Cyprinidae" genus="Alburnoides" kingdom="Animalia" order="Cypriniformes" pageId="6" pageNumber="49" phylum="Chordata" rank="species" species="eichwaldii">
|
||
<emphasis id="B97C707AFFCBFFB0FEABFF4CE188E5D3" box="[294,431,152,176]" italics="true" pageId="6" pageNumber="49">Alburnoides</emphasis>
|
||
cf.
|
||
<emphasis id="B97C707AFFCBFFB0FE56FF4CE277E5D3" box="[475,592,152,176]" italics="true" pageId="6" pageNumber="49">eichwaldii</emphasis>
|
||
</taxonomicName>
|
||
from the Safid River drainage, 4.
|
||
<taxonomicName id="4C08D7EBFFCBFFB0FC5EFF4CE47BE5D3" ID-CoL="RK5" box="[979,1116,152,176]" class="Actinopterygii" family="Cyprinidae" genus="Alburnoides" kingdom="Animalia" order="Cypriniformes" pageId="6" pageNumber="49" phylum="Chordata" rank="genus">
|
||
<emphasis id="B97C707AFFCBFFB0FC5EFF4CE47BE5D3" box="[979,1116,152,176]" italics="true" pageId="6" pageNumber="49">Alburnoides</emphasis>
|
||
</taxonomicName>
|
||
sp. from the Tajan River (in the east of the Safid), 5.
|
||
<taxonomicName id="4C08D7EBFFCBFFB0FE2AFF68E217E5B7" ID-CoL="RK5" box="[423,560,188,212]" class="Actinopterygii" family="Cyprinidae" genus="Alburnoides" kingdom="Animalia" order="Cypriniformes" pageId="6" pageNumber="49" phylum="Chordata" rank="genus">
|
||
<emphasis id="B97C707AFFCBFFB0FE2AFF68E217E5B7" box="[423,560,188,212]" italics="true" pageId="6" pageNumber="49">Alburnoides</emphasis>
|
||
</taxonomicName>
|
||
sp. from the Atrek River drainage, 6.
|
||
<taxonomicName id="4C08D7EBFFCBFFB0FC54FF68E44BE5B7" ID-CoL="BGVV" box="[985,1132,188,212]" class="Actinopterygii" family="Cyprinidae" genus="Alburnoides" kingdom="Animalia" order="Cypriniformes" pageId="6" pageNumber="49" phylum="Chordata" rank="species" species="varentsovi">
|
||
<emphasis id="B97C707AFFCBFFB0FC54FF68E44BE5B7" box="[985,1132,188,212]" italics="true" pageId="6" pageNumber="49">A. varentsovi</emphasis>
|
||
</taxonomicName>
|
||
from rivers of the northern slope of Kopetdag, 7.
|
||
<taxonomicName id="4C08D7EBFFCBFFB0FE02FF34E23FE59B" ID-CoL="RK5" box="[399,536,224,248]" class="Actinopterygii" family="Cyprinidae" genus="Alburnoides" kingdom="Animalia" order="Cypriniformes" pageId="6" pageNumber="49" phylum="Chordata" rank="genus">
|
||
<emphasis id="B97C707AFFCBFFB0FE02FF34E23FE59B" box="[399,536,224,248]" italics="true" pageId="6" pageNumber="49">Alburnoides</emphasis>
|
||
</taxonomicName>
|
||
sp. from the Amu Darya drainage (Tables 2–4,
|
||
<figureCitation id="1333B0EDFFCBFFB0FBBEFF0BE450E594" box="[1075,1143,223,247]" captionStart="FIGURE 3" captionStartId="5.[151,250,1083,1106]" captionTargetBox="[185,1401,193,1062]" captionTargetId="figure@5.[185,1402,193,1062]" captionTargetPageId="5" captionText="FIGURE 3. Map showing the type locality of Alburnoides holciki sp. n.: Hari River system, 34 ° 20 ’ N, 62 ° 12 ’ E (empty star) and a locality of non-type material from Kashaf River (solid star) and sampling sites of comparative material: Alburnoides cf. eichwaldii west of Safid River (empty circles), Alburnoides cf. e ichwaldii Safid River drainage (solid circles), A. cf. eichwaldii Tajan River (empty triangle), Alburnoides sp., Atrek River drainage (solid triangles), A. varentsovi rivers of northern slope of Kopetdag (solid quadrates), Alburnoides sp. Amu Darya drainage (empty quadrates)." httpUri="https://zenodo.org/record/214792/files/figure.png" pageId="6" pageNumber="49">Fig. 3</figureCitation>
|
||
). A UPGMA dendrogram using Kullback-Leibler divergences (
|
||
<figureCitation id="1333B0EDFFCBFFB0FDB5FED0E25BE47F" box="[568,636,260,284]" captionStart="FIGURE 4" captionStartId="6.[151,250,1224,1247]" captionTargetBox="[151,1426,387,1203]" captionTargetId="figure@6.[151,1436,373,1203]" captionTargetPageId="6" captionText="FIGURE 4. UPGMA dendrogram using Kullback-Leibler divergences for six meristic characters (the number of branched anal-fin rays and five vertebral counts)." httpUri="https://zenodo.org/record/214793/files/figure.png" pageId="6" pageNumber="49">Fig. 4</figureCitation>
|
||
) show the grouping of these units based on six meristic characters (the number of branched anal-fin rays and five vertebral counts).
|
||
</paragraph>
|
||
<caption id="DF77FCE0FFCBFFB0FF1AFB1CE200E19E" httpUri="https://zenodo.org/record/214793/files/figure.png" pageId="6" pageNumber="49" targetBox="[151,1426,387,1203]" targetPageId="6">
|
||
<paragraph id="8BB7AC68FFCBFFB0FF1AFB1CE200E19E" blockId="6.[151,1436,1224,1277]" pageId="6" pageNumber="49">
|
||
<emphasis id="B97C707AFFCBFFB0FF1AFB1CE132E1BD" bold="true" box="[151,277,1224,1247]" pageId="6" pageNumber="49">FIGURE 4.</emphasis>
|
||
UPGMA dendrogram using Kullback-Leibler divergences for six meristic characters (the number of branched anal-fin rays and five vertebral counts).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BB7AC68FFCBFFB0FF44FAF1E1E4E0AC" blockId="6.[151,1436,1316,2033]" pageId="6" pageNumber="49">
|
||
<taxonomicName id="4C08D7EBFFCBFFB0FF44FAF1E181E05E" ID-CoL="BGV6" box="[201,422,1317,1341]" class="Actinopterygii" family="Cyprinidae" genus="Alburnoides" kingdom="Animalia" order="Cypriniformes" pageId="6" pageNumber="49" phylum="Chordata" rank="species" species="holciki">
|
||
<emphasis id="B97C707AFFCBFFB0FF44FAF1E181E05E" box="[201,422,1317,1341]" italics="true" pageId="6" pageNumber="49">Alburnoides holciki</emphasis>
|
||
</taxonomicName>
|
||
is most similar to
|
||
<taxonomicName id="4C08D7EBFFCBFFB0FDF4FAF1E325E05E" ID-CoL="RK5" box="[633,770,1317,1341]" class="Actinopterygii" family="Cyprinidae" genus="Alburnoides" kingdom="Animalia" order="Cypriniformes" pageId="6" pageNumber="49" phylum="Chordata" rank="genus">
|
||
<emphasis id="B97C707AFFCBFFB0FDF4FAF1E325E05E" box="[633,770,1317,1341]" italics="true" pageId="6" pageNumber="49">Alburnoides</emphasis>
|
||
</taxonomicName>
|
||
sp. from the Amu Darya drainage (
|
||
<figureCitation id="1333B0EDFFCBFFB0FB1AFAF0E4FDE05F" box="[1175,1242,1316,1340]" captionStart="FIGURE 5" captionStartId="10.[151,250,676,699]" captionTargetBox="[156,1436,193,650]" captionTargetId="figure@10.[156,1441,193,651]" captionTargetPageId="10" captionText="FIGURE 5. Alburnoides sp., ZISP 10336, 65.8 mm SL, Samarkand (Zeravshan River system, Amu Darya drainage, Uzbekistan)." httpUri="https://zenodo.org/record/214794/files/figure.png" pageId="6" pageNumber="49">Fig. 5</figureCitation>
|
||
) by the vertebral counts (Tables 3, 4), the presence of a sharp, scaleless ventral keel, 6–8 gill rakers, and 4 pharyngeal teeth in the long row on the 5th ceratobranchial, but differs from it in having 13–16½, usually 14–15½, branched anal-fin rays (vs. 12–14½, usually 13½), and 46–51(55) lateral-line scales to posterior margin of hypurals (vs.
|
||
<quantity id="4CF0018DFFCBFFB0FB67FA47E569E0C8" box="[1258,1358,1427,1451]" metricMagnitude="0" metricUnit="m" metricValue="1.143" metricValueMax="1.1938" metricValueMin="1.0922" pageId="6" pageNumber="49" unit="in" value="45.0" valueMax="47.0" valueMin="43.0">43–47 in</quantity>
|
||
examined Amu Darya samples).
|
||
</paragraph>
|
||
<paragraph id="8BB7AC68FFCBFFB0FF44FA08E207E292" blockId="6.[151,1436,1316,2033]" pageId="6" pageNumber="49">
|
||
<taxonomicName id="4C08D7EBFFCBFFB0FF44FA08E180E097" ID-CoL="BGV6" box="[201,423,1500,1524]" class="Actinopterygii" family="Cyprinidae" genus="Alburnoides" kingdom="Animalia" order="Cypriniformes" pageId="6" pageNumber="49" phylum="Chordata" rank="species" species="holciki">
|
||
<emphasis id="B97C707AFFCBFFB0FF44FA08E180E097" box="[201,423,1500,1524]" italics="true" pageId="6" pageNumber="49">Alburnoides holciki</emphasis>
|
||
</taxonomicName>
|
||
is also similar to the Atrek riffle minnow in having a sharp, scaleless keel, an upturned, terminal mouth (
|
||
<figureCitation id="1333B0EDFFCBFFB0FED4FA2BE186E374" box="[345,417,1535,1559]" captionStart="FIGURE 6" captionStartId="10.[151,250,1193,1216]" captionTargetBox="[151,1435,779,1172]" captionTargetId="figure@10.[151,1436,779,1172]" captionTargetPageId="10" captionText="FIGURE 6. Alburnoides sp., ZISP 25785, 61.8 mm SL, Atrek River drainage (Turkmenistan)." httpUri="https://zenodo.org/record/214795/files/figure.png" pageId="6" pageNumber="49">Fig. 6</figureCitation>
|
||
), and 46–51(55) lateral-line scales to posterior margin of hypurals (
|
||
<quantity id="4CF0018DFFCBFFB0FB30FA2BE502E374" box="[1213,1317,1535,1559]" metricMagnitude="0" metricUnit="m" metricValue="1.2192" metricValueMax="1.2953999999999999" metricValueMin="1.143" pageId="6" pageNumber="49" unit="in" value="48.0" valueMax="51.0" valueMin="45.0">45–51 in</quantity>
|
||
the Atrek samples). However, the new species differs from the Atrek fish in usually having 4 pharyngeal teeth in the long row on the 5th ceratobranchial (vs. 5); 14–15½ branched anal-fin rays (vs. a mode of 13½); 6–8 gill rakers (vs. 9–10); 40–42, usually 41, total vertebrae (vs. 40); (12)13–14 predorsal vertebrae (vs. 12–13 with a mode of 12); a longer caudal vertebral region, containing (20)21–22 vertebrae (vs. 20–21 with a mode of 20); and 20+21 most usual vertebral formula (vs. 20+20).
|
||
<taxonomicName id="4C08D7EBFFCBFFB0FE79F962E25AE3AD" ID-CoL="RK5" box="[500,637,1718,1742]" class="Actinopterygii" family="Cyprinidae" genus="Alburnoides" kingdom="Animalia" order="Cypriniformes" pageId="6" pageNumber="49" phylum="Chordata" rank="genus">
|
||
<emphasis id="B97C707AFFCBFFB0FE79F962E25AE3AD" box="[500,637,1718,1742]" italics="true" pageId="6" pageNumber="49">Alburnoides</emphasis>
|
||
</taxonomicName>
|
||
sp. from the Atrek drainage is morphologically very close to the riffle minnow from the Tajan River by anal-fin and vertebral counts (
|
||
<figureCitation id="1333B0EDFFCBFFB0FCD5F90EE3BCE391" box="[856,923,1754,1778]" captionStart="FIGURE 4" captionStartId="6.[151,250,1224,1247]" captionTargetBox="[151,1426,387,1203]" captionTargetId="figure@6.[151,1436,373,1203]" captionTargetPageId="6" captionText="FIGURE 4. UPGMA dendrogram using Kullback-Leibler divergences for six meristic characters (the number of branched anal-fin rays and five vertebral counts)." httpUri="https://zenodo.org/record/214793/files/figure.png" pageId="6" pageNumber="49">Fig. 4</figureCitation>
|
||
, Tables 2–4), the number of lateral-line scales to posterior margin of hypurals (
|
||
<quantity id="4CF0018DFFCBFFB0FD83F92AE252E275" box="[526,629,1790,1814]" metricMagnitude="0" metricUnit="m" metricValue="1.2192" metricValueMax="1.2953999999999999" metricValueMin="1.143" pageId="6" pageNumber="49" unit="in" value="48.0" valueMax="51.0" valueMin="45.0">45–51 in</quantity>
|
||
the Atrek samples and
|
||
<quantity id="4CF0018DFFCBFFB0FC07F92AE3C8E275" box="[906,1007,1790,1814]" metricMagnitude="0" metricUnit="m" metricValue="1.2573" metricValueMax="1.3462" metricValueMin="1.1684" pageId="6" pageNumber="49" unit="in" value="49.5" valueMax="53.0" valueMin="46.0">46–53 in</quantity>
|
||
the Tajan sample), but differs by the shape of the mouth which is upturned with a pointed snout and a tip of the mouth cleft on a level with the upper half of the pupil (
|
||
<figureCitation id="1333B0EDFFCBFFB0FEAAF892E14CE23D" box="[295,363,1862,1886]" captionStart="FIGURE 6" captionStartId="10.[151,250,1193,1216]" captionTargetBox="[151,1435,779,1172]" captionTargetId="figure@10.[151,1436,779,1172]" captionTargetPageId="10" captionText="FIGURE 6. Alburnoides sp., ZISP 25785, 61.8 mm SL, Atrek River drainage (Turkmenistan)." httpUri="https://zenodo.org/record/214795/files/figure.png" pageId="6" pageNumber="49">Fig. 6</figureCitation>
|
||
) (vs. a more horizontal, slightly curved mouth with a rounded snout and a tip of the mouth on a level with the lower margin of the pupil,
|
||
<figureCitation id="1333B0EDFFCBFFB0FD09F8BEE2ECE2E1" box="[644,715,1898,1922]" captionStart="FIGURE 7" captionStartId="10.[151,250,1750,1773]" captionTargetBox="[151,1435,1265,1728]" captionTargetId="figure@10.[151,1436,1265,1729]" captionTargetPageId="10" captionText="FIGURE 7. Alburnoides sp., CMNFI 1979 - 0493, 82.8 mm SL, Tajan River (Iran)." httpUri="https://zenodo.org/record/214796/files/figure.png" pageId="6" pageNumber="49">Fig. 7</figureCitation>
|
||
), a sharp and commonly scaleless ventral keel (vs. sharp but variably scaled), and 5 pharyngeal teeth in the long row on the 5th ceratobranchial (vs. usually 4). This is why we do not consider these samples as conspecific though the Tajan and the Atrek are geographically very close flowing into the south-eastern Caspian Sea.
|
||
</paragraph>
|
||
<caption id="DF77FCE0FFC7FFBCFF1AFD70E0E8E7BA" httpUri="https://zenodo.org/record/214794/files/figure.png" pageId="10" pageNumber="53" targetBox="[156,1436,193,650]" targetPageId="10">
|
||
<paragraph id="8BB7AC68FFC7FFBCFF1AFD70E0E8E7BA" blockId="10.[151,1436,676,729]" pageId="10" pageNumber="53">
|
||
<emphasis id="B97C707AFFC7FFBCFF1AFD70E133E7D9" bold="true" box="[151,276,676,699]" pageId="10" pageNumber="53">FIGURE 5.</emphasis>
|
||
<taxonomicName id="4C08D7EBFFC7FFBCFE96FD71E1B0E7D8" ID-CoL="RK5" box="[283,407,677,699]" class="Actinopterygii" family="Cyprinidae" genus="Alburnoides" kingdom="Animalia" order="Cypriniformes" pageId="10" pageNumber="53" phylum="Chordata" rank="genus">
|
||
<emphasis id="B97C707AFFC7FFBCFE96FD71E1B0E7D8" box="[283,407,677,699]" italics="true" pageId="10" pageNumber="53">Alburnoides</emphasis>
|
||
</taxonomicName>
|
||
sp., ZISP 10336, 65.8 mm SL, Samarkand (Zeravshan River system, Amu Darya drainage, Uzbekistan).
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF77FCE0FFC7FFBCFF1AFB7DE472E1DC" box="[151,1109,1193,1216]" httpUri="https://zenodo.org/record/214795/files/figure.png" pageId="10" pageNumber="53" targetBox="[151,1435,779,1172]" targetPageId="10">
|
||
<paragraph id="8BB7AC68FFC7FFBCFF1AFB7DE472E1DC" blockId="10.[151,1109,1193,1216]" box="[151,1109,1193,1216]" pageId="10" pageNumber="53">
|
||
<emphasis id="B97C707AFFC7FFBCFF1AFB7DE135E1DC" bold="true" box="[151,274,1193,1216]" pageId="10" pageNumber="53">FIGURE 6.</emphasis>
|
||
<taxonomicName id="4C08D7EBFFC7FFBCFE94FB7DE1B2E1DC" ID-CoL="RK5" box="[281,405,1193,1215]" class="Actinopterygii" family="Cyprinidae" genus="Alburnoides" kingdom="Animalia" order="Cypriniformes" pageId="10" pageNumber="53" phylum="Chordata" rank="genus">
|
||
<emphasis id="B97C707AFFC7FFBCFE94FB7DE1B2E1DC" box="[281,405,1193,1215]" italics="true" pageId="10" pageNumber="53">Alburnoides</emphasis>
|
||
</taxonomicName>
|
||
sp., ZISP 25785, 61.8 mm SL, Atrek River drainage (Turkmenistan).
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF77FCE0FFC7FFBCFF1AF902E3FAE38E" box="[151,989,1750,1773]" httpUri="https://zenodo.org/record/214796/files/figure.png" pageId="10" pageNumber="53" targetBox="[151,1435,1265,1728]" targetPageId="10">
|
||
<paragraph id="8BB7AC68FFC7FFBCFF1AF902E3FAE38E" blockId="10.[151,989,1750,1773]" box="[151,989,1750,1773]" pageId="10" pageNumber="53">
|
||
<emphasis id="B97C707AFFC7FFBCFF1AF902E134E38F" bold="true" box="[151,275,1750,1773]" pageId="10" pageNumber="53">FIGURE 7.</emphasis>
|
||
<taxonomicName id="4C08D7EBFFC7FFBCFE94F903E1B2E38E" ID-CoL="RK5" box="[281,405,1751,1773]" class="Actinopterygii" family="Cyprinidae" genus="Alburnoides" kingdom="Animalia" order="Cypriniformes" pageId="10" pageNumber="53" phylum="Chordata" rank="genus">
|
||
<emphasis id="B97C707AFFC7FFBCFE94F903E1B2E38E" box="[281,405,1751,1773]" italics="true" pageId="10" pageNumber="53">Alburnoides</emphasis>
|
||
</taxonomicName>
|
||
sp., CMNFI 1979-0493, 82.8 mm SL, Tajan River (Iran).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BB7AC68FFC7FFBDFF44F8E7E0F5E45C" blockId="10.[151,1437,1843,2011]" lastBlockId="11.[151,1436,151,572]" lastPageId="11" lastPageNumber="54" pageId="10" pageNumber="53">
|
||
The new species is morphologically close to another geographically close species,
|
||
<taxonomicName id="4C08D7EBFFC7FFBCFBEEF8E7E13CE20C" ID-CoL="BGVV" authority="Bogutskaya & Coad, 2009" authorityName="Bogutskaya & Coad" authorityYear="2009" class="Actinopterygii" family="Cyprinidae" genus="Alburnoides" kingdom="Animalia" order="Cypriniformes" pageId="10" pageNumber="53" phylum="Chordata" rank="species" species="varentsovi">
|
||
<emphasis id="B97C707AFFC7FFBCFBEEF8E7E4D2E228" box="[1123,1269,1843,1867]" italics="true" pageId="10" pageNumber="53">A. varentsovi</emphasis>
|
||
Bogutskaya & Coad, 2009
|
||
</taxonomicName>
|
||
, in having a large eye (the orbit diameter larger than the snout length and about equal to the interorbital width), a triangular-shaped head, and a clearly forked caudal-fin with moderately pointed lobes. But
|
||
<taxonomicName id="4C08D7EBFFC7FFBCFA99F8AFE5A7E2F0" ID-CoL="BGV6" box="[1300,1408,1915,1939]" class="Actinopterygii" family="Cyprinidae" genus="Alburnoides" kingdom="Animalia" order="Cypriniformes" pageId="10" pageNumber="53" phylum="Chordata" rank="species" species="holciki">
|
||
<emphasis id="B97C707AFFC7FFBCFA99F8AFE5A7E2F0" box="[1300,1408,1915,1939]" italics="true" pageId="10" pageNumber="53">A. holciki</emphasis>
|
||
</taxonomicName>
|
||
is distinguishable from the latter species in having neither lower jaw nor upper jaw protruding relative to each other (vs. a lower jaw slightly protruding over the upper jaw), a sharp scaleless ventral keel (vs. smooth and partly scaleless), most often 2.5–4.2 pharyngeal teeth (vs. commonly 2.5–5.2), 13–16½, usually 14–15½, branched anal-fin rays (vs. 10–14½, usually 12–13½), 40–42, usually 41, total vertebrae (vs. 39–41(42) with a mode of 40), 12–14, with a mode of 13, predorsal vertebrae (vs. 11–13, with a mode of 12), and a caudal vertebral region longer than the abdominal region, most frequent vertebral formulae 20+21, 20+22 and 19+21 (vs. usually equal, 20+20) (Table 2–4).
|
||
</paragraph>
|
||
<paragraph id="8BB7AC68FFC6FFBDFF44FE98E1B3E75F" blockId="11.[151,1436,151,572]" pageId="11" pageNumber="54">
|
||
<taxonomicName id="4C08D7EBFFC6FFBDFF44FE98E181E407" ID-CoL="BGV6" box="[201,422,332,356]" class="Actinopterygii" family="Cyprinidae" genus="Alburnoides" kingdom="Animalia" order="Cypriniformes" pageId="11" pageNumber="54" phylum="Chordata" rank="species" species="holciki">
|
||
<emphasis id="B97C707AFFC6FFBDFF44FE98E181E407" box="[201,422,332,356]" italics="true" pageId="11" pageNumber="54">Alburnoides holciki</emphasis>
|
||
</taxonomicName>
|
||
is distinguishable from
|
||
<taxonomicName id="4C08D7EBFFC6FFBDFD36FE98E377E407" ID-CoL="66875" box="[699,848,332,356]" class="Actinopterygii" family="Cyprinidae" genus="Alburnoides" kingdom="Animalia" order="Cypriniformes" pageId="11" pageNumber="54" phylum="Chordata" rank="species" species="eichwaldii">
|
||
<emphasis id="B97C707AFFC6FFBDFD36FE98E377E407" box="[699,848,332,356]" italics="true" pageId="11" pageNumber="54">A. eichwaldii</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C08D7EBFFC6FFBDFC07FE98E434E407" ID-CoL="RK5" box="[906,1043,332,356]" class="Actinopterygii" family="Cyprinidae" genus="Alburnoides" kingdom="Animalia" order="Cypriniformes" pageId="11" pageNumber="54" phylum="Chordata" rank="genus">
|
||
<emphasis id="B97C707AFFC6FFBDFC07FE98E434E407" box="[906,1043,332,356]" italics="true" pageId="11" pageNumber="54">Alburnoides</emphasis>
|
||
</taxonomicName>
|
||
sp. from rivers in the south of the Talysh Mountains and the Safid River drainage (
|
||
<figureCitation id="1333B0EDFFC6FFBDFD31FEBBE2D8E4E4" box="[700,767,367,391]" captionStart="FIGURE 8" captionStartId="11.[151,250,1088,1111]" captionTargetBox="[151,1436,628,1067]" captionTargetId="figure@11.[151,1436,628,1068]" captionTargetPageId="11" captionText="FIGURE 8. Alburnoides sp., radiograph, CMNFI 1970 - 0537, 77.8 mm SL, Shah River (Safid River drainage, Iran). Shows a more horizontal mouth, a rounded upper profile of the snout, 12 predorsal vertebrae, 20 abdominal vertebrae, and 21 caudal vertebrae." httpUri="https://zenodo.org/record/214797/files/figure.png" pageId="11" pageNumber="54">Fig. 8</figureCitation>
|
||
) in having an upturned mouth with a tip of the mouth cleft on a level with the upper half of the pupil (vs. a more horizontal, slightly curved or straight but never upturned mouth, with a tip of the mouth cleft on a level with the middle of the eye or below), a pointed snout with a straight upper profile (vs. slightly to markedly rounded), a sharp scaleless keel (vs. smooth and variably scaled, often completely scaled), and usually 14–15½ branched anal-fin rays (vs. usually 12½ or 13½) Vertebral counts are summarised in Tables 3–4.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |