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<document id="A9F8B849852DF1E66567716652282E9B" ID-DOI="10.11646/zootaxa.3936.3.6" ID-GBIF-Dataset="073d48e4-8e26-428b-b8cd-bac6a606fcbf" ID-ISSN="1175-5326" ID-Zenodo-Dep="238286" ID-ZooBank="9B737E02-FC85-4589-AA38-A3C18DEB6727" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461015848076" checkinUser="plazi" docAuthor="Müller, Andreas" docDate="2015" docId="0967711CFFA1FFBCFF324F23FE562622" docLanguage="en" docName="zt03936p420.pdf" docOrigin="Zootaxa 3936 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Chelostoma (Gyrodromella) handlirschi Schletterer 1889" docType="treatment" docVersion="10" lastPageNumber="416" masterDocId="F55E0964FFA9FFB5FFA54B04FFAF273F" masterDocTitle="Palaearctic Chelostoma bees of the subgenus Gyrodromella (Megachilidae, Osmiini): biology, taxonomy and key to species" masterLastPageNumber="420" masterPageNumber="408" pageNumber="416" updateTime="1698568755497" updateUser="plazi">
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<mods:title id="82C0FE96AA077D37A930504626F6152B">Palaearctic Chelostoma bees of the subgenus Gyrodromella (Megachilidae, Osmiini): biology, taxonomy and key to species</mods:title>
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<mods:namePart id="C9918DB85EBA5151EAC892EDDFE54177">Müller, Andreas</mods:namePart>
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<treatment id="0967711CFFA1FFBCFF324F23FE562622" ID-DOI="http://doi.org/10.5281/zenodo.6106222" ID-GBIF-Taxon="119601566" ID-Zenodo-Dep="6106222" LSID="urn:lsid:plazi:treatment:0967711CFFA1FFBCFF324F23FE562622" httpUri="http://treatment.plazi.org/id/0967711CFFA1FFBCFF324F23FE562622" lastPageId="9" lastPageNumber="416" pageId="8" pageNumber="416">
<subSubSection id="C9D49381FFA1FFBDFF324F23FCFD237D" box="[151,850,1063,1090]" pageId="8" pageNumber="416" type="nomenclature">
<paragraph id="8171C00AFFA1FFBDFF324F23FCFD237D" blockId="8.[151,850,1063,1090]" box="[151,850,1063,1090]" pageId="8" pageNumber="416">
<heading id="DA397766FFA1FFBDFF324F23FCFD237D" bold="true" box="[151,850,1063,1090]" fontSize="11" level="1" pageId="8" pageNumber="416" reason="1">
<taxonomicName id="46CEBB89FFA1FFBDFF324F23FCFD237D" ID-CoL="TWPM" authority="Schletterer, 1889" authorityName="Schletterer" authorityYear="1889" box="[151,850,1063,1090]" class="Insecta" family="Megachilidae" genus="Chelostoma" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="416" phylum="Arthropoda" rank="species" species="handlirschi" subGenus="Gyrodromella">
<emphasis id="B3BA1C18FFA1FFBDFF324F23FCFD237D" bold="true" box="[151,850,1063,1090]" pageId="8" pageNumber="416">
<emphasis id="B3BA1C18FFA1FFBDFF324F23FDD5237E" bold="true" box="[151,634,1063,1089]" italics="true" pageId="8" pageNumber="416">Chelostoma (Gyrodromella) handlirschi</emphasis>
<bibRefCitation id="E55FBDFBFFA1FFBDFD274F2CFCFD237D" author="Schletterer" box="[642,850,1064,1090]" pageId="8" pageNumber="416" refString="Schletterer, A. (1889) Monographie der Bienen-Gattungen Chelostoma Latr. und Heriades Spin. Zoologisches Jahrbuch fur Systematik, 4, 591 - 691." type="journal article" year="1889">Schletterer, 1889</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C9D49381FFA1FFBDFF324F6AFB8023C0" pageId="8" pageNumber="416" type="reference_group">
<paragraph id="8171C00AFFA1FFBDFF324F6AFD62239C" blockId="8.[151,1436,1134,1279]" pageId="8" pageNumber="416">
<treatmentCitationGroup id="A1DEE724FFA1FFBDFF324F6AFD62239C" pageId="8" pageNumber="416">
<taxonomicName id="46CEBB89FFA1FFBDFF324F6AFD3E23BB" ID-CoL="TWPM" authority="Schletterer, 1889: 624" authorityName="Schletterer" authorityPageNumber="624" authorityYear="1889" box="[151,657,1134,1156]" class="Insecta" family="Megachilidae" genus="Chelostoma" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="416" phylum="Arthropoda" rank="species" species="handlirschi">
<emphasis id="B3BA1C18FFA1FFBDFF324F6AFE3E23BB" box="[151,401,1134,1156]" italics="true" pageId="8" pageNumber="416">Chelostoma handlirschi</emphasis>
<treatmentCitation id="006FE61BFFA1FFBDFE3A4F6AFD3E23BB" author="Schletterer" box="[415,657,1134,1156]" page="624" pageId="8" pageNumber="416" year="1889">
<bibRefCitation id="E55FBDFBFFA1FFBDFE3A4F6AFDF923BB" author="Schletterer" box="[415,598,1134,1156]" pageId="8" pageNumber="416" refString="Schletterer, A. (1889) Monographie der Bienen-Gattungen Chelostoma Latr. und Heriades Spin. Zoologisches Jahrbuch fur Systematik, 4, 591 - 691." type="journal article" year="1889">Schletterer, 1889</bibRefCitation>
: 624
</treatmentCitation>
</taxonomicName>
.
<typeStatus id="5E757EA8FFA1FFBDFD074F6BFD7B23BB" box="[674,724,1135,1156]" pageId="8" pageNumber="416">Type</typeStatus>
material:
<typeStatus id="5E757EA8FFA1FFBDFCEF4F6BFC0023BB" box="[842,943,1135,1156]" pageId="8" pageNumber="416" type="lectotype">Lectotype</typeStatus>
♂, by present designation, “Mehadia 1859” (
<collectingCountry id="F9D9809AFFA1FFBDFF6B4F89FE80239C" box="[206,303,1165,1187]" name="Romania" pageId="8" pageNumber="416">Romania</collectingCountry>
), Entomological Collection ETH Zurich.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8171C00AFFA1FFBDFF324FA9FB8023C0" blockId="8.[151,1436,1134,1279]" pageId="8" pageNumber="416">
New records.
<collectingCountry id="F9D9809AFFA1FFBDFE8F4FA8FE1E23FD" box="[298,433,1196,1218]" name="Bulgaria" pageId="8" pageNumber="416">BULGARIA</collectingCountry>
: Arkutino S of Burgas, 19.6.
<date id="F570E6CAFFA1FFBDFD4F4FA8FCFB23FD" box="[746,852,1196,1218]" pageId="8" pageNumber="416" value="1970-07-14">14.7.1970</date>
, 28.5.
<date id="F570E6CAFFA1FFBDFC3A4FA8FC5423FD" box="[927,1019,1196,1218]" pageId="8" pageNumber="416" value="1971-06-09">9.6.1971</date>
; Harmanli,
<date id="F570E6CAFFA1FFBDFBD24FA8FB7023FD" box="[1143,1247,1196,1218]" pageId="8" pageNumber="416" value="1979-05-17">17.5.1979</date>
; Sozopol, 6.1987; Slancev Brjag, 2.6.
<date id="F570E6CAFFA1FFBDFE3D4FCEFE5A23DF" box="[408,501,1226,1248]" pageId="8" pageNumber="416" value="2008-07-06">6.7.2008</date>
;
<quantity id="46366DEFFFA1FFBDFDA74FCEFD8123DF" box="[514,558,1226,1248]" metricMagnitude="3" metricUnit="m" metricValue="5.0" pageId="8" pageNumber="416" unit="km" value="5.0">5km</quantity>
NE of Harmandli,
<date id="F570E6CAFFA1FFBDFD5E4FCEFCCC23DF" box="[763,867,1226,1248]" pageId="8" pageNumber="416" value="2008-06-14">14.6.2008</date>
; Lozenec,
<date id="F570E6CAFFA1FFBDFC714FCEFB9223DF" box="[980,1085,1226,1248]" pageId="8" pageNumber="416" value="2008-06-16">16.6.2008</date>
;
<quantity id="46366DEFFFA1FFBDFBEC4FCEFB2C23DF" box="[1097,1155,1226,1248]" metricMagnitude="4" metricUnit="m" metricValue="1.5" pageId="8" pageNumber="416" unit="km" value="15.0">15km</quantity>
SW of Elchovo, Knazevo,
<date id="F570E6CAFFA1FFBDFF624FEDFE9F23C0" box="[199,304,1257,1279]" pageId="8" pageNumber="416" value="2008-06-19">19.6.2008</date>
; Sinemorec,
<date id="F570E6CAFFA1FFBDFE174FEDFDB423C0" box="[434,539,1257,1279]" pageId="8" pageNumber="416" value="1998-06-22">22.6.1998</date>
.
<collectingCountry id="F9D9809AFFA1FFBDFD814FEDFD0E23C0" box="[548,673,1257,1279]" name="Romania" pageId="8" pageNumber="416">ROMANIA</collectingCountry>
: Cozla W of Turnu Severin,
<date id="F570E6CAFFA1FFBDFC664FEDFB8423C0" box="[963,1067,1257,1279]" pageId="8" pageNumber="416" value="2002-05-26">26.5.2002</date>
.
</paragraph>
</subSubSection>
<subSubSection id="C9D49381FFA1FFBCFF324E28FE562622" lastPageId="9" lastPageNumber="417" pageId="8" pageNumber="416" type="distribution">
<paragraph id="8171C00AFFA1FFBDFF324E28FDAE228F" blockId="8.[151,1437,1324,2033]" pageId="8" pageNumber="416">
<emphasis id="B3BA1C18FFA1FFBDFF324E28FE9F227A" bold="true" box="[151,304,1324,1349]" pageId="8" pageNumber="416">Distribution.</emphasis>
Reliable records exist only for
<collectingCountry id="F9D9809AFFA1FFBDFD154E28FCBA227A" box="[688,789,1324,1349]" name="Romania" pageId="8" pageNumber="416">Romania</collectingCountry>
and
<collectingCountry id="F9D9809AFFA1FFBDFCF34E28FC15227A" box="[854,954,1324,1349]" name="Bulgaria" pageId="8" pageNumber="416">Bulgaria</collectingCountry>
. The records of
<taxonomicName id="46CEBB89FFA1FFBDFB214E29FA85227A" box="[1156,1322,1324,1349]" class="Insecta" family="Megachilidae" genus="Chelostoma" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="416" phylum="Arthropoda" rank="species" species="handlirschi">
<emphasis id="B3BA1C18FFA1FFBDFB214E29FA85227A" box="[1156,1322,1324,1349]" italics="true" pageId="8" pageNumber="416">C. handlirschi</emphasis>
</taxonomicName>
given by
<bibRefCitation id="E55FBDFBFFA1FFBDFF324E4BFE912257" author="Zanden" box="[151,318,1359,1384]" pageId="8" pageNumber="416" refString="Zanden, G. van der (1984) Beitrag zur Megachiliden-Fauna der Volksrepublik Mazedonien in Jugoslawien (Hymenoptera, Apoidea, Megachilidae). Mitteilungen aus dem zoologischen Museums in Berlin, 60, 219 - 223." type="journal article" year="1984">Zanden (1984)</bibRefCitation>
for
<collectingCountry id="F9D9809AFFA1FFBDFED44E4BFE412257" box="[369,494,1359,1384]" name="Macedonia" pageId="8" pageNumber="416">Macedonia</collectingCountry>
and by
<bibRefCitation id="E55FBDFBFFA1FFBDFDE84E4BFCB52257" author="Schletterer" box="[589,794,1359,1384]" pageId="8" pageNumber="416" refString="Schletterer, A. (1889) Monographie der Bienen-Gattungen Chelostoma Latr. und Heriades Spin. Zoologisches Jahrbuch fur Systematik, 4, 591 - 691." type="journal article" year="1889">Schletterer (1889)</bibRefCitation>
for
<collectingCountry id="F9D9809AFFA1FFBDFCE94E4BFC332257" box="[844,924,1359,1384]" name="Turkey" pageId="8" pageNumber="416">Turkey</collectingCountry>
are not free of doubt as the identification of these individuals could not be checked (see Notes). The occurrence of
<taxonomicName id="46CEBB89FFA1FFBDFC164E71FBFC22B2" box="[947,1107,1396,1421]" class="Insecta" family="Megachilidae" genus="Chelostoma" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="416" phylum="Arthropoda" rank="species" species="handlirschi">
<emphasis id="B3BA1C18FFA1FFBDFC164E71FBFC22B2" box="[947,1107,1396,1421]" italics="true" pageId="8" pageNumber="416">C. handlirschi</emphasis>
</taxonomicName>
in northern
<collectingCountry id="F9D9809AFFA1FFBDFB7B4E70FABD22B2" box="[1246,1298,1396,1421]" name="Italy" pageId="8" pageNumber="416">Italy</collectingCountry>
(
<bibRefCitation id="E55FBDFBFFA1FFBDFA844E70FF7F228F" author="Schletterer" pageId="8" pageNumber="416" refString="Schletterer, A. (1889) Monographie der Bienen-Gattungen Chelostoma Latr. und Heriades Spin. Zoologisches Jahrbuch fur Systematik, 4, 591 - 691." type="journal article" year="1889">Schletterer 1889</bibRefCitation>
) seems highly improbable.
</paragraph>
<paragraph id="8171C00AFFA1FFBDFF624EB8FD5522C7" blockId="8.[151,1437,1324,2033]" pageId="8" pageNumber="416">
<emphasis id="B3BA1C18FFA1FFBDFF624EB8FEF122EA" bold="true" box="[199,350,1468,1493]" pageId="8" pageNumber="416">Pollen hosts.</emphasis>
Probably oligolectic on
<taxonomicName id="46CEBB89FFA1FFBDFDDE4EB8FC6522EA" box="[635,970,1468,1493]" class="Magnoliopsida" family="Campanulaceae" genus="Campanula" kingdom="Plantae" order="Asterales" pageId="8" pageNumber="416" phylum="Tracheophyta" rank="genus">
<emphasis id="B3BA1C18FFA1FFBDFDDE4EB8FD5122EA" box="[635,766,1468,1493]" italics="true" pageId="8" pageNumber="416">Campanula</emphasis>
(Campanulaceae)
</taxonomicName>
and possibly also closely related genera (based on the microscopical analysis of 1 pollen load).
</paragraph>
<paragraph id="8171C00AFFA1FFBDFF624D00FDAD2122" blockId="8.[151,1437,1324,2033]" box="[199,514,1540,1565]" pageId="8" pageNumber="416">
<emphasis id="B3BA1C18FFA1FFBDFF624D00FE2A2122" bold="true" box="[199,389,1540,1565]" pageId="8" pageNumber="416">Nesting biology.</emphasis>
Unknown.
</paragraph>
<paragraph id="8171C00AFFA1FFBDFF624D23FF522193" blockId="8.[151,1437,1324,2033]" pageId="8" pageNumber="416">
<emphasis id="B3BA1C18FFA1FFBDFF624D23FEBE217F" bold="true" box="[199,273,1575,1600]" pageId="8" pageNumber="416">Notes.</emphasis>
<taxonomicName id="46CEBB89FFA1FFBDFEBD4D2CFE16217F" box="[280,441,1575,1600]" class="Insecta" family="Megachilidae" genus="Chelostoma" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="416" phylum="Arthropoda" rank="species" species="handlirschi">
<emphasis id="B3BA1C18FFA1FFBDFEBD4D2CFE16217F" box="[280,441,1575,1600]" italics="true" pageId="8" pageNumber="416">C. handlirschi</emphasis>
</taxonomicName>
was obviously unknown to most European bee taxonomists including K. Warncke, who treated
<taxonomicName id="46CEBB89FFA1FFBDFF4E4D49FED3215A" box="[235,380,1612,1637]" class="Insecta" family="Megachilidae" genus="Chelostoma" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="416" phylum="Arthropoda" rank="species" species="hebraeum">
<emphasis id="B3BA1C18FFA1FFBDFF4E4D49FED3215A" box="[235,380,1612,1637]" italics="true" pageId="8" pageNumber="416">C. hebraeum</emphasis>
</taxonomicName>
and
<taxonomicName id="46CEBB89FFA1FFBDFE104D49FD99215B" box="[437,566,1613,1636]" class="Insecta" family="Megachilidae" genus="Chelostoma" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="416" phylum="Arthropoda" rank="species" species="nasutum">
<emphasis id="B3BA1C18FFA1FFBDFE104D49FD99215B" box="[437,566,1613,1636]" italics="true" pageId="8" pageNumber="416">C. nasutum</emphasis>
</taxonomicName>
as subspecies of
<taxonomicName id="46CEBB89FFA1FFBDFD594D49FC32215A" box="[764,925,1612,1637]" class="Insecta" family="Megachilidae" genus="Chelostoma" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="416" phylum="Arthropoda" rank="species" species="handlirschi">
<emphasis id="B3BA1C18FFA1FFBDFD594D49FC32215A" box="[764,925,1612,1637]" italics="true" pageId="8" pageNumber="416">C. handlirschi</emphasis>
</taxonomicName>
. Thus, literature records of
<taxonomicName id="46CEBB89FFA1FFBDFB7E4D49FAD2215A" box="[1243,1405,1612,1637]" class="Insecta" family="Megachilidae" genus="Chelostoma" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="416" phylum="Arthropoda" rank="species" species="handlirschi">
<emphasis id="B3BA1C18FFA1FFBDFB7E4D49FAD2215A" box="[1243,1405,1612,1637]" italics="true" pageId="8" pageNumber="416">C. handlirschi</emphasis>
</taxonomicName>
as well as specimens labelled as
<taxonomicName id="46CEBB89FFA1FFBDFE594D74FD0F21B7" box="[508,672,1647,1672]" class="Insecta" family="Megachilidae" genus="Chelostoma" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="416" phylum="Arthropoda" rank="species" species="handlirschi">
<emphasis id="B3BA1C18FFA1FFBDFE594D74FD0F21B7" box="[508,672,1647,1672]" italics="true" pageId="8" pageNumber="416">C. handlirschi</emphasis>
</taxonomicName>
are expected to often refer to
<taxonomicName id="46CEBB89FFA1FFBDFBAA4D74FB0221B7" box="[1039,1197,1648,1672]" class="Insecta" family="Megachilidae" genus="Chelostoma" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="416" phylum="Arthropoda" rank="species" species="aegaeicum">
<emphasis id="B3BA1C18FFA1FFBDFBAA4D74FB0221B7" box="[1039,1197,1648,1672]" italics="true" pageId="8" pageNumber="416">C. aegaeicum</emphasis>
</taxonomicName>
,
<taxonomicName id="46CEBB89FFA1FFBDFB1A4D74FAFB21B7" box="[1215,1364,1647,1672]" class="Insecta" family="Megachilidae" genus="Chelostoma" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="416" phylum="Arthropoda" rank="species" species="hebraeum">
<emphasis id="B3BA1C18FFA1FFBDFB1A4D74FAFB21B7" box="[1215,1364,1647,1672]" italics="true" pageId="8" pageNumber="416">C. hebraeum</emphasis>
</taxonomicName>
or
<taxonomicName id="46CEBB89FFA1FFBDFA274D74FF592193" class="Insecta" family="Megachilidae" genus="Chelostoma" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="416" phylum="Arthropoda" rank="species" species="nasutum">
<emphasis id="B3BA1C18FFA1FFBDFA274D74FF592193" italics="true" pageId="8" pageNumber="416">C. nasutum</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8171C00AFFA1FFBDFF624DB3FE412097" blockId="8.[151,1437,1324,2033]" pageId="8" pageNumber="416">
The
<typeStatus id="5E757EA8FFA1FFBDFF5E4DBCFEFD21EF" box="[251,338,1720,1744]" pageId="8" pageNumber="416" type="syntype">syntype</typeStatus>
series of
<taxonomicName id="46CEBB89FFA1FFBDFE674DBCFDCC21EF" box="[450,611,1719,1744]" class="Insecta" family="Megachilidae" genus="Chelostoma" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="416" phylum="Arthropoda" rank="species" species="handlirschi">
<emphasis id="B3BA1C18FFA1FFBDFE674DBCFDCC21EF" box="[450,611,1719,1744]" italics="true" pageId="8" pageNumber="416">C. handlirschi</emphasis>
</taxonomicName>
consisting of males and females from “Nord-Italien,
<collectingCountry id="F9D9809AFFA1FFBDFB614DBCFAB721EF" box="[1220,1304,1720,1744]" name="Hungary" pageId="8" pageNumber="416">Ungarn</collectingCountry>
(Mehadia), Kleinasien” (
<bibRefCitation id="E55FBDFBFFA1FFBDFE8C4DD8FE4521CB" author="Schletterer" box="[297,490,1756,1781]" pageId="8" pageNumber="416" refString="Schletterer, A. (1889) Monographie der Bienen-Gattungen Chelostoma Latr. und Heriades Spin. Zoologisches Jahrbuch fur Systematik, 4, 591 - 691." type="journal article" year="1889">Schletterer, 1889</bibRefCitation>
) could be located neither in the Natural History Museum Wien nor in the Hungarian Natural History Museum Budapest (M. Vizek and Z. Vas, personal communication), suggesting that Schletterer did not deposit the
<typeStatus id="5E757EA8FFA1FFBDFEE44C21FE0C2002" box="[321,419,1829,1853]" pageId="8" pageNumber="416" type="syntype">syntypes</typeStatus>
in these two museums as mentioned in the original publication. However, a male specimen collected at Mehadia in 1859 and labelled as
<taxonomicName id="46CEBB89FFA1FFBDFD294C4CFC84205F" box="[652,811,1863,1888]" class="Insecta" family="Megachilidae" genus="Chelostoma" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="416" phylum="Arthropoda" rank="species" species="handlirschi">
<emphasis id="B3BA1C18FFA1FFBDFD294C4CFC84205F" box="[652,811,1863,1888]" italics="true" pageId="8" pageNumber="416">C. handlirschi</emphasis>
</taxonomicName>
with the handwriting probably of Schletterer was found in the Schulthess collection at ETH Zürich. This male, which most likely belongs to the
<typeStatus id="5E757EA8FFA1FFBDFB3B4C69FB5A20BA" box="[1182,1269,1901,1925]" pageId="8" pageNumber="416" type="syntype">syntype</typeStatus>
series, is here newly designated as
<typeStatus id="5E757EA8FFA1FFBDFEDA4C8BFE462097" box="[383,489,1935,1960]" pageId="8" pageNumber="416" type="lectotype">lectotype</typeStatus>
.
</paragraph>
<paragraph id="8171C00AFFA1FFBCFF624CB0FE562622" blockId="8.[151,1437,1324,2033]" lastBlockId="9.[151,1436,151,285]" lastPageId="9" lastPageNumber="417" pageId="8" pageNumber="416">
The description of the female of
<taxonomicName id="46CEBB89FFA1FFBDFD9B4CB1FD7020F2" box="[574,735,1972,1997]" class="Insecta" family="Megachilidae" genus="Chelostoma" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="416" phylum="Arthropoda" rank="species" species="handlirschi">
<emphasis id="B3BA1C18FFA1FFBDFD9B4CB1FD7020F2" box="[574,735,1972,1997]" italics="true" pageId="8" pageNumber="416">C. handlirschi</emphasis>
</taxonomicName>
by
<bibRefCitation id="E55FBDFBFFA1FFBDFCA84CB0FC7620F3" author="Schletterer" box="[781,985,1972,1997]" pageId="8" pageNumber="416" refString="Schletterer, A. (1889) Monographie der Bienen-Gattungen Chelostoma Latr. und Heriades Spin. Zoologisches Jahrbuch fur Systematik, 4, 591 - 691." type="journal article" year="1889">Schletterer (1889)</bibRefCitation>
does not agree with the morphology of typical
<taxonomicName id="46CEBB89FFA1FFBDFF4E4CDDFE2320CE" box="[235,396,2008,2033]" class="Insecta" family="Megachilidae" genus="Chelostoma" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="416" phylum="Arthropoda" rank="species" species="handlirschi">
<emphasis id="B3BA1C18FFA1FFBDFF4E4CDDFE2320CE" box="[235,396,2008,2033]" italics="true" pageId="8" pageNumber="416">C. handlirschi</emphasis>
</taxonomicName>
females collected together with males identical to the
<typeStatus id="5E757EA8FFA1FFBDFC534CDCFBF220CE" box="[1014,1117,2008,2033]" pageId="8" pageNumber="416" type="lectotype">lectotype</typeStatus>
of
<taxonomicName id="46CEBB89FFA1FFBDFB264CDDFA8C20CE" box="[1155,1315,2008,2033]" class="Insecta" family="Megachilidae" genus="Chelostoma" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="416" phylum="Arthropoda" rank="species" species="handlirschi">
<emphasis id="B3BA1C18FFA1FFBDFB264CDDFA8C20CE" box="[1155,1315,2008,2033]" italics="true" pageId="8" pageNumber="416">C. handlirschi</emphasis>
</taxonomicName>
. Thus, the two sexes described by Schletterer are not conspecific, which suggests that the information Schletterer gives on the species distribution range might be partly erroneous (see Distribution).
<collectingCountry id="F9D9809AFFA0FFBCFC604BB8FC4B27EB" box="[965,996,188,212]" name="American Samoa" pageId="9" pageNumber="417">As</collectingCountry>
no female
<typeStatus id="5E757EA8FFA0FFBCFBC64BB9FB1527EA" box="[1123,1210,189,213]" pageId="9" pageNumber="417" type="syntype">syntype</typeStatus>
could be found, the species identity of Schletterers females remains unclear, the characters listed by Schletterer, however, fit best to
<taxonomicName id="46CEBB89FFA0FFBCFA274BE4FE562622" authority="Muller." authorityName="Muller." class="Magnoliopsida" family="Campanulaceae" genus="Campanula" kingdom="Plantae" order="Asterales" pageId="9" pageNumber="417" phylum="Tracheophyta" rank="species" species="comosum" subGenus="Chelostoma">
<emphasis id="B3BA1C18FFA0FFBCFA274BE4FA3327C8" box="[1410,1436,224,247]" italics="true" pageId="9" pageNumber="417">C.</emphasis>
(
<emphasis id="B3BA1C18FFA0FFBCFF054A00FE892622" box="[160,294,260,285]" italics="true" pageId="9" pageNumber="417">Chelostoma</emphasis>
)
<emphasis id="B3BA1C18FFA0FFBCFE934A01FE0F2623" box="[310,416,261,284]" italics="true" pageId="9" pageNumber="417">comosum</emphasis>
Müller.
</taxonomicName>
</paragraph>
</subSubSection>
</treatment>
</document>