294 lines
25 KiB
XML
294 lines
25 KiB
XML
<document id="BE6B256B815385A1623D2250A64DAB40" ID-CLB-Dataset="28501" ID-DOI="http://dx.doi.org/10.3897/zookeys.826.26488" ID-GBIF-Dataset="9d62af83-e4f5-423b-8e12-7dd567f69921" ID-PMC="PMC6405737" ID-Pensoft-Pub="1313-2970-826-1" ID-PubMed="30858752" ID-ZooBank="B1550A3AE54744509A44BC4366D5E110" ModsDocAuthor="" ModsDocDate="2019" ModsDocID="1313-2970-826-1" ModsDocOrigin="ZooKeys 826" ModsDocTitle="Scarabaeinae dung beetles from Ecuador: a catalog, nomenclatural acts, and distribution records" checkinTime="1551446249571" checkinUser="pensoft" docAuthor="Chamorro, William, Marin-Armijos, Diego, senjo, Angelico & Vaz-De-Mello, Fernando Z." docDate="2019" docId="75EB0ED927A5F755BE0E6DE7CABF5FE7" docLanguage="en" docName="ZooKeys 826: 1-343" docOrigin="ZooKeys 826" docSource="http://dx.doi.org/10.3897/zookeys.826.26488" docTitle="Coprophanaeus (Coprophanaeus) morenoi Arnaud 1982" docType="treatment" docVersion="7" lastPageNumber="71" masterDocId="FFECFFBAFFFC2D25D00B9843FFD67749" masterDocTitle="Scarabaeinae dung beetles from Ecuador: a catalog, nomenclatural acts, and distribution records" masterLastPageNumber="343" masterPageNumber="1" pageNumber="69" updateTime="1701379963039" updateUser="ExternalLinkService">
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<mods:title id="C7518C9FF9DB369BCE44C1693A7FCAC6">Scarabaeinae dung beetles from Ecuador: a catalog, nomenclatural acts, and distribution records</mods:title>
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<mods:namePart id="90C3E342D8A6ED30486746D317458AEA">Chamorro, William</mods:namePart>
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<mods:namePart id="D3391B54260AD9E5099DA54819DD3AE4">Marin-Armijos, Diego</mods:namePart>
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<mods:namePart id="EE3FE8B64D30C90468FC5A154D7263ED">senjo, Angelico</mods:namePart>
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<mods:namePart id="293E00710C790FC1ADA6B9B84EB75348">Vaz-De-Mello, Fernando Z.</mods:namePart>
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<mods:date id="56873E3964782BD1782F2693C612E1ED">2019</mods:date>
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<mods:number id="02CFC01879B7451CD1258E784DF30B81">826</mods:number>
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<mods:classification id="D680390D6AD8B5EC7533B233B83D43FA">journal article</mods:classification>
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<mods:identifier id="BDBE07E175DD62B5B629D21B47FAEBA6" type="DOI">http://dx.doi.org/10.3897/zookeys.826.26488</mods:identifier>
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<treatment id="75EB0ED927A5F755BE0E6DE7CABF5FE7" ID-GBIF-Taxon="154127695" LSID="urn:lsid:plazi:treatment:75EB0ED927A5F755BE0E6DE7CABF5FE7" httpUri="http://treatment.plazi.org/id/75EB0ED927A5F755BE0E6DE7CABF5FE7" lastPageId="70" lastPageNumber="71" pageId="68" pageNumber="69">
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<subSubSection id="0FCD72EE2D56689A249F36FCDAD01FA7" pageId="68" pageNumber="69" type="nomenclature">
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<paragraph id="F7E22616C8413676C9F1BABEBE73DCE7" pageId="68" pageNumber="69">
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<taxonomicName id="927467F79FA301CBC7C72DD6D72882DC" ID-CoL="7ZMPR" authority="Arnaud, 1982" authorityName="Arnaud" authorityYear="1982" class="Insecta" family="Scarabaeidae" genus="Coprophanaeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Coprophanaeus (Coprophanaeus) morenoi" order="Coleoptera" pageId="68" pageNumber="69" phylum="Arthropoda" rank="species" species="morenoi" subGenus="Coprophanaeus">Coprophanaeus (Coprophanaeus) morenoi Arnaud, 1982</taxonomicName>
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Plate 16C
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</paragraph>
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</subSubSection>
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<subSubSection id="9240AFCAFED9326B73D81E541DA21272" pageId="69" pageNumber="70" type="reference_group">
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<paragraph id="1E5C8AE5F2EEEB20EAB5D286F7E6878D" pageId="69" pageNumber="70">
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<taxonomicName id="279F56E218F7D6A9F36E35799BC1300A" class="Insecta" family="Scarabaeidae" genus="Coprophanaeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Coprophanaeus morenoi" order="Coleoptera" pageId="69" pageNumber="70" phylum="Arthropoda" rank="species" species="morenoi">
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<pageBreakToken id="2BB7E64FCCBD60A4158AD27B3330E21D" pageId="69" pageNumber="70" start="start">Coprophanaeus</pageBreakToken>
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morenoi
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</taxonomicName>
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Arnaud, 1982b: 121 (original description. Type locality: Equateur [= Ecuador], km 36 route Sto Domingo-Quevedo).
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</paragraph>
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<paragraph id="B9F209CD808831C8A1362E5EFB05217C" pageId="69" pageNumber="70">
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<taxonomicName id="80FD772535156E9CF58BF9590BF9860E" class="Insecta" family="Scarabaeidae" genus="Coprophanaeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Coprophanaeus morenoi" order="Coleoptera" pageId="69" pageNumber="70" phylum="Arthropoda" rank="species" species="morenoi">Coprophanaeus morenoi</taxonomicName>
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:
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<bibRefCitation id="1F157CD15A95445FD18E0FC2B4C1BBBB" author="Vitolo, AL" journalOrPublisher="Revista de la Academia Colombiana de Ciencias Exactas Fisicas y Naturales" pageId="278" pageNumber="279" pagination="591 - 601" title="Clave para la identificacion de los generos y especies Phanaeinas de Colombia." volume="24" year="2000">Vitolo 2000</bibRefCitation>
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: 599 (characters in key);
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<bibRefCitation id="D74AF0C5699274012CE7469F591E0710" author="Medina, CA" journalOrPublisher="Biota Colombiana" pageId="274" pageNumber="275" pagination="131 - 144" title="Escarabajos coprofagos (Coleoptera: Scarabaeinae) de Colombia." volume="2" year="2001">Medina et al. 2001</bibRefCitation>
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: 140 (cited for Colombia);
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<bibRefCitation id="0AC019B5CEB324B00188A6EF51461D42" author="Arnaud, P" journalOrPublisher="Hillside Books, Canterbury, England" pageId="261" pageNumber="262" title="Les Coleopteres du Monde, Vol. 28. Phanaeini. Dendropaemon, Tetramereia, Homalotarsus, Megatharsis, Diabroctis, Coprophanaeus, Oxysternon, Phanaeus, Sulcophanaeus." year="2002 a">Arnaud 2002a</bibRefCitation>
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: 42 (diagnosis);
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<bibRefCitation id="EAB7ABC233637E51795F9A909EC038B9" author="Vitolo, A" editor="Fernandez, F" journalOrPublisher="Facultad de Ciencias, Universidad Nacional de Colombia, Bogota" pageId="278" pageNumber="279" pagination="277 - 318" title="Escarabajos estercoleros de la tribu Phanaeini de Colombia (Coleoptera: Scarabaeoidea: Scarabaeidae)." volume="3" volumeTitle="Insectos de Colombia." year="2004">Vitolo 2004</bibRefCitation>
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: 290 (redescription);
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<bibRefCitation id="7147A1BC65043777AB60932ED4666573" author="Donoso, DA" journalOrPublisher="Annales de la Societe entomologique de France" pageId="266" pageNumber="267" pagination="455 - 469" title="Diversity and distribuccion of type specimens deposited in the invertebrate section of the museum of zoology QCAZ, Quito, Ecuador." url="https://doi.org/10.1080/00379271.2009.10697628" volume="45" year="2009">Donoso et al. 2009</bibRefCitation>
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: Appendix II. 16 (catalog of type MQCAZ);
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<bibRefCitation id="17F4051F2C2CDFBFE028785F1243791E" author="Edmonds, WD" journalOrPublisher="Insecta Mundi" pageId="266" pageNumber="267" pagination="1 - 111" title="A taxonomic review of the neotropical genus Coprophanaeus Olsoufieff, 1924 (Coleoptera: Scarabaeidae, Scarabaeinae)." volume="129" year="2010">
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Edmonds and
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<normalizedToken id="2098849282D5366B42BEF25E4C022FB3" originalValue="Zídek">Zidek</normalizedToken>
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2010
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</bibRefCitation>
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: 57 (characters in key), 60 (redescription);
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<bibRefCitation id="A19FEB859864C2E4807405A5F4858664" pageId="69" pageNumber="70">
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Kohlmann and
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<normalizedToken id="5A9ACAFD6C0BFAA5395780FDD6724CCD" originalValue="Solís">Solis</normalizedToken>
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2012
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</bibRefCitation>
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: 46 (cited for Ecuador);
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<bibRefCitation id="74BFEE860D8CF98DCA67FD706283006F" pageId="69" pageNumber="70">Krajcik 2012</bibRefCitation>
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: 204 (complete list of species, cited as subgenus of
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<taxonomicName id="42F7A3119FC47CC2AAAA560C6AD503A1" class="Insecta" family="Scarabaeidae" genus="Phanaeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Phanaeus" order="Coleoptera" pageId="69" pageNumber="70" phylum="Arthropoda" rank="genus">Phanaeus</taxonomicName>
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Macleay, 1819);
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</paragraph>
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<paragraph id="41CC8D67D2E11EC582C75197754A05BB" pageId="69" pageNumber="70">
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<taxonomicName id="F048242B58DECAD4581FC84710EB06B4" class="Insecta" family="Scarabaeidae" genus="Coprophanaeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Coprophanaeus (Coprophanaeus) morenoi" order="Coleoptera" pageId="69" pageNumber="70" phylum="Arthropoda" rank="species" species="morenoi" subGenus="Coprophanaeus">Coprophanaeus (Coprophanaeus) morenoi</taxonomicName>
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:
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<bibRefCitation id="79E8B3906BC2AC3F5385C29E538BBD10" author="Carvajal, V" journalOrPublisher="Quito, Ecuado" pageId="264" pageNumber="265" title="Escarabajos del Ecuador. Principales generos. Instituto de Ciencias Biologicas. Escuela Politecnica Nacional. Serie Entomologia, Nro. 1." year="2011">Carvajal et al. 2011</bibRefCitation>
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: 320-321 (cited for Ecuador);
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<bibRefCitation id="7B0C60001B7F077216AD528076327057" author="Chamorro, W" journalOrPublisher="Revista Colombiana de Entomologi ́ a" pageId="264" pageNumber="265" pagination="72 - 100" title="Listado de especies y clave de ge ́ neros y subge ́ neros de escarabajos estercoleros (Coleoptera: Scarabaei- dae: Scarabaeinae) presentes y presuntos para Ecuador." url="https://doi.org/10.25100/socolen.v44i1.6545" volume="44" year="2018">Chamorro et al. 2018</bibRefCitation>
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: 83 (figure 6C), 93 (cited for Ecuador).
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</paragraph>
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</subSubSection>
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<subSubSection id="5E9E682B65F63E5FF0B81339BE822ABE" pageId="69" pageNumber="70" type="type specimens">
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<paragraph id="99A954D1D899F7624DD74E45796683A0" pageId="69" pageNumber="70">Type specimens.</paragraph>
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<paragraph id="755B7CB6AC9577FCFFA1764A755168CA" pageId="69" pageNumber="70">
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<taxonomicName id="EC88FB61C7B0BEBC1A25516E3143CBA2" class="Insecta" family="Scarabaeidae" genus="Coprophanaeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Coprophanaeus morenoi" order="Coleoptera" pageId="69" pageNumber="70" phylum="Arthropoda" rank="species" species="morenoi">Coprophanaeus morenoi</taxonomicName>
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Arnaud, 1982. The holotype (♂) is deposited at the MNHN. Locality: Equateur [= Ecuador], Sto. Domingo km 36, examined. One Paratype is deposited in the MQCAZ, examined.
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</paragraph>
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<paragraph id="B0BBCE02A8EA318FFCC58CA921C81429" pageId="69" pageNumber="70">Holotype (♂): "EQUATEUR / STO DOMINGO K. 36 / JANV. 1982 / P&L ARNAUD coll [p and hw, black margin]", "Coprophanaeus / morenoi Mihi / HOLOTYPE ♂ / P. ARNAUD DET 1982 [p and hw, pink margin]", "HOLOTYPE [p, red label]".</paragraph>
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<paragraph id="9A2F7D3509191E82671734FACDE3776A" pageId="69" pageNumber="70">Paratype (♀): "ECUADOR (PICH) / TINA LANDIA / I. 82 650 m / P & L ARNAUD leg [p]", "Coprophanaeus / morenoi Mihi / P. ARNAUD DET / PARATYPE ♀ [p and hw, pink margin]".</paragraph>
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</subSubSection>
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<subSubSection id="89BF896F734428E8F60B960C5D4E1BD9" pageId="69" pageNumber="70" type="distribution">
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<paragraph id="E0356096A17F53762ADDE885709915FD" pageId="69" pageNumber="70">Distribution.</paragraph>
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<paragraph id="C7C4997B6CC84F17D3CC6BE6645ABFB4" pageId="69" pageNumber="70">Colombia, Costa Rica, Ecuador, Nicaragua, and Panama.</paragraph>
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</subSubSection>
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<subSubSection id="F02A7A40F6817E78FBC274951CD4E2F7" lastPageId="70" lastPageNumber="71" pageId="69" pageNumber="70" type="records examined">
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<paragraph id="D9292735718A354B5ED43CC84FB562D7" pageId="69" pageNumber="70">Records examined.</paragraph>
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<paragraph id="E84E7F2197E38837409B1D78F8ED57AB" lastPageId="70" lastPageNumber="71" pageId="69" pageNumber="70">
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CARCHI: Tobar Donoso, 300 m (8 specimens MECN). COTOPAXI: Guasaganda La Mana, 500 m (1 specimen CEMT; 3 specimens MQCAZ). ESMERALDAS:
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<normalizedToken id="F805F0AD3AC1E6A5FA9BF9B8D2DE639E" originalValue="Colón">Colon</normalizedToken>
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del Onzole (17 specimens MECN); Charco Vicente (13 specimens MGO-UC; 28 specimens MECN); Gualpi (15 specimens MECN); Gualpi Pajonal (17 specimens MECN); Palma Real (1 specimen MGO-UC; 15 specimens
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<pageBreakToken id="C713EA0D19ED4285C15B1438E78DD36F" pageId="70" pageNumber="71" start="start">MECN</pageBreakToken>
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); Playa de Oro (1 specimen MUTPL; 10 speciemens MECN); Playa de Oro, Padre Santo (19 specimens MGO-UC; 31 specimens MECN); Playa de Oro, Estero Pote, 200 m (3 specimen CEMT; 16 specimens MECN); road Ibarra-San Lorenzo, El Placer, 670 m (1 specimen CEMT). PICHINCHA: Guayabilla
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<normalizedToken id="AFCC509FB8D7CBC7C9CE2B56B8BA6EBB" originalValue="Río">Rio</normalizedToken>
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Guayllabamba, Manduriacus, 520 m (2 specimens MUTPL); Llurimaguas
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<normalizedToken id="3F102CD021F4553E1828959EB1B911C0" originalValue="Río">Rio</normalizedToken>
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Guayllabamba, Pedro Vicente Maldonado, 290 m (1 specimen MUTPL); Mangaloma, San Miguel de los Bancos, 820 m (1 specimen MUTPL); Tortugo
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<normalizedToken id="073CE93B6156CB061EFF5CB776143631" originalValue="Río">Rio</normalizedToken>
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Guayllabamba, Pedro Vicente Maldonado, 450 m (1 specimen MUTPL). SANTO DOMINGO DE LOS
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<normalizedToken id="E8A18EAE6DB5D4CBE9475BAC07BB1281" originalValue="TSÁCHILAS">TSACHILAS</normalizedToken>
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: Tinalandia, 650 m (1 specimen MQCAZ).
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</paragraph>
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</subSubSection>
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<subSubSection id="C64F9F789435D6790EFDBB098FC5C35E" pageId="70" pageNumber="71" type="literature records">
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<paragraph id="44012902D8DD5D630EE6171C8EF0A3BC" pageId="70" pageNumber="71">Literature records.</paragraph>
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<paragraph id="5065268A998FFC655D94E3918EB6FAD1" pageId="70" pageNumber="71">
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ESMERALDAS: 11 km SE San Lorenzo, La Chiquita Sta, 5 m (
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<bibRefCitation id="0F6898C6BF0B42E7C11E3D207445D01F" author="Edmonds, WD" journalOrPublisher="Insecta Mundi" pageId="266" pageNumber="267" pagination="1 - 111" title="A taxonomic review of the neotropical genus Coprophanaeus Olsoufieff, 1924 (Coleoptera: Scarabaeidae, Scarabaeinae)." volume="129" year="2010">
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Edmonds and
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<normalizedToken id="60E11D70B8FECDB5AD8546B24C3C6D18" originalValue="Zídek">Zidek</normalizedToken>
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2010
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: 62;
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<bibRefCitation id="93683745FB4757CA6B7CD3BC089494C8" pageId="70" pageNumber="71">
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Kohlmann and
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<normalizedToken id="479D185D4FACE3E7BC89205E34C4ED8E" originalValue="Solís">Solis</normalizedToken>
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2012
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</bibRefCitation>
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: 46); Yalare (
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<bibRefCitation id="2E4E96EE63220A41A9AA072686C73224" author="Edmonds, WD" journalOrPublisher="Insecta Mundi" pageId="266" pageNumber="267" pagination="1 - 111" title="A taxonomic review of the neotropical genus Coprophanaeus Olsoufieff, 1924 (Coleoptera: Scarabaeidae, Scarabaeinae)." volume="129" year="2010">
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Edmonds and
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<normalizedToken id="66864D78B4AF58B6E206A6CA212E7119" originalValue="Zídek">Zidek</normalizedToken>
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2010
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</bibRefCitation>
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: 62;
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<bibRefCitation id="75F225ED1CAF1B6DDC9E937ABDA03396" pageId="70" pageNumber="71">
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Kohlmann and
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<normalizedToken id="69BA20E8C6E3768E53FD9DB76B2B972F" originalValue="Solís">Solis</normalizedToken>
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2012
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</bibRefCitation>
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: 46); Punta Venado (
|
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<bibRefCitation id="78E242186F09559ED90BB2ABB1014089" author="Edmonds, WD" journalOrPublisher="Insecta Mundi" pageId="266" pageNumber="267" pagination="1 - 111" title="A taxonomic review of the neotropical genus Coprophanaeus Olsoufieff, 1924 (Coleoptera: Scarabaeidae, Scarabaeinae)." volume="129" year="2010">
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Edmonds and
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<normalizedToken id="E574DF753D2611918DF51A9B467BFB5E" originalValue="Zídek">Zidek</normalizedToken>
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2010
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</bibRefCitation>
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: 62;
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<bibRefCitation id="E06FEEC5C0CD46530B08D7F0E88A5BA9" pageId="70" pageNumber="71">
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Kohlmann and
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<normalizedToken id="6E19DD7385F9A7E5B51037A86338D154" originalValue="Solís">Solis</normalizedToken>
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2012
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</bibRefCitation>
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: 46); Playa de Oro (
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<bibRefCitation id="FD18EF165214AFEBFA746A31D8449C4B" author="Edmonds, WD" journalOrPublisher="Insecta Mundi" pageId="266" pageNumber="267" pagination="1 - 111" title="A taxonomic review of the neotropical genus Coprophanaeus Olsoufieff, 1924 (Coleoptera: Scarabaeidae, Scarabaeinae)." volume="129" year="2010">
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Edmonds and
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<normalizedToken id="FF8D671C17B876BD61EDD2C5FF3ED81F" originalValue="Zídek">Zidek</normalizedToken>
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2010
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</bibRefCitation>
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: 62;
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<bibRefCitation id="FA4049B8BF1408070D48225C1064570F" pageId="70" pageNumber="71">
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Kohlmann and
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<normalizedToken id="30E5C76B7E65190AED623D7B539CD99D" originalValue="Solís">Solis</normalizedToken>
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2012
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</bibRefCitation>
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: 46); La Concordia (
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<bibRefCitation id="B6427B18984597A4A4A4F2AF2122EAAF" author="Edmonds, WD" journalOrPublisher="Insecta Mundi" pageId="266" pageNumber="267" pagination="1 - 111" title="A taxonomic review of the neotropical genus Coprophanaeus Olsoufieff, 1924 (Coleoptera: Scarabaeidae, Scarabaeinae)." volume="129" year="2010">
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Edmonds and
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<normalizedToken id="F167EA5BB40A84EE9D0A02D78B1C2B25" originalValue="Zídek">Zidek</normalizedToken>
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2010
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</bibRefCitation>
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: 62); Palma Real (
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<bibRefCitation id="FBD0DB069D2311E7F1C59C025F9FA606" author="Edmonds, WD" journalOrPublisher="Insecta Mundi" pageId="266" pageNumber="267" pagination="1 - 111" title="A taxonomic review of the neotropical genus Coprophanaeus Olsoufieff, 1924 (Coleoptera: Scarabaeidae, Scarabaeinae)." volume="129" year="2010">
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Edmonds and
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<normalizedToken id="CBE55FAB90A97CD74CA2283EAEEAE461" originalValue="Zídek">Zidek</normalizedToken>
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2010
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</bibRefCitation>
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: 62). GUAYAS: Los Ceibos (
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<bibRefCitation id="19AAFAD66682B69D1FC155F3F8EA7C4B" pageId="70" pageNumber="71">
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Kohlmann and
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<normalizedToken id="ED64C3055EC9C95A71438E363639C15B" originalValue="Solís">Solis</normalizedToken>
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2012
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</bibRefCitation>
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: 46);
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<normalizedToken id="6C1DCEA9E18ED0B3936656282615E4BF" originalValue="Borbón">Borbon</normalizedToken>
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, 25 m (
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<bibRefCitation id="15AD00EC8D77556EAF1B2F1600077C81" author="Edmonds, WD" journalOrPublisher="Insecta Mundi" pageId="266" pageNumber="267" pagination="1 - 111" title="A taxonomic review of the neotropical genus Coprophanaeus Olsoufieff, 1924 (Coleoptera: Scarabaeidae, Scarabaeinae)." volume="129" year="2010">
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Edmonds and
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<normalizedToken id="8BCD9D7E596961079CC6720AEE34EF53" originalValue="Zídek">Zidek</normalizedToken>
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2010
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</bibRefCitation>
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: 62); San Miguel (
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<bibRefCitation id="F82C46BD9BD1C6CA20310A1164204113" author="Edmonds, WD" journalOrPublisher="Insecta Mundi" pageId="266" pageNumber="267" pagination="1 - 111" title="A taxonomic review of the neotropical genus Coprophanaeus Olsoufieff, 1924 (Coleoptera: Scarabaeidae, Scarabaeinae)." volume="129" year="2010">
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Edmonds and
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<normalizedToken id="FC62E1A1545FC254970AC30A6FE821A1" originalValue="Zídek">Zidek</normalizedToken>
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2010
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</bibRefCitation>
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: 62). PICHINCHA: [= LOS
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<normalizedToken id="4FEE146256DB6514B8506C24DC217CD4" originalValue="RÍOS">RIOS</normalizedToken>
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]: Station
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<normalizedToken id="24FBE4B9AA22C293DA833DB4557F17CA" originalValue="Río">Rio</normalizedToken>
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Palenque (
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<bibRefCitation id="ECA6BE4C532BAF7E49725CA8FBB8CCB4" author="Arnaud, P" journalOrPublisher="Miscellanea entomologica" pageId="260" pageNumber="261" pagination="121 - 123" title="Description de deux nouvelles speces de Phanaeini (Col: Scarabaeide)." volume="49" year="1982 b">Arnaud 1982b</bibRefCitation>
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: 122); 47 km S Sto. Domingo,
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<normalizedToken id="C22B692FDD494400BA875EF083821D5F" originalValue="Río">Rio</normalizedToken>
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Palenque (
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<bibRefCitation id="34CFE4121791A40CF67515CB11FB8960" pageId="70" pageNumber="71">
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Kohlmann and
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<normalizedToken id="2033DA34C5D3452B9C3E9A6283C228E0" originalValue="Solís">Solis</normalizedToken>
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2012
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</bibRefCitation>
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: 46); 113 km NW Quito on Puerto Quito road, 800 m (
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<bibRefCitation id="2AE4425D8AB200A8ACDA4F89231D5E9F" author="Edmonds, WD" journalOrPublisher="Insecta Mundi" pageId="266" pageNumber="267" pagination="1 - 111" title="A taxonomic review of the neotropical genus Coprophanaeus Olsoufieff, 1924 (Coleoptera: Scarabaeidae, Scarabaeinae)." volume="129" year="2010">
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Edmonds and
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<normalizedToken id="9F20D9F30CCF87F93056F253E643F5BE" originalValue="Zídek">Zidek</normalizedToken>
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2010
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</bibRefCitation>
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: 62). LOS
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<normalizedToken id="E3FA8E94959CAD8CD8F96224A1DE9772" originalValue="RÍOS">RIOS</normalizedToken>
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: Quevedo (
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<bibRefCitation id="D061CB23A4B40AF8B404B1FEB4D613F8" author="Edmonds, WD" journalOrPublisher="Insecta Mundi" pageId="266" pageNumber="267" pagination="1 - 111" title="A taxonomic review of the neotropical genus Coprophanaeus Olsoufieff, 1924 (Coleoptera: Scarabaeidae, Scarabaeinae)." volume="129" year="2010">
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Edmonds and
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<normalizedToken id="9BB613F24B358A22AA44F1A22DB4AE8D" originalValue="Zídek">Zidek</normalizedToken>
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2010
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</bibRefCitation>
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: 62); Quevedo Pichilinge (
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<bibRefCitation id="A6018D4DD78D3279BCA5AC3777E77496" pageId="70" pageNumber="71">
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Kohlmann and
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<normalizedToken id="207C09C8211368A1C39C7F492CC15008" originalValue="Solís">Solis</normalizedToken>
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2012
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</bibRefCitation>
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: 46);
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<normalizedToken id="3E41435054A992DFDED0E681587E93C0" originalValue="Río">Rio</normalizedToken>
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Palenque Research Station, 200 m (
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<bibRefCitation id="CA128A6E09FA84A4C520E8061FB6E920" author="Edmonds, WD" journalOrPublisher="Insecta Mundi" pageId="266" pageNumber="267" pagination="1 - 111" title="A taxonomic review of the neotropical genus Coprophanaeus Olsoufieff, 1924 (Coleoptera: Scarabaeidae, Scarabaeinae)." volume="129" year="2010">
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Edmonds and
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<normalizedToken id="93B8DD563C6EB399321466C89793FF28" originalValue="Zídek">Zidek</normalizedToken>
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2010
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</bibRefCitation>
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: 62). SANTO DOMINGO DE LOS
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<normalizedToken id="4C963FBE317CE71E01CC70D56DA44131" originalValue="TSÁCHILAS">TSACHILAS</normalizedToken>
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: km 36 route Sto. Domingo-Quevedo (
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<bibRefCitation id="9A25B1F37707A9C017BF9DCB9D067CCD" author="Arnaud, P" journalOrPublisher="Miscellanea entomologica" pageId="260" pageNumber="261" pagination="121 - 123" title="Description de deux nouvelles speces de Phanaeini (Col: Scarabaeide)." volume="49" year="1982 b">Arnaud 1982b</bibRefCitation>
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: 122); Tina Landia [= Tinalandia] (Env. Sto. Domingo) (
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<bibRefCitation id="550946CABF2AFF6D7FB8AD30A8B9829E" author="Arnaud, P" journalOrPublisher="Miscellanea entomologica" pageId="260" pageNumber="261" pagination="121 - 123" title="Description de deux nouvelles speces de Phanaeini (Col: Scarabaeide)." volume="49" year="1982 b">Arnaud 1982b</bibRefCitation>
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: 122); i1 km E Tinalandia, 600 m (
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<bibRefCitation id="7A1F3473B7F3A5DE120D9FDDCBA94FDF" author="Edmonds, WD" journalOrPublisher="Insecta Mundi" pageId="266" pageNumber="267" pagination="1 - 111" title="A taxonomic review of the neotropical genus Coprophanaeus Olsoufieff, 1924 (Coleoptera: Scarabaeidae, Scarabaeinae)." volume="129" year="2010">
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Edmonds and
|
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<normalizedToken id="3222CF8260C38FB37F2D3EAC312EF464" originalValue="Zídek">Zidek</normalizedToken>
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2010
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</bibRefCitation>
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: 62) 16 km E Sto. Domingo, Tinalandia (
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<bibRefCitation id="E3021892C74D94C8273BFB1C5098C0C7" pageId="70" pageNumber="71">
|
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Kohlmann and
|
|
<normalizedToken id="4F28562190D542A2D0B3DBA8B4795C2C" originalValue="Solís">Solis</normalizedToken>
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2012
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</bibRefCitation>
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: 46).
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</paragraph>
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</subSubSection>
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<subSubSection id="D47F05EAD75D0D0962C08571CD317C5C" pageId="70" pageNumber="71" type="temporal data">
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<paragraph id="83E845D12FE6FC02E574652C2F1C2C86" pageId="70" pageNumber="71">Temporal data.</paragraph>
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<paragraph id="CDB19D1AC24E259C07B43CE37A2D1F50" pageId="70" pageNumber="71">Collected every month of the year.</paragraph>
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</subSubSection>
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<subSubSection id="1A6DD96AB52F2A123866AAEDA49B066A" pageId="70" pageNumber="71" type="remarks">
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<paragraph id="AF14AC103C80BC5DDDC9D0CB3E57C9CF" pageId="70" pageNumber="71">Remarks.</paragraph>
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<paragraph id="E5A617400B4F9E625524C782D070A2AE" pageId="70" pageNumber="71">Inhabits coastal lowland evergreen forests and coastal evergreen foothill forests from 5-850 m a.s.l. Species was collected with flight interception traps and pitfall traps baited with carrion and human feces.</paragraph>
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</subSubSection>
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</treatment>
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</document> |