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<document id="8C193F2A5E53F1AE3379F7E01D07A47A" ID="10.11646/zootaxa.4269.4.11SLASH11137" ID-CLB-Dataset="32978" ID-DOI="10.11646/zootaxa.4269.4.11" ID-GBIF-Dataset="2a914627-2a79-491c-a3b0-d3977ea56e9d" ID-ISSN="1175-5326" ID-Zenodo-Dep="582809" ID-ZooBank="459BFE80-A87B-4922-8C16-FC3BA437704C" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1495616729728" checkinUser="plazi" docAuthor="Lo, Yik-Fui Philip, Li, Fei &amp; Ding, Liang" docDate="2017" docId="03D687C7FF98DD25FF7CFCCCFAE7BB6C" docLanguage="en" docName="zootaxa.4269.4.11SLASH11137.pdf" docOrigin="Zootaxa 4269 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Talicada nyseus subsp. lami Lo, Li &amp; Ding, subsp. nov." docType="treatment" docVersion="7" lastPageNumber="589" masterDocId="FFEFFFBFFF99DD26FFEBFF8DFF9FBE1D" masterDocTitle="Description of a new subspecies of Talicada nyseus (Guérin-Méneville, 1843) from Hainan, China (Lepidoptera: Lycaenidae), with notes on the genus Talicada Moore, 1881" masterLastPageNumber="592" masterPageNumber="586" pageNumber="587" updateTime="1698439884378" updateUser="ExternalLinkService">
<mods:mods id="20A576EE5863CB6F83CB4ED159470F4E" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="5F350EE456E4CA816ADF077F16AE9722">
<mods:title id="D4CF50F2FF0F90C72F1078977103D290">Description of a new subspecies of Talicada nyseus (Guérin-Méneville, 1843) from Hainan, China (Lepidoptera: Lycaenidae), with notes on the genus Talicada Moore, 1881</mods:title>
</mods:titleInfo>
<mods:name id="725782BFE8DFFCF76FFD24F115AB802C" type="personal">
<mods:role id="33541936572718B181CCC3F916CE4184">
<mods:roleTerm id="FF0AA1DF4E5A531E1638CD1C8080E57B">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="10E091220D63B7261D7B5BDB657A88D5">Lo, Yik-Fui Philip</mods:namePart>
</mods:name>
<mods:name id="21423BC051F3D59F39648D65A2D930D6" type="personal">
<mods:role id="BC700106C01B0D12552CEBF92CEA493A">
<mods:roleTerm id="B0EE128B3B29B1E3F4DD175D51109BF9">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="43865577FC0D5B08975E9D2FFB29EEA2">Li, Fei</mods:namePart>
</mods:name>
<mods:name id="E30E90ACB7CF4095166ADEE5B71A6E29" type="personal">
<mods:role id="6A627A5136C72ACC4821F6B5D9627782">
<mods:roleTerm id="9B4B3E8C53AEE97545DCC7C6F3CF1869">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="613BC87577B5ACAFA47CC9547DC3D926">Ding, Liang</mods:namePart>
</mods:name>
<mods:typeOfResource id="97166D8C4F3DE17D9A73A7BB619EFD38">text</mods:typeOfResource>
<mods:relatedItem id="7F7E9B3CA0D365BE88138987F33C6E26" type="host">
<mods:titleInfo id="DB47CD31E68E005B970EA1F4ADA0F0F2">
<mods:title id="8E16629788030BCB2E16BD97F82B1BE4">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="0D48DF7C59502A45C615A1031E14BFB1">
<mods:date id="69AFC6EB5208AC41466E64A894E41C23">2017</mods:date>
<mods:detail id="A0B8818444FC0B166E2E7449A93A45D6" type="volume">
<mods:number id="0605B39CC0227AD6F8357BBAAB1F97B0">4269</mods:number>
</mods:detail>
<mods:detail id="029DFD3422261FC718B155CFBD95E817" type="issue">
<mods:number id="1B305D4A6154E507AA6F059F7F4C7F20">4</mods:number>
</mods:detail>
<mods:extent id="2B14594EF5F34B3CE2E7DD94DC8E7829" unit="page">
<mods:start id="05FB98CDD844C76FAC043AE17CCF4EA3">586</mods:start>
<mods:end id="3E87DC9388AC735ED304A7536881D2FC">592</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="A43A17D6A6CBE1F01400C6A4B610A9EA">journal article</mods:classification>
<mods:identifier id="2766F928B280E393A9EA2FB94724C58B" type="CLB-Dataset">32978</mods:identifier>
<mods:identifier id="3623604D3020E7B0731A3576E188FD1D" type="DOI">10.11646/zootaxa.4269.4.11</mods:identifier>
<mods:identifier id="5165152A8F5F7181EAB4AE103961053E" type="GBIF-Dataset">2a914627-2a79-491c-a3b0-d3977ea56e9d</mods:identifier>
<mods:identifier id="8DB59366E2ABC7E92F7EFB11D4968FEF" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="EF3DE57DAB01A12A10FBEF60098148E2" type="Zenodo-Dep">582809</mods:identifier>
<mods:identifier id="1512595DBA790C8EA3D3B419BD52B7B9" type="ZooBank">459BFE80-A87B-4922-8C16-FC3BA437704C</mods:identifier>
</mods:mods>
<treatment id="03D687C7FF98DD25FF7CFCCCFAE7BB6C" ID-DOI="http://doi.org/10.5281/zenodo.6032776" ID-GBIF-Taxon="130844729" ID-Zenodo-Dep="6032776" LSID="urn:lsid:plazi:treatment:03D687C7FF98DD25FF7CFCCCFAE7BB6C" httpUri="http://treatment.plazi.org/id/03D687C7FF98DD25FF7CFCCCFAE7BB6C" lastPageId="3" lastPageNumber="589" pageId="1" pageNumber="587">
<subSubSection id="C365655AFF98DD27FF7CFCCCFD43BD46" box="[151,732,833,859]" pageId="1" pageNumber="587" type="nomenclature">
<paragraph id="8BC036D1FF98DD27FF7CFCCCFD43BD46" blockId="1.[151,732,833,859]" box="[151,732,833,859]" pageId="1" pageNumber="587">
<heading id="D08881BDFF98DD27FF7CFCCCFD43BD46" bold="true" box="[151,732,833,859]" fontSize="11" level="1" pageId="1" pageNumber="587" reason="1">
<emphasis id="B90BEAC3FF98DD27FF7CFCCCFD43BD46" bold="true" box="[151,732,833,859]" pageId="1" pageNumber="587">
<taxonomicName id="4C7F4D52FF98DD27FF7CFCCCFDD4BD46" authority="Lo, Li &amp; Ding" authorityName="Lo, Li &amp; Ding" box="[151,587,833,859]" class="Insecta" family="Lycaenidae" genus="Talicada" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="587" phylum="Arthropoda" rank="subSpecies" species="nyseus" status="subsp. nov." subSpecies="lami">
<emphasis id="B90BEAC3FF98DD27FF7CFCCCFE0CBD46" bold="true" box="[151,403,833,859]" italics="true" pageId="1" pageNumber="587">Talicada nyseus lami</emphasis>
Lo, Li &amp; Ding
</taxonomicName>
<taxonomicNameLabel id="A23857B8FF98DD27FDB9FCCCFD43BD46" box="[594,732,833,859]" pageId="1" pageNumber="587" rank="subSpecies">subsp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C365655AFF98DD27FF7CFC0AFCAFB9DB" pageId="1" pageNumber="587" type="materials_examined">
<paragraph id="8BC036D1FF98DD27FF7CFC0AFE16BB31" blockId="1.[151,1437,903,1990]" pageId="1" pageNumber="587">
<materialsCitation id="3B173C8CFF98DD27FF7CFC0AFC34BDD8" ID-GBIF-Occurrence="1503738100" collectingDate="2007-12-15" collectingDateMax="2007-12-16" collectingDateMin="2007-12-15" collectorName="Coll. Y. F." country="China" county="Baisha County" location="Gaofeng River" municipality="Yinggeling" pageId="1" pageNumber="587" specimenCount="1" specimenCount-male="1" stateProvince="Hainan" typeStatus="holotype">
<emphasis id="B90BEAC3FF98DD27FF7CFC0AFED6BDBD" bold="true" box="[151,329,903,928]" pageId="1" pageNumber="587">
<typeStatus id="54C48873FF98DD27FF7CFC0AFF4DBDBD" box="[151,210,903,928]" pageId="1" pageNumber="587">Type</typeStatus>
material.
</emphasis>
<typeStatus id="54C48873FF98DD27FEBFFC05FE73BDBD" box="[340,492,904,928]" pageId="1" pageNumber="587" type="holotype">HOLOTYPE</typeStatus>
:
<specimenCount id="9D79FD58FF98DD27FE11FC05FD8EBD82" box="[506,529,904,927]" pageId="1" pageNumber="587" type="male">♂</specimenCount>
,
<collectingCountry id="F3687641FF98DD27FDCAFC0AFDF7BDBD" box="[545,616,903,928]" name="China" pageId="1" pageNumber="587">China</collectingCountry>
:
<collectingRegion id="49BBF833FF98DD27FD9CFC0AFD57BDBD" box="[631,712,903,928]" country="China" name="Hainan" pageId="1" pageNumber="587">Hainan</collectingRegion>
Island,
<collectingCounty id="62A14E5DFF98DD27FCC2FC0AFC4CBDBD" box="[809,979,903,928]" pageId="1" pageNumber="587">Baisha County</collectingCounty>
,
<collectingMunicipality id="6BA4ACABFF98DD27FC08FC0AFBFFBDBD" box="[995,1120,903,928]" pageId="1" pageNumber="587">Yinggeling</collectingMunicipality>
NNR,
<location id="8EA0600AFF98DD27FB53FC0AFAFCBDBD" LSID="urn:lsid:plazi:treatment:03D687C7FF98DD25FF7CFCCCFAE7BB6C:8EA0600AFF98DD27FB53FC0AFAFCBDBD" box="[1208,1379,903,928]" country="China" county="Baisha County" municipality="Yinggeling" name="Gaofeng River" pageId="1" pageNumber="587" stateProvince="Hainan">Gaofeng River</location>
,
<collectingDate id="EF85E9F9FF98DD27FA99FC0AFEBCBDD8" pageId="1" pageNumber="587" value="2007-12-15" valueMax="2007-12-16" valueMin="2007-12-15">
<date id="FFC11011FF98DD27FA99FC0AFEBCBDD8" pageId="1" pageNumber="587" value="2007-12-15" valueMax="2007-12-16" valueMin="2007-12-15">15– 16.XII.2007</date>
</collectingDate>
,
<collectorName id="268A5307FF98DD27FED9FC21FE33BDD9" box="[306,428,940,965]" pageId="1" pageNumber="587">Coll. Y. F.</collectorName>
Lo, pupa
<collectedFrom id="3863752CFF98DD27FDC1FC20FCC8BDD9" box="[554,855,940,965]" pageId="1" pageNumber="587">
on
<taxonomicName id="4C7F4D52FF98DD27FDBAFC21FCC8BDD9" box="[593,855,940,965]" class="Gymnostomatea" family="Spathidiidae" genus="Bryophyllum" kingdom="Plantae" order="Spathidiida" pageId="1" pageNumber="587" phylum="Ciliophora" rank="species" species="pinnatum">
<emphasis id="B90BEAC3FF98DD27FDBAFC21FCC8BDD9" box="[593,855,940,965]" italics="true" pageId="1" pageNumber="587">Bryophyllum pinnatum</emphasis>
</taxonomicName>
</collectedFrom>
, emgd
</materialsCitation>
.
<date id="FFC11011FF98DD27FC51FC21FBDCBDD9" box="[954,1091,940,964]" pageId="1" pageNumber="587" value="2007-12-18">18.XII.2007</date>
(IOZ). PARATYPES: 2♂1♀, locality and date as holotype, Coll. Y. F. Lo (IOZ, YFLC) (1♂ dissected, genitalia YFL ly0015). 4♂3♀, locality and date as holotype, Coll. Y. F. Lo, reared from
<taxonomicName id="4C7F4D52FF98DD27FD29FC79FC0BBA11" box="[706,916,1012,1037]" class="Magnoliopsida" family="Crassulaceae" genus="Kalanchoe" kingdom="Plantae" order="Saxifragales" pageId="1" pageNumber="587" phylum="Tracheophyta" rank="species" species="integra">
<emphasis id="B90BEAC3FF98DD27FD29FC79FC0BBA11" box="[706,916,1012,1037]" italics="true" pageId="1" pageNumber="587">Kalanchoe integra</emphasis>
</taxonomicName>
, emgd.
<date id="FFC11011FF98DD27FC04FC79FB1ABA10" box="[1007,1157,1012,1037]" pageId="1" pageNumber="587" value="2008-01-09" valueMax="2008-01-15" valueMin="2008-01-09">09–15.I.2008</date>
(YFLC, KFBG). 2♂1♀, locality and date as holotype, Coll. Y. F. Lo, reared from
<taxonomicName id="4C7F4D52FF98DD27FCBEFB9AFBC2BA2D" box="[853,1117,1047,1072]" class="Gymnostomatea" family="Spathidiidae" genus="Bryophyllum" kingdom="Plantae" order="Spathidiida" pageId="1" pageNumber="587" phylum="Ciliophora" rank="species" species="pinnatum">
<emphasis id="B90BEAC3FF98DD27FCBEFB9AFBC2BA2D" box="[853,1117,1047,1072]" italics="true" pageId="1" pageNumber="587">Bryophyllum pinnatum</emphasis>
</taxonomicName>
, emgd.
<date id="FFC11011FF98DD27FB28FB9AFAA8BA2D" box="[1219,1335,1047,1072]" pageId="1" pageNumber="587" value="2008-02-16">16.II.2008</date>
(YFLC, BMNH). 3♂, locality as holotype,
<date id="FFC11011FF98DD27FDD7FBB1FD43BA48" box="[572,732,1084,1109]" pageId="1" pageNumber="587" value="2008-05-14" valueMax="2008-05-16" valueMin="2008-05-14">14–16.V.2008</date>
, Coll. Y. F. Lo (YFLC, NTNU). 1♂, locality as holotype,
<date id="FFC11011FF98DD27FF7CFBEDFE9CBA65" box="[151,259,1120,1144]" pageId="1" pageNumber="587" value="2009-01-11">11.I.2009</date>
, Coll. Y. F. Lo (YFLC).
<materialsCitation id="3B173C8CFF98DD27FDC8FBEDFDBDBA81" ID-GBIF-Occurrence="1503738099" collectingDate="2009-04-26" collectionCode="YFLC" collectorName="Coll. Y. F." country="China" county="Changjiang County" location="Huangdi Cave" municipality="Nanyao River" pageId="1" pageNumber="587" specimenCount="1" specimenCount-male="1" stateProvince="Hainan" typeStatus="holotype">
<specimenCount id="9D79FD58FF98DD27FDC8FBEDFDD8BA65" box="[547,583,1120,1144]" pageId="1" pageNumber="587" type="male">1♂</specimenCount>
,
<collectingCountry id="F3687641FF98DD27FDBDFBD2FD02BA65" box="[598,669,1119,1144]" name="China" pageId="1" pageNumber="587">China</collectingCountry>
:
<collectingRegion id="49BBF833FF98DD27FD41FBD2FD64BA65" box="[682,763,1119,1144]" country="China" name="Hainan" pageId="1" pageNumber="587">Hainan</collectingRegion>
Island,
<collectingCounty id="62A14E5DFF98DD27FCB2FBD2FBA8BA65" box="[857,1079,1119,1144]" pageId="1" pageNumber="587">Changjiang County</collectingCounty>
,
<collectingMunicipality id="6BA4ACABFF98DD27FBAFFBEDFB7ABA65" box="[1092,1253,1119,1144]" pageId="1" pageNumber="587">Nanyao River</collectingMunicipality>
,
<location id="8EA0600AFF98DD27FB19FBD2FA07BA65" LSID="urn:lsid:plazi:treatment:03D687C7FF98DD25FF7CFCCCFAE7BB6C:8EA0600AFF98DD27FB19FBD2FA07BA65" box="[1266,1432,1119,1144]" country="China" county="Changjiang County" municipality="Nanyao River" name="Huangdi Cave" pageId="1" pageNumber="587" stateProvince="Hainan">Huangdi Cave</location>
,
<collectingDate id="EF85E9F9FF98DD27FF7CFB09FE89BA80" box="[151,278,1156,1181]" pageId="1" pageNumber="587" value="2009-04-26">
<date id="FFC11011FF98DD27FF7CFB09FE89BA80" box="[151,278,1156,1181]" pageId="1" pageNumber="587" value="2009-04-26">26.IV.2009</date>
</collectingDate>
,
<collectorName id="268A5307FF98DD27FEC9FB09FE09BA81" box="[290,406,1156,1181]" pageId="1" pageNumber="587">Coll. Y. F.</collectorName>
Lo (
<collectionCode id="ED6EAE14FF98DD27FE27FB09FD85BA81" box="[460,538,1156,1180]" pageId="1" pageNumber="587">YFLC</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B173C8CFF98DD27FDC5FB09FDB4BADD" ID-GBIF-Occurrence="1503738101" collectingDate="2009-06-06" collectionCode="YFLC" collectorName="Coll. Y. F." country="China" county="Changjiang County" location="Huangdi Cave" municipality="Nanyao River" pageId="1" pageNumber="587" specimenCount="2" specimenCount-male="2" stateProvince="Hainan" typeStatus="holotype">
<specimenCount id="9D79FD58FF98DD27FDC5FB09FDCCBA81" box="[558,595,1156,1180]" pageId="1" pageNumber="587" type="male">2♂</specimenCount>
,
<collectingCountry id="F3687641FF98DD27FD8BFB09FD38BA80" box="[608,679,1156,1181]" name="China" pageId="1" pageNumber="587">China</collectingCountry>
:
<collectingRegion id="49BBF833FF98DD27FD59FB09FC9CBA80" box="[690,771,1156,1181]" country="China" name="Hainan" pageId="1" pageNumber="587">Hainan</collectingRegion>
Island,
<collectingCounty id="62A14E5DFF98DD27FCB4FB09FBA4BA81" box="[863,1083,1156,1181]" pageId="1" pageNumber="587">Changjiang County</collectingCounty>
,
<collectingMunicipality id="6BA4ACABFF98DD27FBA3FB08FB78BA80" box="[1096,1255,1156,1181]" pageId="1" pageNumber="587">Nanyao River</collectingMunicipality>
,
<location id="8EA0600AFF98DD27FB18FB09FA07BA81" LSID="urn:lsid:plazi:treatment:03D687C7FF98DD25FF7CFCCCFAE7BB6C:8EA0600AFF98DD27FB18FB09FA07BA81" box="[1267,1432,1156,1181]" country="China" county="Changjiang County" municipality="Nanyao River" name="Huangdi Cave" pageId="1" pageNumber="587" stateProvince="Hainan">Huangdi Cave</location>
,
<collectingDate id="EF85E9F9FF98DD27FF7CFB2AFE86BADD" box="[151,281,1191,1216]" pageId="1" pageNumber="587" value="2009-06-06">
<date id="FFC11011FF98DD27FF7CFB2AFE86BADD" box="[151,281,1191,1216]" pageId="1" pageNumber="587" value="2009-06-06">06.VI.2009</date>
</collectingDate>
,
<collectorName id="268A5307FF98DD27FECDFB2AFE03BAA2" box="[294,412,1191,1216]" pageId="1" pageNumber="587">Coll. Y. F.</collectorName>
Lo (
<collectionCode id="ED6EAE14FF98DD27FE3EFB25FDBCBADD" box="[469,547,1192,1216]" pageId="1" pageNumber="587">YFLC</collectionCode>
)
</materialsCitation>
. 1♂1♀, locality as holotype,
<date id="FFC11011FF98DD27FC95FB25FB9FBADD" box="[894,1024,1192,1216]" pageId="1" pageNumber="587" value="2010-09-03">03.IX.2010</date>
, Coll. F. Li, reared from
<taxonomicName id="4C7F4D52FF98DD27FAC8FB2AFF77BAF9" class="Magnoliopsida" family="Crassulaceae" genus="Kalanchoe" kingdom="Plantae" order="Saxifragales" pageId="1" pageNumber="587" phylum="Tracheophyta" rank="species" species="integra">
<emphasis id="B90BEAC3FF98DD27FAC8FB2AFF77BAF9" italics="true" pageId="1" pageNumber="587">Kalanchoe integra</emphasis>
</taxonomicName>
, emgd.
<date id="FFC11011FF98DD27FEA9FB41FE00BAF9" box="[322,415,1228,1252]" pageId="1" pageNumber="587" value="2010-11">XI.2010</date>
(YFLC).
<materialsCitation id="3B173C8CFF98DD27FDE5FB41FF6ABB15" ID-GBIF-Occurrence="1503738102" collectingDate="2012-01-14" collectionCode="YFLC" collectorName="Coll. Y. F." country="China" county="Dongfeng County" location="Exianling" pageId="1" pageNumber="587" specimenCount="1" specimenCount-male="1" stateProvince="Hainan" typeStatus="holotype">
<specimenCount id="9D79FD58FF98DD27FDE5FB41FDADBAF9" box="[526,562,1228,1252]" pageId="1" pageNumber="587" type="male">1♂</specimenCount>
,
<collectingCountry id="F3687641FF98DD27FDD4FB41FD19BAF8" box="[575,646,1228,1253]" name="China" pageId="1" pageNumber="587">China</collectingCountry>
:
<collectingRegion id="49BBF833FF98DD27FD7BFB41FD7EBAF8" box="[656,737,1228,1253]" country="China" name="Hainan" pageId="1" pageNumber="587">Hainan</collectingRegion>
Island,
<collectingCounty id="62A14E5DFF98DD27FCD0FB41FB99BAF9" box="[827,1030,1228,1253]" pageId="1" pageNumber="587">Dongfeng County</collectingCounty>
,
<location id="8EA0600AFF98DD27FBFAFB41FB1EBAF8" LSID="urn:lsid:plazi:treatment:03D687C7FF98DD25FF7CFCCCFAE7BB6C:8EA0600AFF98DD27FBFAFB41FB1EBAF8" box="[1041,1153,1228,1253]" country="China" county="Dongfeng County" name="Exianling" pageId="1" pageNumber="587" stateProvince="Hainan">Exianling</location>
,
<collectingDate id="EF85E9F9FF98DD27FB60FB41FB66BAF9" box="[1163,1273,1228,1252]" pageId="1" pageNumber="587" value="2012-01-14">
<date id="FFC11011FF98DD27FB60FB41FB66BAF9" box="[1163,1273,1228,1252]" pageId="1" pageNumber="587" value="2012-01-14">14.I.2012</date>
</collectingDate>
,
<collectorName id="268A5307FF98DD27FAEEFB41FAE8BAF9" box="[1285,1399,1228,1253]" pageId="1" pageNumber="587">Coll. Y. F.</collectorName>
Lo (
<collectionCode id="ED6EAE14FF98DD27FF74FB7DFF72BB15" box="[159,237,1264,1288]" pageId="1" pageNumber="587">YFLC</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B173C8CFF98DD27FEEBFB7DFE1ABB31" ID-GBIF-Occurrence="1503738098" collectingDate="2013-11-25" collectionCode="YFLC, SYSU" collectorName="Coll. Y. F. &amp; F. Li" country="China" county="Changjiang County" location="Donger" municipality="Bawangling" pageId="1" pageNumber="587" specimenCount="3" specimenCount-male="3" stateProvince="Hainan" typeStatus="holotype">
<specimenCount id="9D79FD58FF98DD27FEEBFB7DFEBABB15" box="[256,293,1264,1288]" pageId="1" pageNumber="587" type="male">3♂</specimenCount>
,
<collectingCountry id="F3687641FF98DD27FED9FB62FEE6BB15" box="[306,377,1263,1288]" name="China" pageId="1" pageNumber="587">China</collectingCountry>
:
<collectingRegion id="49BBF833FF98DD27FE6EFB62FE49BB15" box="[389,470,1263,1288]" country="China" name="Hainan" pageId="1" pageNumber="587">Hainan</collectingRegion>
Island,
<collectingCounty id="62A14E5DFF98DD27FDDAFB62FC91BB15" box="[561,782,1263,1288]" pageId="1" pageNumber="587">Changjiang County</collectingCounty>
,
<collectingMunicipality id="6BA4ACABFF98DD27FCF1FB62FC3EBB15" box="[794,929,1263,1288]" pageId="1" pageNumber="587">Bawangling</collectingMunicipality>
NNR,
<location id="8EA0600AFF98DD27FC18FB7DFBD6BB15" LSID="urn:lsid:plazi:treatment:03D687C7FF98DD25FF7CFCCCFAE7BB6C:8EA0600AFF98DD27FC18FB7DFBD6BB15" box="[1011,1097,1264,1288]" country="China" county="Changjiang County" municipality="Bawangling" name="Donger" pageId="1" pageNumber="587" stateProvince="Hainan">Donger</location>
,
<collectingDate id="EF85E9F9FF98DD27FBBEFB62FB47BB15" box="[1109,1240,1263,1288]" pageId="1" pageNumber="587" value="2013-11-25">
<date id="FFC11011FF98DD27FBBEFB62FB47BB15" box="[1109,1240,1263,1288]" pageId="1" pageNumber="587" value="2013-11-25">25.XI.2013</date>
</collectingDate>
,
<collectorName id="268A5307FF98DD27FB0FFB62FAC7BB1A" box="[1252,1368,1263,1288]" pageId="1" pageNumber="587">Coll. Y. F.</collectorName>
Lo &amp;
<collectorName id="268A5307FF98DD27FF7CFA98FF54BB30" box="[151,203,1300,1325]" pageId="1" pageNumber="587">F. Li</collectorName>
(
<collectionCode id="ED6EAE14FF98DD27FF31FA99FEB9BB31" box="[218,294,1300,1324]" pageId="1" pageNumber="587">YFLC</collectionCode>
,
<collectionCode id="ED6EAE14FF98DD27FEDAFA99FEE2BB31" box="[305,381,1300,1324]" country="Taiwan" httpUri="http://biocol.org/urn:lsid:biocol.org:col:13728" name="National Sun Yat-Sen University, Department of Biological Sciences" pageId="1" pageNumber="587">SYSU</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="8BC036D1FF98DD27FF2CFABAFB50B960" blockId="1.[151,1437,903,1990]" pageId="1" pageNumber="587">
Male (
<figureCitation id="13442A54FF98DD27FEE4FABAFEE3BB4D" box="[271,380,1335,1360]" captionStart="FIGURES 1 – 4" captionStartId="2.[151,264,1184,1207]" captionTargetBox="[150,1436,195,1160]" captionTargetId="figure@2.[150,1436,195,1160]" captionTargetPageId="2" captionText="FIGURES 1 – 4. Talicada nyseus lami subsp. n .. 1, holotype ♂, upperside; 2, ditto, underside; 3, paratype ♀, upperside; 4, ditto, underside. (scale bar = 10 mm)" httpUri="https://zenodo.org/record/582810/files/figure.png" pageId="1" pageNumber="587">Figs. 1–2</figureCitation>
): Length of antenna
<quantity id="4C879B34FF98DD27FD8AFABAFD7FBB4D" box="[609,736,1335,1360]" metricMagnitude="-3" metricUnit="m" metricValue="7.2" metricValueMax="7.7" metricValueMin="6.7" pageId="1" pageNumber="587" unit="mm" value="7.2" valueMax="7.7" valueMin="6.7">6.7–7.7mm</quantity>
(
<quantity id="4C879B34FF98DD27FD05FAB5FCA1BB4D" box="[750,830,1336,1360]" metricMagnitude="-3" metricUnit="m" metricValue="7.2" pageId="1" pageNumber="587" unit="mm" value="7.2">7.2mm</quantity>
±
<quantity id="4C879B34FF98DD27FCB0FAB5FC32BB4D" box="[859,941,1336,1360]" metricMagnitude="-4" metricUnit="m" metricValue="3.0" pageId="1" pageNumber="587" unit="mm" value="0.3">0.3mm</quantity>
, n=17). Wingspan
<quantity id="4C879B34FF98DD27FB95FAB5FA86BB4D" box="[1150,1305,1336,1360]" metricMagnitude="-2" metricUnit="m" metricValue="2.5799999999999996" metricValueMax="2.77" metricValueMin="2.3899999999999997" pageId="1" pageNumber="587" unit="mm" value="25.799999999999997" valueMax="27.7" valueMin="23.9">23.9–27.7mm</quantity>
(
<quantity id="4C879B34FF98DD27FAC3FABAFA19BB4D" box="[1320,1414,1335,1360]" metricMagnitude="-2" metricUnit="m" metricValue="2.58" pageId="1" pageNumber="587" unit="mm" value="25.8">25.8mm</quantity>
±
<quantity id="4C879B34FF98DD27FF7CFAD1FF77BB69" box="[151,232,1372,1396]" metricMagnitude="-3" metricUnit="m" metricValue="1.2" pageId="1" pageNumber="587" unit="mm" value="1.2">1.2mm</quantity>
, n=13). Forewing: Upperside ground color dark brown. Underside ground color white.
<collectingRegion id="49BBF833FF98DD27FB3AFAD1FABBBB68" box="[1233,1316,1372,1397]" country="Sri Lanka" name="Central" pageId="1" pageNumber="587">Central</collectingRegion>
symmetry system with distal band forming a series of dark bars. The bar in cell M2 twice as width as others while the bars completely dislocated inward in cell CuA1 and bent outwards in cell CuA2. Proximal band represented by 1 to 2 dark dots in discoidal cell. Discal spot represented by a dark bar. Element “g” prominent forming a continuous dark band with dark scales extending to marginal band along each vein. Marginal ocelli present in each cell as a series of barely connected dark oval spots. Parafocal element forming a continuous dark line along the margin. Hindwing: Upperside ground color dark brown. Bright orange band with wavy outer margin present at outer half of postdiscal and marginal area from cell M1 to A. “Tail-like” projection of CuA2 short, dark with white tip. Underside ground color white.
<collectingRegion id="49BBF833FF98DD27FEC4F909FE1EB880" box="[303,385,1668,1693]" country="Sri Lanka" name="Central" pageId="1" pageNumber="587">Central</collectingRegion>
symmetry system with distal band forming a series of dark bars, completely dislocated outward in cells M1 to M3, displaced inward in cells CuA1 and CuA2, becoming a stand alone dark spot in cell 2A and also in 3A in some individuals. Proximal band forming two dark spots in Sc+R1 cell and discoidal cell. Discal spot represented by a dark bar. Element “g” prominent represented by a continuous band with color scales extending to marginal band along each vein, dark in cells Sc+R1 and R2 while bright orange from cells M1 to 2A. Marginal ocelli present in each cell as a series of dark oval spots. Parafocal element forming a continuous narrow dark line along the margin. Fringes of both wings distinctly chequered with dark and white cilia.
</paragraph>
<paragraph id="8BC036D1FF98DD27FF2CF805FCAFB9DB" blockId="1.[151,1437,903,1990]" pageId="1" pageNumber="587">
Female (
<figureCitation id="13442A54FF98DD27FEC2F805FE08B9BC" box="[297,407,1928,1953]" captionStart="FIGURES 1 – 4" captionStartId="2.[151,264,1184,1207]" captionTargetBox="[150,1436,195,1160]" captionTargetId="figure@2.[150,1436,195,1160]" captionTargetPageId="2" captionText="FIGURES 1 – 4. Talicada nyseus lami subsp. n .. 1, holotype ♂, upperside; 2, ditto, underside; 3, paratype ♀, upperside; 4, ditto, underside. (scale bar = 10 mm)" httpUri="https://zenodo.org/record/582810/files/figure.png" pageId="1" pageNumber="587">Figs. 3–4</figureCitation>
): Length of antenna
<quantity id="4C879B34FF98DD27FD94F805FD60B9BC" box="[639,767,1928,1953]" metricMagnitude="-3" metricUnit="m" metricValue="6.95" metricValueMax="7.7" metricValueMin="6.2" pageId="1" pageNumber="587" unit="mm" value="6.95" valueMax="7.7" valueMin="6.2">6.2–7.7mm</quantity>
(
<quantity id="4C879B34FF98DD27FCE6F805FCC2B9BC" box="[781,861,1928,1953]" metricMagnitude="-3" metricUnit="m" metricValue="6.9" pageId="1" pageNumber="587" unit="mm" value="6.9">6.9mm</quantity>
±
<quantity id="4C879B34FF98DD27FC90F805FC53B9BC" box="[891,972,1928,1953]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="1" pageNumber="587" unit="mm" value="0.5">0.5mm</quantity>
, n=6). Wingspan
<quantity id="4C879B34FF98DD27FB78F805FAB0B9BC" box="[1171,1327,1928,1953]" metricMagnitude="-2" metricUnit="m" metricValue="2.695" metricValueMax="2.87" metricValueMin="2.52" pageId="1" pageNumber="587" unit="mm" value="26.95" valueMax="28.7" valueMin="25.2">25.2–28.7mm</quantity>
(
<quantity id="4C879B34FF98DD27FAD5F805FA03B9BC" box="[1342,1436,1928,1953]" metricMagnitude="-2" metricUnit="m" metricValue="2.66" pageId="1" pageNumber="587" unit="mm" value="26.6">26.6mm</quantity>
±
<quantity id="4C879B34FF98DD27FF46F820FF61B9D8" box="[173,254,1965,1989]" metricMagnitude="-3" metricUnit="m" metricValue="1.2" pageId="1" pageNumber="587" unit="mm" value="1.2">1.2mm</quantity>
, n=6). Wing pattern resembles to that of the male.
</paragraph>
</subSubSection>
<caption id="DF006659FF9BDD24FF7CFB2DFE4DBAC8" httpUri="https://zenodo.org/record/582810/files/figure.png" pageId="2" pageNumber="588" targetBox="[150,1436,195,1160]" targetPageId="2">
<paragraph id="8BC036D1FF9BDD24FF7CFB2DFE4DBAC8" blockId="2.[151,1436,1184,1237]" pageId="2" pageNumber="588">
<emphasis id="B90BEAC3FF9BDD24FF7CFB2DFEACBAA8" bold="true" box="[151,307,1184,1207]" pageId="2" pageNumber="588">FIGURES 1–4</emphasis>
.
<taxonomicName id="4C7F4D52FF9BDD24FEABFB2DFD8BBAAB" box="[320,532,1184,1207]" class="Insecta" family="Lycaenidae" genus="Talicada" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="588" phylum="Arthropoda" rank="subSpecies" species="nyseus" status="subsp. nov." subSpecies="lami">
<emphasis id="B90BEAC3FF9BDD24FEABFB2DFD8BBAAB" box="[320,532,1184,1207]" italics="true" pageId="2" pageNumber="588">Talicada nyseus lami</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23857B8FF9BDD24FDF1FB2CFDF1BAAB" box="[538,622,1185,1207]" pageId="2" pageNumber="588" rank="subSpecies">subsp. n</taxonomicNameLabel>
.. 1, holotype ♂, upperside; 2, ditto, underside; 3, paratype ♀, upperside; 4, ditto, underside. (scale bar = 10 mm)
</paragraph>
</caption>
<subSubSection id="C365655AFF9BDD24FF2CFB71FCFEBB41" pageId="2" pageNumber="588" type="distribution">
<paragraph id="8BC036D1FF9BDD24FF2CFB71FCFEBB41" blockId="2.[151,1437,1276,2036]" pageId="2" pageNumber="588">
<emphasis id="B90BEAC3FF9BDD24FF2CFB71FE6DBB08" bold="true" box="[199,498,1276,1301]" pageId="2" pageNumber="588">Distribution and habitat.</emphasis>
This subspecies is known only from well preserved natural forests in mountainous area of Baisha, Changjiang and Dongfang counties on
<collectingRegion id="49BBF833FF9BDD24FD11FAADFCD4BB24" box="[762,843,1312,1337]" country="China" name="Hainan" pageId="2" pageNumber="588">Hainan</collectingRegion>
<collectingCountry id="F3687641FF9BDD24FCB9FAADFC06BB24" box="[850,921,1312,1337]" name="Iceland" pageId="2" pageNumber="588">Island</collectingCountry>
,
<collectingCountry id="F3687641FF9BDD24FC4EFAADFC73BB24" box="[933,1004,1312,1337]" name="China" pageId="2" pageNumber="588">China</collectingCountry>
. It prefers shrubs adjacent to bare rock in karst landscape or along stream, where its hostplants present.
</paragraph>
</subSubSection>
<subSubSection id="C365655AFF9BDD24FF2CFAE5FA10B828" pageId="2" pageNumber="588" type="biology_ecology">
<paragraph id="8BC036D1FF9BDD24FF2CFAE5FAF1BBB8" blockId="2.[151,1437,1276,2036]" pageId="2" pageNumber="588">
<emphasis id="B90BEAC3FF9BDD24FF2CFAE5FED4BB9C" bold="true" box="[199,331,1384,1409]" pageId="2" pageNumber="588">Host plant.</emphasis>
The immature of this subspecies is apparently associated with the family
<taxonomicName id="4C7F4D52FF9BDD24FB68FAE5FA86BB9C" box="[1155,1305,1384,1409]" class="Magnoliopsida" family="Crassulaceae" kingdom="Plantae" order="Saxifragales" pageId="2" pageNumber="588" phylum="Tracheophyta" rank="family">Crassulaceae</taxonomicName>
.
<taxonomicName id="4C7F4D52FF9BDD24FAC8FAE5FF77BBB9" class="Magnoliopsida" family="Crassulaceae" genus="Kalanchoe" kingdom="Plantae" order="Saxifragales" pageId="2" pageNumber="588" phylum="Tracheophyta" rank="species" species="integra">
<emphasis id="B90BEAC3FF9BDD24FAC8FAE5FF77BBB9" italics="true" pageId="2" pageNumber="588">Kalanchoe integra</emphasis>
</taxonomicName>
and
<taxonomicName id="4C7F4D52FF9BDD24FEF5FA01FDBFBBB9" box="[286,544,1420,1445]" class="Gymnostomatea" family="Spathidiidae" genus="Bryophyllum" kingdom="Plantae" order="Spathidiida" pageId="2" pageNumber="588" phylum="Ciliophora" rank="species" species="pinnatum">
<emphasis id="B90BEAC3FF9BDD24FEF5FA01FDBFBBB9" box="[286,544,1420,1445]" italics="true" pageId="2" pageNumber="588">Bryophyllum pinnatum</emphasis>
</taxonomicName>
, a naturalised plant with African origin, were utilized by larvae in the wild.
</paragraph>
<paragraph id="8BC036D1FF9BDD24FF2CFA3DFA10B828" blockId="2.[151,1437,1276,2036]" pageId="2" pageNumber="588">
<emphasis id="B90BEAC3FF9BDD24FF2CFA3DFE57BBD4" bold="true" box="[199,456,1456,1481]" pageId="2" pageNumber="588">Immature bionomics.</emphasis>
ovum (
<figureCitation id="13442A54FF9BDD24FDC9FA3DFDFABBD4" box="[546,613,1456,1481]" captionStart="FIGURES 5 – 8" captionStartId="3.[151,264,1178,1201]" captionTargetBox="[150,1436,193,1158]" captionTargetId="figure@3.[150,1436,193,1158]" captionTargetPageId="3" captionText="FIGURES 5 – 8. Immature stages of Talicada nyseus lami subsp. n .. 5, ovum on underside surface of Bryophyllum pinnatum; 6, last-instar larva (epidermis of the hostplant removed); 7, pupa on underside surface of Kalanchoe integra, lateral view; 8, ditto, dorsal view." httpUri="https://zenodo.org/record/582811/files/figure.png" pageId="2" pageNumber="588">Fig. 5</figureCitation>
) is laid singly on the underside of leaf blade. Once hatched, larva bores into leaf of hostplant as leaf miner and feeds on tissue in mesophyll layer (
<figureCitation id="13442A54FF9BDD24FC39FA59FB89BBF0" box="[978,1046,1492,1517]" captionStart="FIGURES 5 – 8" captionStartId="3.[151,264,1178,1201]" captionTargetBox="[150,1436,193,1158]" captionTargetId="figure@3.[150,1436,193,1158]" captionTargetPageId="3" captionText="FIGURES 5 – 8. Immature stages of Talicada nyseus lami subsp. n .. 5, ovum on underside surface of Bryophyllum pinnatum; 6, last-instar larva (epidermis of the hostplant removed); 7, pupa on underside surface of Kalanchoe integra, lateral view; 8, ditto, dorsal view." httpUri="https://zenodo.org/record/582811/files/figure.png" pageId="2" pageNumber="588">Fig. 6</figureCitation>
). Larva does not emerge unless the original leaf is completely consumed. Mature larva pupates on the underside of leaf of hostplant (
<figureCitation id="13442A54FF9BDD24FB05FA75FAC4B80C" box="[1262,1371,1528,1553]" captionStart="FIGURES 5 – 8" captionStartId="3.[151,264,1178,1201]" captionTargetBox="[150,1436,193,1158]" captionTargetId="figure@3.[150,1436,193,1158]" captionTargetPageId="3" captionText="FIGURES 5 – 8. Immature stages of Talicada nyseus lami subsp. n .. 5, ovum on underside surface of Bryophyllum pinnatum; 6, last-instar larva (epidermis of the hostplant removed); 7, pupa on underside surface of Kalanchoe integra, lateral view; 8, ditto, dorsal view." httpUri="https://zenodo.org/record/582811/files/figure.png" pageId="2" pageNumber="588">Figs. 7–8</figureCitation>
). The entire immature stage, from egg to emergence of adult, takes 61–62 days (n=3) under indoor uncontrolled condition.
</paragraph>
</subSubSection>
<subSubSection id="C365655AFF9BDD24FF2CF9CDFC8CB844" box="[199,787,1600,1625]" pageId="2" pageNumber="588" type="description">
<paragraph id="8BC036D1FF9BDD24FF2CF9CDFC8CB844" blockId="2.[151,1437,1276,2036]" box="[199,787,1600,1625]" pageId="2" pageNumber="588">
<emphasis id="B90BEAC3FF9BDD24FF2CF9CDFEDFB844" bold="true" box="[199,320,1600,1625]" pageId="2" pageNumber="588">Voltinism.</emphasis>
Multivoltine, adult occurs all year round.
</paragraph>
</subSubSection>
<subSubSection id="C365655AFF9BDD24FF2CF9E9FC2DB8BD" pageId="2" pageNumber="588" type="etymology">
<paragraph id="8BC036D1FF9BDD24FF2CF9E9FC2DB8BD" blockId="2.[151,1437,1276,2036]" pageId="2" pageNumber="588">
<emphasis id="B90BEAC3FF9BDD24FF2CF9E9FED1B860" bold="true" box="[199,334,1636,1661]" pageId="2" pageNumber="588">Etymology.</emphasis>
The name of the subspecies is dedicated to Alexander Lam’s family in Boston,
<collectingCountry id="F3687641FF9BDD24FB2EF9E8FA9FB860" box="[1221,1280,1637,1661]" name="United States of America" pageId="2" pageNumber="588">USA</collectingCountry>
, for their love and care to the senior author during his undergraduate study in Boston.
</paragraph>
</subSubSection>
<subSubSection id="C365655AFF9BDD25FF2CF921FAE7BB6C" lastPageId="3" lastPageNumber="589" pageId="2" pageNumber="588" type="diagnosis">
<paragraph id="8BC036D1FF9BDD24FF2CF921FEDAB9E9" blockId="2.[151,1437,1276,2036]" pageId="2" pageNumber="588">
<emphasis id="B90BEAC3FF9BDD24FF2CF921FEDEB8D8" bold="true" box="[199,321,1708,1733]" pageId="2" pageNumber="588">Diagnosis.</emphasis>
The male genitalia (
<figureCitation id="13442A54FF9BDD24FDD2F921FD29B8D8" box="[569,694,1708,1733]" captionStart-0="FIGURES 9 – 11" captionStart-1="FIGURE 12" captionStartId-0="4.[151,264,1178,1201]" captionStartId-1="4.[151,250,1240,1262]" captionTargetBox-0="[151,1436,227,1158]" captionTargetId-0="figure@4.[150,1436,193,1158]" captionTargetPageId-0="4" captionText-0="FIGURES 9 – 11. Male genitalia of Talicada nyseus lami subsp. n .. 9, lateral view; 10, dorsal view; 11, ventral view. (scale bar = 0.5 mm)" captionText-1="FIGURE 12. Enlarged view of male genitalia of Talicada nyseus lami subsp. n .. A, right valva; B, aedeagus. (scale bar = 0.5 mm)" httpUri-0="https://zenodo.org/record/582812/files/figure.png" pageId="2" pageNumber="588">Figs. 9–12</figureCitation>
) of
<taxonomicName id="4C7F4D52FF9BDD24FD07F923FCC1B8D8" box="[748,862,1708,1733]" class="Insecta" family="Lycaenidae" genus="Talicada" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="588" phylum="Arthropoda" rank="subSpecies" species="nyseus" subSpecies="lami">
<emphasis id="B90BEAC3FF9BDD24FD07F923FCC1B8D8" box="[748,862,1708,1733]" italics="true" pageId="2" pageNumber="588">T. n. lami</emphasis>
</taxonomicName>
resembles that of nominate subspecies and
<taxonomicName id="4C7F4D52FF9BDD24FA8CF923FE98B8FB" class="Insecta" family="Lycaenidae" genus="Talicada" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="588" phylum="Arthropoda" rank="subSpecies" species="nyseus" subSpecies="delhiensis">
<emphasis id="B90BEAC3FF9BDD24FA8CF923FE98B8FB" italics="true" pageId="2" pageNumber="588">T. n. delhiensis</emphasis>
</taxonomicName>
in general, with long and tapering uncus, prominent brachium, broad valvae with harpe modified to long spine-shaped projection. The size of
<taxonomicName id="4C7F4D52FF9BDD24FDDCF97DFD47B915" box="[567,728,1775,1800]" class="Insecta" family="Lycaenidae" genus="Talicada" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="588" phylum="Arthropoda" rank="subSpecies" species="nyseus" subSpecies="lami">
<emphasis id="B90BEAC3FF9BDD24FDDCF97DFD47B915" box="[567,728,1775,1800]" italics="true" pageId="2" pageNumber="588">T. nyseus lami</emphasis>
</taxonomicName>
is similar to
<emphasis id="B90BEAC3FF9BDD24FC85F97DFBB9B91A" box="[878,1062,1776,1799]" italics="true" pageId="2" pageNumber="588">T. n. annamitica</emphasis>
and both are the smallest among all subspecies. Interestingly, the wing pattern of
<taxonomicName id="4C7F4D52FF9BDD24FD0DF89CFC0EB934" box="[742,913,1808,1833]" class="Insecta" family="Lycaenidae" genus="Talicada" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="588" phylum="Arthropoda" rank="subSpecies" species="nyseus" subSpecies="lami">
<emphasis id="B90BEAC3FF9BDD24FD0DF89CFC0EB934" box="[742,913,1808,1833]" italics="true" pageId="2" pageNumber="588">T. nyseus lami</emphasis>
</taxonomicName>
seems like a combination of several other subspecies. First of all, the wavy outer margin of the orange band on upper hindwing and dark marginal ocelli from cell M1 to 2A on underside hindwing are features only found in three eastern subspecies,
<emphasis id="B90BEAC3FF9BDD24FB2AF8D9FACBB976" box="[1217,1364,1876,1899]" italics="true" pageId="2" pageNumber="588">
T. n.
<taxonomicName id="4C7F4D52FF9BDD24FAE9F8D9FACBB976" box="[1282,1364,1876,1899]" class="Insecta" family="Lycaenidae" genus="Talicada" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="588" phylum="Arthropoda" rank="species" species="metana">metana</taxonomicName>
</emphasis>
,
<emphasis id="B90BEAC3FF9BDD24FA8CF8D9FE8AB992" italics="true" pageId="2" pageNumber="588">T. n. annamitica</emphasis>
and
<taxonomicName id="4C7F4D52FF9BDD24FEBFF8F5FE5BB98D" box="[340,452,1911,1936]" class="Insecta" family="Lycaenidae" genus="Talicada" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="588" phylum="Arthropoda" rank="subSpecies" species="nyseus" subSpecies="lami">
<emphasis id="B90BEAC3FF9BDD24FEBFF8F5FE5BB98D" box="[340,452,1911,1936]" italics="true" pageId="2" pageNumber="588">T. n. lami</emphasis>
</taxonomicName>
. However, dark spots in the middle of discoidal cell is a trait only shared by
<emphasis id="B90BEAC3FF9BDD24FA8CF8F5FF61B9AC" italics="true" pageId="2" pageNumber="588">T. n. macbethi</emphasis>
and
<taxonomicName id="4C7F4D52FF9BDD24FED3F817FE3BB9AC" box="[312,420,1944,1969]" class="Insecta" family="Lycaenidae" genus="Talicada" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="588" phylum="Arthropoda" rank="subSpecies" species="nyseus" subSpecies="lami">
<emphasis id="B90BEAC3FF9BDD24FED3F817FE3BB9AC" box="[312,420,1944,1969]" italics="true" pageId="2" pageNumber="588">T. n. lami</emphasis>
</taxonomicName>
. Furthermore,
<taxonomicName id="4C7F4D52FF9BDD24FDB8F817FD21B9AC" box="[595,702,1944,1969]" class="Insecta" family="Lycaenidae" genus="Talicada" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="588" phylum="Arthropoda" rank="subSpecies" species="nyseus" subSpecies="lami">
<emphasis id="B90BEAC3FF9BDD24FDB8F817FD21B9AC" box="[595,702,1944,1969]" italics="true" pageId="2" pageNumber="588">T. n. lami</emphasis>
</taxonomicName>
is similar to
<emphasis id="B90BEAC3FF9BDD24FCB7F817FC62B9AC" box="[860,1021,1944,1969]" italics="true" pageId="2" pageNumber="588">T. n. burmana</emphasis>
the most in that the arrangement of distal band of central symmetry system, but all the spots are detached from the band of element “g” as defined by
<bibRefCitation id="EFEE4B20FF9BDD24FF7CF856FEDDB9E9" author="Nijhout" box="[151,322,2011,2036]" pageId="2" pageNumber="588" refString="Nijhout, H. F. (1991) The development and evolution of butterfly wing patterns. Smithsonian Institution Press, Washington &amp; London, xvi + 297 pp." type="book" year="1991">Nijhout (1991)</bibRefCitation>
.
</paragraph>
<caption id="DF006659FF9ADD25FF7CFB17FE90BAF3" httpUri="https://zenodo.org/record/582811/files/figure.png" pageId="3" pageNumber="589" targetBox="[150,1436,193,1158]" targetPageId="3">
<paragraph id="8BC036D1FF9ADD25FF7CFB17FE90BAF3" blockId="3.[151,1436,1178,1262]" pageId="3" pageNumber="589">
<emphasis id="B90BEAC3FF9ADD25FF7CFB17FEA3BAAD" bold="true" box="[151,316,1178,1201]" pageId="3" pageNumber="589">FIGURES 5–8.</emphasis>
Immature stages of
<taxonomicName id="4C7F4D52FF9ADD25FDE1FB16FD42BAAC" box="[522,733,1179,1201]" class="Insecta" family="Lycaenidae" genus="Talicada" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="589" phylum="Arthropoda" rank="subSpecies" species="nyseus" status="subsp. nov." subSpecies="lami">
<emphasis id="B90BEAC3FF9ADD25FDE1FB16FD42BAAC" box="[522,733,1179,1201]" italics="true" pageId="3" pageNumber="589">Talicada nyseus lami</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23857B8FF9ADD25FD0FFB16FCA7BAAC" box="[740,824,1179,1201]" pageId="3" pageNumber="589" rank="subSpecies">subsp. n</taxonomicNameLabel>
.. 5, ovum on underside surface of
<taxonomicName id="4C7F4D52FF9ADD25FB7DFB16FAE4BAAC" box="[1174,1403,1179,1201]" class="Gymnostomatea" family="Spathidiidae" genus="Bryophyllum" kingdom="Plantae" order="Spathidiida" pageId="3" pageNumber="589" phylum="Ciliophora" rank="species" species="pinnatum">
<emphasis id="B90BEAC3FF9ADD25FB7DFB16FAE4BAAC" box="[1174,1403,1179,1201]" italics="true" pageId="3" pageNumber="589">Bryophyllum pinnatum</emphasis>
</taxonomicName>
; 6, last-instar larva (epidermis of the hostplant removed); 7, pupa on underside surface of
<taxonomicName id="4C7F4D52FF9ADD25FBE9FB37FB23BACD" box="[1026,1212,1210,1232]" class="Magnoliopsida" family="Crassulaceae" genus="Kalanchoe" kingdom="Plantae" order="Saxifragales" pageId="3" pageNumber="589" phylum="Tracheophyta" rank="species" species="integra">
<emphasis id="B90BEAC3FF9ADD25FBE9FB37FB23BACD" box="[1026,1212,1210,1232]" italics="true" pageId="3" pageNumber="589">Kalanchoe integra</emphasis>
</taxonomicName>
, lateral view; 8, ditto, dorsal view.
</paragraph>
</caption>
<paragraph id="8BC036D1FF9ADD25FF2CFA98FAE7BB6C" blockId="3.[151,1436,1301,1393]" pageId="3" pageNumber="589">
Six distinctive underside patterns, assigned as A to F hereinto, can be identified among the nine subspecies: three subspecies,
<emphasis id="B90BEAC3FF9ADD25FEF7FABAFE1FBB4D" box="[284,384,1335,1360]" italics="true" pageId="3" pageNumber="589">macbethi</emphasis>
(A),
<emphasis id="B90BEAC3FF9ADD25FE55FABAFDBEBB4D" box="[446,545,1335,1360]" italics="true" pageId="3" pageNumber="589">burmana</emphasis>
(B) and
<emphasis id="B90BEAC3FF9ADD25FD62FABAFD25BB4D" box="[649,698,1335,1360]" italics="true" pageId="3" pageNumber="589">lami</emphasis>
(C), have unique pattern and the rest three patterns are shared by three subspecies pairs,
<taxonomicName id="4C7F4D52FF9ADD25FE79FAD4FE44BB6D" box="[402,475,1369,1392]" class="Insecta" family="Lycaenidae" genus="Talicada" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="589" phylum="Arthropoda" rank="species" species="nyseus">
<emphasis id="B90BEAC3FF9ADD25FE79FAD4FE44BB6D" box="[402,475,1369,1392]" italics="true" pageId="3" pageNumber="589">nyseus</emphasis>
</taxonomicName>
&amp;
<emphasis id="B90BEAC3FF9ADD25FE15FAD5FDF4BB6C" box="[510,619,1368,1393]" italics="true" pageId="3" pageNumber="589">delhiensis</emphasis>
(D);
<emphasis id="B90BEAC3FF9ADD25FD4DFAD5FC96BB6C" box="[678,777,1368,1393]" italics="true" pageId="3" pageNumber="589">khasiana</emphasis>
&amp;
<emphasis id="B90BEAC3FF9ADD25FCC7FAD4FC0EBB6D" box="[812,913,1369,1392]" italics="true" pageId="3" pageNumber="589">assamica</emphasis>
(E);
<taxonomicName id="4C7F4D52FF9ADD25FC21FAD4FB84BB6D" box="[970,1051,1369,1392]" class="Insecta" family="Lycaenidae" genus="Talicada" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="589" phylum="Arthropoda" rank="species" species="metana">
<emphasis id="B90BEAC3FF9ADD25FC21FAD4FB84BB6D" box="[970,1051,1369,1392]" italics="true" pageId="3" pageNumber="589">metana</emphasis>
</taxonomicName>
&amp;
<emphasis id="B90BEAC3FF9ADD25FBD6FAD4FB27BB6D" box="[1085,1208,1369,1392]" italics="true" pageId="3" pageNumber="589">annamitica</emphasis>
(F), respectively.
</paragraph>
</subSubSection>
</treatment>
</document>
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