840 lines
128 KiB
XML
840 lines
128 KiB
XML
<document id="4A9EF2364B2475B6BDA22C19D19F50BF" ID-CLB-Dataset="27943" ID-DOI="10.11646/zootaxa.4523.1.1" ID-GBIF-Dataset="23903efe-ba7a-438c-a04f-d4b08680b640" ID-ISSN="1175-5326" ID-Zenodo-Dep="2610202" ID-ZooBank="96B7B9E3-9F49-4983-A46C-D29CD6B2EE49" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1553624654688" checkinUser="plazi" docAuthor="Mahony, Stephen, Kamei, Rachunliu G. & Teeling, Emma C." docDate="2018" docId="03D6878AFFE4020DFF73FE9EFD51FF19" docLanguage="en" docName="zootaxa.4523.1.1.pdf" docOrigin="Zootaxa 4523 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Megophrys (Xenophrys) periosa Mahony & Kamei & Teeling 2018, sp. nov." docType="treatment" docVersion="7" lastPageNumber="63" masterDocId="FFEFFFF2FFDE0232FFE4FFD2FFDEFFCC" masterDocTitle="Cryptic diversity within the Megophrys major species group (Amphibia: Megophryidae) of the Asian Horned Frogs: Phylogenetic perspectives and a taxonomic revision of South Asian taxa, with descriptions of four new species" masterLastPageNumber="96" masterPageNumber="1" pageNumber="59" updateTime="1698713983112" updateUser="ExternalLinkService">
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<mods:title id="7431517C1B90B38A27A0C7291DE8A988">Cryptic diversity within the Megophrys major species group (Amphibia: Megophryidae) of the Asian Horned Frogs: Phylogenetic perspectives and a taxonomic revision of South Asian taxa, with descriptions of four new species</mods:title>
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<mods:namePart id="43AF6551650DC93557115AF92DFC9FF7">Mahony, Stephen</mods:namePart>
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<mods:namePart id="069D31D4E6583D1AD9ED246BB3837158">Kamei, Rachunliu G.</mods:namePart>
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<mods:namePart id="4A8CABBCB6A0C6A585AA222821C39199">Teeling, Emma C.</mods:namePart>
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<mods:title id="B08C00E68381A023BCAD33DDA9948031">Zootaxa</mods:title>
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<mods:date id="F8BF59A4561CC43A3FAEB8254755C3B1">2018</mods:date>
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<mods:number id="FE5BA33A3F42D1702816B141513BCBEF">2018-11-19</mods:number>
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<mods:number id="150696085EBCBC6C0912F9090EA1D1B7">4523</mods:number>
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<treatment id="03D6878AFFE4020DFF73FE9EFD51FF19" ID-DOI="http://doi.org/10.5281/zenodo.5958841" ID-GBIF-Taxon="154499351" ID-Zenodo-Dep="5958841" LSID="urn:lsid:plazi:treatment:03D6878AFFE4020DFF73FE9EFD51FF19" httpUri="http://treatment.plazi.org/id/03D6878AFFE4020DFF73FE9EFD51FF19" lastPageId="63" lastPageNumber="63" pageId="58" pageNumber="59">
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<subSubSection id="C3656517FFE40208FF73FE9EFDA1FEAA" box="[151,639,332,358]" pageId="58" pageNumber="59" type="nomenclature">
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<paragraph id="8BC0369CFFE40208FF73FE9EFDA1FEAA" blockId="58.[151,639,332,391]" box="[151,639,332,358]" pageId="58" pageNumber="59">
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<heading id="D08881F0FFE40208FF73FE9EFDA1FEAA" bold="true" box="[151,639,332,358]" fontSize="11" level="1" pageId="58" pageNumber="59" reason="1">
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<emphasis id="B90BEA8EFFE40208FF73FE9EFDA1FEAA" bold="true" box="[151,639,332,358]" pageId="58" pageNumber="59">
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<taxonomicName id="4C7F4D1FFFE40208FF73FE9EFDC4FEAA" authority="Mahony & Kamei & Teeling, 2018" authorityName="Mahony & Kamei & Teeling" authorityYear="2018" box="[151,538,332,358]" class="Amphibia" family="Megophryidae" genus="Megophrys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="58" pageNumber="59" phylum="Chordata" rank="subGenus" status="sp. nov." subGenus="Xenophrys">
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<emphasis id="B90BEA8EFFE40208FF73FE9EFEC0FEAA" bold="true" box="[151,286,332,358]" italics="true" pageId="58" pageNumber="59">Megophrys</emphasis>
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(
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<emphasis id="B90BEA8EFFE40208FECBFE9EFE6FFEAA" bold="true" box="[303,433,332,358]" italics="true" pageId="58" pageNumber="59">Xenophrys</emphasis>
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)
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<emphasis id="B90BEA8EFFE40208FE26FE9EFDC4FEAA" bold="true" box="[450,538,332,358]" italics="true" pageId="58" pageNumber="59">periosa</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A23857F5FFE40208FDC6FE9EFDA1FEAA" box="[546,639,332,358]" pageId="58" pageNumber="59" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C3656517FFE40208FF73FEBCFE1CFE4B" box="[151,450,366,391]" pageId="58" pageNumber="59" type="description">
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<paragraph id="8BC0369CFFE40208FF73FEBCFE1CFE4B" blockId="58.[151,639,332,391]" box="[151,450,366,391]" pageId="58" pageNumber="59">
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(
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<figureCitation id="13442A19FFE40208FF7AFEBCFEC9FE4B" box="[158,279,366,391]" captionStart="FIGURE 22" captionStartId="59.[151,250,1767,1789]" captionTargetBox="[151,1436,301,1746]" captionTargetId="figure@59.[151,1436,301,1746]" captionTargetPageId="59" captionText="FIGURE 22. Megophrys periosa sp. nov. holotype: adult male (BNHS 6055: SVL 93.8 mm) in preservation: A. dorsal view; B. ventral view; C. profile view of head; D. ventral view of hand; E. ventral view of foot." httpUri="https://zenodo.org/record/2610246/files/figure.png" pageId="58" pageNumber="59">Figures 22</figureCitation>
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&
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<figureCitation id="13442A19FFE40208FEDFFEBDFE84FE4B" box="[315,346,367,391]" captionStart="FIGURE 23" captionStartId="61.[151,250,1796,1818]" captionTargetBox="[240,1344,193,1781]" captionTargetId="figure@61.[240,1344,193,1782]" captionTargetPageId="61" captionText="FIGURE 23. Megophrys periosa sp. nov.: A. lateral view of adult male holotype (BNHS 6055: SVL 93.8 mm) in life. B. lateral view of adult male paratype (BNHS 6057: SVL 80.7 mm) in life. C–E. dorsolateral views of adult males in life showing typical variation: C. referred specimen (SDBDU 2009.1190: SVL 73.6 mm); D. referred specimen (SDBDU 2009.1189: SVL 71.3 mm); E. paratype (BNHS 6061: SVL 87.7 mm). F. dorsolateral view of adult female paratype (BNHS 6056: SVL 112.0 mm) in life. G. ventral view of two adult male paratypes immediately after euthanisation, showing variation of markings (left, BNHS 6064: SVL 88.8 mm; right BNHS 6063: SVL 81.3 mm). H. ventral view of three adult male specimens in preservation showing variation of makings (left, paratype BNHS 6059: SVL 81.9 mm; centre, paratype BNHS 6060: SVL 80.2 mm; right, holotype BNHS 6055: SVL 93.8 mm)." httpUri="https://zenodo.org/record/2610248/files/figure.png" pageId="58" pageNumber="59">23</figureCitation>
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;
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<tableCitation id="C6FD0327FFE40208FE82FEBCFE69FE4B" box="[358,439,366,391]" captionStart="TABLE 1" captionStartId="22.[147,225,525,545]" captionText="TABLE 1. Morphometric details for specimens new and known included in the Taxonomic review section. Refer to Methods and materials section for explanation of measurement abbreviations and museum acronyms. HT = holotype; PT = paratype; LT = lectotype; PLT = paralectotype; NT = neotype; RS = referred specimen; M = adult male; F = adult female; SA = subadult; J = juvenile [post metamorphic]; † = BMNH voucher numbers; # = SDBDU voucher numbers; ind = character limits insufficiently distinct to reliably measure." pageId="58" pageNumber="59">Table 1</tableCitation>
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)
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</paragraph>
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<subSubSection id="C3656517FFE4020EFF73FE64FEECFBB1" lastPageId="60" lastPageNumber="61" pageId="58" pageNumber="59" type="materials_examined">
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<paragraph id="8BC0369CFFE40208FF73FE64FBE0FDDB" blockId="58.[151,1437,438,2011]" pageId="58" pageNumber="59">
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<materialsCitation id="3B173CC1FFE40208FF73FE64FBE0FDDB" ID-GBIF-Occurrence="2028521246" collectingDate="2009-07-26" collectionCode="BNHS, SDBDU" country="India" county="University" elevation="450" latitude="28.209435" location="Pangin town" longLatPrecision="1" longitude="94.986115" municipality="Systematics Lab" pageId="58" pageNumber="59" specimenCode="BNHS 6055, SDBDU 2009.793" specimenCount="1" stateProvince="Arunachal Pradesh" typeStatus="holotype">
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<emphasis id="B90BEA8EFFE40208FF73FE64FED5FE03" bold="true" box="[151,267,438,463]" pageId="58" pageNumber="59">
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<typeStatus id="54C4883EFFE40208FF73FE64FED8FE03" box="[151,262,438,463]" pageId="58" pageNumber="59" type="holotype">Holotype</typeStatus>
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.
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</emphasis>
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||
Adult male (
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<specimenCode id="DBD99EE7FFE40208FE42FE65FDEDFE03" box="[422,563,438,463]" collectionCode="BNHS" country="India" httpUri="http://grbio.org/cool/8g20-8f2r" name="Bombay Natural History Society" pageId="58" pageNumber="59">BNHS 6055</specimenCode>
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[field no.
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<specimenCode id="DBD99EE7FFE40208FD54FE65FC56FE03" box="[688,904,439,463]" collectionCode="SDBDU" pageId="58" pageNumber="59">SDBDU 2009.793</specimenCode>
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]:
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<figureCitation id="13442A19FFE40208FC78FE64FBC6FE03" box="[924,1048,438,463]" captionStart="FIGURE 22" captionStartId="59.[151,250,1767,1789]" captionTargetBox="[151,1436,301,1746]" captionTargetId="figure@59.[151,1436,301,1746]" captionTargetPageId="59" captionText="FIGURE 22. Megophrys periosa sp. nov. holotype: adult male (BNHS 6055: SVL 93.8 mm) in preservation: A. dorsal view; B. ventral view; C. profile view of head; D. ventral view of hand; E. ventral view of foot." httpUri="https://zenodo.org/record/2610246/files/figure.png" pageId="58" pageNumber="59">Figures 22</figureCitation>
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, & 23A & H), from
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<location id="8EA06047FFE40208FAE9FE64FA42FE02" LSID="urn:lsid:plazi:treatment:03D6878AFFE4020DFF73FE9EFD51FF19:8EA06047FFE40208FAE9FE64FA42FE02" box="[1293,1436,438,463]" country="India" county="University" latitude="28.209435" longLatPrecision="1" longitude="94.986115" municipality="Systematics Lab" name="Pangin town" pageId="58" pageNumber="59" stateProvince="Arunachal Pradesh">Pangin town</location>
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(
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,
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<geoCoordinate id="EE4B505BFFE40208FEB9FE08FDDAFE3F" box="[349,516,474,499]" direction="east" orientation="longitude" pageId="58" pageNumber="59" precision="1" value="94.986115">94°59'10.02"E</geoCoordinate>
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,
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<elevation id="0052D1AFFFE40208FDF3FE08FD46FE3F" box="[535,664,474,499]" metricMagnitude="2" metricUnit="m" metricValue="4.5" pageId="58" pageNumber="59" unit="m" value="450.0">
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asl
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</elevation>
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.),
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<location id="8EA06047FFE40208FD5EFE09FC47FE3F" LSID="urn:lsid:plazi:treatment:03D6878AFFE4020DFF73FE9EFD51FF19:8EA06047FFE40208FD5EFE09FC47FE3F" box="[698,921,474,499]" country="India" county="University" latitude="28.209435" longLatPrecision="1" longitude="94.986115" municipality="Systematics Lab" name="East Siang district" pageId="58" pageNumber="59" stateProvince="Arunachal Pradesh">East Siang district</location>
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,
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<collectingRegion id="49BBF87EFFE40208FC4FFE08FB56FE3F" box="[939,1160,474,499]" country="India" name="Arunachal Pradesh" pageId="58" pageNumber="59">Arunachal Pradesh</collectingRegion>
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state,
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<location id="8EA06047FFE40208FB04FE08FA93FE3F" LSID="urn:lsid:plazi:treatment:03D6878AFFE4020DFF73FE9EFD51FF19:8EA06047FFE40208FB04FE08FA93FE3F" box="[1248,1357,474,499]" country="India" county="University" latitude="28.209435" longLatPrecision="1" longitude="94.986115" municipality="Systematics Lab" name="Northeast" pageId="58" pageNumber="59" stateProvince="Arunachal Pradesh">Northeast</location>
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<collectingCountry id="F368760CFFE40208FAB8FE08FA46FE3F" box="[1372,1432,474,499]" name="India" pageId="58" pageNumber="59">India</collectingCountry>
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, collected by members of the
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<collectingMunicipality id="6BA4ACE6FFE40208FE3FFE2CFD48FDDB" box="[475,662,510,535]" pageId="58" pageNumber="59">Systematics Lab</collectingMunicipality>
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,
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<collectingCounty id="62A14E10FFE40208FD46FE2CFCC6FDDB" box="[674,792,510,535]" pageId="58" pageNumber="59">University</collectingCounty>
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of
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<collectingRegion id="49BBF87EFFE40208FCD9FE2CFCA5FDDB" box="[829,891,510,535]" country="India" name="Delhi" pageId="58" pageNumber="59">Delhi</collectingRegion>
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on
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<collectingDate id="EF85E9B4FFE40208FC42FE2CFBE4FDDB" box="[934,1082,510,535]" pageId="58" pageNumber="59" value="2009-07-26">
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<date id="FFC1105CFFE40208FC42FE2CFBE4FDDB" box="[934,1082,510,535]" pageId="58" pageNumber="59" value="2009-07-26">26 July 2009</date>
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</collectingDate>
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.
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</materialsCitation>
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</paragraph>
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<paragraph id="8BC0369CFFE40208FF23FDF0FB0FFCDF" blockId="58.[151,1437,438,2011]" pageId="58" pageNumber="59">
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<materialsCitation id="3B173CC1FFE40208FF23FDF0FD4EFD4F" ID-GBIF-Occurrence="2028521239" collectingDate="2009-07-26" collectingDateMax="2009-07-28" collectingDateMin="2009-07-26" collectionCode="BNHS, SDBDU" location="Systematics Lab" municipality="University" pageId="58" pageNumber="59" specimenCode="BNHS 6057, SDBDU 2009.794, BNHS 6058-6060, SDBDU 2009.1170, 1172" specimenCount="2" stateProvince="Delhi" typeStatus="paratype">
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<emphasis id="B90BEA8EFFE40208FF23FDF0FE98FDF7" bold="true" box="[199,326,546,571]" pageId="58" pageNumber="59">
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<typeStatus id="54C4883EFFE40208FF23FDF0FE9FFDF7" box="[199,321,546,571]" pageId="58" pageNumber="59" type="paratype">Paratypes</typeStatus>
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.
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</emphasis>
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Four adult males (
|
||
<specimenCode id="DBD99EE7FFE40208FDF3FDF1FD7CFDF7" box="[535,674,546,571]" collectionCode="BNHS" country="India" httpUri="http://grbio.org/cool/8g20-8f2r" name="Bombay Natural History Society" pageId="58" pageNumber="59">BNHS 6057</specimenCode>
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[field no.
|
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<specimenCode id="DBD99EE7FFE40208FCF2FDF1FC35FDF7" box="[790,1003,547,571]" collectionCode="SDBDU" pageId="58" pageNumber="59">SDBDU 2009.794</specimenCode>
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]:
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<figureCitation id="13442A19FFE40208FC18FDF0FBA0FDF7" box="[1020,1150,546,571]" captionStart="FIGURE 23" captionStartId="61.[151,250,1796,1818]" captionTargetBox="[240,1344,193,1781]" captionTargetId="figure@61.[240,1344,193,1782]" captionTargetPageId="61" captionText="FIGURE 23. Megophrys periosa sp. nov.: A. lateral view of adult male holotype (BNHS 6055: SVL 93.8 mm) in life. B. lateral view of adult male paratype (BNHS 6057: SVL 80.7 mm) in life. C–E. dorsolateral views of adult males in life showing typical variation: C. referred specimen (SDBDU 2009.1190: SVL 73.6 mm); D. referred specimen (SDBDU 2009.1189: SVL 71.3 mm); E. paratype (BNHS 6061: SVL 87.7 mm). F. dorsolateral view of adult female paratype (BNHS 6056: SVL 112.0 mm) in life. G. ventral view of two adult male paratypes immediately after euthanisation, showing variation of markings (left, BNHS 6064: SVL 88.8 mm; right BNHS 6063: SVL 81.3 mm). H. ventral view of three adult male specimens in preservation showing variation of makings (left, paratype BNHS 6059: SVL 81.9 mm; centre, paratype BNHS 6060: SVL 80.2 mm; right, holotype BNHS 6055: SVL 93.8 mm)." httpUri="https://zenodo.org/record/2610248/files/figure.png" pageId="58" pageNumber="59">Figure 23B</figureCitation>
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||
;
|
||
<specimenCode id="DBD99EE7FFE40208FB6DFDF1FA87FDF7" box="[1161,1369,546,571]" collectionCode="BNHS" country="India" httpUri="http://grbio.org/cool/8g20-8f2r" name="Bombay Natural History Society" pageId="58" pageNumber="59">BNHS 6058–6060</specimenCode>
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||
[field nos.
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<specimenCode id="DBD99EE7FFE40208FF2AFD95FE6DFD93" box="[206,435,583,607]" collectionCode="SDBDU" pageId="58" pageNumber="59">SDBDU 2009.1170</specimenCode>
|
||
–
|
||
<specimenCode id="DBD99EE7FFE40208FE26FD95FE27FD93" box="[450,505,583,607]" collectionCode="SDBDU" pageId="58" pageNumber="59">1172</specimenCode>
|
||
]:
|
||
<figureCitation id="13442A19FFE40208FDE9FD94FD4DFD93" box="[525,659,582,607]" captionStart="FIGURE 23" captionStartId="61.[151,250,1796,1818]" captionTargetBox="[240,1344,193,1781]" captionTargetId="figure@61.[240,1344,193,1782]" captionTargetPageId="61" captionText="FIGURE 23. Megophrys periosa sp. nov.: A. lateral view of adult male holotype (BNHS 6055: SVL 93.8 mm) in life. B. lateral view of adult male paratype (BNHS 6057: SVL 80.7 mm) in life. C–E. dorsolateral views of adult males in life showing typical variation: C. referred specimen (SDBDU 2009.1190: SVL 73.6 mm); D. referred specimen (SDBDU 2009.1189: SVL 71.3 mm); E. paratype (BNHS 6061: SVL 87.7 mm). F. dorsolateral view of adult female paratype (BNHS 6056: SVL 112.0 mm) in life. G. ventral view of two adult male paratypes immediately after euthanisation, showing variation of markings (left, BNHS 6064: SVL 88.8 mm; right BNHS 6063: SVL 81.3 mm). H. ventral view of three adult male specimens in preservation showing variation of makings (left, paratype BNHS 6059: SVL 81.9 mm; centre, paratype BNHS 6060: SVL 80.2 mm; right, holotype BNHS 6055: SVL 93.8 mm)." httpUri="https://zenodo.org/record/2610248/files/figure.png" pageId="58" pageNumber="59">Figure 23H</figureCitation>
|
||
), collected along with the
|
||
<typeStatus id="54C4883EFFE40208FC2CFD94FBF4FD93" box="[968,1066,582,607]" pageId="58" pageNumber="59" type="holotype">holotype</typeStatus>
|
||
by members of the
|
||
<location id="8EA06047FFE40208FAF3FD94FF1BFD4F" LSID="urn:lsid:plazi:treatment:03D6878AFFE4020DFF73FE9EFD51FF19:8EA06047FFE40208FAF3FD94FF1BFD4F" municipality="University" name="Systematics Lab" pageId="58" pageNumber="59" stateProvince="Delhi">Systematics Lab</location>
|
||
,
|
||
<collectingMunicipality id="6BA4ACE6FFE40208FF34FDB8FE98FD4F" box="[208,326,618,643]" pageId="58" pageNumber="59">University</collectingMunicipality>
|
||
of
|
||
<collectingRegion id="49BBF87EFFE40208FE8FFDB8FE77FD4F" box="[363,425,618,643]" country="India" name="Delhi" pageId="58" pageNumber="59">Delhi</collectingRegion>
|
||
on
|
||
<collectingDate id="EF85E9B4FFE40208FE37FDB8FD4EFD4F" box="[467,656,618,643]" pageId="58" pageNumber="59" value="2009-07-26" valueMax="2009-07-28" valueMin="2009-07-26">
|
||
<date id="FFC1105CFFE40208FE37FDB8FD4EFD4F" box="[467,656,618,643]" pageId="58" pageNumber="59" value="2009-07-26" valueMax="2009-07-28" valueMin="2009-07-26">26–28 July 2009</date>
|
||
</collectingDate>
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="3B173CC1FFE40208FD78FDB8FB10FCDF" ID-GBIF-Occurrence="2028521211" collectingDate="2009-08-07" collectingDateMax="2009-08-09" collectingDateMin="2009-08-07" collectionCode="BNHS, SDBDU" country="India" county="University" elevation="1110" latitude="27.101116" location="Sessa River" longLatPrecision="1" longitude="92.52737" municipality="Systematics Lab" pageId="58" pageNumber="59" specimenCode="BNHS 6061, SDBDU 2009.1243, BNHS 6062, SDBDU 2009.1244, BNHS 6063-6064, SDBDU 2009.1265, 1266, BNHS 6056, SDBDU 2009.1285" specimenCount="1" stateProvince="Arunachal Pradesh" typeStatus="paratype">
|
||
four adult males (
|
||
<specimenCode id="DBD99EE7FFE40208FC84FDB9FC35FD4F" box="[864,1003,618,643]" collectionCode="BNHS" country="India" httpUri="http://grbio.org/cool/8g20-8f2r" name="Bombay Natural History Society" pageId="58" pageNumber="59">BNHS 6061</specimenCode>
|
||
[field no.
|
||
<specimenCode id="DBD99EE7FFE40208FBBBFDB9FA9DFD4F" box="[1119,1347,619,643]" collectionCode="SDBDU" pageId="58" pageNumber="59">SDBDU 2009.1243</specimenCode>
|
||
]:
|
||
<figureCitation id="13442A19FFE40208FAB7FDB8FF19FD6B" captionStart="FIGURE 23" captionStartId="61.[151,250,1796,1818]" captionTargetBox="[240,1344,193,1781]" captionTargetId="figure@61.[240,1344,193,1782]" captionTargetPageId="61" captionText="FIGURE 23. Megophrys periosa sp. nov.: A. lateral view of adult male holotype (BNHS 6055: SVL 93.8 mm) in life. B. lateral view of adult male paratype (BNHS 6057: SVL 80.7 mm) in life. C–E. dorsolateral views of adult males in life showing typical variation: C. referred specimen (SDBDU 2009.1190: SVL 73.6 mm); D. referred specimen (SDBDU 2009.1189: SVL 71.3 mm); E. paratype (BNHS 6061: SVL 87.7 mm). F. dorsolateral view of adult female paratype (BNHS 6056: SVL 112.0 mm) in life. G. ventral view of two adult male paratypes immediately after euthanisation, showing variation of markings (left, BNHS 6064: SVL 88.8 mm; right BNHS 6063: SVL 81.3 mm). H. ventral view of three adult male specimens in preservation showing variation of makings (left, paratype BNHS 6059: SVL 81.9 mm; centre, paratype BNHS 6060: SVL 80.2 mm; right, holotype BNHS 6055: SVL 93.8 mm)." httpUri="https://zenodo.org/record/2610248/files/figure.png" pageId="58" pageNumber="59">Figure 23E</figureCitation>
|
||
;
|
||
<specimenCode id="DBD99EE7FFE40208FF37FD5DFE83FD6B" box="[211,349,654,679]" collectionCode="BNHS" country="India" httpUri="http://grbio.org/cool/8g20-8f2r" name="Bombay Natural History Society" pageId="58" pageNumber="59">BNHS 6062</specimenCode>
|
||
[field no.
|
||
<specimenCode id="DBD99EE7FFE40208FE36FD5DFD6BFD6B" box="[466,693,655,679]" collectionCode="SDBDU" pageId="58" pageNumber="59">SDBDU 2009.1244</specimenCode>
|
||
],
|
||
<specimenCode id="DBD99EE7FFE40208FD23FD5DFC49FD6B" box="[711,919,654,679]" collectionCode="BNHS" country="India" httpUri="http://grbio.org/cool/8g20-8f2r" name="Bombay Natural History Society" pageId="58" pageNumber="59">BNHS 6063–6064</specimenCode>
|
||
[field nos.
|
||
<specimenCode id="DBD99EE7FFE40208FBFCFD5DFB25FD6B" box="[1048,1275,654,679]" collectionCode="SDBDU" pageId="58" pageNumber="59">SDBDU 2009.1265</specimenCode>
|
||
–
|
||
<specimenCode id="DBD99EE7FFE40208FAEEFD5CFA9CFD6B" box="[1290,1346,654,679]" collectionCode="SDBDU" pageId="58" pageNumber="59">1266</specimenCode>
|
||
]:
|
||
<figureCitation id="13442A19FFE40208FAB7FD5CFF14FD07" captionStart="FIGURE 23" captionStartId="61.[151,250,1796,1818]" captionTargetBox="[240,1344,193,1781]" captionTargetId="figure@61.[240,1344,193,1782]" captionTargetPageId="61" captionText="FIGURE 23. Megophrys periosa sp. nov.: A. lateral view of adult male holotype (BNHS 6055: SVL 93.8 mm) in life. B. lateral view of adult male paratype (BNHS 6057: SVL 80.7 mm) in life. C–E. dorsolateral views of adult males in life showing typical variation: C. referred specimen (SDBDU 2009.1190: SVL 73.6 mm); D. referred specimen (SDBDU 2009.1189: SVL 71.3 mm); E. paratype (BNHS 6061: SVL 87.7 mm). F. dorsolateral view of adult female paratype (BNHS 6056: SVL 112.0 mm) in life. G. ventral view of two adult male paratypes immediately after euthanisation, showing variation of markings (left, BNHS 6064: SVL 88.8 mm; right BNHS 6063: SVL 81.3 mm). H. ventral view of three adult male specimens in preservation showing variation of makings (left, paratype BNHS 6059: SVL 81.9 mm; centre, paratype BNHS 6060: SVL 80.2 mm; right, holotype BNHS 6055: SVL 93.8 mm)." httpUri="https://zenodo.org/record/2610248/files/figure.png" pageId="58" pageNumber="59">Figure 23G</figureCitation>
|
||
), and one adult female (
|
||
<specimenCode id="DBD99EE7FFE40208FE31FD61FDBFFD07" box="[469,609,690,715]" collectionCode="BNHS" country="India" httpUri="http://grbio.org/cool/8g20-8f2r" name="Bombay Natural History Society" pageId="58" pageNumber="59">BNHS 6056</specimenCode>
|
||
[field no.
|
||
<specimenCode id="DBD99EE7FFE40208FD30FD61FC66FD07" box="[724,952,690,715]" collectionCode="SDBDU" pageId="58" pageNumber="59">SDBDU 2009.1285</specimenCode>
|
||
]:
|
||
<figureCitation id="13442A19FFE40208FC2CFD60FB99FD07" box="[968,1095,690,715]" captionStart="FIGURE 23" captionStartId="61.[151,250,1796,1818]" captionTargetBox="[240,1344,193,1781]" captionTargetId="figure@61.[240,1344,193,1782]" captionTargetPageId="61" captionText="FIGURE 23. Megophrys periosa sp. nov.: A. lateral view of adult male holotype (BNHS 6055: SVL 93.8 mm) in life. B. lateral view of adult male paratype (BNHS 6057: SVL 80.7 mm) in life. C–E. dorsolateral views of adult males in life showing typical variation: C. referred specimen (SDBDU 2009.1190: SVL 73.6 mm); D. referred specimen (SDBDU 2009.1189: SVL 71.3 mm); E. paratype (BNHS 6061: SVL 87.7 mm). F. dorsolateral view of adult female paratype (BNHS 6056: SVL 112.0 mm) in life. G. ventral view of two adult male paratypes immediately after euthanisation, showing variation of markings (left, BNHS 6064: SVL 88.8 mm; right BNHS 6063: SVL 81.3 mm). H. ventral view of three adult male specimens in preservation showing variation of makings (left, paratype BNHS 6059: SVL 81.9 mm; centre, paratype BNHS 6060: SVL 80.2 mm; right, holotype BNHS 6055: SVL 93.8 mm)." httpUri="https://zenodo.org/record/2610248/files/figure.png" pageId="58" pageNumber="59">Figure 23F</figureCitation>
|
||
), from the
|
||
<location id="8EA06047FFE40208FB25FD61FA98FD07" LSID="urn:lsid:plazi:treatment:03D6878AFFE4020DFF73FE9EFD51FF19:8EA06047FFE40208FB25FD61FA98FD07" box="[1217,1350,690,715]" country="India" county="University" latitude="27.101116" longLatPrecision="1" longitude="92.52737" municipality="Systematics Lab" name="Sessa River" pageId="58" pageNumber="59" stateProvince="Arunachal Pradesh">Sessa River</location>
|
||
, nearby
|
||
<location id="8EA06047FFE40208FF73FD05FEF6FD23" LSID="urn:lsid:plazi:treatment:03D6878AFFE4020DFF73FE9EFD51FF19:8EA06047FFE40208FF73FD05FEF6FD23" box="[151,296,726,751]" country="India" county="University" latitude="27.101116" longLatPrecision="1" longitude="92.52737" municipality="Systematics Lab" name="Sessa village" pageId="58" pageNumber="59" stateProvince="Arunachal Pradesh">Sessa village</location>
|
||
(
|
||
<geoCoordinate id="EE4B505BFFE40208FED2FD04FE1BFD23" box="[310,453,726,751]" direction="north" orientation="latitude" pageId="58" pageNumber="59" precision="1" value="27.101116">27°6'4.02"N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE4B505BFFE40208FE34FD04FDA8FD23" box="[464,630,726,751]" direction="east" orientation="longitude" pageId="58" pageNumber="59" precision="1" value="92.52737">92°31'38.52"E</geoCoordinate>
|
||
,
|
||
<elevation id="0052D1AFFFE40208FD65FD05FD23FD23" box="[641,765,726,751]" metricMagnitude="3" metricUnit="m" metricValue="1.1099999999999999" pageId="58" pageNumber="59" unit="m" value="1110.0">
|
||
<quantity id="4C879B79FFE40208FD65FD05FD0DFD22" box="[641,723,727,751]" metricMagnitude="3" metricUnit="m" metricValue="1.1099999999999999" pageId="58" pageNumber="59" unit="m" value="1110.0">1110 m</quantity>
|
||
asl
|
||
</elevation>
|
||
.),
|
||
<location id="8EA06047FFE40208FCF2FD05FBD4FD23" LSID="urn:lsid:plazi:treatment:03D6878AFFE4020DFF73FE9EFD51FF19:8EA06047FFE40208FCF2FD05FBD4FD23" box="[790,1034,726,751]" country="India" county="University" latitude="27.101116" longLatPrecision="1" longitude="92.52737" municipality="Systematics Lab" name="West Kameng district" pageId="58" pageNumber="59" stateProvince="Arunachal Pradesh">West Kameng district</location>
|
||
,
|
||
<collectingRegion id="49BBF87EFFE40208FBF0FD04FB37FD23" box="[1044,1257,726,751]" country="India" name="Arunachal Pradesh" pageId="58" pageNumber="59">Arunachal Pradesh</collectingRegion>
|
||
state,
|
||
<location id="8EA06047FFE40208FACBFD04FA42FD23" LSID="urn:lsid:plazi:treatment:03D6878AFFE4020DFF73FE9EFD51FF19:8EA06047FFE40208FACBFD04FA42FD23" box="[1327,1436,726,751]" country="India" county="University" latitude="27.101116" longLatPrecision="1" longitude="92.52737" municipality="Systematics Lab" name="Northeast" pageId="58" pageNumber="59" stateProvince="Arunachal Pradesh">Northeast</location>
|
||
<collectingCountry id="F368760CFFE40208FF73FD28FF0DFCDF" box="[151,211,762,787]" name="India" pageId="58" pageNumber="59">India</collectingCountry>
|
||
, collected by members of the
|
||
<collectingMunicipality id="6BA4ACE6FFE40208FDC7FD28FD00FCDF" box="[547,734,762,787]" pageId="58" pageNumber="59">Systematics Lab</collectingMunicipality>
|
||
,
|
||
<collectingCounty id="62A14E10FFE40208FD0DFD28FC81FCDF" box="[745,863,762,787]" pageId="58" pageNumber="59">University</collectingCounty>
|
||
of
|
||
<collectingRegion id="49BBF87EFFE40208FC61FD28FC1DFCDF" box="[901,963,762,787]" country="India" name="Delhi" pageId="58" pageNumber="59">Delhi</collectingRegion>
|
||
on
|
||
<collectingDate id="EF85E9B4FFE40208FC09FD29FB10FCDF" box="[1005,1230,763,787]" pageId="58" pageNumber="59" value="2009-08-07" valueMax="2009-08-09" valueMin="2009-08-07">
|
||
<date id="FFC1105CFFE40208FC09FD29FB10FCDF" box="[1005,1230,763,787]" pageId="58" pageNumber="59" value="2009-08-07" valueMax="2009-08-09" valueMin="2009-08-07">07–09 August 2009</date>
|
||
</collectingDate>
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BC0369CFFE40208FF23FCCCFE3AFC27" blockId="58.[151,1437,438,2011]" pageId="58" pageNumber="59">
|
||
<emphasis id="B90BEA8EFFE40208FF23FCCCFE62FCFB" bold="true" box="[199,444,798,823]" pageId="58" pageNumber="59">Referred specimens.</emphasis>
|
||
Two adult males (SDBDU 2009.1189:
|
||
<figureCitation id="13442A19FFE40208FC62FCCCFBD2FCFB" box="[902,1036,798,823]" captionStart="FIGURE 23" captionStartId="61.[151,250,1796,1818]" captionTargetBox="[240,1344,193,1781]" captionTargetId="figure@61.[240,1344,193,1782]" captionTargetPageId="61" captionText="FIGURE 23. Megophrys periosa sp. nov.: A. lateral view of adult male holotype (BNHS 6055: SVL 93.8 mm) in life. B. lateral view of adult male paratype (BNHS 6057: SVL 80.7 mm) in life. C–E. dorsolateral views of adult males in life showing typical variation: C. referred specimen (SDBDU 2009.1190: SVL 73.6 mm); D. referred specimen (SDBDU 2009.1189: SVL 71.3 mm); E. paratype (BNHS 6061: SVL 87.7 mm). F. dorsolateral view of adult female paratype (BNHS 6056: SVL 112.0 mm) in life. G. ventral view of two adult male paratypes immediately after euthanisation, showing variation of markings (left, BNHS 6064: SVL 88.8 mm; right BNHS 6063: SVL 81.3 mm). H. ventral view of three adult male specimens in preservation showing variation of makings (left, paratype BNHS 6059: SVL 81.9 mm; centre, paratype BNHS 6060: SVL 80.2 mm; right, holotype BNHS 6055: SVL 93.8 mm)." httpUri="https://zenodo.org/record/2610248/files/figure.png" pageId="58" pageNumber="59">Figure 23D</figureCitation>
|
||
; SDBDU 2009.1190:
|
||
<figureCitation id="13442A19FFE40208FAEFFCCCFA51FCFB" box="[1291,1423,798,823]" captionStart="FIGURE 23" captionStartId="61.[151,250,1796,1818]" captionTargetBox="[240,1344,193,1781]" captionTargetId="figure@61.[240,1344,193,1782]" captionTargetPageId="61" captionText="FIGURE 23. Megophrys periosa sp. nov.: A. lateral view of adult male holotype (BNHS 6055: SVL 93.8 mm) in life. B. lateral view of adult male paratype (BNHS 6057: SVL 80.7 mm) in life. C–E. dorsolateral views of adult males in life showing typical variation: C. referred specimen (SDBDU 2009.1190: SVL 73.6 mm); D. referred specimen (SDBDU 2009.1189: SVL 71.3 mm); E. paratype (BNHS 6061: SVL 87.7 mm). F. dorsolateral view of adult female paratype (BNHS 6056: SVL 112.0 mm) in life. G. ventral view of two adult male paratypes immediately after euthanisation, showing variation of markings (left, BNHS 6064: SVL 88.8 mm; right BNHS 6063: SVL 81.3 mm). H. ventral view of three adult male specimens in preservation showing variation of makings (left, paratype BNHS 6059: SVL 81.9 mm; centre, paratype BNHS 6060: SVL 80.2 mm; right, holotype BNHS 6055: SVL 93.8 mm)." httpUri="https://zenodo.org/record/2610248/files/figure.png" pageId="58" pageNumber="59">Figure 23C</figureCitation>
|
||
), from Rigo Village (
|
||
<geoCoordinate id="EE4B505BFFE40208FE95FC90FDD3FC97" box="[369,525,834,859]" direction="north" orientation="latitude" pageId="58" pageNumber="59" precision="1" value="28.1596">28°9'34.56"N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE4B505BFFE40208FDFDFC91FD1EFC97" box="[537,704,835,859]" direction="east" orientation="longitude" pageId="58" pageNumber="59" precision="1" value="94.78872">94°47'19.38"E</geoCoordinate>
|
||
,
|
||
<quantity id="4C879B79FFE40208FD2EFC90FCCFFC96" box="[714,785,834,859]" metricMagnitude="2" metricUnit="m" metricValue="2.6" pageId="58" pageNumber="59" unit="m" value="260.0">260 m</quantity>
|
||
asl.), Along town, West Siang district,
|
||
<collectingRegion id="49BBF87EFFE40208FB2CFC90FA43FC97" box="[1224,1437,834,859]" country="India" name="Arunachal Pradesh" pageId="58" pageNumber="59">Arunachal Pradesh</collectingRegion>
|
||
state, Northeast
|
||
<collectingCountry id="F368760CFFE40208FEB8FCB4FE46FCB3" box="[348,408,870,895]" name="India" pageId="58" pageNumber="59">India</collectingCountry>
|
||
, collected by Systematics Lab members on
|
||
<date id="FFC1105CFFE40208FC5EFCB5FB83FCB3" box="[954,1117,870,895]" pageId="58" pageNumber="59" value="2009-07-30">30 July 2009</date>
|
||
;
|
||
<materialsCitation id="3B173CC1FFE40208FB95FCB5FE3EFC27" ID-GBIF-Occurrence="2028521216" collectingDate="2009-08-07" collectingDateMax="2009-08-09" collectingDateMin="2009-08-07" collectionCode="SDBDU" country="India" county="University" elevation="1110" latitude="27.101116" location="Sessa River" longLatPrecision="1" longitude="92.52737" municipality="Systematics Lab" pageId="58" pageNumber="59" specimenCode="SDBDU 2009.1267" specimenCount="1" stateProvince="Arunachal Pradesh">
|
||
one adult male (
|
||
<specimenCode id="DBD99EE7FFE40208FAD8FCB5FECDFC6F" collectionCode="SDBDU" pageId="58" pageNumber="59">SDBDU 2009.1267</specimenCode>
|
||
), from the
|
||
<location id="8EA06047FFE40208FE6BFC59FDCBFC6F" LSID="urn:lsid:plazi:treatment:03D6878AFFE4020DFF73FE9EFD51FF19:8EA06047FFE40208FE6BFC59FDCBFC6F" box="[399,533,906,931]" country="India" county="University" latitude="27.101116" longLatPrecision="1" longitude="92.52737" municipality="Systematics Lab" name="Sessa River" pageId="58" pageNumber="59" stateProvince="Arunachal Pradesh">Sessa River</location>
|
||
, nearby
|
||
<location id="8EA06047FFE40208FD91FC59FCD9FC6F" LSID="urn:lsid:plazi:treatment:03D6878AFFE4020DFF73FE9EFD51FF19:8EA06047FFE40208FD91FC59FCD9FC6F" box="[629,775,906,931]" country="India" county="University" latitude="27.101116" longLatPrecision="1" longitude="92.52737" municipality="Systematics Lab" name="Sessa village" pageId="58" pageNumber="59" stateProvince="Arunachal Pradesh">Sessa village</location>
|
||
(
|
||
<geoCoordinate id="EE4B505BFFE40208FCF3FC58FC7BFC6F" box="[791,933,906,931]" direction="north" orientation="latitude" pageId="58" pageNumber="59" precision="1" value="27.101116">27°6'4.02"N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE4B505BFFE40208FC56FC58FB87FC6F" box="[946,1113,906,931]" direction="east" orientation="longitude" pageId="58" pageNumber="59" precision="1" value="92.52737">92°31'38.52"E</geoCoordinate>
|
||
,
|
||
<elevation id="0052D1AFFFE40208FB81FC59FB3AFC6F" box="[1125,1252,906,931]" metricMagnitude="3" metricUnit="m" metricValue="1.1099999999999999" pageId="58" pageNumber="59" unit="m" value="1110.0">
|
||
<quantity id="4C879B79FFE40208FB81FC59FB66FC6E" box="[1125,1208,907,931]" metricMagnitude="3" metricUnit="m" metricValue="1.1099999999999999" pageId="58" pageNumber="59" unit="m" value="1110.0">1110 m</quantity>
|
||
asl
|
||
</elevation>
|
||
.),
|
||
<location id="8EA06047FFE40208FB1AFC59FF36FC0B" LSID="urn:lsid:plazi:treatment:03D6878AFFE4020DFF73FE9EFD51FF19:8EA06047FFE40208FB1AFC59FF36FC0B" country="India" county="University" latitude="27.101116" longLatPrecision="1" longitude="92.52737" municipality="Systematics Lab" name="West Kameng district" pageId="58" pageNumber="59" stateProvince="Arunachal Pradesh">West Kameng district</location>
|
||
,
|
||
<collectingRegion id="49BBF87EFFE40208FF12FC7CFE11FC0B" box="[246,463,942,967]" country="India" name="Arunachal Pradesh" pageId="58" pageNumber="59">Arunachal Pradesh</collectingRegion>
|
||
state,
|
||
<location id="8EA06047FFE40208FDFAFC7CFD55FC0B" LSID="urn:lsid:plazi:treatment:03D6878AFFE4020DFF73FE9EFD51FF19:8EA06047FFE40208FDFAFC7CFD55FC0B" box="[542,651,942,967]" country="India" county="University" latitude="27.101116" longLatPrecision="1" longitude="92.52737" municipality="Systematics Lab" name="Northeast" pageId="58" pageNumber="59" stateProvince="Arunachal Pradesh">Northeast</location>
|
||
<collectingCountry id="F368760CFFE40208FD71FC7CFD0FFC0B" box="[661,721,942,967]" name="India" pageId="58" pageNumber="59">India</collectingCountry>
|
||
, collected by members of the
|
||
<collectingMunicipality id="6BA4ACE6FFE40208FBD3FC7CFB2BFC0B" box="[1079,1269,942,967]" pageId="58" pageNumber="59">Systematics Lab</collectingMunicipality>
|
||
,
|
||
<collectingCounty id="62A14E10FFE40208FAE0FC7CFAA4FC0B" box="[1284,1402,942,967]" pageId="58" pageNumber="59">University</collectingCounty>
|
||
of
|
||
<collectingRegion id="49BBF87EFFE40208FF73FC00FF0BFC27" box="[151,213,978,1003]" country="India" name="Delhi" pageId="58" pageNumber="59">Delhi</collectingRegion>
|
||
on
|
||
<collectingDate id="EF85E9B4FFE40208FF1BFC01FE3EFC27" box="[255,480,979,1003]" pageId="58" pageNumber="59" value="2009-08-07" valueMax="2009-08-09" valueMin="2009-08-07">
|
||
<date id="FFC1105CFFE40208FF1BFC01FE3EFC27" box="[255,480,979,1003]" pageId="58" pageNumber="59" value="2009-08-07" valueMax="2009-08-09" valueMin="2009-08-07">07–09 August 2009</date>
|
||
</collectingDate>
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BC0369CFFE40208FF23FC24FE89FB2B" blockId="58.[151,1437,438,2011]" pageId="58" pageNumber="59">
|
||
<emphasis id="B90BEA8EFFE40208FF23FC24FCD5FBC3" bold="true" box="[199,779,1014,1039]" pageId="58" pageNumber="59">Provisionally referred specimens (see Remarks).</emphasis>
|
||
One juvenile male (SDBDU 2009.132), and two juvenile females (SDBDU 2009.133 & SDBDU 2009.134), from Pangkava ju (
|
||
<geoCoordinate id="EE4B505BFFE40208FC5DFBC8FB88FBFF" box="[953,1110,1050,1075]" direction="north" orientation="latitude" pageId="58" pageNumber="59" precision="1" value="24.672667">24°40'21.6"N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE4B505BFFE40208FB86FBC8FAD7FBFF" box="[1122,1289,1050,1075]" direction="east" orientation="longitude" pageId="58" pageNumber="59" precision="1" value="94.47218">94°28'19.86"E</geoCoordinate>
|
||
,
|
||
<quantity id="4C879B79FFE40208FAF1FBC9FA80FBFE" box="[1301,1374,1051,1075]" metricMagnitude="2" metricUnit="m" metricValue="8.2" pageId="58" pageNumber="59" unit="m" value="820.0">820 m</quantity>
|
||
asl.), Kangpat Khullen, Kamjong sub-division, Ukhrul district,
|
||
<collectingRegion id="49BBF87EFFE40208FCC9FBECFC53FB9B" box="[813,909,1086,1111]" country="India" name="Manipur" pageId="58" pageNumber="59">Manipur</collectingRegion>
|
||
state, Northeast
|
||
<collectingCountry id="F368760CFFE40208FBB5FBECFB53FB9B" box="[1105,1165,1086,1111]" name="India" pageId="58" pageNumber="59">India</collectingCountry>
|
||
, collected by RGK and SDB on
|
||
<date id="FFC1105CFFE40208FF1CFBB1FE4CFBB7" box="[248,402,1123,1147]" pageId="58" pageNumber="59" value="2009-05-21">21 May 2009</date>
|
||
;
|
||
<materialsCitation id="3B173CC1FFE40208FE7BFBB0FE8DFB2B" ID-GBIF-Occurrence="2028521233" collectingDate="2003-05-14" collectionCode="CAS" collectorName="G. O. U. Wogan & Wogan, J. A & Wilkinson, J. V & Vindum, H & Win, T & K. S. Lwin & A. K. Shein & H. Tun" country="Myanmar" county="Myitkyina district" latitude="25.090334" location="Hepu village" longLatPrecision="1" longitude="96.40367" municipality="Moe Nyin township" pageId="58" pageNumber="59" specimenCode="CAS 232938" specimenCount="1" stateProvince="Kachin">
|
||
adult female (
|
||
<specimenCode id="DBD99EE7FFE40208FDDEFBB1FD10FBB7" box="[570,718,1122,1147]" collectionCode="CAS" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="58" pageNumber="59" type="Museum">CAS 232938</specimenCode>
|
||
––tissue only), from Hepu Stream (
|
||
<geoCoordinate id="EE4B505BFFE40208FBB8FBB0FB34FBB7" box="[1116,1258,1122,1147]" direction="north" orientation="latitude" pageId="58" pageNumber="59" precision="1" value="25.090334">25°5'25.2"N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE4B505BFFE40208FB12FBB0FA4EFBB7" box="[1270,1424,1122,1147]" direction="east" orientation="longitude" pageId="58" pageNumber="59" precision="1" value="96.40367">96°24'13.2"E</geoCoordinate>
|
||
),
|
||
<location id="8EA06047FFE40208FF73FB55FEF1FB53" LSID="urn:lsid:plazi:treatment:03D6878AFFE4020DFF73FE9EFD51FF19:8EA06047FFE40208FF73FB55FEF1FB53" box="[151,303,1158,1183]" country="Myanmar" county="Myitkyina district" latitude="25.090334" longLatPrecision="1" longitude="96.40367" municipality="Moe Nyin township" name="Hepu village" pageId="58" pageNumber="59" stateProvince="Kachin">Hepu village</location>
|
||
,
|
||
<collectingMunicipality id="6BA4ACE6FFE40208FEA4FB55FDF1FB53" box="[320,559,1158,1183]" pageId="58" pageNumber="59">Moe Nyin township</collectingMunicipality>
|
||
,
|
||
<collectingCounty id="62A14E10FFE40208FDA4FB54FCCDFB53" box="[576,787,1158,1183]" pageId="58" pageNumber="59">Myitkyina district</collectingCounty>
|
||
,
|
||
<collectingRegion id="49BBF87EFFE40208FCC7FB54FCAAFB53" box="[803,884,1158,1183]" country="Myanmar" name="Kachin" pageId="58" pageNumber="59">Kachin</collectingRegion>
|
||
state, northern
|
||
<collectingCountry id="F368760CFFE40208FBD0FB55FB7DFB53" box="[1076,1187,1159,1183]" name="Myanmar" pageId="58" pageNumber="59">Myanmar</collectingCountry>
|
||
, collected by
|
||
<collectorName id="268A534AFFE40208FAAAFB55FF37FB0F" pageId="58" pageNumber="59">G.O.U. Wogan</collectorName>
|
||
, J.A.
|
||
<collectorName id="268A534AFFE40208FEC3FB78FE13FB0E" box="[295,461,1194,1219]" pageId="58" pageNumber="59">Wilkinson, J.V</collectorName>
|
||
.
|
||
<collectorName id="268A534AFFE40208FE3CFB78FD89FB0E" box="[472,599,1194,1219]" pageId="58" pageNumber="59">Vindum, H</collectorName>
|
||
.
|
||
<collectorName id="268A534AFFE40208FD85FB78FD6EFB0E" box="[609,688,1194,1219]" pageId="58" pageNumber="59">Win, T</collectorName>
|
||
. Thin [“Additional collectors:
|
||
<collectorName id="268A534AFFE40208FBE2FB79FBA2FB0F" box="[1030,1148,1194,1219]" pageId="58" pageNumber="59">K.S. Lwin</collectorName>
|
||
,
|
||
<collectorName id="268A534AFFE40208FB63FB79FADAFB0F" box="[1159,1284,1194,1219]" pageId="58" pageNumber="59">A.K. Shein</collectorName>
|
||
and
|
||
<collectorName id="268A534AFFE40208FADDFB79FA52FB0F" box="[1337,1420,1195,1219]" pageId="58" pageNumber="59">H. Tun</collectorName>
|
||
”] on
|
||
<collectingDate id="EF85E9B4FFE40208FF5EFB1DFE8DFB2B" box="[186,339,1231,1255]" pageId="58" pageNumber="59" value="2003-05-14">
|
||
<date id="FFC1105CFFE40208FF5EFB1DFE8DFB2B" box="[186,339,1231,1255]" pageId="58" pageNumber="59" value="2003-05-14">14 May 2003</date>
|
||
</collectingDate>
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BC0369CFFE40208FF23FB20FDE8F977" blockId="58.[151,1437,438,2011]" pageId="58" pageNumber="59">
|
||
<emphasis id="B90BEA8EFFE40208FF23FB20FD30FAC7" bold="true" box="[199,750,1266,1291]" pageId="58" pageNumber="59">
|
||
<typeStatus id="54C4883EFFE40208FF23FB20FEEAFAC7" box="[199,308,1266,1291]" pageId="58" pageNumber="59" type="holotype">Holotype</typeStatus>
|
||
description (measurements in mm).
|
||
</emphasis>
|
||
Mature male (SVL 93.8) (
|
||
<figureCitation id="13442A19FFE40208FBCEFB20FB77FAC7" box="[1066,1193,1266,1291]" captionStart="FIGURE 22" captionStartId="59.[151,250,1767,1789]" captionTargetBox="[151,1436,301,1746]" captionTargetId="figure@59.[151,1436,301,1746]" captionTargetPageId="59" captionText="FIGURE 22. Megophrys periosa sp. nov. holotype: adult male (BNHS 6055: SVL 93.8 mm) in preservation: A. dorsal view; B. ventral view; C. profile view of head; D. ventral view of hand; E. ventral view of foot." httpUri="https://zenodo.org/record/2610246/files/figure.png" pageId="58" pageNumber="59">Figures 22</figureCitation>
|
||
, & 23A & H). Head moderately large, wider than long (HW 35.0, HL 33.6, IFE 15.5, IBE 25.3); snout bluntly pointed in dorsal view, obtusely protruding beyond mandible in lateral view, without rostral appendage (
|
||
<figureCitation id="13442A19FFE40208FBCDFAE8FB71FA9F" box="[1065,1199,1338,1363]" captionStart="FIGURE 22" captionStartId="59.[151,250,1767,1789]" captionTargetBox="[151,1436,301,1746]" captionTargetId="figure@59.[151,1436,301,1746]" captionTargetPageId="59" captionText="FIGURE 22. Megophrys periosa sp. nov. holotype: adult male (BNHS 6055: SVL 93.8 mm) in preservation: A. dorsal view; B. ventral view; C. profile view of head; D. ventral view of hand; E. ventral view of foot." httpUri="https://zenodo.org/record/2610246/files/figure.png" pageId="58" pageNumber="59">Figure 22C</figureCitation>
|
||
); loreal region acute, concave; canthus rostralis angular; dorsal surface of snout very slightly concave; eye diameter twice maximum diameter of visible portion of tympanum, and shorter than snout length (EL 10.7, TYD 5.4, SL 11.7); eyetympanum distance (TYE 8.2) less than twice diameter of visible portion of tympanum; tympanum oval-shaped, oblique (
|
||
<figureCitation id="13442A19FFE40208FF1BFA18FE57FA2F" box="[255,393,1482,1507]" captionStart="FIGURE 22" captionStartId="59.[151,250,1767,1789]" captionTargetBox="[151,1436,301,1746]" captionTargetId="figure@59.[151,1436,301,1746]" captionTargetPageId="59" captionText="FIGURE 22. Megophrys periosa sp. nov. holotype: adult male (BNHS 6055: SVL 93.8 mm) in preservation: A. dorsal view; B. ventral view; C. profile view of head; D. ventral view of hand; E. ventral view of foot." httpUri="https://zenodo.org/record/2610246/files/figure.png" pageId="58" pageNumber="59">Figure 22C</figureCitation>
|
||
), with upper ~15% concealed by supratympanic ridge; pupil vertically elliptical; nostril positioned laterally, closer to eye than to snout (EN 5.2, NS 7.1); internarial distance greater than eyelid width, and equal to narrowest point between upper eyelids (IN 10.5, UEW 8.7, IUE 10.5); pineal ocellus not visible externally; vomerine ridge present, medium sized, ovoid, moderately raised, orientated acutely, positioned between to slightly posterior to choanae, equidistant from choanae and each other; vomerine teeth short; maxillary teeth present; tongue not observed due to fixation of jaw and in interest of preventing potential damage to jaws by forcing open mouth wide enough for examination.
|
||
</paragraph>
|
||
<paragraph id="8BC0369CFFE40209FF23F914FE9FFF3B" blockId="58.[151,1437,438,2011]" lastBlockId="59.[151,1437,151,248]" lastPageId="59" lastPageNumber="60" pageId="58" pageNumber="59">
|
||
Forelimbs long, thin (
|
||
<figureCitation id="13442A19FFE40208FE58F914FD5AF912" box="[444,644,1734,1759]" captionStart="FIGURE 22" captionStartId="59.[151,250,1767,1789]" captionTargetBox="[151,1436,301,1746]" captionTargetId="figure@59.[151,1436,301,1746]" captionTargetPageId="59" captionText="FIGURE 22. Megophrys periosa sp. nov. holotype: adult male (BNHS 6055: SVL 93.8 mm) in preservation: A. dorsal view; B. ventral view; C. profile view of head; D. ventral view of hand; E. ventral view of foot." httpUri="https://zenodo.org/record/2610246/files/figure.png" pageId="58" pageNumber="59">Figures 22A & B</figureCitation>
|
||
, &
|
||
<figureCitation id="13442A19FFE40208FD4AF915FD3FF913" box="[686,737,1735,1759]" captionStart="FIGURE 23" captionStartId="61.[151,250,1796,1818]" captionTargetBox="[240,1344,193,1781]" captionTargetId="figure@61.[240,1344,193,1782]" captionTargetPageId="61" captionText="FIGURE 23. Megophrys periosa sp. nov.: A. lateral view of adult male holotype (BNHS 6055: SVL 93.8 mm) in life. B. lateral view of adult male paratype (BNHS 6057: SVL 80.7 mm) in life. C–E. dorsolateral views of adult males in life showing typical variation: C. referred specimen (SDBDU 2009.1190: SVL 73.6 mm); D. referred specimen (SDBDU 2009.1189: SVL 71.3 mm); E. paratype (BNHS 6061: SVL 87.7 mm). F. dorsolateral view of adult female paratype (BNHS 6056: SVL 112.0 mm) in life. G. ventral view of two adult male paratypes immediately after euthanisation, showing variation of markings (left, BNHS 6064: SVL 88.8 mm; right BNHS 6063: SVL 81.3 mm). H. ventral view of three adult male specimens in preservation showing variation of makings (left, paratype BNHS 6059: SVL 81.9 mm; centre, paratype BNHS 6060: SVL 80.2 mm; right, holotype BNHS 6055: SVL 93.8 mm)." httpUri="https://zenodo.org/record/2610248/files/figure.png" pageId="58" pageNumber="59">23A</figureCitation>
|
||
), forearm moderately enlarged relative to upper forelimb, and shorter than hand length (FAL 20.6––left side; right side deformed; HAL 24.0); fingers long, narrow, without lateral fringes (
|
||
<figureCitation id="13442A19FFE40208FEADF8DCFE0AF8EB" box="[329,468,1806,1831]" captionStart="FIGURE 22" captionStartId="59.[151,250,1767,1789]" captionTargetBox="[151,1436,301,1746]" captionTargetId="figure@59.[151,1436,301,1746]" captionTargetPageId="59" captionText="FIGURE 22. Megophrys periosa sp. nov. holotype: adult male (BNHS 6055: SVL 93.8 mm) in preservation: A. dorsal view; B. ventral view; C. profile view of head; D. ventral view of hand; E. ventral view of foot." httpUri="https://zenodo.org/record/2610246/files/figure.png" pageId="58" pageNumber="59">Figure 22D</figureCitation>
|
||
), finger length formula IV<II<I<III (FIL 13.7, FIIL 10.9, FIIIL 15.5, FIVL 10.3); interdigital webbing, subarticular and supernumerary tubercles absent; thenar and outer metacarpal tubercles weakly developed; finger tips flattened, not expanded relative to digit width, and with subcircular pads, terminal grooves on pads absent. Hindlimbs long, thin (
|
||
<figureCitation id="13442A19FFE40208FD43F8A8FCB1F85E" box="[679,879,1914,1939]" captionStart="FIGURE 22" captionStartId="59.[151,250,1767,1789]" captionTargetBox="[151,1436,301,1746]" captionTargetId="figure@59.[151,1436,301,1746]" captionTargetPageId="59" captionText="FIGURE 22. Megophrys periosa sp. nov. holotype: adult male (BNHS 6055: SVL 93.8 mm) in preservation: A. dorsal view; B. ventral view; C. profile view of head; D. ventral view of hand; E. ventral view of foot." httpUri="https://zenodo.org/record/2610246/files/figure.png" pageId="58" pageNumber="59">Figures 22A & B</figureCitation>
|
||
, &
|
||
<figureCitation id="13442A19FFE40208FC7EF8A9FC10F85F" box="[922,974,1915,1939]" captionStart="FIGURE 23" captionStartId="61.[151,250,1796,1818]" captionTargetBox="[240,1344,193,1781]" captionTargetId="figure@61.[240,1344,193,1782]" captionTargetPageId="61" captionText="FIGURE 23. Megophrys periosa sp. nov.: A. lateral view of adult male holotype (BNHS 6055: SVL 93.8 mm) in life. B. lateral view of adult male paratype (BNHS 6057: SVL 80.7 mm) in life. C–E. dorsolateral views of adult males in life showing typical variation: C. referred specimen (SDBDU 2009.1190: SVL 73.6 mm); D. referred specimen (SDBDU 2009.1189: SVL 71.3 mm); E. paratype (BNHS 6061: SVL 87.7 mm). F. dorsolateral view of adult female paratype (BNHS 6056: SVL 112.0 mm) in life. G. ventral view of two adult male paratypes immediately after euthanisation, showing variation of markings (left, BNHS 6064: SVL 88.8 mm; right BNHS 6063: SVL 81.3 mm). H. ventral view of three adult male specimens in preservation showing variation of makings (left, paratype BNHS 6059: SVL 81.9 mm; centre, paratype BNHS 6060: SVL 80.2 mm; right, holotype BNHS 6055: SVL 93.8 mm)." httpUri="https://zenodo.org/record/2610248/files/figure.png" pageId="58" pageNumber="59">23A</figureCitation>
|
||
); thighs equal to shanks, and longer than feet (TL 47.4, SHL 50.2, FOL 45.8); toes long, dorsoventrally flattened, without lateral fringes (
|
||
<figureCitation id="13442A19FFE40208FAECF84CFA4EF87B" box="[1288,1424,1950,1975]" captionStart="FIGURE 22" captionStartId="59.[151,250,1767,1789]" captionTargetBox="[151,1436,301,1746]" captionTargetId="figure@59.[151,1436,301,1746]" captionTargetPageId="59" captionText="FIGURE 22. Megophrys periosa sp. nov. holotype: adult male (BNHS 6055: SVL 93.8 mm) in preservation: A. dorsal view; B. ventral view; C. profile view of head; D. ventral view of hand; E. ventral view of foot." httpUri="https://zenodo.org/record/2610246/files/figure.png" pageId="58" pageNumber="59">Figure 22E</figureCitation>
|
||
), relative toe lengths I<II<V<III<IV; toe tips flattened, not dilated, but with distinct oval-shaped pads, terminal grooves on pads absent; webbing rudimentary; inner metatarsal tubercles weakly defined, longitudinally ovalshaped; subarticular, supernumerary and outer metatarsal tubercles absent; ridge of callous tissue absent on ventral surface of toes.
|
||
</paragraph>
|
||
<caption id="DF006614FFE50209FF73F935FBFCF8D0" httpUri="https://zenodo.org/record/2610246/files/figure.png" pageId="59" pageNumber="60" startId="59.[151,250,1767,1789]" targetBox="[151,1436,301,1746]" targetPageId="59">
|
||
<paragraph id="8BC0369CFFE50209FF73F935FBFCF8D0" blockId="59.[151,1436,1767,1820]" pageId="59" pageNumber="60">
|
||
<emphasis id="B90BEA8EFFE50209FF73F935FEFEF931" bold="true" box="[151,288,1767,1789]" pageId="59" pageNumber="60">FIGURE 22.</emphasis>
|
||
<taxonomicName id="4C7F4D1FFFE50209FEC3F935FE32F932" authority="Mahony & Kamei & Teeling, 2018" authorityName="Mahony & Kamei & Teeling" authorityYear="2018" box="[295,492,1767,1790]" class="Amphibia" family="Megophryidae" genus="Megophrys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="59" pageNumber="60" phylum="Chordata" rank="genus" status="sp. nov.">
|
||
<emphasis id="B90BEA8EFFE50209FEC3F935FE32F932" box="[295,492,1767,1790]" italics="true" pageId="59" pageNumber="60">Megophrys periosa</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A23857F5FFE50209FE17F93AFD9AF931" box="[499,580,1768,1789]" pageId="59" pageNumber="60" rank="species">
|
||
<emphasis id="B90BEA8EFFE50209FE17F93AFD9AF931" bold="true" box="[499,580,1768,1789]" pageId="59" pageNumber="60">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
holotype: adult male (BNHS 6055: SVL 93.8 mm) in preservation:
|
||
<emphasis id="B90BEA8EFFE50209FB1BF935FACFF930" bold="true" box="[1279,1297,1767,1788]" pageId="59" pageNumber="60">A</emphasis>
|
||
. dorsal view;
|
||
<emphasis id="B90BEA8EFFE50209FF73F8D5FF76F8D0" bold="true" box="[151,168,1799,1820]" pageId="59" pageNumber="60">B</emphasis>
|
||
. ventral view;
|
||
<emphasis id="B90BEA8EFFE50209FEDBF8D4FE8FF8D0" bold="true" box="[319,337,1798,1820]" pageId="59" pageNumber="60">C</emphasis>
|
||
. profile view of head;
|
||
<emphasis id="B90BEA8EFFE50209FDDCF8D5FD94F8D0" bold="true" box="[568,586,1799,1820]" pageId="59" pageNumber="60">D</emphasis>
|
||
. ventral view of hand;
|
||
<emphasis id="B90BEA8EFFE50209FCD1F8D5FC98F8D0" bold="true" box="[821,838,1799,1820]" pageId="59" pageNumber="60">E</emphasis>
|
||
. ventral view of foot.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BC0369CFFE5020EFF23F89BFBDBFE1C" blockId="59.[151,1437,1865,2034]" lastBlockId="60.[151,1437,151,2000]" lastPageId="60" lastPageNumber="61" pageId="59" pageNumber="60">Skin of dorsal and lateral surfaces of head, body and limbs primarily smooth, sparsely covered with small, weak granules; tympanum with borders raised relative to surrounding region; outer edge of upper eyelids with a broad pointed bump; supratympanic ridges narrow anteriorly, gradually expanding posterior to tympanum to become moderately enlarged, glandular, extending from orbit, curving downward abruptly at posterior upper border of tympanum, terminating above forelimb insertions; flanks sparsely covered with small to medium sized unevenly scattered pustular tubercles; dorsolateral ridges thin, weakly developed, extending posteriorly from behind supratympanic ridges to ~90% distance to vent; parietoscapular-sacral ridges weakly developed, “>–<” configuration, composed primarily of single row of closely spaced asperities; dorsal surfaces of forearms, thighs and shanks with short transverse ridges; posterior thighs with small scattered pustular tubercles; gular region, chest, abdomen and ventral surfaces of limbs smooth; pectoral glands small, flat, level with axilla on chest; femoral glands medium sized, flat, on posterior surface of thighs, slightly closer to knees than to cloaca; small white dermal asperities form broad dense band circummarginally on gular region, sparse on tympanic regions between tympanum and eyes, on dorsal surfaces of body and upper posterior surface of flanks, increasing in density posteriorly, present along all dorsal ridges, absent from all remaining surfaces.</paragraph>
|
||
<paragraph id="8BC0369CFFE2020EFF23FE0EFAA4FBF9" blockId="60.[151,1437,151,2000]" pageId="60" pageNumber="61">
|
||
<emphasis id="B90BEA8EFFE2020EFF23FE0EFE8EFE39" box="[199,336,476,501]" italics="true" pageId="60" pageNumber="61">Colouration</emphasis>
|
||
:
|
||
<emphasis id="B90BEA8EFFE2020EFE85FE0FFDD3FE38" box="[353,525,477,500]" italics="true" pageId="60" pageNumber="61">In preservative</emphasis>
|
||
(
|
||
<figureCitation id="13442A19FFE2020EFDFAFE0EFD4CFE38" box="[542,658,476,501]" captionStart="FIGURE 22" captionStartId="59.[151,250,1767,1789]" captionTargetBox="[151,1436,301,1746]" captionTargetId="figure@59.[151,1436,301,1746]" captionTargetPageId="59" captionText="FIGURE 22. Megophrys periosa sp. nov. holotype: adult male (BNHS 6055: SVL 93.8 mm) in preservation: A. dorsal view; B. ventral view; C. profile view of head; D. ventral view of hand; E. ventral view of foot." httpUri="https://zenodo.org/record/2610246/files/figure.png" pageId="60" pageNumber="61">Figure 22</figureCitation>
|
||
): Dorsal surfaces of body and head primarily greyish-brown; lightedged, darker brown triangular marking between eyes; parietoscapular-sacral and dorsolateral ridges narrowly bordered by dark brown; flanks grey, darker dorsally, tubercles on flanks primarily dark brown or black, many with small lighter tips; large dark brown blotch covers tympanic region, another extends from lower posterior border of orbits, additional dark brown bar extends vertically from lower orbital border to edge of upper lips; lateral surfaces of canthus rostralis black, front of snout dark brown; remaining surfaces of upper lips and area between dark blotches on lateral surfaces of head greyish-white; dorsal and lateral surfaces of forelimbs primarily light grey with large dark brown transverse blotches on forearms; dorsal surface of fingers with dark brown blotches; dorsal and lateral surfaces of hindlimbs brown with distinct dark brown transverse crossbars; throat, chest and anterior abdomen primarily brown with white-edged darker brown blotches and mottling; posterior half of abdomen and ventral surfaces of forelimbs and hindlimbs primarily creamish-white with dark brown blotches; ventral surfaces of tarsi dark brown; area surrounding vent and posterior surfaces of thighs dark brown; ventral surfaces of feet and hands greyish-brown; pectoral and femoral glands creamish-white.
|
||
<emphasis id="B90BEA8EFFE2020EFC72FC5FFC04FC69" box="[918,986,908,933]" italics="true" pageId="60" pageNumber="61">In life</emphasis>
|
||
(
|
||
<figureCitation id="13442A19FFE2020EFC0FFC5EFBADFC68" box="[1003,1139,908,933]" captionStart="FIGURE 23" captionStartId="61.[151,250,1796,1818]" captionTargetBox="[240,1344,193,1781]" captionTargetId="figure@61.[240,1344,193,1782]" captionTargetPageId="61" captionText="FIGURE 23. Megophrys periosa sp. nov.: A. lateral view of adult male holotype (BNHS 6055: SVL 93.8 mm) in life. B. lateral view of adult male paratype (BNHS 6057: SVL 80.7 mm) in life. C–E. dorsolateral views of adult males in life showing typical variation: C. referred specimen (SDBDU 2009.1190: SVL 73.6 mm); D. referred specimen (SDBDU 2009.1189: SVL 71.3 mm); E. paratype (BNHS 6061: SVL 87.7 mm). F. dorsolateral view of adult female paratype (BNHS 6056: SVL 112.0 mm) in life. G. ventral view of two adult male paratypes immediately after euthanisation, showing variation of markings (left, BNHS 6064: SVL 88.8 mm; right BNHS 6063: SVL 81.3 mm). H. ventral view of three adult male specimens in preservation showing variation of makings (left, paratype BNHS 6059: SVL 81.9 mm; centre, paratype BNHS 6060: SVL 80.2 mm; right, holotype BNHS 6055: SVL 93.8 mm)." httpUri="https://zenodo.org/record/2610248/files/figure.png" pageId="60" pageNumber="61">Figure 23A</figureCitation>
|
||
): Dorsal surfaces of head, body and hindlimbs more olive-brown, distinctly paler on flanks and forelimbs; ventral surfaces generally lighter, with darker brown blotches distinct and revealing distinct wide brown longitudinal stripe on gular region; dark blotches on chest; ventral surfaces of limbs dark to light purple/mauve; distinct orange speckles present on dorsal and ventral surfaces, most noticeable on dorsal surfaces of toes and fingers and on flanks; iris very dark orange.
|
||
</paragraph>
|
||
<paragraph id="8BC0369CFFE2020EFF23FBEDFEECFBB1" blockId="60.[151,1437,151,2000]" pageId="60" pageNumber="61">
|
||
Refer to
|
||
<figureCitation id="13442A19FFE2020EFED6FBEDFE73FB94" box="[306,429,1087,1112]" captionStart="FIGURE 22" captionStartId="59.[151,250,1767,1789]" captionTargetBox="[151,1436,301,1746]" captionTargetId="figure@59.[151,1436,301,1746]" captionTargetPageId="59" captionText="FIGURE 22. Megophrys periosa sp. nov. holotype: adult male (BNHS 6055: SVL 93.8 mm) in preservation: A. dorsal view; B. ventral view; C. profile view of head; D. ventral view of hand; E. ventral view of foot." httpUri="https://zenodo.org/record/2610246/files/figure.png" pageId="60" pageNumber="61">Figures 22</figureCitation>
|
||
and
|
||
<figureCitation id="13442A19FFE2020EFE09FB92FDC3FB94" box="[493,541,1088,1112]" captionStart="FIGURE 23" captionStartId="61.[151,250,1796,1818]" captionTargetBox="[240,1344,193,1781]" captionTargetId="figure@61.[240,1344,193,1782]" captionTargetPageId="61" captionText="FIGURE 23. Megophrys periosa sp. nov.: A. lateral view of adult male holotype (BNHS 6055: SVL 93.8 mm) in life. B. lateral view of adult male paratype (BNHS 6057: SVL 80.7 mm) in life. C–E. dorsolateral views of adult males in life showing typical variation: C. referred specimen (SDBDU 2009.1190: SVL 73.6 mm); D. referred specimen (SDBDU 2009.1189: SVL 71.3 mm); E. paratype (BNHS 6061: SVL 87.7 mm). F. dorsolateral view of adult female paratype (BNHS 6056: SVL 112.0 mm) in life. G. ventral view of two adult male paratypes immediately after euthanisation, showing variation of markings (left, BNHS 6064: SVL 88.8 mm; right BNHS 6063: SVL 81.3 mm). H. ventral view of three adult male specimens in preservation showing variation of makings (left, paratype BNHS 6059: SVL 81.9 mm; centre, paratype BNHS 6060: SVL 80.2 mm; right, holotype BNHS 6055: SVL 93.8 mm)." httpUri="https://zenodo.org/record/2610248/files/figure.png" pageId="60" pageNumber="61">23A</figureCitation>
|
||
for further details of colouration and markings for
|
||
<typeStatus id="54C4883EFFE2020EFB66FBEDFB3AFB94" box="[1154,1252,1087,1112]" pageId="60" pageNumber="61" type="holotype">holotype</typeStatus>
|
||
(in life, and in preservation).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3656517FFE2020CFF23FB55FB07FBF9" lastPageId="62" lastPageNumber="63" pageId="60" pageNumber="61" type="description">
|
||
<paragraph id="8BC0369CFFE2020EFF23FB55FAC5F930" blockId="60.[151,1437,151,2000]" pageId="60" pageNumber="61">
|
||
<emphasis id="B90BEA8EFFE2020EFF23FB55FEE1FB6C" bold="true" box="[199,319,1159,1184]" pageId="60" pageNumber="61">Variation.</emphasis>
|
||
Refer to
|
||
<tableCitation id="C6FD0327FFE2020EFE58FB55FDC8FB6C" box="[444,534,1159,1184]" captionStart="TABLE 1" captionStartId="22.[147,225,525,545]" captionText="TABLE 1. Morphometric details for specimens new and known included in the Taxonomic review section. Refer to Methods and materials section for explanation of measurement abbreviations and museum acronyms. HT = holotype; PT = paratype; LT = lectotype; PLT = paralectotype; NT = neotype; RS = referred specimen; M = adult male; F = adult female; SA = subadult; J = juvenile [post metamorphic]; † = BMNH voucher numbers; # = SDBDU voucher numbers; ind = character limits insufficiently distinct to reliably measure." pageId="60" pageNumber="61">Table 1</tableCitation>
|
||
for morphometric variation within the type series and referred specimens, consisting of
|
||
<specimenCount id="9D79FD15FFE2020EFED6FB7EFE50FB08" box="[306,398,1195,1220]" pageId="60" pageNumber="61" type="adult">12 adult</specimenCount>
|
||
males and one adult female. The
|
||
<typeStatus id="54C4883EFFE2020EFCF2FB7EFC5CFB08" box="[790,898,1196,1220]" pageId="60" pageNumber="61" type="paratype">paratypes</typeStatus>
|
||
and referred specimens generally resemble the
|
||
<typeStatus id="54C4883EFFE2020EFF73FB1CFF27FB2B" box="[151,249,1230,1255]" pageId="60" pageNumber="61" type="holotype">holotype</typeStatus>
|
||
for most morphological characters with the following exceptions: Lateral fringes on toes typically absent, but some specimens have very narrow fringes on all toes; relative finger lengths vary between individuals, including IV<II<I<III for most specimens, but IV=II<I<III observed on three (SDBDU 2009.1189, BNHS 6061, BNHS 6064); posterior edge of tongue varies from weakly, to moderately bifurcated, likely due to varying fixation conditions; dorsolateral ridges vary from weak to moderately well developed, always extending>75% of the trunk length (
|
||
<figureCitation id="13442A19FFE2020EFF08FA52FE85FA54" box="[236,347,1408,1433]" captionStart="FIGURE 23" captionStartId="61.[151,250,1796,1818]" captionTargetBox="[240,1344,193,1781]" captionTargetId="figure@61.[240,1344,193,1782]" captionTargetPageId="61" captionText="FIGURE 23. Megophrys periosa sp. nov.: A. lateral view of adult male holotype (BNHS 6055: SVL 93.8 mm) in life. B. lateral view of adult male paratype (BNHS 6057: SVL 80.7 mm) in life. C–E. dorsolateral views of adult males in life showing typical variation: C. referred specimen (SDBDU 2009.1190: SVL 73.6 mm); D. referred specimen (SDBDU 2009.1189: SVL 71.3 mm); E. paratype (BNHS 6061: SVL 87.7 mm). F. dorsolateral view of adult female paratype (BNHS 6056: SVL 112.0 mm) in life. G. ventral view of two adult male paratypes immediately after euthanisation, showing variation of markings (left, BNHS 6064: SVL 88.8 mm; right BNHS 6063: SVL 81.3 mm). H. ventral view of three adult male specimens in preservation showing variation of makings (left, paratype BNHS 6059: SVL 81.9 mm; centre, paratype BNHS 6060: SVL 80.2 mm; right, holotype BNHS 6055: SVL 93.8 mm)." httpUri="https://zenodo.org/record/2610248/files/figure.png" pageId="60" pageNumber="61">Figure 23</figureCitation>
|
||
A–F); typically only upper border of tympanum is concealed by supratympanic ridge, though up to ~25% of tympanum appears to be concealed on some specimens; parietoscapular-sacral ridge configuration varies considerably amongst individuals, i.e., “>- (”, “>––<”, “> <”, “> |”, “>- <”, or only V-shaped parietoscapular ridge present; coverage of dermal asperities varies mostly in density between individual males in comparison with
|
||
<typeStatus id="54C4883EFFE2020EFEBCF9DCFE63F9EB" box="[344,445,1550,1575]" pageId="60" pageNumber="61" type="holotype">holotype</typeStatus>
|
||
, however, on some specimens both black (or brown) and white asperities were observed, may sparsely cover upper lips, loreal region, front of snout, posterior dorsal surface of upper eyelids, head, and dorsal surface of tibia-tarsus joints; asperities white on female, restricted to dorsal surface of body, sparse on middorsum, increasing in density posteriorly to above cloaca; outer metacarpal tubercles not visible on most specimens; tubercle cover on flanks varies considerably, some with only sparse scattering of small tubercles, others with moderately dense cover of heterogeneous (large to small) sized tubercles (
|
||
<figureCitation id="13442A19FFE2020EFBF0F96DFB5CF914" box="[1044,1154,1727,1752]" captionStart="FIGURE 23" captionStartId="61.[151,250,1796,1818]" captionTargetBox="[240,1344,193,1781]" captionTargetId="figure@61.[240,1344,193,1782]" captionTargetPageId="61" captionText="FIGURE 23. Megophrys periosa sp. nov.: A. lateral view of adult male holotype (BNHS 6055: SVL 93.8 mm) in life. B. lateral view of adult male paratype (BNHS 6057: SVL 80.7 mm) in life. C–E. dorsolateral views of adult males in life showing typical variation: C. referred specimen (SDBDU 2009.1190: SVL 73.6 mm); D. referred specimen (SDBDU 2009.1189: SVL 71.3 mm); E. paratype (BNHS 6061: SVL 87.7 mm). F. dorsolateral view of adult female paratype (BNHS 6056: SVL 112.0 mm) in life. G. ventral view of two adult male paratypes immediately after euthanisation, showing variation of markings (left, BNHS 6064: SVL 88.8 mm; right BNHS 6063: SVL 81.3 mm). H. ventral view of three adult male specimens in preservation showing variation of makings (left, paratype BNHS 6059: SVL 81.9 mm; centre, paratype BNHS 6060: SVL 80.2 mm; right, holotype BNHS 6055: SVL 93.8 mm)." httpUri="https://zenodo.org/record/2610248/files/figure.png" pageId="60" pageNumber="61">Figure 23</figureCitation>
|
||
A–F). Dorsal and ventral markings vary considerably between individuals (see
|
||
<figureCitation id="13442A19FFE2020EFD15F931FC83F937" box="[753,861,1763,1788]" captionStart="FIGURE 23" captionStartId="61.[151,250,1796,1818]" captionTargetBox="[240,1344,193,1781]" captionTargetId="figure@61.[240,1344,193,1782]" captionTargetPageId="61" captionText="FIGURE 23. Megophrys periosa sp. nov.: A. lateral view of adult male holotype (BNHS 6055: SVL 93.8 mm) in life. B. lateral view of adult male paratype (BNHS 6057: SVL 80.7 mm) in life. C–E. dorsolateral views of adult males in life showing typical variation: C. referred specimen (SDBDU 2009.1190: SVL 73.6 mm); D. referred specimen (SDBDU 2009.1189: SVL 71.3 mm); E. paratype (BNHS 6061: SVL 87.7 mm). F. dorsolateral view of adult female paratype (BNHS 6056: SVL 112.0 mm) in life. G. ventral view of two adult male paratypes immediately after euthanisation, showing variation of markings (left, BNHS 6064: SVL 88.8 mm; right BNHS 6063: SVL 81.3 mm). H. ventral view of three adult male specimens in preservation showing variation of makings (left, paratype BNHS 6059: SVL 81.9 mm; centre, paratype BNHS 6060: SVL 80.2 mm; right, holotype BNHS 6055: SVL 93.8 mm)." httpUri="https://zenodo.org/record/2610248/files/figure.png" pageId="60" pageNumber="61">Figure 23</figureCitation>
|
||
which represent extremes in variation).
|
||
</paragraph>
|
||
<paragraph id="8BC0369CFFE2020EFF23F8D4FD4CF81C" blockId="60.[151,1437,151,2000]" pageId="60" pageNumber="61">
|
||
<emphasis id="B90BEA8EFFE2020EFF23F8D4FDF8F8D3" bold="true" box="[199,550,1798,1823]" pageId="60" pageNumber="61">Secondary sexual characters.</emphasis>
|
||
<emphasis id="B90BEA8EFFE2020EFDCAF8D4FDACF8D3" box="[558,626,1798,1823]" italics="true" pageId="60" pageNumber="61">Males</emphasis>
|
||
: nuptial pads present, weakly raised, covered with brown/black microasperities on freshly collected specimens, covering most of dorsal surface of Finger I; nuptial pad on Finger II medium sized, oval, positioned on base of digit on inner dorsal side, extending almost to base of distal phalange; external vocal sac indistinct; large internal vocal slits present on floor of mouth near rear of mandible, on each side; forearms enlarged relative to upper forelimbs.
|
||
<emphasis id="B90BEA8EFFE2020EFD49F846FCDFF861" box="[685,769,1940,1965]" italics="true" pageId="60" pageNumber="61">Female</emphasis>
|
||
: mature ova without pigmented poles; nuptial pads, vocal sac, vocal slits, enlarged forearms, all absent.
|
||
</paragraph>
|
||
<caption id="DF006614FFE3020FFF73F8D6FDC1F83D" httpUri="https://zenodo.org/record/2610248/files/figure.png" pageId="61" pageNumber="62" startId="61.[151,250,1796,1818]" targetBox="[240,1344,193,1781]" targetPageId="61">
|
||
<paragraph id="8BC0369CFFE3020FFF73F8D6FDC1F83D" blockId="61.[151,1436,1796,2033]" pageId="61" pageNumber="62">
|
||
<emphasis id="B90BEA8EFFE3020FFF73F8D6FEFCF8D5" bold="true" box="[151,290,1796,1818]" pageId="61" pageNumber="62">FIGURE 23.</emphasis>
|
||
<taxonomicName id="4C7F4D1FFFE3020FFECFF8D6FE2CF8D6" authority="Mahony & Kamei & Teeling, 2018" authorityName="Mahony & Kamei & Teeling" authorityYear="2018" box="[299,498,1796,1818]" class="Amphibia" family="Megophryidae" genus="Megophrys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="61" pageNumber="62" phylum="Chordata" rank="genus" status="sp. nov.">
|
||
<emphasis id="B90BEA8EFFE3020FFECFF8D6FE2CF8D6" box="[299,498,1796,1818]" italics="true" pageId="61" pageNumber="62">Megophrys periosa</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A23857F5FFE3020FFE1FF8D7FD90F8D6" box="[507,590,1797,1818]" pageId="61" pageNumber="62" rank="species">
|
||
<emphasis id="B90BEA8EFFE3020FFE1FF8D7FD90F8D6" bold="true" box="[507,590,1797,1818]" pageId="61" pageNumber="62">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
:
|
||
<emphasis id="B90BEA8EFFE3020FFDBAF8D6FDAEF8D5" bold="true" box="[606,624,1796,1817]" pageId="61" pageNumber="62">A</emphasis>
|
||
. lateral view of adult male holotype (BNHS 6055: SVL 93.8 mm) in life.
|
||
<emphasis id="B90BEA8EFFE3020FFA60F8D7FA4BF8D6" bold="true" box="[1412,1429,1797,1818]" pageId="61" pageNumber="62">B</emphasis>
|
||
. lateral view of adult male paratype (BNHS 6057: SVL 80.7 mm) in life.
|
||
<emphasis id="B90BEA8EFFE3020FFC90F8F1FC7AF8F4" bold="true" box="[884,932,1827,1849]" pageId="61" pageNumber="62">C–E</emphasis>
|
||
. dorsolateral views of adult males in life showing typical variation:
|
||
<emphasis id="B90BEA8EFFE3020FFEAFF893FE83F89B" bold="true" box="[331,349,1857,1879]" pageId="61" pageNumber="62">C</emphasis>
|
||
. referred specimen (SDBDU 2009.1190: SVL 73.6 mm);
|
||
<emphasis id="B90BEA8EFFE3020FFC54F890FC1CF89B" bold="true" box="[944,962,1858,1879]" pageId="61" pageNumber="62">D</emphasis>
|
||
. referred specimen (SDBDU 2009.1189: SVL 71.3 mm);
|
||
<emphasis id="B90BEA8EFFE3020FFEEDF8B2FEC4F8B9" bold="true" box="[265,282,1888,1909]" pageId="61" pageNumber="62">E</emphasis>
|
||
. paratype (BNHS 6061: SVL 87.7 mm).
|
||
<emphasis id="B90BEA8EFFE3020FFD20F8B2FD0FF8B9" bold="true" box="[708,721,1888,1909]" pageId="61" pageNumber="62">F</emphasis>
|
||
. dorsolateral view of adult female paratype (BNHS 6056: SVL 112.0 mm) in life.
|
||
<emphasis id="B90BEA8EFFE3020FFEF3F8ADFEF6F859" bold="true" box="[279,296,1919,1941]" pageId="61" pageNumber="62">G</emphasis>
|
||
. ventral view of two adult male paratypes immediately after euthanisation, showing variation of markings (left, BNHS 6064: SVL 88.8 mm; right BNHS 6063: SVL 81.3 mm).
|
||
<emphasis id="B90BEA8EFFE3020FFCCCF84CFCE2F87F" bold="true" box="[808,828,1950,1971]" pageId="61" pageNumber="62">H</emphasis>
|
||
. ventral view of three adult male specimens in preservation showing variation of makings (left, paratype BNHS 6059: SVL 81.9 mm; centre, paratype BNHS 6060: SVL 80.2 mm; right, holotype BNHS 6055: SVL 93.8 mm).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BC0369CFFE0020CFF23FF45FC1BFF19" blockId="62.[151,1437,151,2013]" pageId="62" pageNumber="63">
|
||
<emphasis id="B90BEA8EFFE0020CFF23FF45FDCDFF7C" bold="true" box="[199,531,151,176]" pageId="62" pageNumber="63">Morphological comparison.</emphasis>
|
||
Characters used for comparing
|
||
<taxonomicName id="4C7F4D1FFFE0020CFC67FF45FB80FF7C" authority="Mahony & Kamei & Teeling, 2018" authorityName="Mahony & Kamei & Teeling" authorityYear="2018" box="[899,1118,151,176]" class="Amphibia" family="Megophryidae" genus="Megophrys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="62" pageNumber="63" phylum="Chordata" rank="genus" status="sp. nov.">
|
||
<emphasis id="B90BEA8EFFE0020CFC67FF45FB80FF7C" box="[899,1118,151,176]" italics="true" pageId="62" pageNumber="63">Megophrys periosa</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A23857F5FFE0020CFB83FF45FB1DFF7C" box="[1127,1219,151,176]" pageId="62" pageNumber="63" rank="species">
|
||
<emphasis id="B90BEA8EFFE0020CFB83FF45FB1DFF7C" bold="true" box="[1127,1219,151,176]" pageId="62" pageNumber="63">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
with its congeners do not include those from the provisionally assigned referred specimens.
|
||
</paragraph>
|
||
<paragraph id="8BC0369CFFE0020CFF23FF0DFB46FD00" blockId="62.[151,1437,151,2013]" pageId="62" pageNumber="63">
|
||
<taxonomicName id="4C7F4D1FFFE0020CFF23FF0DFE7CFF34" authority="Mahony & Kamei & Teeling, 2018" authorityName="Mahony & Kamei & Teeling" authorityYear="2018" box="[199,418,223,248]" class="Amphibia" family="Megophryidae" genus="Megophrys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="62" pageNumber="63" phylum="Chordata" rank="genus" status="sp. nov.">
|
||
<emphasis id="B90BEA8EFFE0020CFF23FF0DFE7CFF34" box="[199,418,223,248]" italics="true" pageId="62" pageNumber="63">Megophrys periosa</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A23857F5FFE0020CFE48FF0DFDD9FF34" box="[428,519,223,248]" pageId="62" pageNumber="63" rank="species">
|
||
<emphasis id="B90BEA8EFFE0020CFE48FF0DFDD9FF34" bold="true" box="[428,519,223,248]" pageId="62" pageNumber="63">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
(adult males,
|
||
<emphasis id="B90BEA8EFFE0020CFD49FF33FD1EFF34" box="[685,704,225,248]" italics="true" pageId="62" pageNumber="63">N</emphasis>
|
||
=12, adult female,
|
||
<emphasis id="B90BEA8EFFE0020CFC7EFF33FC73FF34" box="[922,941,225,248]" italics="true" pageId="62" pageNumber="63">N</emphasis>
|
||
=1) differs from
|
||
<taxonomicName id="4C7F4D1FFFE0020CFB8BFF33FAC7FF34" authorityName="Fei, Ye and Huang" authorityYear="1983" box="[1135,1305,223,248]" class="Amphibia" family="Megophryidae" genus="Megophrys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="62" pageNumber="63" phylum="Chordata" rank="species" species="medogensis">
|
||
<emphasis id="B90BEA8EFFE0020CFB8BFF33FAC7FF34" box="[1135,1305,223,248]" italics="true" pageId="62" pageNumber="63">M. medogensis</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C7F4D1FFFE0020CFACEFF33FA42FF34" authorityName="Ye and Fei" authorityYear="1992" box="[1322,1436,223,248]" class="Amphibia" family="Megophryidae" genus="Megophrys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="62" pageNumber="63" phylum="Chordata" rank="species" species="zhangi">
|
||
<emphasis id="B90BEA8EFFE0020CFACEFF33FA42FF34" box="[1322,1436,223,248]" italics="true" pageId="62" pageNumber="63">M. zhangi</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C7F4D1FFFE0020CFF2CFED7FEBEFED1" authorityName="" sensu Anderson" authorityYear="1871" box="[200,352,260,285]" class="Amphibia" family="Megophryidae" genus="Xenophrys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="62" pageNumber="63" phylum="Chordata" rank="species" species="monticola">
|
||
<emphasis id="B90BEA8EFFE0020CFF2CFED7FEBEFED1" box="[200,352,260,285]" italics="true" pageId="62" pageNumber="63">M. monticola</emphasis>
|
||
</taxonomicName>
|
||
by its larger adult body size, male SVL
|
||
<quantity id="4C879B79FFE0020CFCD0FED6FC05FED0" box="[820,987,260,284]" metricMagnitude="-2" metricUnit="m" metricValue="8.254999999999999" metricValueMax="9.379999999999999" metricValueMin="7.13" pageId="62" pageNumber="63" unit="mm" value="82.55" valueMax="93.8" valueMin="71.3">71.3–93.8 mm</quantity>
|
||
, female SVL 112.0 mm (vs. male SVL 57.2–68.0 mm,
|
||
<emphasis id="B90BEA8EFFE0020CFEA9FEFBFEBEFE8C" box="[333,352,297,320]" italics="true" pageId="62" pageNumber="63">N</emphasis>
|
||
=17; male SVL
|
||
<quantity id="4C879B79FFE0020CFDF8FEF5FD1DFEF3" box="[540,707,295,320]" metricMagnitude="-2" metricUnit="m" metricValue="3.485" metricValueMax="3.72" metricValueMin="3.25" pageId="62" pageNumber="63" unit="mm" value="34.85" valueMax="37.2" valueMin="32.5">32.5–37.2 mm</quantity>
|
||
,
|
||
<emphasis id="B90BEA8EFFE0020CFD35FEFBFD3AFE8C" box="[721,740,297,320]" italics="true" pageId="62" pageNumber="63">N</emphasis>
|
||
=3; male SVL
|
||
<quantity id="4C879B79FFE0020CFC76FEF5FBE4FEF3" box="[914,1082,295,320]" metricMagnitude="-2" metricUnit="m" metricValue="4.385" metricValueMax="4.95" metricValueMin="3.82" pageId="62" pageNumber="63" unit="mm" value="43.85" valueMax="49.5" valueMin="38.2">38.2–49.5 mm</quantity>
|
||
,
|
||
<emphasis id="B90BEA8EFFE0020CFBACFEFBFB85FE8C" box="[1096,1115,297,320]" italics="true" pageId="62" pageNumber="63">N</emphasis>
|
||
=17, female SVL
|
||
<quantity id="4C879B79FFE0020CFAC8FEF5FF1BFEA8" metricMagnitude="-2" metricUnit="m" metricValue="4.83" metricValueMax="5.61" metricValueMin="4.05" pageId="62" pageNumber="63" unit="mm" value="48.3" valueMax="56.1" valueMin="40.5">40.5–56.1 mm</quantity>
|
||
,
|
||
<emphasis id="B90BEA8EFFE0020CFF37FE9FFF38FEA8" box="[211,230,333,356]" italics="true" pageId="62" pageNumber="63">N</emphasis>
|
||
=6, respectively); differs from
|
||
<taxonomicName id="4C7F4D1FFFE0020CFDA2FE9FFD1DFEA9" authorityName=", Rai and Anders" authorityYear="2002" box="[582,707,332,357]" class="Amphibia" family="Megophryidae" genus="Xenophrys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="62" pageNumber="63" phylum="Chordata" rank="species" species="robusta">
|
||
<emphasis id="B90BEA8EFFE0020CFDA2FE9FFD1DFEA9" box="[582,707,332,357]" italics="true" pageId="62" pageNumber="63">M. robusta</emphasis>
|
||
</taxonomicName>
|
||
by absence of black dermal asperities on posterior abdomen of adult males (vs. usually present), Finger II<I in length,
|
||
<emphasis id="B90BEA8EFFE0020CFCC6FEA3FCEBFE44" box="[802,821,369,392]" italics="true" pageId="62" pageNumber="63">N</emphasis>
|
||
=13 (vs. Finger I=II,
|
||
<emphasis id="B90BEA8EFFE0020CFBD4FEA3FB9DFE44" box="[1072,1091,369,392]" italics="true" pageId="62" pageNumber="63">N</emphasis>
|
||
=10); differs from
|
||
<taxonomicName id="4C7F4D1FFFE0020CFAFAFEBDFEF5FE61" authority="Mahony & Kamei & Teeling, 2018" authorityName="Mahony & Kamei & Teeling" authorityYear="2018" class="Amphibia" family="Megophryidae" genus="Megophrys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="62" pageNumber="63" phylum="Chordata" rank="genus" status="sp. nov.">
|
||
<emphasis id="B90BEA8EFFE0020CFAFAFEBDFEF5FE61" italics="true" pageId="62" pageNumber="63">Megophrys flavipunctata</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A23857F5FFE0020CFED6FE46FE55FE61" box="[306,395,404,429]" pageId="62" pageNumber="63" rank="species">
|
||
<emphasis id="B90BEA8EFFE0020CFED6FE46FE55FE61" bold="true" box="[306,395,404,429]" pageId="62" pageNumber="63">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
,
|
||
<taxonomicName id="4C7F4D1FFFE0020CFE7DFE46FD46FE60" authority="Mahony & Kamei & Teeling, 2018" authorityName="Mahony & Kamei & Teeling" authorityYear="2018" box="[409,664,404,429]" class="Amphibia" family="Megophryidae" genus="Megophrys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="62" pageNumber="63" phylum="Chordata" rank="genus" status="sp. nov.">
|
||
<emphasis id="B90BEA8EFFE0020CFE7DFE46FD46FE60" box="[409,664,404,429]" italics="true" pageId="62" pageNumber="63">Megophrys oreocrypta</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A23857F5FFE0020CFD7BFE46FD27FE61" box="[671,761,404,429]" pageId="62" pageNumber="63" rank="species">
|
||
<emphasis id="B90BEA8EFFE0020CFD7BFE46FD27FE61" bold="true" box="[671,761,404,429]" pageId="62" pageNumber="63">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
,
|
||
<taxonomicName id="4C7F4D1FFFE0020CFCE3FE47FCB1FE60" authority="Mahony & Kamei & Teeling, 2018" authorityName="Mahony & Kamei & Teeling" authorityYear="2018" box="[775,879,405,428]" class="Amphibia" family="Megophryidae" genus="Megalophrys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="62" pageNumber="63" phylum="Chordata" rank="species" sensu="stricto" species="major">
|
||
<emphasis id="B90BEA8EFFE0020CFCE3FE47FCB1FE60" box="[775,879,405,428]" italics="true" pageId="62" pageNumber="63">M. major</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A23857F5FFE0020CFC92FE47FC44FE60" box="[886,922,405,428]" pageId="62" pageNumber="63" sensu="stricto">s.s.</taxonomicNameLabel>
|
||
and
|
||
<taxonomicName id="4C7F4D1FFFE0020CFC35FE47FB7FFE61" authorityName="Fei & Ye" authorityYear="1990" box="[977,1185,404,429]" class="Amphibia" family="Megophryidae" genus="Megophrys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="62" pageNumber="63" phylum="Chordata" rank="species" species="mangshanensis">
|
||
<emphasis id="B90BEA8EFFE0020CFC35FE47FB7FFE61" box="[977,1185,404,429]" italics="true" pageId="62" pageNumber="63">M. mangshanensis</emphasis>
|
||
</taxonomicName>
|
||
by absence of distinct white, cream or light coloured stripe along upper lips (vs. present); further differs from
|
||
<taxonomicName id="4C7F4D1FFFE0020CFB65FE65FA42FE1C" authority="Mahony & Kamei & Teeling, 2018" authorityName="Mahony & Kamei & Teeling" authorityYear="2018" box="[1153,1436,439,464]" class="Amphibia" family="Megophryidae" genus="Megophrys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="62" pageNumber="63" phylum="Chordata" rank="genus" status="sp. nov.">
|
||
<emphasis id="B90BEA8EFFE0020CFB65FE65FA42FE1C" box="[1153,1436,439,464]" italics="true" pageId="62" pageNumber="63">Megophrys flavipunctata</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A23857F5FFE0020CFF73FE0EFF2EFE39" box="[151,240,476,501]" pageId="62" pageNumber="63" rank="species">
|
||
<emphasis id="B90BEA8EFFE0020CFF73FE0EFF2EFE39" bold="true" box="[151,240,476,501]" pageId="62" pageNumber="63">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
and
|
||
<taxonomicName id="4C7F4D1FFFE0020CFEC3FE0FFE29FE39" authorityName="Fei & Ye" authorityYear="1990" box="[295,503,476,501]" class="Amphibia" family="Megophryidae" genus="Megophrys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="62" pageNumber="63" phylum="Chordata" rank="species" species="mangshanensis">
|
||
<emphasis id="B90BEA8EFFE0020CFEC3FE0FFE29FE39" box="[295,503,476,501]" italics="true" pageId="62" pageNumber="63">M. mangshanensis</emphasis>
|
||
</taxonomicName>
|
||
by its larger adult body size, male SVL
|
||
<quantity id="4C879B79FFE0020CFC58FE0EFBBFFE38" box="[956,1121,476,500]" metricMagnitude="-2" metricUnit="m" metricValue="8.254999999999999" metricValueMax="9.379999999999999" metricValueMin="7.13" pageId="62" pageNumber="63" unit="mm" value="82.55" valueMax="93.8" valueMin="71.3">71.3–93.8 mm</quantity>
|
||
, female SVL 112.0 mm (vs. male SVL
|
||
<quantity id="4C879B79FFE0020CFEF6FE2DFE66FDDB" box="[274,440,511,536]" metricMagnitude="-2" metricUnit="m" metricValue="6.265000000000001" metricValueMax="6.840000000000001" metricValueMin="5.6899999999999995" pageId="62" pageNumber="63" unit="mm" value="62.65" valueMax="68.4" valueMin="56.9">56.9–68.4 mm</quantity>
|
||
,
|
||
<emphasis id="B90BEA8EFFE0020CFE20FDD3FE09FDD4" box="[452,471,513,536]" italics="true" pageId="62" pageNumber="63">N</emphasis>
|
||
=4, female SVL 68.0–
|
||
<quantity id="4C879B79FFE0020CFD30FE2DFCE4FDDB" box="[724,826,511,536]" metricMagnitude="-2" metricUnit="m" metricValue="7.459999999999999" pageId="62" pageNumber="63" unit="mm" value="74.6">74.6 mm</quantity>
|
||
,
|
||
<emphasis id="B90BEA8EFFE0020CFCA3FDD3FC84FDD4" box="[839,858,513,536]" italics="true" pageId="62" pageNumber="63">N</emphasis>
|
||
=3; male SVL
|
||
<quantity id="4C879B79FFE0020CFBE6FE2DFBB7FDDB" box="[1026,1129,511,536]" metricMagnitude="-2" metricUnit="m" metricValue="6.25" pageId="62" pageNumber="63" unit="mm" value="62.5">62.5 mm</quantity>
|
||
,
|
||
<emphasis id="B90BEA8EFFE0020CFB91FDD3FB56FDD4" box="[1141,1160,513,536]" italics="true" pageId="62" pageNumber="63">N</emphasis>
|
||
=1, female SVL
|
||
<quantity id="4C879B79FFE0020CFAA2FDD2FA49FDDB" box="[1350,1431,512,536]" metricMagnitude="-2" metricUnit="m" metricValue="7.3" pageId="62" pageNumber="63" unit="mm" value="73.0">73 mm</quantity>
|
||
,
|
||
<emphasis id="B90BEA8EFFE0020CFF73FDF7FF74FDF0" box="[151,170,549,572]" italics="true" pageId="62" pageNumber="63">N</emphasis>
|
||
=1, respectively); further from
|
||
<taxonomicName id="4C7F4D1FFFE0020CFDEFFDF7FDABFDF0" authority="Mahony & Kamei & Teeling, 2018" authorityName="Mahony & Kamei & Teeling" authorityYear="2018" box="[523,629,549,572]" class="Amphibia" family="Megophryidae" genus="Megalophrys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="62" pageNumber="63" phylum="Chordata" rank="species" sensu="stricto" species="major">
|
||
<emphasis id="B90BEA8EFFE0020CFDEFFDF7FDABFDF0" box="[523,629,549,572]" italics="true" pageId="62" pageNumber="63">M. major</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A23857F5FFE0020CFD99FDF7FD7FFDF0" box="[637,673,549,572]" pageId="62" pageNumber="63" sensu="stricto">s.s.</taxonomicNameLabel>
|
||
by dark spots associated with flank tubercles typically present (vs. absent), toe tips not expanded relative to adjacent toe width (vs. expanded); differs from
|
||
<taxonomicName id="4C7F4D1FFFE0020CFB75FD95FA42FDAC" authority="Mahony & Kamei & Teeling, 2018" authorityName="Mahony & Kamei & Teeling" authorityYear="2018" box="[1169,1436,583,608]" class="Amphibia" family="Megophryidae" genus="Megophrys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="62" pageNumber="63" phylum="Chordata" rank="genus" status="sp. nov.">
|
||
<emphasis id="B90BEA8EFFE0020CFB75FD95FA42FDAC" box="[1169,1436,583,608]" italics="true" pageId="62" pageNumber="63">Megophrys himalayana</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A23857F5FFE0020CFF73FDBEFF2CFD49" box="[151,242,620,645]" pageId="62" pageNumber="63" rank="species">
|
||
<emphasis id="B90BEA8EFFE0020CFF73FDBEFF2CFD49" bold="true" box="[151,242,620,645]" pageId="62" pageNumber="63">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
by its typically larger adult body size, male SVL
|
||
<quantity id="4C879B79FFE0020CFCD7FDBEFC04FD48" box="[819,986,620,644]" metricMagnitude="-2" metricUnit="m" metricValue="8.254999999999999" metricValueMax="9.379999999999999" metricValueMin="7.13" pageId="62" pageNumber="63" unit="mm" value="82.55" valueMax="93.8" valueMin="71.3">71.3–93.8 mm</quantity>
|
||
, female SVL 112.0 mm (vs. male SVL 68.0–
|
||
<quantity id="4C879B79FFE0020CFF32FD5DFEE3FD6B" box="[214,317,655,680]" metricMagnitude="-2" metricUnit="m" metricValue="7.35" pageId="62" pageNumber="63" unit="mm" value="73.5">73.5 mm</quantity>
|
||
,
|
||
<emphasis id="B90BEA8EFFE0020CFEAFFD43FE80FD64" box="[331,350,657,680]" italics="true" pageId="62" pageNumber="63">N</emphasis>
|
||
=7, female SVL
|
||
<quantity id="4C879B79FFE0020CFDFAFD42FD5BFD6B" box="[542,645,656,680]" metricMagnitude="-2" metricUnit="m" metricValue="8.39" pageId="62" pageNumber="63" unit="mm" value="83.9">83.9 mm</quantity>
|
||
,
|
||
<emphasis id="B90BEA8EFFE0020CFD76FD43FD7BFD64" box="[658,677,657,680]" italics="true" pageId="62" pageNumber="63">N</emphasis>
|
||
=1), typically smaller eye diameter to snout length ratio for males, ED/ES 76.6–91.5%, mean 84.3 ± 4.2% (vs. ED/ES 87.6–101.1%,
|
||
<emphasis id="B90BEA8EFFE0020CFC99FD67FC4EFD00" box="[893,912,693,716]" italics="true" pageId="62" pageNumber="63">N</emphasis>
|
||
=7, mean 94.2 ± 4.6%).
|
||
</paragraph>
|
||
<paragraph id="8BC0369CFFE0020CFF23FD05FDF9FCD8" blockId="62.[151,1437,151,2013]" pageId="62" pageNumber="63">
|
||
<emphasis id="B90BEA8EFFE0020CFF23FD05FE69FD3C" bold="true" box="[199,439,727,752]" pageId="62" pageNumber="63">Systematic position.</emphasis>
|
||
This taxon represents
|
||
<taxonomicName id="4C7F4D1FFFE0020CFD59FD0BFC93FD3C" authorityName="Mahony & Kamei & Teeling" authorityYear="2018" box="[701,845,727,752]" class="Amphibia" family="Megophryidae" genus="Megalophrys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="62" pageNumber="63" phylum="Chordata" rank="species" species="major">
|
||
<emphasis id="B90BEA8EFFE0020CFD59FD0BFD02FD3C" box="[701,732,729,752]" italics="true" pageId="62" pageNumber="63">M.</emphasis>
|
||
cf.
|
||
<emphasis id="B90BEA8EFFE0020CFCEEFD0BFC93FD3C" box="[778,845,729,752]" italics="true" pageId="62" pageNumber="63">major</emphasis>
|
||
</taxonomicName>
|
||
4 (OTU 12) in the molecular analyses, and “
|
||
<taxonomicName id="4C7F4D1FFFE0020CFABCFD0BFF04FCD8" authorityName="Mahony & Kamei & Teeling" authorityYear="2018" class="Amphibia" family="Megophryidae" genus="Megalophrys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="62" pageNumber="63" phylum="Chordata" rank="species" species="major">
|
||
<emphasis id="B90BEA8EFFE0020CFABCFD0BFAA9FD3C" box="[1368,1399,729,752]" italics="true" pageId="62" pageNumber="63">M.</emphasis>
|
||
cf.
|
||
<emphasis id="B90BEA8EFFE0020CFF73FD2FFF04FCD8" box="[151,218,765,788]" italics="true" pageId="62" pageNumber="63">major</emphasis>
|
||
</taxonomicName>
|
||
[5]” in
|
||
<bibRefCitation id="EFEE4B6DFFE0020CFED6FD2EFDFDFCD8" author="Mahony, S. & Foley, N. C. & Biju, S. D. & Teeling, E. C." box="[306,547,764,789]" pageId="62" pageNumber="63" pagination="744 - 771" refId="ref55439" refString="Mahony, S., Foley, N. C., Biju, S. D. & Teeling, E. C. (2017) Evolutionary history of the Asian Horned Frogs (Megophryinae): integrative approaches to timetree dating in the absence of a fossil record. Molecular Biology and Evolution, 34 (3), 744 - 771 + S. I. 33 pp. https: // doi. org / 10.1093 / molbev / msw 267" type="book chapter" year="2017">
|
||
Mahony
|
||
<emphasis id="B90BEA8EFFE0020CFE72FD2FFE10FCD9" box="[406,462,764,789]" italics="true" pageId="62" pageNumber="63">et al.</emphasis>
|
||
(2017)
|
||
</bibRefCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BC0369CFFE0020CFF23FCCDFB07FBF9" blockId="62.[151,1437,151,2013]" pageId="62" pageNumber="63">
|
||
<taxonomicName id="4C7F4D1FFFE0020CFF23FCCDFE7FFCF4" authority="Mahony & Kamei & Teeling, 2018" authorityName="Mahony & Kamei & Teeling" authorityYear="2018" box="[199,417,799,824]" class="Amphibia" family="Megophryidae" genus="Megophrys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="62" pageNumber="63" phylum="Chordata" rank="genus" status="sp. nov.">
|
||
<emphasis id="B90BEA8EFFE0020CFF23FCCDFE7FFCF4" box="[199,417,799,824]" italics="true" pageId="62" pageNumber="63">Megophrys periosa</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A23857F5FFE0020CFE4DFCCDFDDAFCF4" box="[425,516,799,824]" pageId="62" pageNumber="63" rank="species">
|
||
<emphasis id="B90BEA8EFFE0020CFE4DFCCDFDDAFCF4" bold="true" box="[425,516,799,824]" pageId="62" pageNumber="63">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
consistently formed a sister taxa relationship with two populations found east of the Brahmaputra River in Manipur and
|
||
<collectingCountry id="F368760CFFE0020CFDB6FC97FD61FC91" box="[594,703,837,861]" name="Myanmar" pageId="62" pageNumber="63">Myanmar</collectingCountry>
|
||
referred to herein as
|
||
<taxonomicName id="4C7F4D1FFFE0020CFC49FC96FB77FC90" authorityName="Mahony & Kamei & Teeling" authorityYear="2018" box="[941,1193,836,861]" class="Amphibia" family="Megophryidae" genus="Megophrys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="62" pageNumber="63" phylum="Chordata" rank="species" species="periosa">
|
||
<emphasis id="B90BEA8EFFE0020CFC49FC96FBF5FC91" box="[941,1067,836,861]" italics="true" pageId="62" pageNumber="63">Megophrys</emphasis>
|
||
cf.
|
||
<emphasis id="B90BEA8EFFE0020CFBB1FC97FB77FC90" box="[1109,1193,837,860]" italics="true" pageId="62" pageNumber="63">periosa</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<taxonomicName id="4C7F4D1FFFE0020CFB5EFC97FA98FC90" authorityName="Mahony & Kamei & Teeling" authorityYear="2018" box="[1210,1350,836,861]" class="Amphibia" family="Megophryidae" genus="Megalophrys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="62" pageNumber="63" phylum="Chordata" rank="species" species="major">
|
||
<emphasis id="B90BEA8EFFE0020CFB5EFC97FB06FC90" box="[1210,1240,837,860]" italics="true" pageId="62" pageNumber="63">M.</emphasis>
|
||
cf.
|
||
<emphasis id="B90BEA8EFFE0020CFAE7FC97FA98FC90" box="[1283,1350,837,860]" italics="true" pageId="62" pageNumber="63">major</emphasis>
|
||
</taxonomicName>
|
||
6/OTU
|
||
<quantity id="4C879B79FFE0020CFF73FCBAFF0CFC4C" box="[151,210,871,896]" metricMagnitude="-1" metricUnit="m" metricValue="3.556" pageId="62" pageNumber="63" unit="in" value="14.0">14 in</quantity>
|
||
molecular analyses). Uncorrected
|
||
<emphasis id="B90BEA8EFFE0020CFDB9FCBBFDB5FC4C" box="[605,619,873,896]" italics="true" pageId="62" pageNumber="63">p</emphasis>
|
||
-distances for the 16S rRNA gene between
|
||
<taxonomicName id="4C7F4D1FFFE0020CFBB9FCB5FAE6FC4C" authority="Mahony & Kamei & Teeling, 2018" authorityName="Mahony & Kamei & Teeling" authorityYear="2018" box="[1117,1336,871,896]" class="Amphibia" family="Megophryidae" genus="Megophrys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="62" pageNumber="63" phylum="Chordata" rank="genus" status="sp. nov.">
|
||
<emphasis id="B90BEA8EFFE0020CFBB9FCB5FAE6FC4C" box="[1117,1336,871,896]" italics="true" pageId="62" pageNumber="63">Megophrys periosa</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A23857F5FFE0020CFAA5FCB5FA42FC4C" box="[1345,1436,871,896]" pageId="62" pageNumber="63" rank="species">
|
||
<emphasis id="B90BEA8EFFE0020CFAA5FCB5FA42FC4C" bold="true" box="[1345,1436,871,896]" pageId="62" pageNumber="63">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
and
|
||
<taxonomicName id="4C7F4D1FFFE0020CFF22FC5FFEBDFC68" authorityName="Mahony & Kamei & Teeling" authorityYear="2018" box="[198,355,908,933]" class="Amphibia" family="Megophryidae" genus="Megophrys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="62" pageNumber="63" phylum="Chordata" rank="species" species="periosa">
|
||
<emphasis id="B90BEA8EFFE0020CFF22FC5FFF3BFC68" box="[198,229,909,932]" italics="true" pageId="62" pageNumber="63">M.</emphasis>
|
||
cf.
|
||
<emphasis id="B90BEA8EFFE0020CFEEBFC5FFEBDFC68" box="[271,355,909,932]" italics="true" pageId="62" pageNumber="63">periosa</emphasis>
|
||
</taxonomicName>
|
||
was 1.6–2.0% (Appendix I, Table 6). The systematic position of this clade within the MMC was not fully resolved (see Systematic position section for
|
||
<taxonomicName id="4C7F4D1FFFE0020CFD10FC7DFC23FC04" authority="Mahony & Kamei & Teeling, 2018" authorityName="Mahony & Kamei & Teeling" authorityYear="2018" box="[756,1021,943,968]" class="Amphibia" family="Megophryidae" genus="Megophrys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="62" pageNumber="63" phylum="Chordata" rank="genus" status="sp. nov.">
|
||
<emphasis id="B90BEA8EFFE0020CFD10FC7DFC23FC04" box="[756,1021,943,968]" italics="true" pageId="62" pageNumber="63">Megophrys himalayana</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A23857F5FFE0020CFBE7FC7DFB82FC04" box="[1027,1116,943,968]" pageId="62" pageNumber="63" rank="species">
|
||
<emphasis id="B90BEA8EFFE0020CFBE7FC7DFB82FC04" bold="true" box="[1027,1116,943,968]" pageId="62" pageNumber="63">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
for further details), however most analyses placed this species as the sister taxon of
|
||
<taxonomicName id="4C7F4D1FFFE0020CFCE1FC06FBD1FC21" authority="Mahony & Kamei & Teeling, 2018" authorityName="Mahony & Kamei & Teeling" authorityYear="2018" box="[773,1039,980,1005]" class="Amphibia" family="Megophryidae" genus="Megophrys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="62" pageNumber="63" phylum="Chordata" rank="genus" status="sp. nov.">
|
||
<emphasis id="B90BEA8EFFE0020CFCE1FC06FBD1FC21" box="[773,1039,980,1005]" italics="true" pageId="62" pageNumber="63">Megophrys himalayana</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A23857F5FFE0020CFBF2FC06FBAEFC21" box="[1046,1136,980,1005]" pageId="62" pageNumber="63" rank="species">
|
||
<emphasis id="B90BEA8EFFE0020CFBF2FC06FBAEFC21" bold="true" box="[1046,1136,980,1005]" pageId="62" pageNumber="63">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
(
|
||
<figureCitation id="13442A19FFE0020CFB64FC06FB35FC20" box="[1152,1259,980,1005]" captionStart="FIGURE 2" captionStartId="11.[151,250,1148,1170]" captionTargetBox="[241,1332,252,1087]" captionTargetId="figure@11.[151,1436,193,1127]" captionTargetPageId="11" captionText="FIGURE 2. BI concatenated gene tree based on the 47 taxa concatenated mt+nuDNA alignment (Dataset C), with ML node values provided. Posterior probability/bootstrap values>95% in both analyses are represented by a black spot on the node, values <95% in one or both analyses are provided next to relevant nodes, where * is>95%, and – is given for nodes that were not resolved on the ML tree. Taxon names in pink represent the eastern clade, blue represents the western clade (southern subclade), and green represents the western clade (northern subclade). Revised names for taxa based on the taxonomic review are provided in parentheses." httpUri="https://zenodo.org/record/2610206/files/figure.png" pageId="62" pageNumber="63">Figures 2</figureCitation>
|
||
&
|
||
<figureCitation id="13442A19FFE0020CFAF4FC06FAFFFC20" box="[1296,1313,980,1004]" captionStart="FIGURE 4" captionStartId="13.[151,250,1115,1137]" captionTargetBox="[155,1430,193,1084]" captionTargetId="figure@13.[150,1436,193,1094]" captionTargetPageId="13" captionText="FIGURE 4. RAxML tree for the combined mt+nuDNA dataset expanded to include the Chen et al. (2017) sequences (Dataset J). Bootstrap support values are provided adjacent to the associated nodes. Refer to Appendix I, Table 2 for revised names for taxa based on the taxonomic review." httpUri="https://zenodo.org/record/2610210/files/figure.png" pageId="62" pageNumber="63">4</figureCitation>
|
||
; Appendix I, Table 3; Appendix II,
|
||
<figureCitation id="13442A19FFE0020CFE54FC25FDC5FBDC" box="[432,539,1015,1040]" captionStart="FIGURE 1" captionStartId="10.[151,250,1250,1272]" captionTargetBox="[163,1425,384,1202]" captionTargetId="figure@10.[150,1437,373,1228]" captionTargetPageId="10" captionText="FIGURE 1. Ultrametric guide tree topologies used for BPP analyses: A. topology based on *BEAST analyses of phased nuclear alignments, with posterior probability values for nodes; B. alternative topology guide tree based on the results of concatenated mt+nuDNA gene tree analyses (Dataset E); C. alternative topology based on *BEAST analyses of unphased nuclear alignments, with posterior probability values for nodes. Taxon names in pink represent the eastern clade, blue represents the western clade (southern subclade), and green represents the western clade (northern subclade). Refer to Appendix I, Table 2 for revised names for taxa based on the taxonomic review." httpUri="https://zenodo.org/record/2610204/files/figure.png" pageId="62" pageNumber="63">Figures 1</figureCitation>
|
||
,
|
||
<figureCitation id="13442A19FFE0020CFDCFFC2AFDE7FBDC" box="[555,569,1016,1040]" captionStart="FIGURE 2" captionStartId="11.[151,250,1148,1170]" captionTargetBox="[241,1332,252,1087]" captionTargetId="figure@11.[151,1436,193,1127]" captionTargetPageId="11" captionText="FIGURE 2. BI concatenated gene tree based on the 47 taxa concatenated mt+nuDNA alignment (Dataset C), with ML node values provided. Posterior probability/bootstrap values>95% in both analyses are represented by a black spot on the node, values <95% in one or both analyses are provided next to relevant nodes, where * is>95%, and – is given for nodes that were not resolved on the ML tree. Taxon names in pink represent the eastern clade, blue represents the western clade (southern subclade), and green represents the western clade (northern subclade). Revised names for taxa based on the taxonomic review are provided in parentheses." httpUri="https://zenodo.org/record/2610206/files/figure.png" pageId="62" pageNumber="63">2</figureCitation>
|
||
&
|
||
<figureCitation id="13442A19FFE0020CFD85FC25FDAFFBDC" box="[609,625,1015,1040]" captionStart="FIGURE 5" captionStartId="13.[151,250,1843,1865]" captionTargetBox="[162,1426,1248,1806]" captionTargetId="figure@13.[151,1436,1248,1822]" captionTargetPageId="13" captionText="FIGURE 5. DensiTree cloudogram representations of *BEAST analyses, removing the first 10% of trees as burn-in, produced from the four nuDNA gene alignments of Dataset E: A. original unphased; B. phased. Dark blue branches represent the most frequently occurring topology, light blue branches represent the consensus trees with intensity proportional to the frequency of the topology in the dataset, and green areas represent all topologies in the dataset. Refer to Appendix I, Table 2 for revised names for taxa based on the taxonomic review." httpUri="https://zenodo.org/record/2610212/files/figure.png" pageId="62" pageNumber="63">5</figureCitation>
|
||
;
|
||
<bibRefCitation id="EFEE4B6DFFE0020CFD9BFC25FCBBFBDC" author="Mahony, S. & Foley, N. C. & Biju, S. D. & Teeling, E. C." box="[639,869,1015,1040]" pageId="62" pageNumber="63" pagination="744 - 771" refId="ref55439" refString="Mahony, S., Foley, N. C., Biju, S. D. & Teeling, E. C. (2017) Evolutionary history of the Asian Horned Frogs (Megophryinae): integrative approaches to timetree dating in the absence of a fossil record. Molecular Biology and Evolution, 34 (3), 744 - 771 + S. I. 33 pp. https: // doi. org / 10.1093 / molbev / msw 267" type="book chapter" year="2017">
|
||
Mahony
|
||
<emphasis id="B90BEA8EFFE0020CFD01FC2BFCC1FBDC" box="[741,799,1015,1040]" italics="true" pageId="62" pageNumber="63">et al.</emphasis>
|
||
2017
|
||
</bibRefCitation>
|
||
). Refer to Appendix I, Table 6 for uncorrected
|
||
<emphasis id="B90BEA8EFFE0020CFA60FC2BFA4CFBDC" box="[1412,1426,1017,1040]" italics="true" pageId="62" pageNumber="63">p</emphasis>
|
||
- distances for the 16S rRNA gene between
|
||
<taxonomicName id="4C7F4D1FFFE0020CFD97FBCEFC92FBF8" authority="Mahony & Kamei & Teeling, 2018" authorityName="Mahony & Kamei & Teeling" authorityYear="2018" box="[627,844,1052,1077]" class="Amphibia" family="Megophryidae" genus="Megophrys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="62" pageNumber="63" phylum="Chordata" rank="genus" status="sp. nov.">
|
||
<emphasis id="B90BEA8EFFE0020CFD97FBCEFC92FBF8" box="[627,844,1052,1077]" italics="true" pageId="62" pageNumber="63">Megophrys periosa</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A23857F5FFE0020CFCB7FBCEFC73FBF9" box="[851,941,1052,1077]" pageId="62" pageNumber="63" rank="species">
|
||
<emphasis id="B90BEA8EFFE0020CFCB7FBCEFC73FBF9" bold="true" box="[851,941,1052,1077]" pageId="62" pageNumber="63">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
and other MMSG species.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3656517FFE0020CFF23FBEDFCBAFB6C" pageId="62" pageNumber="63" type="etymology">
|
||
<paragraph id="8BC0369CFFE0020CFF23FBEDFA8DFBB0" blockId="62.[151,1437,151,2013]" pageId="62" pageNumber="63">
|
||
<emphasis id="B90BEA8EFFE0020CFF23FBEDFE90FB94" bold="true" box="[199,334,1087,1112]" pageId="62" pageNumber="63">Etymology.</emphasis>
|
||
The species epithet “
|
||
<taxonomicName id="4C7F4D1FFFE0020CFDA6FB93FD48FB94" authorityName="Mahony & Kamei & Teeling" authorityYear="2018" box="[578,662,1089,1112]" class="Amphibia" family="Megophryidae" genus="Megophrys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="62" pageNumber="63" phylum="Chordata" rank="species" species="periosa">
|
||
<emphasis id="B90BEA8EFFE0020CFDA6FB93FD48FB94" box="[578,662,1089,1112]" italics="true" pageId="62" pageNumber="63">periosa</emphasis>
|
||
</taxonomicName>
|
||
” is a Latinised Greek adjective meaning “immense”, in reference to the very large size that this species attains, even compared to otherwise generally large species in the MMC.
|
||
</paragraph>
|
||
<paragraph id="8BC0369CFFE0020CFF23FB55FCBAFB6C" blockId="62.[151,1437,151,2013]" box="[199,868,1159,1184]" pageId="62" pageNumber="63">
|
||
<emphasis id="B90BEA8EFFE0020CFF23FB55FE20FB6C" bold="true" box="[199,510,1159,1184]" pageId="62" pageNumber="63">Suggested common name:</emphasis>
|
||
Giant Himalayan Horned Frog.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3656517FFE0020CFF23FB7EFBAAF92C" pageId="62" pageNumber="63" type="distribution">
|
||
<paragraph id="8BC0369CFFE0020CFF23FB7EFB48FAB4" blockId="62.[151,1437,151,2013]" pageId="62" pageNumber="63">
|
||
<emphasis id="B90BEA8EFFE0020CFF23FB7EFEBEFB09" bold="true" box="[199,352,1196,1221]" pageId="62" pageNumber="63">Distribution.</emphasis>
|
||
<taxonomicName id="4C7F4D1FFFE0020CFE8DFB7EFD9AFB08" authority="Mahony & Kamei & Teeling, 2018" authorityName="Mahony & Kamei & Teeling" authorityYear="2018" box="[361,580,1196,1221]" class="Amphibia" family="Megophryidae" genus="Megophrys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="62" pageNumber="63" phylum="Chordata" rank="genus" status="sp. nov.">
|
||
<emphasis id="B90BEA8EFFE0020CFE8DFB7EFD9AFB08" box="[361,580,1196,1221]" italics="true" pageId="62" pageNumber="63">Megophrys periosa</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A23857F5FFE0020CFDA9FB7EFD76FB08" box="[589,680,1196,1221]" pageId="62" pageNumber="63" rank="species">
|
||
<emphasis id="B90BEA8EFFE0020CFDA9FB7EFD7FFB09" bold="true" box="[589,673,1196,1221]" pageId="62" pageNumber="63">sp. nov</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
is so far confirmed from three localities in the state of
|
||
<collectingRegion id="49BBF87EFFE0020CFAC3FB7EFF2DFB24" country="India" name="Arunachal Pradesh" pageId="62" pageNumber="63">Arunachal Pradesh</collectingRegion>
|
||
, Northeast
|
||
<collectingCountry id="F368760CFFE0020CFE9CFB1DFE6BFB24" box="[376,437,1231,1256]" name="India" pageId="62" pageNumber="63">India</collectingCountry>
|
||
. The currently known distribution ranges from East Siang district, west through West Siang district, to at least West Kameng district, between 260 and
|
||
<quantity id="4C879B79FFE0020CFC9CFB26FC12FAC0" box="[888,972,1268,1292]" metricMagnitude="3" metricUnit="m" metricValue="1.1099999999999999" pageId="62" pageNumber="63" unit="m" value="1110.0">1110 m</quantity>
|
||
asl. (
|
||
<figureCitation id="13442A19FFE0020CFBEDFB26FB5FFAC0" box="[1033,1153,1268,1293]" captionStart="FIGURE 8" captionStartId="21.[151,250,1742,1764]" captionTargetBox="[161,1425,1085,1720]" captionTargetId="figure@21.[159,1427,1083,1721]" captionTargetPageId="21" captionText="FIGURE 8. Distribution map of the western MMSG taxa: A. All taxa (excluding M. monticola) from Northeast India and surrounding countries: diamonds represent type localities, circles represent referred localities based on molecularly sampled specimens: orange = M. zhangi; yellow = M. robusta; black = M. medogensis; dark blue = M. major; pink = Megophrys flavipunctata sp. nov.; dark green = Megophrys oreocrypta sp. nov.; light blue = Megophrys himalayana sp. nov.; light green = Megophrys periosa sp. nov.; pale blue = M. cf. periosa; purple = M. glandulosa. B. Inset rectangle in Figure A enlarged to show the distribution map for M. monticola from Darjeeling and Sikkim region, Northeast India, showing distribution of named mitochondrial haplotypes: stars represent the type localities, circles represent referred localities confirmed from sequence data: red = M. “sanu” comb. nov., light blue = M. “katabhako” comb. nov. Blue lines represent major rivers; grey lines represent international borders." httpUri="https://zenodo.org/record/2610218/files/figure.png" pageId="62" pageNumber="63">Figure 8A</figureCitation>
|
||
). An improved and more extensive sampling is necessary to define the east-west limits of this species’ geographic range. Specimens assigned here to
|
||
<taxonomicName id="4C7F4D1FFFE0020CFE80FAEEFDB0FA98" authorityName="Mahony & Kamei & Teeling" authorityYear="2018" box="[356,622,1340,1365]" class="Amphibia" family="Megophryidae" genus="Megophrys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="62" pageNumber="63" phylum="Chordata" rank="species" species="periosa">
|
||
<emphasis id="B90BEA8EFFE0020CFE80FAEEFE3CFA99" box="[356,482,1340,1365]" italics="true" pageId="62" pageNumber="63">Megophrys</emphasis>
|
||
cf.
|
||
<emphasis id="B90BEA8EFFE0020CFDFEFAEFFDB0FA98" box="[538,622,1341,1364]" italics="true" pageId="62" pageNumber="63">periosa</emphasis>
|
||
</taxonomicName>
|
||
were collected from Ukhrul district (at
|
||
<quantity id="4C879B79FFE0020CFBB3FAEEFB7AFA98" box="[1111,1188,1340,1364]" metricMagnitude="2" metricUnit="m" metricValue="8.2" pageId="62" pageNumber="63" unit="m" value="820.0">820 m</quantity>
|
||
asl.),
|
||
<collectingRegion id="49BBF87EFFE0020CFB10FAEEFA8BFA99" box="[1268,1365,1340,1365]" country="India" name="Manipur" pageId="62" pageNumber="63">Manipur</collectingRegion>
|
||
state, Northeast
|
||
<collectingCountry id="F368760CFFE0020CFEEFFA8DFE99FAB4" box="[267,327,1375,1400]" name="India" pageId="62" pageNumber="63">India</collectingCountry>
|
||
, and from Myitkyina district,
|
||
<collectingRegion id="49BBF87EFFE0020CFD73FA8DFD36FAB4" box="[663,744,1375,1400]" country="Myanmar" name="Kachin" pageId="62" pageNumber="63">Kachin</collectingRegion>
|
||
state, northern
|
||
<collectingCountry id="F368760CFFE0020CFC72FAB3FBDDFAB5" box="[918,1027,1377,1401]" name="Myanmar" pageId="62" pageNumber="63">Myanmar</collectingCountry>
|
||
(
|
||
<figureCitation id="13442A19FFE0020CFBF5FA8DFB57FAB4" box="[1041,1161,1375,1400]" captionStart="FIGURE 8" captionStartId="21.[151,250,1742,1764]" captionTargetBox="[161,1425,1085,1720]" captionTargetId="figure@21.[159,1427,1083,1721]" captionTargetPageId="21" captionText="FIGURE 8. Distribution map of the western MMSG taxa: A. All taxa (excluding M. monticola) from Northeast India and surrounding countries: diamonds represent type localities, circles represent referred localities based on molecularly sampled specimens: orange = M. zhangi; yellow = M. robusta; black = M. medogensis; dark blue = M. major; pink = Megophrys flavipunctata sp. nov.; dark green = Megophrys oreocrypta sp. nov.; light blue = Megophrys himalayana sp. nov.; light green = Megophrys periosa sp. nov.; pale blue = M. cf. periosa; purple = M. glandulosa. B. Inset rectangle in Figure A enlarged to show the distribution map for M. monticola from Darjeeling and Sikkim region, Northeast India, showing distribution of named mitochondrial haplotypes: stars represent the type localities, circles represent referred localities confirmed from sequence data: red = M. “sanu” comb. nov., light blue = M. “katabhako” comb. nov. Blue lines represent major rivers; grey lines represent international borders." httpUri="https://zenodo.org/record/2610218/files/figure.png" pageId="62" pageNumber="63">Figure 8A</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BC0369CFFE0020CFF23FA56FBAAF92C" blockId="62.[151,1437,151,2013]" pageId="62" pageNumber="63">
|
||
<emphasis id="B90BEA8EFFE0020CFF23FA56FDC6FA51" bold="true" box="[199,536,1412,1437]" pageId="62" pageNumber="63">Habitat and natural history.</emphasis>
|
||
All males were in breeding condition, and the female contained well-developed ova within her ovaries, indicative that the breeding season extends at least through late-July and early-August. All specimens were collected after dusk, typically perched on the rocky banks of small to large (
|
||
<quantity id="4C879B79FFE0020CFB12FA1EFA8EFA28" box="[1270,1360,1484,1509]" metricMagnitude="0" metricUnit="m" metricValue="8.0" metricValueMax="15.0" metricValueMin="1.0" pageId="62" pageNumber="63" unit="m" value="8.0" valueMax="15.0" valueMin="1.0">1–15 m</quantity>
|
||
wide) moderately fast flowing mountain streams flowing through dense mature secondary/primary forest (e.g.,
|
||
<figureCitation id="13442A19FFE0020CFACEFA3DFF19F9E0" captionStart="FIGURE 9" captionStartId="26.[151,250,1801,1823]" captionTargetBox="[329,1255,196,1769]" captionTargetId="figure@26.[329,1255,195,1769]" captionTargetPageId="26" captionText="FIGURE 9. Habitat images for collection localities of MMSG species in Northeast India. A. Sessa River, near Sessa village (1110 m asl.), West Kameng dist., Arunachal Pradesh, collection locality of Megophrys robusta and Megophrys periosa sp. nov.; B. and D. Mawphlang Sacred Grove (~1810 m asl.), near Mawphlang village, East Khasi Hills dist., Meghalaya, type locality of Megophrys flavipunctata sp. nov.; C. Senchal Wildlife Sanctuary (2220 m asl.), Rambi (East Range), Darjeeling Sadar subdiv., Darjeeling dist., West Bengal, collection locality of M. monticola; E. Sessa River near Elephant village (370 m asl.), West Kameng dist., Arunachal Pradesh, off which smaller side streams were the type locality of Megophrys himalayana sp. nov.; F. stream near Pangin town (450 m asl.), East Siang dist., Arunachal Pradesh, type locality of Megophrys periosa sp. nov." httpUri="https://zenodo.org/record/2610220/files/figure.png" pageId="62" pageNumber="63">Figure 9A & F</figureCitation>
|
||
). Males spaced themselves along the banks of the streams at least
|
||
<quantity id="4C879B79FFE0020CFC4FF9C6FC3DF9E0" box="[939,995,1556,1580]" metricMagnitude="1" metricUnit="m" metricValue="1.0" pageId="62" pageNumber="63" unit="m" value="10.0">10 m</quantity>
|
||
apart. Vocalisations typically consisted of a succession of a few notes followed by an extended silence, but were rarely heard. Healed injuries were observed on two specimens; the right hand of the
|
||
<typeStatus id="54C4883EFFE0020CFD26F98EFCFAF9B9" box="[706,804,1628,1653]" pageId="62" pageNumber="63" type="holotype">holotype</typeStatus>
|
||
appears to have been badly dislocated or broken (
|
||
<figureCitation id="13442A19FFE0020CFAB6F98EFF23F954" captionStart="FIGURE 22" captionStartId="59.[151,250,1767,1789]" captionTargetBox="[151,1436,301,1746]" captionTargetId="figure@59.[151,1436,301,1746]" captionTargetPageId="59" captionText="FIGURE 22. Megophrys periosa sp. nov. holotype: adult male (BNHS 6055: SVL 93.8 mm) in preservation: A. dorsal view; B. ventral view; C. profile view of head; D. ventral view of hand; E. ventral view of foot." httpUri="https://zenodo.org/record/2610246/files/figure.png" pageId="62" pageNumber="63">Figure 22B & E</figureCitation>
|
||
). This large mature male had noticeably less well-developed nuptial pads than the other males, possibly as a result of lower overall fitness, although it otherwise appeared to be in good health. One other specimen (BNHS 6064) had a large swelling on Finger II on the right hand, the cause for which is unclear.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3656517FFE0020DFF23F93EFD51FF19" lastPageId="63" lastPageNumber="64" pageId="62" pageNumber="63" type="discussion">
|
||
<paragraph id="8BC0369CFFE0020DFF23F93EFD51FF19" blockId="62.[151,1437,151,2013]" lastBlockId="63.[151,1436,151,213]" lastPageId="63" lastPageNumber="64" pageId="62" pageNumber="63">
|
||
<emphasis id="B90BEA8EFFE0020CFF23F93EFEE5F8C9" bold="true" box="[199,315,1772,1797]" pageId="62" pageNumber="63">Remarks.</emphasis>
|
||
Specimens reported as
|
||
<taxonomicName id="4C7F4D1FFFE0020CFDA7F93FFD1BF8C9" authorityName="Anderson" authorityYear="1871" box="[579,709,1772,1797]" class="Amphibia" family="Rhacophoridae" genus="Ixalus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="62" pageNumber="63" phylum="Chordata" rank="species" species="lateralis">
|
||
<emphasis id="B90BEA8EFFE0020CFDA7F93FFD1BF8C9" box="[579,709,1772,1797]" italics="true" pageId="62" pageNumber="63">M. lateralis</emphasis>
|
||
</taxonomicName>
|
||
and/or
|
||
<taxonomicName id="4C7F4D1FFFE0020CFCFEF93FFC4AF8C9" authorityName=", Rai and Anders" authorityYear="2002" box="[794,916,1772,1797]" class="Amphibia" family="Megophryidae" genus="Xenophrys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="62" pageNumber="63" phylum="Chordata" rank="species" species="robusta">
|
||
<emphasis id="B90BEA8EFFE0020CFCFEF93FFC4AF8C9" box="[794,916,1772,1797]" italics="true" pageId="62" pageNumber="63">M. robusta</emphasis>
|
||
</taxonomicName>
|
||
by
|
||
<bibRefCitation id="EFEE4B6DFFE0020CFC59F93EFB30F8C8" author="Bordoloi, S. & Borah, M. M." box="[957,1262,1772,1797]" pageId="62" pageNumber="63" pagination="191 - 193" refId="ref50702" refString="Bordoloi, S. & Borah, M. M. (2001) First record of a frog Megophrys robusta (Boulenger 1908) from Arunachal Pradesh, India. Cotton College Research Journal, 1, 191 - 193." type="journal article" year="2001">Borah and Bordoloi (2001)</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="EFEE4B6DFFE0020CFB1EF93EFF3FF8E4" author="Bordoloi, S. & Borah, M. M. & Sharma, K. K. & Kalita, S. N." pageId="62" pageNumber="63" pagination="363 - 371" refId="ref50744" refString="Bordoloi, S., Borah, M. M., Sharma, K. K. & Kalita, S. N. (2000) An overview of amphibian fauna of Arunachal Pradesh, India. Journal of the Assam Science Society, 41 (4), 363 - 371." type="journal article" year="2000">
|
||
Bordoloi
|
||
<emphasis id="B90BEA8EFFE0020CFA80F93FFA42F8C9" box="[1380,1436,1772,1797]" italics="true" pageId="62" pageNumber="63">et al.</emphasis>
|
||
(2000)
|
||
</bibRefCitation>
|
||
and
|
||
<bibRefCitation id="EFEE4B6DFFE0020CFEC3F8DDFDE2F8E4" author="Sarkar, A. K. & Ray, S." box="[295,572,1807,1833]" pageId="62" pageNumber="63" pagination="285 - 316" refId="ref57338" refString="Sarkar, A. K. & Ray, S. (2006) Amphibia. In: Alfred, J. R. B. (Ed.), Fauna of Arunachal Pradesh. State Fauna Series. 13 (1). Zoological Survey of India, Kolkata, pp. 285 - 316." type="book chapter" year="2006">Sarkar and Ray (2006)</bibRefCitation>
|
||
from
|
||
<collectingRegion id="49BBF87EFFE0020CFD74F8DDFCB2F8E4" box="[656,876,1807,1832]" country="India" name="Arunachal Pradesh" pageId="62" pageNumber="63">Arunachal Pradesh</collectingRegion>
|
||
may apply to this species, and/or
|
||
<taxonomicName id="4C7F4D1FFFE0020CFAFAF8DDFEC2F881" authority="Mahony & Kamei & Teeling, 2018" authorityName="Mahony & Kamei & Teeling" authorityYear="2018" class="Amphibia" family="Megophryidae" genus="Megophrys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="62" pageNumber="63" phylum="Chordata" rank="genus" status="sp. nov.">
|
||
<emphasis id="B90BEA8EFFE0020CFAFAF8DDFEC2F881" italics="true" pageId="62" pageNumber="63">Megophrys himalayana</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A23857F5FFE0020CFEC2F8E6FE5AF880" box="[294,388,1844,1869]" pageId="62" pageNumber="63" rank="species">
|
||
<emphasis id="B90BEA8EFFE0020CFEC2F8E6FEA0F881" bold="true" box="[294,382,1844,1869]" pageId="62" pageNumber="63">sp. nov</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
The specimens reported in these studies should be re-examined to determine the correct identities. Populations provisionally assigned to this taxa (as
|
||
<taxonomicName id="4C7F4D1FFFE0020CFCA0F88BFC3FF8BC" authorityName="Mahony & Kamei & Teeling" authorityYear="2018" box="[836,993,1879,1904]" class="Amphibia" family="Megophryidae" genus="Megophrys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="62" pageNumber="63" phylum="Chordata" rank="species" species="periosa">
|
||
<emphasis id="B90BEA8EFFE0020CFCA0F88BFCBDF8BC" box="[836,867,1881,1904]" italics="true" pageId="62" pageNumber="63">M.</emphasis>
|
||
cf.
|
||
<emphasis id="B90BEA8EFFE0020CFC69F88BFC3FF8BC" box="[909,993,1881,1904]" italics="true" pageId="62" pageNumber="63">periosa</emphasis>
|
||
</taxonomicName>
|
||
) from eastern
|
||
<collectingRegion id="49BBF87EFFE0020CFB62F885FB39F8BC" box="[1158,1255,1879,1904]" country="India" name="Manipur" pageId="62" pageNumber="63">Manipur</collectingRegion>
|
||
state, Northeast
|
||
<collectingCountry id="F368760CFFE0020CFF73F8AEFF0FF859" box="[151,209,1916,1941]" name="India" pageId="62" pageNumber="63">India</collectingCountry>
|
||
and northern
|
||
<collectingCountry id="F368760CFFE0020CFE89F8AFFE02F859" box="[365,476,1917,1941]" name="Myanmar" pageId="62" pageNumber="63">Myanmar</collectingCountry>
|
||
, were found to be the sister taxon to
|
||
<taxonomicName id="4C7F4D1FFFE0020CFC93F8AEFB8EF858" authority="Mahony & Kamei & Teeling, 2018" authorityName="Mahony & Kamei & Teeling" authorityYear="2018" box="[887,1104,1916,1941]" class="Amphibia" family="Megophryidae" genus="Megophrys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="62" pageNumber="63" phylum="Chordata" rank="genus" status="sp. nov.">
|
||
<emphasis id="B90BEA8EFFE0020CFC93F8AEFB8EF858" box="[887,1104,1916,1941]" italics="true" pageId="62" pageNumber="63">Megophrys periosa</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A23857F5FFE0020CFBB3F8AEFB6EF859" box="[1111,1200,1916,1941]" pageId="62" pageNumber="63" rank="species">
|
||
<emphasis id="B90BEA8EFFE0020CFBB3F8AEFB6EF859" bold="true" box="[1111,1200,1916,1941]" pageId="62" pageNumber="63">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
(
|
||
<bibRefCitation id="EFEE4B6DFFE0020CFB5BF8AEFA42F858" author="Mahony, S. & Foley, N. C. & Biju, S. D. & Teeling, E. C." box="[1215,1436,1916,1941]" pageId="62" pageNumber="63" pagination="744 - 771" refId="ref55439" refString="Mahony, S., Foley, N. C., Biju, S. D. & Teeling, E. C. (2017) Evolutionary history of the Asian Horned Frogs (Megophryinae): integrative approaches to timetree dating in the absence of a fossil record. Molecular Biology and Evolution, 34 (3), 744 - 771 + S. I. 33 pp. https: // doi. org / 10.1093 / molbev / msw 267" type="book chapter" year="2017">
|
||
Mahony
|
||
<emphasis id="B90BEA8EFFE0020CFAC1F8AFFA83F859" box="[1317,1373,1916,1941]" italics="true" pageId="62" pageNumber="63">et al.</emphasis>
|
||
2017
|
||
</bibRefCitation>
|
||
[as
|
||
<taxonomicName id="4C7F4D1FFFE0020CFF25F873FE8DF874" authorityName="Mahony & Kamei & Teeling" authorityYear="2018" box="[193,339,1952,1977]" class="Amphibia" family="Megophryidae" genus="Megalophrys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="62" pageNumber="63" phylum="Chordata" rank="species" species="major">
|
||
<emphasis id="B90BEA8EFFE0020CFF25F873FF3EF874" box="[193,224,1953,1976]" italics="true" pageId="62" pageNumber="63">M.</emphasis>
|
||
cf.
|
||
<emphasis id="B90BEA8EFFE0020CFEF4F873FE8DF874" box="[272,339,1953,1976]" italics="true" pageId="62" pageNumber="63">major</emphasis>
|
||
</taxonomicName>
|
||
[6]];
|
||
<figureCitation id="13442A19FFE0020CFE7DF872FDD8F874" box="[409,518,1952,1977]" captionStart="FIGURE 2" captionStartId="11.[151,250,1148,1170]" captionTargetBox="[241,1332,252,1087]" captionTargetId="figure@11.[151,1436,193,1127]" captionTargetPageId="11" captionText="FIGURE 2. BI concatenated gene tree based on the 47 taxa concatenated mt+nuDNA alignment (Dataset C), with ML node values provided. Posterior probability/bootstrap values>95% in both analyses are represented by a black spot on the node, values <95% in one or both analyses are provided next to relevant nodes, where * is>95%, and – is given for nodes that were not resolved on the ML tree. Taxon names in pink represent the eastern clade, blue represents the western clade (southern subclade), and green represents the western clade (northern subclade). Revised names for taxa based on the taxonomic review are provided in parentheses." httpUri="https://zenodo.org/record/2610206/files/figure.png" pageId="62" pageNumber="63">Figures 2</figureCitation>
|
||
,
|
||
<figureCitation id="13442A19FFE0020CFDF1F872FDFDF874" box="[533,547,1952,1976]" captionStart="FIGURE 4" captionStartId="13.[151,250,1115,1137]" captionTargetBox="[155,1430,193,1084]" captionTargetId="figure@13.[150,1436,193,1094]" captionTargetPageId="13" captionText="FIGURE 4. RAxML tree for the combined mt+nuDNA dataset expanded to include the Chen et al. (2017) sequences (Dataset J). Bootstrap support values are provided adjacent to the associated nodes. Refer to Appendix I, Table 2 for revised names for taxa based on the taxonomic review." httpUri="https://zenodo.org/record/2610210/files/figure.png" pageId="62" pageNumber="63">4</figureCitation>
|
||
&
|
||
<figureCitation id="13442A19FFE0020CFDA9F872FD83F875" box="[589,605,1952,1977]" captionStart="FIGURE 5" captionStartId="13.[151,250,1843,1865]" captionTargetBox="[162,1426,1248,1806]" captionTargetId="figure@13.[151,1436,1248,1822]" captionTargetPageId="13" captionText="FIGURE 5. DensiTree cloudogram representations of *BEAST analyses, removing the first 10% of trees as burn-in, produced from the four nuDNA gene alignments of Dataset E: A. original unphased; B. phased. Dark blue branches represent the most frequently occurring topology, light blue branches represent the consensus trees with intensity proportional to the frequency of the topology in the dataset, and green areas represent all topologies in the dataset. Refer to Appendix I, Table 2 for revised names for taxa based on the taxonomic review." httpUri="https://zenodo.org/record/2610212/files/figure.png" pageId="62" pageNumber="63">5</figureCitation>
|
||
). Regardless of BPP results indicating that this lineage may represent a distinct species-level taxon (Appendix I, Table 5), we recommend that comparable series of adult specimens be studied to further determine the taxonomic status of this lineage. For now, it may provisionally be regarded as conspecific with
|
||
<taxonomicName id="4C7F4D1FFFE1020DFEB2FF6EFDF1FF18" authority="Mahony & Kamei & Teeling, 2018" authorityName="Mahony & Kamei & Teeling" authorityYear="2018" box="[342,559,188,213]" class="Amphibia" family="Megophryidae" genus="Megophrys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="63" pageNumber="64" phylum="Chordata" rank="genus" status="sp. nov.">
|
||
<emphasis id="B90BEA8EFFE1020DFEB2FF6EFDF1FF18" box="[342,559,188,213]" italics="true" pageId="63" pageNumber="64">Megophrys periosa</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A23857F5FFE1020DFDD2FF6EFD51FF19" box="[566,655,188,213]" pageId="63" pageNumber="64" rank="species">
|
||
<emphasis id="B90BEA8EFFE1020DFDD2FF6EFD51FF19" bold="true" box="[566,655,188,213]" pageId="63" pageNumber="64">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |