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<mods:title id="FABF60D7F1078C02CBF04FBC03A89A0A">A revision of male ants of the Malagasy region (Hymenoptera: Formicidae): Key to genera of the subfamily Dolichoderinae</mods:title>
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<treatment id="03F81D2F900AFFB8FF0BBD06C65267F8" ID-DOI="http://doi.org/10.5281/zenodo.5689431" ID-GBIF-Taxon="119393159" ID-Zenodo-Dep="5689431" LSID="urn:lsid:plazi:treatment:03F81D2F900AFFB8FF0BBD06C65267F8" httpUri="http://treatment.plazi.org/id/03F81D2F900AFFB8FF0BBD06C65267F8" lastPageId="14" lastPageNumber="15" pageId="13" pageNumber="14">
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<taxonomicName id="4C51D7BA900AFFBBFF0BBD06C3926551" ID-CoL="7T8H" authority="Mayr, 1872" authorityName="Mayr" authorityYear="1872" box="[151,484,835,862]" class="Insecta" family="Formicidae" genus="Technomyrmex" kingdom="Animalia" order="Hymenoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="genus">
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<emphasis id="B925702B900AFFBBFF0BBD06C3276552" bold="true" box="[151,337,836,862]" italics="true" pageId="13" pageNumber="14">Technomyrmex</emphasis>
Mayr, 1872
</emphasis>
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<paragraph id="8BEEAC39900AFFBBFF0BBD24C14D6572" blockId="13.[151,827,835,894]" box="[151,827,870,894]" pageId="13" pageNumber="14">
(
<figureCitation id="136AB0BC900AFFBBFF03BD24C2916572" box="[159,231,870,894]" captionStart="FIGURES 1 5" captionStartId="16.[151,264,1953,1976]" captionTargetBox="[202,1381,195,1911]" captionTargetId="figure@16.[202,1382,195,1927]" captionTargetPageId="16" captionText="FIGURES 1 5. Head of dolichoderine males in full-face view. 1, Ochetellus glaber (CASENT 0191065); 2, Aptinoma mangabe (CASENT 0173594); 3, Ravavy miafina (CASENT 0474633); 4, Tapinoma subtile (CASENT 0136850); 5, Technomyrmex difficilis (CASENT 0049968)." httpUri="https://zenodo.org/record/276994/files/figure.png" pageId="13" pageNumber="14">Figs 5</figureCitation>
,
<figureCitation id="136AB0BC900AFFBBFF6FBD24C3676572" box="[243,273,870,894]" captionStart="FIGURES 6 10" captionStartId="17.[151,264,1954,1977]" captionTargetBox="[199,1377,193,1913]" captionTargetId="figure@17.[199,1378,193,1927]" captionTargetPageId="17" captionText="FIGURES 6 10. Head to abdomen of dolichoderine males in lateral view. 6, Ravavy miafina (CASENT 0474633); 7, Aptinoma mangabe (CASENT 0173594); 8, Ochetellus glaber (CASENT 0191065); 9, Tapinoma subtile (CASENT 0136850); 10, Technomyrmex difficilis (CASENT 0049968)." httpUri="https://zenodo.org/record/276995/files/figure.png" pageId="13" pageNumber="14">10</figureCitation>
,
<figureCitation id="136AB0BC900AFFBBFE81BD24C34D6572" box="[285,315,870,894]" captionStart="FIGURES 11 15" captionStartId="18.[151,264,1929,1952]" captionTargetBox="[187,1384,193,1883]" captionTargetId="figure@18.[187,1385,193,1900]" captionTargetPageId="18" captionText="FIGURES 11 15. Head to abdomen of dolichoderine males in dorsal view. 11, Aptinoma mangabe (CASENT 0173594); 12, Ochetellus glaber (CASENT 0191065); 13, Ravavy miafina (CASENT 0474633); 14, Tapinoma subtile (CASENT 0136850); 15, Technomyrmex difficilis (CASENT 0049968)." httpUri="https://zenodo.org/record/276996/files/figure.png" pageId="13" pageNumber="14">15</figureCitation>
,
<figureCitation id="136AB0BC900AFFBBFEDBBD24C3136572" box="[327,357,870,894]" captionStart="FIGURES 16 23" captionStartId="19.[151,264,1847,1870]" captionTargetBox="[237,1345,193,1821]" captionTargetId="figure@19.[237,1346,193,1822]" captionTargetPageId="19" captionText="FIGURES 16 23. Males of Dolichoderinae. 16, Aptinoma mangabe (CASENT 0173594); 17, Aptinoma mangabe (CASENT 0175002); 18, 20, Ochetellus glaber (CASENT 0179489); 19, 21, Ravavy miafina (CASENT 0080308); 22, Technomyrmex curiosus (paratype: CASENT 0070364); 23, Tapinoma mg 10 (CASENT 0115650). 16, distal portion of abdomen in oblique posterior view; 17, petiole in lateral view; 18, 19, petiole and abdominal segments in ventral view; 20, petiole and abdominal segments in dorsal view; 21, petiole and abdominal segments in oblique frontal view; 22, 23, genitalia in ventral view." httpUri="https://zenodo.org/record/276997/files/figure.png" pageId="13" pageNumber="14">22</figureCitation>
,
<figureCitation id="136AB0BC900AFFBBFEEDBD24C3F96572" box="[369,399,870,894]" captionStart="FIGURES 24 28" captionStartId="20.[151,264,1912,1935]" captionTargetBox="[195,1392,193,1875]" captionTargetId="figure@20.[195,1393,193,1891]" captionTargetPageId="20" captionText="FIGURES 24 28. Abdominal sternum IX of dolichoderine males in ectal view. 24, Aptinoma mangabe (CASENT 0175002); 25, Ochetellus glaber (CASENT 0179489); 26, Ravavy miafina (CASENT 0179530); 27, Tapinoma melanocephalum (CASENT 0492372); 28, Technomyrmex madecassus (CASENT 0179526)." httpUri="https://zenodo.org/record/276998/files/figure.png" pageId="13" pageNumber="14">28</figureCitation>
,
<figureCitation id="136AB0BC900AFFBBFE07BD24C3CF6572" box="[411,441,870,894]" captionStart="FIGURES 29 33" captionStartId="21.[151,264,1944,1967]" captionTargetBox="[204,1379,193,1905]" captionTargetId="figure@21.[204,1380,193,1921]" captionTargetPageId="21" captionText="FIGURES 29 33. Genital capsules of dolichoderine males. 29, Tapinoma mg 11 (CASENT 0179494); 30, Aptinoma mangabe (CASENT 0175002); 31, Ochetellus glaber (CASENT 0179489); 32, Ravavy miafina (CASENT 0179533); 33, Technomyrmex madecassus (CASENT 0179526). Right side, ventral view; left side, dorsal view." httpUri="https://zenodo.org/record/276999/files/figure.png" pageId="13" pageNumber="14">33</figureCitation>
,
<figureCitation id="136AB0BC900AFFBBFE59BD24C3956572" box="[453,483,870,894]" captionStart="FIGURES 34 38" captionStartId="22.[151,264,1945,1968]" captionTargetBox="[213,1384,193,1897]" captionTargetId="figure@22.[207,1386,193,1926]" captionTargetPageId="22" captionText="FIGURES 34 38. Genital capsules of dolichoderine males in lateral view. 34, Ochetellus glaber (CASENT 0179489); 35, Aptinoma mangabe (CASENT 0175002); 36, Ravavy miafina (CASENT 0179533); 37, Tapinoma mg 11 (CASENT 0179494); 38, Technomyrmex madecassus (CASENT 0179526)." httpUri="https://zenodo.org/record/277000/files/figure.png" pageId="13" pageNumber="14">38</figureCitation>
,
<figureCitation id="136AB0BC900AFFBBFE73BD24C07B6572" box="[495,525,870,894]" captionStart="FIGURES 41 45" captionStartId="24.[151,264,1945,1968]" captionTargetBox="[200,1379,193,1913]" captionTargetId="figure@24.[200,1380,193,1928]" captionTargetPageId="24" captionText="FIGURES 41 45. Aedeagal plates of dolichoderine males in lateral view. 41, Ravavy miafina (CASENT 0179530); 42, Aptinoma mangabe (CASENT 0175002); 43, Ochetellus glaber (CASENT 0179489); 44, Tapinoma mg 11 (CASENT 0179494); 45, Technomyrmex madecassus (CASENT 0179526)." httpUri="https://zenodo.org/record/277002/files/figure.png" pageId="13" pageNumber="14">45</figureCitation>
,
<figureCitation id="136AB0BC900AFFBBFD85BD24C0416572" box="[537,567,870,894]" captionStart="FIGURES 46 48" captionStartId="25.[151,264,1971,1994]" captionTargetBox="[163,1436,193,1909]" captionTargetId="figure@25.[163,1448,193,1940]" captionTargetPageId="25" captionText="FIGURES 46 48. Volsella of dolichoderine males in oblique posterior view. 46, Ochetellus glaber (CASENT 0179489); 47, Tapinoma mg 11 (CASENT 0179494); 48, Technomyrmex madecassus (CASENT 0179526)." httpUri="https://zenodo.org/record/277003/files/figure.png" pageId="13" pageNumber="14">48</figureCitation>
,
<figureCitation id="136AB0BC900AFFBBFDDFBD24C0146572" box="[579,610,870,894]" captionStart="FIGURES 52 54" captionStartId="27.[151,264,1973,1996]" captionTargetBox="[221,1390,193,1923]" captionTargetId="figure@27.[195,1393,193,1952]" captionTargetPageId="27" captionText="FIGURES 52 54. Forewings of male ants. 52, Ravavy miafina (CASENT 0179530); 53, Tapinoma mg 10 (CASENT 0179493); 54, Technomyrmex madecassus (CASENT 0179526)." httpUri="https://zenodo.org/record/277005/files/figure.png" pageId="13" pageNumber="14">54</figureCitation>
,
<figureCitation id="136AB0BC900AFFBBFDF1BD24C0FD6572" box="[621,651,870,894]" captionStart="FIGURES 58 61" captionStartId="29.[151,264,1972,1995]" captionTargetBox="[188,1384,193,1901]" captionTargetId="figure@29.[188,1385,193,1924]" captionTargetPageId="29" captionText="FIGURES 58 61. Hindwings of dolichoderine males. 58, Ravavy miafina (CASENT 0179530); 59, Tapinoma mg 10 (CASENT 0179493); 60, Tapinoma subtile (CASENT 0179490); 61, Technomyrmex madecassus (CASENT 0179526)." httpUri="https://zenodo.org/record/277007/files/figure.png" pageId="13" pageNumber="14">61</figureCitation>
,
<figureCitation id="136AB0BC900AFFBBFD0BBD24C0C36572" box="[663,693,870,894]" captionStart="FIGURES 62 66" captionStartId="30.[151,264,1944,1967]" captionTargetBox="[162,1421,193,1904]" captionTargetId="figure@30.[162,1422,193,1910]" captionTargetPageId="30" captionText="FIGURES 62 66. Median hypostoma of dolichoderine males in ectal view. 62, Aptinoma mangabe (CASENT 0175002); 63, Ochetellus glaber (CASENT 0179489); 64, Ravavy miafina (CASENT 0179533); 65, Tapinoma mg 11 (CASENT 0179494); 66, Technomyrmex pallipes (CASENT 0179528)." httpUri="https://zenodo.org/record/277008/files/figure.png" pageId="13" pageNumber="14">66</figureCitation>
,
<figureCitation id="136AB0BC900AFFBBFD5DBD24C0A96572" box="[705,735,870,894]" captionStart="FIGURES 67 71" captionStartId="31.[151,264,1946,1969]" captionTargetBox="[199,1392,193,1931]" captionTargetId="figure@31.[199,1393,193,1938]" captionTargetPageId="31" captionText="FIGURES 67 71. Labrums of dolichoderine males. 67, Aptinoma mangabe (CASENT 0175002); 68, Ochetellus glaber (CASENT 0179489); 69, Ravavy miafina (CASENT 0179530); 70, Tapinoma mg 11 (CASENT 0179494); 71, Technomyrmex pallipes (CASENT 0179528). 67, 68, 70, in ectal view; 69, 71, in mesal view." httpUri="https://zenodo.org/record/277009/files/figure.png" pageId="13" pageNumber="14">71</figureCitation>
,
<figureCitation id="136AB0BC900AFFBBFD77BD24C17C6572" box="[747,778,870,894]" captionStart="FIGURES 72 76" captionStartId="32.[151,264,1873,1896]" captionTargetBox="[151,1434,193,1844]" captionTargetId="figure@32.[151,1436,193,1853]" captionTargetPageId="32" captionText="FIGURES 72 76. Mandibles of dolichoderine males. 72, Aptinoma mangabe (CASENT 0175002); 73, Ochetellus glaber (CASENT 0179489); 74, Ravavy miafina (CASENT 0179530); 75, Tapinoma mg 11 (CASENT 0179494); 76, Technomyrmex pallipes (CASENT 0179528)." httpUri="https://zenodo.org/record/277010/files/figure.png" pageId="13" pageNumber="14">76</figureCitation>
,
<figureCitation id="136AB0BC900AFFBBFC89BD24C1446572" box="[789,818,870,894]" captionStart="FIGURES 77 81" captionStartId="33.[151,264,1274,1297]" captionTargetBox="[151,1436,193,1236]" captionTargetId="figure@33.[151,1436,193,1253]" captionTargetPageId="33" captionText="FIGURES 77 81. Mouthparts of dolichoderine males. 77, Aptinoma mangabe (CASENT 0175002); 78, Ochetellus glaber (CASENT 0179489); 79, Ravavy miafina (CASENT 0179530); 80, Tapinoma subtile (CASENT 0179490); 81, Technomyrmex pallipes (CASENT 0179528). 77, 79 81, in mesal view; 78, in lateral view." httpUri="https://zenodo.org/record/277011/files/figure.png" pageId="13" pageNumber="14">81</figureCitation>
)
</paragraph>
<paragraph id="8BEEAC39900AFFBBFF0BBDECC16A6322" blockId="13.[151,1437,942,2011]" pageId="13" pageNumber="14">
With characters of
<taxonomicName id="4C51D7BA900AFFBBFEF3BDECC06965CA" box="[367,543,942,966]" class="Insecta" family="Formicidae" kingdom="Animalia" order="Hymenoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="subFamily" subFamily="Dolichoderinae">Dolichoderinae</taxonomicName>
. Ergatoid males known, but all males in
<collectingCountry id="F346ECA9900AFFBBFC73BDECC62A65CA" box="[1007,1116,942,966]" name="Madagascar" pageId="13" pageNumber="14">Malagasy</collectingCountry>
region alate. Medial hypostoma present (
<figureCitation id="136AB0BC900AFFBBFEABBD90C3FA65E6" box="[311,396,978,1002]" captionStart="FIGURES 62 66" captionStartId="30.[151,264,1944,1967]" captionTargetBox="[162,1421,193,1904]" captionTargetId="figure@30.[162,1422,193,1910]" captionTargetPageId="30" captionText="FIGURES 62 66. Median hypostoma of dolichoderine males in ectal view. 62, Aptinoma mangabe (CASENT 0175002); 63, Ochetellus glaber (CASENT 0179489); 64, Ravavy miafina (CASENT 0179533); 65, Tapinoma mg 11 (CASENT 0179494); 66, Technomyrmex pallipes (CASENT 0179528)." httpUri="https://zenodo.org/record/277008/files/figure.png" pageId="13" pageNumber="14">Fig. 66</figureCitation>
). Mandible triangular, but its basal angle indistinct. Basal margin and masticatory margin of mandible wholly covered with serrate denticles (
<figureCitation id="136AB0BC900AFFBBFD29BDB4C17E6202" box="[693,776,1014,1038]" captionStart="FIGURES 72 76" captionStartId="32.[151,264,1873,1896]" captionTargetBox="[151,1434,193,1844]" captionTargetId="figure@32.[151,1436,193,1853]" captionTargetPageId="32" captionText="FIGURES 72 76. Mandibles of dolichoderine males. 72, Aptinoma mangabe (CASENT 0175002); 73, Ochetellus glaber (CASENT 0179489); 74, Ravavy miafina (CASENT 0179530); 75, Tapinoma mg 11 (CASENT 0179494); 76, Technomyrmex pallipes (CASENT 0179528)." httpUri="https://zenodo.org/record/277010/files/figure.png" pageId="13" pageNumber="14">Fig. 76</figureCitation>
). Apical tooth on masticatory margin longer than subapical one. Palpal formula 6,4 (one specimen of each of eight species dissected:
<figureCitation id="136AB0BC900AFFBBFC48BA58C650623E" box="[980,1062,1050,1074]" captionStart="FIGURES 77 81" captionStartId="33.[151,264,1274,1297]" captionTargetBox="[151,1436,193,1236]" captionTargetId="figure@33.[151,1436,193,1253]" captionTargetPageId="33" captionText="FIGURES 77 81. Mouthparts of dolichoderine males. 77, Aptinoma mangabe (CASENT 0175002); 78, Ochetellus glaber (CASENT 0179489); 79, Ravavy miafina (CASENT 0179530); 80, Tapinoma subtile (CASENT 0179490); 81, Technomyrmex pallipes (CASENT 0179528). 77, 79 81, in mesal view; 78, in lateral view." httpUri="https://zenodo.org/record/277011/files/figure.png" pageId="13" pageNumber="14">Fig. 81</figureCitation>
). Third maxillary palpal segment shorter than or as long as fourth. Concavity on distal margin of labrum
<taxonomicName id="4C51D7BA900AFFBBFC4FBA7CC66C625A" box="[979,1050,1086,1110]" class="Insecta" family="Formicidae" genus="Chronoxenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="absent">absent</taxonomicName>
in most cases; when visible, it is much reduced in size and longest setae located distant from apices of lobes (
<figureCitation id="136AB0BC900AFFBBFC79BA20C64E6276" box="[997,1080,1122,1146]" captionStart="FIGURES 67 71" captionStartId="31.[151,264,1946,1969]" captionTargetBox="[199,1392,193,1931]" captionTargetId="figure@31.[199,1393,193,1938]" captionTargetPageId="31" captionText="FIGURES 67 71. Labrums of dolichoderine males. 67, Aptinoma mangabe (CASENT 0175002); 68, Ochetellus glaber (CASENT 0179489); 69, Ravavy miafina (CASENT 0179530); 70, Tapinoma mg 11 (CASENT 0179494); 71, Technomyrmex pallipes (CASENT 0179528). 67, 68, 70, in ectal view; 69, 71, in mesal view." httpUri="https://zenodo.org/record/277009/files/figure.png" pageId="13" pageNumber="14">Fig. 71</figureCitation>
). Scape excluding its basal condyle shorter than length of flagellar segments 1+2. Pedicel conical (
<figureCitation id="136AB0BC900AFFBBFC3BBAC4C1986292" box="[935,1006,1158,1182]" captionStart="FIGURES 1 5" captionStartId="16.[151,264,1953,1976]" captionTargetBox="[202,1381,195,1911]" captionTargetId="figure@16.[202,1382,195,1927]" captionTargetPageId="16" captionText="FIGURES 1 5. Head of dolichoderine males in full-face view. 1, Ochetellus glaber (CASENT 0191065); 2, Aptinoma mangabe (CASENT 0173594); 3, Ravavy miafina (CASENT 0474633); 4, Tapinoma subtile (CASENT 0136850); 5, Technomyrmex difficilis (CASENT 0049968)." httpUri="https://zenodo.org/record/276994/files/figure.png" pageId="13" pageNumber="14">Fig. 5</figureCitation>
). First and second flagellar segments straight. Axillae medially compressed, anterior and posterior margins not parallel. Petiolar node strongly inclined anteriorly, its anterior margin much shorter than posterior margin in lateral view, not much expanded laterally. Petiole narrowly attached to abdominal segment III. Anterior surface of abdominal segment III with indentation that fits posterior surface of petiolar node. Pygostyles present.
</paragraph>
<paragraph id="8BEEAC39900AFFBBFF59BB78C77263B2" blockId="13.[151,1437,942,2011]" pageId="13" pageNumber="14">
Distal portion of abdominal sternum IX bilobed, its distal margin widely concave (
<figureCitation id="136AB0BC900AFFBBFBF7BB78C6C9635E" box="[1131,1215,1338,1362]" captionStart="FIGURES 24 28" captionStartId="20.[151,264,1912,1935]" captionTargetBox="[195,1392,193,1875]" captionTargetId="figure@20.[195,1393,193,1891]" captionTargetPageId="20" captionText="FIGURES 24 28. Abdominal sternum IX of dolichoderine males in ectal view. 24, Aptinoma mangabe (CASENT 0175002); 25, Ochetellus glaber (CASENT 0179489); 26, Ravavy miafina (CASENT 0179530); 27, Tapinoma melanocephalum (CASENT 0492372); 28, Technomyrmex madecassus (CASENT 0179526)." httpUri="https://zenodo.org/record/276998/files/figure.png" pageId="13" pageNumber="14">Fig. 28</figureCitation>
). Apicoventral portion of basimere without projection. Harpago moderate in size, separated from basimere only by suture, without membranous region between them (
<figureCitation id="136AB0BC900AFFBBFDB3BBC0C0F16396" box="[559,647,1410,1434]" captionStart="FIGURES 29 33" captionStartId="21.[151,264,1944,1967]" captionTargetBox="[204,1379,193,1905]" captionTargetId="figure@21.[204,1380,193,1921]" captionTargetPageId="21" captionText="FIGURES 29 33. Genital capsules of dolichoderine males. 29, Tapinoma mg 11 (CASENT 0179494); 30, Aptinoma mangabe (CASENT 0175002); 31, Ochetellus glaber (CASENT 0179489); 32, Ravavy miafina (CASENT 0179533); 33, Technomyrmex madecassus (CASENT 0179526). Right side, ventral view; left side, dorsal view." httpUri="https://zenodo.org/record/276999/files/figure.png" pageId="13" pageNumber="14">Figs 33</figureCitation>
,
<figureCitation id="136AB0BC900AFFBBFD09BBC0C0C46396" box="[661,690,1410,1434]" captionStart="FIGURES 34 38" captionStartId="22.[151,264,1945,1968]" captionTargetBox="[213,1384,193,1897]" captionTargetId="figure@22.[207,1386,193,1926]" captionTargetPageId="22" captionText="FIGURES 34 38. Genital capsules of dolichoderine males in lateral view. 34, Ochetellus glaber (CASENT 0179489); 35, Aptinoma mangabe (CASENT 0175002); 36, Ravavy miafina (CASENT 0179533); 37, Tapinoma mg 11 (CASENT 0179494); 38, Technomyrmex madecassus (CASENT 0179526)." httpUri="https://zenodo.org/record/277000/files/figure.png" pageId="13" pageNumber="14">38</figureCitation>
),
<emphasis id="B925702B900AFFBBFD56BBC1C30C63B2" italics="true" pageId="13" pageNumber="14">expanded mesally in ventral view, forming distinct and more or less flat ventral face</emphasis>
(
<figureCitation id="136AB0BC900AFFBBFE14BBE4C3AB63B2" box="[392,477,1446,1470]" captionStart="FIGURES 16 23" captionStartId="19.[151,264,1847,1870]" captionTargetBox="[237,1345,193,1821]" captionTargetId="figure@19.[237,1346,193,1822]" captionTargetPageId="19" captionText="FIGURES 16 23. Males of Dolichoderinae. 16, Aptinoma mangabe (CASENT 0173594); 17, Aptinoma mangabe (CASENT 0175002); 18, 20, Ochetellus glaber (CASENT 0179489); 19, 21, Ravavy miafina (CASENT 0080308); 22, Technomyrmex curiosus (paratype: CASENT 0070364); 23, Tapinoma mg 10 (CASENT 0115650). 16, distal portion of abdomen in oblique posterior view; 17, petiole in lateral view; 18, 19, petiole and abdominal segments in ventral view; 20, petiole and abdominal segments in dorsal view; 21, petiole and abdominal segments in oblique frontal view; 22, 23, genitalia in ventral view." httpUri="https://zenodo.org/record/276997/files/figure.png" pageId="13" pageNumber="14">Fig. 22</figureCitation>
). Basal portion of aedeagus does not bear distinct ventral lobe (
<figureCitation id="136AB0BC900AFFBBFB3FBBE4C68163B2" box="[1187,1271,1446,1470]" captionStart="FIGURES 41 45" captionStartId="24.[151,264,1945,1968]" captionTargetBox="[200,1379,193,1913]" captionTargetId="figure@24.[200,1380,193,1928]" captionTargetPageId="24" captionText="FIGURES 41 45. Aedeagal plates of dolichoderine males in lateral view. 41, Ravavy miafina (CASENT 0179530); 42, Aptinoma mangabe (CASENT 0175002); 43, Ochetellus glaber (CASENT 0179489); 44, Tapinoma mg 11 (CASENT 0179494); 45, Technomyrmex madecassus (CASENT 0179526)." httpUri="https://zenodo.org/record/277002/files/figure.png" pageId="13" pageNumber="14">Fig. 45</figureCitation>
).
</paragraph>
<paragraph id="8BEEAC39900AFFBBFF59BB88C6726026" blockId="13.[151,1437,942,2011]" pageId="13" pageNumber="14">
Forewing not extremely elongated apical to wing stigma, its radial sector reaches to costal margin (
<figureCitation id="136AB0BC900AFFBBFAA5BB88C7F963EE" box="[1337,1423,1482,1506]" captionStart="FIGURES 52 54" captionStartId="27.[151,264,1973,1996]" captionTargetBox="[221,1390,193,1923]" captionTargetId="figure@27.[195,1393,193,1952]" captionTargetPageId="27" captionText="FIGURES 52 54. Forewings of male ants. 52, Ravavy miafina (CASENT 0179530); 53, Tapinoma mg 10 (CASENT 0179493); 54, Technomyrmex madecassus (CASENT 0179526)." httpUri="https://zenodo.org/record/277005/files/figure.png" pageId="13" pageNumber="14">Fig. 54</figureCitation>
), media partially
<taxonomicName id="4C51D7BA900AFFBBFED6BBACC3E4600A" box="[330,402,1518,1542]" class="Insecta" family="Formicidae" genus="Chronoxenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="absent">absent</taxonomicName>
between Rs+M and 2rs-m, and 1m-cu
<taxonomicName id="4C51D7BA900AFFBBFCCFBBACC1EC600A" box="[851,922,1518,1542]" class="Insecta" family="Formicidae" genus="Chronoxenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="absent">absent</taxonomicName>
in many cases. On hindwing,
<emphasis id="B925702B900AFFBBFB64BBADC7E3600A" box="[1272,1429,1518,1542]" italics="true" pageId="13" pageNumber="14">
M+
<taxonomicName id="4C51D7BA900AFFBBFABFBBADC7E3600A" box="[1315,1429,1518,1542]" class="Insecta" family="Formicidae" genus="Chronoxenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="absent">Cu absent</taxonomicName>
</emphasis>
, M+1m-cu, free sections of radial sector and cubitus, and cu-a
<taxonomicName id="4C51D7BA900AFFBBFCD1B850C1E36026" box="[845,917,1554,1578]" class="Insecta" family="Formicidae" genus="Chronoxenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="absent">absent</taxonomicName>
(
<figureCitation id="136AB0BC900AFFBBFC38B850C1816026" box="[932,1015,1554,1578]" captionStart="FIGURES 58 61" captionStartId="29.[151,264,1972,1995]" captionTargetBox="[188,1384,193,1901]" captionTargetId="figure@29.[188,1385,193,1924]" captionTargetPageId="29" captionText="FIGURES 58 61. Hindwings of dolichoderine males. 58, Ravavy miafina (CASENT 0179530); 59, Tapinoma mg 10 (CASENT 0179493); 60, Tapinoma subtile (CASENT 0179490); 61, Technomyrmex madecassus (CASENT 0179526)." httpUri="https://zenodo.org/record/277007/files/figure.png" pageId="13" pageNumber="14">Fig. 61</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C34BFFB2900AFFB8FF59B875C65267F8" lastPageId="14" lastPageNumber="15" pageId="13" pageNumber="14" type="discussion">
<paragraph id="8BEEAC39900AFFBBFF59B875C1966162" blockId="13.[151,1437,942,2011]" pageId="13" pageNumber="14">
<emphasis id="B925702B900AFFBBFF59B875C34F6043" bold="true" box="[197,313,1591,1615]" pageId="13" pageNumber="14">Remarks.</emphasis>
Males of nine species were examined. They are distinguished easily from those of the four other
<collectingCountry id="F346ECA9900AFFBBFF0BB818C372607E" box="[151,260,1626,1650]" name="Madagascar" pageId="13" pageNumber="14">Malagasy</collectingCountry>
dolichoderine genera by the ventral portion of the harpago, which is expanded mesally to form a ventral face (
<figureCitation id="136AB0BC900AFFBBFF4AB83CC35D609A" box="[214,299,1662,1686]" captionStart="FIGURES 16 23" captionStartId="19.[151,264,1847,1870]" captionTargetBox="[237,1345,193,1821]" captionTargetId="figure@19.[237,1346,193,1822]" captionTargetPageId="19" captionText="FIGURES 16 23. Males of Dolichoderinae. 16, Aptinoma mangabe (CASENT 0173594); 17, Aptinoma mangabe (CASENT 0175002); 18, 20, Ochetellus glaber (CASENT 0179489); 19, 21, Ravavy miafina (CASENT 0080308); 22, Technomyrmex curiosus (paratype: CASENT 0070364); 23, Tapinoma mg 10 (CASENT 0115650). 16, distal portion of abdomen in oblique posterior view; 17, petiole in lateral view; 18, 19, petiole and abdominal segments in ventral view; 20, petiole and abdominal segments in dorsal view; 21, petiole and abdominal segments in oblique frontal view; 22, 23, genitalia in ventral view." httpUri="https://zenodo.org/record/276997/files/figure.png" pageId="13" pageNumber="14">Fig. 22</figureCitation>
), and the absence of M+Cu on the hindwing (
<figureCitation id="136AB0BC900AFFBBFCAAB83CC1FD609A" box="[822,907,1662,1686]" captionStart="FIGURES 58 61" captionStartId="29.[151,264,1972,1995]" captionTargetBox="[188,1384,193,1901]" captionTargetId="figure@29.[188,1385,193,1924]" captionTargetPageId="29" captionText="FIGURES 58 61. Hindwings of dolichoderine males. 58, Ravavy miafina (CASENT 0179530); 59, Tapinoma mg 10 (CASENT 0179493); 60, Tapinoma subtile (CASENT 0179490); 61, Technomyrmex madecassus (CASENT 0179526)." httpUri="https://zenodo.org/record/277007/files/figure.png" pageId="13" pageNumber="14">Fig. 61</figureCitation>
).
<taxonomicName id="4C51D7BA900AFFBBFC3CB83CC63E609A" box="[928,1096,1662,1686]" class="Insecta" family="Formicidae" genus="Technomyrmex" kingdom="Animalia" order="Hymenoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="genus">
<emphasis id="B925702B900AFFBBFC3CB83CC63E609A" box="[928,1096,1662,1686]" italics="true" pageId="13" pageNumber="14">Technomyrmex</emphasis>
</taxonomicName>
and
<taxonomicName id="4C51D7BA900AFFBBFB1EB83CC699609A" box="[1154,1263,1662,1686]" class="Insecta" family="Formicidae" genus="Aptinoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="genus">
<emphasis id="B925702B900AFFBBFB1EB83CC699609A" box="[1154,1263,1662,1686]" italics="true" pageId="13" pageNumber="14">Aptinoma</emphasis>
</taxonomicName>
uniquely share the following characters: the basal margin of the mandible is wholly covered with serrate denticles (
<figureCitation id="136AB0BC900AFFBBFB77B8E0C74A60B6" box="[1259,1340,1698,1722]" captionStart="FIGURES 72 76" captionStartId="32.[151,264,1873,1896]" captionTargetBox="[151,1434,193,1844]" captionTargetId="figure@32.[151,1436,193,1853]" captionTargetPageId="32" captionText="FIGURES 72 76. Mandibles of dolichoderine males. 72, Aptinoma mangabe (CASENT 0175002); 73, Ochetellus glaber (CASENT 0179489); 74, Ravavy miafina (CASENT 0179530); 75, Tapinoma mg 11 (CASENT 0179494); 76, Technomyrmex pallipes (CASENT 0179528)." httpUri="https://zenodo.org/record/277010/files/figure.png" pageId="13" pageNumber="14">Fig. 76</figureCitation>
) and the concavity on the distal margin of the labrum is reduced (
<figureCitation id="136AB0BC900AFFBBFC96B884C12A60D2" box="[778,860,1734,1758]" captionStart="FIGURES 67 71" captionStartId="31.[151,264,1946,1969]" captionTargetBox="[199,1392,193,1931]" captionTargetId="figure@31.[199,1393,193,1938]" captionTargetPageId="31" captionText="FIGURES 67 71. Labrums of dolichoderine males. 67, Aptinoma mangabe (CASENT 0175002); 68, Ochetellus glaber (CASENT 0179489); 69, Ravavy miafina (CASENT 0179530); 70, Tapinoma mg 11 (CASENT 0179494); 71, Technomyrmex pallipes (CASENT 0179528). 67, 68, 70, in ectal view; 69, 71, in mesal view." httpUri="https://zenodo.org/record/277009/files/figure.png" pageId="13" pageNumber="14">Fig. 71</figureCitation>
). In
<taxonomicName id="4C51D7BA900AFFBBFC12B884C64160D2" box="[910,1079,1734,1758]" class="Insecta" family="Formicidae" genus="Technomyrmex" kingdom="Animalia" order="Hymenoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="genus">
<emphasis id="B925702B900AFFBBFC12B884C64160D2" box="[910,1079,1734,1758]" italics="true" pageId="13" pageNumber="14">Technomyrmex</emphasis>
</taxonomicName>
and
<taxonomicName id="4C51D7BA900AFFBBFBF0B88AC6AF60D3" box="[1132,1241,1736,1759]" class="Insecta" family="Formicidae" genus="Tapinoma" kingdom="Animalia" order="Hymenoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="genus">
<emphasis id="B925702B900AFFBBFBF0B88AC6AF60D3" box="[1132,1241,1736,1759]" italics="true" pageId="13" pageNumber="14">Tapinoma</emphasis>
</taxonomicName>
the petiolar node is inclined. The central notch of the labrum is reduced in size or
<taxonomicName id="4C51D7BA900AFFBBFCEDB8A8C1CD610E" box="[881,955,1770,1794]" class="Insecta" family="Formicidae" genus="Chronoxenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="absent">absent</taxonomicName>
, although it is still visible in several males of
<taxonomicName id="4C51D7BA900AFFBBFF2AB94CC328612A" box="[182,350,1806,1830]" class="Insecta" family="Formicidae" genus="Technomyrmex" kingdom="Animalia" order="Hymenoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="genus">
<emphasis id="B925702B900AFFBBFF2AB94CC328612A" box="[182,350,1806,1830]" italics="true" pageId="13" pageNumber="14">Technomyrmex</emphasis>
</taxonomicName>
. When the notch is visible in
<taxonomicName id="4C51D7BA900AFFBBFD2DB94CC12F612A" box="[689,857,1806,1830]" class="Insecta" family="Formicidae" genus="Technomyrmex" kingdom="Animalia" order="Hymenoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="genus">
<emphasis id="B925702B900AFFBBFD2DB94CC12F612A" box="[689,857,1806,1830]" italics="true" pageId="13" pageNumber="14">Technomyrmex</emphasis>
</taxonomicName>
, the longest setae on the labrum are located distant from the apices of the lobes formed by the central notch, while in the other dolichoderine genera with bilobed labrum, the setae are mostly located on the apex of each lobe (
<figureCitation id="136AB0BC900AFFBBFCCCB914C1A56162" box="[848,979,1878,1902]" captionStart="FIGURES 67 71" captionStartId="31.[151,264,1946,1969]" captionTargetBox="[199,1392,193,1931]" captionTargetId="figure@31.[199,1393,193,1938]" captionTargetPageId="31" captionText="FIGURES 67 71. Labrums of dolichoderine males. 67, Aptinoma mangabe (CASENT 0175002); 68, Ochetellus glaber (CASENT 0179489); 69, Ravavy miafina (CASENT 0179530); 70, Tapinoma mg 11 (CASENT 0179494); 71, Technomyrmex pallipes (CASENT 0179528). 67, 68, 70, in ectal view; 69, 71, in mesal view." httpUri="https://zenodo.org/record/277009/files/figure.png" pageId="13" pageNumber="14">Figs 6870</figureCitation>
).
</paragraph>
<paragraph id="8BEEAC39900AFFB8FF59B938C34D67C3" blockId="13.[151,1437,942,2011]" lastBlockId="14.[151,1436,151,500]" lastPageId="14" lastPageNumber="15" pageId="13" pageNumber="14">
Several of the present results for
<taxonomicName id="4C51D7BA900AFFBBFDA0B939C093619F" box="[572,741,1915,1939]" class="Insecta" family="Formicidae" genus="Technomyrmex" kingdom="Animalia" order="Hymenoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="genus">
<emphasis id="B925702B900AFFBBFDA0B939C093619F" box="[572,741,1915,1939]" italics="true" pageId="13" pageNumber="14">Technomyrmex</emphasis>
</taxonomicName>
disagree with previous studies. A median notch or groove on the anterior margin of the median hypostoma is found in all males of
<taxonomicName id="4C51D7BA900AFFBBFC3DB9DCC63C61BA" box="[929,1098,1950,1974]" class="Insecta" family="Formicidae" genus="Technomyrmex" kingdom="Animalia" order="Hymenoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="genus">
<emphasis id="B925702B900AFFBBFC3DB9DCC63C61BA" box="[929,1098,1950,1974]" italics="true" pageId="13" pageNumber="14">Technomyrmex</emphasis>
</taxonomicName>
(
<figureCitation id="136AB0BC900AFFBBFBC5B9DCC6DB61BA" box="[1113,1197,1950,1974]" captionStart="FIGURES 62 66" captionStartId="30.[151,264,1944,1967]" captionTargetBox="[162,1421,193,1904]" captionTargetId="figure@30.[162,1422,193,1910]" captionTargetPageId="30" captionText="FIGURES 62 66. Median hypostoma of dolichoderine males in ectal view. 62, Aptinoma mangabe (CASENT 0175002); 63, Ochetellus glaber (CASENT 0179489); 64, Ravavy miafina (CASENT 0179533); 65, Tapinoma mg 11 (CASENT 0179494); 66, Technomyrmex pallipes (CASENT 0179528)." httpUri="https://zenodo.org/record/277008/files/figure.png" pageId="13" pageNumber="14">Fig. 66</figureCitation>
), while the margin in
<bibRefCitation id="EFC0D1C8900AFFBBFF0BB980C32F61D6" author="Shattuck" box="[151,345,1986,2010]" pageId="13" pageNumber="14" refString="Shattuck, S. O. (1992 a) Generic revision of the ant subfamily Dolichoderinae (Hymenoptera: Formicidae). Sociobiology, 21, 1 - 181." type="journal article" year="1992" yearSuffix="a">Shattuck (1992a)</bibRefCitation>
is regarded as entire. The inner margin of the eye in full-face view is slightly concave in most species, while in
<bibRefCitation id="EFC0D1C89009FFB8FEC2BED5C05666A3" author="Shattuck" box="[350,544,151,175]" pageId="14" pageNumber="15" refString="Shattuck, S. O. (1992 a) Generic revision of the ant subfamily Dolichoderinae (Hymenoptera: Formicidae). Sociobiology, 21, 1 - 181." type="journal article" year="1992" yearSuffix="a">Shattuck (1992a)</bibRefCitation>
the margin is regarded as flat. The anteromedial margin of the clypeus is not concave in several species (
<taxonomicName id="4C51D7BA9009FFB8FE43BEFFC02D66D8" box="[479,603,188,212]" class="Insecta" family="Formicidae" genus="Technomyrmex" kingdom="Animalia" order="Hymenoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="difficilis">
<emphasis id="B925702B9009FFB8FE43BEFFC39A66D8" box="[479,492,189,212]" italics="true" pageId="14" pageNumber="15">T</emphasis>
.
<emphasis id="B925702B9009FFB8FE63BEFEC02D66D8" box="[511,603,188,212]" italics="true" pageId="14" pageNumber="15">difficilis</emphasis>
</taxonomicName>
,
<taxonomicName id="4C51D7BA9009FFB8FDF1BEFFC0A266D8" box="[621,724,188,212]" class="Insecta" family="Formicidae" genus="Technomyrmex" kingdom="Animalia" order="Hymenoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="fisheri">
<emphasis id="B925702B9009FFB8FDF1BEFFC00C66D8" box="[621,634,189,212]" italics="true" pageId="14" pageNumber="15">T</emphasis>
.
<emphasis id="B925702B9009FFB8FD11BEFEC0A266D8" box="[653,724,188,212]" italics="true" pageId="14" pageNumber="15">fisheri</emphasis>
</taxonomicName>
, and
<taxonomicName id="4C51D7BA9009FFB8FC87BEFFC1E966D8" box="[795,927,189,212]" class="Insecta" family="Formicidae" genus="Technomyrmex" kingdom="Animalia" order="Hymenoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="innocens">
<emphasis id="B925702B9009FFB8FC87BEFFC15E66D8" box="[795,808,189,212]" italics="true" pageId="14" pageNumber="15">T</emphasis>
.
<emphasis id="B925702B9009FFB8FCA7BEFFC1E966D8" box="[827,927,189,212]" italics="true" pageId="14" pageNumber="15">innocens</emphasis>
</taxonomicName>
); the margin of
<taxonomicName id="4C51D7BA9009FFB8FBF9BEFEC77866D8" box="[1125,1294,188,212]" class="Insecta" family="Formicidae" genus="Technomyrmex" kingdom="Animalia" order="Hymenoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="genus">
<emphasis id="B925702B9009FFB8FBF9BEFEC77866D8" box="[1125,1294,188,212]" italics="true" pageId="14" pageNumber="15">Technomyrmex</emphasis>
</taxonomicName>
in
<bibRefCitation id="EFC0D1C89009FFB8FAA6BEFEC29166FB" author="Shattuck" pageId="14" pageNumber="15" refString="Shattuck, S. O. (1992 a) Generic revision of the ant subfamily Dolichoderinae (Hymenoptera: Formicidae). Sociobiology, 21, 1 - 181." type="journal article" year="1992" yearSuffix="a">Shattuck (1992a</bibRefCitation>
,
<bibRefCitation id="EFC0D1C89009FFB8FF6FBE9DC35A66FB" author="Shattuck" box="[243,300,223,247]" pageId="14" pageNumber="15" refString="Shattuck, S. O. (1995) Generic-level relationships within the ant subfamily Dolichoderinae (Hymenoptera: Formicidae). Systematic Entomology, 20, 217 - 228." type="journal article" year="1995">1995</bibRefCitation>
) and
<bibRefCitation id="EFC0D1C89009FFB8FEF7BE9DC02A66FB" box="[363,604,223,248]" pageId="14" pageNumber="15" refString="Brandao, C. R. F., Baroni Urbani, C., Wagensberg, J. &amp; Yamamoto, C. I. (1999 (1998 )) New Technomyrmex in Dominican amber (Hymenoptera: Formicidae), with a reappraisal of Dolichoderinae phylogeny. Entomologica Scandinavica, 29, 411 - 428." type="journal article">
Brandão
<emphasis id="B925702B9009FFB8FE4DBEA3C07466F4" box="[465,514,224,248]" italics="true" pageId="14" pageNumber="15">et al</emphasis>
. (1999)
</bibRefCitation>
is treated as broadly concave. The relative length of the first flagellomere compared with its width varies from less than two times to three times (see
<tableCitation id="C6D399829009FFB8FC6EBF46C6316710" box="[1010,1095,260,284]" captionStart="TABLE 4" captionStartId="10.[151,235,152,173]" captionText="TABLE 4. Character matrix for males of Malagasy Dolichoderinae. Intra-generic variations were observed in these characters, making them useful for distinguishing among species rather than genera. For the six characters, character states are shown as 0, 1, 2; as?, for states difficult to judge; or 0 / 1, if both states 0 and 1 were observed for each genus. Number of species in which the character states were observed is given in parentheses following character state. Character states have been confirmed by direct observation or by dissection." pageId="14" pageNumber="15">Table 4</tableCitation>
), while the length in
<bibRefCitation id="EFC0D1C89009FFB8FAA6BF46C2986733" author="Shattuck" pageId="14" pageNumber="15" refString="Shattuck, S. O. (1992 a) Generic revision of the ant subfamily Dolichoderinae (Hymenoptera: Formicidae). Sociobiology, 21, 1 - 181." type="journal article" year="1992" yearSuffix="a">Shattuck (1992a)</bibRefCitation>
is regarded as less than two times. Abdominal segment III does not always cover the petiole completely by overhanging it anteriorly in
<taxonomicName id="4C51D7BA9009FFB8FE40BF0EC0F26768" box="[476,644,332,356]" class="Insecta" family="Formicidae" genus="Technomyrmex" kingdom="Animalia" order="Hymenoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="genus">
<emphasis id="B925702B9009FFB8FE40BF0EC0F26768" box="[476,644,332,356]" italics="true" pageId="14" pageNumber="15">Technomyrmex</emphasis>
</taxonomicName>
(e.g.
<taxonomicName id="4C51D7BA9009FFB8FD56BF0FC15F6768" box="[714,809,333,356]" class="Insecta" family="Formicidae" genus="Technomyrmex" kingdom="Animalia" order="Hymenoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="mayri">
<emphasis id="B925702B9009FFB8FD56BF0FC0AE6768" box="[714,728,333,356]" italics="true" pageId="14" pageNumber="15">T</emphasis>
.
<emphasis id="B925702B9009FFB8FD74BF0FC15F6768" box="[744,809,333,356]" italics="true" pageId="14" pageNumber="15">mayri</emphasis>
</taxonomicName>
), although this segment in
<taxonomicName id="4C51D7BA9009FFB8FBF4BF0EC7676768" box="[1128,1297,332,356]" class="Insecta" family="Formicidae" genus="Technomyrmex" kingdom="Animalia" order="Hymenoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="genus">
<emphasis id="B925702B9009FFB8FBF4BF0EC7676768" box="[1128,1297,332,356]" italics="true" pageId="14" pageNumber="15">Technomyrmex</emphasis>
</taxonomicName>
in
<bibRefCitation id="EFC0D1C89009FFB8FAA6BF0EC291678B" author="Shattuck" pageId="14" pageNumber="15" refString="Shattuck, S. O. (1992 a) Generic revision of the ant subfamily Dolichoderinae (Hymenoptera: Formicidae). Sociobiology, 21, 1 - 181." type="journal article" year="1992" yearSuffix="a">Shattuck (1992a</bibRefCitation>
,
<bibRefCitation id="EFC0D1C89009FFB8FF6FBF2DC35A678B" author="Shattuck" box="[243,300,367,391]" pageId="14" pageNumber="15" refString="Shattuck, S. O. (1995) Generic-level relationships within the ant subfamily Dolichoderinae (Hymenoptera: Formicidae). Systematic Entomology, 20, 217 - 228." type="journal article" year="1995">1995</bibRefCitation>
) is regarded as overhanging the petiole so that the latter is invisible in dorsal view. The cuspis on the volsella is
<taxonomicName id="4C51D7BA9009FFB8FE93BFD6C32167A0" box="[271,343,404,428]" class="Insecta" family="Formicidae" genus="Chronoxenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="absent">absent</taxonomicName>
in one species (
<taxonomicName id="4C51D7BA9009FFB8FD90BFD7C0D867A0" box="[524,686,404,428]" class="Insecta" family="Formicidae" genus="Technomyrmex" kingdom="Animalia" order="Hymenoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="madecassus">
<emphasis id="B925702B9009FFB8FD90BFD7C06F67A0" box="[524,537,405,428]" italics="true" pageId="14" pageNumber="15">T</emphasis>
.
<emphasis id="B925702B9009FFB8FDB4BFD6C0D867A0" box="[552,686,404,428]" italics="true" pageId="14" pageNumber="15">madecassus</emphasis>
</taxonomicName>
:
<figureCitation id="136AB0BC9009FFB8FD21BFD6C16667A0" box="[701,784,404,428]" captionStart="FIGURES 46 48" captionStartId="25.[151,264,1971,1994]" captionTargetBox="[163,1436,193,1909]" captionTargetId="figure@25.[163,1448,193,1940]" captionTargetPageId="25" captionText="FIGURES 46 48. Volsella of dolichoderine males in oblique posterior view. 46, Ochetellus glaber (CASENT 0179489); 47, Tapinoma mg 11 (CASENT 0179494); 48, Technomyrmex madecassus (CASENT 0179526)." httpUri="https://zenodo.org/record/277003/files/figure.png" pageId="14" pageNumber="15">Fig. 48</figureCitation>
), while regarded as present in
<taxonomicName id="4C51D7BA9009FFB8FBFBBFD6C76667A0" box="[1127,1296,404,428]" class="Insecta" family="Formicidae" genus="Technomyrmex" kingdom="Animalia" order="Hymenoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="genus">
<emphasis id="B925702B9009FFB8FBFBBFD6C76667A0" box="[1127,1296,404,428]" italics="true" pageId="14" pageNumber="15">Technomyrmex</emphasis>
</taxonomicName>
by
<bibRefCitation id="EFC0D1C89009FFB8FAA6BFD6C29167C3" author="Shattuck" pageId="14" pageNumber="15" refString="Shattuck, S. O. (1992 a) Generic revision of the ant subfamily Dolichoderinae (Hymenoptera: Formicidae). Sociobiology, 21, 1 - 181." type="journal article" year="1992" yearSuffix="a">Shattuck (1992a</bibRefCitation>
,
<bibRefCitation id="EFC0D1C89009FFB8FF6FBFF5C35967C3" author="Shattuck" box="[243,303,439,463]" pageId="14" pageNumber="15" refString="Shattuck, S. O. (1995) Generic-level relationships within the ant subfamily Dolichoderinae (Hymenoptera: Formicidae). Systematic Entomology, 20, 217 - 228." type="journal article" year="1995">1995</bibRefCitation>
).
</paragraph>
<paragraph id="8BEEAC399009FFB8FF59BF9EC65267F8" blockId="14.[151,1436,151,500]" box="[197,1060,476,500]" pageId="14" pageNumber="15">
Additional discussion of characters is included in the remarks for
<taxonomicName id="4C51D7BA9009FFB8FC3ABF9EC66867F8" box="[934,1054,476,500]" class="Insecta" family="Formicidae" genus="Ochetellus" kingdom="Animalia" order="Hymenoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="genus">
<emphasis id="B925702B9009FFB8FC3ABF9EC66867F8" box="[934,1054,476,500]" italics="true" pageId="14" pageNumber="15">Ochetellus</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>