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<document id="7881E15F1F2CD0300C941D868208B28A" ID-DOI="10.11646/zootaxa.3630.1.2" ID-GBIF-Dataset="28703b77-1579-49e7-885f-5c2edb895f83" ID-ISSN="1175-5326" ID-Zenodo-Dep="283467" ID-ZooBank="A338EE37-E8AC-4AC3-8D6B-D0177FB680BF" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460582983698" checkinUser="plazi" docAuthor="Jałoszyński, Paweł" docDate="2013" docId="03D687E5B0739A58BDD14643165FF844" docLanguage="en" docName="zt03630p079.pdf" docOrigin="Zootaxa 3630 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Scydmaenilla (Scydmaenilla) thompsoniana Franz" docType="treatment" docVersion="7" lastPageNumber="74" masterDocId="FFEFFF9DB0509A7BBD46427C1313FF91" masterDocTitle="Revision of subgenera of Stenichnus Thomson, with review of Australo-Pacific species (Coleoptera, Staphylinidae, Scydmaeninae)" masterLastPageNumber="79" masterPageNumber="39" pageNumber="74" updateTime="1698324670604" updateUser="plazi">
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<mods:title id="9676FD6B8DB217A92E7C5F05E22E643F">Revision of subgenera of Stenichnus Thomson, with review of Australo-Pacific species (Coleoptera, Staphylinidae, Scydmaeninae)</mods:title>
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<mods:roleTerm id="D3BC49BBA1D9C862A365D782F377E8EC">Author</mods:roleTerm>
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<mods:namePart id="B29705F89EF030190FF59824F2208F5A">Jałoszyński, Paweł</mods:namePart>
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<mods:date id="7432FBD85BF2734D8AD30FE6C6D76E05">2013</mods:date>
<mods:detail id="F740218CFB31C018685971660FB50EBA" type="volume">
<mods:number id="8960E68A32B03F8AA4208652546678D3">3630</mods:number>
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<mods:classification id="2B1A0533E246C00BC2A406CF88D5C613">journal article</mods:classification>
<mods:identifier id="90CB90B8101747067B266E4A7B6E42F4" type="DOI">10.11646/zootaxa.3630.1.2</mods:identifier>
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<treatment id="03D687E5B0739A58BDD14643165FF844" ID-DOI="http://doi.org/10.5281/zenodo.5696238" ID-GBIF-Taxon="127663491" ID-Zenodo-Dep="5696238" LSID="urn:lsid:plazi:treatment:03D687E5B0739A58BDD14643165FF844" httpUri="http://treatment.plazi.org/id/03D687E5B0739A58BDD14643165FF844" lastPageNumber="74" pageId="35" pageNumber="74">
<subSubSection id="C3656578B0739A58BDD1464311EFFB51" pageId="35" pageNumber="74" type="nomenclature">
<paragraph id="8BC036F3B0739A58BDD1464311EDFBC8" blockId="35.[151,766,1087,1146]" box="[151,766,1087,1113]" pageId="35" pageNumber="74">
<heading id="D088819FB0739A58BDD1464311EDFBC8" bold="true" box="[151,766,1087,1113]" fontSize="11" level="1" pageId="35" pageNumber="74" reason="1">
<taxonomicName id="4C7F4D70B0739A58BDD1464311EDFBC8" ID-CoL="79ZTL" authority="Franz" authorityName="Franz" box="[151,766,1087,1113]" class="Insecta" family="Staphylinidae" genus="Scydmaenilla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="35" pageNumber="74" phylum="Arthropoda" rank="species" species="thompsoniana" subGenus="Scydmaenilla">
<emphasis id="B90BEAE1B0739A58BDD1464311EDFBC8" bold="true" box="[151,766,1087,1113]" pageId="35" pageNumber="74">
<emphasis id="B90BEAE1B0739A58BDD146431229FBC8" bold="true" box="[151,314,1087,1113]" italics="true" pageId="35" pageNumber="74">Scydmaenilla</emphasis>
(
<emphasis id="B90BEAE1B0739A58BC0D464312FDFBC8" bold="true" box="[331,494,1087,1113]" italics="true" pageId="35" pageNumber="74">Scydmaenilla</emphasis>
)
<emphasis id="B90BEAE1B0739A58BCB8464311B9FBC8" bold="true" box="[510,682,1087,1113]" italics="true" pageId="35" pageNumber="74">thompsoniana</emphasis>
Franz
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="8BC036F3B0739A58BDD1461E1260FBEB" blockId="35.[151,766,1087,1146]" box="[151,371,1122,1146]" pageId="35" pageNumber="74">
(
<figureCitation id="13442A76B0739A58BDD9461E13EEFBEB" box="[159,253,1122,1146]" captionStart="FIGURES 93 – 98" captionStartId="28.[151,264,1922,1945]" captionTargetBox="[163,1419,179,1880]" captionTargetId="figure@28.[151,1436,171,1902]" captionTargetPageId="28" captionText="FIGURES 93 – 98. Dorsal habitus of Scydmaenilla. Scydmaenilla (s. str.) constricta Lea, paralectotype female (93), Scydmaenilla (s. str.) queenslandica Franz, holotype male (94), Scydmaenilla (s. str.) thompsoniana, holotype female (95), Scydmaenilla (s. str.) brisbanensis, holotype female (96), Scydmaenilla (s. str.) sydneyana, holotype female (97), Scydmaenilla (Scydmaenillunia) adelaidensis Franz, holotype female (98)." httpUri="https://zenodo.org/record/283485/files/figure.png" pageId="35" pageNumber="74">Figs. 95</figureCitation>
,
<figureCitation id="13442A76B0739A58BC4E461E1227FBEB" box="[264,308,1122,1146]" captionStart="FIGURES 99 – 104" captionStartId="32.[151,264,1887,1910]" captionTargetBox="[200,1391,182,1857]" captionTargetId="figure@32.[185,1399,171,1872]" captionTargetPageId="32" captionText="FIGURES 99 – 104. Ven t er of Scydmaenilla. Scydmaenilla (s. str.) constricta Lea, paralectotype female (99), Scydmaenilla (s. str.) queenslandica Franz, holotype male (100), Scydmaenilla (s. str.) thompsoniana, holotype female (101), Scydmaenilla (s. str.) brisbanensis, holotype female (102), Scydmaenilla (s. str.) sydneyana, holotype female (103), Scydmaenilla (Scydmaenillunia) adelaidensis Franz, holotype female (104). Abbreviations: ai, asetose impression of mesoventrite; lmtc, lateral metaventral carina; pl, posterior lobe of mesocoxal projection; si, setose impression of mesoventrite." httpUri="https://zenodo.org/record/283486/files/figure.png" pageId="35" pageNumber="74">101</figureCitation>
,
<figureCitation id="13442A76B0739A58BC06461E1278FBEB" box="[320,363,1122,1146]" captionStart="FIGURES 105 – 110" captionStartId="34.[151,260,1329,1352]" captionTargetBox="[171,1427,201,1296]" captionTargetId="figure@34.[151,1436,193,1308]" captionTargetPageId="34" captionText="FIGURES 105 – 110. Original sets of labels. Scydmaenilla (s. str.) constricta Lea, lectotype (105), Scydmaenilla (s. str.) queenslandica Franz, holotype (106), Scydmaenilla (s. str.) thompsoniana, holotype (107), Scydmaenilla (s. str.) brisbanensis, holotype (108), Scydmaenilla (s. str.) sydneyana, holotype (109), Scydmaenilla (Scydmaenillunia) adelaidensis Franz, holotype (110)." httpUri="https://zenodo.org/record/283487/files/figure.png" pageId="35" pageNumber="74">107</figureCitation>
)
</paragraph>
<paragraph id="8BC036F3B0739A58BDD146D611EFFB51" blockId="35.[151,764,1194,1216]" box="[151,764,1194,1216]" pageId="35" pageNumber="74">
<taxonomicName id="4C7F4D70B0739A58BDD146D611EBFB51" authority="Franz, 1975: 139" authorityName="Franz" authorityPageNumber="139" authorityYear="1975" box="[151,760,1194,1216]" class="Insecta" family="Staphylinidae" genus="Stenichnus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="35" pageNumber="74" phylum="Arthropoda" rank="species" species="thompsonianus" subGenus="Scydmaenilla">
<emphasis id="B90BEAE1B0739A58BDD146D61217FB51" box="[151,260,1194,1216]" italics="true" pageId="35" pageNumber="74">Stenichnus</emphasis>
(
<emphasis id="B90BEAE1B0739A58BC5446D61289FB51" box="[274,410,1194,1216]" italics="true" pageId="35" pageNumber="74">Scydmaenilla</emphasis>
)
<emphasis id="B90BEAE1B0739A58BCEF46D61151FB51" box="[425,578,1194,1216]" italics="true" pageId="35" pageNumber="74">thompsonianus</emphasis>
Franz, 1975: 139
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C3656578B0739A58BDD1469113FDFAE0" pageId="35" pageNumber="74" type="materials_examined">
<paragraph id="8BC036F3B0739A58BDD1469113FDFAE0" blockId="35.[151,1437,1261,2005]" pageId="35" pageNumber="74">
<emphasis id="B90BEAE1B0739A58BDD1469112B1FA97" bold="true" box="[151,418,1261,1286]" pageId="35" pageNumber="74">
<typeStatus id="54C48851B0739A58BDD1469113C1FA94" box="[151,210,1261,1285]" pageId="35" pageNumber="74">Type</typeStatus>
material studied
</emphasis>
.
<typeStatus id="54C48851B0739A58BCF546911133FA94" box="[435,544,1261,1285]" pageId="35" pageNumber="74" type="holotype">Holotype</typeStatus>
: Ƥ: three labels (
<figureCitation id="13442A76B0739A58BFAF4691105CFA94" box="[745,847,1261,1285]" captionStart="FIGURES 105 – 110" captionStartId="34.[151,260,1329,1352]" captionTargetBox="[171,1427,201,1296]" captionTargetId="figure@34.[151,1436,193,1308]" captionTargetPageId="34" captionText="FIGURES 105 – 110. Original sets of labels. Scydmaenilla (s. str.) constricta Lea, lectotype (105), Scydmaenilla (s. str.) queenslandica Franz, holotype (106), Scydmaenilla (s. str.) thompsoniana, holotype (107), Scydmaenilla (s. str.) brisbanensis, holotype (108), Scydmaenilla (s. str.) sydneyana, holotype (109), Scydmaenilla (Scydmaenillunia) adelaidensis Franz, holotype (110)." httpUri="https://zenodo.org/record/283487/files/figure.png" pageId="35" pageNumber="74">Fig. 107</figureCitation>
): &quot;Umg. Maipoton / Queensld., Austr. / lg.H.Franz 1970&quot; and &quot;Pa 76&quot; on the reverse side [white, printed, reverse handwritten in blue], &quot;
<taxonomicName id="4C7F4D70B0739A58B908476D17D8FAB8" box="[1102,1227,1297,1321]" class="Insecta" family="Staphylinidae" genus="Stenichnus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="35" pageNumber="79" phylum="Arthropoda" rank="genus">Stenichnus</taxonomicName>
/ (
<taxonomicName id="4C7F4D70B0739A58B9AF476D1696FAB8" box="[1257,1413,1297,1321]" class="Insecta" family="Staphylinidae" genus="Scydmaenilla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="35" pageNumber="74" phylum="Arthropoda" rank="genus">Scydmaenilla</taxonomicName>
) /
<taxonomicName id="4C7F4D70B0739A58BDD147491227FADC" box="[151,308,1333,1357]" class="Insecta" family="Staphylinidae" genus="Scydmaenilla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="35" pageNumber="74" phylum="Arthropoda" rank="species" species="thompsoniana">thompsoniana</taxonomicName>
[sic!] m. / det.H.Franz&quot; [white, printed and handwritten in blue], &quot;
<typeStatus id="54C48851B0739A58B96447491778FADC" box="[1058,1131,1333,1357]" pageId="35" pageNumber="74">Typus</typeStatus>
&quot; [red, handwritten in blue] (SAM).
</paragraph>
</subSubSection>
<subSubSection id="C3656578B0739A58BD81470110CDF9FC" pageId="35" pageNumber="74" type="diagnosis">
<paragraph id="8BC036F3B0739A58BD81470111EDF990" blockId="35.[151,1437,1261,2005]" pageId="35" pageNumber="74">
<emphasis id="B90BEAE1B0739A58BD8147011252FA07" bold="true" box="[199,321,1405,1430]" pageId="35" pageNumber="74">Diagnosis.</emphasis>
Body length
<quantity id="4C879B16B0739A58BC9C47011152FA04" box="[474,577,1405,1429]" metricMagnitude="-3" metricUnit="m" metricValue="1.15" pageId="35" pageNumber="74" unit="mm" value="1.15">1.15 mm</quantity>
; elytra stout, EI 1.47; antennal club with slightly asymmetrical antennomere X; metatibiae with short and slender subapical setal brushes; lateral metaventral carinae absent; short median longitudinal groove on posterior pronotal collar present; lateral marginal pronotal carinae barely discernible. Indistinguishable from
<taxonomicName id="4C7F4D70B0739A58BCDA47961149F990" box="[412,602,1513,1537]" class="Insecta" family="Staphylinidae" genus="Scydmaenilla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="35" pageNumber="74" phylum="Arthropoda" rank="species" species="queenslandica">
<emphasis id="B90BEAE1B0739A58BCDA47961149F990" box="[412,602,1513,1537]" italics="true" pageId="35" pageNumber="74">S. queenslandica</emphasis>
</taxonomicName>
; see Remarks.
</paragraph>
<paragraph id="8BC036F3B0739A58BD81447110CDF9FC" blockId="35.[151,1437,1261,2005]" pageId="35" pageNumber="74">
<emphasis id="B90BEAE1B0739A58BD8144711267F9B7" bold="true" box="[199,372,1549,1574]" pageId="35" pageNumber="74">Redescription.</emphasis>
The
<typeStatus id="54C48851B0739A58BCEB4471111CF9B4" box="[429,527,1549,1573]" pageId="35" pageNumber="74" type="holotype">holotype</typeStatus>
female (
<figureCitation id="13442A76B0739A58BF36447111DEF9B4" box="[624,717,1549,1573]" captionStart="FIGURES 93 – 98" captionStartId="28.[151,264,1922,1945]" captionTargetBox="[163,1419,179,1880]" captionTargetId="figure@28.[151,1436,171,1902]" captionTargetPageId="28" captionText="FIGURES 93 – 98. Dorsal habitus of Scydmaenilla. Scydmaenilla (s. str.) constricta Lea, paralectotype female (93), Scydmaenilla (s. str.) queenslandica Franz, holotype male (94), Scydmaenilla (s. str.) thompsoniana, holotype female (95), Scydmaenilla (s. str.) brisbanensis, holotype female (96), Scydmaenilla (s. str.) sydneyana, holotype female (97), Scydmaenilla (Scydmaenillunia) adelaidensis Franz, holotype female (98)." httpUri="https://zenodo.org/record/283485/files/figure.png" pageId="35" pageNumber="74">Figs. 95</figureCitation>
,
<figureCitation id="13442A76B0739A58BF9E44711010F9B4" box="[728,771,1549,1573]" captionStart="FIGURES 99 – 104" captionStartId="32.[151,264,1887,1910]" captionTargetBox="[200,1391,182,1857]" captionTargetId="figure@32.[185,1399,171,1872]" captionTargetPageId="32" captionText="FIGURES 99 – 104. Ven t er of Scydmaenilla. Scydmaenilla (s. str.) constricta Lea, paralectotype female (99), Scydmaenilla (s. str.) queenslandica Franz, holotype male (100), Scydmaenilla (s. str.) thompsoniana, holotype female (101), Scydmaenilla (s. str.) brisbanensis, holotype female (102), Scydmaenilla (s. str.) sydneyana, holotype female (103), Scydmaenilla (Scydmaenillunia) adelaidensis Franz, holotype female (104). Abbreviations: ai, asetose impression of mesoventrite; lmtc, lateral metaventral carina; pl, posterior lobe of mesocoxal projection; si, setose impression of mesoventrite." httpUri="https://zenodo.org/record/283486/files/figure.png" pageId="35" pageNumber="74">101</figureCitation>
) differs from the
<typeStatus id="54C48851B0739A58BE814471173AF9B4" box="[967,1065,1549,1573]" pageId="35" pageNumber="74" type="holotype">holotype</typeStatus>
male of
<taxonomicName id="4C7F4D70B0739A58B9CC44721654F9B4" box="[1162,1351,1549,1573]" class="Insecta" family="Staphylinidae" genus="Scydmaenilla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="35" pageNumber="74" phylum="Arthropoda" rank="species" species="queenslandica">
<emphasis id="B90BEAE1B0739A58B9CC44721654F9B4" box="[1162,1351,1549,1573]" italics="true" pageId="35" pageNumber="74">S. queenslandica</emphasis>
</taxonomicName>
only in more lightly pigmented body and slightly different measurements: BL
<quantity id="4C879B16B0739A58BEF6444D1704F9D8" box="[944,1047,1585,1609]" metricMagnitude="-3" metricUnit="m" metricValue="1.15" pageId="35" pageNumber="74" unit="mm" value="1.15">1.15 mm</quantity>
; HL
<quantity id="4C879B16B0739A58B909444D17A5F9D8" box="[1103,1206,1585,1609]" metricMagnitude="-4" metricUnit="m" metricValue="2.0" pageId="35" pageNumber="74" unit="mm" value="0.2">0.20 mm</quantity>
, HW
<quantity id="4C879B16B0739A58B9B1444D164EF9D8" box="[1271,1373,1585,1609]" metricMagnitude="-4" metricUnit="m" metricValue="2.3" pageId="35" pageNumber="74" unit="mm" value="0.23">0.23 mm</quantity>
, AnL
<quantity id="4C879B16B0739A58BDD1442913EDF9FC" box="[151,254,1621,1645]" metricMagnitude="-4" metricUnit="m" metricValue="3.3" pageId="35" pageNumber="74" unit="mm" value="0.33">0.33 mm</quantity>
; PL
<quantity id="4C879B16B0739A58BC7744291284F9FC" box="[305,407,1621,1645]" metricMagnitude="-4" metricUnit="m" metricValue="3.3" pageId="35" pageNumber="74" unit="mm" value="0.33">0.33 mm</quantity>
, PW
<quantity id="4C879B16B0739A58BC9544291129F9FC" box="[467,570,1621,1645]" metricMagnitude="-4" metricUnit="m" metricValue="2.9" pageId="35" pageNumber="74" unit="mm" value="0.29">0.29 mm</quantity>
; EL
<quantity id="4C879B16B0739A58BF29442911C6F9FC" box="[623,725,1621,1645]" metricMagnitude="-4" metricUnit="m" metricValue="6.3" pageId="35" pageNumber="74" unit="mm" value="0.63">0.63 mm</quantity>
, EW
<quantity id="4C879B16B0739A58BE5544291069F9FC" box="[787,890,1621,1645]" metricMagnitude="-4" metricUnit="m" metricValue="4.3" pageId="35" pageNumber="74" unit="mm" value="0.43">0.43 mm</quantity>
, EI 1.47.
</paragraph>
</subSubSection>
<subSubSection id="C3656578B0739A58BD814406129BF900" box="[199,392,1657,1682]" pageId="35" pageNumber="74" type="description">
<paragraph id="8BC036F3B0739A58BD814406129BF900" blockId="35.[151,1437,1261,2005]" box="[199,392,1657,1682]" pageId="35" pageNumber="74">
<emphasis id="B90BEAE1B0739A58BD8144061217F903" bold="true" box="[199,260,1658,1682]" pageId="35" pageNumber="74">Male</emphasis>
. Unknown.
</paragraph>
</subSubSection>
<subSubSection id="C3656578B0739A58BD8144E11045F924" box="[199,854,1693,1718]" pageId="35" pageNumber="74" type="distribution">
<paragraph id="8BC036F3B0739A58BD8144E11045F924" blockId="35.[151,1437,1261,2005]" box="[199,854,1693,1718]" pageId="35" pageNumber="74">
<emphasis id="B90BEAE1B0739A58BD8144E11273F927" bold="true" box="[199,352,1693,1718]" pageId="35" pageNumber="74">Distribution.</emphasis>
Eastern
<collectingCountry id="F3687663B0739A58BC8444E1113EF924" box="[450,557,1693,1717]" name="Australia" pageId="35" pageNumber="74">Australia</collectingCountry>
: southeastern Queensland.
</paragraph>
</subSubSection>
<subSubSection id="C3656578B0739A58BD8144BE165FF844" pageId="35" pageNumber="74" type="discussion">
<paragraph id="8BC036F3B0739A58BD8144BE165FF844" blockId="35.[151,1437,1261,2005]" pageId="35" pageNumber="74">
<emphasis id="B90BEAE1B0739A58BD8144BE1228F94B" bold="true" box="[199,315,1730,1754]" pageId="35" pageNumber="74">Remarks.</emphasis>
Franz (1975) described
<taxonomicName id="4C7F4D70B0739A58BF1244BE1003F948" box="[596,784,1729,1753]" class="Insecta" family="Staphylinidae" genus="Scydmaenilla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="35" pageNumber="74" phylum="Arthropoda" rank="species" species="thompsoniana">
<emphasis id="B90BEAE1B0739A58BF1244BE1003F948" box="[596,784,1729,1753]" italics="true" pageId="35" pageNumber="74">S. thompsoniana</emphasis>
</taxonomicName>
on the basis of a single female, and
<taxonomicName id="4C7F4D70B0739A58B9F144BE1664F948" box="[1207,1399,1729,1753]" class="Insecta" family="Staphylinidae" genus="Scydmaenilla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="35" pageNumber="74" phylum="Arthropoda" rank="species" species="queenslandica">
<emphasis id="B90BEAE1B0739A58B9F144BE1664F948" box="[1207,1399,1729,1753]" italics="true" pageId="35" pageNumber="74">S. queenslandica</emphasis>
</taxonomicName>
on the basis of a single male, both collected in the same locality. In the identification key for
<taxonomicName id="4C7F4D70B0739A58B9F24499168FF96C" authority="Franz" authorityName="Franz" box="[1204,1436,1765,1789]" class="Insecta" family="Staphylinidae" genus="Scydmaenilla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="35" pageNumber="74" phylum="Arthropoda" rank="genus">
<emphasis id="B90BEAE1B0739A58B9F244991658F96C" box="[1204,1355,1765,1789]" italics="true" pageId="35" pageNumber="74">Scydmaenilla</emphasis>
, Franz
</taxonomicName>
(1975; p. 140–141) differentiated these species on the basis of the proportions of the pronotum, body shape and elytral punctation. However, examination of the
<typeStatus id="54C48851B0739A58BF82455211E7F8D7" box="[708,756,1838,1862]" pageId="35" pageNumber="74">type</typeStatus>
specimens revealed that they have nearly identical PL and PW, share a similar body shape, and the difference in the elytral punctation is barely noticeable. The
<typeStatus id="54C48851B0739A58B846452D1671F8F8" box="[1280,1378,1873,1897]" pageId="35" pageNumber="74" type="holotype">holotype</typeStatus>
of
<taxonomicName id="4C7F4D70B0739A58B8C1452E1225F81C" class="Insecta" family="Staphylinidae" genus="Scydmaenilla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="35" pageNumber="74" phylum="Arthropoda" rank="species" species="thompsoniana">
<emphasis id="B90BEAE1B0739A58B8C1452E1225F81C" italics="true" pageId="35" pageNumber="74">S. thompsoniana</emphasis>
</taxonomicName>
is lighter in color and has slightly narrower head. The observed differences can be attributed to a sexual dimorphism or to inter-individual variation within species. However, I prefer not to synonymize these names until a larger sample has become available and the morphological variability has been better studied.
</paragraph>
</subSubSection>
</treatment>
</document>
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