715 lines
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715 lines
80 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zse.96.48312" ID-GBIF-Dataset="7de3ea60-ba35-4831-ae1d-048dc362f8b0" ID-GBIF-Taxon="161939044" ID-Pensoft-Pub="1860-0743-1-25" ID-Pensoft-UUID="4EDA9B5B777551B7BBB0152DD188E5EF" ID-ZooBank="F2C8585A1268443693348B64AE20F6EE" ModsDocID="1860-0743-1-25" checkinTime="1580744411276" checkinUser="pensoft" docAuthor="Sands, Arthur F, Gloeer, Peter, Guerlek, Mustafa E, Albrecht, Christian & Neubauer, Thomas A" docDate="2020" docId="AF57794BB4205489A2DB32969731D52C" docLanguage="en" docName="ZoosystEvol 96(1): 25-66" docOrigin="Zoosystematics and Evolution 96 (1)" docSource="http://dx.doi.org/10.3897/zse.96.48312" docTitle="Theodoxus baeticus" docType="treatment" docVersion="6" id="4EDA9B5B777551B7BBB0152DD188E5EF" lastPageNumber="25" masterDocId="4EDA9B5B777551B7BBB0152DD188E5EF" masterDocTitle="A revision of the extant species of Theodoxus (Gastropoda, Neritidae) in Asia, with the description of three new species" masterLastPageNumber="66" masterPageNumber="25" pageId="0" pageNumber="25" updateTime="1635972787372" updateUser="ExternalLinkService">
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<mods:title>A revision of the extant species of Theodoxus (Gastropoda, Neritidae) in Asia, with the description of three new species</mods:title>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Sands, Arthur F</mods:namePart>
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<mods:affiliation>Justus Liebig Univrsity Giessen, Giessen, Germany</mods:affiliation>
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<mods:namePart>Gloeer, Peter</mods:namePart>
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<mods:affiliation>Biodiversity Research Laboratory, Hetlingen, Germany</mods:affiliation>
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<mods:name type="personal">
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Guerlek, Mustafa E</mods:namePart>
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<mods:affiliation>Mehmet Akif Ersoy niversity, Burdur, Turkey</mods:affiliation>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Albrecht, Christian</mods:namePart>
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<mods:affiliation>Justus Liebig University Giessen, Giessen, Germany</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Neubauer, Thomas A</mods:namePart>
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<mods:affiliation>Justus Liebig University Giessen, Giessen, Germany & Naturalis Biodiversity Centre, Leiden, Netherlands</mods:affiliation>
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<mods:title>Zoosystematics and Evolution</mods:title>
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<mods:part>
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<mods:date>2020</mods:date>
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<mods:number>96</mods:number>
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<mods:number>1</mods:number>
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<mods:start>25</mods:start>
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<mods:end>66</mods:end>
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<mods:url>http://dx.doi.org/10.3897/zse.96.48312</mods:url>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zse.96.48312</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1860-0743-1-25</mods:identifier>
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<treatment ID-GBIF-Taxon="161939044" LSID="urn:lsid:plazi:treatment:AF57794BB4205489A2DB32969731D52C" httpUri="http://treatment.plazi.org/id/AF57794BB4205489A2DB32969731D52C" lastPageNumber="25" pageId="0" pageNumber="25">
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<subSubSection pageId="0" pageNumber="25" type="nomenclature">
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<paragraph pageId="0" pageNumber="25">
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<taxonomicName LSID="AF57794B-B420-5489-A2DB-32969731D52C" authority="(Lamarck, 1822)" baseAuthorityName="Lamarck" baseAuthorityYear="1822" class="Gastropoda" family="Neritidae" genus="Theodoxus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Theodoxus baeticus" order="Cycloneritida" pageId="0" pageNumber="25" phylum="Mollusca" rank="species" species="baeticus">Theodoxus baeticus (Lamarck, 1822)</taxonomicName>
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<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Theodoxus baeticus (Lamarck, 1822). A - C. Paralectotype of N. baetica from Andalusia, Spain (MHNG-MOLL- 51319). D) Operculum of a paralectotype of N. baetica from Andalusia (MHNG-MOLL- 51319); E - G. Lectotype of N. baetica from Andalusia (MHNG-MOLL- 51319); H-J; K; L; M; N. Five syntypes of N. meridionalis from Sicily (ZMB, without coll. no.); O - R. Topotype of N. guadianensis collected in the Guadiana River near Mertola, Portugal incorporated into the phylogeny (UGSB 22084); S - V. Specimen from Soller, Balearic Islands used in the phylogeny (UGSB 19162). Scale bars: 1 mm." figureDoi="10.3897/zse.96.48312.figure6" httpUri="https://binary.pensoft.net/fig/376455" pageId="0" pageNumber="25">Figures 6A-V</figureCitation>
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<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Theodoxus baeticus (Lamarck, 1822). A - D. Specimen (T. callosus - morphoype) collected in Akyaka, Turkey (UGSB 19118); E - H. Specimen (T. callosus - morphotype) collected in Kato Tragana, Greece (UGSB 2488); I - L. Specimen (T. varius - morphotype) collected at the Blue Eye Nature Monument, Albania (UGSB 24170); M - P. Specimen (T. meridionalis - morphotype) collected at the Ketana Oasis, Gabes, Tunisia (UGSB 18111). All specimens, except A-H, were used in the phylogeny (Fig. 2). Scale bars: 1 mm." figureDoi="10.3897/zse.96.48312.figure7" httpUri="https://binary.pensoft.net/fig/376456" pageId="0" pageNumber="25">, 7A-P</figureCitation>
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<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Theodoxus baeticus (Lamarck, 1822). A - D. Specimen from Cuquillo, Granada, Andalusia, Spain (UGSB 22089); E - H. Specimen (T. meridionalis - morphotype) collected in the Palancia River, Navajas, Spain (UGSB 22090); I - L. Specimen (T. elongatulus - morphotype) from Arrancada, Pombal, Portugal (UGSB 22080); M - P. Specimen (T. velascoi - morphotype) from Montanejos, Castellon, Spain (UGSB 22082). Theodoxus cf. valentinus (Graells, 1846). Q - S. Specimen conforming to T. valentinus from the Verd River, Masalaves, Spain (UGSB 21787); T. Operculum of a specimen conforming to T. valentinus from the same locality as Q-S (UGSB 22087). All specimens, except Q-S, were used in the phylogeny (Fig. 2). Scale bars: 1 mm." figureDoi="10.3897/zse.96.48312.figure8" httpUri="https://binary.pensoft.net/fig/376457" pageId="0" pageNumber="25">, 8A-P</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="25" type="reference_group">
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<paragraph pageId="0" pageNumber="25">
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<taxonomicName authorityName="Lamarck" authorityYear="1822" class="Gastropoda" family="Neritidae" genus="Neritina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Neritina Baetica" order="Cycloneritida" pageId="0" pageNumber="25" phylum="Mollusca" rank="species" species="Baetica">Neritina Baetica</taxonomicName>
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<bibRefCitation author="Lamarck, J-BPAdMd" journalOrPublisher="Privately published, Paris" pageId="0" pageNumber="25" refId="B86" refString="Lamarck, J-BPAdMd, 1822. Histoire naturelle des animaux sans vertebres, presentant les caracteres generaux et particuliers de ces animaux, leur distribution, leurs classes, leurs familles, leurs genres, et la citation des principales especes qui s'y rapportent; precedee d'une introduction offrant la determination des caracteres essentiels de l'animal, sa distinction du vegetal et des autres corps naturels, enfin, l'exposition des principes fondomentaux de la zoologie. Tome sixieme, deuxieme partie. Privately published, Paris" title="Histoire naturelle des animaux sans vertebres, presentant les caracteres generaux et particuliers de ces animaux, leur distribution, leurs classes, leurs familles, leurs genres, et la citation des principales especes qui s'y rapportent; precedee d'une introduction offrant la determination des caracteres essentiels de l'animal, sa distinction du vegetal et des autres corps naturels, enfin, l'exposition des principes fondomentaux de la zoologie. Tome sixieme, deuxieme partie." year="1822">Lamarck 1822</bibRefCitation>
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: 188;
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<bibRefCitation author="Martens, Ev" journalOrPublisher="Bauer & Raspe, Nuernberg" pageId="0" pageNumber="25" publicationUrl="https://biodiversitylibrary.org/page/52484536" refId="B99" refString="Martens, Ev, 1879. Die Gattung Neritina. - Systematisches Conchylien-Cabinet von Martini und Chemnitz 2 (10). Bauer & Raspe, Nuernberg, https://biodiversitylibrary.org/page/52484536" title="Die Gattung Neritina. - Systematisches Conchylien-Cabinet von Martini und Chemnitz 2 (10)." url="https://biodiversitylibrary.org/page/52484536" year="1879">Martens 1879</bibRefCitation>
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: 234-235, pl. 23, figs 1, 2;
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<bibRefCitation author="Kobelt, W" journalOrPublisher="Aquila" pageId="0" pageNumber="25" publicationUrl="https://biodiversitylibrary.org/page/16299729" refId="B83" refString="Kobelt, W, 1899. Iconographie der Land- & Suesswasser-Mollusken mit vorzueglicher Beruecksichtigung der europaeischen noch nicht abgebildeten Arten von E. A. Rossmaessler. VIII. Band. C. W. Kreidel, Wiesbaden, 1-11. [pls 211-240] https://biodiversitylibrary.org/page/16299729" title="Iconographie der Land- & Suesswasser-Mollusken mit vorzueglicher Beruecksichtigung der europaeischen noch nicht abgebildeten Arten von E. A. Rossmaessler. VIII. Band. C. W. Kreidel, Wiesbaden, 1 - 11. [pls 211 - 240]" url="https://biodiversitylibrary.org/page/16299729" year="1899">Kobelt 1899</bibRefCitation>
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: 13-14, pl. 214, figs 1350, 1356.
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</paragraph>
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<paragraph pageId="0" pageNumber="25">
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<taxonomicName class="Gastropoda" family="Neritidae" genus="Neritina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Neritina varia" order="Cycloneritida" pageId="0" pageNumber="25" phylum="Mollusca" rank="species" species="varia">Neritina varia</taxonomicName>
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:
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<bibRefCitation author="Menke, CT" journalOrPublisher="Gelpke, Pyrmonti" pageId="0" pageNumber="25" publicationUrl="https://biodiversitylibrary.org/page/13119590" refId="B100" refString="Menke, CT, 1828. Synopsis methodica molluscorum generum omnium et specierum earum, quae in museo Menkeano adservantur; cum synonymia critica et novarum specierum diagnosibus. Gelpke, Pyrmonti, https://biodiversitylibrary.org/page/13119590" title="Synopsis methodica molluscorum generum omnium et specierum earum, quae in museo Menkeano adservantur; cum synonymia critica et novarum specierum diagnosibus." url="https://biodiversitylibrary.org/page/13119590" year="1828">Menke 1828</bibRefCitation>
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: 27 (nomen nudum).
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</paragraph>
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<paragraph pageId="0" pageNumber="25">
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<taxonomicName authorityName="Deshayes" authorityYear="1835" class="Gastropoda" family="Neritidae" genus="Neritina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Neritina callosa" order="Cycloneritida" pageId="0" pageNumber="25" phylum="Mollusca" rank="species" species="callosa">Neritina callosa</taxonomicName>
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Deshayes in
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<bibRefCitation author="Geoffroy Saint-Hilaire, I" journalOrPublisher="Levrault, Paris" pageId="0" pageNumber="25" publicationUrl="http://reader.digitale-sammlungen.de/de/fs3/object/display/bsb10360027_00005.html" refId="B58" refString="Geoffroy Saint-Hilaire, I, Geoffroy Saint-Hilaire, E, Deshayes, GP, Bibron, G, Bory de Saint-Vincent, J-B, 1832-1836. Expedition scientifique de Moree, Tome III - 1.re Partie. Zoologie. Premiere Section. - Animaux vertebres, Mollusques et Polypiers. F. G. Levrault, Paris, http://reader.digitale-sammlungen.de/de/fs3/object/display/bsb10360027_00005.html" title="Expedition scientifique de Moree, Tome III - 1. re Partie. Zoologie. Premiere Section. - Animaux vertebres, Mollusques et Polypiers. F. G." url="http://reader.digitale-sammlungen.de/de/fs3/object/display/bsb10360027_00005.html" year="1832 - 1836">Geoffroy Saint-Hilaire et al. 1835</bibRefCitation>
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: 156, pl. 19, figs 16-18;
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<bibRefCitation author="Martens, Ev" journalOrPublisher="Bauer & Raspe, Nuernberg" pageId="0" pageNumber="25" publicationUrl="https://biodiversitylibrary.org/page/52484536" refId="B99" refString="Martens, Ev, 1879. Die Gattung Neritina. - Systematisches Conchylien-Cabinet von Martini und Chemnitz 2 (10). Bauer & Raspe, Nuernberg, https://biodiversitylibrary.org/page/52484536" title="Die Gattung Neritina. - Systematisches Conchylien-Cabinet von Martini und Chemnitz 2 (10)." url="https://biodiversitylibrary.org/page/52484536" year="1879">Martens 1879</bibRefCitation>
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: 232-233, pl. 22, figs 27-30;
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<bibRefCitation author="Kobelt, W" journalOrPublisher="Aquila" pageId="0" pageNumber="25" publicationUrl="https://biodiversitylibrary.org/page/16299729" refId="B83" refString="Kobelt, W, 1899. Iconographie der Land- & Suesswasser-Mollusken mit vorzueglicher Beruecksichtigung der europaeischen noch nicht abgebildeten Arten von E. A. Rossmaessler. VIII. Band. C. W. Kreidel, Wiesbaden, 1-11. [pls 211-240] https://biodiversitylibrary.org/page/16299729" title="Iconographie der Land- & Suesswasser-Mollusken mit vorzueglicher Beruecksichtigung der europaeischen noch nicht abgebildeten Arten von E. A. Rossmaessler. VIII. Band. C. W. Kreidel, Wiesbaden, 1 - 11. [pls 211 - 240]" url="https://biodiversitylibrary.org/page/16299729" year="1899">Kobelt 1899</bibRefCitation>
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: 11-12, pl. 213, figs 1345, 1346.
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</paragraph>
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<paragraph pageId="0" pageNumber="25">
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<taxonomicName class="Gastropoda" family="Neritidae" genus="Neritina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Neritina varia" order="Cycloneritida" pageId="0" pageNumber="25" phylum="Mollusca" rank="species" species="varia">Neritina varia</taxonomicName>
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<normalizedToken originalValue="Rossmässler">Rossmaessler</normalizedToken>
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1835: 18;
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<bibRefCitation author="Martens, Ev" journalOrPublisher="Bauer & Raspe, Nuernberg" pageId="0" pageNumber="25" publicationUrl="https://biodiversitylibrary.org/page/52484536" refId="B99" refString="Martens, Ev, 1879. Die Gattung Neritina. - Systematisches Conchylien-Cabinet von Martini und Chemnitz 2 (10). Bauer & Raspe, Nuernberg, https://biodiversitylibrary.org/page/52484536" title="Die Gattung Neritina. - Systematisches Conchylien-Cabinet von Martini und Chemnitz 2 (10)." url="https://biodiversitylibrary.org/page/52484536" year="1879">Martens 1879</bibRefCitation>
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: 225-226, pl. 21, figs 17-19;
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<bibRefCitation author="Kobelt, W" journalOrPublisher="Aquila" pageId="0" pageNumber="25" publicationUrl="https://biodiversitylibrary.org/page/16299729" refId="B83" refString="Kobelt, W, 1899. Iconographie der Land- & Suesswasser-Mollusken mit vorzueglicher Beruecksichtigung der europaeischen noch nicht abgebildeten Arten von E. A. Rossmaessler. VIII. Band. C. W. Kreidel, Wiesbaden, 1-11. [pls 211-240] https://biodiversitylibrary.org/page/16299729" title="Iconographie der Land- & Suesswasser-Mollusken mit vorzueglicher Beruecksichtigung der europaeischen noch nicht abgebildeten Arten von E. A. Rossmaessler. VIII. Band. C. W. Kreidel, Wiesbaden, 1 - 11. [pls 211 - 240]" url="https://biodiversitylibrary.org/page/16299729" year="1899">Kobelt 1899</bibRefCitation>
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: 9, pl. 213, figs 1338.
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</paragraph>
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<paragraph pageId="0" pageNumber="25">
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<taxonomicName authorityName="Philippi" authorityYear="1836" class="Gastropoda" family="Neritidae" genus="Nerita" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Nerita meridionalis" order="Cycloneritida" pageId="0" pageNumber="25" phylum="Mollusca" rank="species" species="meridionalis">Nerita meridionalis</taxonomicName>
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<bibRefCitation author="Philippi, RA" journalOrPublisher="Simon Schropp, Berlin" pageId="0" pageNumber="25" publicationUrl="http://reader.digitale-sammlungen.de/de/fs2/object/display/bsb10231737_00007.html" refId="B121" refString="Philippi, RA, 1836. Enumeratio Molluscorum Siciliae cum viventium tum in tellure teriaria fossilium, quae in itinere suo observavit. Vol. 1. Simon Schropp, Berlin, http://reader.digitale-sammlungen.de/de/fs2/object/display/bsb10231737_00007.html" title="Enumeratio Molluscorum Siciliae cum viventium tum in tellure teriaria fossilium, quae in itinere suo observavit. Vol. 1." url="http://reader.digitale-sammlungen.de/de/fs2/object/display/bsb10231737_00007.html" year="1836">Philippi 1836</bibRefCitation>
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: 159-160, pl. 9, fig. 13.
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</paragraph>
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<paragraph pageId="0" pageNumber="25">
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<taxonomicName class="Gastropoda" family="Neritidae" genus="Nerita" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Nerita Philippii" order="Cycloneritida" pageId="0" pageNumber="25" phylum="Mollusca" rank="species" species="Philippii">Nerita Philippii</taxonomicName>
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<bibRefCitation author="Recluz, CA" journalOrPublisher="Systematic Biology" pageId="0" pageNumber="25" publicationUrl="https://biodiversitylibrary.org/page/2361985" refId="B131" refString="Recluz, CA, 1841. Description de quelques especes de Nerites vivantes. (2e partie, suite et fin.). Revue Zoologique par la Societe Cuvierienne 1841(4): 102-109; 1841(5): 147-152; 1841(9): 273-278; 1841(10): 310-318; 1841(11): 337-343. https://biodiversitylibrary.org/page/2361985" title="Description de quelques especes de Nerites vivantes. (2 e partie, suite et fin.). Revue Zoologique par la Societe Cuvierienne 1841 (4): 102 - 109; 1841 (5): 147 - 152; 1841 (9): 273 - 278; 1841 (10): 310 - 318; 1841 (11): 337 - 343." url="https://biodiversitylibrary.org/page/2361985" year="1841">
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<normalizedToken originalValue="Récluz">Recluz</normalizedToken>
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1841
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</bibRefCitation>
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: 341-342.
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</paragraph>
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<paragraph pageId="0" pageNumber="25">
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<taxonomicName authorityName="Morelet" authorityYear="1845" class="Gastropoda" family="Neritidae" genus="Neritina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Neritina elongatula" order="Cycloneritida" pageId="0" pageNumber="25" phylum="Mollusca" rank="species" species="elongatula">Neritina elongatula</taxonomicName>
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<bibRefCitation author="Morelet, A" journalOrPublisher="Bailliere, Paris" pageId="0" pageNumber="25" publicationUrl="https://biodiversitylibrary.org/page/12988736" refId="B108" refString="Morelet, A, 1845. Description des mollusques terrestres et fluviatiles du Portugal. Bailliere, Paris, https://biodiversitylibrary.org/page/12988736" title="Description des mollusques terrestres et fluviatiles du Portugal." url="https://biodiversitylibrary.org/page/12988736" year="1845">Morelet 1845</bibRefCitation>
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: 96-97, pl. 9, fig. 4;
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<bibRefCitation author="Martens, Ev" journalOrPublisher="Bauer & Raspe, Nuernberg" pageId="0" pageNumber="25" publicationUrl="https://biodiversitylibrary.org/page/52484536" refId="B99" refString="Martens, Ev, 1879. Die Gattung Neritina. - Systematisches Conchylien-Cabinet von Martini und Chemnitz 2 (10). Bauer & Raspe, Nuernberg, https://biodiversitylibrary.org/page/52484536" title="Die Gattung Neritina. - Systematisches Conchylien-Cabinet von Martini und Chemnitz 2 (10)." url="https://biodiversitylibrary.org/page/52484536" year="1879">Martens 1879</bibRefCitation>
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: 228-229, pl. 22, figs 16-19;
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<bibRefCitation author="Kobelt, W" journalOrPublisher="Aquila" pageId="0" pageNumber="25" publicationUrl="https://biodiversitylibrary.org/page/16299729" refId="B83" refString="Kobelt, W, 1899. Iconographie der Land- & Suesswasser-Mollusken mit vorzueglicher Beruecksichtigung der europaeischen noch nicht abgebildeten Arten von E. A. Rossmaessler. VIII. Band. C. W. Kreidel, Wiesbaden, 1-11. [pls 211-240] https://biodiversitylibrary.org/page/16299729" title="Iconographie der Land- & Suesswasser-Mollusken mit vorzueglicher Beruecksichtigung der europaeischen noch nicht abgebildeten Arten von E. A. Rossmaessler. VIII. Band. C. W. Kreidel, Wiesbaden, 1 - 11. [pls 211 - 240]" url="https://biodiversitylibrary.org/page/16299729" year="1899">Kobelt 1899</bibRefCitation>
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: 10-11, pl. 213, figs 1341, 1342.
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||
</paragraph>
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<paragraph pageId="0" pageNumber="25">
|
||
<taxonomicName authorityName="Morelet" authorityYear="1845" class="Gastropoda" family="Neritidae" genus="Neritina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Neritina inquinata" order="Cycloneritida" pageId="0" pageNumber="25" phylum="Mollusca" rank="species" species="inquinata">Neritina inquinata</taxonomicName>
|
||
<bibRefCitation author="Morelet, A" journalOrPublisher="Bailliere, Paris" pageId="0" pageNumber="25" publicationUrl="https://biodiversitylibrary.org/page/12988736" refId="B108" refString="Morelet, A, 1845. Description des mollusques terrestres et fluviatiles du Portugal. Bailliere, Paris, https://biodiversitylibrary.org/page/12988736" title="Description des mollusques terrestres et fluviatiles du Portugal." url="https://biodiversitylibrary.org/page/12988736" year="1845">Morelet 1845</bibRefCitation>
|
||
: 93-94, pl. 9, fig. 2.
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="25">
|
||
<taxonomicName authorityName="Morelet" authorityYear="1845" class="Gastropoda" family="Neritidae" genus="Neritina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Neritina guadianensis" order="Cycloneritida" pageId="0" pageNumber="25" phylum="Mollusca" rank="species" species="guadianensis">Neritina guadianensis</taxonomicName>
|
||
<bibRefCitation author="Morelet, A" journalOrPublisher="Bailliere, Paris" pageId="0" pageNumber="25" publicationUrl="https://biodiversitylibrary.org/page/12988736" refId="B108" refString="Morelet, A, 1845. Description des mollusques terrestres et fluviatiles du Portugal. Bailliere, Paris, https://biodiversitylibrary.org/page/12988736" title="Description des mollusques terrestres et fluviatiles du Portugal." url="https://biodiversitylibrary.org/page/12988736" year="1845">Morelet 1845</bibRefCitation>
|
||
: 95-96, pl. 9, fig. 3;
|
||
<bibRefCitation author="Martens, Ev" journalOrPublisher="Bauer & Raspe, Nuernberg" pageId="0" pageNumber="25" publicationUrl="https://biodiversitylibrary.org/page/52484536" refId="B99" refString="Martens, Ev, 1879. Die Gattung Neritina. - Systematisches Conchylien-Cabinet von Martini und Chemnitz 2 (10). Bauer & Raspe, Nuernberg, https://biodiversitylibrary.org/page/52484536" title="Die Gattung Neritina. - Systematisches Conchylien-Cabinet von Martini und Chemnitz 2 (10)." url="https://biodiversitylibrary.org/page/52484536" year="1879">Martens 1879</bibRefCitation>
|
||
: 231-232, pl. 22, figs 22-26;
|
||
<bibRefCitation author="Kobelt, W" journalOrPublisher="Aquila" pageId="0" pageNumber="25" publicationUrl="https://biodiversitylibrary.org/page/16299729" refId="B83" refString="Kobelt, W, 1899. Iconographie der Land- & Suesswasser-Mollusken mit vorzueglicher Beruecksichtigung der europaeischen noch nicht abgebildeten Arten von E. A. Rossmaessler. VIII. Band. C. W. Kreidel, Wiesbaden, 1-11. [pls 211-240] https://biodiversitylibrary.org/page/16299729" title="Iconographie der Land- & Suesswasser-Mollusken mit vorzueglicher Beruecksichtigung der europaeischen noch nicht abgebildeten Arten von E. A. Rossmaessler. VIII. Band. C. W. Kreidel, Wiesbaden, 1 - 11. [pls 211 - 240]" url="https://biodiversitylibrary.org/page/16299729" year="1899">Kobelt 1899</bibRefCitation>
|
||
: 16, pl. 215, figs 1357, 1358.
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="25">
|
||
<taxonomicName baseAuthorityName="Gmelin" baseAuthorityYear="1791" class="Gastropoda" family="Neritidae" genus="Neritina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Neritina violacea" order="Cycloneritida" pageId="0" pageNumber="25" phylum="Mollusca" rank="species" species="violacea">Neritina violacea</taxonomicName>
|
||
<bibRefCitation author="Morelet, A" journalOrPublisher="Bailliere, Paris" pageId="0" pageNumber="25" publicationUrl="https://biodiversitylibrary.org/page/12988736" refId="B108" refString="Morelet, A, 1845. Description des mollusques terrestres et fluviatiles du Portugal. Bailliere, Paris, https://biodiversitylibrary.org/page/12988736" title="Description des mollusques terrestres et fluviatiles du Portugal." url="https://biodiversitylibrary.org/page/12988736" year="1845">Morelet 1845</bibRefCitation>
|
||
: 92-93, pl. 9, fig. 1.
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="25">
|
||
<taxonomicName class="Gastropoda" family="Neritidae" genus="Neritina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Neritina" order="Cycloneritida" pageId="0" pageNumber="25" phylum="Mollusca" rank="genus">Neritina</taxonomicName>
|
||
Velascoi
|
||
<bibRefCitation author="Graells, MdlP" journalOrPublisher="Viuda e Hijos de Don Antonia Calleja; Calleja, Ojea y Compania, Madrid, Lima" pageId="0" pageNumber="25" publicationUrl="https://biodiversitylibrary.org/page/14891373" refId="B66" refString="Graells, MdlP, 1846. Catalogo de los moluscos terrestres y de agua dulce observado en Espana, y descripcion y notas de algunas especies nuevas o poco conocidas del mismo pais. Viuda e Hijos de Don Antonia Calleja; Calleja, Ojea y Compania, Madrid, Lima, https://biodiversitylibrary.org/page/14891373" title="Catalogo de los moluscos terrestres y de agua dulce observado en Espana, y descripcion y notas de algunas especies nuevas o poco conocidas del mismo pais." url="https://biodiversitylibrary.org/page/14891373" year="1846">Graells 1846</bibRefCitation>
|
||
: 20-21, unnumbered plate, figs 25-30.
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="25">
|
||
<taxonomicName class="Gastropoda" family="Neritidae" genus="Neritina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Neritina Anatensis" order="Cycloneritida" pageId="0" pageNumber="25" phylum="Mollusca" rank="species" species="Anatensis">Neritina Anatensis</taxonomicName>
|
||
G.B. Sowerby
|
||
<date value="1849-02">II 1849</date>
|
||
: 535, pl. 116, figs 247, 248.
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="25">
|
||
<taxonomicName class="Gastropoda" family="Neritidae" genus="Neritina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Neritina meridionalis" order="Cycloneritida" pageId="0" pageNumber="25" phylum="Mollusca" rank="species" species="meridionalis">Neritina meridionalis</taxonomicName>
|
||
:
|
||
<bibRefCitation author="Martens, Ev" journalOrPublisher="Bauer & Raspe, Nuernberg" pageId="0" pageNumber="25" publicationUrl="https://biodiversitylibrary.org/page/52484536" refId="B99" refString="Martens, Ev, 1879. Die Gattung Neritina. - Systematisches Conchylien-Cabinet von Martini und Chemnitz 2 (10). Bauer & Raspe, Nuernberg, https://biodiversitylibrary.org/page/52484536" title="Die Gattung Neritina. - Systematisches Conchylien-Cabinet von Martini und Chemnitz 2 (10)." url="https://biodiversitylibrary.org/page/52484536" year="1879">Martens 1879</bibRefCitation>
|
||
: 227-228, pl. 4, figs 27-29, pl. 22, figs 11-15;
|
||
<bibRefCitation author="Kobelt, W" journalOrPublisher="Aquila" pageId="0" pageNumber="25" publicationUrl="https://biodiversitylibrary.org/page/16299729" refId="B83" refString="Kobelt, W, 1899. Iconographie der Land- & Suesswasser-Mollusken mit vorzueglicher Beruecksichtigung der europaeischen noch nicht abgebildeten Arten von E. A. Rossmaessler. VIII. Band. C. W. Kreidel, Wiesbaden, 1-11. [pls 211-240] https://biodiversitylibrary.org/page/16299729" title="Iconographie der Land- & Suesswasser-Mollusken mit vorzueglicher Beruecksichtigung der europaeischen noch nicht abgebildeten Arten von E. A. Rossmaessler. VIII. Band. C. W. Kreidel, Wiesbaden, 1 - 11. [pls 211 - 240]" url="https://biodiversitylibrary.org/page/16299729" year="1899">Kobelt 1899</bibRefCitation>
|
||
: 9-10, pl. 213, figs 1339, 1340.
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="25">
|
||
<taxonomicName authorityName="Crosse" authorityYear="1880" class="Gastropoda" family="Neritidae" genus="Neritina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Neritina Hidalgoi" order="Cycloneritida" pageId="0" pageNumber="25" phylum="Mollusca" rank="species" species="Hidalgoi">Neritina Hidalgoi</taxonomicName>
|
||
<bibRefCitation author="Crosse, H" journalOrPublisher="Journal de Conchyliologie" pageId="0" pageNumber="25" pagination="320 - 322" publicationUrl="https://biodiversitylibrary.org/page/15919433" refId="B37" refString="Crosse, H, 1880. Description d'un nouvelle espece de Neritina d'Espagne. Journal de Conchyliologie 28: 320 - 322, https://biodiversitylibrary.org/page/15919433" title="Description d'un nouvelle espece de Neritina d'Espagne." url="https://biodiversitylibrary.org/page/15919433" volume="28" year="1880">Crosse 1880</bibRefCitation>
|
||
: 320-322;
|
||
<bibRefCitation author="Kobelt, W" journalOrPublisher="Aquila" pageId="0" pageNumber="25" publicationUrl="https://biodiversitylibrary.org/page/16299729" refId="B83" refString="Kobelt, W, 1899. Iconographie der Land- & Suesswasser-Mollusken mit vorzueglicher Beruecksichtigung der europaeischen noch nicht abgebildeten Arten von E. A. Rossmaessler. VIII. Band. C. W. Kreidel, Wiesbaden, 1-11. [pls 211-240] https://biodiversitylibrary.org/page/16299729" title="Iconographie der Land- & Suesswasser-Mollusken mit vorzueglicher Beruecksichtigung der europaeischen noch nicht abgebildeten Arten von E. A. Rossmaessler. VIII. Band. C. W. Kreidel, Wiesbaden, 1 - 11. [pls 211 - 240]" url="https://biodiversitylibrary.org/page/16299729" year="1899">Kobelt 1899</bibRefCitation>
|
||
: 12, pl. 214, fig. 1347.
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="25">
|
||
<taxonomicName authority="callosus" class="Gastropoda" family="Neritidae" genus="Theodoxus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Theodoxus varius" order="Cycloneritida" pageId="0" pageNumber="25" phylum="Mollusca" rank="species" species="varius">Theodoxus (Neritaea) varius callosus</taxonomicName>
|
||
:
|
||
<bibRefCitation author="Bank, RA" journalOrPublisher="Heldia" pageId="0" pageNumber="25" pagination="51 - 86" refId="B21" refString="Bank, RA, 2006. Towards a catalogue and bibliography of the freshwater Mollusca of Greece. Heldia 6 (1): 51 - 86" title="Towards a catalogue and bibliography of the freshwater Mollusca of Greece." volume="6" year="2006">Bank 2006</bibRefCitation>
|
||
: 52.
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="25">
|
||
<taxonomicName authority="varius" class="Gastropoda" family="Neritidae" genus="Theodoxus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Theodoxus varius" order="Cycloneritida" pageId="0" pageNumber="25" phylum="Mollusca" rank="species" species="varius">Theodoxus (Neritaea) varius varius</taxonomicName>
|
||
:
|
||
<bibRefCitation author="Bank, RA" journalOrPublisher="Heldia" pageId="0" pageNumber="25" pagination="51 - 86" refId="B21" refString="Bank, RA, 2006. Towards a catalogue and bibliography of the freshwater Mollusca of Greece. Heldia 6 (1): 51 - 86" title="Towards a catalogue and bibliography of the freshwater Mollusca of Greece." volume="6" year="2006">Bank 2006</bibRefCitation>
|
||
: 52.
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="25">
|
||
<taxonomicName baseAuthorityName="Lamarck" baseAuthorityYear="1822" class="Gastropoda" family="Neritidae" genus="Theodoxus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Theodoxus baeticus" order="Cycloneritida" pageId="0" pageNumber="25" phylum="Mollusca" rank="species" species="baeticus">Theodoxus baeticus</taxonomicName>
|
||
:
|
||
<bibRefCitation author="Gloeer, P" journalOrPublisher="Ecologica Montenegrina" pageId="0" pageNumber="25" pagination="133 - 137" refId="B61" refString="Gloeer, P, 2018. On the identity of Neritina baetica Lamarck, 1822 and Nerita meridionalis Philippi, 1836 (Gastropoda: Neritidae) from the Iberian Peninsula. Ecologica Montenegrina 18: 133 - 137" title="On the identity of Neritina baetica Lamarck, 1822 and Nerita meridionalis Philippi, 1836 (Gastropoda: Neritidae) from the Iberian Peninsula." volume="18" year="2018">
|
||
<normalizedToken originalValue="Glöer">Gloeer</normalizedToken>
|
||
2018
|
||
</bibRefCitation>
|
||
: 135-136, figs 5-16;
|
||
<bibRefCitation author="Gloeer, P" journalOrPublisher="Peter Gloeer, Hetlingen, Germany" pageId="0" pageNumber="25" refId="B62" refString="Gloeer, P, 2019. The freshwater gastropods of the West-Palaearctis. Vol. 1. Fresh- and brackish waters except spring and subterranean snails. Peter Gloeer, Hetlingen, Germany" title="The freshwater gastropods of the West-Palaearctis. Vol. 1. Fresh- and brackish waters except spring and subterranean snails." year="2019">
|
||
<normalizedToken originalValue="Glöer">Gloeer</normalizedToken>
|
||
2019
|
||
</bibRefCitation>
|
||
: 38-39, figs 21, 22.
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="25">
|
||
<taxonomicName baseAuthorityName="Morelet" baseAuthorityYear="1854" class="Gastropoda" family="Neritidae" genus="Theodoxus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Theodoxus elongatulus" order="Cycloneritida" pageId="0" pageNumber="25" phylum="Mollusca" rank="species" species="elongatulus">Theodoxus elongatulus</taxonomicName>
|
||
:
|
||
<bibRefCitation author="Welter-Schultes, FW" journalOrPublisher="Planet Poster Editions, Goettingen" pageId="0" pageNumber="25" refId="B158" refString="Welter-Schultes, FW, 2012. European non-marine molluscs, a guide for species identification. Planet Poster Editions, Goettingen" title="European non-marine molluscs, a guide for species identification." year="2012">Welter-Schultes 2012</bibRefCitation>
|
||
: 27, unnumbered text figures.
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="25">
|
||
<taxonomicName baseAuthorityName="Philippi" baseAuthorityYear="1836" class="Gastropoda" family="Neritidae" genus="Theodoxus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Theodoxus meridionalis" order="Cycloneritida" pageId="0" pageNumber="25" phylum="Mollusca" rank="species" species="meridionalis">Theodoxus meridionalis</taxonomicName>
|
||
:
|
||
<bibRefCitation author="Alba, DM" journalOrPublisher="Spira" pageId="0" pageNumber="25" pagination="41 - 65" refId="B6" refString="Alba, DM, Corbella, J, Guillen, G, Prats, L, Tarruella, A, 2016. Presence of two different species of Theodoxus Montfort, 1810 (Gastropoda: Neritidae) in Catalonia. Spira 6: 41 - 65" title="Presence of two different species of Theodoxus Montfort, 1810 (Gastropoda: Neritidae) in Catalonia." volume="6" year="2016">Alba et al. 2016</bibRefCitation>
|
||
: 48, figs 4, 9;
|
||
<bibRefCitation author="Gloeer, P" journalOrPublisher="Peter Gloeer, Hetlingen, Germany" pageId="0" pageNumber="25" refId="B62" refString="Gloeer, P, 2019. The freshwater gastropods of the West-Palaearctis. Vol. 1. Fresh- and brackish waters except spring and subterranean snails. Peter Gloeer, Hetlingen, Germany" title="The freshwater gastropods of the West-Palaearctis. Vol. 1. Fresh- and brackish waters except spring and subterranean snails." year="2019">
|
||
<normalizedToken originalValue="Glöer">Gloeer</normalizedToken>
|
||
2019
|
||
</bibRefCitation>
|
||
: 46-47, fig. 33.
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="25">
|
||
<taxonomicName baseAuthorityName="Graells" baseAuthorityYear="1846" class="Gastropoda" family="Neritidae" genus="Theodoxus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Theodoxus velascoi" order="Cycloneritida" pageId="0" pageNumber="25" phylum="Mollusca" rank="species" species="velascoi">Theodoxus velascoi</taxonomicName>
|
||
:
|
||
<bibRefCitation author="Welter-Schultes, FW" journalOrPublisher="Planet Poster Editions, Goettingen" pageId="0" pageNumber="25" refId="B158" refString="Welter-Schultes, FW, 2012. European non-marine molluscs, a guide for species identification. Planet Poster Editions, Goettingen" title="European non-marine molluscs, a guide for species identification." year="2012">Welter-Schultes 2012</bibRefCitation>
|
||
: 30, unnumbered text figures.
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="25">
|
||
<taxonomicName baseAuthorityName="Philippi" baseAuthorityYear="1836" class="Gastropoda" family="Neritidae" genus="Theodoxus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Theodoxus meridionalis" order="Cycloneritida" pageId="0" pageNumber="25" phylum="Mollusca" rank="species" species="meridionalis">Theodoxus (Theodoxus) cf. meridionalis</taxonomicName>
|
||
:
|
||
<bibRefCitation author="Alba, DM" journalOrPublisher="Spira" pageId="0" pageNumber="25" pagination="41 - 65" refId="B6" refString="Alba, DM, Corbella, J, Guillen, G, Prats, L, Tarruella, A, 2016. Presence of two different species of Theodoxus Montfort, 1810 (Gastropoda: Neritidae) in Catalonia. Spira 6: 41 - 65" title="Presence of two different species of Theodoxus Montfort, 1810 (Gastropoda: Neritidae) in Catalonia." volume="6" year="2016">Alba et al. 2016</bibRefCitation>
|
||
: 44-52, figs 2, 3, 5.
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="25">
|
||
<taxonomicName baseAuthorityName="Deshayes" baseAuthorityYear="1835" class="Gastropoda" family="Neritidae" genus="Theodoxus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Theodoxus callosus" order="Cycloneritida" pageId="0" pageNumber="25" phylum="Mollusca" rank="species" species="callosus">Theodoxus callosus</taxonomicName>
|
||
:
|
||
<bibRefCitation author="Gloeer, P" journalOrPublisher="Peter Gloeer, Hetlingen, Germany" pageId="0" pageNumber="25" refId="B62" refString="Gloeer, P, 2019. The freshwater gastropods of the West-Palaearctis. Vol. 1. Fresh- and brackish waters except spring and subterranean snails. Peter Gloeer, Hetlingen, Germany" title="The freshwater gastropods of the West-Palaearctis. Vol. 1. Fresh- and brackish waters except spring and subterranean snails." year="2019">
|
||
<normalizedToken originalValue="Glöer">Gloeer</normalizedToken>
|
||
2019
|
||
</bibRefCitation>
|
||
: 39-40, fig. 23.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="25" type="type locality">
|
||
<paragraph pageId="0" pageNumber="25">
|
||
<typeStatus>Type</typeStatus>
|
||
locality.
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="25">Freshwaters of Andalusia, Spain (no precise locality given).</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="25" type="type material">
|
||
<paragraph pageId="0" pageNumber="25">
|
||
<typeStatus>Type</typeStatus>
|
||
material.
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="25">
|
||
<materialsCitation collectionCode="MHNG" country="Turkey" typeStatus="syntypes">
|
||
Two
|
||
<typeStatus>syntypes</typeStatus>
|
||
of
|
||
<taxonomicName genus="N." lsidName="N. baetica" pageId="0" pageNumber="25" rank="species" species="baetica">
|
||
<emphasis italics="true" pageId="0" pageNumber="25">N. baetica</emphasis>
|
||
</taxonomicName>
|
||
are stored in
|
||
<collectionCode collectionName="Switzerland, Geneva, Museum d'Histoire Naturelle" country="Switzerland" httpUri="http://grbio.org/cool/2jyg-xmd2" name="Museum d'Histoire Naturelle" type="Museum">MHNG</collectionCode>
|
||
(coll. no. MHNG-MOLL-51319)
|
||
</materialsCitation>
|
||
.
|
||
<materialsCitation country="Turkey" typeStatus="syntype">
|
||
We designate here the
|
||
<typeStatus>syntype</typeStatus>
|
||
illustrated in Figure
|
||
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Theodoxus baeticus (Lamarck, 1822). A - C. Paralectotype of N. baetica from Andalusia, Spain (MHNG-MOLL- 51319). D) Operculum of a paralectotype of N. baetica from Andalusia (MHNG-MOLL- 51319); E - G. Lectotype of N. baetica from Andalusia (MHNG-MOLL- 51319); H-J; K; L; M; N. Five syntypes of N. meridionalis from Sicily (ZMB, without coll. no.); O - R. Topotype of N. guadianensis collected in the Guadiana River near Mertola, Portugal incorporated into the phylogeny (UGSB 22084); S - V. Specimen from Soller, Balearic Islands used in the phylogeny (UGSB 19162). Scale bars: 1 mm." figureDoi="10.3897/zse.96.48312.figure6" httpUri="https://binary.pensoft.net/fig/376455" pageId="0" pageNumber="25">6E-G</figureCitation>
|
||
as
|
||
<typeStatus>lectotype</typeStatus>
|
||
(MHNG-MOLL-51319)
|
||
</materialsCitation>
|
||
.
|
||
<materialsCitation collectionCode="ZMB" country="Turkey" typeStatus="syntypes">
|
||
Forty
|
||
<typeStatus>syntypes</typeStatus>
|
||
of
|
||
<taxonomicName genus="N." lsidName="N. meridionalis" pageId="0" pageNumber="25" rank="species" species="meridionalis">
|
||
<emphasis italics="true" pageId="0" pageNumber="25">N. meridionalis</emphasis>
|
||
</taxonomicName>
|
||
, consisting mainly of empty shells in addition to a few specimens with operculum, are housed in
|
||
<collectionCode collectionName="Germany, Berlin, Museum fuer Naturkunde der Humboldt-Universitaet" country="Germany" httpUri="http://grbio.org/cool/8syf-d21i" name="Museum f<>r Naturkunde Berlin (Zoological Collections)" type="Museum">ZMB</collectionCode>
|
||
(without coll. no.;
|
||
<bibRefCitation author="Gloeer, P" journalOrPublisher="Ecologica Montenegrina" pageId="0" pageNumber="25" pagination="133 - 137" refId="B61" refString="Gloeer, P, 2018. On the identity of Neritina baetica Lamarck, 1822 and Nerita meridionalis Philippi, 1836 (Gastropoda: Neritidae) from the Iberian Peninsula. Ecologica Montenegrina 18: 133 - 137" title="On the identity of Neritina baetica Lamarck, 1822 and Nerita meridionalis Philippi, 1836 (Gastropoda: Neritidae) from the Iberian Peninsula." volume="18" year="2018">
|
||
<normalizedToken originalValue="Glöer">Gloeer</normalizedToken>
|
||
2018
|
||
</bibRefCitation>
|
||
)
|
||
</materialsCitation>
|
||
.
|
||
<materialsCitation collectionCode="UCBL" country="Turkey" typeStatus="holotype">
|
||
The
|
||
<typeStatus>holotype</typeStatus>
|
||
(by monotypy) of
|
||
<taxonomicName genus="N." lsidName="N. callosa" pageId="0" pageNumber="25" rank="species" species="callosa">
|
||
<emphasis italics="true" pageId="0" pageNumber="25">N. callosa</emphasis>
|
||
</taxonomicName>
|
||
is supposed to be stored in
|
||
<collectionCode collectionName="UCBL" country="France" httpUri="http://grbio.org/cool/2dzr-gk4t" name="Centre de Paleontologie Stratigraphique et Paleoecologie" type="">UCBL</collectionCode>
|
||
(coll. no. UCBL-EM 33336), but it could not be found (E. Robert pers. comm. 07/2018)
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="25" type="remarks">
|
||
<paragraph pageId="0" pageNumber="25">Remarks.</paragraph>
|
||
<paragraph pageId="0" pageNumber="25">
|
||
Across the Mediterranean region a number of widely used nominal species exist that show considerable variation in shell shape and periostracum patterning, yet relatively similar operculum structures (including the presence of a pseudo-apophysis, ivory colouration of the calcareous base of the operculum, and a minimal rib-shield; Figs
|
||
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Theodoxus baeticus (Lamarck, 1822). A - C. Paralectotype of N. baetica from Andalusia, Spain (MHNG-MOLL- 51319). D) Operculum of a paralectotype of N. baetica from Andalusia (MHNG-MOLL- 51319); E - G. Lectotype of N. baetica from Andalusia (MHNG-MOLL- 51319); H-J; K; L; M; N. Five syntypes of N. meridionalis from Sicily (ZMB, without coll. no.); O - R. Topotype of N. guadianensis collected in the Guadiana River near Mertola, Portugal incorporated into the phylogeny (UGSB 22084); S - V. Specimen from Soller, Balearic Islands used in the phylogeny (UGSB 19162). Scale bars: 1 mm." figureDoi="10.3897/zse.96.48312.figure6" httpUri="https://binary.pensoft.net/fig/376455" pageId="0" pageNumber="25">6</figureCitation>
|
||
-
|
||
<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Theodoxus baeticus (Lamarck, 1822). A - D. Specimen from Cuquillo, Granada, Andalusia, Spain (UGSB 22089); E - H. Specimen (T. meridionalis - morphotype) collected in the Palancia River, Navajas, Spain (UGSB 22090); I - L. Specimen (T. elongatulus - morphotype) from Arrancada, Pombal, Portugal (UGSB 22080); M - P. Specimen (T. velascoi - morphotype) from Montanejos, Castellon, Spain (UGSB 22082). Theodoxus cf. valentinus (Graells, 1846). Q - S. Specimen conforming to T. valentinus from the Verd River, Masalaves, Spain (UGSB 21787); T. Operculum of a specimen conforming to T. valentinus from the same locality as Q-S (UGSB 22087). All specimens, except Q-S, were used in the phylogeny (Fig. 2). Scale bars: 1 mm." figureDoi="10.3897/zse.96.48312.figure8" httpUri="https://binary.pensoft.net/fig/376457" pageId="0" pageNumber="25">8</figureCitation>
|
||
). These species include
|
||
<taxonomicName genus="T." lsidName="T. baeticus" pageId="0" pageNumber="25" rank="species" species="baeticus">
|
||
<emphasis italics="true" pageId="0" pageNumber="25">T. baeticus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName genus="T." lsidName="T. callosus" pageId="0" pageNumber="25" rank="species" species="callosus">
|
||
<emphasis italics="true" pageId="0" pageNumber="25">T. callosus</emphasis>
|
||
</taxonomicName>
|
||
(Deshayes in
|
||
<bibRefCitation author="Geoffroy Saint-Hilaire, I" journalOrPublisher="Levrault, Paris" pageId="0" pageNumber="25" publicationUrl="http://reader.digitale-sammlungen.de/de/fs3/object/display/bsb10360027_00005.html" refId="B58" refString="Geoffroy Saint-Hilaire, I, Geoffroy Saint-Hilaire, E, Deshayes, GP, Bibron, G, Bory de Saint-Vincent, J-B, 1832-1836. Expedition scientifique de Moree, Tome III - 1.re Partie. Zoologie. Premiere Section. - Animaux vertebres, Mollusques et Polypiers. F. G. Levrault, Paris, http://reader.digitale-sammlungen.de/de/fs3/object/display/bsb10360027_00005.html" title="Expedition scientifique de Moree, Tome III - 1. re Partie. Zoologie. Premiere Section. - Animaux vertebres, Mollusques et Polypiers. F. G." url="http://reader.digitale-sammlungen.de/de/fs3/object/display/bsb10360027_00005.html" year="1832 - 1836">Geoffroy Saint-Hilaire et al. 1835</bibRefCitation>
|
||
),
|
||
<taxonomicName genus="T." lsidName="T. varius" pageId="0" pageNumber="25" rank="species" species="varius">
|
||
<emphasis italics="true" pageId="0" pageNumber="25">T. varius</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<normalizedToken originalValue="Rossmässler">Rossmaessler</normalizedToken>
|
||
, 1835),
|
||
<taxonomicName genus="T." lsidName="T. meridionalis" pageId="0" pageNumber="25" rank="species" species="meridionalis">
|
||
<emphasis italics="true" pageId="0" pageNumber="25">T. meridionalis</emphasis>
|
||
</taxonomicName>
|
||
(Philippi, 1836),
|
||
<taxonomicName genus="T." lsidName="T. elongatulus" pageId="0" pageNumber="25" rank="species" species="elongatulus">
|
||
<emphasis italics="true" pageId="0" pageNumber="25">T. elongatulus</emphasis>
|
||
</taxonomicName>
|
||
(Morelet, 1845),
|
||
<taxonomicName genus="T." lsidName="T. guadianensis" pageId="0" pageNumber="25" rank="species" species="guadianensis">
|
||
<emphasis italics="true" pageId="0" pageNumber="25">T. guadianensis</emphasis>
|
||
</taxonomicName>
|
||
(Morelet, 1845),
|
||
<taxonomicName genus="T." lsidName="T. velascoi" pageId="0" pageNumber="25" rank="species" species="velascoi">
|
||
<emphasis italics="true" pageId="0" pageNumber="25">T. velascoi</emphasis>
|
||
</taxonomicName>
|
||
(Graells, 1846), and
|
||
<taxonomicName genus="T." lsidName="T. valentinus" pageId="0" pageNumber="25" rank="species" species="valentinus">
|
||
<emphasis italics="true" pageId="0" pageNumber="25">T. valentinus</emphasis>
|
||
</taxonomicName>
|
||
(Graells, 1846), as well as a number of species already synonymised under the above, including
|
||
<taxonomicName class="Gastropoda" family="Neritidae" genus="Nerita" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Nerita philippii" order="Cycloneritida" pageId="0" pageNumber="25" phylum="Mollusca" rank="species" species="philippii">
|
||
<emphasis italics="true" pageId="0" pageNumber="25">Nerita philippii</emphasis>
|
||
</taxonomicName>
|
||
<normalizedToken originalValue="Récluz">Recluz</normalizedToken>
|
||
, 1841,
|
||
<taxonomicName authorityName="Morelet" authorityYear="1845" class="Gastropoda" family="Neritidae" genus="Neritina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Neritina inquinata" order="Cycloneritida" pageId="0" pageNumber="25" phylum="Mollusca" rank="species" species="inquinata">
|
||
<emphasis italics="true" pageId="0" pageNumber="25">Neritina inquinata</emphasis>
|
||
</taxonomicName>
|
||
Morelet, 1845,
|
||
<taxonomicName baseAuthorityName="Gmelin" baseAuthorityYear="1791" class="Gastropoda" family="Neritidae" genus="Neritina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Neritina violacea" order="Cycloneritida" pageId="0" pageNumber="25" phylum="Mollusca" rank="species" species="violacea">
|
||
<emphasis italics="true" pageId="0" pageNumber="25">Neritina violacea</emphasis>
|
||
</taxonomicName>
|
||
Morelet, 1845,
|
||
<taxonomicName class="Gastropoda" family="Neritidae" genus="Neritina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Neritina anatensis" order="Cycloneritida" pageId="0" pageNumber="25" phylum="Mollusca" rank="species" species="anatensis">
|
||
<emphasis italics="true" pageId="0" pageNumber="25">Neritina anatensis</emphasis>
|
||
</taxonomicName>
|
||
G.B. Sowerby II, 1849, and
|
||
<taxonomicName authorityName="Crosse" authorityYear="1880" class="Gastropoda" family="Neritidae" genus="Neritina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Neritina hidalgoi" order="Cycloneritida" pageId="0" pageNumber="25" phylum="Mollusca" rank="species" species="hidalgoi">
|
||
<emphasis italics="true" pageId="0" pageNumber="25">Neritina hidalgoi</emphasis>
|
||
</taxonomicName>
|
||
Crosse, 1880. Most of these species were described from the Iberian Peninsula. Recent insights from molecular phylogenetics and morphological reviews indicate the presence of either a single species (
|
||
<taxonomicName genus="T." lsidName="T. meridionalis" pageId="0" pageNumber="25" rank="species" species="meridionalis">
|
||
<emphasis italics="true" pageId="0" pageNumber="25">T. meridionalis</emphasis>
|
||
</taxonomicName>
|
||
) or two species (
|
||
<taxonomicName genus="T." lsidName="T. meridionalis" pageId="0" pageNumber="25" rank="species" species="meridionalis">
|
||
<emphasis italics="true" pageId="0" pageNumber="25">T. meridionalis</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName genus="T." lsidName="T. valentinus" pageId="0" pageNumber="25" rank="species" species="valentinus">
|
||
<emphasis italics="true" pageId="0" pageNumber="25">T. valentinus</emphasis>
|
||
</taxonomicName>
|
||
), while all others may be considered junior synonyms (
|
||
<bibRefCitation DOI="https://doi.org/10.1016/j.ympev.2006.06.026" author="Bunje, PME" journalOrPublisher="Molecular Phylogenetics and Evolution" pageId="0" pageNumber="25" pagination="373 - 387" refId="B35" refString="Bunje, PME, Lindberg, DR, 2007. Lineage divergence of a freshwater snail clade associated with post-Tethys marine basin development. Molecular Phylogenetics and Evolution 42 (2): 373 - 387, DOI: https://doi.org/10.1016/j.ympev.2006.06.026" title="Lineage divergence of a freshwater snail clade associated with post-Tethys marine basin development." url="https://doi.org/10.1016/j.ympev.2006.06.026" volume="42" year="2007">Bunje and Lindberg 2007</bibRefCitation>
|
||
;
|
||
<bibRefCitation author="Ramos, A" journalOrPublisher="Systematic Biology" pageId="0" pageNumber="25" publicationUrl="https://www.miteco.gob.es/es/biodiversidad/temas/conservacion-de-especies/dictamen_theodoxus_velascoi_tcm30-201183.pdf" refId="B130" refString="Ramos, A, 2014. CC 08/2014. Ministerio de Agricultura, Alimentacion y Medio Ambiente, Madrid. https://www.miteco.gob.es/es/biodiversidad/temas/conservacion-de-especies/dictamen_theodoxus_velascoi_tcm30-201183.pdf" title="CC 08 / 2014. Ministerio de Agricultura, Alimentacion y Medio Ambiente, Madrid." url="https://www.miteco.gob.es/es/biodiversidad/temas/conservacion-de-especies/dictamen_theodoxus_velascoi_tcm30-201183.pdf" year="2014">Ramos 2014</bibRefCitation>
|
||
;
|
||
<bibRefCitation author="Martinez-Orti, A" journalOrPublisher="Privately published, Tehran" pageId="0" pageNumber="25" refId="B97" refString="Martinez-Orti, A, Pedrola-Monfort, J, Robles, F, Madeira, MJ, 2015. Revision taxonomica, mediante estudios moleculares, de las especies de Theodoxus Montfort, 1810 (Gastropoda, Neritopsina) descritas en la Peninsula Iberica. In Foro Malacologico. Sociedad Espanola De Malacologia, Malaga, 25-26." title="Revision taxonomica, mediante estudios moleculares, de las especies de Theodoxus Montfort, 1810 (Gastropoda, Neritopsina) descritas en la Peninsula Iberica. In Foro Malacologico. Sociedad Espanola De Malacologia, Malaga, 25 - 26." year="2015">
|
||
<normalizedToken originalValue="Martínez-Ortí">Martinez-Orti</normalizedToken>
|
||
et al. 2015
|
||
</bibRefCitation>
|
||
;
|
||
<bibRefCitation author="Alba, DM" journalOrPublisher="Spira" pageId="0" pageNumber="25" pagination="41 - 65" refId="B6" refString="Alba, DM, Corbella, J, Guillen, G, Prats, L, Tarruella, A, 2016. Presence of two different species of Theodoxus Montfort, 1810 (Gastropoda: Neritidae) in Catalonia. Spira 6: 41 - 65" title="Presence of two different species of Theodoxus Montfort, 1810 (Gastropoda: Neritidae) in Catalonia." volume="6" year="2016">Alba et al. 2016</bibRefCitation>
|
||
;
|
||
<bibRefCitation DOI="https://doi.org/10.1111/jbi.13590" author="Sands, AF" journalOrPublisher="Journal of Biogeography" pageId="0" pageNumber="25" pagination="1274 - 1286" refId="B138" refString="Sands, AF, Sereda, SV, Stelbrink, B, Neubauer, TA, Lazarev, S, Wilke, T, Albrecht, C, 2019a. Contributions of biogeographical functions to species accumulation may change over time in refugial regions. Journal of Biogeography 46 (6): 1274 - 1286, DOI: https://doi.org/10.1111/jbi.13590" title="Contributions of biogeographical functions to species accumulation may change over time in refugial regions." url="https://doi.org/10.1111/jbi.13590" volume="46" year="2019 a">Sands et al. 2019a</bibRefCitation>
|
||
). Based on a reinvestigation of
|
||
<typeStatus>type</typeStatus>
|
||
material,
|
||
<bibRefCitation author="Gloeer, P" journalOrPublisher="Ecologica Montenegrina" pageId="0" pageNumber="25" pagination="133 - 137" refId="B61" refString="Gloeer, P, 2018. On the identity of Neritina baetica Lamarck, 1822 and Nerita meridionalis Philippi, 1836 (Gastropoda: Neritidae) from the Iberian Peninsula. Ecologica Montenegrina 18: 133 - 137" title="On the identity of Neritina baetica Lamarck, 1822 and Nerita meridionalis Philippi, 1836 (Gastropoda: Neritidae) from the Iberian Peninsula." volume="18" year="2018">
|
||
<normalizedToken originalValue="Glöer">Gloeer</normalizedToken>
|
||
(2018)
|
||
</bibRefCitation>
|
||
recently showed that
|
||
<taxonomicName genus="T." lsidName="T. meridionalis" pageId="0" pageNumber="25" rank="species" species="meridionalis">
|
||
<emphasis italics="true" pageId="0" pageNumber="25">T. meridionalis</emphasis>
|
||
</taxonomicName>
|
||
is a junior synonym of
|
||
<taxonomicName genus="T." lsidName="T. baeticus" pageId="0" pageNumber="25" rank="species" species="baeticus">
|
||
<emphasis italics="true" pageId="0" pageNumber="25">T. baeticus</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
<taxonomicName baseAuthorityName="Lamarck" baseAuthorityYear="1822" class="Gastropoda" family="Neritidae" genus="Theodoxus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Theodoxus baeticus" order="Cycloneritida" pageId="0" pageNumber="25" phylum="Mollusca" rank="species" species="baeticus">
|
||
<emphasis italics="true" pageId="0" pageNumber="25">Theodoxus baeticus</emphasis>
|
||
</taxonomicName>
|
||
has previously been considered a junior synonym of
|
||
<taxonomicName genus="T." lsidName="T. fluviatilis" pageId="0" pageNumber="25" rank="species" species="fluviatilis">
|
||
<emphasis italics="true" pageId="0" pageNumber="25">T. fluviatilis</emphasis>
|
||
</taxonomicName>
|
||
, a view that was first rejected on the basis of molecular data by
|
||
<bibRefCitation DOI="https://doi.org/10.1016/j.ympev.2006.06.026" author="Bunje, PME" journalOrPublisher="Molecular Phylogenetics and Evolution" pageId="0" pageNumber="25" pagination="373 - 387" refId="B35" refString="Bunje, PME, Lindberg, DR, 2007. Lineage divergence of a freshwater snail clade associated with post-Tethys marine basin development. Molecular Phylogenetics and Evolution 42 (2): 373 - 387, DOI: https://doi.org/10.1016/j.ympev.2006.06.026" title="Lineage divergence of a freshwater snail clade associated with post-Tethys marine basin development." url="https://doi.org/10.1016/j.ympev.2006.06.026" volume="42" year="2007">Bunje and Lindberg (2007)</bibRefCitation>
|
||
. Here, we consider all species listed above except for
|
||
<taxonomicName genus="T." lsidName="T. valentinus" pageId="0" pageNumber="25" rank="species" species="valentinus">
|
||
<emphasis italics="true" pageId="0" pageNumber="25">T. valentinus</emphasis>
|
||
</taxonomicName>
|
||
as synonyms of
|
||
<taxonomicName genus="T." lsidName="T. baeticus" pageId="0" pageNumber="25" rank="species" species="baeticus">
|
||
<emphasis italics="true" pageId="0" pageNumber="25">T. baeticus</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
<taxonomicName baseAuthorityName="Graells" baseAuthorityYear="1846" class="Gastropoda" family="Neritidae" genus="Theodoxus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Theodoxus valentinus" order="Cycloneritida" pageId="0" pageNumber="25" phylum="Mollusca" rank="species" species="valentinus">
|
||
<emphasis italics="true" pageId="0" pageNumber="25">Theodoxus valentinus</emphasis>
|
||
</taxonomicName>
|
||
differs from
|
||
<taxonomicName genus="T." lsidName="T. baeticus" pageId="0" pageNumber="25" rank="species" species="baeticus">
|
||
<emphasis italics="true" pageId="0" pageNumber="25">T. baeticus</emphasis>
|
||
</taxonomicName>
|
||
in its unique shell shape, showing a high spire and two bulgy keels. Specimens with similar morphology obtained from nearby the
|
||
<typeStatus>type</typeStatus>
|
||
locality of
|
||
<taxonomicName genus="T." lsidName="T. valentinus" pageId="0" pageNumber="25" rank="species" species="valentinus">
|
||
<emphasis italics="true" pageId="0" pageNumber="25">T. valentinus</emphasis>
|
||
</taxonomicName>
|
||
(near
|
||
<normalizedToken originalValue="Xàtiva">Xativa</normalizedToken>
|
||
, Spain) grouped with
|
||
<taxonomicName genus="T." lsidName="T. baeticus" pageId="0" pageNumber="25" rank="species" species="baeticus">
|
||
<emphasis italics="true" pageId="0" pageNumber="25">T. baeticus</emphasis>
|
||
</taxonomicName>
|
||
in our analyses (Figs
|
||
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Dated phylogeny of the genus Theodoxus constructed with BEAST, based on existing GenBank and newly incorporated data for COI, 16 S and ATPα (Suppl. material 1: Table S 1). Morphospecies where associations are queried are highlighted by a question mark after the species name (see the " Systematic checklist " for more information). Small green (PP ≥ 0.95) and red (PP <0.95) dots on nodes indicate the relative posterior probability support of divergence events. Node labels (1 - 32) correspond to divergence dates and HPD intervals in millions of years ago (Mya) for selected nodes (see Suppl. material 1: Table S 3). Dashed dark blue and light blue lines represent the minimum interspecific and maximum intraspecific variation determined by Sands et al. (2019 a) to aid Theodoxus spp. delimitation (see see Methods, Phylogenetics and species delimitation and that study for more information on the delimitation methods)." figureDoi="10.3897/zse.96.48312.figure2" httpUri="https://binary.pensoft.net/fig/376451" pageId="0" pageNumber="25">2</figureCitation>
|
||
,
|
||
<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Theodoxus baeticus (Lamarck, 1822). A - D. Specimen from Cuquillo, Granada, Andalusia, Spain (UGSB 22089); E - H. Specimen (T. meridionalis - morphotype) collected in the Palancia River, Navajas, Spain (UGSB 22090); I - L. Specimen (T. elongatulus - morphotype) from Arrancada, Pombal, Portugal (UGSB 22080); M - P. Specimen (T. velascoi - morphotype) from Montanejos, Castellon, Spain (UGSB 22082). Theodoxus cf. valentinus (Graells, 1846). Q - S. Specimen conforming to T. valentinus from the Verd River, Masalaves, Spain (UGSB 21787); T. Operculum of a specimen conforming to T. valentinus from the same locality as Q-S (UGSB 22087). All specimens, except Q-S, were used in the phylogeny (Fig. 2). Scale bars: 1 mm." figureDoi="10.3897/zse.96.48312.figure8" httpUri="https://binary.pensoft.net/fig/376457" pageId="0" pageNumber="25">8Q-T</figureCitation>
|
||
; Suppl. material 1: Table S1), which may suggest the two species are synonyms. We refrain here from a final conclusion on the matter until topotypic material of
|
||
<taxonomicName genus="T." lsidName="T. valentinus" pageId="0" pageNumber="25" rank="species" species="valentinus">
|
||
<emphasis italics="true" pageId="0" pageNumber="25">T. valentinus</emphasis>
|
||
</taxonomicName>
|
||
is genetically studied.
|
||
</paragraph>
|
||
<caption doi="10.3897/zse.96.48312.figure6" httpUri="https://binary.pensoft.net/fig/376455" pageId="0" pageNumber="25" start="Figure 6" startId="F6">
|
||
<paragraph pageId="0" pageNumber="25">
|
||
<emphasis bold="true" pageId="0" pageNumber="25">Figure 6.</emphasis>
|
||
<taxonomicName baseAuthorityName="Lamarck" baseAuthorityYear="1822" class="Gastropoda" family="Neritidae" genus="Theodoxus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Theodoxus baeticus" order="Cycloneritida" pageId="0" pageNumber="25" phylum="Mollusca" rank="species" species="baeticus">
|
||
<emphasis italics="true" pageId="0" pageNumber="25">Theodoxus baeticus</emphasis>
|
||
</taxonomicName>
|
||
(Lamarck, 1822).
|
||
<emphasis bold="true" pageId="0" pageNumber="25">A</emphasis>
|
||
-
|
||
<emphasis bold="true" pageId="0" pageNumber="25">C</emphasis>
|
||
.
|
||
<typeStatus>Paralectotype</typeStatus>
|
||
of
|
||
<taxonomicName genus="N." lsidName="N. baetica" pageId="0" pageNumber="25" rank="species" species="baetica">
|
||
<emphasis italics="true" pageId="0" pageNumber="25">N. baetica</emphasis>
|
||
</taxonomicName>
|
||
from Andalusia, Spain (MHNG-MOLL-51319).
|
||
<emphasis bold="true" pageId="0" pageNumber="25">D</emphasis>
|
||
) Operculum of a
|
||
<typeStatus>paralectotype</typeStatus>
|
||
of
|
||
<taxonomicName genus="N." lsidName="N. baetica" pageId="0" pageNumber="25" rank="species" species="baetica">
|
||
<emphasis italics="true" pageId="0" pageNumber="25">N. baetica</emphasis>
|
||
</taxonomicName>
|
||
from Andalusia (MHNG-MOLL-51319);
|
||
<emphasis bold="true" pageId="0" pageNumber="25">E</emphasis>
|
||
-
|
||
<emphasis bold="true" pageId="0" pageNumber="25">G</emphasis>
|
||
.
|
||
<typeStatus>Lectotype</typeStatus>
|
||
of
|
||
<taxonomicName genus="N." lsidName="N. baetica" pageId="0" pageNumber="25" rank="species" species="baetica">
|
||
<emphasis italics="true" pageId="0" pageNumber="25">N. baetica</emphasis>
|
||
</taxonomicName>
|
||
from Andalusia (MHNG-MOLL-51319);
|
||
<emphasis bold="true" pageId="0" pageNumber="25">H-J; K; L; M; N</emphasis>
|
||
. Five
|
||
<typeStatus>syntypes</typeStatus>
|
||
of
|
||
<taxonomicName genus="N." lsidName="N. meridionalis" pageId="0" pageNumber="25" rank="species" species="meridionalis">
|
||
<emphasis italics="true" pageId="0" pageNumber="25">N. meridionalis</emphasis>
|
||
</taxonomicName>
|
||
from Sicily (
|
||
<collectionCode collectionName="Germany, Berlin, Museum fuer Naturkunde der Humboldt-Universitaet" country="Germany" httpUri="http://grbio.org/cool/8syf-d21i" name="Museum f<>r Naturkunde Berlin (Zoological Collections)" type="Museum">ZMB</collectionCode>
|
||
, without coll. no.);
|
||
<emphasis bold="true" pageId="0" pageNumber="25">O</emphasis>
|
||
-
|
||
<emphasis bold="true" pageId="0" pageNumber="25">R</emphasis>
|
||
. Topotype of
|
||
<taxonomicName genus="N." lsidName="N. guadianensis" pageId="0" pageNumber="25" rank="species" species="guadianensis">
|
||
<emphasis italics="true" pageId="0" pageNumber="25">N. guadianensis</emphasis>
|
||
</taxonomicName>
|
||
collected in the Guadiana River near
|
||
<normalizedToken originalValue="Mértola">Mertola</normalizedToken>
|
||
, Portugal incorporated into the phylogeny (
|
||
<collectionCode collectionName="UGSB">UGSB</collectionCode>
|
||
22084);
|
||
<emphasis bold="true" pageId="0" pageNumber="25">S</emphasis>
|
||
-
|
||
<emphasis bold="true" pageId="0" pageNumber="25">V</emphasis>
|
||
. Specimen from
|
||
<normalizedToken originalValue="Sóller">Soller</normalizedToken>
|
||
, Balearic Islands used in the phylogeny (
|
||
<collectionCode collectionName="UGSB">UGSB</collectionCode>
|
||
19162). Scale bars:
|
||
<quantity metricMagnitude="-3" metricUnit="m" metricValue="1.0" unit="mm" value="1.0">1 mm</quantity>
|
||
.
|
||
</paragraph>
|
||
</caption>
|
||
<caption doi="10.3897/zse.96.48312.figure7" httpUri="https://binary.pensoft.net/fig/376456" pageId="0" pageNumber="25" start="Figure 7" startId="F7">
|
||
<paragraph pageId="0" pageNumber="25">
|
||
<emphasis bold="true" pageId="0" pageNumber="25">Figure 7.</emphasis>
|
||
<taxonomicName baseAuthorityName="Lamarck" baseAuthorityYear="1822" class="Gastropoda" family="Neritidae" genus="Theodoxus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Theodoxus baeticus" order="Cycloneritida" pageId="0" pageNumber="25" phylum="Mollusca" rank="species" species="baeticus">
|
||
<emphasis italics="true" pageId="0" pageNumber="25">Theodoxus baeticus</emphasis>
|
||
</taxonomicName>
|
||
(Lamarck, 1822).
|
||
<emphasis bold="true" pageId="0" pageNumber="25">A</emphasis>
|
||
-
|
||
<emphasis bold="true" pageId="0" pageNumber="25">D</emphasis>
|
||
. Specimen (
|
||
<taxonomicName genus="T." lsidName="T. callosus" pageId="0" pageNumber="25" rank="species" species="callosus">
|
||
<emphasis italics="true" pageId="0" pageNumber="25">T. callosus</emphasis>
|
||
</taxonomicName>
|
||
-morphoype) collected in Akyaka, Turkey (
|
||
<collectionCode collectionName="UGSB">UGSB</collectionCode>
|
||
19118);
|
||
<emphasis bold="true" pageId="0" pageNumber="25">E</emphasis>
|
||
-
|
||
<emphasis bold="true" pageId="0" pageNumber="25">H</emphasis>
|
||
. Specimen (
|
||
<taxonomicName genus="T." lsidName="T. callosus" pageId="0" pageNumber="25" rank="species" species="callosus">
|
||
<emphasis italics="true" pageId="0" pageNumber="25">T. callosus</emphasis>
|
||
</taxonomicName>
|
||
-morphotype) collected in
|
||
<normalizedToken originalValue="Káto">Kato</normalizedToken>
|
||
<normalizedToken originalValue="Tragána">Tragana</normalizedToken>
|
||
, Greece (
|
||
<collectionCode collectionName="UGSB">UGSB</collectionCode>
|
||
2488);
|
||
<emphasis bold="true" pageId="0" pageNumber="25">I</emphasis>
|
||
-
|
||
<emphasis bold="true" pageId="0" pageNumber="25">L</emphasis>
|
||
. Specimen (
|
||
<taxonomicName genus="T." lsidName="T. varius" pageId="0" pageNumber="25" rank="species" species="varius">
|
||
<emphasis italics="true" pageId="0" pageNumber="25">T. varius</emphasis>
|
||
</taxonomicName>
|
||
-morphotype) collected at the Blue Eye Nature Monument, Albania (
|
||
<collectionCode collectionName="UGSB">UGSB</collectionCode>
|
||
24170);
|
||
<emphasis bold="true" pageId="0" pageNumber="25">M</emphasis>
|
||
-
|
||
<emphasis bold="true" pageId="0" pageNumber="25">P</emphasis>
|
||
. Specimen (
|
||
<taxonomicName genus="T." lsidName="T. meridionalis" pageId="0" pageNumber="25" rank="species" species="meridionalis">
|
||
<emphasis italics="true" pageId="0" pageNumber="25">T. meridionalis</emphasis>
|
||
</taxonomicName>
|
||
-morphotype) collected at the Ketana Oasis, Gabes, Tunisia (
|
||
<collectionCode collectionName="UGSB">UGSB</collectionCode>
|
||
18111). All specimens, except A-H, were used in the phylogeny (Fig.
|
||
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Dated phylogeny of the genus Theodoxus constructed with BEAST, based on existing GenBank and newly incorporated data for COI, 16 S and ATPα (Suppl. material 1: Table S 1). Morphospecies where associations are queried are highlighted by a question mark after the species name (see the " Systematic checklist " for more information). Small green (PP ≥ 0.95) and red (PP <0.95) dots on nodes indicate the relative posterior probability support of divergence events. Node labels (1 - 32) correspond to divergence dates and HPD intervals in millions of years ago (Mya) for selected nodes (see Suppl. material 1: Table S 3). Dashed dark blue and light blue lines represent the minimum interspecific and maximum intraspecific variation determined by Sands et al. (2019 a) to aid Theodoxus spp. delimitation (see see Methods, Phylogenetics and species delimitation and that study for more information on the delimitation methods)." figureDoi="10.3897/zse.96.48312.figure2" httpUri="https://binary.pensoft.net/fig/376451" pageId="0" pageNumber="25">2</figureCitation>
|
||
). Scale bars:
|
||
<quantity metricMagnitude="-3" metricUnit="m" metricValue="1.0" unit="mm" value="1.0">1 mm</quantity>
|
||
.
|
||
</paragraph>
|
||
</caption>
|
||
<caption doi="10.3897/zse.96.48312.figure8" httpUri="https://binary.pensoft.net/fig/376457" pageId="0" pageNumber="25" start="Figure 8" startId="F8">
|
||
<paragraph pageId="0" pageNumber="25">
|
||
<emphasis bold="true" pageId="0" pageNumber="25">Figure 8.</emphasis>
|
||
<taxonomicName baseAuthorityName="Lamarck" baseAuthorityYear="1822" class="Gastropoda" family="Neritidae" genus="Theodoxus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Theodoxus baeticus" order="Cycloneritida" pageId="0" pageNumber="25" phylum="Mollusca" rank="species" species="baeticus">
|
||
<emphasis italics="true" pageId="0" pageNumber="25">Theodoxus baeticus</emphasis>
|
||
</taxonomicName>
|
||
(Lamarck, 1822).
|
||
<emphasis bold="true" pageId="0" pageNumber="25">A</emphasis>
|
||
-
|
||
<emphasis bold="true" pageId="0" pageNumber="25">D</emphasis>
|
||
. Specimen from Cuquillo, Granada, Andalusia, Spain (
|
||
<collectionCode collectionName="UGSB">UGSB</collectionCode>
|
||
22089);
|
||
<emphasis bold="true" pageId="0" pageNumber="25">E</emphasis>
|
||
-
|
||
<emphasis bold="true" pageId="0" pageNumber="25">H</emphasis>
|
||
. Specimen (
|
||
<taxonomicName genus="T." lsidName="T. meridionalis" pageId="0" pageNumber="25" rank="species" species="meridionalis">
|
||
<emphasis italics="true" pageId="0" pageNumber="25">T. meridionalis</emphasis>
|
||
</taxonomicName>
|
||
-morphotype) collected in the Palancia River, Navajas, Spain (
|
||
<collectionCode collectionName="UGSB">UGSB</collectionCode>
|
||
22090);
|
||
<emphasis bold="true" pageId="0" pageNumber="25">I</emphasis>
|
||
-
|
||
<emphasis bold="true" pageId="0" pageNumber="25">L</emphasis>
|
||
. Specimen (
|
||
<taxonomicName genus="T." lsidName="T. elongatulus" pageId="0" pageNumber="25" rank="species" species="elongatulus">
|
||
<emphasis italics="true" pageId="0" pageNumber="25">T. elongatulus</emphasis>
|
||
</taxonomicName>
|
||
-morphotype) from Arrancada, Pombal, Portugal (
|
||
<collectionCode collectionName="UGSB">UGSB</collectionCode>
|
||
22080);
|
||
<emphasis bold="true" pageId="0" pageNumber="25">M</emphasis>
|
||
-
|
||
<emphasis bold="true" pageId="0" pageNumber="25">P</emphasis>
|
||
. Specimen (
|
||
<taxonomicName genus="T." lsidName="T. velascoi" pageId="0" pageNumber="25" rank="species" species="velascoi">
|
||
<emphasis italics="true" pageId="0" pageNumber="25">T. velascoi</emphasis>
|
||
</taxonomicName>
|
||
-morphotype) from Montanejos,
|
||
<normalizedToken originalValue="Castellón">Castellon</normalizedToken>
|
||
, Spain (
|
||
<collectionCode collectionName="UGSB">UGSB</collectionCode>
|
||
22082).
|
||
<taxonomicName baseAuthorityName="Graells" baseAuthorityYear="1846" class="Gastropoda" family="Neritidae" genus="Theodoxus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Theodoxus valentinus" order="Cycloneritida" pageId="0" pageNumber="25" phylum="Mollusca" rank="species" species="valentinus">Theodoxus cf. valentinus</taxonomicName>
|
||
(Graells, 1846).
|
||
<emphasis bold="true" pageId="0" pageNumber="25">Q</emphasis>
|
||
-
|
||
<emphasis bold="true" pageId="0" pageNumber="25">S</emphasis>
|
||
. Specimen conforming to
|
||
<taxonomicName genus="T." lsidName="T. valentinus" pageId="0" pageNumber="25" rank="species" species="valentinus">
|
||
<emphasis italics="true" pageId="0" pageNumber="25">T. valentinus</emphasis>
|
||
</taxonomicName>
|
||
from the Verd River,
|
||
<normalizedToken originalValue="Masalavés">Masalaves</normalizedToken>
|
||
, Spain (
|
||
<collectionCode collectionName="UGSB">UGSB</collectionCode>
|
||
21787);
|
||
<emphasis bold="true" pageId="0" pageNumber="25">T</emphasis>
|
||
. Operculum of a specimen conforming to
|
||
<taxonomicName genus="T." lsidName="T. valentinus" pageId="0" pageNumber="25" rank="species" species="valentinus">
|
||
<emphasis italics="true" pageId="0" pageNumber="25">T. valentinus</emphasis>
|
||
</taxonomicName>
|
||
from the same locality as Q-S (
|
||
<collectionCode collectionName="UGSB">UGSB</collectionCode>
|
||
22087). All specimens, except Q-S, were used in the phylogeny (Fig.
|
||
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Dated phylogeny of the genus Theodoxus constructed with BEAST, based on existing GenBank and newly incorporated data for COI, 16 S and ATPα (Suppl. material 1: Table S 1). Morphospecies where associations are queried are highlighted by a question mark after the species name (see the " Systematic checklist " for more information). Small green (PP ≥ 0.95) and red (PP <0.95) dots on nodes indicate the relative posterior probability support of divergence events. Node labels (1 - 32) correspond to divergence dates and HPD intervals in millions of years ago (Mya) for selected nodes (see Suppl. material 1: Table S 3). Dashed dark blue and light blue lines represent the minimum interspecific and maximum intraspecific variation determined by Sands et al. (2019 a) to aid Theodoxus spp. delimitation (see see Methods, Phylogenetics and species delimitation and that study for more information on the delimitation methods)." figureDoi="10.3897/zse.96.48312.figure2" httpUri="https://binary.pensoft.net/fig/376451" pageId="0" pageNumber="25">2</figureCitation>
|
||
). Scale bars:
|
||
<quantity metricMagnitude="-3" metricUnit="m" metricValue="1.0" unit="mm" value="1.0">1 mm</quantity>
|
||
.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph pageId="0" pageNumber="25">
|
||
Our phylogenetic analysis indicate that
|
||
<taxonomicName genus="T." lsidName="T. baeticus" pageId="0" pageNumber="25" rank="species" species="baeticus">
|
||
<emphasis italics="true" pageId="0" pageNumber="25">T. baeticus</emphasis>
|
||
</taxonomicName>
|
||
diverged from other species likely during the early Pleistocene and shares a close relationship with an undescribed species from Spain (see
|
||
<bibRefCitation DOI="https://doi.org/10.1111/jbi.13590" author="Sands, AF" journalOrPublisher="Journal of Biogeography" pageId="0" pageNumber="25" pagination="1274 - 1286" refId="B138" refString="Sands, AF, Sereda, SV, Stelbrink, B, Neubauer, TA, Lazarev, S, Wilke, T, Albrecht, C, 2019a. Contributions of biogeographical functions to species accumulation may change over time in refugial regions. Journal of Biogeography 46 (6): 1274 - 1286, DOI: https://doi.org/10.1111/jbi.13590" title="Contributions of biogeographical functions to species accumulation may change over time in refugial regions." url="https://doi.org/10.1111/jbi.13590" volume="46" year="2019 a">Sands et al. 2019a</bibRefCitation>
|
||
) and
|
||
<taxonomicName genus="T." lsidName="T. marteli" pageId="0" pageNumber="25" rank="species" species="marteli">
|
||
<emphasis italics="true" pageId="0" pageNumber="25">T. marteli</emphasis>
|
||
</taxonomicName>
|
||
(Pallary, 1918) from northern Africa (Fig.
|
||
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Dated phylogeny of the genus Theodoxus constructed with BEAST, based on existing GenBank and newly incorporated data for COI, 16 S and ATPα (Suppl. material 1: Table S 1). Morphospecies where associations are queried are highlighted by a question mark after the species name (see the " Systematic checklist " for more information). Small green (PP ≥ 0.95) and red (PP <0.95) dots on nodes indicate the relative posterior probability support of divergence events. Node labels (1 - 32) correspond to divergence dates and HPD intervals in millions of years ago (Mya) for selected nodes (see Suppl. material 1: Table S 3). Dashed dark blue and light blue lines represent the minimum interspecific and maximum intraspecific variation determined by Sands et al. (2019 a) to aid Theodoxus spp. delimitation (see see Methods, Phylogenetics and species delimitation and that study for more information on the delimitation methods)." figureDoi="10.3897/zse.96.48312.figure2" httpUri="https://binary.pensoft.net/fig/376451" pageId="0" pageNumber="25">2</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="25" type="distribution">
|
||
<paragraph pageId="0" pageNumber="25">Distribution.</paragraph>
|
||
<paragraph pageId="0" pageNumber="25">
|
||
Considering the synonymies proposed here,
|
||
<taxonomicName genus="T." lsidName="T. baeticus" pageId="0" pageNumber="25" rank="species" species="baeticus">
|
||
<emphasis italics="true" pageId="0" pageNumber="25">T. baeticus</emphasis>
|
||
</taxonomicName>
|
||
is a widespread species throughout the Mediterranean region. It is present across the Iberian Peninsula, the Balearic Islands, Sicily, Tunisia, and the Balkans (
|
||
<bibRefCitation author="Bănărescu, P" journalOrPublisher="Proceedings of the Academy of Natural Sciences of Philadelphia" pageId="0" pageNumber="25" refId="B19" refString="Bănărescu, P, 1991. Zoogeography of Fresh Waters. Vol. 2. Distribution and dispersal of freshwater animals in North America and Eurasia. AULA-Verlag, Wiesbaden, Germany, 519-1091." title="Zoogeography of Fresh Waters. Vol. 2. Distribution and dispersal of freshwater animals in North America and Eurasia. AULA-Verlag, Wiesbaden, Germany, 519 - 1091." year="1991">Bănărescu 1991</bibRefCitation>
|
||
;
|
||
<bibRefCitation author="Bandel, K" journalOrPublisher="Mitteilungen aus dem geologisch-palaeontologischen Institut der Universitaet Hamburg" pageId="0" pageNumber="25" pagination="65 - 164" refId="B20" refString="Bandel, K, 2001. The history of Theodoxus and Neritina connected with description and systematic evaluation of related Neritimorpha (Gastropoda). Mitteilungen aus dem geologisch-palaeontologischen Institut der Universitaet Hamburg 85: 65 - 164" title="The history of Theodoxus and Neritina connected with description and systematic evaluation of related Neritimorpha (Gastropoda)." volume="85" year="2001">Bandel 2001</bibRefCitation>
|
||
;
|
||
<bibRefCitation author="Bank, RA" journalOrPublisher="Heldia" pageId="0" pageNumber="25" pagination="51 - 86" refId="B21" refString="Bank, RA, 2006. Towards a catalogue and bibliography of the freshwater Mollusca of Greece. Heldia 6 (1): 51 - 86" title="Towards a catalogue and bibliography of the freshwater Mollusca of Greece." volume="6" year="2006">Bank 2006</bibRefCitation>
|
||
;
|
||
<bibRefCitation author="Feher, Z" journalOrPublisher="Schriften zur Malakozoologie aus dem Haus der Natur-Cismar" pageId="0" pageNumber="25" pagination="22 - 38" refId="B52" refString="Feher, Z, Eross, ZP, 2009. Checklist of the Albanian mollusc fauna. Schriften zur Malakozoologie aus dem Haus der Natur-Cismar 25: 22 - 38" title="Checklist of the Albanian mollusc fauna." volume="25" year="2009">
|
||
<normalizedToken originalValue="Fehér">Feher</normalizedToken>
|
||
and
|
||
<normalizedToken originalValue="Erőss">Eross</normalizedToken>
|
||
2009
|
||
</bibRefCitation>
|
||
;
|
||
<bibRefCitation DOI="https://doi.org/10.2305/iucn.uk.2010-4.rlts.t165354a6005925.en" author="Zettler, ML" journalOrPublisher="Mollusca" pageId="0" pageNumber="25" refId="B167" refString="Zettler, ML, van Damme, D, 2010. Theodoxusmeridionalis. The IUCN Red List of Threatened Species 2010: e.T165354A6005925. https://doi.org/10.2305/iucn.uk.2010-4.rlts.t165354a6005925.en" title="Theodoxusmeridionalis. The IUCN Red List of Threatened Species 2010: e. T 165354 A 6005925." url="https://doi.org/10.2305/iucn.uk.2010-4.rlts.t165354a6005925.en" year="2010">Zettler and van Damme 2010</bibRefCitation>
|
||
;
|
||
<bibRefCitation author="Martinez-Orti, A" journalOrPublisher="Privately published, Tehran" pageId="0" pageNumber="25" refId="B97" refString="Martinez-Orti, A, Pedrola-Monfort, J, Robles, F, Madeira, MJ, 2015. Revision taxonomica, mediante estudios moleculares, de las especies de Theodoxus Montfort, 1810 (Gastropoda, Neritopsina) descritas en la Peninsula Iberica. In Foro Malacologico. Sociedad Espanola De Malacologia, Malaga, 25-26." title="Revision taxonomica, mediante estudios moleculares, de las especies de Theodoxus Montfort, 1810 (Gastropoda, Neritopsina) descritas en la Peninsula Iberica. In Foro Malacologico. Sociedad Espanola De Malacologia, Malaga, 25 - 26." year="2015">
|
||
<normalizedToken originalValue="Martínez-Ortí">Martinez-Orti</normalizedToken>
|
||
et al. 2015
|
||
</bibRefCitation>
|
||
;
|
||
<bibRefCitation DOI="https://doi.org/10.1111/jbi.13590" author="Sands, AF" journalOrPublisher="Journal of Biogeography" pageId="0" pageNumber="25" pagination="1274 - 1286" refId="B138" refString="Sands, AF, Sereda, SV, Stelbrink, B, Neubauer, TA, Lazarev, S, Wilke, T, Albrecht, C, 2019a. Contributions of biogeographical functions to species accumulation may change over time in refugial regions. Journal of Biogeography 46 (6): 1274 - 1286, DOI: https://doi.org/10.1111/jbi.13590" title="Contributions of biogeographical functions to species accumulation may change over time in refugial regions." url="https://doi.org/10.1111/jbi.13590" volume="46" year="2019 a">Sands et al. 2019a</bibRefCitation>
|
||
). Within our study region, the species is only noted from the freshwater drainages of the Gulf of
|
||
<normalizedToken originalValue="Gökova">Goekova</normalizedToken>
|
||
in south-west Anatolia (Fig.
|
||
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Occurrence map for Theodoxus spp. in Asia. All points conform to specimens used in the phylogenetic analyses (Suppl. material 1: Table S 1), except for T. gloeri for which no molecular data are available and only the type and paratype localities are indicated. Species are partitioned across four identical maps (A - D) to limit overlap of points as far as possible. Barring T. gloeri, closer related species are grouped together (Fig. 2)." figureDoi="10.3897/zse.96.48312.figure3" httpUri="https://binary.pensoft.net/fig/376452" pageId="0" pageNumber="25">3A</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |