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<document id="E5F1595D5D189FBDD77634D943B8648F" ID="10.11646/zootaxa.4258.1.1SLASH10897" ID-CLB-Dataset="33167" ID-DOI="10.11646/zootaxa.4258.1.1" ID-GBIF-Dataset="fdbe32e3-f5b6-4537-b870-6d295865da73" ID-ISSN="1175-5326" ID-Zenodo-Dep="569114" ID-ZooBank="B1F45C56-A43A-4A8C-9190-2B861A37D33C" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1493194184337" checkinUser="plazi" docAuthor="Judson, Mark L. I." docDate="2017" docId="03D65C0B65773341FF333BB26689F90C" docLanguage="en" docName="zootaxa.4258.1.1SLASH10897.pdf" docOrigin="Zootaxa 4258 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Cybella deharvengi Judson, 2017, n. sp." docType="treatment" docUuid="172A8ACA-138C-44CC-9F7E-5CF5EB6D9398" docUuidSource="ZooBank" docVersion="8" lastPageNumber="14" masterDocId="FFEF24736573334CFFA43E146234FFEF" masterDocTitle="A new subfamily of Feaellidae (Arachnida, Chelonethi, Feaelloidea) from Southeast Asia" masterLastPageNumber="33" masterPageNumber="1" pageNumber="5" updateTime="1698439057093" updateUser="ExternalLinkService">
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<mods:title id="38C7F43BE6603CE1FF35510E2035C775">A new subfamily of Feaellidae (Arachnida, Chelonethi, Feaelloidea) from Southeast Asia</mods:title>
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<mods:date id="FCC84F8D8F5A34F3CB44D93C5DA6035F">2017</mods:date>
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<treatment id="03D65C0B65773341FF333BB26689F90C" ID-DOI="http://doi.org/10.5281/zenodo.6023803" ID-GBIF-Taxon="128699666" ID-Zenodo-Dep="6023803" LSID="urn:lsid:zoobank.org:act:172A8ACA-138C-44CC-9F7E-5CF5EB6D9398" httpUri="http://treatment.plazi.org/id/03D65C0B65773341FF333BB26689F90C" lastPageId="13" lastPageNumber="14" pageId="4" pageNumber="5">
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<paragraph id="8BC0ED1D65773348FF333BB263FEFA2F" blockId="4.[151,458,1446,1506]" box="[151,458,1446,1472]" pageId="4" pageNumber="5">
<heading id="D0885A7165773348FF333BB263FEFA2F" bold="true" box="[151,458,1446,1472]" fontSize="11" level="1" pageId="4" pageNumber="5" reason="1">
<emphasis id="B90B310F65773348FF333BB263FEFA2F" bold="true" box="[151,458,1446,1472]" pageId="4" pageNumber="5">
<taxonomicName id="4C7F969E65773348FF333BB263B4FA2F" ID-CoL="32RLJ" box="[151,384,1446,1472]" class="Arachnida" family="Feaellidae" genus="Cybella" kingdom="Animalia" order="Pseudoscorpiones" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="deharvengi" status="sp. nov.">
<emphasis id="B90B310F65773348FF333BB263B4FA2F" bold="true" box="[151,384,1446,1472]" italics="true" pageId="4" pageNumber="5">Cybella deharvengi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2388C7465773348FE2C3BB263FEFA2F" box="[392,458,1446,1472]" pageId="4" pageNumber="5" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BC0ED1D65773348FF333BDD6332FA0D" blockId="4.[151,458,1446,1506]" box="[151,262,1481,1506]" pageId="4" pageNumber="5">
<figureCitation id="1344F19865773348FF333BDD6332FA0D" box="[151,262,1481,1506]" captionStart-0="FIGURES 1 2" captionStart-1="FIGURES 3 7" captionStart-2="FIGURES 8 13" captionStart-3="FIGURES 14 16" captionStart-4="FIGURES 17 22" captionStart-5="FIGURES 23 34" captionStart-6="FIGURES 35 43" captionStart-7="FIGURES 44 47" captionStartId-0="5.[151,264,1190,1213]" captionStartId-1="6.[151,264,1896,1919]" captionStartId-2="7.[151,264,1848,1871]" captionStartId-3="8.[151,264,1719,1742]" captionStartId-4="9.[151,264,1855,1878]" captionStartId-5="10.[151,264,1752,1775]" captionStartId-6="11.[151,264,1850,1873]" captionStartId-7="12.[151,264,844,867]" captionTargetBox-0="[151,1436,519,1166]" captionTargetBox-1="[233,1351,598,1844]" captionTargetBox-2="[236,1334,199,1833]" captionTargetBox-3="[151,1428,202,1688]" captionTargetBox-4="[166,1401,441,1799]" captionTargetBox-5="[232,1344,207,1715]" captionTargetBox-6="[159,1426,202,1817]" captionTargetBox-7="[156,1429,202,816]" captionTargetId-0="figure@5.[151,1436,519,1166]" captionTargetId-1="figure@6.[226,1360,589,1876]" captionTargetId-2="figure@7.[230,1346,193,1838]" captionTargetId-3="figure@8.[151,1436,193,1699]" captionTargetId-4="figure@9.[151,1436,423,1846]" captionTargetId-5="figure@10.[224,1350,193,1724]" captionTargetId-6="figure@11.[151,1436,193,1830]" captionTargetId-7="figure@12.[151,1436,193,823]" captionTargetPageId-0="5" captionTargetPageId-1="6" captionTargetPageId-2="7" captionTargetPageId-3="8" captionTargetPageId-4="9" captionTargetPageId-5="10" captionTargetPageId-6="11" captionTargetPageId-7="12" captionText-0="FIGURES 1 2. Cybella deharvengi n. sp. 1, area of cavity wall on which the types were found (photograph L. Deharveng); 2, habitus of male, dorsal view." captionText-1="FIGURES 3 7. Cybella deharvengi n. sp., male. 3, pleurum at level of segments VIII X, ventral view; 4, carapace and anterior tergites, dorsal view; 5, carapace, lateral view; 6, opisthosoma, anterior view, prosoma removed; 7, lateral tergal plate of segment I, seen in same orientation as Fig. 6. Abbreviations: den, denticulate ornamentation; gp, gland pore; ly, lyrifissure; s, setae; T II, tergite II. Scale lines 0.2 mm (Figs 1 6), 0.05 mm (Fig. 7)." captionText-2="FIGURES 8 13. Cybella deharvengi n. sp., male. 8, coxae, ventral view; 9, left palp coxa, lateral view; 10, median maxillary lyrifissure; 11, right coxae II IV, dorsal (internal) view; 12, median part of coxae I, showing pit and coxal spines; 13, manducatory process of right palp coxa, ventral view. Abbreviations: cliv, clivus; fbII, border of internal foramen of coxa II, fbIII, border of internal foramen of coxa III, fbIƑ, border of internal foramen of coxa IV, k, condyle; pcs, primary coxal spine; pmIII, posterior margin of coxa III. Scale lines: 0.2 mm (Fig. 9) 0.1 mm (Figs 11 12), 0.05 mm (Figs 10, 13), 0.03 mm (Fig. 10)." captionText-3="FIGURES 14 16. Cybella deharvengi n. sp., male. 14, genitalia, ventral view; 15, posterior genital plate, ventral view 16 right side of opisthosomal segments II V, ventral view. Abbreviations: apl, atrial plate; da, dorsal apodeme; dpdg, ducts of posterior dorsal gland; ejca, ejaculatory canal atrium; gp, gland pore; inv, invagination at side of atrial plate; lgs, lateral genital sac; ly, lyrifissure; mgs, median genital sac; pap, papilla on atrial wall; sp, spiracle; sps, spatulate seta; sr, sclerotized ring. Scale lines: 0.1 mm (Figs 14, 16), 0.05 mm (Fig. 15)." captionText-4="FIGURES 17 22. Cybella deharvengi n. sp., male, left chelicera. 17, whole chelicera, antiaxial view; 18, tip of movable finger, antiaxial view; 19, distal part of chelicera, ventral view; 20, spinneret, ventral view; 21, distal part of chelicera (movable finger omitted), paraxial view; 22, chelicera in posterior view (dorsal at top, paraxial side to left). Abbreviations: ds, ducts of serrula interior; f, foramen; gs, galeal seta; ly, lyrifissure; pr, distal process of serrula exterior; r, rallum; s. ext, serrula exterior; s. int., serrula interior. Other abbreviations (b, es, gs, is, ls, sb) are designations of setae. Scale lines 0.05 mm (Figs 18 20 share same scale line)." captionText-5="FIGURES 23 34. Cybella deharvengi n. sp., male right palp. 23, chela, dorsal view; 24, palp minus chela, dorsal view (reticulate ornamentation drawn in part on femur); 25, tip of fixed chelal finger, dorsal view; 26, basal tuberculate tooth of fixed finger, dorsal view (relative to chela); 27, tip of movable chelal finger, paraxial view; 28, basal tuberculate tooth of movable chelal finger, ventral view (relative to chela), distal end to left; 29, chela, paraxial view (setae omitted); 30, chela, antiaxial view; 31, coupled sensilla; 32, distal sensillum; 33, distal end of movable finger, ventral view; 34, grouped chemosensory setae, antiaxial view. Abbreviations: af, apical sensillum of fixed finger; am 1, apical sensilla of movable finger; dx, duplex trichobothria; ld, probable lamina defensor; ly, lyrifissure; pc, coupled sensilla. Other abbreviations (eb, esb, est, et, ib, isb, ist, it, b, sb, st, t) are standard designations of trichobothria. Scale lines: 0.2 mm (Figs 23 24, 29), 0.1 mm (Fig. 30), 0.05 mm (Figs 25 28, 33 34), 0.01 mm (Figs 31 32)." captionText-6="FIGURES 35 43. Cybella deharvengi n. sp., male right legs. 35, leg I, antiaxial view; 36, trochanter and femur of leg I, dorsal view; 37, trochanter and femur of leg I, paraxial view; 38, leg IV, antiaxial view; 39, patella of leg I, dorsal view; 40, patella of leg IV, dorsal view; 41, leg III, antiaxial view; 42, arolium of leg III, ventral; 43 apotele of leg I, antiaxial view. Abbreviations: ly, lyrifissure;. Scale lines 0.2 mm (Figs 35 41), 0.05 (Figs 42 43)." captionText-7="FIGURES 44 47. Cybella deharvengi n. sp., paratype tritonymph. 44, chelicera, antiaxial view; 45, tip of movable cheliceral finger, antiaxial view; 46, right chela, antiaxial view; 47 grouped chemosensory setae, antiaxial view. Abbreviations: am 1, apical sensillum of movable finger; pr, distal process of serrula exterior; pc, coupled sensilla. Scale lines 0.1 mm (Figs 44, 46), 0.05 mm (Fig. 45), 0.02 mm (Fig. 47)." httpUri-0="https://zenodo.org/record/569115/files/figure.png" httpUri-1="https://zenodo.org/record/569116/files/figure.png" httpUri-2="https://zenodo.org/record/569117/files/figure.png" httpUri-3="https://zenodo.org/record/569118/files/figure.png" httpUri-4="https://zenodo.org/record/569119/files/figure.png" httpUri-5="https://zenodo.org/record/569120/files/figure.png" httpUri-6="https://zenodo.org/record/569121/files/figure.png" httpUri-7="https://zenodo.org/record/569122/files/figure.png" pageId="4" pageNumber="5">Figs 247</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C365BE9665773348FF33380560B9F9C5" box="[151,653,1553,1578]" pageId="4" pageNumber="5" type="diagnosis">
<paragraph id="8BC0ED1D65773348FF33380560B9F9C5" blockId="4.[151,1436,1553,2010]" box="[151,653,1553,1578]" pageId="4" pageNumber="5">
<emphasis id="B90B310F65773348FF3338056325F9C5" bold="true" box="[151,273,1553,1578]" pageId="4" pageNumber="5">Diagnosis.</emphasis>
See under
<taxonomicName id="4C7F969E65773348FE2A3805607CF9C5" box="[398,584,1553,1578]" class="Arachnida" family="Feaellidae" genus="Cybella" kingdom="Animalia" order="Pseudoscorpiones" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="bedosae" status="sp. nov.">
<emphasis id="B90B310F65773348FE2A3805607CF9C5" box="[398,584,1553,1578]" italics="true" pageId="4" pageNumber="5">Cybella bedosae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2388C7465773348FDEB380560B9F9C5" box="[591,653,1553,1578]" pageId="4" pageNumber="5" rank="species">
<emphasis id="B90B310F65773348FDEB380560B9F9C5" bold="true" box="[591,653,1553,1578]" pageId="4" pageNumber="5">n. sp.</emphasis>
</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="C365BE9665773348FF6338216372F979" pageId="4" pageNumber="5" type="etymology">
<paragraph id="8BC0ED1D65773348FF6338216372F979" blockId="4.[151,1436,1553,2010]" pageId="4" pageNumber="5">
<emphasis id="B90B310F65773348FF633821637AF9A1" bold="true" box="[199,334,1589,1614]" pageId="4" pageNumber="5">Etymology.</emphasis>
This species is named after Louis Deharveng (MNHN), who discovered the cavity in which the
<typeStatus id="54C453BF65773348FF33384E62E6F99D" box="[151,210,1626,1650]" pageId="4" pageNumber="5">types</typeStatus>
were found and who has made many important contributions to our knowledge of soil and cave faunas in Southeast Asia.
</paragraph>
</subSubSection>
<subSubSection id="C365BE9665773348FF6338B5631BF835" pageId="4" pageNumber="5" type="materials_examined">
<paragraph id="8BC0ED1D65773348FF6338B56331F8A5" blockId="4.[151,1436,1553,2010]" pageId="4" pageNumber="5">
<materialsCitation id="3B17E74065773348FF6338B5608BF8ED" ID-GBIF-Occurrence="1493295235" collectingDate="2008-06-02" collectionCode="MNHN" collectorName="M. Judson" country="Vietnam" county="Feaellidae'" elevation="32" location="Kien Luong" municipality="Mo So Lon" pageId="4" pageNumber="5" specimenCount="1" specimenCount-male="1" stateProvince="Kien Giang" typeStatus="holotype">
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<typeStatus id="54C453BF65773348FF6338B56336F955" box="[199,258,1697,1722]" pageId="4" pageNumber="5">Type</typeStatus>
material.
</emphasis>
<typeStatus id="54C453BF65773348FEDE38B563D6F955" box="[378,482,1697,1722]" pageId="4" pageNumber="5" type="holotype">Holotype</typeStatus>
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,
<collectingCountry id="F368AD8D65773348FDAF38B56059F955" box="[523,621,1697,1722]" name="Vietnam" pageId="4" pageNumber="5">Vietnam</collectingCountry>
,
<collectingRegion id="49BB23FF65773348FDD338B560C8F955" box="[631,764,1697,1722]" country="Vietnam" name="Kien Giang" pageId="4" pageNumber="5">Kien Giang</collectingRegion>
,
<location id="8EA0BBC665773348FCA238B561BBF955" LSID="urn:lsid:plazi:treatment:03D65C0B65773341FF333BB26689F90C:8EA0BBC665773348FCA238B561BBF955" box="[774,911,1697,1722]" country="Vietnam" county="Feaellidae'" municipality="Mo So Lon" name="Kien Luong" pageId="4" pageNumber="5" stateProvince="Kien Giang">Kien Luong</location>
,
<location id="8EA0BBC665773348FC3E38B6662AF955" LSID="urn:lsid:plazi:treatment:03D65C0B65773341FF333BB26689F90C:8EA0BBC665773348FC3E38B6662AF955" box="[922,1054,1697,1722]" country="Vietnam" county="Feaellidae'" municipality="Mo So Lon" name="Hon Chong" pageId="4" pageNumber="5" stateProvince="Kien Giang">Hon Chong</location>
,
<location id="8EA0BBC665773348FB8D38B56698F955" LSID="urn:lsid:plazi:treatment:03D65C0B65773341FF333BB26689F90C:8EA0BBC665773348FB8D38B56698F955" box="[1065,1196,1697,1722]" country="Vietnam" county="Feaellidae'" municipality="Mo So Lon" name="Nui Bai Voi" pageId="4" pageNumber="5" stateProvince="Kien Giang">Nui Bai Voi</location>
(
<collectingMunicipality id="6BA4776765773348FB1F38B6670BF955" box="[1211,1343,1698,1722]" pageId="4" pageNumber="5">Mo So Lon</collectingMunicipality>
), grotte des
<collectingCounty id="62A1959165773348FF6238D16371F931" box="[198,325,1733,1758]" pageId="4" pageNumber="5">Feaellidae</collectingCounty>
, 1013ʹ25.81ʺN 10436ʹ57.35ʺE,
<elevation id="00520A2E65773348FD1438D26118F931" box="[688,812,1733,1758]" metricMagnitude="1" metricUnit="m" metricValue="3.2" pageId="4" pageNumber="5" unit="m" value="32.0">
<quantity id="4C8740F865773348FD1438D260D9F932" box="[688,749,1734,1758]" metricMagnitude="1" metricUnit="m" metricValue="3.2" pageId="4" pageNumber="5" unit="m" value="32.0">32 m</quantity>
a.s.l.
</elevation>
, on wall of limestone cavity in near-total obscurity,
<collectingDate id="EF85323565773348FA2A38D26324F8ED" pageId="4" pageNumber="5" value="2008-06-02">
<date id="FFC1CBDD65773348FA2A38D26324F8ED" pageId="4" pageNumber="5" value="2008-06-02">2 June 2008</date>
</collectingDate>
, leg.
<collectorName id="268A88CB65773348FEF038FE63F9F8ED" box="[340,461,1769,1794]" pageId="4" pageNumber="5">M. Judson</collectorName>
(
<collectionCode id="ED6E75D865773348FE7B38FE6003F8ED" box="[479,567,1770,1794]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="4" pageNumber="5">MNHN</collectionCode>
Ps-704.43)
</materialsCitation>
.
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<typeStatus id="54C453BF65773348FD6A38FE6174F8ED" box="[718,832,1770,1794]" pageId="4" pageNumber="5" type="paratype">Paratypes</typeStatus>
:
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, 1 tritonymph, same details as
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(
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,
<collectionCode id="ED6E75D865773348FF33391A62D8F8CA" box="[151,236,1806,1829]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="4" pageNumber="5">MNHN</collectionCode>
Ps-704.4445, and tritonymph,
<collectionCode id="ED6E75D865773348FDFF391A6084F8CA" box="[603,688,1806,1829]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="4" pageNumber="5">MNHN</collectionCode>
Ps-704.46;
<specimenCount id="9D79269465773348FC9E391A6151F8CA" box="[826,869,1806,1830]" pageId="4" pageNumber="5" type="male">1 ♂</specimenCount>
deposited in
<location id="8EA0BBC665773348FC5B3919666FF8C9" LSID="urn:lsid:plazi:treatment:03D65C0B65773341FF333BB26689F90C:8EA0BBC665773348FC5B3919666FF8C9" box="[1023,1115,1805,1830]" country="Vietnam" municipality="Tropical Biology" name="Institute" pageId="4" pageNumber="5" stateProvince="Hanoi">Institute</location>
of
<collectingMunicipality id="6BA4776765773348FB2739196771F8C9" box="[1155,1349,1805,1830]" pageId="4" pageNumber="5">Tropical Biology</collectingMunicipality>
,
<collectingRegion id="49BB23FF65773348FAF5391967ACF8C9" box="[1361,1432,1805,1830]" country="Vietnam" name="Ha Noi" pageId="4" pageNumber="5">Hanoi</collectingRegion>
,
<collectingCountry id="F368AD8D65773348FF33392562CEF8A5" box="[151,250,1841,1866]" name="Vietnam" pageId="4" pageNumber="5">Vietnam</collectingCountry>
)
</materialsCitation>
.
</paragraph>
<paragraph id="8BC0ED1D65773348FF633941631BF835" blockId="4.[151,1436,1553,2010]" pageId="4" pageNumber="5">
<materialsCitation id="3B17E74065773348FF633941631BF835" ID-GBIF-Occurrence="1493295183" collectingDate="2003-01-16" collectionCode="MNHN" collectorName="Grottehopital de Mo So &amp; Berlese &amp; Le Cong Man" country="Vietnam" county="Non-type" elevation="29" location="Hon Chong" municipality="Kien Luong" pageId="4" pageNumber="5" specimenCount="1" stateProvince="Kien Giang">
<emphasis id="B90B310F65773348FF633941639DF881" bold="true" box="[199,425,1877,1902]" pageId="4" pageNumber="5">
<collectingCounty id="62A1959165773348FF6339416301F881" box="[199,309,1877,1902]" pageId="4" pageNumber="5">Non-type</collectingCounty>
material.
</emphasis>
1 tritonymph,
<collectingCountry id="F368AD8D65773348FDF739416083F881" box="[595,695,1877,1902]" name="Vietnam" pageId="4" pageNumber="5">Vietnam</collectingCountry>
:
<collectingRegion id="49BB23FF65773348FD6639416173F881" box="[706,839,1877,1902]" country="Vietnam" name="Kien Giang" pageId="4" pageNumber="5">Kien Giang</collectingRegion>
,
<collectingMunicipality id="6BA4776765773348FCF0394161EAF881" box="[852,990,1877,1902]" pageId="4" pageNumber="5">Kien Luong</collectingMunicipality>
,
<location id="8EA0BBC665773348FC4E39426644F881" LSID="urn:lsid:plazi:treatment:03D65C0B65773341FF333BB26689F90C:8EA0BBC665773348FC4E39426644F881" box="[1002,1136,1877,1902]" country="Vietnam" county="Non-type" municipality="Kien Luong" name="Hon Chong" pageId="4" pageNumber="5" stateProvince="Kien Giang">Hon Chong</location>
,
<location id="8EA0BBC665773348FBD839416733F881" LSID="urn:lsid:plazi:treatment:03D65C0B65773341FF333BB26689F90C:8EA0BBC665773348FBD839416733F881" box="[1148,1287,1877,1902]" country="Vietnam" county="Non-type" municipality="Kien Luong" name="Nui Bai Voi" pageId="4" pageNumber="5" stateProvince="Kien Giang">Nui Bai Voi</location>
, near
<collectorName id="268A88CB65773348FAEF3942636AF87D" pageId="4" pageNumber="5">
<location id="8EA0BBC665773348FAEF3942636AF87D" LSID="urn:lsid:plazi:treatment:03D65C0B65773341FF333BB26689F90C:8EA0BBC665773348FAEF3942636AF87D" country="Vietnam" county="Non-type" municipality="Kien Luong" name="Grottehopital de Mo So" pageId="4" pageNumber="5" stateProvince="Kien Giang">Grottehôpital de Mo So</location>
</collectorName>
[=Hang Moi Chau or Hang Mo So], 1013ʹ31.6ʺN 10436ʹ59.2ʺE,
<elevation id="00520A2E65773348FBF4396E66FDF87D" box="[1104,1225,1913,1938]" metricMagnitude="1" metricUnit="m" metricValue="2.9" pageId="4" pageNumber="5" unit="m" value="29.0">
<quantity id="4C8740F865773348FBF4396E66BFF87E" box="[1104,1163,1914,1938]" metricMagnitude="1" metricUnit="m" metricValue="2.9" pageId="4" pageNumber="5" unit="m" value="29.0">29 m</quantity>
a.s.l.
</elevation>
,
<collectingDate id="EF85323565773348FB7C396D67ACF87D" box="[1240,1432,1913,1938]" pageId="4" pageNumber="5" value="2003-01-16">
<date id="FFC1CBDD65773348FB7C396D67ACF87D" box="[1240,1432,1913,1938]" pageId="4" pageNumber="5" value="2003-01-16">16 January 2003</date>
</collectingDate>
, litter,
<collectorName id="268A88CB65773348FF7E3989631AF859" box="[218,302,1949,1974]" pageId="4" pageNumber="5">Berlese</collectorName>
extraction, leg.
<collectorName id="268A88CB65773348FE43398A60BCF85A" box="[487,648,1950,1974]" pageId="4" pageNumber="5">Le Cong Man</collectorName>
, collectors code VIET-ManKG23 (
<collectionCode id="ED6E75D865773348FBB9398A6641F859" box="[1053,1141,1950,1974]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="4" pageNumber="5">MNHN</collectionCode>
Ps-704.47). Tarsus of left leg I missing.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C365BE9665763341FF633E8360AFFB07" lastPageId="13" lastPageNumber="14" pageId="5" pageNumber="6" type="description">
<paragraph id="8BC0ED1D65763349FF633E83674FFE3F" blockId="5.[151,1436,151,464]" pageId="5" pageNumber="6">
<emphasis id="B90B310F65763349FF633E8363F7FF5F" bold="true" box="[199,451,151,176]" pageId="5" pageNumber="6">Description of male.</emphasis>
Sclerotized parts amber-brown (
<figureCitation id="1344F19865763349FCEC3E8361A7FF5F" box="[840,915,151,176]" captionStart="FIGURES 1 2" captionStartId="5.[151,264,1190,1213]" captionTargetBox="[151,1436,519,1166]" captionTargetId="figure@5.[151,1436,519,1166]" captionTargetPageId="5" captionText="FIGURES 1 2. Cybella deharvengi n. sp. 1, area of cavity wall on which the types were found (photograph L. Deharveng); 2, habitus of male, dorsal view." httpUri="https://zenodo.org/record/569115/files/figure.png" pageId="5" pageNumber="6">Fig. 2</figureCitation>
); body and appendages covered with finely granular cerotegument, apart from non-reticulate areas, such as the fingers of chelicerae and chelae. Vestitural setae very small and simple, only chelal fingers, chelicera and manducatory process with larger setae. Carapace (
<figureCitation id="1344F19865763349FAF43ECB67A0FF17" box="[1360,1428,223,248]" captionStart="FIGURES 3 7" captionStartId="6.[151,264,1896,1919]" captionTargetBox="[233,1351,598,1844]" captionTargetId="figure@6.[226,1360,589,1876]" captionTargetPageId="6" captionText="FIGURES 3 7. Cybella deharvengi n. sp., male. 3, pleurum at level of segments VIII X, ventral view; 4, carapace and anterior tergites, dorsal view; 5, carapace, lateral view; 6, opisthosoma, anterior view, prosoma removed; 7, lateral tergal plate of segment I, seen in same orientation as Fig. 6. Abbreviations: den, denticulate ornamentation; gp, gland pore; ly, lyrifissure; s, setae; T II, tergite II. Scale lines 0.2 mm (Figs 1 6), 0.05 mm (Fig. 7)." httpUri="https://zenodo.org/record/569116/files/figure.png" pageId="5" pageNumber="6">Fig. 4</figureCitation>
) longer than broad, distinctly broadened proximally. Anterolateral corners truncate, not projecting anteriorly, median projections small, directed slightly towards midline, two raised mounds between eyes and an unpaired mound in middle of mesozone, behind which is a moderate ridge that is only weakly produced laterally. Two pairs of large eyes, with reflective tapeta. One pair of lyrifissures near eyes, one lateral pair behind posterior eyes, and 2 5 posteriorly; one specimen with a single lyrifissure on mesozone (total number of lyrifissures 69). Carapace with about 37 setae (N = 1), of which 6 on posterior margin and 4 laterally on each side, below level of eyes (
<figureCitation id="1344F19865763349FA8D3FA36759FE3F" box="[1321,1389,439,464]" captionStart="FIGURES 3 7" captionStartId="6.[151,264,1896,1919]" captionTargetBox="[233,1351,598,1844]" captionTargetId="figure@6.[226,1360,589,1876]" captionTargetPageId="6" captionText="FIGURES 3 7. Cybella deharvengi n. sp., male. 3, pleurum at level of segments VIII X, ventral view; 4, carapace and anterior tergites, dorsal view; 5, carapace, lateral view; 6, opisthosoma, anterior view, prosoma removed; 7, lateral tergal plate of segment I, seen in same orientation as Fig. 6. Abbreviations: den, denticulate ornamentation; gp, gland pore; ly, lyrifissure; s, setae; T II, tergite II. Scale lines 0.2 mm (Figs 1 6), 0.05 mm (Fig. 7)." httpUri="https://zenodo.org/record/569116/files/figure.png" pageId="5" pageNumber="6">Fig. 5</figureCitation>
).
</paragraph>
<caption id="DF00BD9565763349FF333AB2638FFB34" httpUri="https://zenodo.org/record/569115/files/figure.png" pageId="5" pageNumber="6" targetBox="[151,1436,519,1166]" targetPageId="5">
<paragraph id="8BC0ED1D65763349FF333AB2638FFB34" blockId="5.[151,1435,1190,1243]" pageId="5" pageNumber="6">
<emphasis id="B90B310F65763349FF333AB2630EFB53" bold="true" box="[151,314,1190,1213]" pageId="5" pageNumber="6">FIGURES 12.</emphasis>
<taxonomicName id="4C7F969E65763349FEE43AB36033FB52" box="[320,519,1191,1213]" class="Arachnida" family="Feaellidae" genus="Cybella" kingdom="Animalia" order="Pseudoscorpiones" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="deharvengi" status="sp. nov.">
<emphasis id="B90B310F65763349FEE43AB36033FB52" box="[320,519,1191,1213]" italics="true" pageId="5" pageNumber="6">Cybella deharvengi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2388C7465763349FDA93AB36073FB53" box="[525,583,1191,1212]" pageId="5" pageNumber="6" rank="species">
<emphasis id="B90B310F65763349FDA93AB36073FB53" bold="true" box="[525,583,1191,1212]" pageId="5" pageNumber="6">n. sp.</emphasis>
</taxonomicNameLabel>
1, area of cavity wall on which the types were found (photograph L. Deharveng); 2, habitus of male, dorsal view.
</paragraph>
</caption>
<paragraph id="8BC0ED1D65763349FF633B176009F9F8" blockId="5.[151,1437,1283,2028]" pageId="5" pageNumber="6">
Tergite I (
<figureCitation id="1344F19865763349FE923B17634EFAF4" box="[310,378,1283,1308]" captionStart="FIGURES 3 7" captionStartId="6.[151,264,1896,1919]" captionTargetBox="[233,1351,598,1844]" captionTargetId="figure@6.[226,1360,589,1876]" captionTargetPageId="6" captionText="FIGURES 3 7. Cybella deharvengi n. sp., male. 3, pleurum at level of segments VIII X, ventral view; 4, carapace and anterior tergites, dorsal view; 5, carapace, lateral view; 6, opisthosoma, anterior view, prosoma removed; 7, lateral tergal plate of segment I, seen in same orientation as Fig. 6. Abbreviations: den, denticulate ornamentation; gp, gland pore; ly, lyrifissure; s, setae; T II, tergite II. Scale lines 0.2 mm (Figs 1 6), 0.05 mm (Fig. 7)." httpUri="https://zenodo.org/record/569116/files/figure.png" pageId="5" pageNumber="6">Fig. 4</figureCitation>
) distinctly shelved to allow posterior margin of carapace to pass over distal portion; posterior margin desclerotized; undivided medially, but flanked on each side by a small, sclerotized plate, each bearing 1 or 2 setae (no seta on main part of tergite I); lyrifissures absent. Membrane between tergites I and II extensive and rounded over posterior part of I. Tergites IIIX divided medially; setae per half-tergite 8: 1011:
<date id="FFC1CBDD65763349FB4E3B7B6704FA68" box="[1258,1328,1390,1415]" pageId="5" pageNumber="6" value="2015-03-01">1 315</date>
: 1315: 1415: 1416: 1316: 1214; tergite X undivided, with 21 setae. Segment XI with 1011 posterior setae plus 3 around anal mound (total 1314). Tergite XII with 2 setae. Tergite I without lyrifissures; half-tergite II with 1 anterolateral and 1 submedian lyrifissure; half-tergites IIIIX with 1 anterolateral and 34 posterior lyrifissures; tergite X with a total of 5 lyrifissures.
</paragraph>
<paragraph id="8BC0ED1D65763349FF63383761E8F948" blockId="5.[151,1437,1283,2028]" pageId="5" pageNumber="6">
Pleural membranes very broad, with two longitudinal folds (
<figureCitation id="1344F19865763349FCD7383761ECF9D3" box="[883,984,1571,1596]" captionStart="FIGURES 3 7" captionStartId="6.[151,264,1896,1919]" captionTargetBox="[233,1351,598,1844]" captionTargetId="figure@6.[226,1360,589,1876]" captionTargetPageId="6" captionText="FIGURES 3 7. Cybella deharvengi n. sp., male. 3, pleurum at level of segments VIII X, ventral view; 4, carapace and anterior tergites, dorsal view; 5, carapace, lateral view; 6, opisthosoma, anterior view, prosoma removed; 7, lateral tergal plate of segment I, seen in same orientation as Fig. 6. Abbreviations: den, denticulate ornamentation; gp, gland pore; ly, lyrifissure; s, setae; T II, tergite II. Scale lines 0.2 mm (Figs 1 6), 0.05 mm (Fig. 7)." httpUri="https://zenodo.org/record/569116/files/figure.png" pageId="5" pageNumber="6">Figs 3, 6</figureCitation>
) on each side; dorsal fold with setae on segments IIIXI and lyrifissures on VVII; ventral fold with setae on segments VIXI and lyrifissures on VIIIX; lyrifissures set in slight depressions in folds; simple gland pores opening ventrally on folds (
<figureCitation id="1344F19865763349FB0D387F66D9F96C" box="[1193,1261,1643,1668]" captionStart="FIGURES 3 7" captionStartId="6.[151,264,1896,1919]" captionTargetBox="[233,1351,598,1844]" captionTargetId="figure@6.[226,1360,589,1876]" captionTargetPageId="6" captionText="FIGURES 3 7. Cybella deharvengi n. sp., male. 3, pleurum at level of segments VIII X, ventral view; 4, carapace and anterior tergites, dorsal view; 5, carapace, lateral view; 6, opisthosoma, anterior view, prosoma removed; 7, lateral tergal plate of segment I, seen in same orientation as Fig. 6. Abbreviations: den, denticulate ornamentation; gp, gland pore; ly, lyrifissure; s, setae; T II, tergite II. Scale lines 0.2 mm (Figs 1 6), 0.05 mm (Fig. 7)." httpUri="https://zenodo.org/record/569116/files/figure.png" pageId="5" pageNumber="6">Fig. 3</figureCitation>
); pleural plates absent (only tiny areas of sclerotization around pores and areoles of setae).
</paragraph>
<paragraph id="8BC0ED1D65763349FF6338A766B3F84C" blockId="5.[151,1437,1283,2028]" pageId="5" pageNumber="6">
Palp coxa elongate, almost as long as combined length of leg coxae IIV (
<figureCitation id="1344F19865763349FBB638A76662F924" box="[1042,1110,1715,1740]" captionStart="FIGURES 8 13" captionStartId="7.[151,264,1848,1871]" captionTargetBox="[236,1334,199,1833]" captionTargetId="figure@7.[230,1346,193,1838]" captionTargetPageId="7" captionText="FIGURES 8 13. Cybella deharvengi n. sp., male. 8, coxae, ventral view; 9, left palp coxa, lateral view; 10, median maxillary lyrifissure; 11, right coxae II IV, dorsal (internal) view; 12, median part of coxae I, showing pit and coxal spines; 13, manducatory process of right palp coxa, ventral view. Abbreviations: cliv, clivus; fbII, border of internal foramen of coxa II, fbIII, border of internal foramen of coxa III, fbIƑ, border of internal foramen of coxa IV, k, condyle; pcs, primary coxal spine; pmIII, posterior margin of coxa III. Scale lines: 0.2 mm (Fig. 9) 0.1 mm (Figs 11 12), 0.05 mm (Figs 10, 13), 0.03 mm (Fig. 10)." httpUri="https://zenodo.org/record/569117/files/figure.png" pageId="5" pageNumber="6">Fig. 8</figureCitation>
). Manducatory process (
<figureCitation id="1344F19865763349FACB38A76280F900" captionStart="FIGURES 8 13" captionStartId="7.[151,264,1848,1871]" captionTargetBox="[236,1334,199,1833]" captionTargetId="figure@7.[230,1346,193,1838]" captionTargetPageId="7" captionText="FIGURES 8 13. Cybella deharvengi n. sp., male. 8, coxae, ventral view; 9, left palp coxa, lateral view; 10, median maxillary lyrifissure; 11, right coxae II IV, dorsal (internal) view; 12, median part of coxae I, showing pit and coxal spines; 13, manducatory process of right palp coxa, ventral view. Abbreviations: cliv, clivus; fbII, border of internal foramen of coxa II, fbIII, border of internal foramen of coxa III, fbIƑ, border of internal foramen of coxa IV, k, condyle; pcs, primary coxal spine; pmIII, posterior margin of coxa III. Scale lines: 0.2 mm (Fig. 9) 0.1 mm (Figs 11 12), 0.05 mm (Figs 10, 13), 0.03 mm (Fig. 10)." httpUri="https://zenodo.org/record/569117/files/figure.png" pageId="5" pageNumber="6">Fig. 13</figureCitation>
) with 1 small apical seta and a large subapical seta, both simple; another large seta is present at base of manducatory process. Clivus (
<figureCitation id="1344F19865763349FE5338EF606BF8FC" box="[503,607,1787,1812]" captionStart="FIGURES 8 13" captionStartId="7.[151,264,1848,1871]" captionTargetBox="[236,1334,199,1833]" captionTargetId="figure@7.[230,1346,193,1838]" captionTargetPageId="7" captionText="FIGURES 8 13. Cybella deharvengi n. sp., male. 8, coxae, ventral view; 9, left palp coxa, lateral view; 10, median maxillary lyrifissure; 11, right coxae II IV, dorsal (internal) view; 12, median part of coxae I, showing pit and coxal spines; 13, manducatory process of right palp coxa, ventral view. Abbreviations: cliv, clivus; fbII, border of internal foramen of coxa II, fbIII, border of internal foramen of coxa III, fbIƑ, border of internal foramen of coxa IV, k, condyle; pcs, primary coxal spine; pmIII, posterior margin of coxa III. Scale lines: 0.2 mm (Fig. 9) 0.1 mm (Figs 11 12), 0.05 mm (Figs 10, 13), 0.03 mm (Fig. 10)." httpUri="https://zenodo.org/record/569117/files/figure.png" pageId="5" pageNumber="6">Figs 89</figureCitation>
) well marked, but not projecting laterally. A single, normal lyrifissure behind foramen of palp coxa (
<figureCitation id="1344F19865763349FE4B390A6007F8D8" box="[495,563,1822,1847]" captionStart="FIGURES 8 13" captionStartId="7.[151,264,1848,1871]" captionTargetBox="[236,1334,199,1833]" captionTargetId="figure@7.[230,1346,193,1838]" captionTargetPageId="7" captionText="FIGURES 8 13. Cybella deharvengi n. sp., male. 8, coxae, ventral view; 9, left palp coxa, lateral view; 10, median maxillary lyrifissure; 11, right coxae II IV, dorsal (internal) view; 12, median part of coxae I, showing pit and coxal spines; 13, manducatory process of right palp coxa, ventral view. Abbreviations: cliv, clivus; fbII, border of internal foramen of coxa II, fbIII, border of internal foramen of coxa III, fbIƑ, border of internal foramen of coxa IV, k, condyle; pcs, primary coxal spine; pmIII, posterior margin of coxa III. Scale lines: 0.2 mm (Fig. 9) 0.1 mm (Figs 11 12), 0.05 mm (Figs 10, 13), 0.03 mm (Fig. 10)." httpUri="https://zenodo.org/record/569117/files/figure.png" pageId="5" pageNumber="6">Fig. 9</figureCitation>
) (similar to position of posterior maxillary lyrifissure in Iocheirata). Median maxillary lyrifissure (
<figureCitation id="1344F19865763349FE2E395763E8F8B4" box="[394,476,1859,1884]" captionStart="FIGURES 8 13" captionStartId="7.[151,264,1848,1871]" captionTargetBox="[236,1334,199,1833]" captionTargetId="figure@7.[230,1346,193,1838]" captionTargetPageId="7" captionText="FIGURES 8 13. Cybella deharvengi n. sp., male. 8, coxae, ventral view; 9, left palp coxa, lateral view; 10, median maxillary lyrifissure; 11, right coxae II IV, dorsal (internal) view; 12, median part of coxae I, showing pit and coxal spines; 13, manducatory process of right palp coxa, ventral view. Abbreviations: cliv, clivus; fbII, border of internal foramen of coxa II, fbIII, border of internal foramen of coxa III, fbIƑ, border of internal foramen of coxa IV, k, condyle; pcs, primary coxal spine; pmIII, posterior margin of coxa III. Scale lines: 0.2 mm (Fig. 9) 0.1 mm (Figs 11 12), 0.05 mm (Figs 10, 13), 0.03 mm (Fig. 10)." httpUri="https://zenodo.org/record/569117/files/figure.png" pageId="5" pageNumber="6">Fig. 10</figureCitation>
) circular, central disk porous; internal apodeme present and attached to muscle. Coxa I (
<figureCitation id="1344F19865763349FF1439726336F890" box="[176,258,1894,1919]" captionStart="FIGURES 8 13" captionStartId="7.[151,264,1848,1871]" captionTargetBox="[236,1334,199,1833]" captionTargetId="figure@7.[230,1346,193,1838]" captionTargetPageId="7" captionText="FIGURES 8 13. Cybella deharvengi n. sp., male. 8, coxae, ventral view; 9, left palp coxa, lateral view; 10, median maxillary lyrifissure; 11, right coxae II IV, dorsal (internal) view; 12, median part of coxae I, showing pit and coxal spines; 13, manducatory process of right palp coxa, ventral view. Abbreviations: cliv, clivus; fbII, border of internal foramen of coxa II, fbIII, border of internal foramen of coxa III, fbIƑ, border of internal foramen of coxa IV, k, condyle; pcs, primary coxal spine; pmIII, posterior margin of coxa III. Scale lines: 0.2 mm (Fig. 9) 0.1 mm (Figs 11 12), 0.05 mm (Figs 10, 13), 0.03 mm (Fig. 10)." httpUri="https://zenodo.org/record/569117/files/figure.png" pageId="5" pageNumber="6">Fig. 12</figureCitation>
) with 1 primary and 815 secondary spines arranged around rim of pit. Coxa III with enlarged, irregular granules anteromedially (
<figureCitation id="1344F19865763349FE10399F63CCF84C" box="[436,504,1931,1956]" captionStart="FIGURES 8 13" captionStartId="7.[151,264,1848,1871]" captionTargetBox="[236,1334,199,1833]" captionTargetId="figure@7.[230,1346,193,1838]" captionTargetPageId="7" captionText="FIGURES 8 13. Cybella deharvengi n. sp., male. 8, coxae, ventral view; 9, left palp coxa, lateral view; 10, median maxillary lyrifissure; 11, right coxae II IV, dorsal (internal) view; 12, median part of coxae I, showing pit and coxal spines; 13, manducatory process of right palp coxa, ventral view. Abbreviations: cliv, clivus; fbII, border of internal foramen of coxa II, fbIII, border of internal foramen of coxa III, fbIƑ, border of internal foramen of coxa IV, k, condyle; pcs, primary coxal spine; pmIII, posterior margin of coxa III. Scale lines: 0.2 mm (Fig. 9) 0.1 mm (Figs 11 12), 0.05 mm (Figs 10, 13), 0.03 mm (Fig. 10)." httpUri="https://zenodo.org/record/569117/files/figure.png" pageId="5" pageNumber="6">Fig. 8</figureCitation>
). Setae of coxae: P ca
<date id="FFC1CBDD65763349FD52399F6169F84B" box="[758,861,1931,1956]" pageId="5" pageNumber="6" value="1930-01-06" valueMax="1930-01-07" valueMin="1930-01-06">30, I 67</date>
, II 7
<date id="FFC1CBDD65763349FC04399F6623F84B" box="[928,1047,1931,1956]" pageId="5" pageNumber="6" value="2010-03-06" valueMax="2010-03-08" valueMin="2010-03-06">10, III 68</date>
,
<date id="FFC1CBDD65763349FB87399866B3F84C" box="[1059,1159,1931,1955]" pageId="5" pageNumber="6">IV 914.</date>
</paragraph>
<paragraph id="8BC0ED1D6576334AFF6339BB666AFEAF" blockId="5.[151,1437,1283,2028]" lastBlockId="6.[151,1436,151,536]" lastPageId="6" lastPageNumber="7" pageId="5" pageNumber="6">
Anterior genital sternite (
<figureCitation id="1344F19865763349FE4439BB6005F827" box="[480,561,1967,1992]" captionStart="FIGURES 14 16" captionStartId="8.[151,264,1719,1742]" captionTargetBox="[151,1428,202,1688]" captionTargetId="figure@8.[151,1436,193,1699]" captionTargetPageId="8" captionText="FIGURES 14 16. Cybella deharvengi n. sp., male. 14, genitalia, ventral view; 15, posterior genital plate, ventral view 16 right side of opisthosomal segments II V, ventral view. Abbreviations: apl, atrial plate; da, dorsal apodeme; dpdg, ducts of posterior dorsal gland; ejca, ejaculatory canal atrium; gp, gland pore; inv, invagination at side of atrial plate; lgs, lateral genital sac; ly, lyrifissure; mgs, median genital sac; pap, papilla on atrial wall; sp, spiracle; sps, spatulate seta; sr, sclerotized ring. Scale lines: 0.1 mm (Figs 14, 16), 0.05 mm (Fig. 15)." httpUri="https://zenodo.org/record/569118/files/figure.png" pageId="5" pageNumber="6">Fig. 16</figureCitation>
) with 8 median setae and a pair of lyrifissures; posterior genital sternite with a median pair of lyrifissures, 911 median setae and a single pair of lateral setae (total 1113). Sternites IVX divided medially. Darkened bars weakly developed on sternites VIVIII, stronger on IX. Denticulation on sternites VIIIXI (weak on VIII and XI). Setae of half-sternites (IVX) 810: 810: 9: 911: 710: 69: 7; all setae posterior. Segment XI undivided, with 11 setae anteriad of anal opening. Sternite XII with 2 setae, closer together than those of tergite XII. Each half-sternite usually with 1 paraxial and 1 antiaxial lyrifissure, but occasionally with up to 4; segment XI with only antiaxial lyrifissures and a pair in front of anal opening.
</paragraph>
<paragraph id="8BC0ED1D6575334AFF633F5861D2FDF7" blockId="6.[151,1436,151,536]" pageId="6" pageNumber="7">
Male genitalia (
<figureCitation id="1344F1986575334AFEDB3F586036FE8A" box="[383,514,332,357]" captionStart="FIGURES 14 16" captionStartId="8.[151,264,1719,1742]" captionTargetBox="[151,1428,202,1688]" captionTargetId="figure@8.[151,1436,193,1699]" captionTargetPageId="8" captionText="FIGURES 14 16. Cybella deharvengi n. sp., male. 14, genitalia, ventral view; 15, posterior genital plate, ventral view 16 right side of opisthosomal segments II V, ventral view. Abbreviations: apl, atrial plate; da, dorsal apodeme; dpdg, ducts of posterior dorsal gland; ejca, ejaculatory canal atrium; gp, gland pore; inv, invagination at side of atrial plate; lgs, lateral genital sac; ly, lyrifissure; mgs, median genital sac; pap, papilla on atrial wall; sp, spiracle; sps, spatulate seta; sr, sclerotized ring. Scale lines: 0.1 mm (Figs 14, 16), 0.05 mm (Fig. 15)." httpUri="https://zenodo.org/record/569118/files/figure.png" pageId="6" pageNumber="7">Figs 1415</figureCitation>
) relatively simple; atrial plate weakly sclerotized and without strongly marked ornamentation, but with a distinct tubular pocket laterally (
<figureCitation id="1344F1986575334AFC913F7B61BFFE67" box="[821,907,367,392]" captionStart="FIGURES 14 16" captionStartId="8.[151,264,1719,1742]" captionTargetBox="[151,1428,202,1688]" captionTargetId="figure@8.[151,1436,193,1699]" captionTargetPageId="8" captionText="FIGURES 14 16. Cybella deharvengi n. sp., male. 14, genitalia, ventral view; 15, posterior genital plate, ventral view 16 right side of opisthosomal segments II V, ventral view. Abbreviations: apl, atrial plate; da, dorsal apodeme; dpdg, ducts of posterior dorsal gland; ejca, ejaculatory canal atrium; gp, gland pore; inv, invagination at side of atrial plate; lgs, lateral genital sac; ly, lyrifissure; mgs, median genital sac; pap, papilla on atrial wall; sp, spiracle; sps, spatulate seta; sr, sclerotized ring. Scale lines: 0.1 mm (Figs 14, 16), 0.05 mm (Fig. 15)." httpUri="https://zenodo.org/record/569118/files/figure.png" pageId="6" pageNumber="7">Fig. 14</figureCitation>
:
<emphasis id="B90B310F6575334AFC333F65618DFE67" box="[919,953,369,392]" italics="true" pageId="6" pageNumber="7">inv</emphasis>
); lateral sacs of moderate size, without an apical constriction; median sac reduced, but with a distinct sclerotized ring at end of duct (
<figureCitation id="1344F1986575334AFB023F8066C8FE42" box="[1190,1276,404,429]" captionStart="FIGURES 14 16" captionStartId="8.[151,264,1719,1742]" captionTargetBox="[151,1428,202,1688]" captionTargetId="figure@8.[151,1436,193,1699]" captionTargetPageId="8" captionText="FIGURES 14 16. Cybella deharvengi n. sp., male. 14, genitalia, ventral view; 15, posterior genital plate, ventral view 16 right side of opisthosomal segments II V, ventral view. Abbreviations: apl, atrial plate; da, dorsal apodeme; dpdg, ducts of posterior dorsal gland; ejca, ejaculatory canal atrium; gp, gland pore; inv, invagination at side of atrial plate; lgs, lateral genital sac; ly, lyrifissure; mgs, median genital sac; pap, papilla on atrial wall; sp, spiracle; sps, spatulate seta; sr, sclerotized ring. Scale lines: 0.1 mm (Figs 14, 16), 0.05 mm (Fig. 15)." httpUri="https://zenodo.org/record/569118/files/figure.png" pageId="6" pageNumber="7">Fig. 15</figureCitation>
:
<emphasis id="B90B310F6575334AFAAD3F81672BFE43" box="[1289,1311,405,428]" italics="true" pageId="6" pageNumber="7">sr</emphasis>
); posterior wall of atrium with a small number of papillae (
<figureCitation id="1344F1986575334AFD643FA36122FE3F" box="[704,790,439,464]" captionStart="FIGURES 14 16" captionStartId="8.[151,264,1719,1742]" captionTargetBox="[151,1428,202,1688]" captionTargetId="figure@8.[151,1436,193,1699]" captionTargetPageId="8" captionText="FIGURES 14 16. Cybella deharvengi n. sp., male. 14, genitalia, ventral view; 15, posterior genital plate, ventral view 16 right side of opisthosomal segments II V, ventral view. Abbreviations: apl, atrial plate; da, dorsal apodeme; dpdg, ducts of posterior dorsal gland; ejca, ejaculatory canal atrium; gp, gland pore; inv, invagination at side of atrial plate; lgs, lateral genital sac; ly, lyrifissure; mgs, median genital sac; pap, papilla on atrial wall; sp, spiracle; sps, spatulate seta; sr, sclerotized ring. Scale lines: 0.1 mm (Figs 14, 16), 0.05 mm (Fig. 15)." httpUri="https://zenodo.org/record/569118/files/figure.png" pageId="6" pageNumber="7">Fig. 14</figureCitation>
:
<emphasis id="B90B310F6575334AFC863FAD6178FE3F" box="[802,844,441,464]" italics="true" pageId="6" pageNumber="7">pap</emphasis>
) that are not associated with glands; posterior plate with 34 internal setae on each side, of which the posterior pair is enlarged and slightly spatulate. Mature, encysted spermatozoa (observed in
<emphasis id="B90B310F6575334AFE193C15607CFDF7" box="[445,584,511,536]" italics="true" pageId="6" pageNumber="7">vas deferens</emphasis>
) with a maximum diameter of 4 µm.
</paragraph>
<caption id="DF00BD956575334AFF33397C6156F834" httpUri="https://zenodo.org/record/569116/files/figure.png" pageId="6" pageNumber="7" targetBox="[233,1351,598,1844]" targetPageId="6">
<paragraph id="8BC0ED1D6575334AFF33397C6156F834" blockId="6.[151,1436,1896,2011]" pageId="6" pageNumber="7">
<emphasis id="B90B310F6575334AFF33397C630AF891" bold="true" box="[151,318,1896,1919]" pageId="6" pageNumber="7">FIGURES 37.</emphasis>
<taxonomicName id="4C7F969E6575334AFEEC397D6027F890" box="[328,531,1897,1919]" class="Arachnida" family="Feaellidae" genus="Cybella" kingdom="Animalia" order="Pseudoscorpiones" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="deharvengi" status="sp. nov.">
<emphasis id="B90B310F6575334AFEEC397D6027F890" box="[328,531,1897,1919]" italics="true" pageId="6" pageNumber="7">Cybella deharvengi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2388C746575334AFDB9397D606EF891" box="[541,602,1897,1918]" pageId="6" pageNumber="7" rank="species">
<emphasis id="B90B310F6575334AFDB9397D606EF891" bold="true" box="[541,602,1897,1918]" pageId="6" pageNumber="7">n. sp.</emphasis>
</taxonomicNameLabel>
, male. 3, pleurum at level of segments VIIIX, ventral view; 4, carapace and anterior tergites, dorsal view; 5, carapace, lateral view; 6, opisthosoma, anterior view, prosoma removed; 7, lateral tergal plate of segment I, seen in same orientation as Fig. 6. Abbreviations:
<emphasis id="B90B310F6575334AFCBD39B26109F853" box="[793,829,1958,1980]" italics="true" pageId="6" pageNumber="7">den</emphasis>
, denticulate ornamentation;
<emphasis id="B90B310F6575334AFBFA39B36643F853" box="[1118,1143,1959,1980]" italics="true" pageId="6" pageNumber="7">gp</emphasis>
, gland pore;
<emphasis id="B90B310F6575334AFB5F39B26738F853" box="[1275,1292,1958,1980]" italics="true" pageId="6" pageNumber="7">ly</emphasis>
, lyrifissure;
<emphasis id="B90B310F6575334AFA2839B367A2F853" box="[1420,1430,1959,1980]" italics="true" pageId="6" pageNumber="7">s</emphasis>
, setae;
<emphasis id="B90B310F6575334AFF7339D26336F834" box="[215,258,1990,2011]" italics="true" pageId="6" pageNumber="7">TII</emphasis>
, tergite II. Scale lines 0.2 mm (Figs 16), 0.05 mm (Fig. 7).
</paragraph>
</caption>
<caption id="DF00BD956574334BFF33392C628BF807" httpUri="https://zenodo.org/record/569117/files/figure.png" pageId="7" pageNumber="8" targetBox="[236,1334,199,1833]" targetPageId="7">
<paragraph id="8BC0ED1D6574334BFF33392C628BF807" blockId="7.[151,1436,1848,2024]" pageId="7" pageNumber="8">
<emphasis id="B90B310F6574334BFF33392C6373F8A1" bold="true" box="[151,327,1848,1871]" pageId="7" pageNumber="8">FIGURES 813.</emphasis>
<taxonomicName id="4C7F969E6574334BFEE9392D6021F8A0" box="[333,533,1849,1871]" class="Arachnida" family="Feaellidae" genus="Cybella" kingdom="Animalia" order="Pseudoscorpiones" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="deharvengi" status="sp. nov.">
<emphasis id="B90B310F6574334BFEE9392D6021F8A0" box="[333,533,1849,1871]" italics="true" pageId="7" pageNumber="8">Cybella deharvengi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2388C746574334BFDB8392D6061F8A1" box="[540,597,1849,1870]" pageId="7" pageNumber="8" rank="species">
<emphasis id="B90B310F6574334BFDB8392D6061F8A1" bold="true" box="[540,597,1849,1870]" pageId="7" pageNumber="8">n. sp.</emphasis>
</taxonomicNameLabel>
, male. 8, coxae, ventral view; 9, left palp coxa, lateral view; 10, median maxillary lyrifissure; 11, right coxae IIIV, dorsal (internal) view; 12, median part of coxae I, showing pit and coxal spines; 13, manducatory process of right palp coxa, ventral view. Abbreviations:
<emphasis id="B90B310F6574334BFCCC396261BFF863" box="[872,907,1910,1932]" italics="true" pageId="7" pageNumber="8">cliv</emphasis>
, clivus;
<emphasis id="B90B310F6574334BFC403962663CF863" box="[996,1032,1910,1932]" italics="true" pageId="7" pageNumber="8">fbII</emphasis>
, border of internal foramen of coxa II,
<emphasis id="B90B310F6574334BFF33398162F0F844" box="[151,196,1941,1963]" italics="true" pageId="7" pageNumber="8">fbIII</emphasis>
, border of internal foramen of coxa III,
<emphasis id="B90B310F6574334BFDC5398160BDF844" box="[609,649,1941,1963]" italics="true" pageId="7" pageNumber="8">fbIƑ</emphasis>
, border of internal foramen of coxa IV,
<emphasis id="B90B310F6574334BFB803981661BF844" box="[1060,1071,1941,1963]" italics="true" pageId="7" pageNumber="8">k</emphasis>
, condyle;
<emphasis id="B90B310F6574334BFB3F39826689F844" box="[1179,1213,1942,1963]" italics="true" pageId="7" pageNumber="8">pcs</emphasis>
, primary coxal spine;
<emphasis id="B90B310F6574334BFF3339A162FBF825" box="[151,207,1973,1994]" italics="true" pageId="7" pageNumber="8">pmIII</emphasis>
, posterior margin of coxa III. Scale lines: 0.2 mm (Fig. 9) 0.1 mm (Figs 1112), 0.05 mm (Figs 10, 13), 0.03 mm (Fig. 10).
</paragraph>
</caption>
<caption id="DF00BD95657B3344FF3338A3600DF8A6" httpUri="https://zenodo.org/record/569118/files/figure.png" pageId="8" pageNumber="9" targetBox="[151,1428,202,1688]" targetPageId="8">
<paragraph id="8BC0ED1D657B3344FF3338A3600DF8A6" blockId="8.[151,1437,1719,1865]" pageId="8" pageNumber="9">
<emphasis id="B90B310F657B3344FF3338A36367F922" bold="true" box="[151,339,1719,1742]" pageId="8" pageNumber="9">FIGURES 1416.</emphasis>
<taxonomicName id="4C7F969E657B3344FEFD38AC602BF921" box="[345,543,1720,1742]" class="Arachnida" family="Feaellidae" genus="Cybella" kingdom="Animalia" order="Pseudoscorpiones" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="deharvengi" status="sp. nov.">
<emphasis id="B90B310F657B3344FEFD38AC602BF921" box="[345,543,1720,1742]" italics="true" pageId="8" pageNumber="9">Cybella deharvengi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2388C74657B3344FD8138AC606AF922" box="[549,606,1720,1741]" pageId="8" pageNumber="9" rank="species">
<emphasis id="B90B310F657B3344FD8138AC606AF922" bold="true" box="[549,606,1720,1741]" pageId="8" pageNumber="9">n. sp.</emphasis>
</taxonomicNameLabel>
, male. 14, genitalia, ventral view; 15, posterior genital plate, ventral view 16 right side of opisthosomal segments IIV, ventral view. Abbreviations:
<emphasis id="B90B310F657B3344FC9638C36166F902" box="[818,850,1751,1773]" italics="true" pageId="8" pageNumber="9">apl</emphasis>
, atrial plate;
<emphasis id="B90B310F657B3344FC7C38C361C5F902" box="[984,1009,1751,1773]" italics="true" pageId="8" pageNumber="9">da</emphasis>
, dorsal apodeme;
<emphasis id="B90B310F657B3344FB0F38C366EAF902" box="[1195,1246,1751,1773]" italics="true" pageId="8" pageNumber="9">dpdg</emphasis>
, ducts of posterior dorsal gland;
<emphasis id="B90B310F657B3344FE8738E36379F8E3" box="[291,333,1783,1804]" italics="true" pageId="8" pageNumber="9">ejca</emphasis>
, ejaculatory canal atrium;
<emphasis id="B90B310F657B3344FDC538E3604EF8E3" box="[609,634,1783,1804]" italics="true" pageId="8" pageNumber="9">gp</emphasis>
, gland pore;
<emphasis id="B90B310F657B3344FCA038E36115F8E3" box="[772,801,1783,1804]" italics="true" pageId="8" pageNumber="9">inv</emphasis>
, invagination at side of atrial plate;
<emphasis id="B90B310F657B3344FB3338E16680F8E4" box="[1175,1204,1781,1803]" italics="true" pageId="8" pageNumber="9">lgs</emphasis>
, lateral genital sac;
<emphasis id="B90B310F657B3344FA2138E167A2F8E4" box="[1413,1430,1781,1803]" italics="true" pageId="8" pageNumber="9">ly</emphasis>
, lyrifissure;
<emphasis id="B90B310F657B3344FEAF39016300F8C5" box="[267,308,1813,1834]" italics="true" pageId="8" pageNumber="9">mgs</emphasis>
, median genital sac;
<emphasis id="B90B310F657B3344FDAF39016005F8C5" box="[523,561,1813,1834]" italics="true" pageId="8" pageNumber="9">pap</emphasis>
, papilla on atrial wall;
<emphasis id="B90B310F657B3344FCB839016106F8C5" box="[796,818,1813,1834]" italics="true" pageId="8" pageNumber="9">sp</emphasis>
, spiracle;
<emphasis id="B90B310F657B3344FC3839016188F8C5" box="[924,956,1813,1834]" italics="true" pageId="8" pageNumber="9">sps</emphasis>
, spatulate seta;
<emphasis id="B90B310F657B3344FBFA39016645F8C5" box="[1118,1137,1813,1834]" italics="true" pageId="8" pageNumber="9">sr</emphasis>
, sclerotized ring. Scale lines: 0.1 mm (Figs 14, 16), 0.05 mm (Fig. 15).
</paragraph>
</caption>
<paragraph id="8BC0ED1D657B3345FF63396161DAFE67" blockId="8.[151,1436,1909,2006]" lastBlockId="9.[151,1437,151,392]" lastPageId="9" lastPageNumber="10" pageId="8" pageNumber="9">
Chelicera with reticulation ending fairly abruptly in a raised crest; palm with 5 large and 67 small setae; 2 lyrifissures on antiaxial face (
<figureCitation id="1344F198657B3344FE4A398D6077F85D" box="[494,579,1945,1970]" captionStart="FIGURES 17 22" captionStartId="9.[151,264,1855,1878]" captionTargetBox="[166,1401,441,1799]" captionTargetId="figure@9.[151,1436,423,1846]" captionTargetPageId="9" captionText="FIGURES 17 22. Cybella deharvengi n. sp., male, left chelicera. 17, whole chelicera, antiaxial view; 18, tip of movable finger, antiaxial view; 19, distal part of chelicera, ventral view; 20, spinneret, ventral view; 21, distal part of chelicera (movable finger omitted), paraxial view; 22, chelicera in posterior view (dorsal at top, paraxial side to left). Abbreviations: ds, ducts of serrula interior; f, foramen; gs, galeal seta; ly, lyrifissure; pr, distal process of serrula exterior; r, rallum; s. ext, serrula exterior; s. int., serrula interior. Other abbreviations (b, es, gs, is, ls, sb) are designations of setae. Scale lines 0.05 mm (Figs 18 20 share same scale line)." httpUri="https://zenodo.org/record/569119/files/figure.png" pageId="8" pageNumber="9">Fig. 17</figureCitation>
), 1 on paraxial face at base of fixed finger (
<figureCitation id="1344F198657B3344FBEC398D66AAF85D" box="[1096,1182,1945,1970]" captionStart="FIGURES 17 22" captionStartId="9.[151,264,1855,1878]" captionTargetBox="[166,1401,441,1799]" captionTargetId="figure@9.[151,1436,423,1846]" captionTargetPageId="9" captionText="FIGURES 17 22. Cybella deharvengi n. sp., male, left chelicera. 17, whole chelicera, antiaxial view; 18, tip of movable finger, antiaxial view; 19, distal part of chelicera, ventral view; 20, spinneret, ventral view; 21, distal part of chelicera (movable finger omitted), paraxial view; 22, chelicera in posterior view (dorsal at top, paraxial side to left). Abbreviations: ds, ducts of serrula interior; f, foramen; gs, galeal seta; ly, lyrifissure; pr, distal process of serrula exterior; r, rallum; s. ext, serrula exterior; s. int., serrula interior. Other abbreviations (b, es, gs, is, ls, sb) are designations of setae. Scale lines 0.05 mm (Figs 18 20 share same scale line)." httpUri="https://zenodo.org/record/569119/files/figure.png" pageId="8" pageNumber="9">Fig. 21</figureCitation>
). Base of fixed finger concave on paraxial side. Movable finger with a single, large tooth subdistally (
<figureCitation id="1344F198657B3344FB8839A966B5F839" box="[1068,1153,1981,2006]" captionStart="FIGURES 17 22" captionStartId="9.[151,264,1855,1878]" captionTargetBox="[166,1401,441,1799]" captionTargetId="figure@9.[151,1436,423,1846]" captionTargetPageId="9" captionText="FIGURES 17 22. Cybella deharvengi n. sp., male, left chelicera. 17, whole chelicera, antiaxial view; 18, tip of movable finger, antiaxial view; 19, distal part of chelicera, ventral view; 20, spinneret, ventral view; 21, distal part of chelicera (movable finger omitted), paraxial view; 22, chelicera in posterior view (dorsal at top, paraxial side to left). Abbreviations: ds, ducts of serrula interior; f, foramen; gs, galeal seta; ly, lyrifissure; pr, distal process of serrula exterior; r, rallum; s. ext, serrula exterior; s. int., serrula interior. Other abbreviations (b, es, gs, is, ls, sb) are designations of setae. Scale lines 0.05 mm (Figs 18 20 share same scale line)." httpUri="https://zenodo.org/record/569119/files/figure.png" pageId="8" pageNumber="9">Fig. 18</figureCitation>
). Spinneret short (1823 µm) simple, tubular (not flattened), but flared at base (
<figureCitation id="1344F198657A3345FCA23E8361BFFF5F" box="[774,907,151,176]" captionStart="FIGURES 17 22" captionStartId="9.[151,264,1855,1878]" captionTargetBox="[166,1401,441,1799]" captionTargetId="figure@9.[151,1436,423,1846]" captionTargetPageId="9" captionText="FIGURES 17 22. Cybella deharvengi n. sp., male, left chelicera. 17, whole chelicera, antiaxial view; 18, tip of movable finger, antiaxial view; 19, distal part of chelicera, ventral view; 20, spinneret, ventral view; 21, distal part of chelicera (movable finger omitted), paraxial view; 22, chelicera in posterior view (dorsal at top, paraxial side to left). Abbreviations: ds, ducts of serrula interior; f, foramen; gs, galeal seta; ly, lyrifissure; pr, distal process of serrula exterior; r, rallum; s. ext, serrula exterior; s. int., serrula interior. Other abbreviations (b, es, gs, is, ls, sb) are designations of setae. Scale lines 0.05 mm (Figs 18 20 share same scale line)." httpUri="https://zenodo.org/record/569119/files/figure.png" pageId="9" pageNumber="10">Figs 1820</figureCitation>
); 3 internal silk ducts, these lying side by side and thus only evident in ventral view. Serrula exterior (
<figureCitation id="1344F198657A3345FD5B3EA8617BFF3B" box="[767,847,188,213]" captionStart="FIGURES 17 22" captionStartId="9.[151,264,1855,1878]" captionTargetBox="[166,1401,441,1799]" captionTargetId="figure@9.[151,1436,423,1846]" captionTargetPageId="9" captionText="FIGURES 17 22. Cybella deharvengi n. sp., male, left chelicera. 17, whole chelicera, antiaxial view; 18, tip of movable finger, antiaxial view; 19, distal part of chelicera, ventral view; 20, spinneret, ventral view; 21, distal part of chelicera (movable finger omitted), paraxial view; 22, chelicera in posterior view (dorsal at top, paraxial side to left). Abbreviations: ds, ducts of serrula interior; f, foramen; gs, galeal seta; ly, lyrifissure; pr, distal process of serrula exterior; r, rallum; s. ext, serrula exterior; s. int., serrula interior. Other abbreviations (b, es, gs, is, ls, sb) are designations of setae. Scale lines 0.05 mm (Figs 18 20 share same scale line)." httpUri="https://zenodo.org/record/569119/files/figure.png" pageId="9" pageNumber="10">Fig. 19</figureCitation>
) with 1821 blades, arranged in an arc, plus a small, distally-directed apical process (
<figureCitation id="1344F198657A3345FDAD3ECB6068FF17" box="[521,604,223,248]" captionStart="FIGURES 17 22" captionStartId="9.[151,264,1855,1878]" captionTargetBox="[166,1401,441,1799]" captionTargetId="figure@9.[151,1436,423,1846]" captionTargetPageId="9" captionText="FIGURES 17 22. Cybella deharvengi n. sp., male, left chelicera. 17, whole chelicera, antiaxial view; 18, tip of movable finger, antiaxial view; 19, distal part of chelicera, ventral view; 20, spinneret, ventral view; 21, distal part of chelicera (movable finger omitted), paraxial view; 22, chelicera in posterior view (dorsal at top, paraxial side to left). Abbreviations: ds, ducts of serrula interior; f, foramen; gs, galeal seta; ly, lyrifissure; pr, distal process of serrula exterior; r, rallum; s. ext, serrula exterior; s. int., serrula interior. Other abbreviations (b, es, gs, is, ls, sb) are designations of setae. Scale lines 0.05 mm (Figs 18 20 share same scale line)." httpUri="https://zenodo.org/record/569119/files/figure.png" pageId="9" pageNumber="10">Fig. 18</figureCitation>
) (total 2022). Serrula interior (
<figureCitation id="1344F198657A3345FC763ECB6612FF17" box="[978,1062,223,248]" captionStart="FIGURES 17 22" captionStartId="9.[151,264,1855,1878]" captionTargetBox="[166,1401,441,1799]" captionTargetId="figure@9.[151,1436,423,1846]" captionTargetPageId="9" captionText="FIGURES 17 22. Cybella deharvengi n. sp., male, left chelicera. 17, whole chelicera, antiaxial view; 18, tip of movable finger, antiaxial view; 19, distal part of chelicera, ventral view; 20, spinneret, ventral view; 21, distal part of chelicera (movable finger omitted), paraxial view; 22, chelicera in posterior view (dorsal at top, paraxial side to left). Abbreviations: ds, ducts of serrula interior; f, foramen; gs, galeal seta; ly, lyrifissure; pr, distal process of serrula exterior; r, rallum; s. ext, serrula exterior; s. int., serrula interior. Other abbreviations (b, es, gs, is, ls, sb) are designations of setae. Scale lines 0.05 mm (Figs 18 20 share same scale line)." httpUri="https://zenodo.org/record/569119/files/figure.png" pageId="9" pageNumber="10">Fig. 21</figureCitation>
) with about 1315 blades, basal blades enlarged and difficult to distinguish individually (counts based on internal canals). Rallum (
<figureCitation id="1344F198657A3345FAAA3F1067A0FEF3" box="[1294,1428,260,285]" captionStart="FIGURES 17 22" captionStartId="9.[151,264,1855,1878]" captionTargetBox="[166,1401,441,1799]" captionTargetId="figure@9.[151,1436,423,1846]" captionTargetPageId="9" captionText="FIGURES 17 22. Cybella deharvengi n. sp., male, left chelicera. 17, whole chelicera, antiaxial view; 18, tip of movable finger, antiaxial view; 19, distal part of chelicera, ventral view; 20, spinneret, ventral view; 21, distal part of chelicera (movable finger omitted), paraxial view; 22, chelicera in posterior view (dorsal at top, paraxial side to left). Abbreviations: ds, ducts of serrula interior; f, foramen; gs, galeal seta; ly, lyrifissure; pr, distal process of serrula exterior; r, rallum; s. ext, serrula exterior; s. int., serrula interior. Other abbreviations (b, es, gs, is, ls, sb) are designations of setae. Scale lines 0.05 mm (Figs 18 20 share same scale line)." httpUri="https://zenodo.org/record/569119/files/figure.png" pageId="9" pageNumber="10">Figs 19, 21</figureCitation>
) with two long (anterior 26 µm, posterior 25 µm), simple blades, closely appressed and thus difficult to distinguish, tips blunt, but this is only evident when they are observed at high magnification; in close proximity to basal blade of serrula exterior. Galeal seta (
<emphasis id="B90B310F657A3345FE5C3F656025FE67" box="[504,529,369,392]" italics="true" pageId="9" pageNumber="10">gs</emphasis>
) barely extending beyond tip of spinneret.
</paragraph>
<caption id="DF00BD95657A3345FF33392B630BF800" httpUri="https://zenodo.org/record/569119/files/figure.png" pageId="9" pageNumber="10" targetBox="[166,1401,441,1799]" targetPageId="9">
<paragraph id="8BC0ED1D657A3345FF33392B630BF800" blockId="9.[151,1436,1855,2031]" pageId="9" pageNumber="10">
<emphasis id="B90B310F657A3345FF33392B6363F8BA" bold="true" box="[151,343,1855,1878]" pageId="9" pageNumber="10">FIGURES 1722.</emphasis>
<taxonomicName id="4C7F969E657A3345FEC43954601FF8B9" box="[352,555,1856,1878]" class="Arachnida" family="Feaellidae" genus="Cybella" kingdom="Animalia" order="Pseudoscorpiones" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="deharvengi" status="sp. nov.">
<emphasis id="B90B310F657A3345FEC43954601FF8B9" box="[352,555,1856,1878]" italics="true" pageId="9" pageNumber="10">Cybella deharvengi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2388C74657A3345FD9039546045F8BA" box="[564,625,1856,1877]" pageId="9" pageNumber="10" rank="species">
<emphasis id="B90B310F657A3345FD9039546045F8BA" bold="true" box="[564,625,1856,1877]" pageId="9" pageNumber="10">n. sp.</emphasis>
</taxonomicNameLabel>
, male, left chelicera. 17, whole chelicera, antiaxial view; 18, tip of movable finger, antiaxial view; 19, distal part of chelicera, ventral view; 20, spinneret, ventral view; 21, distal part of chelicera (movable finger omitted), paraxial view; 22, chelicera in posterior view (dorsal at top, paraxial side to left). Abbreviations:
<emphasis id="B90B310F657A3345FA813969670FF87C" box="[1317,1339,1917,1939]" italics="true" pageId="9" pageNumber="10">ds</emphasis>
, ducts of serrula interior;
<emphasis id="B90B310F657A3345FE9F39886376F85D" box="[315,322,1948,1970]" italics="true" pageId="9" pageNumber="10">f</emphasis>
, foramen;
<emphasis id="B90B310F657A3345FE16398963FCF85D" box="[434,456,1949,1970]" italics="true" pageId="9" pageNumber="10">gs</emphasis>
, galeal seta;
<emphasis id="B90B310F657A3345FDEA3988606BF85D" box="[590,607,1948,1970]" italics="true" pageId="9" pageNumber="10">ly</emphasis>
, lyrifissure;
<emphasis id="B90B310F657A3345FD45398960C2F85D" box="[737,758,1949,1970]" italics="true" pageId="9" pageNumber="10">pr</emphasis>
, distal process of serrula exterior;
<emphasis id="B90B310F657A3345FBFD39896656F85D" box="[1113,1122,1949,1970]" italics="true" pageId="9" pageNumber="10">r</emphasis>
, rallum;
<emphasis id="B90B310F657A3345FB1B398966D9F85D" box="[1215,1261,1949,1970]" italics="true" pageId="9" pageNumber="10">s.ext</emphasis>
, serrula exterior;
<emphasis id="B90B310F657A3345FF3339AF62FDF83F" box="[151,201,1979,2000]" italics="true" pageId="9" pageNumber="10">s.int.</emphasis>
, serrula interior. Other abbreviations (
<emphasis id="B90B310F657A3345FDE939AE606DF83F" box="[589,601,1978,2000]" italics="true" pageId="9" pageNumber="10">b</emphasis>
,
<emphasis id="B90B310F657A3345FDC339AF6048F83F" box="[615,636,1979,2000]" italics="true" pageId="9" pageNumber="10">es</emphasis>
,
<emphasis id="B90B310F657A3345FD2D39AF60ABF83F" box="[649,671,1979,2000]" italics="true" pageId="9" pageNumber="10">gs</emphasis>
,
<emphasis id="B90B310F657A3345FD0839AF6089F83F" box="[684,701,1979,2000]" italics="true" pageId="9" pageNumber="10">is</emphasis>
,
<emphasis id="B90B310F657A3345FD6E39AE60EFF83F" box="[714,731,1978,2000]" italics="true" pageId="9" pageNumber="10">ls</emphasis>
,
<emphasis id="B90B310F657A3345FD4339AE60C9F83F" box="[743,765,1978,2000]" italics="true" pageId="9" pageNumber="10">sb</emphasis>
) are designations of setae. Scale lines 0.05 mm (Figs 1820 share same scale line).
</paragraph>
</caption>
<caption id="DF00BD9565793346FF3338CC6015F809" httpUri="https://zenodo.org/record/569120/files/figure.png" pageId="10" pageNumber="11" targetBox="[232,1344,207,1715]" targetPageId="10">
<paragraph id="8BC0ED1D65793346FF3338CC6015F809" blockId="10.[151,1436,1752,2022]" pageId="10" pageNumber="11">
<emphasis id="B90B310F65793346FF3338CC636CF902" bold="true" box="[151,344,1752,1775]" pageId="10" pageNumber="11">FIGURES 2334.</emphasis>
<taxonomicName id="4C7F969E65793346FEC638CC6019F901" box="[354,557,1752,1774]" class="Arachnida" family="Feaellidae" genus="Cybella" kingdom="Animalia" order="Pseudoscorpiones" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="deharvengi" status="sp. nov.">
<emphasis id="B90B310F65793346FEC638CC6019F901" box="[354,557,1752,1774]" italics="true" pageId="10" pageNumber="11">Cybella deharvengi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2388C7465793346FD9338CD6041F901" box="[567,629,1753,1774]" pageId="10" pageNumber="11" rank="species">
<emphasis id="B90B310F65793346FD9338CD6041F901" bold="true" box="[567,629,1753,1774]" pageId="10" pageNumber="11">n. sp.</emphasis>
</taxonomicNameLabel>
, male right palp. 23, chela, dorsal view; 24, palp minus chela, dorsal view (reticulate ornamentation drawn in part on femur); 25, tip of fixed chelal finger, dorsal view; 26, basal tuberculate tooth of fixed finger, dorsal view (relative to chela); 27, tip of movable chelal finger, paraxial view; 28, basal tuberculate tooth of movable chelal finger, ventral view (relative to chela), distal end to left; 29, chela, paraxial view (setae omitted); 30, chela, antiaxial view; 31, coupled sensilla; 32, distal sensillum; 33, distal end of movable finger, ventral view; 34, grouped chemosensory setae, antiaxial view. Abbreviations:
<emphasis id="B90B310F65793346FE46396663C1F867" box="[482,501,1906,1928]" italics="true" pageId="10" pageNumber="11">af</emphasis>
, apical sensillum of fixed finger;
<emphasis id="B90B310F65793346FCD3396761A2F867" box="[887,918,1907,1928]" italics="true" pageId="10" pageNumber="11">am</emphasis>
1, apical sensilla of movable finger;
<emphasis id="B90B310F65793346FA8F39666776F867" box="[1323,1346,1906,1928]" italics="true" pageId="10" pageNumber="11">dx</emphasis>
, duplex trichobothria;
<emphasis id="B90B310F65793346FE833987630FF846" box="[295,315,1939,1961]" italics="true" pageId="10" pageNumber="11">ld</emphasis>
, probable lamina defensor;
<emphasis id="B90B310F65793346FDFC3987605DF846" box="[600,617,1939,1961]" italics="true" pageId="10" pageNumber="11">ly</emphasis>
, lyrifissure;
<emphasis id="B90B310F65793346FD4E39806136F846" box="[746,770,1940,1961]" italics="true" pageId="10" pageNumber="11">pc</emphasis>
, coupled sensilla. Other abbreviations (
<emphasis id="B90B310F65793346FB323987669AF846" box="[1174,1198,1939,1961]" italics="true" pageId="10" pageNumber="11">eb</emphasis>
,
<emphasis id="B90B310F65793346FB1F398766E9F846" box="[1211,1245,1939,1961]" italics="true" pageId="10" pageNumber="11">esb</emphasis>
,
<emphasis id="B90B310F65793346FB4E39806732F846" box="[1258,1286,1940,1961]" italics="true" pageId="10" pageNumber="11">est</emphasis>
,
<emphasis id="B90B310F65793346FAB739806711F846" box="[1299,1317,1940,1961]" italics="true" pageId="10" pageNumber="11">et</emphasis>
,
<emphasis id="B90B310F65793346FA9639876772F846" box="[1330,1350,1939,1961]" italics="true" pageId="10" pageNumber="11">ib</emphasis>
,
<emphasis id="B90B310F65793346FAF739876744F846" box="[1363,1392,1939,1961]" italics="true" pageId="10" pageNumber="11">isb</emphasis>
,
<emphasis id="B90B310F65793346FAD9398067A1F846" box="[1405,1429,1940,1961]" italics="true" pageId="10" pageNumber="11">ist</emphasis>
,
<emphasis id="B90B310F65793346FF3339A76291F827" box="[151,165,1971,1992]" italics="true" pageId="10" pageNumber="11">it</emphasis>
,
<emphasis id="B90B310F65793346FF1539A66289F827" box="[177,189,1970,1992]" italics="true" pageId="10" pageNumber="11">b</emphasis>
,
<emphasis id="B90B310F65793346FF6E39A662D4F827" box="[202,224,1970,1992]" italics="true" pageId="10" pageNumber="11">sb</emphasis>
,
<emphasis id="B90B310F65793346FF4939A762CAF827" box="[237,254,1971,1992]" italics="true" pageId="10" pageNumber="11">st</emphasis>
,
<emphasis id="B90B310F65793346FEAE39A76325F827" box="[266,273,1971,1992]" italics="true" pageId="10" pageNumber="11">t</emphasis>
) are standard designations of trichobothria. Scale lines: 0.2 mm (Figs 2324, 29), 0.1 mm (Fig. 30), 0.05 mm (Figs 2528, 3334), 0.01 mm (Figs 3132).
</paragraph>
</caption>
<caption id="DF00BD9565783347FF33392E6174F842" httpUri="https://zenodo.org/record/569121/files/figure.png" pageId="11" pageNumber="12" targetBox="[159,1426,202,1817]" targetPageId="11">
<paragraph id="8BC0ED1D65783347FF33392E6174F842" blockId="11.[151,1437,1850,1965]" pageId="11" pageNumber="12">
<emphasis id="B90B310F65783347FF33392E6367F8BF" bold="true" box="[151,339,1850,1873]" pageId="11" pageNumber="12">FIGURES 3543.</emphasis>
<taxonomicName id="4C7F969E65783347FEFD392F6014F8BE" box="[345,544,1851,1873]" class="Arachnida" family="Feaellidae" genus="Cybella" kingdom="Animalia" order="Pseudoscorpiones" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="deharvengi" status="sp. nov.">
<emphasis id="B90B310F65783347FEFD392F6014F8BE" box="[345,544,1851,1873]" italics="true" pageId="11" pageNumber="12">Cybella deharvengi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2388C7465783347FD82392F606BF8BF" box="[550,607,1851,1872]" pageId="11" pageNumber="12" rank="species">
<emphasis id="B90B310F65783347FD82392F606BF8BF" bold="true" box="[550,607,1851,1872]" pageId="11" pageNumber="12">n. sp.</emphasis>
</taxonomicNameLabel>
, male right legs. 35, leg I, antiaxial view; 36, trochanter and femur of leg I, dorsal view; 37, trochanter and femur of leg I, paraxial view; 38, leg IV, antiaxial view; 39, patella of leg I, dorsal view; 40, patella of leg IV, dorsal view; 41, leg III, antiaxial view; 42, arolium of leg III, ventral; 43 apotele of leg I, antiaxial view. Abbreviations:
<emphasis id="B90B310F65783347FF333983629CF842" box="[151,168,1943,1965]" italics="true" pageId="11" pageNumber="12">ly</emphasis>
, lyrifissure;. Scale lines 0.2 mm (Figs 3541), 0.05 (Figs 4243).
</paragraph>
</caption>
<caption id="DF00BD95657F3340FF333D586028FC2E" httpUri="https://zenodo.org/record/569122/files/figure.png" pageId="12" pageNumber="13" targetBox="[156,1429,202,816]" targetPageId="12">
<paragraph id="8BC0ED1D657F3340FF333D586028FC2E" blockId="12.[151,1436,844,961]" pageId="12" pageNumber="13">
<emphasis id="B90B310F657F3340FF333D586360FC8E" bold="true" box="[151,340,844,867]" pageId="12" pageNumber="13">FIGURES 4447.</emphasis>
<taxonomicName id="4C7F969E657F3340FEFF3D586016FC8D" box="[347,546,844,866]" class="Arachnida" family="Feaellidae" genus="Cybella" kingdom="Animalia" order="Pseudoscorpiones" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="deharvengi" status="sp. nov.">
<emphasis id="B90B310F657F3340FEFF3D586016FC8D" box="[347,546,844,866]" italics="true" pageId="12" pageNumber="13">Cybella deharvengi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2388C74657F3340FD8D3D596056FC8D" box="[553,610,845,866]" pageId="12" pageNumber="13" rank="species">
<emphasis id="B90B310F657F3340FD8D3D596056FC8D" bold="true" box="[553,610,845,866]" pageId="12" pageNumber="13">n. sp.</emphasis>
</taxonomicNameLabel>
, paratype tritonymph. 44, chelicera, antiaxial view; 45, tip of movable cheliceral finger, antiaxial view; 46, right chela, antiaxial view; 47 grouped chemosensory setae, antiaxial view. Abbreviations:
<emphasis id="B90B310F657F3340FAD43D7867BAFC6E" box="[1392,1422,876,897]" italics="true" pageId="12" pageNumber="13">am</emphasis>
1, apical sensillum of movable finger;
<emphasis id="B90B310F657F3340FDA03D99602DFC4D" box="[516,537,909,930]" italics="true" pageId="12" pageNumber="13">pr</emphasis>
, distal process of serrula exterior;
<emphasis id="B90B310F657F3340FCDC3D9961A4FC4D" box="[888,912,909,930]" italics="true" pageId="12" pageNumber="13">pc</emphasis>
, coupled sensilla. Scale lines 0.1 mm (Figs 44, 46), 0.05 mm (Fig. 45), 0.02 mm (Fig. 47).
</paragraph>
</caption>
<paragraph id="8BC0ED1D657F3340FF633DFC60E2F921" blockId="12.[151,1437,1000,2029]" pageId="12" pageNumber="13">
Palps of typical feaellid facies, femur very robust, patella and chela small (
<figureCitation id="1344F198657F3340FB973DFC668EFBEF" box="[1075,1210,1000,1025]" captionStart="FIGURES 23 34" captionStartId="10.[151,264,1752,1775]" captionTargetBox="[232,1344,207,1715]" captionTargetId="figure@10.[224,1350,193,1724]" captionTargetPageId="10" captionText="FIGURES 23 34. Cybella deharvengi n. sp., male right palp. 23, chela, dorsal view; 24, palp minus chela, dorsal view (reticulate ornamentation drawn in part on femur); 25, tip of fixed chelal finger, dorsal view; 26, basal tuberculate tooth of fixed finger, dorsal view (relative to chela); 27, tip of movable chelal finger, paraxial view; 28, basal tuberculate tooth of movable chelal finger, ventral view (relative to chela), distal end to left; 29, chela, paraxial view (setae omitted); 30, chela, antiaxial view; 31, coupled sensilla; 32, distal sensillum; 33, distal end of movable finger, ventral view; 34, grouped chemosensory setae, antiaxial view. Abbreviations: af, apical sensillum of fixed finger; am 1, apical sensilla of movable finger; dx, duplex trichobothria; ld, probable lamina defensor; ly, lyrifissure; pc, coupled sensilla. Other abbreviations (eb, esb, est, et, ib, isb, ist, it, b, sb, st, t) are standard designations of trichobothria. Scale lines: 0.2 mm (Figs 23 24, 29), 0.1 mm (Fig. 30), 0.05 mm (Figs 25 28, 33 34), 0.01 mm (Figs 31 32)." httpUri="https://zenodo.org/record/569120/files/figure.png" pageId="12" pageNumber="13">Figs 2324</figureCitation>
). Trochanter with a strong, thick, dorso-ventrally flattened, anterodistal projection. Femur with a moderate anterobasal projection and a dorsobasal mound; reticulation becoming strongly raised anteriorly, producing a tooth-like appearance in dorsal view. Patella with 1 large and 1 small lyrifissure dorsally near base, and 1 small dorsal lyrifissure apically. Chela with reticulate cuticle extending onto base of fixed finger, but not raised distally. Fixed finger with 4 lyrifissures: 1 antiaxial and 1 paraxial at base; 1 antiaxial behind trichobothria
<emphasis id="B90B310F657F3340FCC73A8D614AFB5D" box="[867,894,1177,1202]" italics="true" pageId="12" pageNumber="13">eb</emphasis>
and
<emphasis id="B90B310F657F3340FC173A8D61ECFB5D" box="[947,984,1177,1202]" italics="true" pageId="12" pageNumber="13">esb</emphasis>
; 1 dorsal, just in front of trichobothrium
<emphasis id="B90B310F657F3340FF333AAA6293FB3A" box="[151,167,1214,1237]" italics="true" pageId="12" pageNumber="13">it</emphasis>
. Movable finger with 2 ventral lyrifissures, one near middle and the other just in front of trichobothrium
<emphasis id="B90B310F657F3340FAE23AAA677AFB3A" box="[1350,1358,1214,1237]" italics="true" pageId="12" pageNumber="13">t</emphasis>
. Fixed finger with apodens plus 1516 marginal, 79 paraxioventral, 5 paraxial and 1 tuberculate basal-paraxial tooth (total 2931). Tuberculate tooth of fixed finger with an ordinary seta at base (
<figureCitation id="1344F198657F3340FC5B3B106667FAF2" box="[1023,1107,1284,1309]" captionStart="FIGURES 23 34" captionStartId="10.[151,264,1752,1775]" captionTargetBox="[232,1344,207,1715]" captionTargetId="figure@10.[224,1350,193,1724]" captionTargetPageId="10" captionText="FIGURES 23 34. Cybella deharvengi n. sp., male right palp. 23, chela, dorsal view; 24, palp minus chela, dorsal view (reticulate ornamentation drawn in part on femur); 25, tip of fixed chelal finger, dorsal view; 26, basal tuberculate tooth of fixed finger, dorsal view (relative to chela); 27, tip of movable chelal finger, paraxial view; 28, basal tuberculate tooth of movable chelal finger, ventral view (relative to chela), distal end to left; 29, chela, paraxial view (setae omitted); 30, chela, antiaxial view; 31, coupled sensilla; 32, distal sensillum; 33, distal end of movable finger, ventral view; 34, grouped chemosensory setae, antiaxial view. Abbreviations: af, apical sensillum of fixed finger; am 1, apical sensilla of movable finger; dx, duplex trichobothria; ld, probable lamina defensor; ly, lyrifissure; pc, coupled sensilla. Other abbreviations (eb, esb, est, et, ib, isb, ist, it, b, sb, st, t) are standard designations of trichobothria. Scale lines: 0.2 mm (Figs 23 24, 29), 0.1 mm (Fig. 30), 0.05 mm (Figs 25 28, 33 34), 0.01 mm (Figs 31 32)." httpUri="https://zenodo.org/record/569120/files/figure.png" pageId="12" pageNumber="13">Fig. 26</figureCitation>
), that of movable finger with a seta nearby, but not on tubercle (
<figureCitation id="1344F198657F3340FDB13B33605DFAAF" box="[533,617,1319,1344]" captionStart="FIGURES 23 34" captionStartId="10.[151,264,1752,1775]" captionTargetBox="[232,1344,207,1715]" captionTargetId="figure@10.[224,1350,193,1724]" captionTargetPageId="10" captionText="FIGURES 23 34. Cybella deharvengi n. sp., male right palp. 23, chela, dorsal view; 24, palp minus chela, dorsal view (reticulate ornamentation drawn in part on femur); 25, tip of fixed chelal finger, dorsal view; 26, basal tuberculate tooth of fixed finger, dorsal view (relative to chela); 27, tip of movable chelal finger, paraxial view; 28, basal tuberculate tooth of movable chelal finger, ventral view (relative to chela), distal end to left; 29, chela, paraxial view (setae omitted); 30, chela, antiaxial view; 31, coupled sensilla; 32, distal sensillum; 33, distal end of movable finger, ventral view; 34, grouped chemosensory setae, antiaxial view. Abbreviations: af, apical sensillum of fixed finger; am 1, apical sensilla of movable finger; dx, duplex trichobothria; ld, probable lamina defensor; ly, lyrifissure; pc, coupled sensilla. Other abbreviations (eb, esb, est, et, ib, isb, ist, it, b, sb, st, t) are standard designations of trichobothria. Scale lines: 0.2 mm (Figs 23 24, 29), 0.1 mm (Fig. 30), 0.05 mm (Figs 25 28, 33 34), 0.01 mm (Figs 31 32)." httpUri="https://zenodo.org/record/569120/files/figure.png" pageId="12" pageNumber="13">Fig. 28</figureCitation>
). Tuberculate tooth of movable finger in a more distal position relative to that of fixed finger (
<figureCitation id="1344F198657F3340FE273B5F63EEFA8C" box="[387,474,1355,1380]" captionStart="FIGURES 23 34" captionStartId="10.[151,264,1752,1775]" captionTargetBox="[232,1344,207,1715]" captionTargetId="figure@10.[224,1350,193,1724]" captionTargetPageId="10" captionText="FIGURES 23 34. Cybella deharvengi n. sp., male right palp. 23, chela, dorsal view; 24, palp minus chela, dorsal view (reticulate ornamentation drawn in part on femur); 25, tip of fixed chelal finger, dorsal view; 26, basal tuberculate tooth of fixed finger, dorsal view (relative to chela); 27, tip of movable chelal finger, paraxial view; 28, basal tuberculate tooth of movable chelal finger, ventral view (relative to chela), distal end to left; 29, chela, paraxial view (setae omitted); 30, chela, antiaxial view; 31, coupled sensilla; 32, distal sensillum; 33, distal end of movable finger, ventral view; 34, grouped chemosensory setae, antiaxial view. Abbreviations: af, apical sensillum of fixed finger; am 1, apical sensilla of movable finger; dx, duplex trichobothria; ld, probable lamina defensor; ly, lyrifissure; pc, coupled sensilla. Other abbreviations (eb, esb, est, et, ib, isb, ist, it, b, sb, st, t) are standard designations of trichobothria. Scale lines: 0.2 mm (Figs 23 24, 29), 0.1 mm (Fig. 30), 0.05 mm (Figs 25 28, 33 34), 0.01 mm (Figs 31 32)." httpUri="https://zenodo.org/record/569120/files/figure.png" pageId="12" pageNumber="13">Fig. 29</figureCitation>
). Fixed finger with a minute dorsal seta near apex (
<figureCitation id="1344F198657F3340FB953B5F66B3FA8B" box="[1073,1159,1355,1380]" captionStart="FIGURES 23 34" captionStartId="10.[151,264,1752,1775]" captionTargetBox="[232,1344,207,1715]" captionTargetId="figure@10.[224,1350,193,1724]" captionTargetPageId="10" captionText="FIGURES 23 34. Cybella deharvengi n. sp., male right palp. 23, chela, dorsal view; 24, palp minus chela, dorsal view (reticulate ornamentation drawn in part on femur); 25, tip of fixed chelal finger, dorsal view; 26, basal tuberculate tooth of fixed finger, dorsal view (relative to chela); 27, tip of movable chelal finger, paraxial view; 28, basal tuberculate tooth of movable chelal finger, ventral view (relative to chela), distal end to left; 29, chela, paraxial view (setae omitted); 30, chela, antiaxial view; 31, coupled sensilla; 32, distal sensillum; 33, distal end of movable finger, ventral view; 34, grouped chemosensory setae, antiaxial view. Abbreviations: af, apical sensillum of fixed finger; am 1, apical sensilla of movable finger; dx, duplex trichobothria; ld, probable lamina defensor; ly, lyrifissure; pc, coupled sensilla. Other abbreviations (eb, esb, est, et, ib, isb, ist, it, b, sb, st, t) are standard designations of trichobothria. Scale lines: 0.2 mm (Figs 23 24, 29), 0.1 mm (Fig. 30), 0.05 mm (Figs 25 28, 33 34), 0.01 mm (Figs 31 32)." httpUri="https://zenodo.org/record/569120/files/figure.png" pageId="12" pageNumber="13">Fig. 25</figureCitation>
), movable finger with a similar seta ventrally near apex (
<figureCitation id="1344F198657F3340FD913B7A60FFFA68" box="[565,715,1390,1415]" captionStart="FIGURES 23 34" captionStartId="10.[151,264,1752,1775]" captionTargetBox="[232,1344,207,1715]" captionTargetId="figure@10.[224,1350,193,1724]" captionTargetPageId="10" captionText="FIGURES 23 34. Cybella deharvengi n. sp., male right palp. 23, chela, dorsal view; 24, palp minus chela, dorsal view (reticulate ornamentation drawn in part on femur); 25, tip of fixed chelal finger, dorsal view; 26, basal tuberculate tooth of fixed finger, dorsal view (relative to chela); 27, tip of movable chelal finger, paraxial view; 28, basal tuberculate tooth of movable chelal finger, ventral view (relative to chela), distal end to left; 29, chela, paraxial view (setae omitted); 30, chela, antiaxial view; 31, coupled sensilla; 32, distal sensillum; 33, distal end of movable finger, ventral view; 34, grouped chemosensory setae, antiaxial view. Abbreviations: af, apical sensillum of fixed finger; am 1, apical sensilla of movable finger; dx, duplex trichobothria; ld, probable lamina defensor; ly, lyrifissure; pc, coupled sensilla. Other abbreviations (eb, esb, est, et, ib, isb, ist, it, b, sb, st, t) are standard designations of trichobothria. Scale lines: 0.2 mm (Figs 23 24, 29), 0.1 mm (Fig. 30), 0.05 mm (Figs 25 28, 33 34), 0.01 mm (Figs 31 32)." httpUri="https://zenodo.org/record/569120/files/figure.png" pageId="12" pageNumber="13">Figs 27, 33</figureCitation>
), these perhaps representing the
<emphasis id="B90B310F657F3340FBCE3B7A671EFA68" box="[1130,1322,1390,1415]" italics="true" pageId="12" pageNumber="13">lamina defensor</emphasis>
of other pseudoscorpions. Movable finger weakly S-shaped in ventral view; with 1 apical, 1820 marginal (almost aligned), 58 paraxial and 1 basal tuberculate tooth (total 2728). Sensillum
<emphasis id="B90B310F657F3340FC2B3BA26190FA21" box="[911,932,1462,1486]" italics="true" pageId="12" pageNumber="13">af</emphasis>
<quantity id="4C8740F8657F3340FC003BD061FEFA21" box="[932,970,1461,1490]" metricMagnitude="-2" metricUnit="m" metricValue="2.54" pageId="12" pageNumber="13" unit="in" value="1.0">1 in</quantity>
dorsal position, slightly closer to
<emphasis id="B90B310F657F3340FAEE3BA16751FA21" box="[1354,1381,1461,1486]" italics="true" pageId="12" pageNumber="13">dx</emphasis>
than to
<emphasis id="B90B310F657F3340FF173BC962F6FA1B" box="[179,194,1501,1524]" italics="true" pageId="12" pageNumber="13">it</emphasis>
. Movable finger with coupled sensilla (
<emphasis id="B90B310F657F3340FDDB3BC960ADFA1B" box="[639,665,1501,1524]" italics="true" pageId="12" pageNumber="13">pc</emphasis>
) above trichobothrium
<emphasis id="B90B310F657F3340FC053BCF619BFA1B" box="[929,943,1499,1524]" italics="true" pageId="12" pageNumber="13">b</emphasis>
(
<figureCitation id="1344F198657F3340FC193BCF6674FA1B" box="[957,1088,1499,1524]" captionStart="FIGURES 23 34" captionStartId="10.[151,264,1752,1775]" captionTargetBox="[232,1344,207,1715]" captionTargetId="figure@10.[224,1350,193,1724]" captionTargetPageId="10" captionText="FIGURES 23 34. Cybella deharvengi n. sp., male right palp. 23, chela, dorsal view; 24, palp minus chela, dorsal view (reticulate ornamentation drawn in part on femur); 25, tip of fixed chelal finger, dorsal view; 26, basal tuberculate tooth of fixed finger, dorsal view (relative to chela); 27, tip of movable chelal finger, paraxial view; 28, basal tuberculate tooth of movable chelal finger, ventral view (relative to chela), distal end to left; 29, chela, paraxial view (setae omitted); 30, chela, antiaxial view; 31, coupled sensilla; 32, distal sensillum; 33, distal end of movable finger, ventral view; 34, grouped chemosensory setae, antiaxial view. Abbreviations: af, apical sensillum of fixed finger; am 1, apical sensilla of movable finger; dx, duplex trichobothria; ld, probable lamina defensor; ly, lyrifissure; pc, coupled sensilla. Other abbreviations (eb, esb, est, et, ib, isb, ist, it, b, sb, st, t) are standard designations of trichobothria. Scale lines: 0.2 mm (Figs 23 24, 29), 0.1 mm (Fig. 30), 0.05 mm (Figs 25 28, 33 34), 0.01 mm (Figs 31 32)." httpUri="https://zenodo.org/record/569120/files/figure.png" pageId="12" pageNumber="13">Figs 3031</figureCitation>
); sensillum
<emphasis id="B90B310F657F3340FB613BC966D3FA1B" box="[1221,1255,1501,1524]" italics="true" pageId="12" pageNumber="13">am</emphasis>
1 slightly behind tip, in dorso-antiaxial position; a small pore or sensillum is also present ventrally, immediately in front of trichobothrium
<emphasis id="B90B310F657F3340FEE33832637BF9D2" box="[327,335,1574,1597]" italics="true" pageId="12" pageNumber="13">t</emphasis>
(
<figureCitation id="1344F198657F3340FEC438316381F9D1" box="[352,437,1573,1598]" captionStart="FIGURES 23 34" captionStartId="10.[151,264,1752,1775]" captionTargetBox="[232,1344,207,1715]" captionTargetId="figure@10.[224,1350,193,1724]" captionTargetPageId="10" captionText="FIGURES 23 34. Cybella deharvengi n. sp., male right palp. 23, chela, dorsal view; 24, palp minus chela, dorsal view (reticulate ornamentation drawn in part on femur); 25, tip of fixed chelal finger, dorsal view; 26, basal tuberculate tooth of fixed finger, dorsal view (relative to chela); 27, tip of movable chelal finger, paraxial view; 28, basal tuberculate tooth of movable chelal finger, ventral view (relative to chela), distal end to left; 29, chela, paraxial view (setae omitted); 30, chela, antiaxial view; 31, coupled sensilla; 32, distal sensillum; 33, distal end of movable finger, ventral view; 34, grouped chemosensory setae, antiaxial view. Abbreviations: af, apical sensillum of fixed finger; am 1, apical sensilla of movable finger; dx, duplex trichobothria; ld, probable lamina defensor; ly, lyrifissure; pc, coupled sensilla. Other abbreviations (eb, esb, est, et, ib, isb, ist, it, b, sb, st, t) are standard designations of trichobothria. Scale lines: 0.2 mm (Figs 23 24, 29), 0.1 mm (Fig. 30), 0.05 mm (Figs 25 28, 33 34), 0.01 mm (Figs 31 32)." httpUri="https://zenodo.org/record/569120/files/figure.png" pageId="12" pageNumber="13">Fig. 33</figureCitation>
). Fixed chelal finger with 13 chemosensory setae in antiaxial row. Movable finger with 1314 bacilliform setae (2024 µm long) in pit (
<figureCitation id="1344F198657F3340FD66385C617CF98E" box="[706,840,1608,1633]" captionStart="FIGURES 23 34" captionStartId="10.[151,264,1752,1775]" captionTargetBox="[232,1344,207,1715]" captionTargetId="figure@10.[224,1350,193,1724]" captionTargetPageId="10" captionText="FIGURES 23 34. Cybella deharvengi n. sp., male right palp. 23, chela, dorsal view; 24, palp minus chela, dorsal view (reticulate ornamentation drawn in part on femur); 25, tip of fixed chelal finger, dorsal view; 26, basal tuberculate tooth of fixed finger, dorsal view (relative to chela); 27, tip of movable chelal finger, paraxial view; 28, basal tuberculate tooth of movable chelal finger, ventral view (relative to chela), distal end to left; 29, chela, paraxial view (setae omitted); 30, chela, antiaxial view; 31, coupled sensilla; 32, distal sensillum; 33, distal end of movable finger, ventral view; 34, grouped chemosensory setae, antiaxial view. Abbreviations: af, apical sensillum of fixed finger; am 1, apical sensilla of movable finger; dx, duplex trichobothria; ld, probable lamina defensor; ly, lyrifissure; pc, coupled sensilla. Other abbreviations (eb, esb, est, et, ib, isb, ist, it, b, sb, st, t) are standard designations of trichobothria. Scale lines: 0.2 mm (Figs 23 24, 29), 0.1 mm (Fig. 30), 0.05 mm (Figs 25 28, 33 34), 0.01 mm (Figs 31 32)." httpUri="https://zenodo.org/record/569120/files/figure.png" pageId="12" pageNumber="13">Figs 30, 34</figureCitation>
); walls of pit finely granular (probably due to a thin lining of exocuticle). Trichobothria typical;
<emphasis id="B90B310F657F3340FD3C38786085F96A" box="[664,689,1644,1669]" italics="true" pageId="12" pageNumber="13">sb</emphasis>
and
<emphasis id="B90B310F657F3340FD54387960CCF96B" box="[752,760,1645,1668]" italics="true" pageId="12" pageNumber="13">t</emphasis>
of movable finger situated on ventral face,
<emphasis id="B90B310F657F3340FAA038786729F96A" box="[1284,1309,1644,1669]" italics="true" pageId="12" pageNumber="13">sb</emphasis>
level with posterior rim of pit containing chemosensory setae. Hair of trichobothrium
<date id="FFC1CBDD657F3340FC5638856633F943" box="[1010,1031,1681,1708]" pageId="12" pageNumber="13">
<emphasis id="B90B310F657F3340FC56388561CAF947" box="[1010,1022,1681,1704]" italics="true" pageId="12" pageNumber="13">x</emphasis>
2
</date>
thicker than that of
<date id="FFC1CBDD657F3340FB5538856731F943" box="[1265,1285,1681,1708]" pageId="12" pageNumber="13">
<emphasis id="B90B310F657F3340FB55388566C9F947" box="[1265,1277,1681,1704]" italics="true" pageId="12" pageNumber="13">x</emphasis>
1
</date>
and abruptly curved at tip; tip of
<date id="FFC1CBDD657F3340FED038A363BDF93C" box="[372,393,1719,1747]" pageId="12" pageNumber="13">
<emphasis id="B90B310F657F3340FED038A363B4F921" box="[372,384,1719,1742]" italics="true" pageId="12" pageNumber="13">x</emphasis>
1
</date>
blunt (very slightly bulbous).
</paragraph>
<paragraph id="8BC0ED1D657F3340FF6338C96167F826" blockId="12.[151,1437,1000,2029]" pageId="12" pageNumber="13">
Legs with reticulate-scaly ornamentation; very few gland pores present. Setae generally short and strongly curved, but those of ventral faces longer. Femur of leg I with a large, oblique, paraxio-basal lyrifissure. Patella of legs IIV with a dorsal lyrifissure orientated parallel to long axis and set in a depression. Trochanters of legs III and IV with a small posterodorsal spur just beyond pedicel; trochanters inflated, with maximum diameter oblique to dorso-ventral axis. Tarsus of leg I (but not other legs) with slight torsion distally, dorsal face orientated slightly anteriorly. Subterminal setae not obviously differentiated from other setae. Arolia simple, broad, distinctly shorter than claws; claws simple, not broadened distally (
<figureCitation id="1344F198657F3340FD6039A56173F826" box="[708,839,1969,1994]" captionStart="FIGURES 35 43" captionStartId="11.[151,264,1850,1873]" captionTargetBox="[159,1426,202,1817]" captionTargetId="figure@11.[151,1436,193,1830]" captionTargetPageId="11" captionText="FIGURES 35 43. Cybella deharvengi n. sp., male right legs. 35, leg I, antiaxial view; 36, trochanter and femur of leg I, dorsal view; 37, trochanter and femur of leg I, paraxial view; 38, leg IV, antiaxial view; 39, patella of leg I, dorsal view; 40, patella of leg IV, dorsal view; 41, leg III, antiaxial view; 42, arolium of leg III, ventral; 43 apotele of leg I, antiaxial view. Abbreviations: ly, lyrifissure;. Scale lines 0.2 mm (Figs 35 41), 0.05 (Figs 42 43)." httpUri="https://zenodo.org/record/569121/files/figure.png" pageId="12" pageNumber="13">Figs 4243</figureCitation>
).
</paragraph>
<paragraph id="8BC0ED1D657F3341FF6339C163D3FE1A" blockId="12.[151,1437,1000,2029]" lastBlockId="13.[151,1437,151,1763]" lastPageId="13" lastPageNumber="14" pageId="12" pageNumber="13">Measurements (in mm, standard ratios in parentheses). Body 1.8×1.21.3. Carapace 0.5460.619×0.4330.447 (1.31.4). Chelicera 0.2020.247×0.1280.154 (1.6), palm 0.1320.175 (1.1), movable finger 0.0950.114 (0.65×palm), spinneret 0.0180.023. Palp femur 0.4930.522×0.2640.281 (1.82.1), patella 0.3960.414×0.155 0.172 (2.42.6), chela 0.5550.571×0.1260.132 (4.34.5), palm 0.1420.148 (1.11.2), movable finger 0.401 0.413 (2.52.9). Chelicera 0.247×0.154 (1.6), palm 0.175 (1.1), movable finger 0.114 (0.65). Leg I (N=3) trochanter 0.150×0.110 (1.4), femur 0.2110.223×0.0750.81 (2.72.8), patella 0.1780.194×0.0880.092 (2.0 2.1), tibia 0.1650.190×0.0630.068 (2.63.0), tarsus 0.2560.263×0.0530.054 (4.84.9). Leg III (N=1) trochanter 0.196×0.112 (1.7), femur 0.212×0.088 (2.4), patella 0.247×0.103 (2.4), tibia 0.220×0.070 (3.2), tarsus 0.326×0.056 (5.9). Leg IV (N=3) trochanter 0.2510.264×0.1260.135 (1.92.0), femur 0.1610.188×0.0750.088 (2.02.2), patella 0.2560.275×0.0940.108 (2.42.7), tibia 0.3370.361×0.0590.066 (5.16.0), tarsus 0.325 0.386×0.0480.055 (6.87.2).</paragraph>
<paragraph id="8BC0ED1D657E3341FF633FEB637BFC4A" blockId="13.[151,1437,151,1763]" pageId="13" pageNumber="14">
<emphasis id="B90B310F657E3341FF633FEB603DFDF7" bold="true" box="[199,521,511,536]" pageId="13" pageNumber="14">Description of tritonymph.</emphasis>
Generally similar to adult and almost of same size, but less sclerotized and thus lighter in colour. Carapace with untanned dorsolateral lines running along its length, giving the impression that the carapace is divided into three plates (one dorsal and two lateral); posterior depression less marked than in adult. Anterolateral plates less sclerotized than in adult and hence less evident. Chelicera (
<figureCitation id="1344F198657E3341FBC63C786683FD6B" box="[1122,1207,620,645]" captionStart="FIGURES 44 47" captionStartId="12.[151,264,844,867]" captionTargetBox="[156,1429,202,816]" captionTargetId="figure@12.[151,1436,193,823]" captionTargetPageId="12" captionText="FIGURES 44 47. Cybella deharvengi n. sp., paratype tritonymph. 44, chelicera, antiaxial view; 45, tip of movable cheliceral finger, antiaxial view; 46, right chela, antiaxial view; 47 grouped chemosensory setae, antiaxial view. Abbreviations: am 1, apical sensillum of movable finger; pr, distal process of serrula exterior; pc, coupled sensilla. Scale lines 0.1 mm (Figs 44, 46), 0.05 mm (Fig. 45), 0.02 mm (Fig. 47)." httpUri="https://zenodo.org/record/569122/files/figure.png" pageId="13" pageNumber="14">Fig. 44</figureCitation>
) with 5 large and 5 small setae on palm; galeal seta 0.78 from base; spinneret (
<figureCitation id="1344F198657E3341FCE83C9B6195FD47" box="[844,929,655,680]" captionStart="FIGURES 44 47" captionStartId="12.[151,264,844,867]" captionTargetBox="[156,1429,202,816]" captionTargetId="figure@12.[151,1436,193,823]" captionTargetPageId="12" captionText="FIGURES 44 47. Cybella deharvengi n. sp., paratype tritonymph. 44, chelicera, antiaxial view; 45, tip of movable cheliceral finger, antiaxial view; 46, right chela, antiaxial view; 47 grouped chemosensory setae, antiaxial view. Abbreviations: am 1, apical sensillum of movable finger; pr, distal process of serrula exterior; pc, coupled sensilla. Scale lines 0.1 mm (Figs 44, 46), 0.05 mm (Fig. 45), 0.02 mm (Fig. 47)." httpUri="https://zenodo.org/record/569122/files/figure.png" pageId="13" pageNumber="14">Fig. 45</figureCitation>
) distinctly longer than in male, tubular and gently curved ventrally, base flared in dorsoventral view, but not in lateral view; rallum with two blades (anterior 23 µm, posterior 22 µm long), as in adult; serrula exterior with 19 blades plus anteriorly-directed process, distal 3 blades; serrula interior obscured. Fixed finger of chela with apodens plus 18 marginal teeth, 6 paraxial teeth and a basal tuberculate tooth (total 26), movable finger with apodens plus 17 teeth in slightly irregular marginal row, 7 paraxial teeth and basal tuberculate tooth (total 26). Movable finger with 13 bacilliform chemosensory setae in pit (
<figureCitation id="1344F198657E3341FF3B3D7362C0FC6F" box="[159,244,871,896]" captionStart="FIGURES 44 47" captionStartId="12.[151,264,844,867]" captionTargetBox="[156,1429,202,816]" captionTargetId="figure@12.[151,1436,193,823]" captionTargetPageId="12" captionText="FIGURES 44 47. Cybella deharvengi n. sp., paratype tritonymph. 44, chelicera, antiaxial view; 45, tip of movable cheliceral finger, antiaxial view; 46, right chela, antiaxial view; 47 grouped chemosensory setae, antiaxial view. Abbreviations: am 1, apical sensillum of movable finger; pr, distal process of serrula exterior; pc, coupled sensilla. Scale lines 0.1 mm (Figs 44, 46), 0.05 mm (Fig. 45), 0.02 mm (Fig. 47)." httpUri="https://zenodo.org/record/569122/files/figure.png" pageId="13" pageNumber="14">Fig. 47</figureCitation>
), surface of pit granular on left chela, but not on right chela. Trichobothria as in adult, except
<emphasis id="B90B310F657E3341FA8D3D73677EFC6F" box="[1321,1354,871,896]" italics="true" pageId="13" pageNumber="14">isb</emphasis>
and
<emphasis id="B90B310F657E3341FA273D7367A8FC6F" box="[1411,1436,871,896]" italics="true" pageId="13" pageNumber="14">sb</emphasis>
absent (
<figureCitation id="1344F198657E3341FF4A3D986376FC4A" box="[238,322,908,933]" captionStart="FIGURES 44 47" captionStartId="12.[151,264,844,867]" captionTargetBox="[156,1429,202,816]" captionTargetId="figure@12.[151,1436,193,823]" captionTargetPageId="12" captionText="FIGURES 44 47. Cybella deharvengi n. sp., paratype tritonymph. 44, chelicera, antiaxial view; 45, tip of movable cheliceral finger, antiaxial view; 46, right chela, antiaxial view; 47 grouped chemosensory setae, antiaxial view. Abbreviations: am 1, apical sensillum of movable finger; pr, distal process of serrula exterior; pc, coupled sensilla. Scale lines 0.1 mm (Figs 44, 46), 0.05 mm (Fig. 45), 0.02 mm (Fig. 47)." httpUri="https://zenodo.org/record/569122/files/figure.png" pageId="13" pageNumber="14">Fig. 46</figureCitation>
).
</paragraph>
<paragraph id="8BC0ED1D657E3341FF633DA46385FB92" blockId="13.[151,1437,151,1763]" pageId="13" pageNumber="14">
Measurements of
<typeStatus id="54C453BF657E3341FE343DA463C4FC27" box="[400,496,944,968]" pageId="13" pageNumber="14" type="paratype">paratype</typeStatus>
tritonymph (in mm, standard ratios in parentheses). Body 1.80×1.1 (1.5). Carapace 0.533×0.416 (1.3). Chelicera 0.182×0.119 (1.5), palm 0.136 (1.15), movable finger 0.087 (0.64). Palp femur 0.459×0.241 (1.9), patella 0.355×0.149 (2.4), chela 0.539×0.126 (4.3), palm 0.138 (1.1), movable finger 0.378 (2.8). Leg I femur 0.192×0.070 (2.8), patella 0.167×0.081 (2.1), tibia 0.159×0.059 (2.7), tarsus 0.234×0.051 (4.6). Leg IV trochanter 0.232×0.121 (1.9), femur 0.161×0.075 (2.1), patella 0.242×0.095 (2.5), tibia 0.304×0.095 (3.2), tarsus 0.344×0.055 (6.3).
</paragraph>
<paragraph id="8BC0ED1D657E3341FF633A9D60AFFB07" blockId="13.[151,1437,151,1763]" pageId="13" pageNumber="14">Measurements of smaller (non-type) tritonymph (in mm, standard ratios in parentheses). Body 1.48×0.96 (1.5). Carapace 0.463×0.370 (1.3). Palp femur 0.394×0.203 (1.9), patella 0.310×0.136 (2.3), chela 0.501×0.110 (4.6), palm 0.122 (1.1), movable finger 0.0.34 (2.8).</paragraph>
</subSubSection>
<subSubSection id="C365BE96657E3341FF633AE06689F90C" pageId="13" pageNumber="14" type="discussion">
<paragraph id="8BC0ED1D657E3341FF633AE066ADF9BD" blockId="13.[151,1437,151,1763]" pageId="13" pageNumber="14">
<emphasis id="B90B310F657E3341FF633AE0630FFAE2" bold="true" box="[199,315,1268,1293]" pageId="13" pageNumber="14">Remarks.</emphasis>
The cavity in which the
<typeStatus id="54C453BF657E3341FDD53AE16098FAE2" box="[625,684,1269,1293]" pageId="13" pageNumber="14">types</typeStatus>
were found was accessible by a vertical opening that was just sufficient for a person to slide through. The feaellids were found on a small, exposed surface (no more than
<quantity id="4C8740F8657E3341FAFB3B0F67BEFADD" box="[1375,1418,1307,1331]" metricMagnitude="0" metricUnit="m" metricValue="1.0" pageId="13" pageNumber="14" unit="m" value="1.0">1 m</quantity>
2) on a wall facing away from the entrance. This surface differed from the rest of the wall in having a pocked appearance and a dark, rust-like colour (
<figureCitation id="1344F198657E3341FDC43B766097FA94" box="[608,675,1378,1403]" captionStart="FIGURES 1 2" captionStartId="5.[151,264,1190,1213]" captionTargetBox="[151,1436,519,1166]" captionTargetId="figure@5.[151,1436,519,1166]" captionTargetPageId="5" captionText="FIGURES 1 2. Cybella deharvengi n. sp. 1, area of cavity wall on which the types were found (photograph L. Deharveng); 2, habitus of male, dorsal view." httpUri="https://zenodo.org/record/569115/files/figure.png" pageId="13" pageNumber="14">Fig. 1</figureCitation>
), presumably due to a metal oxide. It was humid, but not wet. The rest of the walls of the cavity, most of which were wet, were not found to have feaellids on them. The pseudoscorpions were near the entrance and would have been in permanent obscurity, but not total darkness. The floor of the cavity consisted of compact clay-soil, unsuitable for pseudoscorpions. The collecting details accompanying the non-type tritonymph from leaf litter do not specify whether it came from a cavity or a truly epigean habitat. However, given the large number of litter samples made by Le Cong Man, L. Deharveng and A. Bedos at Nui Bai Voi, it seems likely that this species has a preference for cavities or caves.
</paragraph>
<paragraph id="8BC0ED1D657E3341FF63384A6689F90C" blockId="13.[151,1437,151,1763]" pageId="13" pageNumber="14">The identification of the non-type tritonymph is discussed below. The collection details for this specimen suggest that it was found in an epigean habitat, although the possibility that it came from one of the many cavities around the Grotte-hôpital cannot be excluded. Sifting of litter (by the author) and Berlese extractions of litter outside the cavity (by Louis Deharveng and Anne Bedos) did not produced any other feaellids.</paragraph>
</subSubSection>
</treatment>
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