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<document ID-DOI="10.25221/fee.448.1" ID-ISSN="2713-2196" ID-Zenodo-Dep="7166846" approvalRequired="57" approvalRequired_for_textStreams="57" checkinTime="1665256722461" checkinUser="felipe" docAuthor="Khachikov, E. A., Háva, J. &amp; Poushkova, S. V." docDate="2022" docId="03F88337FFF3FFBAFF2F7BD6493EF993" docLanguage="en" docName="FarEastEntomol.448.1-10.pdf" docOrigin="Far Eastern Entomologist 448" docSource="http://dx.doi.org/10.25221/fee.448.1" docStyle="DocumentStyle:68641B8F4A21450A5F9A74164B730F50.1:FarEastEntomol.2017-.journal_article" docStyleId="68641B8F4A21450A5F9A74164B730F50" docStyleName="FarEastEntomol.2017-.journal_article" docStyleVersion="1" docTitle="Phradonoma beskalensis Khachikov, Hava et Poushkova 2022, sp. n." docType="treatment" docVersion="2" lastPageNumber="6" masterDocId="FFC1FB4FFFF7FFBFFFCC792B4A1CFFD2" masterDocTitle="New records of the skin beetles (Coleoptera: Dermestidae) from Russia, Central Asia and Caucasus, with description of a new species from Kazakhstan" masterLastPageNumber="10" masterPageNumber="1" pageNumber="5" updateTime="1665271042267" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>New records of the skin beetles (Coleoptera: Dermestidae) from Russia, Central Asia and Caucasus, with description of a new species from Kazakhstan</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Khachikov, E. A.</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Háva, J.</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Poushkova, S. V.</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>Far Eastern Entomologist</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2022</mods:date>
<mods:detail type="pubDate">
<mods:number>2022-02-01</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>448</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>1</mods:start>
<mods:end>10</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location>
<mods:url>http://dx.doi.org/10.25221/fee.448.1</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">10.25221/fee.448.1</mods:identifier>
<mods:identifier type="ISSN">2713-2196</mods:identifier>
<mods:identifier type="Zenodo-Dep">7166846</mods:identifier>
</mods:mods>
<treatment LSID="urn:lsid:plazi:treatment:03F88337FFF3FFBAFF2F7BD6493EF993" httpUri="http://treatment.plazi.org/id/03F88337FFF3FFBAFF2F7BD6493EF993" lastPageId="5" lastPageNumber="6" pageId="4" pageNumber="5">
<subSubSection pageId="4" pageNumber="5" type="nomenclature">
<paragraph blockId="4.[227,1081,764,821]" box="[227,939,764,789]" pageId="4" pageNumber="5">
<heading box="[227,939,764,789]" fontSize="9" level="2" pageId="4" pageNumber="5" reason="2">
<emphasis bold="true" box="[227,939,764,789]" pageId="4" pageNumber="5">
<taxonomicName authority="Khachikov, Hava et Poushkova" authorityName="Khachikov, Hava et Poushkova" authorityYear="2022" box="[227,867,764,789]" class="Insecta" family="Dermestidae" genus="Phradonoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="beskalensis" status="sp. n.">
<emphasis bold="true" box="[227,501,765,789]" italics="true" pageId="4" pageNumber="5">Phradonoma beskalensis</emphasis>
Khachikov, Háva et Poushkova
</taxonomicName>
,
<taxonomicNameLabel box="[879,939,765,789]" pageId="4" pageNumber="5" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph blockId="4.[227,1081,764,821]" box="[227,1081,799,821]" pageId="4" pageNumber="5">
http://zoobank.org/NomenclaturalActs/
<uuid box="[616,1081,799,821]" pageId="4" pageNumber="5">5EC87AAA-82E9-4F8A-8FA1-2B92E35CCC9B</uuid>
</paragraph>
</subSubSection>
<subSubSection box="[227,440,832,857]" pageId="4" pageNumber="5" type="description">
<paragraph blockId="4.[227,440,832,857]" box="[227,440,832,857]" pageId="4" pageNumber="5">
<figureCitation box="[227,361,832,857]" captionStart="Figs 1–12" captionStartId="2.[265,306,1472,1494]" captionTargetBox="[274,1079,415,1454]" captionTargetId="figure-46@2.[257,1096,407,1457]" captionTargetPageId="2" captionText="Figs 1–12. Antennae and male genitalia of Phradonoma spp. 1–3, 7, 9, 10 – Ph. villosulum:" figureDoi="http://doi.org/10.5281/zenodo.7166850" httpUri="https://zenodo.org/record/7166850/files/figure.png" pageId="4" pageNumber="5">Figs 5, 6, 11</figureCitation>
,
<figureCitation box="[373,440,833,857]" captionStart="Figs 19–21" captionStartId="5.[265,306,1552,1574]" captionTargetBox="[260,1084,647,1500]" captionTargetId="figure-122@5.[227,1111,625,1519]" captionTargetPageId="5" captionText="Figs 19–21. Phradonoma spp. 19, 20 – Ph. beskalensis sp. n.: 19 – dorsal aspect of body;" figureDoi="http://doi.org/10.5281/zenodo.7166854" httpUri="https://zenodo.org/record/7166854/files/figure.png" pageId="4" pageNumber="5">19, 20</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="4" pageNumber="5" type="materials_examined">
<paragraph blockId="4.[227,1127,893,1577]" box="[265,1126,893,917]" pageId="4" pageNumber="5">
<materialsCitation box="[265,1126,893,917]" collectionCode="TYPE, MATERIAL" country="Kazakhstan" location="Guryev Region" pageId="4" pageNumber="5" specimenCount="1" specimenCount-male="1" stateProvince="Atyrau Oblysy" typeStatus="holotype">
<collectionCode box="[265,485,893,917]" pageId="4" pageNumber="5">TYPE MATERIAL</collectionCode>
.
<typeStatus box="[501,600,893,917]" pageId="4" pageNumber="5">Holotype</typeStatus>
–
<specimenCount box="[638,660,893,916]" pageId="4" pageNumber="5" type="male">♂</specimenCount>
,
<emphasis bold="true" box="[677,822,893,917]" pageId="4" pageNumber="5">
<collectingCountry box="[677,817,893,917]" name="Kazakhstan" pageId="4" pageNumber="5">Kazakhstan</collectingCountry>
:
</emphasis>
<location LSID="urn:lsid:plazi:treatment:03F88337FFF3FFBAFF2F7BD6493EF993:8E8E64FAFFF3FFBBFC8F7A5649F0FC47" box="[835,1004,893,917]" country="Kazakhstan" name="Guryev Region" pageId="4" pageNumber="5" stateProvince="Atyrau Oblysy">Guryev Region</location>
[=
<collectingRegion box="[1052,1126,893,917]" country="Kazakhstan" name="Atyrau Oblysy" pageId="4" pageNumber="5">Atyrau</collectingRegion>
</materialsCitation>
</paragraph>
<paragraph blockId="4.[227,1127,893,1577]" pageId="4" pageNumber="5">
Region], Kois tract, Beskala vill. [=Madeniet vill.],
<date box="[790,924,922,947]" pageId="4" pageNumber="5" value="1980-07-05">05.VII 1980</date>
, leg. E.A. Khachikov (
<collectionCode box="[281,331,953,977]" country="Russia" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="4" pageNumber="5" type="Museum">ZIN</collectionCode>
).
</paragraph>
</subSubSection>
<subSubSection pageId="4" pageNumber="5" type="description">
<paragraph blockId="4.[227,1127,893,1577]" pageId="4" pageNumber="5">DESCRIPTION. Body length is 2.1 mm, the top is dark, almost matte, covered with erect reddish setae. Head is transverse, with coarse and rare punctation, with long more or less decumbent yellow-brown setae, maxillary palpi darkish brown,</paragraph>
<paragraph blockId="4.[227,1127,893,1577]" box="[227,1127,1073,1097]" pageId="4" pageNumber="5">eyes large with microsetae. Forehead with ocellus. Antennae 11-segmented with a</paragraph>
<paragraph blockId="4.[227,1127,893,1577]" box="[227,1126,1102,1127]" pageId="4" pageNumber="5">
5-antennomered club (
<figureCitation box="[474,577,1102,1127]" captionStart="Figs 1–12" captionStartId="2.[265,306,1472,1494]" captionTargetBox="[274,1079,415,1454]" captionTargetId="figure-46@2.[257,1096,407,1457]" captionTargetPageId="2" captionText="Figs 1–12. Antennae and male genitalia of Phradonoma spp. 1–3, 7, 9, 10 – Ph. villosulum:" figureDoi="http://doi.org/10.5281/zenodo.7166850" httpUri="https://zenodo.org/record/7166850/files/figure.png" pageId="4" pageNumber="5">Figs 5, 6</figureCitation>
), 11th–9th antennomeres larger than 8th and 7th;
</paragraph>
<paragraph blockId="4.[227,1127,893,1577]" pageId="4" pageNumber="5">the 11th antennomere is apically smoothly narrowed and rounded; 10th antennomere transverse, slightly widened apically and basally rounded; the 9th antennomere</paragraph>
<paragraph blockId="4.[227,1127,893,1577]" box="[227,1127,1193,1217]" pageId="4" pageNumber="5">(main segment of the club) is transverse, in the form of inverted parallelepiped; the</paragraph>
<paragraph blockId="4.[227,1127,893,1577]" box="[227,1127,1223,1247]" pageId="4" pageNumber="5">5th to 8th antennomeres are strongly transverse, slightly rounded; the 1st and 2nd</paragraph>
<paragraph blockId="4.[227,1127,893,1577]" pageId="4" pageNumber="5">
antennomeres are slightly transverse and apically widened. The first antennomere is brown, the club very darkish brown and all other segments are light brown. Pronotum is entirely dark brown, shiny, sparsely and not very coarsely punctate, with yellowbrown setae (
<figureCitation box="[373,454,1343,1367]" captionStart="Figs 19–21" captionStartId="5.[265,306,1552,1574]" captionTargetBox="[260,1084,647,1500]" captionTargetId="figure-122@5.[227,1111,625,1519]" captionTargetPageId="5" captionText="Figs 19–21. Phradonoma spp. 19, 20 – Ph. beskalensis sp. n.: 19 – dorsal aspect of body;" figureDoi="http://doi.org/10.5281/zenodo.7166854" httpUri="https://zenodo.org/record/7166854/files/figure.png" pageId="4" pageNumber="5">Fig. 19</figureCitation>
), apically smoothly narrowed, almost as wide as elytra. Basal margin of pronotum is in the form of an obtuse angle, the apex of which is at its middle. The scutellum is small, dark brown, triangular, without setation or punctu-
</paragraph>
<paragraph blockId="4.[227,1127,893,1577]" pageId="4" pageNumber="5">
res. Elytra dark brown anteriorly, light brown posteriorly, coarsely and sparsely punctate, sparsely covered by yellow-brown setae. Elytra are also gradually tapering towards the posterior margin of the body, and more than 2.5 times as long as the pronotum. The underside of the specimen is finely and more densely punctate than the elytra. Tibiae and tarsi are brown, the femurs darkened. Male genitalia as in
<figureCitation box="[1041,1119,1553,1577]" captionStart="Figs 1–12" captionStartId="2.[265,306,1472,1494]" captionTargetBox="[274,1079,415,1454]" captionTargetId="figure-46@2.[257,1096,407,1457]" captionTargetPageId="2" captionText="Figs 1–12. Antennae and male genitalia of Phradonoma spp. 1–3, 7, 9, 10 – Ph. villosulum:" figureDoi="http://doi.org/10.5281/zenodo.7166850" httpUri="https://zenodo.org/record/7166850/files/figure.png" pageId="4" pageNumber="5">Fig. 11</figureCitation>
.
</paragraph>
</subSubSection>
<subSubSection pageId="5" pageNumber="6" type="diagnosis">
<paragraph blockId="5.[227,1127,264,595]" pageId="5" pageNumber="6">
DIAGNOSIS. The new species is similar to
<taxonomicName baseAuthorityName="Duftschmid" baseAuthorityYear="1825" box="[740,997,264,288]" class="Insecta" family="Dermestidae" genus="Phradonoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="villosulum">
<emphasis box="[740,997,264,288]" italics="true" pageId="5" pageNumber="6">Phradonoma villosulum</emphasis>
</taxonomicName>
but reliably different in the shape of the antennomeres. In
<taxonomicName baseAuthorityName="Duftschmid" baseAuthorityYear="1825" box="[727,868,294,318]" class="Insecta" family="Dermestidae" genus="Phradonoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="villosulum">
<emphasis box="[727,868,294,318]" italics="true" pageId="5" pageNumber="6">P. villosulum</emphasis>
</taxonomicName>
the 9th and 10th antennomeres are much more transverse and widened, while the 8th–6th antennomeres are contrastingly narrower than in
<taxonomicName authority="Khachikov &amp; Háva &amp; Poushkova, 2022" authorityName="Khachikov &amp; Háva &amp; Poushkova" authorityYear="2022" box="[597,750,356,380]" class="Insecta" family="Dermestidae" genus="Phradonoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="beskalensis" status="sp. n.">
<emphasis box="[597,750,356,380]" italics="true" pageId="5" pageNumber="6">P. beskalensis</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[756,814,356,379]" pageId="5" pageNumber="6" rank="species">sp. n.</taxonomicNameLabel>
There are also differences in the structure of the male genitalia, especially in parameres (
<figureCitation box="[858,935,386,411]" captionStart="Figs 1–12" captionStartId="2.[265,306,1472,1494]" captionTargetBox="[274,1079,415,1454]" captionTargetId="figure-46@2.[257,1096,407,1457]" captionTargetPageId="2" captionText="Figs 1–12. Antennae and male genitalia of Phradonoma spp. 1–3, 7, 9, 10 – Ph. villosulum:" figureDoi="http://doi.org/10.5281/zenodo.7166850" httpUri="https://zenodo.org/record/7166850/files/figure.png" pageId="5" pageNumber="6">Fig. 11</figureCitation>
), which are wider in the distal part.
<taxonomicName authorityName="Khachikov &amp; Háva &amp; Poushkova" authorityYear="2022" box="[420,573,417,441]" class="Insecta" family="Dermestidae" genus="Phradonoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="beskalensis">
<emphasis box="[420,573,417,441]" italics="true" pageId="5" pageNumber="6">P. beskalensis</emphasis>
</taxonomicName>
also differs from other species of the “
<taxonomicName baseAuthorityName="Duftschmid" baseAuthorityYear="1825" box="[1004,1115,417,441]" class="Insecta" family="Dermestidae" genus="Phradonoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="villosulum">
<emphasis box="[1004,1115,417,441]" italics="true" pageId="5" pageNumber="6">villosulum</emphasis>
</taxonomicName>
”
</paragraph>
<paragraph blockId="5.[227,1127,264,595]" box="[227,1127,448,472]" pageId="5" pageNumber="6">
group of the genus
<taxonomicName authorityName="Jacquelin du Val" authorityYear="1859" box="[434,573,448,472]" class="Insecta" family="Dermestidae" genus="Phradonoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">
<emphasis box="[434,573,448,472]" italics="true" pageId="5" pageNumber="6">Phradonoma</emphasis>
</taxonomicName>
in the structure of antennomeres and male genitalia
</paragraph>
<paragraph blockId="5.[227,1127,264,595]" box="[227,375,478,503]" pageId="5" pageNumber="6">(Háva, 2006).</paragraph>
</subSubSection>
<subSubSection box="[265,597,509,533]" pageId="5" pageNumber="6" type="distribution">
<paragraph blockId="5.[227,1127,264,595]" box="[265,597,509,533]" pageId="5" pageNumber="6">
DISTRIBUTION.
<collectingCountry box="[465,593,509,533]" name="Kazakhstan" pageId="5" pageNumber="6">Kazakhstan</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection pageId="5" pageNumber="6" type="etymology">
<paragraph blockId="5.[227,1127,264,595]" box="[265,1127,540,564]" pageId="5" pageNumber="6">ETYMOLOGY. A name of new species derived from Beskala village, the envi-</paragraph>
<paragraph blockId="5.[227,1127,264,595]" box="[227,783,570,595]" pageId="5" pageNumber="6">
rons of which being the
<typeStatus box="[486,532,571,595]" pageId="5" pageNumber="6">type</typeStatus>
locality of this species.
</paragraph>
<caption ID-DOI="http://doi.org/10.5281/zenodo.7166854" ID-Zenodo-Dep="7166854" box="[265,1127,1552,1574]" httpUri="https://zenodo.org/record/7166854/files/figure.png" pageId="5" pageNumber="6" startId="5.[265,306,1552,1574]" targetBox="[260,1084,647,1500]" targetPageId="5">
<paragraph blockId="5.[227,1127,1552,1602]" box="[265,1127,1552,1574]" pageId="5" pageNumber="6">
Figs 19–21.
<taxonomicName box="[385,554,1552,1573]" class="Insecta" family="Dermestidae" genus="Phradonoma" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis box="[385,508,1552,1573]" italics="true" pageId="5" pageNumber="6">Phradonoma</emphasis>
spp.
</taxonomicName>
19, 20 –
<taxonomicName authority="Khachikov &amp; Háva &amp; Poushkova, 2022" authorityName="Khachikov &amp; Háva &amp; Poushkova" authorityYear="2022" box="[642,790,1552,1573]" class="Insecta" family="Dermestidae" genus="Phradonoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="beskalensis" status="sp. n.">
<emphasis box="[642,790,1552,1573]" italics="true" pageId="5" pageNumber="6">Ph. beskalensis</emphasis>
</taxonomicName>
<emphasis bold="true" box="[796,851,1552,1573]" pageId="5" pageNumber="6">
<taxonomicNameLabel box="[796,851,1552,1573]" pageId="5" pageNumber="6" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
: 19 – dorsal aspect of body;
</paragraph>
</caption>
<paragraph blockId="5.[227,1127,1552,1602]" box="[227,802,1580,1602]" pageId="5" pageNumber="6">
20 – VIII tergite; 21 –
<taxonomicName authority=", VIII" authorityName="VIII" box="[444,728,1580,1601]" class="Insecta" family="Dermestidae" genus="Phradonoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="villosulum">
<emphasis box="[444,675,1580,1601]" italics="true" pageId="5" pageNumber="6">Phradonoma villosulum</emphasis>
, VIII
</taxonomicName>
tergite.
</paragraph>
</subSubSection>
</treatment>
</document>
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