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<document id="211940575D960931B54B55A801C84ED2" ID-CLB-Dataset="32249" ID-DOI="10.11646/zootaxa.4313.1.1" ID-GBIF-Dataset="d8cc2cd8-8410-49aa-a75d-7a41d9f52b26" ID-ISSN="1175-5326" ID-Zenodo-Dep="883161" ID-ZooBank="A5Fdfba5-F992-44A8-84C2-1756C943C19B" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1504182989959" checkinUser="plazi" docAuthor="Bush, Sarah E." docDate="2017" docId="832187E9FEBCFEFAFF746664FADEFE0E" docLanguage="en" docName="zootaxa.4313.1.1.pdf" docOrigin="Zootaxa 4313 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Couala angulata Bush 2017" docType="treatment" docVersion="17" lastPageNumber="314" masterDocId="7F18FF91FF8AFFC3FFE36200FFC9FFA3" masterDocTitle="Morphological revision of the hyperdiverse Brueelia - complex (Insecta: Phthiraptera: Ischnocera: Philopteridae) with new taxa, checklists and generic key" masterLastPageNumber="443" masterPageNumber="1" pageNumber="311" updateTime="1698467717597" updateUser="ExternalLinkService">
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<mods:title id="866705505BAEAF0E98D95A52A104BDD8">Morphological revision of the hyperdiverse Brueelia - complex (Insecta: Phthiraptera: Ischnocera: Philopteridae) with new taxa, checklists and generic key</mods:title>
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<mods:namePart id="8FACF3A52A0785543B9671D3D7B0F4B1">Bush, Sarah E.</mods:namePart>
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<treatment id="832187E9FEBCFEFAFF746664FADEFE0E" ID-DOI="http://doi.org/10.5281/zenodo.5297077" ID-GBIF-Taxon="133100158" ID-Zenodo-Dep="5297077" LSID="urn:lsid:plazi:treatment:832187E9FEBCFEFAFF746664FADEFE0E" httpUri="http://treatment.plazi.org/id/832187E9FEBCFEFAFF746664FADEFE0E" lastPageId="313" lastPageNumber="314" pageId="310" pageNumber="311">
<subSubSection id="43926574FEBCFEF5FF746664FE8AFB3C" pageId="310" pageNumber="311" type="nomenclature">
<paragraph id="0B3736FFFEBCFEF5FF746664FDDAFBDD" blockId="310.[151,531,1124,1183]" box="[151,531,1124,1150]" pageId="310" pageNumber="311">
<heading id="507F8193FEBCFEF5FF746664FDDAFBDD" bold="true" box="[151,531,1124,1150]" fontSize="11" level="1" pageId="310" pageNumber="311" reason="1">
<taxonomicName id="CC884D7CFEBCFEF5FF746664FDDAFBDD" authority="(Piaget, 1880)" authorityName="Bush" authorityYear="2017" baseAuthorityName="Piaget" baseAuthorityYear="1880" box="[151,531,1124,1150]" class="Insecta" family="Philopteridae" genus="Couala" kingdom="Animalia" order="Phthiraptera" pageId="310" pageNumber="311" phylum="Arthropoda" rank="species" species="angulata">
<emphasis id="39FCEAEDFEBCFEF5FF746664FDDAFBDD" bold="true" box="[151,531,1124,1150]" pageId="310" pageNumber="311">
<emphasis id="39FCEAEDFEBCFEF5FF746664FEA8FBDD" bold="true" box="[151,353,1124,1150]" italics="true" pageId="310" pageNumber="311">Couala angulata</emphasis>
(
<bibRefCitation id="6F194B0EFEBCFEF5FE936664FDC2FBDD" author="Piaget" box="[368,523,1124,1150]" pageId="310" pageNumber="311" refString="Piaget, E. (1880) Les Pediculines. Essai Monographique. Two volumes. E. J. Brill, Leide. xxxix + 714 pp., 56 pls." type="book" year="1880">Piaget, 1880</bibRefCitation>
)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="0B3736FFFEBCFEF5FF746686FE8AFB3C" blockId="310.[151,531,1124,1183]" box="[151,323,1158,1183]" pageId="310" pageNumber="311">
(
<figureCitation id="93B32A7AFEBCFEF5FF7C6686FEF2FB3C" box="[159,315,1158,1183]" captionStart-0="FIGURE 518 519" captionStart-1="FIGURES 520 523" captionStartId-0="311.[151,250,1889,1911]" captionStartId-1="312.[151,264,1947,1969]" captionTargetBox-0="[166,1436,494,1842]" captionTargetBox-1="[194,1343,198,1893]" captionTargetId-0="figure@311.[151,1436,481,1867]" captionTargetId-1="figure@312.[163,1407,193,1902]" captionTargetPageId-0="311" captionTargetPageId-1="312" captionText-0="FIGURE 518 519 Couala angulata (Piaget, 1880) n. comb. ex Coua caerulea: 518, male habitus, dorsal and ventral views. 519, female habitus, dorsal and ventral views." captionText-1="FIGURES 520 523 Couala angulata (Piaget, 1880) n. comb. ex Coua caerulea: 520, male head, dorsal and ventral views. 521, male genitalia, dorsal view. 522, male mesosome and parameres, ventral view. 523, female subgenital plate and vulval margin, ventral view." httpUri-0="https://zenodo.org/record/883436/files/figure.png" httpUri-1="https://zenodo.org/record/883438/files/figure.png" pageId="310" pageNumber="311">Figs 518523</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="43926574FEBCFEF5FF7466D0FCCBFA81" pageId="310" pageNumber="311" type="reference_group">
<paragraph id="0B3736FFFEBCFEF5FF7466D0FDC2FB47" blockId="310.[151,770,1230,1314]" box="[151,523,1230,1253]" pageId="310" pageNumber="311">
<treatmentCitationGroup id="2B9811D1FEBCFEF5FF7466D0FDC2FB47" box="[151,523,1230,1253]" pageId="310" pageNumber="311">
<taxonomicName id="CC884D7CFEBCFEF5FF7466D0FDCEFB47" authority="Piaget, 1880: 134" authorityName="Piaget" authorityPageNumber="134" authorityYear="1880" box="[151,519,1230,1253]" class="Insecta" family="Philopteridae" genus="Nirmus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Phthiraptera" pageId="310" pageNumber="311" phylum="Arthropoda" rank="species" species="angulatus">
<emphasis id="39FCEAEDFEBCFEF5FF7466D0FE85FB47" box="[151,332,1230,1253]" italics="true" pageId="310" pageNumber="311">Nirmus angulatus</emphasis>
<treatmentCitation id="8A2910EEFEBCFEF5FEB166CEFDCEFB47" author="Piaget" box="[338,519,1230,1252]" page="134" pageId="310" pageNumber="311" year="1880">
<bibRefCitation id="6F194B0EFEBCFEF5FEB166CEFE1DFB47" author="Piaget" box="[338,468,1230,1252]" pageId="310" pageNumber="311" refString="Piaget, E. (1880) Les Pediculines. Essai Monographique. Two volumes. E. J. Brill, Leide. xxxix + 714 pp., 56 pls." type="book" year="1880">Piaget, 1880</bibRefCitation>
: 134
</treatmentCitation>
</taxonomicName>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="0B3736FFFEBCFEF5FF7466EDFCCBFA81" blockId="310.[151,770,1230,1314]" pageId="310" pageNumber="311">
<treatmentCitationGroup id="2B9811D1FEBCFEF5FF7466EDFCCBFA81" pageId="310" pageNumber="311">
<taxonomicName id="CC884D7CFEBCFEF5FF7466EDFDCEFAA0" authority="(Piaget, 1880)" baseAuthorityName="Piaget" baseAuthorityYear="1880" box="[151,519,1261,1283]" class="Insecta" family="Philopteridae" genus="Degeeriella" kingdom="Animalia" order="Phthiraptera" pageId="310" pageNumber="311" phylum="Arthropoda" rank="species" species="angulata">
<emphasis id="39FCEAEDFEBCFEF5FF7466EDFEA7FAA0" box="[151,366,1261,1283]" italics="true" pageId="310" pageNumber="311">Degeeriella angulata</emphasis>
(
<bibRefCitation id="6F194B0EFEBCFEF5FE9866EDFDC9FAA0" author="Piaget" box="[379,512,1261,1283]" pageId="310" pageNumber="311" refString="Piaget, E. (1880) Les Pediculines. Essai Monographique. Two volumes. E. J. Brill, Leide. xxxix + 714 pp., 56 pls." type="book" year="1880">Piaget, 1880</bibRefCitation>
)
</taxonomicName>
;
<treatmentCitation id="8A2910EEFEBCFEF5FDF266EDFD29FAA0" author="Harrison" box="[529,736,1261,1283]" page="108" pageId="310" pageNumber="311" year="1916">
<bibRefCitation id="6F194B0EFEBCFEF5FDF266EDFD64FAA0" author="Harrison" box="[529,685,1261,1283]" pageId="310" pageNumber="311" refString="Harrison, L. (1916) The genera and species of Mallophaga. Parasitology, 9, 1 - 156." type="journal article" year="1916">Harrison, 1916</bibRefCitation>
: 108
</treatmentCitation>
.
<taxonomicName id="CC884D7CFEBCFEF5FF74670CFE2FFA81" authority="(Piaget, 1880)" baseAuthorityName="Piaget" baseAuthorityYear="1880" box="[151,486,1292,1314]" class="Insecta" family="Philopteridae" genus="Brueelia" kingdom="Animalia" order="Phthiraptera" pageId="310" pageNumber="311" phylum="Arthropoda" rank="species" species="angulata">
<emphasis id="39FCEAEDFEBCFEF5FF74670CFE87FA81" box="[151,334,1292,1314]" italics="true" pageId="310" pageNumber="311">Brueelia angulata</emphasis>
(
<bibRefCitation id="6F194B0EFEBCFEF5FEB8670CFE29FA81" author="Piaget" box="[347,480,1292,1314]" pageId="310" pageNumber="311" refString="Piaget, E. (1880) Les Pediculines. Essai Monographique. Two volumes. E. J. Brill, Leide. xxxix + 714 pp., 56 pls." type="book" year="1880">Piaget, 1880</bibRefCitation>
)
</taxonomicName>
;
<treatmentCitation id="8A2910EEFEBCFEF5FE12670CFD37FA81" author="Hopkins" box="[497,766,1292,1314]" page="53" pageId="310" pageNumber="311" year="1952">
<bibRefCitation id="6F194B0EFEBCFEF5FE12670CFD11FA81" author="Hopkins" box="[497,728,1292,1314]" pageId="310" pageNumber="311" refString="Hopkins, G. H. &amp; Clay, T. (1952) A check list of the genera &amp; species of Mallophaga. British Museum (Natural History), London. 362 pp." type="book" year="1952">Hopkins &amp; Clay, 1952</bibRefCitation>
: 53
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="43926574FEBCFEF5FF746749FD2AFA3F" pageId="310" pageNumber="311" type="materials_examined">
<paragraph id="0B3736FFFEBCFEF5FF746749FDA3FADD" blockId="310.[151,932,1353,1436]" pageId="310" pageNumber="311">
<emphasis id="39FCEAEDFEBCFEF5FF746749FECCFAFC" bold="true" box="[151,261,1353,1375]" pageId="310" pageNumber="311">Type host.</emphasis>
<taxonomicName id="CC884D7CFEBCFEF5FEEF674AFDBBFAFC" authority="(Pontoppidan, 1763)" baseAuthorityName="Pontoppidan" baseAuthorityYear="1763" box="[268,626,1353,1375]" class="Aves" family="Strigidae" genus="Asio" kingdom="Animalia" order="Strigiformes" pageId="310" pageNumber="311" phylum="Chordata" rank="species" species="flammeus">
<emphasis id="39FCEAEDFEBCFEF5FEEF674AFE69FAFC" box="[268,416,1353,1375]" italics="true" pageId="310" pageNumber="311">Asio flammeus</emphasis>
(Pontoppidan, 1763)
</taxonomicName>
—short-eared owl (in
<emphasis id="39FCEAEDFEBCFEF5FCB46749FC57FAFC" bold="true" box="[855,926,1353,1375]" pageId="310" pageNumber="311">Error)</emphasis>
.
<emphasis id="39FCEAEDFEBCFEF5FF746768FEECFADD" bold="true" box="[151,293,1384,1406]" pageId="310" pageNumber="311">Type locality.</emphasis>
Zoological Garden, Rotterdam.
</paragraph>
<paragraph id="0B3736FFFEBCFEF5FF746786FD2AFA3F" blockId="310.[151,932,1353,1436]" box="[151,739,1414,1436]" pageId="310" pageNumber="311">
<emphasis id="39FCEAEDFEBCFEF5FF746786FED8FA3F" bold="true" box="[151,273,1414,1436]" pageId="310" pageNumber="311">Other host.</emphasis>
<taxonomicName id="CC884D7CFEBCFEF5FEF46787FD97FA3F" authority="(Linnaeus, 1766)" baseAuthorityName="Linnaeus" baseAuthorityYear="1766" box="[279,606,1414,1436]" class="Aves" family="Cuculidae" genus="Coua" kingdom="Animalia" order="Cuculiformes" pageId="310" pageNumber="311" phylum="Chordata" rank="species" species="caerulea">
<emphasis id="39FCEAEDFEBCFEF5FEF46787FE64FA3F" box="[279,429,1414,1436]" italics="true" pageId="310" pageNumber="311">Coua caerulea</emphasis>
(Linnaeus, 1766)
</taxonomicName>
—blue
<taxonomicName id="CC884D7CFEBCFEF5FD486787FD17FA3F" authorityName="Schinz" authorityYear="1821" box="[683,734,1415,1436]" class="Aves" family="Cuculidae" genus="Coua" kingdom="Animalia" order="Cuculiformes" pageId="310" pageNumber="420" phylum="Chordata" rank="genus">coua</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="43926574FEBCFEF4FF7467C9FE3CFE0E" lastPageId="311" lastPageNumber="312" pageId="310" pageNumber="311" type="description">
<paragraph id="0B3736FFFEBCFEF5FF7467C9FC93F919" blockId="310.[151,1436,1481,2010]" pageId="310" pageNumber="311">
<emphasis id="39FCEAEDFEBCFEF5FF7467C9FE78FA42" bold="true" box="[151,433,1481,1506]" pageId="310" pageNumber="311">
Description.
<emphasis id="39FCEAEDFEBCFEF5FED067C9FE78FA42" bold="true" box="[307,433,1481,1506]" italics="true" pageId="310" pageNumber="311">Both sexes</emphasis>
</emphasis>
. Head shape, structure, and chaetotaxy as in genus description and
<figureCitation id="93B32A7AFEBCFEF5FB2867C9FAE6FA41" box="[1227,1327,1481,1506]" captionStart="FIGURES 520 523" captionStartId="312.[151,264,1947,1969]" captionTargetBox="[194,1343,198,1893]" captionTargetId="figure@312.[163,1407,193,1902]" captionTargetPageId="312" captionText="FIGURES 520 523 Couala angulata (Piaget, 1880) n. comb. ex Coua caerulea: 520, male head, dorsal and ventral views. 521, male genitalia, dorsal view. 522, male mesosome and parameres, ventral view. 523, female subgenital plate and vulval margin, ventral view." httpUri="https://zenodo.org/record/883438/files/figure.png" pageId="310" pageNumber="311">Fig. 520</figureCitation>
. Temples acutely angular, occiput deeply concave. Marginal and marginal temporal carinae very broad. Preantennal, preocular, and postocular nodi very large. Ventral carina broad and diffuse anterior to pulvinus. Coni long, reaching well beyond distal margin of scape. Lateral margin of antennal canal clearly demarcated, continuing to posterior margin of head, which it reaches just median to
<emphasis id="39FCEAEDFEBCFEF5FD2E645AFCCBF9D1" box="[717,770,1626,1650]" italics="true" pageId="310" pageNumber="311">
<taxonomicName id="CC884D7CFEBCFEF5FD2E645AFD3DF9D1" box="[717,756,1626,1650]" class="Insecta" family="Philopteridae" genus="Titanomessor" kingdom="Animalia" order="Phthiraptera" pageId="310" pageNumber="311" phylum="Arthropoda" rank="species" species="mts">mts</taxonomicName>
3
</emphasis>
; median margin of antennal canal clearly demarcated for anterior fourth;
<emphasis id="39FCEAEDFEBCFEF5FEA9647EFEB7F935" box="[330,382,1662,1686]" italics="true" pageId="310" pageNumber="311">
<taxonomicName id="CC884D7CFEBCFEF5FEA9647EFEB8F935" box="[330,369,1662,1686]" class="Insecta" family="Philopteridae" genus="Titanomessor" kingdom="Animalia" order="Phthiraptera" pageId="310" pageNumber="311" phylum="Arthropoda" rank="species" species="mts">mts</taxonomicName>
2
</emphasis>
curved, spine-like, just lateral to
<emphasis id="39FCEAEDFEBCFEF5FD18647EFCF9F935" box="[763,816,1662,1686]" italics="true" pageId="310" pageNumber="311">
<taxonomicName id="CC884D7CFEBCFEF5FD18647EFCEBF935" box="[763,802,1662,1686]" class="Insecta" family="Philopteridae" genus="Titanomessor" kingdom="Animalia" order="Phthiraptera" pageId="310" pageNumber="311" phylum="Arthropoda" rank="species" species="mts">mts</taxonomicName>
3
</emphasis>
, and clearly ventral. Gular plate pointed. Thoracic and abdominal segments as in genus description and
<figureCitation id="93B32A7AFEBCFEF5FD5964A1FC9FF919" box="[698,854,1697,1722]" captionStart="FIGURE 518 519" captionStartId="311.[151,250,1889,1911]" captionTargetBox="[166,1436,494,1842]" captionTargetId="figure@311.[151,1436,481,1867]" captionTargetPageId="311" captionText="FIGURE 518 519 Couala angulata (Piaget, 1880) n. comb. ex Coua caerulea: 518, male habitus, dorsal and ventral views. 519, female habitus, dorsal and ventral views." httpUri="https://zenodo.org/record/883436/files/figure.png" pageId="310" pageNumber="311">Figs 518519</figureCitation>
.
</paragraph>
<paragraph id="0B3736FFFEBCFEF4FF2464C5FBBBFF76" blockId="310.[151,1436,1481,2010]" lastBlockId="311.[151,1436,151,429]" lastPageId="311" lastPageNumber="312" pageId="310" pageNumber="311">
<collectingRegion id="C94CF81DFEBCFEF5FF2464C5FECBF97D" box="[199,258,1733,1758]" country="Maldives" name="Maale" pageId="310" pageNumber="311">
<emphasis id="39FCEAEDFEBCFEF5FF2464C5FECBF97D" bold="true" box="[199,258,1733,1758]" italics="true" pageId="310" pageNumber="311">Male</emphasis>
</collectingRegion>
. Subgenital plate (
<figureCitation id="93B32A7AFEBCFEF5FE3C64C5FD8BF97D" box="[479,578,1733,1758]" captionStart="FIGURE 518 519" captionStartId="311.[151,250,1889,1911]" captionTargetBox="[166,1436,494,1842]" captionTargetId="figure@311.[151,1436,481,1867]" captionTargetPageId="311" captionText="FIGURE 518 519 Couala angulata (Piaget, 1880) n. comb. ex Coua caerulea: 518, male habitus, dorsal and ventral views. 519, female habitus, dorsal and ventral views." httpUri="https://zenodo.org/record/883436/files/figure.png" pageId="310" pageNumber="311">Fig. 518</figureCitation>
) trapezoidal, narrowing at posterior extreme. Abdominal chaetotaxy as in
<tableCitation id="460A0344FEBCFEF5FF7464E9FF25F8A1" box="[151,236,1769,1794]" captionStart="TABLE 2" captionStartId="30.[151,239,796,819]" captionText="TABLE 2. Summary of abdominal chaetotaxy of segments II VIII for the Brueelia - complex, including both genera and subgenera. In genera where chaetotaxy differs between species group (marked with *), only that of the type species is given here, and more detailed accounts are given separately under the respective genus entries and in Tables 3 12. Trichoid setae of segment VIII are present in all genera, and are not listed. Abbreviations: ps = paratergal seta; aps = accessory post-spiracular seta; psps = principal post-spiracular seta; tps = tergal posterior seta; ss = sutural seta; sts = sternal seta." pageId="310" pageNumber="311">Table 2</tableCitation>
and
<figureCitation id="93B32A7AFEBCFEF5FEC464E9FE43F8A1" box="[295,394,1769,1794]" captionStart="FIGURE 518 519" captionStartId="311.[151,250,1889,1911]" captionTargetBox="[166,1436,494,1842]" captionTargetId="figure@311.[151,1436,481,1867]" captionTargetPageId="311" captionText="FIGURE 518 519 Couala angulata (Piaget, 1880) n. comb. ex Coua caerulea: 518, male habitus, dorsal and ventral views. 519, female habitus, dorsal and ventral views." httpUri="https://zenodo.org/record/883436/files/figure.png" pageId="310" pageNumber="311">Fig. 518</figureCitation>
.
<collectingRegion id="C94CF81DFEBCFEF5FE7464E9FE18F8A1" box="[407,465,1769,1794]" country="Maldives" name="Maale" pageId="310" pageNumber="311">Male</collectingRegion>
genitalia (
<figureCitation id="93B32A7AFEBCFEF5FDA864E9FD22F8A1" box="[587,747,1769,1794]" captionStart="FIGURES 520 523" captionStartId="312.[151,264,1947,1969]" captionTargetBox="[194,1343,198,1893]" captionTargetId="figure@312.[163,1407,193,1902]" captionTargetPageId="312" captionText="FIGURES 520 523 Couala angulata (Piaget, 1880) n. comb. ex Coua caerulea: 520, male head, dorsal and ventral views. 521, male genitalia, dorsal view. 522, male mesosome and parameres, ventral view. 523, female subgenital plate and vulval margin, ventral view." httpUri="https://zenodo.org/record/883438/files/figure.png" pageId="310" pageNumber="311">Figs 521522</figureCitation>
) distinct. Basal apodeme long, slender. Proximal mesosome fused to and not distinguishable from basal apodeme. Gonopore elongated, open distally, with anterior extension on ventral side (
<figureCitation id="93B32A7AFEBCFEF5FECC6531FE5CF8E9" box="[303,405,1841,1866]" captionStart="FIGURES 520 523" captionStartId="312.[151,264,1947,1969]" captionTargetBox="[194,1343,198,1893]" captionTargetId="figure@312.[163,1407,193,1902]" captionTargetPageId="312" captionText="FIGURES 520 523 Couala angulata (Piaget, 1880) n. comb. ex Coua caerulea: 520, male head, dorsal and ventral views. 521, male genitalia, dorsal view. 522, male mesosome and parameres, ventral view. 523, female subgenital plate and vulval margin, ventral view." httpUri="https://zenodo.org/record/883438/files/figure.png" pageId="310" pageNumber="311">Fig. 522</figureCitation>
). Mesosomal lobes large, quadratic, intensely papillate and rugose on dorsal surface and median, distal, and lateral margins; 2
<emphasis id="39FCEAEDFEBCFEF5FDAB6556FD4BF8CE" box="[584,642,1878,1901]" italics="true" pageId="310" pageNumber="311">ames</emphasis>
sensilla on each side lateral to gonopore; 2
<emphasis id="39FCEAEDFEBCFEF5FB9C6556FB71F8CE" box="[1151,1208,1878,1901]" italics="true" pageId="310" pageNumber="311">pmes</emphasis>
microsetae on each side marginal on postero-lateral corner of rugose area on lobes, where the mesosome is fused to the parameres;
<emphasis id="39FCEAEDFEBCFEF5FF74659EFF18F816" box="[151,209,1950,1973]" italics="true" pageId="310" pageNumber="311">pmes</emphasis>
often hard to see due to papillation. Parameral heads (
<figureCitation id="93B32A7AFEBCFEF5FCD1659DFC5BF815" box="[818,914,1949,1974]" captionStart="FIGURES 520 523" captionStartId="312.[151,264,1947,1969]" captionTargetBox="[194,1343,198,1893]" captionTargetId="figure@312.[163,1407,193,1902]" captionTargetPageId="312" captionText="FIGURES 520 523 Couala angulata (Piaget, 1880) n. comb. ex Coua caerulea: 520, male head, dorsal and ventral views. 521, male genitalia, dorsal view. 522, male mesosome and parameres, ventral view. 523, female subgenital plate and vulval margin, ventral view." httpUri="https://zenodo.org/record/883438/files/figure.png" pageId="310" pageNumber="311">Fig. 522</figureCitation>
) fused to postero-lateral mesosomal lobes, but lateral ends of heads not fused, angular. Parameral blades strongly curved, convergent;
<emphasis id="39FCEAEDFEBCFEF5FB5A65C2FACDF879" box="[1209,1284,1986,2010]" italics="true" pageId="310" pageNumber="311">pst12</emphasis>
as in genus description. Measurements ex
<taxonomicName id="CC884D7CFEBDFEF4FE086298FD59FF13" baseAuthorityName="Linnaeus" baseAuthorityYear="1766" box="[491,656,151,176]" class="Aves" family="Cuculidae" genus="Coua" kingdom="Animalia" order="Cuculiformes" pageId="311" pageNumber="312" phylum="Chordata" rank="species" species="caerulea">
<emphasis id="39FCEAEDFEBDFEF4FE086298FD59FF13" box="[491,656,151,176]" italics="true" pageId="311" pageNumber="312">Coua caerulea</emphasis>
</taxonomicName>
(n = 22): TL = 1.171.30 (1.24); HL = 0.350.39 (0.37); HW = 0.42 0.49 (0.45); PRW = 0.240.27 (0.25); PTW = 0.370.41 (0.39); AW = 0.490.61 (0.54).
</paragraph>
<paragraph id="0B3736FFFEBDFEF4FF2462DFFE3CFE0E" blockId="311.[151,1436,151,429]" pageId="311" pageNumber="312">
<emphasis id="39FCEAEDFEBDFEF4FF2462DFFED7FF5B" bold="true" box="[199,286,223,248]" italics="true" pageId="311" pageNumber="312">Female</emphasis>
. Abdominal chaetotaxy as in
<tableCitation id="460A0344FEBDFEF4FD8C62DFFD0BFF5B" box="[623,706,223,248]" captionStart="TABLE 2" captionStartId="30.[151,239,796,819]" captionText="TABLE 2. Summary of abdominal chaetotaxy of segments II VIII for the Brueelia - complex, including both genera and subgenera. In genera where chaetotaxy differs between species group (marked with *), only that of the type species is given here, and more detailed accounts are given separately under the respective genus entries and in Tables 3 12. Trichoid setae of segment VIII are present in all genera, and are not listed. Abbreviations: ps = paratergal seta; aps = accessory post-spiracular seta; psps = principal post-spiracular seta; tps = tergal posterior seta; ss = sutural seta; sts = sternal seta." pageId="311" pageNumber="312">Table 2</tableCitation>
and
<figureCitation id="93B32A7AFEBDFEF4FD1A62DFFC93FF5B" box="[761,858,223,248]" captionStart="FIGURE 518 519" captionStartId="311.[151,250,1889,1911]" captionTargetBox="[166,1436,494,1842]" captionTargetId="figure@311.[151,1436,481,1867]" captionTargetPageId="311" captionText="FIGURE 518 519 Couala angulata (Piaget, 1880) n. comb. ex Coua caerulea: 518, male habitus, dorsal and ventral views. 519, female habitus, dorsal and ventral views." httpUri="https://zenodo.org/record/883436/files/figure.png" pageId="311" pageNumber="312">Fig. 519</figureCitation>
. Subgenital plate broadly rounded trapezoidal (
<figureCitation id="93B32A7AFEBDFEF4FA8C62DFFF0DFEBE" captionStart="FIGURES 520 523" captionStartId="312.[151,264,1947,1969]" captionTargetBox="[194,1343,198,1893]" captionTargetId="figure@312.[163,1407,193,1902]" captionTargetPageId="312" captionText="FIGURES 520 523 Couala angulata (Piaget, 1880) n. comb. ex Coua caerulea: 520, male head, dorsal and ventral views. 521, male genitalia, dorsal view. 522, male mesosome and parameres, ventral view. 523, female subgenital plate and vulval margin, ventral view." httpUri="https://zenodo.org/record/883438/files/figure.png" pageId="311" pageNumber="312">Fig. 523</figureCitation>
), not approaching vulval margin. Vulval margin (
<figureCitation id="93B32A7AFEBDFEF4FD086304FC83FEBE" box="[747,842,260,285]" captionStart="FIGURES 520 523" captionStartId="312.[151,264,1947,1969]" captionTargetBox="[194,1343,198,1893]" captionTargetId="figure@312.[163,1407,193,1902]" captionTargetPageId="312" captionText="FIGURES 520 523 Couala angulata (Piaget, 1880) n. comb. ex Coua caerulea: 520, male head, dorsal and ventral views. 521, male genitalia, dorsal view. 522, male mesosome and parameres, ventral view. 523, female subgenital plate and vulval margin, ventral view." httpUri="https://zenodo.org/record/883438/files/figure.png" pageId="311" pageNumber="312">Fig. 523</figureCitation>
) with broad, rounded rectangular bulge medianly; 5 8 large, spine-like
<emphasis id="39FCEAEDFEBDFEF4FE866329FE58FEE3" box="[357,401,297,320]" italics="true" pageId="311" pageNumber="312">vms</emphasis>
on each side, and 35 minute
<emphasis id="39FCEAEDFEBDFEF4FD016329FCCDFEE3" box="[738,772,297,320]" italics="true" pageId="311" pageNumber="312">vss</emphasis>
on each side, restricted to median section of vulval margin; 45 long, slender
<emphasis id="39FCEAEDFEBDFEF4FE87634DFE40FEC7" box="[356,393,333,356]" italics="true" pageId="311" pageNumber="312">vos</emphasis>
; 1
<emphasis id="39FCEAEDFEBDFEF4FE4C634DFE1DFEC7" box="[431,468,333,356]" italics="true" pageId="311" pageNumber="312">vos</emphasis>
median to
<emphasis id="39FCEAEDFEBDFEF4FDB5634DFD4BFEC7" box="[598,642,333,356]" italics="true" pageId="311" pageNumber="312">vms</emphasis>
, near vulval margin. Measurements ex
<taxonomicName id="CC884D7CFEBDFEF4FBA6634DFB25FEC6" baseAuthorityName="Linnaeus" baseAuthorityYear="1766" box="[1093,1260,332,357]" class="Aves" family="Cuculidae" genus="Coua" kingdom="Animalia" order="Cuculiformes" pageId="311" pageNumber="312" phylum="Chordata" rank="species" species="caerulea">
<emphasis id="39FCEAEDFEBDFEF4FBA6634DFB25FEC6" box="[1093,1260,332,357]" italics="true" pageId="311" pageNumber="312">Coua caerulea</emphasis>
</taxonomicName>
(n = 25): TL = 1.381.63 (1.50); HL = 0.400.44 (0.42); HW = 0.480.55 (0.51); PRW = 0.270.29 (0.28); PTW = 0.390.45 (0.41); AW = 0.520.68 (0.61).
</paragraph>
</subSubSection>
<caption id="5FF76677FEBDFEF4FF746561FDA1F836" httpUri="https://zenodo.org/record/883436/files/figure.png" pageId="311" pageNumber="312" targetBox="[166,1436,494,1842]" targetPageId="311">
<paragraph id="0B3736FFFEBDFEF4FF746561FDA1F836" blockId="311.[151,1437,1889,1941]" pageId="311" pageNumber="312">
<emphasis id="39FCEAEDFEBDFEF4FF746561FE93F8D4" bold="true" box="[151,346,1889,1911]" pageId="311" pageNumber="312">FIGURE 518519</emphasis>
<taxonomicName id="CC884D7CFEBDFEF4FE826561FD6CF8D4" authority="(Piaget, 1880)" authorityName="Bush" authorityYear="2017" baseAuthorityName="Piaget" baseAuthorityYear="1880" box="[353,677,1889,1911]" class="Insecta" family="Philopteridae" genus="Couala" kingdom="Animalia" order="Phthiraptera" pageId="311" pageNumber="312" phylum="Arthropoda" rank="species" species="angulata" status="comb. nov.">
<emphasis id="39FCEAEDFEBDFEF4FE826561FDC4F8D4" box="[353,525,1889,1911]" italics="true" pageId="311" pageNumber="312">Couala angulata</emphasis>
(Piaget, 1880)
</taxonomicName>
<taxonomicNameLabel id="22CF5796FEBDFEF4FD4E6561FCC2F8D4" box="[685,779,1889,1911]" pageId="311" pageNumber="312">
<emphasis id="39FCEAEDFEBDFEF4FD4E6561FCC2F8D4" bold="true" box="[685,779,1889,1911]" pageId="311" pageNumber="312">n. comb.</emphasis>
</taxonomicNameLabel>
ex
<taxonomicName id="CC884D7CFEBDFEF4FCD16562FC00F8D4" baseAuthorityName="Linnaeus" baseAuthorityYear="1766" box="[818,969,1889,1911]" class="Aves" family="Cuculidae" genus="Coua" kingdom="Animalia" order="Cuculiformes" pageId="311" pageNumber="312" phylum="Chordata" rank="species" species="caerulea">
<emphasis id="39FCEAEDFEBDFEF4FCD16562FC00F8D4" box="[818,969,1889,1911]" italics="true" pageId="311" pageNumber="312">Coua caerulea</emphasis>
</taxonomicName>
:
<emphasis id="39FCEAEDFEBDFEF4FC346561FBCAF8D4" bold="true" box="[983,1027,1889,1911]" pageId="311" pageNumber="312">518,</emphasis>
male habitus, dorsal and ventral views.
<emphasis id="39FCEAEDFEBDFEF4FF74657FFF0AF836" bold="true" box="[151,195,1919,1941]" pageId="311" pageNumber="312">519,</emphasis>
female habitus, dorsal and ventral views.
</paragraph>
</caption>
<caption id="5FF76677FEB2FEFBFF74659BFEA7F84D" httpUri="https://zenodo.org/record/883438/files/figure.png" pageId="312" pageNumber="313" targetBox="[194,1343,198,1893]" targetPageId="312">
<paragraph id="0B3736FFFEB2FEFBFF74659BFEA7F84D" blockId="312.[151,1437,1947,2030]" pageId="312" pageNumber="313">
<emphasis id="39FCEAEDFEB2FEFBFF74659BFEA1F812" bold="true" box="[151,360,1947,1969]" pageId="312" pageNumber="313">FIGURES 520523</emphasis>
<taxonomicName id="CC884D7CFEB2FEFBFE93659BFD7EF812" authority="(Piaget, 1880)" authorityName="Bush" authorityYear="2017" baseAuthorityName="Piaget" baseAuthorityYear="1880" box="[368,695,1947,1969]" class="Insecta" family="Philopteridae" genus="Couala" kingdom="Animalia" order="Phthiraptera" pageId="312" pageNumber="313" phylum="Arthropoda" rank="species" species="angulata" status="comb. nov.">
<emphasis id="39FCEAEDFEB2FEFBFE93659BFE72F812" box="[368,443,1947,1969]" italics="true" pageId="312" pageNumber="313">Couala</emphasis>
<emphasis id="39FCEAEDFEB2FEFBFE20659BFDD4F812" box="[451,541,1947,1969]" italics="true" pageId="312" pageNumber="313">angulata</emphasis>
(Piaget, 1880)
</taxonomicName>
<taxonomicNameLabel id="22CF5796FEB2FEFBFD5C659BFCD7F812" box="[703,798,1947,1969]" pageId="312" pageNumber="313">
<emphasis id="39FCEAEDFEB2FEFBFD5C659BFCD7F812" bold="true" box="[703,798,1947,1969]" pageId="312" pageNumber="313">n. comb.</emphasis>
</taxonomicNameLabel>
ex
<taxonomicName id="CC884D7CFEB2FEFBFCA6659CFC14F812" baseAuthorityName="Linnaeus" baseAuthorityYear="1766" box="[837,989,1947,1969]" class="Aves" family="Cuculidae" genus="Coua" kingdom="Animalia" order="Cuculiformes" pageId="312" pageNumber="313" phylum="Chordata" rank="species" species="caerulea">
<emphasis id="39FCEAEDFEB2FEFBFCA6659CFC14F812" box="[837,989,1947,1969]" italics="true" pageId="312" pageNumber="313">Coua caerulea</emphasis>
</taxonomicName>
:
<emphasis id="39FCEAEDFEB2FEFBFC0F659BFBD0F812" bold="true" box="[1004,1049,1947,1969]" pageId="312" pageNumber="313">520,</emphasis>
male head, dorsal and ventral views.
<emphasis id="39FCEAEDFEB2FEFBFF7465B9FF0AF86C" bold="true" box="[151,195,1977,1999]" pageId="312" pageNumber="313">521,</emphasis>
male genitalia, dorsal view.
<emphasis id="39FCEAEDFEB2FEFBFE1365B9FDD5F86C" bold="true" box="[496,540,1977,1999]" pageId="312" pageNumber="313">522,</emphasis>
male mesosome and parameres, ventral view.
<emphasis id="39FCEAEDFEB2FEFBFBE065B9FBE6F86C" bold="true" box="[1027,1071,1977,1999]" pageId="312" pageNumber="313">523,</emphasis>
female subgenital plate and vulval margin, ventral view.
</paragraph>
</caption>
<subSubSection id="43926574FEB3FEFAFF246297FB81FF5B" pageId="313" pageNumber="314" type="materials_examined">
<paragraph id="0B3736FFFEB3FEFAFF246297FBE6FF13" blockId="313.[151,1436,151,429]" box="[199,1071,151,176]" pageId="313" pageNumber="314">
<emphasis id="39FCEAEDFEB3FEFAFF246297FEBCFF13" bold="true" box="[199,373,151,176]" pageId="313" pageNumber="314">Type material.</emphasis>
Ex
<emphasis id="39FCEAEDFEB3FEFAFE416297FD40FF13" bold="true" box="[418,649,151,176]" pageId="313" pageNumber="314">
<taxonomicName id="CC884D7CFEB3FEFAFE416297FDC5FF0C" baseAuthorityName="Linnaeus" baseAuthorityYear="1758" box="[418,524,151,176]" class="Aves" family="Strigidae" genus="Asio" kingdom="Animalia" order="Strigiformes" pageId="313" pageNumber="314" phylum="Chordata" rank="species" species="otus">
<emphasis id="39FCEAEDFEB3FEFAFE416297FDC5FF0C" bold="true" box="[418,524,151,176]" italics="true" pageId="313" pageNumber="314">Asio otus</emphasis>
</taxonomicName>
: Holotype
</emphasis>
♂, Piaget Collection, 1013 (NHML).
</paragraph>
<paragraph id="0B3736FFFEB3FEFAFF2462BCFD72FF76" blockId="313.[151,1436,151,429]" box="[199,699,188,213]" pageId="313" pageNumber="314">
<emphasis id="39FCEAEDFEB3FEFAFF2462BCFD72FF76" bold="true" box="[199,699,188,213]" pageId="313" pageNumber="314">Additional material examined (non-types)</emphasis>
</paragraph>
<paragraph id="0B3736FFFEB3FEFAFF2462E0FB81FF5B" blockId="313.[151,1436,151,429]" box="[199,1096,223,248]" pageId="313" pageNumber="314">
<materialsCitation id="BBE03CA2FEB3FEFAFF2462E0FB81FF5B" ID-GBIF-Occurrence="1587279368" box="[199,1096,223,248]" collectionCode="NHML" collectorName="R. Meinertzhagen" country="Madagascar" location="Madagascar" pageId="313" pageNumber="314" specimenCount="75" specimenCount-female="38" specimenCount-male="37">
Ex
<emphasis id="39FCEAEDFEB3FEFAFF0E62DFFE57FF5B" bold="true" box="[237,414,223,248]" pageId="313" pageNumber="314">
<taxonomicName id="CC884D7CFEB3FEFAFF0E62DFFE5CFF5B" baseAuthorityName="Linnaeus" baseAuthorityYear="1766" box="[237,405,223,248]" class="Aves" family="Cuculidae" genus="Coua" kingdom="Animalia" order="Cuculiformes" pageId="313" pageNumber="314" phylum="Chordata" rank="species" species="caerulea">
<emphasis id="39FCEAEDFEB3FEFAFF0E62DFFE5CFF5B" bold="true" box="[237,405,223,248]" italics="true" pageId="313" pageNumber="314">Coua caerulea</emphasis>
</taxonomicName>
:
</emphasis>
<specimenCount id="1D8EFD76FEB3FEFAFE4562E0FE10FF5B" box="[422,473,224,248]" pageId="313" pageNumber="314" type="male">37♂</specimenCount>
,
<specimenCount id="1D8EFD76FEB3FEFAFE0662E0FDDEFF5B" box="[485,535,224,248]" pageId="313" pageNumber="314" type="female">38♀</specimenCount>
,
<collectingCountry id="739F766FFEB3FEFAFDC762DFFD64FF5B" box="[548,685,223,248]" name="Madagascar" pageId="313" pageNumber="314">Madagascar</collectingCountry>
,
<collectorName id="A67D5329FEB3FEFAFD5B62E0FC4DFF5B" box="[696,900,223,248]" pageId="313" pageNumber="314">R. Meinertzhagen</collectorName>
, 16648 (
<collectionCode id="6D99AE3AFEB3FEFAFC0762E0FBF4FF5B" box="[996,1085,224,248]" country="Libya" httpUri="http://grbio.org/cool/49nc-28y0" name="Natural History Museum, Tripoli" pageId="313" pageNumber="314">NHML</collectionCode>
).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="43926574FEB3FEFAFF246304FADEFE0E" pageId="313" pageNumber="314" type="discussion">
<paragraph id="0B3736FFFEB3FEFAFF246304FADEFE0E" blockId="313.[151,1436,151,429]" pageId="313" pageNumber="314">
<emphasis id="39FCEAEDFEB3FEFAFF246304FEF2FEBE" bold="true" box="[199,315,260,285]" pageId="313" pageNumber="314">Remarks.</emphasis>
The
<typeStatus id="D433885DFEB3FEFAFE9B6305FE61FEBE" box="[376,424,261,285]" pageId="313" pageNumber="314">type</typeStatus>
host was given by
<bibRefCitation id="6F194B0EFEB3FEFAFD6A6304FC94FEBE" author="Piaget" box="[649,861,260,285]" pageId="313" pageNumber="314" refString="Piaget, E. (1880) Les Pediculines. Essai Monographique. Two volumes. E. J. Brill, Leide. xxxix + 714 pp., 56 pls." type="book" year="1880">Piaget (1880: 135)</bibRefCitation>
as
<taxonomicName id="CC884D7CFEB3FEFAFC646305FBECFEBE" baseAuthorityName="Linnaeus" baseAuthorityYear="1758" box="[903,1061,260,285]" class="Aves" family="Strigidae" genus="Strix" kingdom="Animalia" order="Strigiformes" pageId="313" pageNumber="314" phylum="Chordata" rank="species" species="flammea">
<emphasis id="39FCEAEDFEB3FEFAFC646305FBECFEBE" box="[903,1061,260,285]" italics="true" pageId="313" pageNumber="314">Strix flammea</emphasis>
</taxonomicName>
[=
<taxonomicName id="CC884D7CFEB3FEFAFBAC6304FF13FEE3" authority="(Pontoppidan, 1763)" baseAuthorityName="Pontoppidan" baseAuthorityYear="1763" class="Aves" family="Strigidae" genus="Asio" kingdom="Animalia" order="Strigiformes" pageId="313" pageNumber="314" phylum="Chordata" rank="species" species="flammeus">
<emphasis id="39FCEAEDFEB3FEFAFBAC6304FB3CFEBE" box="[1103,1269,260,285]" italics="true" pageId="313" pageNumber="314">Asio flammeus</emphasis>
(Pontoppidan, 1763)
</taxonomicName>
] from the Zoological Garden in Rotterdam, but
<taxonomicName id="CC884D7CFEB3FEFAFCF36328FC71FEE3" baseAuthorityName="Pontoppidan" baseAuthorityYear="1763" box="[784,952,295,320]" class="Aves" family="Strigidae" genus="Asio" kingdom="Animalia" order="Strigiformes" pageId="313" pageNumber="314" phylum="Chordata" rank="species" species="flammeus">
<emphasis id="39FCEAEDFEB3FEFAFCF36328FC71FEE3" box="[784,952,295,320]" italics="true" pageId="313" pageNumber="314">Asio flammeus</emphasis>
</taxonomicName>
does not occur in
<collectingCountry id="739F766FFEB3FEFAFB756327FAD6FEE3" box="[1174,1311,295,320]" name="Madagascar" pageId="313" pageNumber="314">Madagascar</collectingCountry>
, only
<taxonomicName id="CC884D7CFEB3FEFAFA896328FDE3FEC7" authority="(A. Smith, 1834)" baseAuthorityName="A. Smith" baseAuthorityYear="1834" class="Aves" family="Strigidae" genus="Asio" kingdom="Animalia" order="Strigiformes" pageId="313" pageNumber="314" phylum="Chordata" rank="species" species="madagascariensis">
<emphasis id="39FCEAEDFEB3FEFAFA896328FEA8FEC6" italics="true" pageId="313" pageNumber="314">Asio madagascariensis</emphasis>
(
<collectingRegion id="C94CF81DFEB3FEFAFE92634CFE40FEC7" box="[369,393,332,356]" country="Nigeria" name="Rivers" pageId="313" pageNumber="314">A</collectingRegion>
. Smith, 1834)
</taxonomicName>
. The
<typeStatus id="D433885DFEB3FEFAFD8B634CFD03FEC6" box="[616,714,332,357]" pageId="313" pageNumber="314" type="holotype">holotype</typeStatus>
of
<taxonomicName id="CC884D7CFEB3FEFAFD12634CFC66FEC6" authorityName="Bush" authorityYear="2017" baseAuthorityName="Piaget" baseAuthorityYear="1880" box="[753,943,332,357]" class="Insecta" family="Philopteridae" genus="Couala" kingdom="Animalia" order="Phthiraptera" pageId="313" pageNumber="314" phylum="Arthropoda" rank="species" species="angulata">
<emphasis id="39FCEAEDFEB3FEFAFD12634CFC66FEC6" box="[753,943,332,357]" italics="true" pageId="313" pageNumber="314">Couala angulata</emphasis>
</taxonomicName>
is identical to material from
<taxonomicName id="CC884D7CFEB3FEFAFB15634DFE90FE2B" authority="(Linnaeus, 1766)" baseAuthorityName="Linnaeus" baseAuthorityYear="1766" class="Aves" family="Cuculidae" genus="Coua" kingdom="Animalia" order="Cuculiformes" pageId="313" pageNumber="314" phylum="Chordata" rank="species" species="caerulea">
<emphasis id="39FCEAEDFEB3FEFAFB15634DFA55FEC6" box="[1270,1436,332,357]" italics="true" pageId="313" pageNumber="314">Coua caerulea</emphasis>
(Linnaeus, 1766)
</taxonomicName>
. Considering that all the hosts of species of
<taxonomicName id="CC884D7CFEB3FEFAFCA4636FFC53FE2B" authority="Bush, 2017" authorityName="Bush" authorityYear="2017" box="[839,922,367,392]" class="Insecta" family="Philopteridae" genus="Couala" kingdom="Animalia" order="Phthiraptera" pageId="313" pageNumber="314" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="39FCEAEDFEB3FEFAFCA4636FFC53FE2B" box="[839,922,367,392]" italics="true" pageId="313" pageNumber="314">Couala</emphasis>
</taxonomicName>
<taxonomicNameLabel id="22CF5796FEB3FEFAFC42636FFC39FE2B" box="[929,1008,367,392]" pageId="313" pageNumber="314" rank="genus">
<emphasis id="39FCEAEDFEB3FEFAFC42636FFC39FE2B" bold="true" box="[929,1008,367,392]" pageId="313" pageNumber="314">n. gen.</emphasis>
</taxonomicNameLabel>
are from
<collectingCountry id="739F766FFEB3FEFAFBBD636FFB2EFE2B" box="[1118,1255,367,392]" name="Madagascar" pageId="313" pageNumber="314">Madagascar</collectingCountry>
, we believe that either Piaget was mistaken about the host, or that contamination had occurred at the zoological garden.
</paragraph>
</subSubSection>
</treatment>
</document>