287 lines
40 KiB
XML
287 lines
40 KiB
XML
<document id="4BF5D078CF862698D89F98CED6A1BE68" ID-CLB-Dataset="26738" ID-DOI="10.11646/zootaxa.4608.3.8" ID-GBIF-Dataset="dc1355a6-6693-471c-b91b-ad64e7a9cbcf" ID-ISSN="1175-5326" ID-Zenodo-Dep="3066776" ID-ZooBank="BB206D49-1623-4413-8DB2-6FAED7761321" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1558417247298" checkinUser="plazi" docAuthor="Pilato, Giovanni, D’Urso, Vera, Sabella, Giorgio & Lisi, Oscar" docDate="2019" docId="191487ACE22BFFFCFF1D3EECFEE1FEDB" docLanguage="en" docName="zootaxa.4608.3.8.pdf" docOrigin="Zootaxa 4608 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Isohypsibius occultus Pilato & D’Urso & Sabella & Lisi 2019, sp. nov." docType="treatment" docVersion="8" lastPageNumber="547" masterDocId="E52DFFD4E229FFF8FF8A3E75FFBDFFC4" masterDocTitle="A new species of Isohypsibiid (Eutardigrada) from Sicily" masterLastPageNumber="554" masterPageNumber="543" pageNumber="545" updateTime="1698731362845" updateUser="ExternalLinkService">
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<mods:mods id="E49E9C9298939A23D5A66BF940D0042B" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="76E78FFF6968D575E08484EF99302198">
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<mods:title id="988F1276B68D4F01BA3913CACD0D573C">A new species of Isohypsibiid (Eutardigrada) from Sicily</mods:title>
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</mods:titleInfo>
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<mods:name id="5586810F410B1EFC95D97126E7731967" type="personal">
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<mods:role id="037D6AAD35413A18CB012A87B4049606">
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<mods:roleTerm id="F8229FDD4BA7C09E106C3FA87F67B8EE">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="36BE9E84091F2623B3F1FD13F4AA43F0">Pilato, Giovanni</mods:namePart>
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</mods:name>
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<mods:name id="5517CC0279E48A38E4752EDFE821283D" type="personal">
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<mods:role id="2AC8E2A4674176DE69A43C8C99B9AEF9">
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<mods:roleTerm id="5972F19915985D9C14611D4099BA270A">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="5F7350B80D5D9874CA62F9DF0468BE6B">D’Urso, Vera</mods:namePart>
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</mods:name>
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<mods:name id="B20EF3C24F2068254E4FC0D53A12E5AD" type="personal">
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<mods:roleTerm id="5221798507431F49CA45035DCB644D5D">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="5A200AEA924D2C32351624171C75F2D7">Sabella, Giorgio</mods:namePart>
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</mods:name>
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<mods:name id="FC43FE1C22AFC92D5E1EF41AE9AF10AF" type="personal">
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<mods:role id="0E5A4B2331CE3CECFA0F49D509ACB607">
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<mods:roleTerm id="52E2C59EF40C9A479C1F003C73F01D5C">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="F1D2BF1FE2042919E2964C3EF5EBB3D8">Lisi, Oscar</mods:namePart>
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</mods:name>
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<mods:typeOfResource id="352C9714899AB0F8114A62F73E394846">text</mods:typeOfResource>
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<mods:title id="E816992507A7FFD0EFF8DB3D4E69A723">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="F6030CCB973C3A24F6AE0CF13EA8784E">
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<mods:date id="B86FFB1052DE387759D8554754418C38">2019</mods:date>
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<mods:detail id="595977DDC6B6EBD8B7496038A8749B13" type="pubDate">
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<mods:number id="24EBA13A9A5F5262AD65BD1F28AE230D">2019-05-21</mods:number>
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</mods:detail>
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<mods:detail id="F2F2E4614D7F7AF17A6317C9AB811751" type="volume">
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<mods:number id="C48A3044A22FD8F547A18F62D8818709">4608</mods:number>
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</mods:detail>
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<mods:detail id="157091BA1BB82658ED020758F72E2793" type="issue">
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<mods:number id="23A557A98D591DA8AFA6D4D562353D1E">3</mods:number>
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</mods:detail>
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<mods:classification id="1CD9C97084BA83C46EAE78F91DCC89A2">journal article</mods:classification>
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<mods:identifier id="96748CE908AB09D561284B4E7A705BBC" type="CLB-Dataset">26738</mods:identifier>
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<mods:identifier id="D2D44BCC0C7CB5F127B743AF8A8BCCB3" type="DOI">10.11646/zootaxa.4608.3.8</mods:identifier>
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<treatment id="191487ACE22BFFFCFF1D3EECFEE1FEDB" ID-GBIF-Taxon="159420409" LSID="urn:lsid:plazi:treatment:191487ACE22BFFFCFF1D3EECFEE1FEDB" httpUri="http://treatment.plazi.org/id/191487ACE22BFFFCFF1D3EECFEE1FEDB" lastPageId="4" lastPageNumber="547" pageId="2" pageNumber="545">
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<subSubSection id="D9A76531E22BFFFAFF1D3EECFDBDFF70" box="[151,512,153,180]" pageId="2" pageNumber="545" type="nomenclature">
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<paragraph id="910236BAE22BFFFAFF1D3EECFDBDFF70" blockId="2.[151,1437,153,756]" box="[151,512,153,180]" pageId="2" pageNumber="545">
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<heading id="CA4A81D6E22BFFFAFF1D3EECFDBDFF70" bold="true" box="[151,512,153,180]" fontSize="11" level="1" pageId="2" pageNumber="545" reason="1">
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<emphasis id="A3C9EAA8E22BFFFAFF1D3EECFDBDFF70" bold="true" box="[151,512,153,180]" pageId="2" pageNumber="545">
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<taxonomicName id="56BD4D39E22BFFFAFF1D3EECFE26FF70" authority="Pilato & D’Urso & Sabella & Lisi, 2019" authorityName="Pilato & D’Urso & Sabella & Lisi" authorityYear="2019" box="[151,411,153,180]" class="Eutardigrada" family="Hypsibiidae" genus="Isohypsibius" kingdom="Animalia" order="Parachela" pageId="2" pageNumber="545" phylum="Tardigrada" rank="species" species="occultus" status="sp. nov.">
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<emphasis id="A3C9EAA8E22BFFFAFF1D3EECFE26FF70" bold="true" box="[151,411,153,180]" italics="true" pageId="2" pageNumber="545">Isohypsibius occultus</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="B8FA57D3E22BFFFAFE293EEFFDBDFF70" box="[419,512,154,180]" pageId="2" pageNumber="545" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="D9A76531E22BFFFAFF1D3EC8FEDDFF13" box="[151,352,189,215]" pageId="2" pageNumber="545" type="description">
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||
<paragraph id="910236BAE22BFFFAFF1D3EC8FEDDFF13" blockId="2.[151,1437,153,756]" box="[151,352,189,215]" pageId="2" pageNumber="545">
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<figureCitation id="09862A3FE22BFFFAFF1D3EC8FF5EFF13" box="[151,227,189,215]" captionStart="FIGURE 1" captionStartId="2.[151,250,1828,1853]" captionTargetBox="[242,1358,762,1816]" captionTargetId="figure@2.[259,1329,770,1817]" captionText="FIGURE 1. Isohypsibius occultus sp. nov. A, B, The arrows indicate details of the cuticular ornamentation (A: paratype, slide No. 5548. B: paratype, slide No. 5608). C, Bucco-pharyngeal apparatus in lateral view (paratype, slide No. 1203). The ridge- shaped apophyses for the insertion of the stylet muscles (arrow a), the short and stout stylet supports (arrow b), and the very developed stylet furcae (arrows c) are evident. D, Bucco-pharyngeal apparatus in dorsal view (holotype, slide No. 5608). The well-developed medio-dorsal transversal ridge of the buccal armature (arrow a), the short and stout stylet supports (arrow b) and the well-developed stylet furcae (arrow c) are evident. Scale bars = 10 µm." httpUri="https://zenodo.org/record/3403907/files/figure.png" pageId="2" pageNumber="545">Figs. 1</figureCitation>
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, 2.
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<tableCitation id="DC3F0301E22BFFFAFE873EC8FEDDFF13" box="[269,352,189,215]" captionStart="TABLE 1" captionStartId="4.[151,239,328,352]" captionTargetBox="[161,1407,437,1686]" captionTargetPageId="4" captionText="TABLE 1. Measurements in µm and pt index values of relevant taxonomic characters of five specimens (holotype and four paratypes) of Isohypsibius occultus sp. nov. In the last column the difference between the maximum and the mini- mum value is indicated." httpUri="http://table.plazi.org/id/C5C26632E22DFFFCFF1D3F3DFE2AFE6C" pageId="2" pageNumber="545" tableUuid="C5C26632E22DFFFCFF1D3F3DFE2AFE6C">Table 1</tableCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="D9A76531E22BFFFAFF1D3F73FE05FE4F" pageId="2" pageNumber="545" type="materials_examined">
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<paragraph id="910236BAE22BFFFAFF1D3F73FB67FEDB" blockId="2.[151,1437,153,756]" box="[151,1242,261,288]" pageId="2" pageNumber="545">
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<materialsCitation id="21D53CE7E22BFFFAFF1D3F73FB6BFEDB" ID-GBIF-Occurrence="2397873193" box="[151,1238,261,288]" location="Sicily" municipality="Mirabella Imbaccari" pageId="2" pageNumber="545" specimenCount="1" typeStatus="holotype">
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<emphasis id="A3C9EAA8E22BFFFAFF1D3F73FE8DFEE4" bold="true" box="[151,304,261,288]" pageId="2" pageNumber="545">
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<typeStatus id="4E068818E22BFFFAFF1D3F73FF6FFEE4" box="[151,210,262,288]" pageId="2" pageNumber="545">Type</typeStatus>
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locality
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</emphasis>
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:
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<location id="94626061E22BFFFAFEB53F70FE38FEDB" LSID="urn:lsid:plazi:treatment:191487ACE22BFFFCFF1D3EECFEE1FEDB:94626061E22BFFFAFEB53F70FE38FEDB" box="[319,389,261,287]" municipality="Mirabella Imbaccari" name="Sicily" pageId="2" pageNumber="545">Sicily</location>
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:
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<collectingMunicipality id="7166ACC0E22BFFFAFE053F70FDCAFEDB" box="[399,631,261,287]" pageId="2" pageNumber="545">Mirabella Imbaccari</collectingMunicipality>
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: wet mosses at the side of the river “Fiume del Ferro”
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</materialsCitation>
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.
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</paragraph>
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<paragraph id="910236BAE22BFFFAFF4D3F5CFE05FE4F" blockId="2.[151,1437,153,756]" pageId="2" pageNumber="545">
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<emphasis id="A3C9EAA8E22BFFFAFF4D3F5CFE14FE80" bold="true" box="[199,425,297,324]" pageId="2" pageNumber="545">Material examined</emphasis>
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||
:
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<materialsCitation id="21D53CE7E22BFFFAFE3D3F5CFE05FE4F" ID-GBIF-Occurrence="2397873194" collectingDate="1980-10" location="Flascio River" pageId="2" pageNumber="545" specimenCount="21" typeStatus="holotype">
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<typeStatus id="4E068818E22BFFFAFE3D3F5CFDA2FE87" box="[439,543,297,323]" pageId="2" pageNumber="545" type="holotype">Holotype</typeStatus>
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and one
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<typeStatus id="4E068818E22BFFFAFD083F5FFD5FFE80" box="[642,738,298,324]" pageId="2" pageNumber="545" type="paratype">paratype</typeStatus>
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(slide N. 5608) collected in wet mosses in
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<date id="E503107AE22BFFFAFB353F5CFAE5FE87" box="[1215,1368,297,323]" pageId="2" pageNumber="545" value="1980-10">
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<collectingDate id="F547E992E22BFFFAFB353F5CFAE5FE87" box="[1215,1368,297,323]" pageId="2" pageNumber="545" value="1980-10">
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October
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||
<quantity id="56459B5FE22BFFFAFAAA3F5CFAE5FE87" box="[1312,1368,297,323]" metricMagnitude="1" metricUnit="m" metricValue="5.0291999999999994" pageId="2" pageNumber="545" unit="in" value="1980.0">1980</quantity>
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</collectingDate>
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</date>
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in the type locality;
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<specimenCount id="87BBFD33E22BFFFAFEBB3F38FE02FEAC" box="[305,447,333,360]" pageId="2" pageNumber="545" type="generic" typeStatus="paratypes">21 paratypes</specimenCount>
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(slides Nos.
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<date id="E503107AE22BFFFAFDD83F38FD6EFEA3" box="[594,723,333,359]" pageId="2" pageNumber="545" value="1201" valueMax="1204">1201–1204</date>
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||
; 2774; 5548–5550) extracted from sediment of
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<location id="94626061E22BFFFAFB633F38FAC2FEA3" LSID="urn:lsid:plazi:treatment:191487ACE22BFFFCFF1D3EECFEE1FEDB:94626061E22BFFFAFB633F38FAC2FEA3" box="[1257,1407,333,359]" name="Flascio River" pageId="2" pageNumber="545">Flascio River</location>
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in the territory of Randazzo.
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</materialsCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="D9A76531E22BFFFAFF4D3FE0FB3DFDA7" pageId="2" pageNumber="545" type="diagnosis">
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<paragraph id="910236BAE22BFFFAFF4D3FE0FB3DFDA7" blockId="2.[151,1437,153,756]" pageId="2" pageNumber="545">
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<emphasis id="A3C9EAA8E22BFFFAFF4D3FE0FE26FE74" bold="true" box="[199,411,405,432]" pageId="2" pageNumber="545">Specific diagnosis</emphasis>
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: cuticle smooth or with ornamentation fading to invisibility, often difficult to see, similar to that of
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<emphasis id="A3C9EAA8E22BFFFAFE803FCCFDB6FE17" box="[266,523,441,467]" italics="true" pageId="2" pageNumber="545">Isohypsibius monoicus</emphasis>
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. Eyes present. Buccal apparatus of the
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<emphasis id="A3C9EAA8E22BFFFAFC5E3FCCFBDDFE17" box="[980,1120,441,467]" italics="true" pageId="2" pageNumber="545">Isohypsibius</emphasis>
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type; buccal armature with well-developed medial transverse ridges. Claws, of the
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<emphasis id="A3C9EAA8E22BFFFAFC823FA8FC29FE33" box="[776,916,477,503]" italics="true" pageId="2" pageNumber="545">Isohypsibius</emphasis>
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type, well developed; internal claws I–III and anterior claws IV with the main branch thinning gradually; common basal portion thin; flexible lunules present, often rolled down under the claw base; no cuticular bar is present on the legs but, in some cases, a cuticular bulge is present simulating the presence of a bar-like thickening. Smooth eggs laid in the exuviae.
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</paragraph>
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</subSubSection>
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<subSubSection id="D9A76531E22BFFFBFF4D3C18FD3DFDA7" lastPageId="3" lastPageNumber="546" pageId="2" pageNumber="545" type="description">
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||
<paragraph id="910236BAE22BFFFBFF4D3C18FD3DFDA7" blockId="2.[151,1437,153,756]" lastBlockId="3.[151,1437,153,755]" lastPageId="3" lastPageNumber="546" pageId="2" pageNumber="545">
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<emphasis id="A3C9EAA8E22BFFFAFF4D3C18FDACFD4C" bold="true" box="[199,529,621,648]" pageId="2" pageNumber="545">Description of the holotype:</emphasis>
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Body length 411 µm, yellowish. Eyes present. Cuticle with ornamentation fading to invisibility, very difficult to see, more easily recognizable in the caudal portion of the body; tubercles give to the cuticle, as in
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<emphasis id="A3C9EAA8E22BFFFAFEAD3CC0FD99FD0B" box="[295,548,693,719]" italics="true" pageId="2" pageNumber="545">Isohypsibius monoicus</emphasis>
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, a “Persian lamb”-like pattern (Figs. 1 A, B, arrows). In a paratype 383 µm long, the largest caudal tubercles have a diameter up to 2.7 µm, but most are smaller The buccal apparatus is of the
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<taxonomicName id="56BD4D39E22AFFFBFDA93EECFD12FF77" authorityName="Thulin" authorityYear="1928" box="[547,687,153,179]" class="Eutardigrada" family="Hypsibiidae" genus="Isohypsibius" kingdom="Animalia" order="Parachela" pageId="3" pageNumber="546" phylum="Tardigrada" rank="genus">
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<emphasis id="A3C9EAA8E22AFFFBFDA93EECFD12FF77" box="[547,687,153,179]" italics="true" pageId="3" pageNumber="546">Isohypsibius</emphasis>
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</taxonomicName>
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<typeStatus id="4E068818E22AFFFBFD3D3EECFD57FF77" box="[695,746,153,179]" pageId="3" pageNumber="546">type</typeStatus>
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, i.e. the mouth has no peribuccal lamellae, the buccal tube is rigid without ventral lamina, and with ridge-shaped apophyses for the insertion of the stylet muscles, symmetrical with respect to the frontal plane (
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<figureCitation id="09862A3FE22AFFFBFD823E94FDDBFF3F" box="[520,614,225,251]" captionStart="FIGURE 1" captionStartId="2.[151,250,1828,1853]" captionTargetBox="[242,1358,762,1816]" captionTargetId="figure@2.[259,1329,770,1817]" captionText="FIGURE 1. Isohypsibius occultus sp. nov. A, B, The arrows indicate details of the cuticular ornamentation (A: paratype, slide No. 5548. B: paratype, slide No. 5608). C, Bucco-pharyngeal apparatus in lateral view (paratype, slide No. 1203). The ridge- shaped apophyses for the insertion of the stylet muscles (arrow a), the short and stout stylet supports (arrow b), and the very developed stylet furcae (arrows c) are evident. D, Bucco-pharyngeal apparatus in dorsal view (holotype, slide No. 5608). The well-developed medio-dorsal transversal ridge of the buccal armature (arrow a), the short and stout stylet supports (arrow b) and the well-developed stylet furcae (arrow c) are evident. Scale bars = 10 µm." httpUri="https://zenodo.org/record/3403907/files/figure.png" pageId="3" pageNumber="546">Fig. 1 C</figureCitation>
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, arrows a). Buccal armature (
|
||
<bibRefCitation id="F52C4B4BE22AFFFBFC243E94FB8CFF3F" author="Pilato, G." box="[942,1073,225,251]" pageId="3" pageNumber="546" pagination="43 - 58" refId="ref5652" refString="Pilato, G. (1974) Struttura dell'armatura boccale di alcune specie di Isohypsibius (Eutardigrada). Animalia, 1, 43 - 58." type="journal article" year="1974">Pilato 1974</bibRefCitation>
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||
) without bands of teeth but with a long dorsal and a long ventral transverse ridge in the caudal portion of the buccal cavity; the median portion of those ridges is well developed (
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<figureCitation id="09862A3FE22AFFFBFE713F5CFDE9FE87" box="[507,596,297,323]" captionStart="FIGURE 1" captionStartId="2.[151,250,1828,1853]" captionTargetBox="[242,1358,762,1816]" captionTargetId="figure@2.[259,1329,770,1817]" captionText="FIGURE 1. Isohypsibius occultus sp. nov. A, B, The arrows indicate details of the cuticular ornamentation (A: paratype, slide No. 5548. B: paratype, slide No. 5608). C, Bucco-pharyngeal apparatus in lateral view (paratype, slide No. 1203). The ridge- shaped apophyses for the insertion of the stylet muscles (arrow a), the short and stout stylet supports (arrow b), and the very developed stylet furcae (arrows c) are evident. D, Bucco-pharyngeal apparatus in dorsal view (holotype, slide No. 5608). The well-developed medio-dorsal transversal ridge of the buccal armature (arrow a), the short and stout stylet supports (arrow b) and the well-developed stylet furcae (arrow c) are evident. Scale bars = 10 µm." httpUri="https://zenodo.org/record/3403907/files/figure.png" pageId="3" pageNumber="546">Fig. 1D</figureCitation>
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||
, arrow a), the lateral portions are subdivided into teeth. The buccal tube is
|
||
<quantity id="56459B5FE22AFFFBFF1D3F38FF4BFEA3" box="[151,246,333,359]" metricMagnitude="-2" metricUnit="m" metricValue="4.17" pageId="3" pageNumber="546" unit="mm" value="41.7">41.7 µm</quantity>
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||
long and
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||
<quantity id="56459B5FE22AFFFBFEE23F38FE04FEA3" box="[360,441,333,359]" metricMagnitude="-3" metricUnit="m" metricValue="4.9" pageId="3" pageNumber="546" unit="mm" value="4.9">4.9 µm</quantity>
|
||
wide (
|
||
<emphasis id="A3C9EAA8E22AFFFBFD833F3BFDA2FEA3" box="[521,543,334,359]" italics="true" pageId="3" pageNumber="546">pt</emphasis>
|
||
= 11.7); the stylet supports are short and stout (
|
||
<figureCitation id="09862A3FE22AFFFBFBB43F38FB01FEA3" box="[1086,1212,333,359]" captionStart="FIGURE 1" captionStartId="2.[151,250,1828,1853]" captionTargetBox="[242,1358,762,1816]" captionTargetId="figure@2.[259,1329,770,1817]" captionText="FIGURE 1. Isohypsibius occultus sp. nov. A, B, The arrows indicate details of the cuticular ornamentation (A: paratype, slide No. 5548. B: paratype, slide No. 5608). C, Bucco-pharyngeal apparatus in lateral view (paratype, slide No. 1203). The ridge- shaped apophyses for the insertion of the stylet muscles (arrow a), the short and stout stylet supports (arrow b), and the very developed stylet furcae (arrows c) are evident. D, Bucco-pharyngeal apparatus in dorsal view (holotype, slide No. 5608). The well-developed medio-dorsal transversal ridge of the buccal armature (arrow a), the short and stout stylet supports (arrow b) and the well-developed stylet furcae (arrow c) are evident. Scale bars = 10 µm." httpUri="https://zenodo.org/record/3403907/files/figure.png" pageId="3" pageNumber="546">Figs. 1C,D</figureCitation>
|
||
, arrows b), inserted on the buccal tube at 70.6 percent of its length (
|
||
<emphasis id="A3C9EAA8E22AFFFBFD4D3F07FD60FE4F" box="[711,733,370,395]" italics="true" pageId="3" pageNumber="546">pt</emphasis>
|
||
= 70.6); each stylet has a well-developed furca, the terminal branches of which have very evident swollen apices (
|
||
<figureCitation id="09862A3FE22AFFFBFD733FE0FC3DFE6B" box="[761,896,405,431]" captionStart="FIGURE 1" captionStartId="2.[151,250,1828,1853]" captionTargetBox="[242,1358,762,1816]" captionTargetId="figure@2.[259,1329,770,1817]" captionText="FIGURE 1. Isohypsibius occultus sp. nov. A, B, The arrows indicate details of the cuticular ornamentation (A: paratype, slide No. 5548. B: paratype, slide No. 5608). C, Bucco-pharyngeal apparatus in lateral view (paratype, slide No. 1203). The ridge- shaped apophyses for the insertion of the stylet muscles (arrow a), the short and stout stylet supports (arrow b), and the very developed stylet furcae (arrows c) are evident. D, Bucco-pharyngeal apparatus in dorsal view (holotype, slide No. 5608). The well-developed medio-dorsal transversal ridge of the buccal armature (arrow a), the short and stout stylet supports (arrow b) and the well-developed stylet furcae (arrow c) are evident. Scale bars = 10 µm." httpUri="https://zenodo.org/record/3403907/files/figure.png" pageId="3" pageNumber="546">Figs. 1 C,D</figureCitation>
|
||
, arrows c). The pharyngeal bulb has apophyses and three short and stout macroplacoids without lateral incisions; microplacoid and septulum absent (
|
||
<figureCitation id="09862A3FE22AFFFBFA9A3FCCFA32FE17" box="[1296,1423,441,467]" captionStart="FIGURE 1" captionStartId="2.[151,250,1828,1853]" captionTargetBox="[242,1358,762,1816]" captionTargetId="figure@2.[259,1329,770,1817]" captionText="FIGURE 1. Isohypsibius occultus sp. nov. A, B, The arrows indicate details of the cuticular ornamentation (A: paratype, slide No. 5548. B: paratype, slide No. 5608). C, Bucco-pharyngeal apparatus in lateral view (paratype, slide No. 1203). The ridge- shaped apophyses for the insertion of the stylet muscles (arrow a), the short and stout stylet supports (arrow b), and the very developed stylet furcae (arrows c) are evident. D, Bucco-pharyngeal apparatus in dorsal view (holotype, slide No. 5608). The well-developed medio-dorsal transversal ridge of the buccal armature (arrow a), the short and stout stylet supports (arrow b) and the well-developed stylet furcae (arrow c) are evident. Scale bars = 10 µm." httpUri="https://zenodo.org/record/3403907/files/figure.png" pageId="3" pageNumber="546">Figs. 1C,D</figureCitation>
|
||
). The first macroplacoid, with a short triangular anterior portion, is 5.0 µm (
|
||
<emphasis id="A3C9EAA8E22AFFFBFC593FABFC54FE33" box="[979,1001,478,503]" italics="true" pageId="3" pageNumber="546">pt</emphasis>
|
||
= 12.0), the second
|
||
<quantity id="56459B5FE22AFFFBFB403FA8FAA7FE33" box="[1226,1306,477,503]" metricMagnitude="-3" metricUnit="m" metricValue="4.8" pageId="3" pageNumber="546" unit="mm" value="4.8">4.8 µm</quantity>
|
||
(
|
||
<emphasis id="A3C9EAA8E22AFFFBFAA33FABFA82FE33" box="[1321,1343,478,503]" italics="true" pageId="3" pageNumber="546">pt</emphasis>
|
||
= 11.6), the third
|
||
<quantity id="56459B5FE22AFFFBFF773C75FEF3FDDF" box="[253,334,512,539]" metricMagnitude="-3" metricUnit="m" metricValue="5.9" pageId="3" pageNumber="546" unit="mm" value="5.9">5.9 µm</quantity>
|
||
(
|
||
<emphasis id="A3C9EAA8E22AFFFBFED53C77FEC8FDDF" box="[351,373,514,539]" italics="true" pageId="3" pageNumber="546">pt</emphasis>
|
||
= 14.2), the entire placoid row is
|
||
<quantity id="56459B5FE22AFFFBFD7F3C74FCE9FDDF" box="[757,852,513,539]" metricMagnitude="-2" metricUnit="m" metricValue="1.68" pageId="3" pageNumber="546" unit="mm" value="16.8">16.8 µm</quantity>
|
||
(
|
||
<emphasis id="A3C9EAA8E22AFFFBFCEF3C77FCC6FDDF" box="[869,891,514,539]" italics="true" pageId="3" pageNumber="546">pt</emphasis>
|
||
= 40.5). By coverslip compression, the placoids may assume (
|
||
<figureCitation id="09862A3FE22AFFFBFEB93C50FE0DFDFB" box="[307,432,549,575]" captionStart="FIGURE 1" captionStartId="2.[151,250,1828,1853]" captionTargetBox="[242,1358,762,1816]" captionTargetId="figure@2.[259,1329,770,1817]" captionText="FIGURE 1. Isohypsibius occultus sp. nov. A, B, The arrows indicate details of the cuticular ornamentation (A: paratype, slide No. 5548. B: paratype, slide No. 5608). C, Bucco-pharyngeal apparatus in lateral view (paratype, slide No. 1203). The ridge- shaped apophyses for the insertion of the stylet muscles (arrow a), the short and stout stylet supports (arrow b), and the very developed stylet furcae (arrows c) are evident. D, Bucco-pharyngeal apparatus in dorsal view (holotype, slide No. 5608). The well-developed medio-dorsal transversal ridge of the buccal armature (arrow a), the short and stout stylet supports (arrow b) and the well-developed stylet furcae (arrow c) are evident. Scale bars = 10 µm." httpUri="https://zenodo.org/record/3403907/files/figure.png" pageId="3" pageNumber="546">Figs. 1C,D</figureCitation>
|
||
) the appearance described by
|
||
<bibRefCitation id="F52C4B4BE22AFFFBFC813C50FC63FD84" author="Ramazzotti, G." box="[779,990,549,576]" pageId="3" pageNumber="546" pagination="201 - 207" refId="ref6115" refString="Ramazzotti, G. (1966) Tardigradi del Lago Baikal e descrizione di Hypsibius (I.) granulifer baicalensis var. nov. Memorie dell'Istituto Italiano di Idrobiologia, 20, 201 - 207." type="journal article" year="1966">Ramazzotti (1966)</bibRefCitation>
|
||
for
|
||
<taxonomicName id="56BD4D39E22AFFFBFB9A3C50FA21FDFB" box="[1040,1436,549,575]" class="Eutardigrada" family="Hypsibiidae" genus="Isohypsibius" kingdom="Animalia" order="Parachela" pageId="3" pageNumber="546" phylum="Tardigrada" rank="subSpecies" species="granulifer" subSpecies="baicalensis">
|
||
<emphasis id="A3C9EAA8E22AFFFBFB9A3C50FA21FDFB" box="[1040,1436,549,575]" italics="true" pageId="3" pageNumber="546">Isohypsibius granulifer baicalensis</emphasis>
|
||
</taxonomicName>
|
||
(see
|
||
<bibRefCitation id="F52C4B4BE22AFFFBFF413C3CFE37FDA7" author="Ramazzotti, G." box="[203,394,585,611]" pageId="3" pageNumber="546" pagination="201 - 207" refId="ref6115" refString="Ramazzotti, G. (1966) Tardigradi del Lago Baikal e descrizione di Hypsibius (I.) granulifer baicalensis var. nov. Memorie dell'Istituto Italiano di Idrobiologia, 20, 201 - 207." type="journal article" year="1966">Ramazzotti 1966</bibRefCitation>
|
||
<figureCitation id="09862A3FE22AFFFBFE1B3C3CFE5DFDA7" box="[401,480,585,611]" captionStart="FIGURE 1" captionStartId="2.[151,250,1828,1853]" captionTargetBox="[242,1358,762,1816]" captionTargetId="figure@2.[259,1329,770,1817]" captionText="FIGURE 1. Isohypsibius occultus sp. nov. A, B, The arrows indicate details of the cuticular ornamentation (A: paratype, slide No. 5548. B: paratype, slide No. 5608). C, Bucco-pharyngeal apparatus in lateral view (paratype, slide No. 1203). The ridge- shaped apophyses for the insertion of the stylet muscles (arrow a), the short and stout stylet supports (arrow b), and the very developed stylet furcae (arrows c) are evident. D, Bucco-pharyngeal apparatus in dorsal view (holotype, slide No. 5608). The well-developed medio-dorsal transversal ridge of the buccal armature (arrow a), the short and stout stylet supports (arrow b) and the well-developed stylet furcae (arrow c) are evident. Scale bars = 10 µm." httpUri="https://zenodo.org/record/3403907/files/figure.png" pageId="3" pageNumber="546">Fig.1B</figureCitation>
|
||
, and
|
||
<figureCitation id="09862A3FE22AFFFBFD913C3CFDCFFDA7" box="[539,626,584,611]" captionStart="FIGURE 5" captionStartId="8.[151,250,1893,1918]" captionTargetBox="[156,1428,450,1878]" captionTargetId="figure@8.[151,1435,445,1881]" captionText="FIGURE 5. Isohypsibius baicalensis, A, Detail of the cuticular ornamentation. B, Bucco-pharyngeal apparatus. C, Claws of the second pairs of legs. D, Claws of the hind legs. Scale bars = 10 µm. The photographs of Isohypsibius baicalensis, were kindly furnished by Prof. Roberto Bertolani, from a specimen in the Ramazzotti Collection preserved in the Museum of Natural His- tory of Verona, who had kindly gave permission for the examination and photography." httpUri="https://zenodo.org/record/3404771/files/figure.png" pageId="3" pageNumber="546">Fig. 5B</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<paragraph id="910236BAE22BFFFAFB803CACFBB3FD37" blockId="2.[151,1437,153,756]" box="[1034,1038,729,755]" pageId="2" pageNumber="545">.</paragraph>
|
||
<caption id="C5C26632E22BFFFAFF1D3951FC33F836" httpUri="https://zenodo.org/record/3403907/files/figure.png" pageId="2" pageNumber="545" startId="2.[151,250,1828,1853]" targetBox="[242,1358,762,1816]" targetPageId="2">
|
||
<paragraph id="910236BAE22BFFFAFF1D3951FC33F836" pageId="2" pageNumber="545">
|
||
<emphasis id="A3C9EAA8E22BFFFAFF1D3951FEAEF8F9" bold="true" box="[151,275,1828,1853]" pageId="2" pageNumber="545">FIGURE 1.</emphasis>
|
||
<emphasis id="A3C9EAA8E22BFFFAFE903950FE4EF8F9" box="[282,499,1829,1853]" italics="true" pageId="2" pageNumber="545">Isohypsibius occultus</emphasis>
|
||
<emphasis id="A3C9EAA8E22BFFFAFE703950FD35F8F9" bold="true" box="[506,648,1828,1853]" pageId="2" pageNumber="545">sp. nov. A, B,</emphasis>
|
||
The arrows indicate details of the cuticular ornamentation (
|
||
<emphasis id="A3C9EAA8E22BFFFAFB623951FB47F8F9" bold="true" box="[1256,1274,1828,1853]" pageId="2" pageNumber="545">A</emphasis>
|
||
: paratype, slide No. 5548.
|
||
<emphasis id="A3C9EAA8E22BFFFAFE8E393CFEA8F8A5" bold="true" box="[260,277,1865,1889]" pageId="2" pageNumber="545">B</emphasis>
|
||
: paratype, slide No. 5608).
|
||
<emphasis id="A3C9EAA8E22BFFFAFDBF393DFDFAF8A5" bold="true" box="[565,583,1864,1889]" pageId="2" pageNumber="545">C</emphasis>
|
||
, Bucco-pharyngeal apparatus in lateral view (paratype, slide No. 1203). The ridge- shaped apophyses for the insertion of the stylet muscles (arrow a), the short and stout stylet supports (arrow b), and the very developed stylet furcae (arrows c) are evident.
|
||
<emphasis id="A3C9EAA8E22BFFFAFDF339E4FD36F86D" bold="true" box="[633,651,1937,1961]" pageId="2" pageNumber="545">D</emphasis>
|
||
, Bucco-pharyngeal apparatus in dorsal view (holotype, slide No. 5608). The well-developed medio-dorsal transversal ridge of the buccal armature (arrow a), the short and stout stylet supports (arrow b) and the well-developed stylet furcae (arrow c) are evident. Scale bars = 10 µm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="910236BAE22AFFFBFF4D3C18FD8AFD37" blockId="3.[151,1437,153,755]" pageId="3" pageNumber="546">
|
||
The claws, of the
|
||
<taxonomicName id="56BD4D39E22AFFFBFE1B3C18FDA0FD43" authorityName="Thulin" authorityYear="1928" box="[401,541,621,647]" class="Eutardigrada" family="Hypsibiidae" genus="Isohypsibius" kingdom="Animalia" order="Parachela" pageId="3" pageNumber="546" phylum="Tardigrada" rank="genus">
|
||
<emphasis id="A3C9EAA8E22AFFFBFE1B3C18FDA0FD43" box="[401,541,621,647]" italics="true" pageId="3" pageNumber="546">Isohypsibius</emphasis>
|
||
</taxonomicName>
|
||
<typeStatus id="4E068818E22AFFFBFDAE3C18FDE9FD43" box="[548,596,621,647]" pageId="3" pageNumber="546">type</typeStatus>
|
||
(
|
||
<figureCitation id="09862A3FE22AFFFBFDEE3C18FD14FD43" box="[612,681,621,647]" captionStart="FIGURE 2" captionStartId="3.[151,250,1690,1715]" captionTargetBox="[360,1229,773,1673]" captionTargetId="figure@3.[355,1233,769,1677]" captionTargetPageId="3" captionText="FIGURE 2. Isohypsibius occultus sp. nov. A, Claws of the second pair of legs (paratype, slide No. 2774). B, Claws of the fourth pair of legs of the holotype; the arrow indicates the lunule of the internal claw (slide No. 5608). C, Claws of the second pair of legs (paratype, slide No. 2774); arrow ‘a’ indicates a lunule: arrow ‘b’ a cuticular bulge simulating the presence of a bar-like thickening. D, Claws of the hind legs (paratype. slide No. 1203); the arrow indicates the accessory points of a posterior claw. Scale bars = 10 µm." httpUri="https://zenodo.org/record/3066778/files/figure.png" pageId="3" pageNumber="546">Fig. 2</figureCitation>
|
||
), are well developed. The external claws I are
|
||
<quantity id="56459B5FE22AFFFBFB3D3C19FAA8FD43" box="[1207,1301,620,647]" metricMagnitude="-2" metricUnit="m" metricValue="2.54" pageId="3" pageNumber="546" unit="mm" value="25.4">25.4 µm</quantity>
|
||
(
|
||
<emphasis id="A3C9EAA8E22AFFFBFAAC3C1BFA81FD43" box="[1318,1340,622,647]" italics="true" pageId="3" pageNumber="546">pt</emphasis>
|
||
= 61.2); internal claws I was not measurable; external and internal claws II,
|
||
<quantity id="56459B5FE22AFFFBFC033CE4FC5AFD6F" box="[905,999,657,683]" metricMagnitude="-2" metricUnit="m" metricValue="2.77" pageId="3" pageNumber="546" unit="mm" value="27.7">27.7 µm</quantity>
|
||
(
|
||
<emphasis id="A3C9EAA8E22AFFFBFC7D3CE7FBB0FD6F" box="[1015,1037,658,683]" italics="true" pageId="3" pageNumber="546">pt</emphasis>
|
||
= 66.7) and
|
||
<quantity id="56459B5FE22AFFFBFB113CE4FB44FD6F" box="[1179,1273,657,683]" metricMagnitude="-2" metricUnit="m" metricValue="1.81" pageId="3" pageNumber="546" unit="mm" value="18.1">18.1 µm</quantity>
|
||
(
|
||
<emphasis id="A3C9EAA8E22AFFFBFA833CE7FAA2FD6F" box="[1289,1311,658,683]" italics="true" pageId="3" pageNumber="546">pt</emphasis>
|
||
= 43.6) respectively; external and internal claws III were not measurable; posterior and anterior claws IV, 33.0 µm (
|
||
<emphasis id="A3C9EAA8E22AFFFBFABC3CC3FAF1FD0B" box="[1334,1356,694,719]" italics="true" pageId="3" pageNumber="546">pt</emphasis>
|
||
=79.5) and
|
||
<quantity id="56459B5FE22AFFFBFF4D3CACFE98FD37" box="[199,293,729,755]" metricMagnitude="-2" metricUnit="m" metricValue="2.29" pageId="3" pageNumber="546" unit="mm" value="22.9">22.9 µm</quantity>
|
||
(
|
||
<emphasis id="A3C9EAA8E22AFFFBFEBF3CAFFEF6FD37" box="[309,331,730,755]" italics="true" pageId="3" pageNumber="546">pt</emphasis>
|
||
= 55.2) respectively.
|
||
</paragraph>
|
||
<caption id="C5C26632E22AFFFBFF1D38EFFEDFF887" httpUri="https://zenodo.org/record/3066778/files/figure.png" pageId="3" pageNumber="546" startId="3.[151,250,1690,1715]" targetBox="[360,1229,773,1673]" targetPageId="3">
|
||
<paragraph id="910236BAE22AFFFBFF1D38EFFEDFF887" blockId="3.[151,1437,1690,1859]" pageId="3" pageNumber="546">
|
||
<emphasis id="A3C9EAA8E22AFFFBFF1D38EFFEB1F977" bold="true" box="[151,268,1690,1715]" pageId="3" pageNumber="546">FIGURE 2</emphasis>
|
||
.
|
||
<taxonomicName id="56BD4D39E22AFFFBFE9D38EFFE52F976" authority="Pilato & D’Urso & Sabella & Lisi, 2019" authorityName="Pilato & D’Urso & Sabella & Lisi" authorityYear="2019" box="[279,495,1690,1714]" class="Eutardigrada" family="Hypsibiidae" genus="Isohypsibius" kingdom="Animalia" order="Parachela" pageId="3" pageNumber="546" phylum="Tardigrada" rank="species" species="occultus" status="sp. nov.">
|
||
<emphasis id="A3C9EAA8E22AFFFBFE9D38EFFE52F976" box="[279,495,1690,1714]" italics="true" pageId="3" pageNumber="546">Isohypsibius occultus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="A3C9EAA8E22AFFFBFE7F38EEFDDEF977" bold="true" box="[501,611,1690,1715]" pageId="3" pageNumber="546">
|
||
<taxonomicNameLabel id="B8FA57D3E22AFFFBFE7F38EEFDF8F977" box="[501,581,1691,1715]" pageId="3" pageNumber="546" rank="species">sp. nov.</taxonomicNameLabel>
|
||
A,
|
||
</emphasis>
|
||
Claws of the second pair of legs (paratype, slide No. 2774).
|
||
<emphasis id="A3C9EAA8E22AFFFBFB3438EEFB68F977" bold="true" box="[1214,1237,1691,1715]" pageId="3" pageNumber="546">B,</emphasis>
|
||
Claws of the fourth pair of legs of the holotype; the arrow indicates the lunule of the internal claw (slide No. 5608).
|
||
<emphasis id="A3C9EAA8E22AFFFBFBED38CBFB3DF913" bold="true" box="[1127,1152,1726,1751]" pageId="3" pageNumber="546">C,</emphasis>
|
||
Claws of the second pair of legs (paratype, slide No. 2774); arrow ‘a’ indicates a lunule: arrow ‘b’ a cuticular bulge simulating the presence of a bar-like thickening.
|
||
<emphasis id="A3C9EAA8E22AFFFBFE9A3972FE9FF8DB" bold="true" box="[272,290,1799,1823]" pageId="3" pageNumber="546">D</emphasis>
|
||
, Claws of the hind legs (paratype. slide No. 1203); the arrow indicates the accessory points of a posterior claw. Scale bars = 10 µm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="910236BAE22AFFFBFF4D3904FD8AF833" blockId="3.[151,1436,1905,2039]" pageId="3" pageNumber="546">
|
||
Small, flexible lunules are present (
|
||
<figureCitation id="09862A3FE22AFFFBFDD83904FD0DF84F" box="[594,688,1905,1931]" captionStart="FIGURE 2" captionStartId="3.[151,250,1690,1715]" captionTargetBox="[360,1229,773,1673]" captionTargetId="figure@3.[355,1233,769,1677]" captionTargetPageId="3" captionText="FIGURE 2. Isohypsibius occultus sp. nov. A, Claws of the second pair of legs (paratype, slide No. 2774). B, Claws of the fourth pair of legs of the holotype; the arrow indicates the lunule of the internal claw (slide No. 5608). C, Claws of the second pair of legs (paratype, slide No. 2774); arrow ‘a’ indicates a lunule: arrow ‘b’ a cuticular bulge simulating the presence of a bar-like thickening. D, Claws of the hind legs (paratype. slide No. 1203); the arrow indicates the accessory points of a posterior claw. Scale bars = 10 µm." httpUri="https://zenodo.org/record/3066778/files/figure.png" pageId="3" pageNumber="546">Fig. 2 B</figureCitation>
|
||
, arrow; and 2 C, arrow ‘a’), often rolled down under the claw base, and concealed. No cuticular bar is present on the legs, but in some cases, as described and shown by Dastych in
|
||
<taxonomicName id="56BD4D39E22AFFFBFF1D39CCFED5F817" baseAuthorityName="Murray" baseAuthorityYear="1906" box="[151,360,1977,2003]" class="Eutardigrada" family="Hypsibiidae" genus="Isohypsibius" kingdom="Animalia" order="Parachela" pageId="3" pageNumber="546" phylum="Tardigrada" rank="species" species="asper">
|
||
<emphasis id="A3C9EAA8E22AFFFBFF1D39CCFED5F817" box="[151,360,1977,2003]" italics="true" pageId="3" pageNumber="546">Isohypsibius asper</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<bibRefCitation id="F52C4B4BE22AFFFBFEF239CCFDA2F817" author="Dastych, H." box="[376,543,1977,2003]" pageId="3" pageNumber="546" pagination="21 - 43" refId="ref5011" refString="Dastych, H. (2016) Redescription and taxonomic status of the Antarctic water-bear Isohypsibius tetradactyloides (Richters, 1907), as concluded from the rediscovered type material (Tardigrada). Acta Biologica Benrodis, 18, 21 - 43" type="journal article" year="2016">Dastych, 2016</bibRefCitation>
|
||
: figures 19 and 21), a cuticular bulge is present (
|
||
<figureCitation id="09862A3FE22AFFFBFBC839CCFB1CF817" box="[1090,1185,1977,2003]" captionStart="FIGURE 2" captionStartId="3.[151,250,1690,1715]" captionTargetBox="[360,1229,773,1673]" captionTargetId="figure@3.[355,1233,769,1677]" captionTargetPageId="3" captionText="FIGURE 2. Isohypsibius occultus sp. nov. A, Claws of the second pair of legs (paratype, slide No. 2774). B, Claws of the fourth pair of legs of the holotype; the arrow indicates the lunule of the internal claw (slide No. 5608). C, Claws of the second pair of legs (paratype, slide No. 2774); arrow ‘a’ indicates a lunule: arrow ‘b’ a cuticular bulge simulating the presence of a bar-like thickening. D, Claws of the hind legs (paratype. slide No. 1203); the arrow indicates the accessory points of a posterior claw. Scale bars = 10 µm." httpUri="https://zenodo.org/record/3066778/files/figure.png" pageId="3" pageNumber="546">Fig. 2 C</figureCitation>
|
||
, arrow ‘b’) simulating the presence of a bar-like thickening.
|
||
</paragraph>
|
||
<paragraph id="910236BAE22DFFFCFF4D3EECFEE1FEDB" blockId="4.[151,1436,153,287]" pageId="4" pageNumber="547">
|
||
Smooth eggs laid in the exuviae (we found with up
|
||
<specimenCount id="87BBFD33E22DFFFCFC803EECFCEAFF77" box="[778,855,153,179]" pageId="4" pageNumber="547" type="egg">8 eggs</specimenCount>
|
||
).
|
||
<subSubSection id="D9A76531E22DFFFCFF4D3EC8FEE1FEDB" pageId="4" pageNumber="547" type="discussion">
|
||
<emphasis id="A3C9EAA8E22DFFFCFF4D3EC8FE80FF13" bold="true" box="[199,317,189,215]" pageId="4" pageNumber="547">Remarks:</emphasis>
|
||
The
|
||
<typeStatus id="4E068818E22DFFFCFEF33EC8FE59FF13" box="[377,484,189,215]" pageId="4" pageNumber="547" type="paratype">paratypes</typeStatus>
|
||
are similar to the
|
||
<typeStatus id="4E068818E22DFFFCFD3C3EC8FCA5FF13" box="[694,792,189,215]" pageId="4" pageNumber="547" type="holotype">holotype</typeStatus>
|
||
in both qualitative and quantitative characters (
|
||
<tableCitation id="DC3F0301E22DFFFCFABC3EC8FA36FF13" box="[1334,1419,189,215]" captionStart="TABLE 1" captionStartId="4.[151,239,328,352]" captionTargetBox="[161,1407,437,1686]" captionTargetPageId="4" captionText="TABLE 1. Measurements in µm and pt index values of relevant taxonomic characters of five specimens (holotype and four paratypes) of Isohypsibius occultus sp. nov. In the last column the difference between the maximum and the mini- mum value is indicated." httpUri="http://table.plazi.org/id/C5C26632E22DFFFCFF1D3F3DFE2AFE6C" pageId="4" pageNumber="547" tableUuid="C5C26632E22DFFFCFF1D3F3DFE2AFE6C">Table 1</tableCitation>
|
||
). In the buccal armature of some
|
||
<typeStatus id="4E068818E22DFFFCFE703E94FDD5FF3F" box="[506,616,225,251]" pageId="4" pageNumber="547" type="paratype">paratypes</typeStatus>
|
||
, the medio-dorsal ridge may have the anterior margin with incisions, and appears indented.
|
||
</subSubSection>
|
||
</paragraph>
|
||
</treatment>
|
||
</document> |