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<document id="A9BF7786A7A9B0AB6A0A78470F9433B0" ID-ISSN="1464-5262" IM.materialsCitations_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1629979807253" checkinUser="felipe" docAuthor="Rodney" docDate="2003" docId="03D687A1BE362B69284AD4BCFE077340" docLanguage="en" docName="JNatHist.37.21.2615-2626.pdf" docOrigin="Journal of Natural History 37" docStyle="DocumentStyle:443144A73DAE3E092C51A378E4CD0138.2:JNatHist.2003-2004.journal_article.1cover" docStyleId="443144A73DAE3E092C51A378E4CD0138" docStyleName="JNatHist.2003-2004.journal_article.1cover" docStyleVersion="2" docTitle="Leptandria ecitophila Rodney 2003, sp. n." docType="treatment" docVersion="2" lastPageNumber="2625" masterDocId="FFEFFFD9BE3F2B622A32D528FFBB750A" masterDocTitle="Leptandria gen. n., a monophyletic group of Neotropical aleocharine Staphylinidae (Coleoptera)" masterLastPageNumber="2626" masterPageNumber="2615" pageNumber="2623" updateTime="1699132954488" updateUser="plazi" zenodo-license-document="CLOSED">
<mods:mods id="4B643F447A000AEB045E4E5C0051F81E" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="9C15B2D1729F397AA4370225BF8943B2">
<mods:title id="D48E8DAA301245449600BAF6DF308100">Leptandria gen. n., a monophyletic group of Neotropical aleocharine Staphylinidae (Coleoptera)</mods:title>
</mods:titleInfo>
<mods:name id="C650303E6D5B0409DE48D3A0D397CFD5" type="personal">
<mods:role id="490FC9AD37E7DACD77D99C9D64C3268B">
<mods:roleTerm id="58F45878BE7DF9C0CEC3AB7252933165">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="82B25B5A8631C1AE9BF2A2C570814F10">Rodney</mods:namePart>
</mods:name>
<mods:typeOfResource id="11C431E09BC93352504FA8B99AE7FDC4">text</mods:typeOfResource>
<mods:relatedItem id="F451DA65091B41147184137F6D0F48B9" type="host">
<mods:titleInfo id="6F8CF1946F479C08A5D52E2A8C8549D2">
<mods:title id="8E94959B27EC1D41AFB85A45B35D9FCF">Journal of Natural History</mods:title>
</mods:titleInfo>
<mods:part id="AD1D1595B8A93CCC45123CAB6075CD50">
<mods:date id="9E6D23FDB5841FDA814C16A439DFD184">2003</mods:date>
<mods:detail id="B87F99126818A7FEAC9DE650BAF6EF50" type="pubDate">
<mods:number id="7A44B083B2A2957C363D6BCDBCD68440">2003-10-10</mods:number>
</mods:detail>
<mods:detail id="9B5D13DBAF413F0B0A65BABCC52E04A0" type="volume">
<mods:number id="12D5F1747D0A9995E0539202CCADAE74">37</mods:number>
</mods:detail>
<mods:extent id="0999D5F185C74AC6B79F4DE65B774943" unit="page">
<mods:start id="BCC7FC77487949817FB2EAF56F6E54EB">2615</mods:start>
<mods:end id="0B1A50435DFF1F88CF273A439D365DE2">2626</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="C7F36836FE6D37EAA45419FC4243C9A9">journal article</mods:classification>
<mods:identifier id="F5D65B86C703C610635C3F516B4F524F" type="ISSN">1464-5262</mods:identifier>
</mods:mods>
<treatment id="03D687A1BE362B69284AD4BCFE077340" LSID="urn:lsid:plazi:treatment:03D687A1BE362B69284AD4BCFE077340" httpUri="http://treatment.plazi.org/id/03D687A1BE362B69284AD4BCFE077340" lastPageId="11" lastPageNumber="2625" pageId="9" pageNumber="2623">
<subSubSection id="C365653CBE362B6B284AD4BCFCDE74C6" pageId="9" pageNumber="2623" type="nomenclature">
<paragraph id="8BC036B7BE362B6B284AD4BCFC0F74A6" blockId="9.[632,948,404,460]" box="[632,948,404,428]" pageId="9" pageNumber="2623">
<heading id="D08881DBBE362B6B284AD4BCFC0F74A6" box="[632,948,404,428]" centered="true" fontSize="10" level="2" pageId="9" pageNumber="2623" reason="3">
<taxonomicName id="4C7F4D34BE362B6B284AD4BCFCDD74A6" ID-CoL="3THMB" authority="Rodney, 2003" authorityName="Rodney" authorityYear="2003" box="[632,870,404,428]" class="Insecta" family="Staphylinidae" genus="Leptandria" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="2623" phylum="Arthropoda" rank="species" species="ecitophila" status="sp. nov.">
<emphasis id="B90BEAA5BE362B6B284AD4BCFCDD74A6" bold="true" box="[632,870,404,428]" italics="true" pageId="9" pageNumber="2623">Leptandria ecitophila</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A23857DEBE362B6B2944D4BDFC0F74A6" box="[886,948,405,428]" pageId="9" pageNumber="2623" rank="species">sp. n.</taxonomicNameLabel>
</heading>
</paragraph>
<paragraph id="8BC036B7BE362B6B28F5D49CFCDE74C6" blockId="9.[632,948,404,460]" box="[711,869,435,460]" pageId="9" pageNumber="2623">(Figures 4–5)</paragraph>
</subSubSection>
<subSubSection id="C365653CBE362B692B53D4CCFE077340" lastPageId="11" lastPageNumber="2625" pageId="9" pageNumber="2623" type="materials_examined">
<paragraph id="8BC036B7BE362B682B53D4CCFB4F73C2" blockId="9.[313,1268,484,668]" lastBlockId="10.[313,1269,1232,1736]" lastPageId="10" lastPageNumber="2624" pageId="9" pageNumber="2623">
<emphasis id="B90BEAA5BE362B6B2B53D4CCFE7174F6" box="[353,458,484,508]" italics="true" pageId="9" pageNumber="2623">Material.</emphasis>
H
<collectionCode id="ED6EAE72BE362B6B2811D4CCFD8A74F6" box="[547,561,484,508]" name="Tavera, Department of Geology and Geophysics" pageId="9" pageNumber="2623"></collectionCode>
 male, [‘
<emphasis id="B90BEAA5BE362B6B2881D4CCFC8674F6" bold="true" box="[691,829,484,508]" pageId="9" pageNumber="2623">
<collectingCountry id="F3687627BE362B6B2881D4CCFC8674F6" box="[691,829,484,508]" name="Ecuador" pageId="9" pageNumber="2623">ECUADOR</collectingCountry>
</emphasis>
:
<collectingRegion id="49BBF855BE362B6B2978D4CCFC3674F1" box="[842,909,484,507]" country="Ecuador" name="Napo" pageId="9" pageNumber="2623">Napo</collectingRegion>
, mid. Rio Tiputini, Yasuni Res. Stn., 0
<emphasis id="B90BEAA5BE362B6B2BB4D72CFE2B7716" bold="true" box="[390,400,516,540]" pageId="9" pageNumber="2623">°</emphasis>
40.5
<emphasis id="B90BEAA5BE362B6B2B8DD72CFE7E7716" bold="true" box="[447,453,516,540]" pageId="9" pageNumber="2623">∞</emphasis>
<geoCoordinate id="EE4B5070BE362B6B2BF7D72CFDA47716" box="[453,543,516,540]" degrees="76" direction="south" minutes="24" orientation="latitude" pageId="9" pageNumber="2623" precision="925" value="-76.4">
S 76
<emphasis id="B90BEAA5BE362B6B2BCBD72CFDB87716" bold="true" box="[505,515,516,540]" pageId="9" pageNumber="2623">°</emphasis>
24
</geoCoordinate>
<emphasis id="B90BEAA5BE362B6B282CD72CFD9F7716" bold="true" box="[542,548,516,540]" pageId="9" pageNumber="2623">∞</emphasis>
W;
<date id="FFC11077BE362B6B2863D72CFD527716" box="[593,745,516,540]" pageId="9" pageNumber="2623" value="1999-07-18">18 July 1999</date>
. AKT #083’/‘
<taxonomicName id="4C7F4D34BE362B6B29B9D72DFB887716" box="[907,1075,516,540]" class="Insecta" family="Formicidae" genus="Eciton" kingdom="Animalia" order="Hymenoptera" pageId="9" pageNumber="2623" phylum="Arthropoda" rank="species" species="buchelli">
<emphasis id="B90BEAA5BE362B6B29B9D72DFB887716" box="[907,1075,516,540]" italics="true" pageId="9" pageNumber="2623">Eciton buchelli</emphasis>
</taxonomicName>
colony EC #21. Statary bivouac site just after emigration. A. Tishechkin’/‘
<typeStatus id="54C48815BE362B6B29E2D70CFBD57736" box="[976,1134,548,572]" pageId="9" pageNumber="2623" type="holotype">HOLOTYPE</typeStatus>
,
<taxonomicName id="4C7F4D34BE362B6B2E48D70CFDBA7756" authority="Hanley" authorityName="Hanley" authorityYear="2003" class="Insecta" family="Staphylinidae" genus="Leptandria" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="2623" phylum="Arthropoda" rank="species" species="ecitophila">
<emphasis id="B90BEAA5BE362B6B2E48D70CFE1F7756" italics="true" pageId="9" pageNumber="2623">Leptandria ecitophila</emphasis>
Hanley
</taxonomicName>
, desig.
<collectionCode id="ED6EAE72BE362B6B2865D76CFDD47751" box="[599,623,580,603]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="9" pageNumber="2623">R</collectionCode>
. S. Hanley, 2001’ (red)] (
<collectionCode id="ED6EAE72BE362B6B29A3D76CFC5A7756" box="[913,993,580,604]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34264" name="University of Kansas - Biodiversity Institute" pageId="9" pageNumber="2623" type="Museum">SEMC</collectionCode>
). P
<collectionCode id="ED6EAE72BE362B6B2E29D76CFB937756" box="[1051,1064,580,604]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="9" pageNumber="2623"></collectionCode>
<collectionCode id="ED6EAE72BE362B6B2E07D76CFBF97756" box="[1077,1090,580,604]" name="Tavera, Department of Geology and Geophysics" pageId="9" pageNumber="2623"></collectionCode>
,
<specimenCount id="9D79FD3EBE362B6B2EB7D76CFB4B7756" box="[1157,1264,580,604]" pageId="9" pageNumber="2623" type="male">one male</specimenCount>
,
<specimenCount id="9D79FD3EBE362B6B2B0BD74CFE7E7771" box="[313,453,611,635]" pageId="9" pageNumber="2623" type="female">two females</specimenCount>
(
<collectionCode id="ED6EAE72BE362B6B2BEAD74CFD937776" box="[472,552,612,636]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34264" name="University of Kansas - Biodiversity Institute" pageId="9" pageNumber="2623" type="Museum">SEMC</collectionCode>
),
<specimenCount id="9D79FD3EBE362B6B287BD74CFD0F7771" box="[585,692,611,635]" pageId="9" pageNumber="2623" type="male">one male</specimenCount>
(
<collectionCode id="ED6EAE72BE362B6B28FAD74CFCAF7776" box="[712,788,612,636]" pageId="9" pageNumber="2623">LSUC</collectionCode>
) with same label data as
<typeStatus id="54C48815BE362B6B2E66D74BFB017771" box="[1108,1210,611,635]" pageId="9" pageNumber="2623" type="holotype">holotype</typeStatus>
;
<specimenCount id="9D79FD3EBE362B6B2EF9D74CFE1B7791" pageId="9" pageNumber="2623" type="generic">one specimen</specimenCount>
(
<collectionCode id="ED6EAE72BE362B6B2B8AD7ACFDB37796" box="[440,520,644,668]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34264" name="University of Kansas - Biodiversity Institute" pageId="9" pageNumber="2623" type="Museum">SEMC</collectionCode>
), [‘
<emphasis id="B90BEAA5BE362B6B2808D7ABFD7F7791" bold="true" box="[570,708,643,667]" pageId="9" pageNumber="2623">
<collectingCountry id="F3687627BE362B6B2808D7ABFD7F7791" box="[570,708,643,667]" name="Ecuador" pageId="9" pageNumber="2623">ECUADOR</collectingCountry>
</emphasis>
:
<collectingRegion id="49BBF855BE362B6B28E8D7ACFCA67791" box="[730,797,644,667]" country="Ecuador" name="Napo" pageId="9" pageNumber="2623">Napo</collectingRegion>
, mid. Rio Tiputini, Yasuni Res. Stn., 0
<emphasis id="B90BEAA5BE352B682B74D1F9FEEB71E3" bold="true" box="[326,336,1233,1257]" pageId="10" pageNumber="2624">°</emphasis>
40.5
<emphasis id="B90BEAA5BE352B682B4DD1F9FE3E71E3" bold="true" box="[383,389,1233,1257]" pageId="10" pageNumber="2624">∞</emphasis>
<geoCoordinate id="EE4B5070BE352B682BB7D1F9FE6771E3" box="[389,476,1233,1257]" degrees="76" direction="south" minutes="24" orientation="latitude" pageId="10" pageNumber="2624" precision="925" value="-76.4">
S 76
<emphasis id="B90BEAA5BE352B682B84D1F9FE7B71E3" bold="true" box="[438,448,1233,1257]" pageId="10" pageNumber="2624">°</emphasis>
24
</geoCoordinate>
<emphasis id="B90BEAA5BE352B682BE9D1F9FE5A71E3" bold="true" box="[475,481,1233,1257]" pageId="10" pageNumber="2624">∞</emphasis>
W;
<date id="FFC11077BE352B682838D1F9FD1971E3" box="[522,674,1233,1257]" pageId="10" pageNumber="2624" value="1999-06-18">18 June 1999</date>
. AKT #006’/‘
<taxonomicName id="4C7F4D34BE352B68290FD1FAFC5971E3" box="[829,994,1233,1257]" class="Insecta" family="Formicidae" genus="Eciton" kingdom="Animalia" order="Hymenoptera" pageId="10" pageNumber="2624" phylum="Arthropoda" rank="species" species="buchelli">
<emphasis id="B90BEAA5BE352B68290FD1FAFC5971E3" box="[829,994,1233,1257]" italics="true" pageId="10" pageNumber="2624">Eciton buchelli</emphasis>
</taxonomicName>
colony EC #3. In/along emigration colony; A. Tishechkin’/‘
<typeStatus id="54C48815BE352B6828D5D1D9FCC47003" box="[743,895,1265,1289]" pageId="10" pageNumber="2624" type="paratype">PARATYPE</typeStatus>
,
<taxonomicName id="4C7F4D34BE352B6829A6D1D9FB4B7003" authority="Hanley" authorityName="Hanley" authorityYear="2003" box="[916,1264,1265,1289]" class="Insecta" family="Staphylinidae" genus="Leptandria" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="2624" phylum="Arthropoda" rank="species" species="ecitophila">
<emphasis id="B90BEAA5BE352B6829A6D1D9FB317003" box="[916,1162,1265,1289]" italics="true" pageId="10" pageNumber="2624">Leptandria ecitophila</emphasis>
Hanley
</taxonomicName>
, desig.
<collectionCode id="ED6EAE72BE352B682BBED03AFE1F7023" box="[396,420,1298,1321]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="10" pageNumber="2624">R</collectionCode>
. S. Hanley, 2001’ (yellow)];
<specimenCount id="9D79FD3EBE352B682929D03AFC277023" box="[795,924,1297,1321]" pageId="10" pageNumber="2624" type="male">two males</specimenCount>
,
<specimenCount id="9D79FD3EBE352B682980D03AFBFE7023" box="[946,1093,1297,1321]" pageId="10" pageNumber="2624" type="female">two females</specimenCount>
(
<collectionCode id="ED6EAE72BE352B682E6DD039FB147023" box="[1119,1199,1297,1321]" pageId="10" pageNumber="2624">SMEC</collectionCode>
),
<specimenCount id="9D79FD3EBE352B682EE7D039FEC77043" pageId="10" pageNumber="2624" type="male">six males</specimenCount>
,
<specimenCount id="9D79FD3EBE352B682BB9D019FDA97043" box="[395,530,1329,1353]" pageId="10" pageNumber="2624" type="female">five females</specimenCount>
(
<collectionCode id="ED6EAE72BE352B682811D019FDD47043" box="[547,623,1329,1353]" pageId="10" pageNumber="2624">LSUC</collectionCode>
), [‘
<emphasis id="B90BEAA5BE352B6828AFD019FC9C7043" bold="true" box="[669,807,1329,1353]" pageId="10" pageNumber="2624">
<collectingCountry id="F3687627BE352B6828AFD019FC9C7043" box="[669,807,1329,1353]" name="Ecuador" pageId="10" pageNumber="2624">ECUADOR</collectingCountry>
</emphasis>
:
<collectingRegion id="49BBF855BE352B68290AD019FCC07042" box="[824,891,1329,1352]" country="Ecuador" name="Napo" pageId="10" pageNumber="2624">Napo</collectingRegion>
, mid. Rio Tiputini, Yasuni Res. Stn., 0
<emphasis id="B90BEAA5BE352B682BBED079FE2D7063" bold="true" box="[396,406,1361,1385]" pageId="10" pageNumber="2624">°</emphasis>
40.5
<emphasis id="B90BEAA5BE352B682BF7D079FE707063" bold="true" box="[453,459,1361,1385]" pageId="10" pageNumber="2624">∞</emphasis>
<geoCoordinate id="EE4B5070BE352B682BF9D079FD917063" box="[459,554,1361,1385]" degrees="76" direction="south" minutes="24" orientation="latitude" pageId="10" pageNumber="2624" precision="925" value="-76.4">
S 76
<emphasis id="B90BEAA5BE352B682837D079FDB47063" bold="true" box="[517,527,1361,1385]" pageId="10" pageNumber="2624">°</emphasis>
24
</geoCoordinate>
<emphasis id="B90BEAA5BE352B682818D079FD8B7063" bold="true" box="[554,560,1361,1385]" pageId="10" pageNumber="2624">∞</emphasis>
W;
<date id="FFC11077BE352B682850D079FCBD7063" box="[610,774,1361,1385]" pageId="10" pageNumber="2624" value="1999-07-22">22 July 1999</date>
. AKT #091’/‘
<taxonomicName id="4C7F4D34BE352B682986D07AFBD97063" box="[948,1122,1361,1385]" class="Insecta" family="Formicidae" genus="Eciton" kingdom="Animalia" order="Hymenoptera" pageId="10" pageNumber="2624" phylum="Arthropoda" rank="species" species="buchelli">
<emphasis id="B90BEAA5BE352B682986D07AFBD97063" box="[948,1122,1361,1385]" italics="true" pageId="10" pageNumber="2624">Eciton buchelli</emphasis>
</taxonomicName>
colony EC #21. Nomadic bivouac site just after emigration; A. Tishechkin’/‘
<typeStatus id="54C48815BE352B682E6AD059FB4B7083" box="[1112,1264,1393,1417]" pageId="10" pageNumber="2624" type="paratype">PARATYPE</typeStatus>
,
<taxonomicName id="4C7F4D34BE352B682B0BD0B9FDC570A3" authority="Hanley" authorityName="Hanley" authorityYear="2003" box="[313,638,1425,1449]" class="Insecta" family="Staphylinidae" genus="Leptandria" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="2624" phylum="Arthropoda" rank="species" species="ecitophila">
<emphasis id="B90BEAA5BE352B682B0BD0B9FD9F70A3" box="[313,548,1425,1449]" italics="true" pageId="10" pageNumber="2624">Leptandria ecitophila</emphasis>
Hanley
</taxonomicName>
, desig.
<collectionCode id="ED6EAE72BE352B6828FCD0B9FD5D70A2" box="[718,742,1425,1448]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="10" pageNumber="2624">R</collectionCode>
. S. Hanley, 2001’ (yellow)];
<specimenCount id="9D79FD3EBE352B682E2CD0B9FB3970A3" box="[1054,1154,1425,1449]" pageId="10" pageNumber="2624" type="male">one male</specimenCount>
(
<collectionCode id="ED6EAE72BE352B682EA2D0B9FB5B70A3" box="[1168,1248,1425,1449]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34264" name="University of Kansas - Biodiversity Institute" pageId="10" pageNumber="2624" type="Museum">SEMC</collectionCode>
), [‘
<emphasis id="B90BEAA5BE352B682B75D098FE6A70C2" bold="true" box="[327,465,1456,1480]" pageId="10" pageNumber="2624">
<collectingCountry id="F3687627BE352B682B75D098FE6A70C2" box="[327,465,1456,1480]" name="Ecuador" pageId="10" pageNumber="2624">ECUADOR</collectingCountry>
</emphasis>
:
<collectingRegion id="49BBF855BE352B682BEDD099FD9970C2" box="[479,546,1457,1480]" country="Ecuador" name="Napo" pageId="10" pageNumber="2624">Napo</collectingRegion>
, mid. Rio Tiputini, Yasuni Res. Stn., 0
<emphasis id="B90BEAA5BE352B6829E9D099FC5E70C3" bold="true" box="[987,997,1457,1481]" pageId="10" pageNumber="2624">°</emphasis>
40.5
<emphasis id="B90BEAA5BE352B682E27D099FBA070C3" bold="true" box="[1045,1051,1457,1481]" pageId="10" pageNumber="2624">∞</emphasis>
<geoCoordinate id="EE4B5070BE352B682E28D099FBCB70C3" box="[1050,1136,1456,1481]" degrees="76" direction="south" minutes="24" orientation="latitude" pageId="10" pageNumber="2624" precision="925" value="-76.4">
S 76
<emphasis id="B90BEAA5BE352B682E79D099FBEE70C3" bold="true" box="[1099,1109,1457,1481]" pageId="10" pageNumber="2624">°</emphasis>
24
</geoCoordinate>
<emphasis id="B90BEAA5BE352B682E5DD099FBCE70C3" bold="true" box="[1135,1141,1457,1481]" pageId="10" pageNumber="2624">∞</emphasis>
W;
<date id="FFC11077BE352B682EACD098FEC970E3" pageId="10" pageNumber="2624" value="1999-06-26">26 June 1999</date>
. AKT #021’/‘
<taxonomicName id="4C7F4D34BE352B682821D0F9FD0170E2" box="[531,698,1488,1512]" class="Insecta" family="Formicidae" genus="Eciton" kingdom="Animalia" order="Hymenoptera" pageId="10" pageNumber="2624" phylum="Arthropoda" rank="species" species="buchelli">
<emphasis id="B90BEAA5BE352B682821D0F9FD0170E2" box="[531,698,1488,1512]" italics="true" pageId="10" pageNumber="2624">Eciton buchelli</emphasis>
</taxonomicName>
colony EC #10. Nomadic bivouac site just after emigration; A. Tishechkin’/‘
<typeStatus id="54C48815BE352B6828BFD0D8FC9E7302" box="[653,805,1520,1544]" pageId="10" pageNumber="2624" type="paratype">PARATYPE</typeStatus>
,
<taxonomicName id="4C7F4D34BE352B68290FD0D8FB267302" authority="Hanley" authorityName="Hanley" authorityYear="2003" box="[829,1181,1520,1544]" class="Insecta" family="Staphylinidae" genus="Leptandria" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="2624" phylum="Arthropoda" rank="species" species="ecitophila">
<emphasis id="B90BEAA5BE352B68290FD0D8FB8E7302" box="[829,1077,1520,1544]" italics="true" pageId="10" pageNumber="2624">Leptandria ecitophila</emphasis>
Hanley
</taxonomicName>
, desig.
<collectionCode id="ED6EAE72BE352B682B0BD339FEEA7322" box="[313,337,1553,1576]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="10" pageNumber="2624">R</collectionCode>
. S. Hanley, 2001’ (yellow)];
<specimenCount id="9D79FD3EBE352B682891D339FCB67322" box="[675,781,1552,1576]" pageId="10" pageNumber="2624" type="male">one male</specimenCount>
(
<collectionCode id="ED6EAE72BE352B682912D338FCCB7322" box="[800,880,1552,1576]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34264" name="University of Kansas - Biodiversity Institute" pageId="10" pageNumber="2624" type="Museum">SEMC</collectionCode>
), [‘
<emphasis id="B90BEAA5BE352B6829AFD338FB9C7322" bold="true" box="[925,1063,1552,1576]" pageId="10" pageNumber="2624">
<collectingCountry id="F3687627BE352B6829AFD338FB9C7322" box="[925,1063,1552,1576]" name="Ecuador" pageId="10" pageNumber="2624">ECUADOR</collectingCountry>
</emphasis>
:
<collectingRegion id="49BBF855BE352B682E0BD339FBC77322" box="[1081,1148,1553,1576]" country="Ecuador" name="Napo" pageId="10" pageNumber="2624">Napo</collectingRegion>
, mid. Rio Tiputini, Yasuni Res. Stn., 0
<emphasis id="B90BEAA5BE352B6828A4D318FD1B7342" bold="true" box="[662,672,1584,1608]" pageId="10" pageNumber="2624">°</emphasis>
40.5
<emphasis id="B90BEAA5BE352B6828FDD318FD6E7342" bold="true" box="[719,725,1584,1608]" pageId="10" pageNumber="2624">∞</emphasis>
<geoCoordinate id="EE4B5070BE352B6828E7D318FC897342" box="[725,818,1584,1608]" degrees="76" direction="south" minutes="24" orientation="latitude" pageId="10" pageNumber="2624" precision="925" value="-76.4">
S 76
<emphasis id="B90BEAA5BE352B68293ED318FCAD7342" bold="true" box="[780,790,1584,1608]" pageId="10" pageNumber="2624">°</emphasis>
24
</geoCoordinate>
<emphasis id="B90BEAA5BE352B682903D318FC8C7342" bold="true" box="[817,823,1584,1608]" pageId="10" pageNumber="2624">∞</emphasis>
W;
<date id="FFC11077BE352B682954D318FBBE7342" box="[870,1029,1584,1608]" pageId="10" pageNumber="2624" value="1999-07-27">27 July 1999</date>
. AKT #103’/‘
<taxonomicName id="4C7F4D34BE352B682E9ED319FE1B7362" baseAuthorityName="Fabricius" baseAuthorityYear="1782" class="Insecta" family="Formicidae" genus="Eciton" kingdom="Animalia" order="Hymenoptera" pageId="10" pageNumber="2624" phylum="Arthropoda" rank="species" species="hamatum">
<emphasis id="B90BEAA5BE352B682E9ED319FE1B7362" italics="true" pageId="10" pageNumber="2624">Eciton hamatum</emphasis>
</taxonomicName>
colony EC #25. Ex. emigration column. A. Tishechkin’/‘
<typeStatus id="54C48815BE352B682E6AD378FB4B7362" box="[1112,1264,1616,1640]" pageId="10" pageNumber="2624" type="paratype">PARATYPE</typeStatus>
,
<taxonomicName id="4C7F4D34BE352B682B0BD358FD2A7382" authority="Hanley" authorityName="Hanley" authorityYear="2003" box="[313,657,1648,1672]" class="Insecta" family="Staphylinidae" genus="Leptandria" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="2624" phylum="Arthropoda" rank="species" species="ecitophila">
<emphasis id="B90BEAA5BE352B682B0BD358FD967382" box="[313,557,1648,1672]" italics="true" pageId="10" pageNumber="2624">Leptandria ecitophila</emphasis>
Hanley
</taxonomicName>
, desig.
<collectionCode id="ED6EAE72BE352B6828C1D358FCB0738D" box="[755,779,1648,1671]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="10" pageNumber="2624">R</collectionCode>
. S. Hanley, 2001’ (yellow)];
<specimenCount id="9D79FD3EBE352B682E40D358FB4F7382" box="[1138,1268,1648,1672]" pageId="10" pageNumber="2624" type="female">one female</specimenCount>
(
<collectionCode id="ED6EAE72BE352B682B73D3B8FE2A73A2" box="[321,401,1680,1704]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34264" name="University of Kansas - Biodiversity Institute" pageId="10" pageNumber="2624" type="Museum">SEMC</collectionCode>
), [‘
<emphasis id="B90BEAA5BE352B682BF6D3B8FDF573A2" bold="true" box="[452,590,1680,1704]" pageId="10" pageNumber="2624">
<collectingCountry id="F3687627BE352B682BF6D3B8FDF573A2" box="[452,590,1680,1704]" name="Ecuador" pageId="10" pageNumber="2624">ECUADOR</collectingCountry>
</emphasis>
:
<collectingRegion id="49BBF855BE352B682854D3B8FD1273AD" box="[614,681,1680,1703]" country="Ecuador" name="Napo" pageId="10" pageNumber="2624">Napo</collectingRegion>
, mid. Rio Tiputini, Yasuni Res. Stn., 0
<emphasis id="B90BEAA5BE352B682E94D3B8FB0B73A2" bold="true" box="[1190,1200,1680,1704]" pageId="10" pageNumber="2624">°</emphasis>
40.5
<emphasis id="B90BEAA5BE352B682ED2D3B8FB5D73A2" bold="true" box="[1248,1254,1680,1704]" pageId="10" pageNumber="2624">∞</emphasis>
<geoCoordinate id="EE4B5070BE352B682ED7D3B8FEC273C2" degrees="76" direction="south" minutes="24" orientation="latitude" pageId="10" pageNumber="2624" precision="925" value="-76.4">
S 76
<emphasis id="B90BEAA5BE352B682B66D398FEE573C2" bold="true" box="[340,350,1712,1736]" pageId="10" pageNumber="2624">°</emphasis>
24
</geoCoordinate>
<emphasis id="B90BEAA5BE352B682B4AD398FEC573C2" bold="true" box="[376,382,1712,1736]" pageId="10" pageNumber="2624">∞</emphasis>
W;
<date id="FFC11077BE352B682B98D398FDF973C2" box="[426,578,1712,1736]" pageId="10" pageNumber="2624" value="1999-07-31">31 July 1999</date>
. AKT #111’/‘
<taxonomicName id="4C7F4D34BE352B6828D1D398FC3173C2" box="[739,906,1712,1736]" class="Insecta" family="Formicidae" genus="Eciton" kingdom="Animalia" order="Hymenoptera" pageId="10" pageNumber="2624" phylum="Arthropoda" rank="species" species="buchelli">
<emphasis id="B90BEAA5BE352B6828D1D398FC3173C2" box="[739,906,1712,1736]" italics="true" pageId="10" pageNumber="2624">Eciton buchelli</emphasis>
</taxonomicName>
colony EC #28. Total bivouac
</paragraph>
<caption id="DF00663FBE362B6B2B0BD355FD4173CD" pageId="9" pageNumber="2623" startId="9.[313,329,1661,1682]" targetBox="[313,1268,723,1635]" targetPageId="9">
<paragraph id="8BC036B7BE362B6B2B0BD355FD4173CD" blockId="9.[313,1268,1660,1735]" pageId="9" pageNumber="2623">
F. 4.
<taxonomicName id="4C7F4D34BE362B6B2BA7D354FDD0739B" authority="Rodney, 2003" authorityName="Rodney" authorityYear="2003" box="[405,619,1660,1681]" class="Insecta" family="Staphylinidae" genus="Leptandria" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="2623" phylum="Arthropoda" rank="species" species="ecitophila" status="sp. nov.">
<emphasis id="B90BEAA5BE362B6B2BA7D354FDD0739B" box="[405,619,1660,1681]" italics="true" pageId="9" pageNumber="2623">Leptandria ecitophila</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23857DEBE362B6B2846D355FD0B7398" box="[628,688,1661,1682]" pageId="9" pageNumber="2623" rank="species">sp. n.</taxonomicNameLabel>
: (A) labrum; (B) mandible, dorsal view; (C) mandible, ventral view; (D) maxilla; (E) labium; (F) hypopharynx; (G) antenna; (H) pronotum; (I) right elytron; (J) tarsus, hind leg.
</paragraph>
</caption>
<caption id="DF00663FBE352B682B0BD112FD497195" pageId="10" pageNumber="2624" startId="10.[313,329,1082,1103]" targetBox="[313,1268,177,1055]" targetPageId="10">
<paragraph id="8BC036B7BE352B682B0BD112FD497195" blockId="10.[313,1268,1081,1183]" pageId="10" pageNumber="2624">
F. 5.
<taxonomicName id="4C7F4D34BE352B682BA7D112FDD57145" authority="Rodney, 2003" authorityName="Rodney" authorityYear="2003" box="[405,622,1082,1103]" class="Insecta" family="Staphylinidae" genus="Leptandria" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="2624" phylum="Arthropoda" rank="species" species="ecitophila" status="sp. nov.">
<emphasis id="B90BEAA5BE352B682BA7D112FDD57145" box="[405,622,1082,1103]" italics="true" pageId="10" pageNumber="2624">Leptandria ecitophila</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23857DEBE352B682848D112FD027145" box="[634,697,1082,1103]" pageId="10" pageNumber="2624" rank="species">sp. n.</taxonomicNameLabel>
: (A) meso- and metathoracic region; (B) abdominal sternum VII (male); (C) left elytron (male); (D) abdominal tergum VII (male); (E) genitalic tract (female), dorsal view; (F) aedeagus, lateral view; (G) paramere, lateral view; (H) spermatheca, lateral view.
</paragraph>
</caption>
<paragraph id="8BC036B7BE342B692B0BD59DFD2B75E6" blockId="11.[313,1269,180,1610]" pageId="11" pageNumber="2625">
<materialsCitation id="3B173CEABE342B692B0BD59DFD3E75E6" collectionCode="R" collectorName="A. Tishechkin' &amp; S. Hanley" pageId="11" pageNumber="2625" specimenCount="1" typeStatus="paratype">
sampling;
<collectorName id="268A5361BE342B692B8FD59DFDC975C7" box="[445,626,181,205]" pageId="11" pageNumber="2625">A. Tishechkin’</collectorName>
/‘
<typeStatus id="54C48815BE342B6928B2D59DFCA375C7" box="[640,792,181,205]" pageId="11" pageNumber="2625" type="paratype">PARATYPE</typeStatus>
,
<taxonomicName id="4C7F4D34BE342B692900D59DFB2275C7" authority="Hanley" authorityName="Hanley" authorityYear="2003" box="[818,1177,181,205]" class="Insecta" family="Staphylinidae" genus="Leptandria" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="2625" phylum="Arthropoda" rank="species" species="ecitophila">
<emphasis id="B90BEAA5BE342B692900D59DFB9575C7" box="[818,1070,181,205]" italics="true" pageId="11" pageNumber="2625">Leptandria ecitophila</emphasis>
Hanley
</taxonomicName>
, desig.
<collectionCode id="ED6EAE72BE342B692B0BD5FDFEEA75E6" box="[313,337,213,236]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="11" pageNumber="2625">R</collectionCode>
.
<collectorName id="268A5361BE342B692B6CD5FCFE6A75E6" box="[350,465,212,236]" pageId="11" pageNumber="2625">S. Hanley</collectorName>
, 2001’ (yellow)
</materialsCitation>
].
</paragraph>
<paragraph id="8BC036B7BE342B692B53D5DCFE5377EA" blockId="11.[313,1269,180,1610]" pageId="11" pageNumber="2625">
<emphasis id="B90BEAA5BE342B692B53D5DCFE6E7401" box="[353,469,244,267]" italics="true" pageId="11" pageNumber="2625">Diagnosis.</emphasis>
This species differs from
<taxonomicName id="4C7F4D34BE342B692922D5DCFCD17401" authorityName="Rodney" authorityYear="2003" box="[784,874,243,267]" class="Insecta" family="Staphylinidae" genus="Leptandria" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="2625" phylum="Arthropoda" rank="species" species="ashei">
<emphasis id="B90BEAA5BE342B692922D5DCFCD17401" box="[784,874,243,267]" italics="true" pageId="11" pageNumber="2625">L. ashei</emphasis>
</taxonomicName>
by having very large eyes, overall length about 0.6–0.7 times the length of the temples (in
<taxonomicName id="4C7F4D34BE342B69298ED43BFBAE7421" authorityName="Rodney" authorityYear="2003" box="[956,1045,275,299]" class="Insecta" family="Staphylinidae" genus="Leptandria" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="2625" phylum="Arthropoda" rank="species" species="ashei">
<emphasis id="B90BEAA5BE342B69298ED43BFBAE7421" box="[956,1045,275,299]" italics="true" pageId="11" pageNumber="2625">L. ashei</emphasis>
</taxonomicName>
, the eyes are small, length about 0.3–0.4 times the length of the temples), by having a more or less quadrate pronotum with the area along the midline distinctly extended posteriorly (in
<taxonomicName id="4C7F4D34BE342B692B56D459FE7B7482" authorityName="Rodney" authorityYear="2003" box="[356,448,368,392]" class="Insecta" family="Staphylinidae" genus="Leptandria" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="2625" phylum="Arthropoda" rank="species" species="ashei">
<emphasis id="B90BEAA5BE342B692B56D459FE7B7482" box="[356,448,368,392]" italics="true" pageId="11" pageNumber="2625">L. ashei</emphasis>
</taxonomicName>
, the pronotum is more or less oval with area along the midline only slightly extended posteriorly), and by having abdominal sternum VIII with a rounded, semilunate duct opening with apical hairs arranged into a mostly pointed process (in
<taxonomicName id="4C7F4D34BE342B692BF4D4E7FD9F74EC" authorityName="Rodney" authorityYear="2003" box="[454,548,462,486]" class="Insecta" family="Staphylinidae" genus="Leptandria" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="2625" phylum="Arthropoda" rank="species" species="ashei">
<emphasis id="B90BEAA5BE342B692BF4D4E7FD9F74EC" box="[454,548,462,486]" italics="true" pageId="11" pageNumber="2625">L. ashei</emphasis>
</taxonomicName>
, abdominal sternum VIII does not possess a modified duct opening).
<taxonomicName id="4C7F4D34BE342B692B9FD4C5FD22770F" authorityName="Hanley" authorityYear="2003" box="[429,665,493,517]" class="Insecta" family="Staphylinidae" genus="Leptandria" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="2625" phylum="Arthropoda" rank="species" species="ecitophila">
<emphasis id="B90BEAA5BE342B692B9FD4C5FD22770F" box="[429,665,493,517]" italics="true" pageId="11" pageNumber="2625">Leptandria ecitophila</emphasis>
</taxonomicName>
differs from
<taxonomicName id="4C7F4D34BE342B692918D4C6FC7B770F" authorityName="Hanley" authorityYear="2003" box="[810,960,493,517]" class="Insecta" family="Staphylinidae" genus="Leptandria" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="2625" phylum="Arthropoda" rank="species" species="tishechkini">
<emphasis id="B90BEAA5BE342B692918D4C6FC7B770F" box="[810,960,493,517]" italics="true" pageId="11" pageNumber="2625">L. tishechkini</emphasis>
</taxonomicName>
by having moderately long antennae, typically reaching the middle of the elytra when extended posteriorly (in
<taxonomicName id="4C7F4D34BE342B692B58D704FDBC774E" authorityName="Hanley" authorityYear="2003" box="[362,519,556,580]" class="Insecta" family="Staphylinidae" genus="Leptandria" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="2625" phylum="Arthropoda" rank="species" species="tishechkini">
<emphasis id="B90BEAA5BE342B692B58D704FDBC774E" box="[362,519,556,580]" italics="true" pageId="11" pageNumber="2625">L. tishechkini</emphasis>
</taxonomicName>
, the antennae are short, generally only reaching the apex of the pronotum when extended posteriorly), by possessing various secondary sexual features in the males (in
<taxonomicName id="4C7F4D34BE342B692843D743FCB67788" authorityName="Hanley" authorityYear="2003" box="[625,781,618,642]" class="Insecta" family="Staphylinidae" genus="Leptandria" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="2625" phylum="Arthropoda" rank="species" species="tishechkini">
<emphasis id="B90BEAA5BE342B692843D743FCB67788" box="[625,781,618,642]" italics="true" pageId="11" pageNumber="2625">L. tishechkini</emphasis>
</taxonomicName>
, males lack distinctive secondary sexual features), and by possessing lobe-like structures in the genitalic tract of the females (in
<taxonomicName id="4C7F4D34BE342B692B59D781FDB277CB" authorityName="Hanley" authorityYear="2003" box="[363,521,681,705]" class="Insecta" family="Staphylinidae" genus="Leptandria" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="2625" phylum="Arthropoda" rank="species" species="tishechkini">
<emphasis id="B90BEAA5BE342B692B59D781FDB277CB" box="[363,521,681,705]" italics="true" pageId="11" pageNumber="2625">L. tishechkini</emphasis>
</taxonomicName>
, females lacking any distinctive modifications to the females genitalic tract).
</paragraph>
<paragraph id="8BC036B7BE342B692B53D7C0FDF870C7" blockId="11.[313,1269,180,1610]" pageId="11" pageNumber="2625">
<emphasis id="B90BEAA5BE342B692B53D7C0FE5377F5" box="[353,488,744,767]" italics="true" pageId="11" pageNumber="2625">Description.</emphasis>
Body length
<quantity id="4C879B52BE342B6928A0D7CFFCA377F5" box="[658,792,743,767]" metricMagnitude="-3" metricUnit="m" metricValue="3.8" metricValueMax="4.4" metricValueMin="3.2" pageId="11" pageNumber="2625" unit="mm" value="3.8" valueMax="4.4" valueMin="3.2">3.2–4.4 mm</quantity>
. Head piceous to black, pronotum, elytra, and abdomen brown to dark brown, lateral margins of pronotum and abdominal terga often tan to light brown. Head, pronotum, and elytra covered throughout with short pile of yellow microsetae; abdomen sparsely covered with piceous to black microsetae; integument distinctly punctured, moderately glossy. Head with eyes very large, length about 0.6–0.7 times length of temples, sparse pubescence primarily directed anteriorly towards midline. Antennae (figure 4G) moderate in length, reaching to middle of elytra when extended posteriorly, widest part at segments 8–11, apical segment acute. Pronotum (figure 4H) more or less quadrate with outer apical angles distinctly sinuate, area along midline extended posteriorly with rounded or sharp corners, pubescence more or less sparse, evenly distributed, primarily directed posteriorly, each lateral margin of disc with six large macrosetae. Elytra (figure 4I) with each elytron slightly longer than wide, outer apical margin distinctly sinuate, surface with setiferous punctures, evenly pubescent, with pubescence directed toward apico-lateral angles. Tarsi (figure 4J) with single, long empodial bristle, length about 1.5 times as long as tarsal claws. Mesosternum (figure 5A) without medial carina. Abdominal sternum VIII with rounded, semilunate duct opening with apical hairs typically arranged into moderately pointed process (figure 5B). Males often with secondary sexual features of extended area along midline of pronotum with sharp corners (figure 4H), small, medioapical denticle on each elytron (figure 5F), conical, longitudinal carina on abdominal tergum VII (figure 5G). Females with lobe-like structure in genitalic tract of abdomen with apical setae (figure 5H). Aedeagus (figure 5C), paramere (figure 5D) spermatheca (figure 5E) as depicted.
</paragraph>
<paragraph id="8BC036B7BE342B692B53D0FCFE077340" blockId="11.[313,1269,180,1610]" pageId="11" pageNumber="2625">
<emphasis id="B90BEAA5BE342B692B53D0FCFE7770E6" box="[353,460,1492,1516]" italics="true" pageId="11" pageNumber="2625">Remarks.</emphasis>
This species was referred to as ‘undescribed species 3’ in the phylogeny portion in Hanley (2002). Adults of
<taxonomicName id="4C7F4D34BE342B6928DCD0DCFCC67306" authorityName="Hanley" authorityYear="2003" box="[750,893,1524,1548]" class="Insecta" family="Staphylinidae" genus="Leptandria" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="2625" phylum="Arthropoda" rank="species" species="ecitophila">
<emphasis id="B90BEAA5BE342B6928DCD0DCFCC67306" box="[750,893,1524,1548]" italics="true" pageId="11" pageNumber="2625">L. ecitophila</emphasis>
</taxonomicName>
were collected from the statary and nomadic bivouac sites of the army ant,
<taxonomicName id="4C7F4D34BE342B692976D33CFC4E7321" baseAuthorityName="Westwood" baseAuthorityYear="1842" box="[836,1013,1555,1579]" class="Insecta" family="Formicidae" genus="Eciton" kingdom="Animalia" order="Hymenoptera" pageId="11" pageNumber="2625" phylum="Arthropoda" rank="species" species="burchelli">
<emphasis id="B90BEAA5BE342B692976D33CFC4E7321" box="[836,1013,1555,1579]" italics="true" pageId="11" pageNumber="2625">Eciton burchelli</emphasis>
</taxonomicName>
, just after emigration of the ants.
</paragraph>
</subSubSection>
</treatment>
</document>
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