treatments-xml/data/03/DC/87/03DC87A4157F41753881AC31FCBBF864.xml
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<mods:title id="CF40B9205FBFAB5345D731F930EA1F0C">Nomenclatural changes on some Mediterranean Aphaenogaster Mayr, 1853 taxa (Hymenoptera, Formicidae)</mods:title>
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<mods:affiliation id="D269BF3C6A5386F799AF0EB322C5D4E9">Department of Chemistry, Life Sciences and Environmental Sustainability, University of Parma, 43124 Parma, Italy. https: // orcid. org / 0000 - 0003 - 0684 - 6229</mods:affiliation>
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<taxonomicName id="4C754D31157F41753881AC31FBECF9BE" authority="Dalla Torre, 1893" authorityName="Dalla Torre" authorityYear="1893" box="[151,714,1590,1617]" class="Insecta" family="Formicidae" genus="Aphaenogaster" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="60" phylum="Arthropoda" rank="species" species="mauritanica">
<emphasis id="B901EAA0157F41753881AC31FBECF9BE" bold="true" box="[151,714,1590,1617]" pageId="1" pageNumber="60">
<emphasis id="B901EAA0157F41753881AC31F8CFF9BF" bold="true" box="[151,489,1590,1617]" italics="true" pageId="1" pageNumber="60">Aphaenogaster mauritanica</emphasis>
Dalla Torre, 1893
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<emphasis id="B901EAA0157F41753881AC7CF983F97D" bold="true" box="[151,165,1659,1683]" pageId="1" pageNumber="60">=</emphasis>
<taxonomicName id="4C754D31157F417538B0AC7CF8BFF97D" authority="Cagniant, 1996" authorityName="Cagniant" authorityYear="1996" box="[166,409,1659,1683]" class="Insecta" family="Formicidae" genus="Aphaenogaster" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="60" phylum="Arthropoda" rank="species" species="foreli">
<emphasis id="B901EAA0157F417538B0AC7CF9DEF97D" box="[166,248,1659,1683]" italics="true" pageId="1" pageNumber="60">A. foreli</emphasis>
<bibRefCitation id="EFE44B43157F417538E8AC7CF8BFF97D" author="Cagniant, H." box="[254,409,1659,1683]" pageId="1" pageNumber="60" pagination="67 - 85" refId="ref8347" refString="Cagniant, H. (1996) Les Aphaenogaster du Maroc (Hymenoptera: Formicidae): cle et catalogue des especes. Annales de la Societe Entomologique de France, 32, 67 - 85. https: // doi. org / 10.4000 / books. editionsmsh. 39003" type="journal article" year="1996">Cagniant, 1996</bibRefCitation>
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<taxonomicNameLabel id="A23257DB157F41753989AC7CF8DAF97D" box="[415,508,1659,1683]" pageId="1" pageNumber="60">syn. nov.</taxonomicNameLabel>
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<emphasis id="B901EAA0157F41753881AC9DF983F95C" bold="true" box="[151,165,1690,1714]" pageId="1" pageNumber="60">=</emphasis>
<taxonomicName id="4C754D31157F417538B0AC9DFB29F95C" ID-CoL="5FPLQ" authority="Emery, 1921" authorityName="Emery" authorityYear="1921" box="[166,527,1690,1714]" class="Insecta" family="Formicidae" genus="Aphaenogaster" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="60" phylum="Arthropoda" rank="subSpecies" species="gibbosa" subSpecies="homonyma">
<emphasis id="B901EAA0157F417538B0AC9DF8A1F95C" box="[166,391,1690,1714]" italics="true" pageId="1" pageNumber="60">A. gibbosa homonyma</emphasis>
<bibRefCitation id="EFE44B43157F41753998AC9DFB29F95C" author="Emery, C." box="[398,527,1690,1714]" pageId="1" pageNumber="60" pagination="1 - 94" refId="ref8599" refString="Emery, C. (1921) Hymenoptera. Fam. Formicidae. Subfam. Myrmicinae. In: Wytsman, P., Genera Insectorum, 174, 1 - 94." type="journal article" year="1921">Emery, 1921</bibRefCitation>
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<taxonomicNameLabel id="A23257DB157F41753A03AC9DFB55F95C" box="[533,627,1690,1714]" pageId="1" pageNumber="60">syn. nov.</taxonomicNameLabel>
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<emphasis id="B901EAA0157F41753881ACE7F895F914" bold="true" box="[151,435,1760,1786]" pageId="1" pageNumber="60">Investigated material—</emphasis>
<taxonomicName id="4C754D31157F417539A2ACE7FBC9F914" authorityName="Dalla Torre" authorityYear="1893" box="[436,751,1760,1786]" class="Insecta" family="Formicidae" genus="Aphaenogaster" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="60" phylum="Arthropoda" rank="species" species="mauritanica">
<emphasis id="B901EAA0157F417539A2ACE7FBC9F914" box="[436,751,1760,1786]" italics="true" pageId="1" pageNumber="60">Aphaenogaster mauritanica</emphasis>
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CASENT0904173,
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CASENT
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<bibRefCitation id="EFE44B43157F41753D29AD4BF831F864" author="Schifani, E. &amp; Alicata, A. &amp; Menchetti, M. &amp; Borowiec, L. &amp; Fisher, B. L. &amp; Karaman, K. &amp; Kiran, C. &amp; Oueslati, W. &amp; Salata, S. &amp; Blatrix, R." pageId="1" pageNumber="60" pagination="627 - 648" refId="ref9100" refString="Schifani, E., Alicata, A., Menchetti, M., Borowiec, L., Fisher, B. L., Karaman, K., Kiran, C., Oueslati, W., Salata, S. &amp; Blatrix, R. (2022) Revisiting the morphological species groups of West-Palearctic Aphaenogaster ants (Hymenoptera: Formicidae) from a phylogenetic perspective: toward an evolutionary classification. Arhtropod Systematics and Phylogeny, 80, 627 - 648. https: // doi. org / 10.3897 / asp. 80. e 84428" type="journal article" year="2022">
Schifani
<emphasis id="B901EAA0157F41753881AD76F9F7F864" box="[151,209,1904,1930]" italics="true" pageId="1" pageNumber="60">et al.</emphasis>
2022
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</materialsCitation>
;
<materialsCitation id="3B1D3CEF157F4175393AAD77FCBFF864" ID-GBIF-Occurrence="4525969304" box="[300,1433,1904,1931]" collectingDate="2017-06-07" collectionCode="ESPC" collectorName="W. Oueslati" country="Tunisia" location="Cap Serrat" pageId="1" pageNumber="60" specimenCount="12" specimenCount-female="12" stateProvince="Bizerte" typeStatus="syntype">
<specimenCount id="9D73FD3B157F4175393AAD77F852F864" box="[300,372,1904,1930]" pageId="1" pageNumber="60" type="female">12♀♀</specimenCount>
,
<location id="8EAA6069157F41753994AD77F8D8F864" LSID="urn:lsid:plazi:treatment:03DC87A4157F41753881AC31FCBBF864:8EAA6069157F41753994AD77F8D8F864" box="[386,510,1904,1930]" country="Tunisia" name="Cap Serrat" pageId="1" pageNumber="60" stateProvince="Bizerte">Cap Serrat</location>
,
<collectingRegion id="49B1F850157F41753A1DAD77FB7BF864" box="[523,605,1904,1930]" country="Tunisia" name="Bizerte" pageId="1" pageNumber="60">Biserta</collectingRegion>
,
<collectingCountry id="F3627622157F41753A7CAD77FBE6F864" box="[618,704,1904,1930]" name="Tunisia" pageId="1" pageNumber="60">Tunisia</collectingCountry>
,
<date id="FFCB1072157F41753ADBAD77FA68F864" box="[717,846,1904,1930]" pageId="1" pageNumber="60" value="2017-06-07">
<collectingDate id="EF8FE99A157F41753ADBAD77FA68F864" box="[717,846,1904,1930]" pageId="1" pageNumber="60" value="2017-06-07">07.VI.2017</collectingDate>
</date>
,
<collectorName id="26805364157F41753B4AAD76FAC6F864" box="[860,992,1904,1931]" pageId="1" pageNumber="60">W. Oueslati</collectorName>
legit (
<collectionCode id="ED64AE77157F41753C3AAD77FD54F864" box="[1068,1138,1904,1930]" pageId="1" pageNumber="60">ESPC</collectionCode>
) (in
<bibRefCitation id="EFE44B43157F41753CBDAD77FCB7F864" author="Schifani, E. &amp; Alicata, A. &amp; Menchetti, M. &amp; Borowiec, L. &amp; Fisher, B. L. &amp; Karaman, K. &amp; Kiran, C. &amp; Oueslati, W. &amp; Salata, S. &amp; Blatrix, R." box="[1195,1425,1904,1930]" pageId="1" pageNumber="60" pagination="627 - 648" refId="ref9100" refString="Schifani, E., Alicata, A., Menchetti, M., Borowiec, L., Fisher, B. L., Karaman, K., Kiran, C., Oueslati, W., Salata, S. &amp; Blatrix, R. (2022) Revisiting the morphological species groups of West-Palearctic Aphaenogaster ants (Hymenoptera: Formicidae) from a phylogenetic perspective: toward an evolutionary classification. Arhtropod Systematics and Phylogeny, 80, 627 - 648. https: // doi. org / 10.3897 / asp. 80. e 84428" type="journal article" year="2022">
Schifani
<emphasis id="B901EAA0157F41753D07AD76FC6DF864" box="[1297,1355,1904,1930]" italics="true" pageId="1" pageNumber="60">et al.</emphasis>
2022
</bibRefCitation>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C36F6539157C417738D1AFBDFDFAFE1B" lastPageId="3" lastPageNumber="62" pageId="2" pageNumber="61" type="description">
<paragraph id="8BCA36B2157C417738D1AFBDFDFAFE1B" blockId="2.[151,1437,1466,2032]" lastBlockId="3.[151,1437,151,501]" lastPageId="3" lastPageNumber="62" pageId="2" pageNumber="61">
In the Maghreb, the
<taxonomicName id="4C754D31157C417639B1AFBDF8D0FA3A" ID-CoL="F9VY" baseAuthorityName="Nylander" baseAuthorityYear="1849" box="[423,502,1466,1492]" class="Insecta" family="Formicidae" genus="Aphaenogaster" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="61" phylum="Arthropoda" rank="species" species="pallida">
<emphasis id="B901EAA0157C417639B1AFBDF8D0FA3A" box="[423,502,1466,1492]" italics="true" pageId="2" pageNumber="61">pallida</emphasis>
</taxonomicName>
group includes a morphospecies with short propodeal spines which was temporarily named “
<taxonomicName id="4C754D31157C417638E2AFD9FBB2FA16" authority="sensu Forel, 1890" authorityName="sensu Forel" authorityYear="1890" box="[244,660,1502,1528]" class="Insecta" family="Formicidae" genus="Aphaenogaster" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="61" phylum="Arthropoda" rank="species" species="subterraneoides">
<emphasis id="B901EAA0157C417638E2AFD9FB2DFA19" box="[244,523,1502,1528]" italics="true" pageId="2" pageNumber="61">A. subterraneoides sensu</emphasis>
<bibRefCitation id="EFE44B43157C41763A04AFD9FBB2FA16" author="Forel, A." box="[530,660,1502,1528]" pageId="2" pageNumber="61" pagination="61 - 75" refId="ref8632" refString="Forel, A. (1890) Fourmis de Tunisie et de l'Algerie orientale. Annales de la Societe entomologique de Belgique, 34, 61 - 75." type="journal article" year="1890">Forel, 1890</bibRefCitation>
</taxonomicName>
” by
<bibRefCitation id="EFE44B43157C41763AD1AFD9FD29FA16" author="Cagniant, H. &amp; Espadaler, X." box="[711,1039,1502,1528]" pageId="2" pageNumber="61" pagination="89 - 93" refId="ref8314" refString="Cagniant, H. &amp; Espadaler, X. (1993) Liste des especes de fourmis du Maroc. Actes des Colloques. Insectes Sociaux, 8, 89 - 93." type="journal article" year="1993">Cagniant &amp; Espadaler (1993)</bibRefCitation>
, without providing any information about its identity. Later,
<bibRefCitation id="EFE44B43157C417639A5AC05FB4BF9F2" author="Cagniant, H." box="[435,621,1538,1564]" pageId="2" pageNumber="61" pagination="67 - 85" refId="ref8347" refString="Cagniant, H. (1996) Les Aphaenogaster du Maroc (Hymenoptera: Formicidae): cle et catalogue des especes. Annales de la Societe Entomologique de France, 32, 67 - 85. https: // doi. org / 10.4000 / books. editionsmsh. 39003" type="journal article" year="1996">Cagniant (1996)</bibRefCitation>
claimed that
<bibRefCitation id="EFE44B43157C41763B1BAC05FABAF9F2" author="Forel, A." box="[781,924,1538,1564]" pageId="2" pageNumber="61" pagination="61 - 75" refId="ref8632" refString="Forel, A. (1890) Fourmis de Tunisie et de l'Algerie orientale. Annales de la Societe entomologique de Belgique, 34, 61 - 75." type="journal article" year="1890">Forel (1890)</bibRefCitation>
had treated
<taxonomicName id="4C754D31157C41763C39AC05FDC5F9F2" authorityName="sensu Forel" authorityYear="1890" box="[1071,1251,1538,1564]" class="Insecta" family="Formicidae" genus="Aphaenogaster" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="61" phylum="Arthropoda" rank="species" species="subterraneoides">
<emphasis id="B901EAA0157C41763C39AC05FDC5F9F2" box="[1071,1251,1538,1564]" italics="true" pageId="2" pageNumber="61">subterraneoides</emphasis>
</taxonomicName>
as a subspecies of
<taxonomicName id="4C754D31157C417638AEAC21F80EF9AE" baseAuthorityName="Nylander" baseAuthorityYear="1849" box="[184,296,1574,1600]" class="Insecta" family="Formicidae" genus="Aphaenogaster" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="61" phylum="Arthropoda" rank="species" species="pallida">
<emphasis id="B901EAA0157C417638AEAC21F80EF9AE" box="[184,296,1574,1600]" italics="true" pageId="2" pageNumber="61">A. pallida</emphasis>
</taxonomicName>
and that the name was preoccupied with Emerys description of
<taxonomicName id="4C754D31157C41763C03AC21F9E9F98A" authority="Emery, 1881" authorityName="Emery" authorityYear="1881" class="Insecta" family="Formicidae" genus="Aphaenogaster" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="61" phylum="Arthropoda" rank="subSpecies" species="pallida" subSpecies="subterraneoides">
<emphasis id="B901EAA0157C41763C03AC21FC65F9AE" box="[1045,1347,1574,1600]" italics="true" pageId="2" pageNumber="61">A. pallida subterraneoides</emphasis>
<bibRefCitation id="EFE44B43157C41763D5AAC21F9E9F98A" author="Emery, C." pageId="2" pageNumber="61" pagination="525 - 535" refId="ref8427" refString="Emery, C. (1881) Viaggio ad Assab nel Mar Rosso dei Signori G. Doria ed O. Beccari con il R. Avviso &quot; Esploratore &quot; dal 16 novembre 1879 al 26 febbraio 1880. I. Formiche. Annali del Museo Civico di Storia Naturale Giacomo Doria, 16, 525 - 535." type="journal article" year="1881">Emery, 1881</bibRefCitation>
</taxonomicName>
from
<collectingCountry id="F3627622157C41763901AC4DF840F98A" box="[279,358,1610,1636]" name="Greece" pageId="2" pageNumber="61">Greece</collectingCountry>
(
<bibRefCitation id="EFE44B43157C41763961AC4DFB28F98A" author="Emery, C." box="[375,526,1610,1636]" pageId="2" pageNumber="61" pagination="525 - 535" refId="ref8427" refString="Emery, C. (1881) Viaggio ad Assab nel Mar Rosso dei Signori G. Doria ed O. Beccari con il R. Avviso &quot; Esploratore &quot; dal 16 novembre 1879 al 26 febbraio 1880. I. Formiche. Annali del Museo Civico di Storia Naturale Giacomo Doria, 16, 525 - 535." type="journal article" year="1881">Emery, 1881</bibRefCitation>
). On this basis, he established the name
<taxonomicName id="4C754D31157C41763BF7AC4DFD18F98A" authorityName="Cagniant" authorityYear="1996" box="[993,1086,1610,1636]" class="Insecta" family="Formicidae" genus="Aphaenogaster" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="61" phylum="Arthropoda" rank="species" species="foreli">
<emphasis id="B901EAA0157C41763BF7AC4DFD18F98A" box="[993,1086,1610,1636]" italics="true" pageId="2" pageNumber="61">A. foreli</emphasis>
</taxonomicName>
to replace the taxon described by
<bibRefCitation id="EFE44B43157C417638A8AC69F877F966" author="Forel, A." box="[190,337,1646,1672]" pageId="2" pageNumber="61" pagination="61 - 75" refId="ref8632" refString="Forel, A. (1890) Fourmis de Tunisie et de l'Algerie orientale. Annales de la Societe entomologique de Belgique, 34, 61 - 75." type="journal article" year="1890">Forel (1890)</bibRefCitation>
. However,
<bibRefCitation id="EFE44B43157C417639C0AC69FB40F966" author="Forel, A." box="[470,614,1646,1672]" pageId="2" pageNumber="61" pagination="61 - 75" refId="ref8632" refString="Forel, A. (1890) Fourmis de Tunisie et de l'Algerie orientale. Annales de la Societe entomologique de Belgique, 34, 61 - 75." type="journal article" year="1890">Forel (1890)</bibRefCitation>
had used the spelling
<emphasis id="B901EAA0157C41763B66AC69FD3EF966" box="[880,1048,1646,1672]" italics="true" pageId="2" pageNumber="61">subterranoides</emphasis>
(and not “
<taxonomicName id="4C754D31157C41763C8CAC69FC68F966" authorityName="sensu Forel" authorityYear="1890" box="[1178,1358,1646,1672]" class="Insecta" family="Formicidae" genus="Aphaenogaster" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="61" phylum="Arthropoda" rank="species" species="subterraneoides">
<emphasis id="B901EAA0157C41763C8CAC69FC68F966" box="[1178,1358,1646,1672]" italics="true" pageId="2" pageNumber="61">subterraneoides</emphasis>
</taxonomicName>
”) as a smaller and more brownish variety of
<taxonomicName id="4C754D31157C41763A40AC95FAD9F942" authority="(Roger, 1859)" baseAuthorityName="Roger" baseAuthorityYear="1859" box="[598,1023,1681,1708]" class="Insecta" family="Formicidae" genus="Aphaenogaster" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="61" phylum="Arthropoda" rank="species" species="striola">
<emphasis id="B901EAA0157C41763A40AC95FA75F942" box="[598,851,1682,1708]" italics="true" pageId="2" pageNumber="61">Aphaenogaster striola</emphasis>
(Roger, 1859)
</taxonomicName>
, a junior synonym of the European species
<taxonomicName id="4C754D31157C417638F8ACB1FBE1F93E" authority="(Latreille, 1798) (Emery 1895)" authorityName="Emery" authorityYear="1895" baseAuthorityName="Latreille" baseAuthorityYear="1798" box="[238,711,1717,1744]" class="Insecta" family="Formicidae" genus="Aphaenogaster" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="61" phylum="Arthropoda" rank="species" species="gibbosa">
<emphasis id="B901EAA0157C417638F8ACB1F840F93E" box="[238,358,1718,1744]" italics="true" pageId="2" pageNumber="61">A. gibbosa</emphasis>
(Latreille, 1798) (
<bibRefCitation id="EFE44B43157C41763A26ACB1FB99F93E" author="Emery, C." box="[560,703,1717,1744]" pageId="2" pageNumber="61" pagination="257 - 360" refId="ref8486" refString="Emery, C. (1895) Beitrage zur Kenntniss der nordamerikanischen Ameisenfauna. (Schluss). Zoologische Jahrbucher. Abteilung fur Systematik, Geographie und Biologie der Tiere, 8, 257 - 360." type="journal article" year="1895">Emery 1895</bibRefCitation>
)
</taxonomicName>
. Moreover,
<bibRefCitation id="EFE44B43157C41763B5AACB1FD24F93E" author="Cagniant, H." box="[844,1026,1718,1744]" pageId="2" pageNumber="61" pagination="67 - 85" refId="ref8347" refString="Cagniant, H. (1996) Les Aphaenogaster du Maroc (Hymenoptera: Formicidae): cle et catalogue des especes. Annales de la Societe Entomologique de France, 32, 67 - 85. https: // doi. org / 10.4000 / books. editionsmsh. 39003" type="journal article" year="1996">Cagniant (1996)</bibRefCitation>
did not take into account that
<bibRefCitation id="EFE44B43157C41763D44ACB1F9C4F91A" author="Emery, C." pageId="2" pageNumber="61" pagination="1 - 94" refId="ref8599" refString="Emery, C. (1921) Hymenoptera. Fam. Formicidae. Subfam. Myrmicinae. In: Wytsman, P., Genera Insectorum, 174, 1 - 94." type="journal article" year="1921">Emery (1921)</bibRefCitation>
had already introduced the name
<taxonomicName id="4C754D31157C41763A49ACDDFAACF91A" authorityName="Emery" authorityYear="1921" box="[607,906,1754,1780]" class="Insecta" family="Formicidae" genus="Aphaenogaster" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="61" phylum="Arthropoda" rank="variety" species="gibbosa" variety="homonyma">
<emphasis id="B901EAA0157C41763A49ACDDFBFEF91A" box="[607,728,1754,1780]" italics="true" pageId="2" pageNumber="61">A. gibbosa</emphasis>
var.
<emphasis id="B901EAA0157C41763B19ACDDFAACF91A" box="[783,906,1754,1780]" italics="true" pageId="2" pageNumber="61">homonyma</emphasis>
</taxonomicName>
with the explicit intent of replacing the
<taxonomicName id="4C754D31157C41763D58ACDDF819F8F6" class="Insecta" family="Formicidae" genus="Aphaenogaster" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="61" phylum="Arthropoda" rank="variety" species="striola" variety="subterranoides">
variety
<emphasis id="B901EAA0157C41763881ACF9F819F8F6" box="[151,319,1790,1816]" italics="true" pageId="2" pageNumber="61">subterranoides</emphasis>
</taxonomicName>
described by
<bibRefCitation id="EFE44B43157C417639C8ACF9FB4AF8F6" author="Forel, A." box="[478,620,1790,1816]" pageId="2" pageNumber="61" pagination="61 - 75" refId="ref8632" refString="Forel, A. (1890) Fourmis de Tunisie et de l'Algerie orientale. Annales de la Societe entomologique de Belgique, 34, 61 - 75." type="journal article" year="1890">Forel (1890)</bibRefCitation>
(also there mentioning it as
<taxonomicName id="4C754D31157C41763BA6ACF9FD42F8F6" authorityName="sensu Forel" authorityYear="1890" box="[944,1124,1790,1816]" class="Insecta" family="Formicidae" genus="Aphaenogaster" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="61" phylum="Arthropoda" rank="species" species="subterraneoides">
<emphasis id="B901EAA0157C41763BA6ACF9FD42F8F6" box="[944,1124,1790,1816]" italics="true" pageId="2" pageNumber="61">subterraneoides</emphasis>
</taxonomicName>
). In the description,
<bibRefCitation id="EFE44B43157C41763D44ACF9F9C4F8D2" author="Emery, C." pageId="2" pageNumber="61" pagination="1 - 94" refId="ref8599" refString="Emery, C. (1921) Hymenoptera. Fam. Formicidae. Subfam. Myrmicinae. In: Wytsman, P., Genera Insectorum, 174, 1 - 94." type="journal article" year="1921">Emery (1921)</bibRefCitation>
did not explain how the
<taxonomicName id="4C754D31157C417639E6AD25FB99F8D2" authorityName="Emery" authorityYear="1921" box="[496,703,1826,1852]" class="Insecta" family="Formicidae" genus="Aphaenogaster" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="61" phylum="Arthropoda" rank="variety" species="gibbosa" variety="homonyma">
variety
<emphasis id="B901EAA0157C41763A52AD25FB99F8D2" box="[580,703,1826,1852]" italics="true" pageId="2" pageNumber="61">homonyma</emphasis>
</taxonomicName>
differs from other members of the
<taxonomicName id="4C754D31157C41763C29AD25FDBEF8D2" authorityName="Emery" authorityYear="1895" baseAuthorityName="Latreille" baseAuthorityYear="1798" box="[1087,1176,1826,1852]" class="Insecta" family="Formicidae" genus="Aphaenogaster" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="61" phylum="Arthropoda" rank="species" species="gibbosa">
<emphasis id="B901EAA0157C41763C29AD25FDBEF8D2" box="[1087,1176,1826,1852]" italics="true" pageId="2" pageNumber="61">gibbosa</emphasis>
</taxonomicName>
group. An examination of the
<specimenCount id="9D73FD3B157C417638F7AD41F87FF88E" box="[225,345,1862,1888]" pageId="2" pageNumber="61" type="worker">six worker</specimenCount>
specimens in the collection of Forel labeled “
<taxonomicName id="4C754D31157C41763B70AD41FDA6F88E" box="[870,1152,1862,1888]" class="Insecta" family="Formicidae" genus="Aphaenogaster" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="61" phylum="Arthropoda" rank="subSpecies" species="striola" subSpecies="subterranoides">
<emphasis id="B901EAA0157C41763B70AD41FDA6F88E" box="[870,1152,1862,1888]" italics="true" pageId="2" pageNumber="61">A. striola subterranoides</emphasis>
</taxonomicName>
” immediately confirmed that they belong to the
<taxonomicName id="4C754D31157C4176398BAD6DF8D0F86A" authorityName="Emery" authorityYear="1895" baseAuthorityName="Latreille" baseAuthorityYear="1798" box="[413,502,1898,1924]" class="Insecta" family="Formicidae" genus="Aphaenogaster" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="61" phylum="Arthropoda" rank="species" species="gibbosa">
<emphasis id="B901EAA0157C4176398BAD6DF8D0F86A" box="[413,502,1898,1924]" italics="true" pageId="2" pageNumber="61">gibbosa</emphasis>
</taxonomicName>
group, which is very distinct from the
<taxonomicName id="4C754D31157C41763BA6AD6DFAD9F86A" baseAuthorityName="Nylander" baseAuthorityYear="1849" box="[944,1023,1898,1924]" class="Insecta" family="Formicidae" genus="Aphaenogaster" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="61" phylum="Arthropoda" rank="species" species="pallida">
<emphasis id="B901EAA0157C41763BA6AD6DFAD9F86A" box="[944,1023,1898,1924]" italics="true" pageId="2" pageNumber="61">pallida</emphasis>
</taxonomicName>
group that includes the Greek taxon described by Emery (
<bibRefCitation id="EFE44B43157C41763995AD89FB36F846" author="Emery, C." box="[387,528,1934,1960]" pageId="2" pageNumber="61" pagination="525 - 535" refId="ref8427" refString="Emery, C. (1881) Viaggio ad Assab nel Mar Rosso dei Signori G. Doria ed O. Beccari con il R. Avviso &quot; Esploratore &quot; dal 16 novembre 1879 al 26 febbraio 1880. I. Formiche. Annali del Museo Civico di Storia Naturale Giacomo Doria, 16, 525 - 535." type="journal article" year="1881">Emery 1881</bibRefCitation>
;
<bibRefCitation id="EFE44B43157C41763A0DAD89FBDFF846" author="Schifani, E. &amp; Alicata, A. &amp; Menchetti, M. &amp; Borowiec, L. &amp; Fisher, B. L. &amp; Karaman, K. &amp; Kiran, C. &amp; Oueslati, W. &amp; Salata, S. &amp; Blatrix, R." box="[539,761,1934,1960]" pageId="2" pageNumber="61" pagination="627 - 648" refId="ref9100" refString="Schifani, E., Alicata, A., Menchetti, M., Borowiec, L., Fisher, B. L., Karaman, K., Kiran, C., Oueslati, W., Salata, S. &amp; Blatrix, R. (2022) Revisiting the morphological species groups of West-Palearctic Aphaenogaster ants (Hymenoptera: Formicidae) from a phylogenetic perspective: toward an evolutionary classification. Arhtropod Systematics and Phylogeny, 80, 627 - 648. https: // doi. org / 10.3897 / asp. 80. e 84428" type="journal article" year="2022">
Schifani
<emphasis id="B901EAA0157C41763A69AD89FB90F846" box="[639,694,1934,1960]" italics="true" pageId="2" pageNumber="61">et al.</emphasis>
2022
</bibRefCitation>
). Recently, on AntCat it was proposed to consider
<taxonomicName id="4C754D31157C41763D33AD89FC59F846" authorityName="Cagniant" authorityYear="1996" box="[1317,1407,1934,1960]" class="Insecta" family="Formicidae" genus="Aphaenogaster" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="61" phylum="Arthropoda" rank="species" species="foreli">
<emphasis id="B901EAA0157C41763D33AD89FC59F846" box="[1317,1407,1934,1960]" italics="true" pageId="2" pageNumber="61">A. foreli</emphasis>
</taxonomicName>
as a junior synonym of
<taxonomicName id="4C754D31157C41763969ADB5FB5EF822" authorityName="Emery" authorityYear="1921" box="[383,632,1970,1996]" class="Insecta" family="Formicidae" genus="Aphaenogaster" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="61" phylum="Arthropoda" rank="subSpecies" species="gibbosa" subSpecies="homonyma">
<emphasis id="B901EAA0157C41763969ADB5FB5EF822" box="[383,632,1970,1996]" italics="true" pageId="2" pageNumber="61">A. gibbosa homonyma</emphasis>
</taxonomicName>
, and since
<bibRefCitation id="EFE44B43157C41763AE2ADB5FA8CF822" author="Cagniant, H." box="[756,938,1970,1996]" pageId="2" pageNumber="61" pagination="67 - 85" refId="ref8347" refString="Cagniant, H. (1996) Les Aphaenogaster du Maroc (Hymenoptera: Formicidae): cle et catalogue des especes. Annales de la Societe Entomologique de France, 32, 67 - 85. https: // doi. org / 10.4000 / books. editionsmsh. 39003" type="journal article" year="1996">Cagniant (1996)</bibRefCitation>
had treated
<taxonomicName id="4C754D31157C41763C25ADB5FDABF822" authorityName="Cagniant" authorityYear="1996" box="[1075,1165,1970,1996]" class="Insecta" family="Formicidae" genus="Aphaenogaster" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="61" phylum="Arthropoda" rank="species" species="foreli">
<emphasis id="B901EAA0157C41763C25ADB5FDABF822" box="[1075,1165,1970,1996]" italics="true" pageId="2" pageNumber="61">A. foreli</emphasis>
</taxonomicName>
as a species-rank taxon, to treat it as a full species,
<taxonomicName id="4C754D31157C417639D7ADD1FBD8F81E" authority="(Bolton 2022)" baseAuthorityName="Bolton" baseAuthorityYear="2022" box="[449,766,2006,2032]" class="Insecta" family="Formicidae" genus="Aphaenogaster" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="61" phylum="Arthropoda" rank="species" species="homonyma">
<emphasis id="B901EAA0157C417639D7ADD1FB7DF81E" box="[449,603,2006,2032]" italics="true" pageId="2" pageNumber="61">A. homonyma</emphasis>
(
<bibRefCitation id="EFE44B43157C41763A7CADD1FBDEF81E" author="Bolton, B." box="[618,760,2006,2032]" pageId="2" pageNumber="61" refId="ref8131" refString="Bolton, B. (2022) An online catalog of the ants of the world. Available from https: // antcat. org." type="url" year="2022">Bolton 2022</bibRefCitation>
)
</taxonomicName>
—even though
<bibRefCitation id="EFE44B43157C41763BBBADD1FD45F81E" author="Cagniant, H." box="[941,1123,2006,2032]" pageId="2" pageNumber="61" pagination="67 - 85" refId="ref8347" refString="Cagniant, H. (1996) Les Aphaenogaster du Maroc (Hymenoptera: Formicidae): cle et catalogue des especes. Annales de la Societe Entomologique de France, 32, 67 - 85. https: // doi. org / 10.4000 / books. editionsmsh. 39003" type="journal article" year="1996">Cagniant (1996)</bibRefCitation>
clearly referred to a
<taxonomicName id="4C754D31157C41763D5BADD1FCBAF81E" baseAuthorityName="Nylander" baseAuthorityYear="1849" box="[1357,1436,2006,2032]" class="Insecta" family="Formicidae" genus="Aphaenogaster" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="61" phylum="Arthropoda" rank="species" species="pallida">
<emphasis id="B901EAA0157C41763D5BADD1FCBAF81E" box="[1357,1436,2006,2032]" italics="true" pageId="2" pageNumber="61">pallida</emphasis>
</taxonomicName>
group morphospecies. Based on comparisons between
<typeStatus id="54CE8810157D41773B1CAA90FA6EFF5F" box="[778,840,151,177]" pageId="3" pageNumber="62">types</typeStatus>
, the morphology of the workers from the collection of Forel is fully congruent with the only other
<taxonomicName id="4C754D31157D41773AB1AABCFA26FF3B" authorityName="Emery" authorityYear="1895" baseAuthorityName="Latreille" baseAuthorityYear="1798" box="[679,768,187,213]" class="Insecta" family="Formicidae" genus="Aphaenogaster" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="62" phylum="Arthropoda" rank="species" species="gibbosa">
<emphasis id="B901EAA0157D41773AB1AABCFA26FF3B" box="[679,768,187,213]" italics="true" pageId="3" pageNumber="62">gibbosa</emphasis>
</taxonomicName>
group taxon in the Eastern Maghreb,
<taxonomicName id="4C754D31157D41773CB9AABCF83BFF17" authority="Dalla Torre, 1893" authorityName="Dalla Torre" authorityYear="1893" class="Insecta" family="Formicidae" genus="Aphaenogaster" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="62" phylum="Arthropoda" rank="species" species="mauritanica">
<emphasis id="B901EAA0157D41773CB9AABCFC7EFF3A" box="[1199,1368,187,213]" italics="true" pageId="3" pageNumber="62">A. mauritanica</emphasis>
Dalla Torre, 1893
</taxonomicName>
, and no characters to separate the two have ever been described. They are easily distinguishable from the European
<taxonomicName id="4C754D31157D41773922AB04F88BFEF3" authorityName="Emery" authorityYear="1895" baseAuthorityName="Latreille" baseAuthorityYear="1798" box="[308,429,259,285]" class="Insecta" family="Formicidae" genus="Aphaenogaster" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="62" phylum="Arthropoda" rank="species" species="gibbosa">
<emphasis id="B901EAA0157D41773922AB04F88BFEF3" box="[308,429,259,285]" italics="true" pageId="3" pageNumber="62">A. gibbosa</emphasis>
</taxonomicName>
and from the Moroccan
<taxonomicName id="4C754D31157D41773AD1AB04FAFFFEF3" authority="Santschi, 1923" authorityName="Santschi" authorityYear="1923" box="[711,985,259,285]" class="Insecta" family="Formicidae" genus="Aphaenogaster" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="62" phylum="Arthropoda" rank="species" species="theryi">
<emphasis id="B901EAA0157D41773AD1AB04FA0EFEF3" box="[711,808,259,285]" italics="true" pageId="3" pageNumber="62">A. theryi</emphasis>
Santschi, 1923
</taxonomicName>
, but appear indistinguishable from each other as recognized in the only existing revision of the group (
<bibRefCitation id="EFE44B43157D41773B51AB20FD4AFEAF" author="Salata, S. &amp; Borowiec, L." box="[839,1132,295,321]" pageId="3" pageNumber="62" pagination="010002" refId="ref8884" refString="Salata, S. &amp; Borowiec, L. (2018 a) Redescription of Aphaenogaster muschtaidica Emery, 1908 with a key to gibbosa species group. Asian Myrmecology, 10, e 010002. https: // doi. org / 10.20362 / am. 010002" type="journal article" year="2018">Salata &amp; Borowiec 2018a</bibRefCitation>
). Accordingly, we establish
<taxonomicName id="4C754D31157D41773881AB4CF866FE8A" authorityName="Dalla Torre" authorityYear="1893" box="[151,320,331,357]" class="Insecta" family="Formicidae" genus="Aphaenogaster" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="62" phylum="Arthropoda" rank="species" species="mauritanica">
<emphasis id="B901EAA0157D41773881AB4CF866FE8A" box="[151,320,331,357]" italics="true" pageId="3" pageNumber="62">A. mauritanica</emphasis>
</taxonomicName>
as the senior synonym of both
<taxonomicName id="4C754D31157D41773AB2AB4CFA26FE8B" authorityName="Cagniant" authorityYear="1996" box="[676,768,331,357]" class="Insecta" family="Formicidae" genus="Aphaenogaster" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="62" phylum="Arthropoda" rank="species" species="foreli">
<emphasis id="B901EAA0157D41773AB2AB4CFA26FE8B" box="[676,768,331,357]" italics="true" pageId="3" pageNumber="62">A. foreli</emphasis>
</taxonomicName>
and
<taxonomicName id="4C754D31157D41773B2EAB4CFD15FE8B" authorityName="Emery" authorityYear="1921" box="[824,1075,331,357]" class="Insecta" family="Formicidae" genus="Aphaenogaster" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="62" phylum="Arthropoda" rank="subSpecies" species="gibbosa" subSpecies="homonyma">
<emphasis id="B901EAA0157D41773B2EAB4CFD15FE8B" box="[824,1075,331,357]" italics="true" pageId="3" pageNumber="62">A. gibbosa homonyma</emphasis>
</taxonomicName>
. The Maghrebian
<taxonomicName id="4C754D31157D41773D12AB4CFC75FE8B" baseAuthorityName="Nylander" baseAuthorityYear="1849" box="[1284,1363,331,357]" class="Insecta" family="Formicidae" genus="Aphaenogaster" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="62" phylum="Arthropoda" rank="species" species="pallida">
<emphasis id="B901EAA0157D41773D12AB4CFC75FE8B" box="[1284,1363,331,357]" italics="true" pageId="3" pageNumber="62">pallida</emphasis>
</taxonomicName>
group morphospecies with short spines also needs a formal name: according to available morphological and phylogenetic evidence, it should be conspecific with
<taxonomicName id="4C754D31157D41773A40AB94FA5CFE43" authority="Emery, 1924" authorityName="Emery" authorityYear="1924" box="[598,890,403,429]" class="Insecta" family="Formicidae" genus="Aphaenogaster" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="62" phylum="Arthropoda" rank="species" species="dulcineae">
<emphasis id="B901EAA0157D41773A40AB94FBC5FE43" box="[598,739,403,429]" italics="true" pageId="3" pageNumber="62">A. dulcineae</emphasis>
Emery, 1924
</taxonomicName>
(
<bibRefCitation id="EFE44B43157D41773B9FAB94FD4BFE43" author="Schifani, E. &amp; Alicata, A. &amp; Menchetti, M. &amp; Borowiec, L. &amp; Fisher, B. L. &amp; Karaman, K. &amp; Kiran, C. &amp; Oueslati, W. &amp; Salata, S. &amp; Blatrix, R." box="[905,1133,403,429]" pageId="3" pageNumber="62" pagination="627 - 648" refId="ref9100" refString="Schifani, E., Alicata, A., Menchetti, M., Borowiec, L., Fisher, B. L., Karaman, K., Kiran, C., Oueslati, W., Salata, S. &amp; Blatrix, R. (2022) Revisiting the morphological species groups of West-Palearctic Aphaenogaster ants (Hymenoptera: Formicidae) from a phylogenetic perspective: toward an evolutionary classification. Arhtropod Systematics and Phylogeny, 80, 627 - 648. https: // doi. org / 10.3897 / asp. 80. e 84428" type="journal article" year="2022">
Schifani
<emphasis id="B901EAA0157D41773BE6AB94FD0FFE43" box="[1008,1065,403,429]" italics="true" pageId="3" pageNumber="62">et al.</emphasis>
2022
</bibRefCitation>
;
<figureCitation id="134E2A37157D41773C6EAB94FD9FFE43" box="[1144,1209,403,429]" captionStart="FIGURE 1" captionStartId="2.[152,255,1323,1347]" captionTargetBox="[151,1436,381,1300]" captionTargetId="figure-123@2.[151,1436,381,1300]" captionTargetPageId="2" captionText="FIGURE 1. Workers of Aphaenogaster dulcineae. A,B: specimen from Tunisia (sequenced as A. foreli by Schifani et al. 2022). C,D: Syntype specimen from Spain (Aphaenogaster pallida dulcineae, CASENT0913113, photo from Zach Lieberman available on www.antweb.org)." figureDoi="http://doi.org/10.5281/zenodo.7891707" httpUri="https://zenodo.org/record/7891707/files/figure.png" pageId="3" pageNumber="62">Fig. 1</figureCitation>
). The southernmost range of
<taxonomicName id="4C754D31157D417738EAABB0F8AFFE3F" authorityName="Emery" authorityYear="1924" box="[252,393,439,465]" class="Insecta" family="Formicidae" genus="Aphaenogaster" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="62" phylum="Arthropoda" rank="species" species="dulcineae">
<emphasis id="B901EAA0157D417738EAABB0F8AFFE3F" box="[252,393,439,465]" italics="true" pageId="3" pageNumber="62">A. dulcineae</emphasis>
</taxonomicName>
was hitherto considered to end in southern
<collectingCountry id="F3627622157D41773B60ABB0FA91FE3F" box="[886,951,439,465]" name="Georgia" pageId="3" pageNumber="62">Iberia</collectingCountry>
(Tarifa), while surprisingly no comparison was made until recently with the population inhabiting the Maghreb (
<bibRefCitation id="EFE44B43157D41773B89ABDCFDA6FE1B" author="Schifani, E. &amp; Alicata, A. &amp; Menchetti, M. &amp; Borowiec, L. &amp; Fisher, B. L. &amp; Karaman, K. &amp; Kiran, C. &amp; Oueslati, W. &amp; Salata, S. &amp; Blatrix, R." box="[927,1152,475,501]" pageId="3" pageNumber="62" pagination="627 - 648" refId="ref9100" refString="Schifani, E., Alicata, A., Menchetti, M., Borowiec, L., Fisher, B. L., Karaman, K., Kiran, C., Oueslati, W., Salata, S. &amp; Blatrix, R. (2022) Revisiting the morphological species groups of West-Palearctic Aphaenogaster ants (Hymenoptera: Formicidae) from a phylogenetic perspective: toward an evolutionary classification. Arhtropod Systematics and Phylogeny, 80, 627 - 648. https: // doi. org / 10.3897 / asp. 80. e 84428" type="journal article" year="2022">
Schifani
<emphasis id="B901EAA0157D41773C13ABDCFD1BFE1B" box="[1029,1085,475,501]" italics="true" pageId="3" pageNumber="62">et al.</emphasis>
2022
</bibRefCitation>
;
<figureCitation id="134E2A37157D41773C9DABDCFDEDFE1B" box="[1163,1227,475,501]" captionStart="FIGURE 1" captionStartId="2.[152,255,1323,1347]" captionTargetBox="[151,1436,381,1300]" captionTargetId="figure-123@2.[151,1436,381,1300]" captionTargetPageId="2" captionText="FIGURE 1. Workers of Aphaenogaster dulcineae. A,B: specimen from Tunisia (sequenced as A. foreli by Schifani et al. 2022). C,D: Syntype specimen from Spain (Aphaenogaster pallida dulcineae, CASENT0913113, photo from Zach Lieberman available on www.antweb.org)." figureDoi="http://doi.org/10.5281/zenodo.7891707" httpUri="https://zenodo.org/record/7891707/files/figure.png" pageId="3" pageNumber="62">Fig. 1</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>