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<document id="92856C3E76BC294B43FC33D281C0DD77" ID="10.11646/zootaxa.4258.3.1SLASH10916" ID-CLB-Dataset="33146" ID-DOI="10.11646/zootaxa.4258.3.1" ID-GBIF-Dataset="97942778-cef4-4cf6-b4e4-c077e633daa8" ID-ISSN="1175-5326" ID-Zenodo-Dep="569729" ID-ZooBank="DE89E162-B255-474C-A824-20DFA551DBBD" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1493363817835" checkinUser="plazi" docAuthor="Penz, Carla M., Casagrande, Mirna M., Devries, Phil &amp; Simonsen, Thomas J." docDate="2017" docId="03BB87F4FFA6FF83E1FF23A4BB94F83C" docLanguage="en" docName="zootaxa.4258.3.1SLASH10916.pdf" docOrigin="Zootaxa 4258 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Bia decaerulea Weymer, STAT. NOV." docType="treatment" docVersion="7" lastPageNumber="211" masterDocId="FF82FF8CFFACFF89E1682278BA7DFFFC" masterDocTitle="Documenting diversity in the Amazonian butterfly genus Bia (Lepidoptera, Nymphalidae)" masterLastPageNumber="237" masterPageNumber="201" pageNumber="211" updateTime="1698439134094" updateUser="ExternalLinkService">
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<mods:title id="394621E6B962F415991125AF3B1D0B04">Documenting diversity in the Amazonian butterfly genus Bia (Lepidoptera, Nymphalidae)</mods:title>
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<mods:namePart id="A7B274B1287A0895B3703BFD14036AF9">Penz, Carla M.</mods:namePart>
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<mods:namePart id="5EB8A40BBB3416F43FF53E67A5D0D450">Casagrande, Mirna M.</mods:namePart>
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<mods:namePart id="54851232B3B8D10D7AB254339F550C88">Devries, Phil</mods:namePart>
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<mods:namePart id="47669DEC55B5D6710F60AC4A6705E582">Simonsen, Thomas J.</mods:namePart>
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<treatment id="03BB87F4FFA6FF83E1FF23A4BB94F83C" ID-DOI="http://doi.org/10.5281/zenodo.6017377" ID-GBIF-Taxon="128810411" ID-Zenodo-Dep="6017377" LSID="urn:lsid:plazi:treatment:03BB87F4FFA6FF83E1FF23A4BB94F83C" httpUri="http://treatment.plazi.org/id/03BB87F4FFA6FF83E1FF23A4BB94F83C" lastPageNumber="211" pageId="10" pageNumber="211">
<subSubSection id="C3086569FFA6FF83E1FF23A4BB32FDEB" pageId="10" pageNumber="211" type="nomenclature">
<paragraph id="8BAD36E2FFA6FF83E1FF23A4B81EFE0A" blockId="10.[151,611,476,535]" box="[151,611,476,502]" pageId="10" pageNumber="211">
<heading id="D0E5818EFFA6FF83E1FF23A4B81EFE0A" bold="true" box="[151,611,476,502]" fontSize="11" level="1" pageId="10" pageNumber="211" reason="1">
<emphasis id="B966EAF0FFA6FF83E1FF23A4B81EFE0A" bold="true" box="[151,611,476,502]" pageId="10" pageNumber="211">
<taxonomicName id="4C124D61FFA6FF83E1FF23A4BBC0FE0A" authority="Weymer" authorityName="Weymer" box="[151,445,476,502]" class="Insecta" family="Nymphalidae" genus="Bia" kingdom="Animalia" order="Lepidoptera" pageId="10" pageNumber="211" phylum="Arthropoda" rank="species" species="decaerulea" status="stat. nov.">
<emphasis id="B966EAF0FFA6FF83E1FF23A4BB31FE0A" bold="true" box="[151,332,476,502]" italics="true" pageId="10" pageNumber="211">Bia decaerulea</emphasis>
Weymer
</taxonomicName>
,
<taxonomicNameLabel id="A255578BFFA6FF83E0A123A4B81EFE0A" box="[457,611,476,502]" pageId="10" pageNumber="211">STAT. NOV.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BAD36E2FFA6FF83E1FF2386BB32FDEB" blockId="10.[151,611,476,535]" box="[151,335,510,535]" pageId="10" pageNumber="211">
(
<figureCitation id="13292A67FFA6FF83E1F72386BA9AFDEB" box="[159,231,510,535]" captionStart="FIGURE 4" captionStartId="11.[151,250,1848,1870]" captionTargetBox="[262,1339,234,1759]" captionTargetId="figure@11.[209,1367,195,1798]" captionTargetPageId="11" captionText="FIGURE 4. Adults in dorsal and ventral views. Taxon names appear below each set. Bia decaerulea decaerulea: a, male, Brazil, Amazonas, Manacapuru (CMNH); b, female, same locality (CMNH). Bia decaerulea cayana, Simonsen &amp; Penz, new ssp.: c, male holotype, French Guiana, Mana River (CMNH); d, male paratype, same locality (CMNH); e, male paratype, same locality (CMNH); f, female paratype, French Guiana, Oyapok River (CMNH). Bia decaerulea pallida Casagrande &amp; Penz, new ssp: g, male holotype, Brazil, Amazonas, Barcelos, Rio Padauari (DZUP); h, female paratype, Brazil, Amazonas, Barcelos, Rio Padauari (DZUP)." httpUri="https://zenodo.org/record/569733/files/figure.png" pageId="10" pageNumber="211">Figs 4</figureCitation>
,
<figureCitation id="13292A67FFA6FF83E19B2387BB72FDEB" box="[243,271,511,535]" captionStart="FIGURE 11" captionStartId="25.[151,250,1869,1891]" captionTargetBox="[184,1395,202,1838]" captionTargetId="figure@25.[174,1410,193,1842]" captionTargetPageId="25" captionText="FIGURE 11. Genitalia. a, male actorion, Guyana (dissection Penz 12 - 01, CMNH); b, female actorion, Guyana (dissection Penz 10 - 02, LACM; scale bar different from male illustrations); c, male decaerulea, Lectotype, Brazil, Pará, Faro, (dissection Miller 9047, MfN); d, male rebeli, Peru, San Martín (dissection Penz 10 - 08, USNM); e, male caelestis, paratype, Brazil, Amazonas, São Paulo de Olivença (dissection Penz 10 - 29, CMNH); f, male pucallpa, Peru, Pucallpa (USUSNM 2041410); g, male peruana, Peru, Huanuco, Rio Pichis (dissection Penz 09 - 06, FLMNH)." httpUri="https://zenodo.org/record/569741/files/figure.png" pageId="10" pageNumber="211">11</figureCitation>
c, 12)
</paragraph>
</subSubSection>
<subSubSection id="C3086569FFA6FF83E1FF203EBE48FB13" pageId="10" pageNumber="211" type="description">
<paragraph id="8BAD36E2FFA6FF83E1FF203EBE48FB13" blockId="10.[151,1437,582,1984]" pageId="10" pageNumber="211">
<materialsCitation id="3B7A3CBFFFA6FF83E1FF203EBFE5FD13" ID-GBIF-Occurrence="1499315712" country="Brazil" location="Amazonas" pageId="10" pageNumber="211" specimenCount="1" stateProvince="Amazonas">
<emphasis id="B966EAF0FFA6FF83E1FF203EBBACFDA3" bold="true" box="[151,465,582,607]" pageId="10" pageNumber="211">Diagnostic re-description.</emphasis>
General characters as described for the genus. (1) MF DFW white apical ocelli medium-large. (2) MF DFW orange band somewhat diffuse, generally wide but varying in width, narrower in
<taxonomicName id="4C124D61FFA6FF83E1FF20F6BB10FD5A" box="[151,365,654,678]" class="Insecta" family="Nymphalidae" genus="Bia" kingdom="Animalia" order="Lepidoptera" pageId="10" pageNumber="211" phylum="Arthropoda" rank="subSpecies" species="decaerulea" subSpecies="pallida">
<emphasis id="B966EAF0FFA6FF83E1FF20F6BB10FD5A" box="[151,365,654,678]" italics="true" pageId="10" pageNumber="211">decaerulea pallida</emphasis>
</taxonomicName>
(
<collectingCountry id="F3057672FFA6FF83E0E820F6BBB4FD5B" box="[384,457,654,679]" name="Brazil" pageId="10" pageNumber="211">Brazil</collectingCountry>
,
<collectingRegion id="49D6F800FFA6FF83E0B020F7B82CFD5B" box="[472,593,655,679]" country="Colombia" name="Amazonas" pageId="10" pageNumber="211">Amazonas</collectingRegion>
, Barcelos and nearby localities;
<collectingCountry id="F3057672FFA6FF83E2B920F6BE39FD5B" box="[977,1092,654,679]" name="Colombia" pageId="10" pageNumber="211">Colombia</collectingCountry>
), intermediate in the nominal subspecies (
<collectingCountry id="F3057672FFA6FF83E07520CABB1BFD37" box="[285,358,690,715]" name="Brazil" pageId="10" pageNumber="211">Brazil</collectingCountry>
,
<collectingRegion id="49D6F800FFA6FF83E01920CBBB97FD37" box="[369,490,691,715]" country="Colombia" name="Amazonas" pageId="10" pageNumber="211">Amazonas</collectingRegion>
, Manacapuru;
<collectingCountry id="F3057672FFA6FF83E3E720CBB897FD37" box="[655,746,691,715]" name="Guyana" pageId="10" pageNumber="211">Guyana</collectingCountry>
;
<collectingCountry id="F3057672FFA6FF83E39D20CAB91FFD37" box="[757,866,690,715]" name="Suriname" pageId="10" pageNumber="211">Suriname</collectingCountry>
) and very wide in
<taxonomicName id="4C124D61FFA6FF83E55920CBBF78FD36" box="[1073,1285,691,715]" class="Insecta" family="Nymphalidae" genus="Bia" kingdom="Animalia" order="Lepidoptera" pageId="10" pageNumber="211" phylum="Arthropoda" rank="subSpecies" species="decaerulea" subSpecies="cayana">
<emphasis id="B966EAF0FFA6FF83E55920CBBF78FD36" box="[1073,1285,691,715]" italics="true" pageId="10" pageNumber="211">decaerulea cayana</emphasis>
</taxonomicName>
, reaching the distal edge of the discal cell and sometimes spreading into the discal cell (
<collectingCountry id="F3057672FFA6FF83E56820AEBED3FD13" box="[1024,1198,726,751]" name="French Guiana" pageId="10" pageNumber="211">French Guiana</collectingCountry>
;
<collectingCountry id="F3057672FFA6FF83E5D620AEBF78FD13" box="[1214,1285,726,751]" name="Brazil" pageId="10" pageNumber="211">Brazil</collectingCountry>
, north
<collectingRegion id="49D6F800FFA6FF83E43120AFBFEDFD13" box="[1369,1424,727,751]" country="Suriname" name="Para" pageId="10" pageNumber="211">Pará</collectingRegion>
)
</materialsCitation>
;
<materialsCitation id="3B7A3CBFFFA6FF83E1FF2083B82DFC0D" ID-GBIF-Occurrence="1499315588" country="Brazil" location="French Guiana region" pageId="10" pageNumber="211" specimenCount="1" stateProvince="French Guiana region">
orange scales broadly extended proximally along veins, often lining cross-veins at the distal edge of discal cell. (3) M DFW iridescent band generally absent, but the location of the band might be visible under the scope when the specimen is tilted (a sheen is visible). F DFW blue iridescence well developed but variable; usually extended across the entire DC and diffuse anteriorly in both subspecies; usually visible at the CuA1-CuA2 intersection in both subspecies; below CuA2 the iridescent area is wide and may spread towards the tornus to meet the edge of the orange band in
<taxonomicName id="4C124D61FFA6FF83E02B21CCB86BFC30" box="[323,534,948,972]" class="Insecta" family="Nymphalidae" genus="Bia" kingdom="Animalia" order="Lepidoptera" pageId="10" pageNumber="211" phylum="Arthropoda" rank="subSpecies" species="decaerulea" subSpecies="cayana">
<emphasis id="B966EAF0FFA6FF83E02B21CCB86BFC30" box="[323,534,948,972]" italics="true" pageId="10" pageNumber="211">decaerulea cayana</emphasis>
</taxonomicName>
. F DFW iridescent band smaller than in
<taxonomicName id="4C124D61FFA6FF83E2B321CDBE24FC30" box="[987,1113,949,972]" class="Insecta" family="Nymphalidae" genus="Bia" kingdom="Animalia" order="Lepidoptera" pageId="10" pageNumber="211" phylum="Arthropoda" rank="species" species="actorion">
<emphasis id="B966EAF0FFA6FF83E2B321CDBE24FC30" box="[987,1113,949,972]" italics="true" pageId="10" pageNumber="211">B. actorion</emphasis>
</taxonomicName>
in most localities, but just as large in others (
<collectingRegion id="49D6F800FFA6FF83E02221A0B834FC0D" box="[330,585,984,1009]" country="France" name="Guyane" pageId="10" pageNumber="211">
<collectingRegion id="49D6F800FFA6FF83E02221A0BB8EFC0D" box="[330,499,984,1009]" inferredBySuffix="true" pageId="10" pageNumber="211">French Guiana</collectingRegion>
region
</collectingRegion>
)
</materialsCitation>
. (4) M DFW androconial scales on the CuA-CuA1-CuA2 intersection dark brown, matching scale color of surrounding area and intermingled with orange scales. (5) M DHW discal androconial pad dark brown, nearly black, always darker than associated hairpencil. (6) M DHW discal hairpencil dark brown, but lighter in color than discal androconial pad. (7) F VFW ripple pattern less dense than that of M, with the VFW postmedial area showing a more predominantly yellow color and with one or two pale markings sometimes present across DC. Male genitalia (
<figureCitation id="13292A67FFA6FF83E3CD26F6B890FB5B" box="[677,749,1166,1191]" captionStart="FIGURE 2" captionStartId="4.[151,250,1593,1615]" captionTargetBox="[173,1417,207,1563]" captionTargetId="figure@4.[151,1436,193,1572]" captionTargetPageId="4" captionText="FIGURE 2. Variation in valva and gnathos morphology (lateral view) across the range of Bia, including a sample of various species and subspecies. Note intraspecific variation in series from the same locality outlined by boxes." httpUri="https://zenodo.org/record/569731/files/figure.png" pageId="10" pageNumber="211">Figs 2</figureCitation>
and
<figureCitation id="13292A67FFA6FF83E24D26F7B93FFB5B" box="[805,834,1167,1191]" captionStart="FIGURE 11" captionStartId="25.[151,250,1869,1891]" captionTargetBox="[184,1395,202,1838]" captionTargetId="figure@25.[174,1410,193,1842]" captionTargetPageId="25" captionText="FIGURE 11. Genitalia. a, male actorion, Guyana (dissection Penz 12 - 01, CMNH); b, female actorion, Guyana (dissection Penz 10 - 02, LACM; scale bar different from male illustrations); c, male decaerulea, Lectotype, Brazil, Pará, Faro, (dissection Miller 9047, MfN); d, male rebeli, Peru, San Martín (dissection Penz 10 - 08, USNM); e, male caelestis, paratype, Brazil, Amazonas, São Paulo de Olivença (dissection Penz 10 - 29, CMNH); f, male pucallpa, Peru, Pucallpa (USUSNM 2041410); g, male peruana, Peru, Huanuco, Rio Pichis (dissection Penz 09 - 06, FLMNH)." httpUri="https://zenodo.org/record/569741/files/figure.png" pageId="10" pageNumber="211">11</figureCitation>
c): tip of valva often truncated, varying in height, but sometimes narrow, spiny projection medium-tall, spines larger in specimens from the Guyana-Suriname-French Guiana region; gnathos broad (aforementioned region) to narrow (other localities).
</paragraph>
</subSubSection>
<subSubSection id="C3086569FFA6FF83E1AF2683BE0FF884" pageId="10" pageNumber="211" type="materials_examined">
<paragraph id="8BAD36E2FFA6FF83E1AF2683BE0FF884" blockId="10.[151,1437,582,1984]" pageId="10" pageNumber="211">
<materialsCitation id="3B7A3CBFFFA6FF83E1AF2683BE66F8A8" ID-GBIF-Occurrence="1499315721" collectingDate="2016-08-30" collectionCode="MfN" collectorName="R. I. Vane-Wright &amp; Paul Hahnel &amp; Hahnel &amp; The &amp; Faro &amp; Hnl &amp; Lee D." country="Brazil" county="The" location="The" municipality="Weymer" pageId="10" pageNumber="211" specimenCount="8" stateProvince="Para" typeStatus="lectotype">
<emphasis id="B966EAF0FFA6FF83E1AF2683BB05FAE8" bold="true" box="[199,376,1275,1300]" pageId="10" pageNumber="211">
<typeStatus id="54A98840FFA6FF83E1AF2683BB7FFAE8" box="[199,258,1275,1300]" pageId="10" pageNumber="211">Type</typeStatus>
material.
</emphasis>
<collectingCounty id="62CC4E6EFFA6FF83E0E92683BBD0FAE8" box="[385,429,1275,1300]" pageId="10" pageNumber="211">The</collectingCounty>
description by
<bibRefCitation id="EF834B13FFA6FF83E30A2684B969FAEF" author="Weymer" box="[610,788,1275,1300]" pageId="10" pageNumber="211" refString="Weymer, G. (1911) Family, Satyridae. In: Seitz, A. (Ed.), Die Gross-Schmetterlinge der erde. Vol. 5. Alfred Kernen, Stuttgart, pp. 173 - 283." type="book" year="1911">
<collectingMunicipality id="6BC9AC98FFA6FF83E30A2684B8BDFAE8" box="[610,704,1276,1300]" pageId="10" pageNumber="211">Weymer</collectingMunicipality>
(1911)
</bibRefCitation>
reads as follows: “
<location id="8ECD6039FFA6FF83E29C2683BE5CFAE8" LSID="urn:lsid:plazi:treatment:03BB87F4FFA6FF83E1FF23A4BB94F83C:8ECD6039FFA6FF83E29C2683BE5CFAE8" box="[1012,1057,1275,1300]" country="Brazil" county="The" municipality="Weymer" name="The" pageId="10" pageNumber="211" stateProvince="Para">The</location>
form
<taxonomicName id="4C124D61FFA6FF83E5042683BF5BFAEF" authority="Stgr." authorityName="Stgr." box="[1132,1318,1275,1299]" class="Insecta" family="Nymphalidae" genus="Bia" kingdom="Animalia" order="Lepidoptera" pageId="10" pageNumber="211" phylum="Arthropoda" rank="species" species="decaerulea">
<emphasis id="B966EAF0FFA6FF83E5042683BE94FAEF" box="[1132,1257,1275,1299]" italics="true" pageId="10" pageNumber="211">decaerulea</emphasis>
Stgr.
</taxonomicName>
i.l. (60 e) shows no trace of blue reflection, on the other hand the brownish yellow band is somewhat broader.
<location id="8ECD6039FFA6FF83E45F2758BF35FACB" LSID="urn:lsid:plazi:treatment:03BB87F4FFA6FF83E1FF23A4BB94F83C:8ECD6039FFA6FF83E45F2758BF35FACB" box="[1335,1352,1312,1335]" country="Brazil" county="The" municipality="Weymer" name="It" pageId="10" pageNumber="211" stateProvince="Para">It</location>
occurs among the
<typeStatus id="54A98840FFA6FF83E076273CBB33FAA0" box="[286,334,1348,1372]" pageId="10" pageNumber="211">type</typeStatus>
form.”
<location id="8ECD6039FFA6FF83E0D8273BBBA1FAA0" LSID="urn:lsid:plazi:treatment:03BB87F4FFA6FF83E1FF23A4BB94F83C:8ECD6039FFA6FF83E0D8273BBBA1FAA0" box="[432,476,1347,1372]" country="Brazil" county="The" municipality="Weymer" name="The" pageId="10" pageNumber="211" stateProvince="Para">The</location>
mention of a letter from
<location id="8ECD6039FFA6FF83E27D273BB9F3FAA0" LSID="urn:lsid:plazi:treatment:03BB87F4FFA6FF83E1FF23A4BB94F83C:8ECD6039FFA6FF83E27D273BB9F3FAA0" box="[789,910,1347,1372]" country="Brazil" county="The" municipality="Weymer" name="Staudinger" pageId="10" pageNumber="211" stateProvince="Para">Staudinger</location>
suggests that
<typeStatus id="54A98840FFA6FF83E554273CBE11FAA0" box="[1084,1132,1348,1372]" pageId="10" pageNumber="211">type</typeStatus>
material would be in his collection, currently housed at
<collectionCode id="ED03AE27FFA6FF83E094271FB84BFA7C" box="[508,566,1383,1408]" country="Germany" httpUri="http://grbio.org/cool/jp8y-i5nv" name="Museum f<>r Naturkunde" pageId="10" pageNumber="211">MfN</collectionCode>
.
<location id="8ECD6039FFA6FF83E32B271FB8B2FA7C" LSID="urn:lsid:plazi:treatment:03BB87F4FFA6FF83E1FF23A4BB94F83C:8ECD6039FFA6FF83E32B271FB8B2FA7C" box="[579,719,1383,1408]" country="Brazil" county="The" municipality="Weymer" name="Nonetheless" pageId="10" pageNumber="211" stateProvince="Para">Nonetheless</location>
,
<typeStatus id="54A98840FFA6FF83E3B42710B96AFA7C" box="[732,791,1384,1408]" pageId="10" pageNumber="211">types</typeStatus>
for this taxon were not found.
<location id="8ECD6039FFA6FF83E5ED271FBE92FA7C" LSID="urn:lsid:plazi:treatment:03BB87F4FFA6FF83E1FF23A4BB94F83C:8ECD6039FFA6FF83E5ED271FBE92FA7C" box="[1157,1263,1383,1408]" country="Brazil" county="The" municipality="Weymer" name="Although" pageId="10" pageNumber="211" stateProvince="Para">Although</location>
a
<typeStatus id="54A98840FFA6FF83E4662710BF43FA7C" box="[1294,1342,1384,1408]" pageId="10" pageNumber="211">type</typeStatus>
locality cannot be established with certainty, in 1969
<collectorName id="26E75334FFA6FF83E3FB27F4B929FA58" box="[659,852,1419,1444]" pageId="10" pageNumber="211">R.I. Vane-Wright</collectorName>
selected and labeled a male from
<collectingCountry id="F3057672FFA6FF83E5BE27F3BF60FA58" box="[1238,1309,1419,1444]" name="Brazil" pageId="10" pageNumber="211">Brazil</collectingCountry>
,
<collectingRegion id="49D6F800FFA6FF83E44127F3BF23FA5F" box="[1321,1374,1419,1443]" country="Suriname" name="Para" pageId="10" pageNumber="211">Pará</collectingRegion>
,
<location id="8ECD6039FFA6FF83E40127F4BFE1FA5F" LSID="urn:lsid:plazi:treatment:03BB87F4FFA6FF83E1FF23A4BB94F83C:8ECD6039FFA6FF83E40127F4BFE1FA5F" box="[1385,1436,1420,1443]" country="Brazil" county="The" municipality="Weymer" name="Faro" pageId="10" pageNumber="211" stateProvince="Para">Faro</location>
as
<typeStatus id="54A98840FFA6FF83E1DF27D7BB60FA34" box="[183,285,1455,1480]" pageId="10" pageNumber="211" type="lectotype">lectotype</typeStatus>
of
<taxonomicName id="4C124D61FFA6FF83E02D27C8BBB8FA3B" box="[325,453,1456,1479]" class="Insecta" family="Nymphalidae" genus="Bia" kingdom="Animalia" order="Lepidoptera" pageId="10" pageNumber="211" phylum="Arthropoda" rank="species" species="actorion">
<emphasis id="B966EAF0FFA6FF83E02D27C8BBB8FA3B" box="[325,453,1456,1479]" italics="true" pageId="10" pageNumber="211">B. actorion</emphasis>
</taxonomicName>
form
<taxonomicName id="4C124D61FFA6FF83E36427D7B8F5FA3B" box="[524,648,1455,1479]" class="Insecta" family="Nymphalidae" genus="Bia" kingdom="Animalia" order="Lepidoptera" pageId="10" pageNumber="211" phylum="Arthropoda" rank="species" species="decaerulea">
<emphasis id="B966EAF0FFA6FF83E36427D7B8F5FA3B" box="[524,648,1455,1479]" italics="true" pageId="10" pageNumber="211">decaerulea</emphasis>
</taxonomicName>
.
<location id="8ECD6039FFA6FF83E3F027C8B8B3FA3B" LSID="urn:lsid:plazi:treatment:03BB87F4FFA6FF83E1FF23A4BB94F83C:8ECD6039FFA6FF83E3F027C8B8B3FA3B" box="[664,718,1456,1479]" country="Brazil" county="The" municipality="Weymer" name="Here" pageId="10" pageNumber="211" stateProvince="Para">Here</location>
we formally designate this specimen as
<typeStatus id="54A98840FFA6FF83E5F627D7BF79FA34" box="[1182,1284,1455,1480]" pageId="10" pageNumber="211" type="lectotype">lectotype</typeStatus>
based on the following.
<location id="8ECD6039FFA6FF83E07C27ABBB3DFA10" LSID="urn:lsid:plazi:treatment:03BB87F4FFA6FF83E1FF23A4BB94F83C:8ECD6039FFA6FF83E07C27ABBB3DFA10" box="[276,320,1491,1516]" country="Brazil" county="The" municipality="Weymer" name="The" pageId="10" pageNumber="211" stateProvince="Para">The</location>
specimen was examined by CMP and TS, and it matches the
<taxonomicName id="4C124D61FFA6FF83E56127ABBEFBFA17" box="[1033,1158,1491,1515]" class="Insecta" family="Nymphalidae" genus="Bia" kingdom="Animalia" order="Lepidoptera" pageId="10" pageNumber="211" phylum="Arthropoda" rank="species" species="decaerulea">
<emphasis id="B966EAF0FFA6FF83E56127ABBEFBFA17" box="[1033,1158,1491,1515]" italics="true" pageId="10" pageNumber="211">decaerulea</emphasis>
</taxonomicName>
phenotype described by
<location id="8ECD6039FFA6FF83E1FF2780BA88F9EC" LSID="urn:lsid:plazi:treatment:03BB87F4FFA6FF83E1FF23A4BB94F83C:8ECD6039FFA6FF83E1FF2780BA88F9EC" box="[151,245,1528,1552]" country="Brazil" county="The" municipality="Weymer" name="Weymer" pageId="10" pageNumber="211" stateProvince="Para">Weymer</location>
and illustrated in
<location id="8ECD6039FFA6FF83E0AE278FB8AAF9EC" LSID="urn:lsid:plazi:treatment:03BB87F4FFA6FF83E1FF23A4BB94F83C:8ECD6039FFA6FF83E0AE278FB8AAF9EC" box="[454,727,1527,1552]" country="Brazil" county="The" municipality="Weymer" name="Seitz' Macrolepidoptera" pageId="10" pageNumber="211" stateProvince="Para">Seitz Macrolepidoptera</location>
of the
<location id="8ECD6039FFA6FF83E243278FB9B2F9EC" LSID="urn:lsid:plazi:treatment:03BB87F4FFA6FF83E1FF23A4BB94F83C:8ECD6039FFA6FF83E243278FB9B2F9EC" box="[811,975,1527,1552]" country="Brazil" county="The" municipality="Weymer" name="World Volume" pageId="10" pageNumber="211" stateProvince="Para">World Volume</location>
5 plate 60 e.
<location id="8ECD6039FFA6FF83E5182780BEFCF9F3" LSID="urn:lsid:plazi:treatment:03BB87F4FFA6FF83E1FF23A4BB94F83C:8ECD6039FFA6FF83E5182780BEFCF9F3" box="[1136,1153,1528,1551]" country="Brazil" county="The" municipality="Weymer" name="It" pageId="10" pageNumber="211" stateProvince="Para">It</location>
bears the label “
<location id="8ECD6039FFA6FF83E42A278FBF05F9F3" LSID="urn:lsid:plazi:treatment:03BB87F4FFA6FF83E1FF23A4BB94F83C:8ECD6039FFA6FF83E42A278FBF05F9F3" box="[1346,1400,1527,1551]" country="Brazil" county="The" municipality="Weymer" name="Faro" pageId="10" pageNumber="211" stateProvince="Para">Faro</location>
86
<location id="8ECD6039FFA6FF83E1FF2463BAB8F9C8" LSID="urn:lsid:plazi:treatment:03BB87F4FFA6FF83E1FF23A4BB94F83C:8ECD6039FFA6FF83E1FF2463BAB8F9C8" box="[151,197,1563,1588]" country="Brazil" county="The" municipality="Weymer" name="Hhl" pageId="10" pageNumber="211" stateProvince="Para">Hhl</location>
”, indicating that it was collected by
<collectorName id="26E75334FFA6FF83E3342463B89BF9C8" box="[604,742,1563,1588]" pageId="10" pageNumber="211">Paul Hahnel</collectorName>
in 1886.
<location id="8ECD6039FFA6FF83E2382463B9DCF9C8" LSID="urn:lsid:plazi:treatment:03BB87F4FFA6FF83E1FF23A4BB94F83C:8ECD6039FFA6FF83E2382463B9DCF9C8" box="[848,929,1563,1588]" country="Brazil" county="The" municipality="Weymer" name="Hahnel" pageId="10" pageNumber="211" stateProvince="Para">Hahnel</location>
participated in
<location id="8ECD6039FFA6FF83E5382463BED2F9CF" LSID="urn:lsid:plazi:treatment:03BB87F4FFA6FF83E1FF23A4BB94F83C:8ECD6039FFA6FF83E5382463BED2F9CF" box="[1104,1199,1563,1587]" country="Brazil" county="The" municipality="Weymer" name="Amazon" pageId="10" pageNumber="211" stateProvince="Para">Amazon</location>
expeditions between
<collectingDate id="EFE8E9CAFFA6FF83E1FF2447BBB2F9A4" box="[151,463,1599,1624]" isEnumeration="true" pageId="10" pageNumber="211">
<date id="FFAC1022FFA6FF83E1FF2447BB68F9AB" box="[151,277,1599,1623]" pageId="10" pageNumber="211" value="1879" valueMax="1884">18791884</date>
and
<date id="FFAC1022FFA6FF83E0262447BBB2F9A4" box="[334,463,1599,1624]" pageId="10" pageNumber="211" value="1885" valueMax="1887">18851887</date>
</collectingDate>
, well after the establishment of Faro in 1769. The Hahnel collection was sold to Otto Staudinger and his son-in-law
<collectingRegion id="49D6F800FFA6FF83E099241BB833F980" box="[497,590,1635,1660]" country="Saint Vincent and the Grenadines" name="Saint Andrew" pageId="10" pageNumber="211">Andreas</collectingRegion>
Bang-Hass, possibly after Hahnels death in 1887. This time frame allows for the exchange of correspondence
<locationDeviation id="9A1F2FC0FFA6FF83E34524FFB9E3F95C" box="[557,926,1671,1696]" pageId="10" pageNumber="211">between Weymer and Staudinger</locationDeviation>
(deceased in 1900) prior to the description of
<taxonomicName id="4C124D61FFA6FF83E1FF24D3BB69F93F" box="[151,276,1707,1731]" class="Insecta" family="Nymphalidae" genus="Bia" kingdom="Animalia" order="Lepidoptera" pageId="10" pageNumber="211" phylum="Arthropoda" rank="species" species="decaerulea">
<emphasis id="B966EAF0FFA6FF83E1FF24D3BB69F93F" box="[151,276,1707,1731]" italics="true" pageId="10" pageNumber="211">decaerulea</emphasis>
</taxonomicName>
in 1911. Finally, Faro is located north of the
<location id="8ECD6039FFA6FF83E24924D3B9BAF938" LSID="urn:lsid:plazi:treatment:03BB87F4FFA6FF83E1FF23A4BB94F83C:8ECD6039FFA6FF83E24924D3B9BAF938" box="[801,967,1707,1732]" country="Brazil" county="The" municipality="Weymer" name="Amazon River" pageId="10" pageNumber="211" stateProvince="Para">Amazon River</location>
and east of the
<location id="8ECD6039FFA6FF83E5EA24D3BF39F938" LSID="urn:lsid:plazi:treatment:03BB87F4FFA6FF83E1FF23A4BB94F83C:8ECD6039FFA6FF83E5EA24D3BF39F938" box="[1154,1348,1707,1732]" country="Brazil" county="The" municipality="Weymer" name="Nhamunda River" pageId="10" pageNumber="211" stateProvince="Para">Nhamundá River</location>
mouth, where MMC collected specimens that match the one collected by
<collectorName id="26E75334FFA6FF83E2E824B7B9ACF914" box="[896,977,1743,1768]" pageId="10" pageNumber="211">Hahnel</collectorName>
(
<figureCitation id="13292A67FFA6FF83E28924B7BE4EF91B" box="[993,1075,1743,1768]" captionStart="FIGURE 10" captionStartId="24.[151,250,1758,1780]" captionTargetBox="[159,1426,836,1727]" captionTargetId="figure@24.[151,1436,827,1737]" captionTargetPageId="24" captionText="FIGURE 10. Androconial organs of Bia species and subspecies. Each set of images includes a detail of dorsal forewing CuA- CuA 1 - CuA 2 intersection (image on left), dorsal hind wing discal androconial pad with associated hairpencil (center) and with the hairpencil removed (right). a, actorion actorion, Brazil, Manaus (DZUP 16588); b, actorion ecuatoria, Brazil, Mâncio Lima (DZUP 32829); c, actorion occulta, Brazil, Benevides (DZUP 22417); d, decaerulea decaerulea, Brazil, Amazonas, Nhamundá (DZUP 22356); e, decaerulea pallida, Brazil, Barcelos (DZUP 22506); f, decaerulea cayanna, French Guiana, Vidal (USNM); g, rebeli aegina, Peru, Tambopata Reserve (DZUP 22927); h, rebeli acreana, Brazil, Sta. Rosa do Purus (DZUP 22557); i, rebeli arikeme, Brazil, Ouro Preto d'Oeste (DZUP 22756); j, rebeli cuprea, Brazil, Borba (DZUP 16378); k, caelestis, Brazil, Mâncio Lima (DZUP 32830); l, pucallpa, Peru, Pucallpa. (USNM)." httpUri="https://zenodo.org/record/569740/files/figure.png" pageId="10" pageNumber="211">Fig. 10</figureCitation>
a; a photograph including labels is available at http://butterfliesofamerica.com/L/t/
<taxonomicName id="4C124D61FFA6FF83E3B0248BB883F8F0" box="[728,766,1779,1804]" class="Insecta" family="Nymphalidae" genus="Bia" kingdom="Animalia" order="Lepidoptera" pageId="10" pageNumber="211" phylum="Arthropoda" rank="genus">Bia</taxonomicName>
_a.htm, last accessed
<collectingDate id="EFE8E9CAFFA6FF83E292248BBEC6F8F0" box="[1018,1211,1779,1804]" pageId="10" pageNumber="211" value="2016-08-30">
<date id="FFAC1022FFA6FF83E292248BBEC6F8F0" box="[1018,1211,1779,1804]" pageId="10" pageNumber="211" value="2016-08-30">30 August 2016</date>
</collectingDate>
).
<collectorName id="26E75334FFA6FF83E5BA248BBE83F8F0" box="[1234,1278,1779,1804]" pageId="10" pageNumber="211">The</collectorName>
<typeStatus id="54A98840FFA6FF83E462248BBF0DF8F0" box="[1290,1392,1779,1804]" pageId="10" pageNumber="211" type="lectotype">lectotype</typeStatus>
M, deposited in the
<collectionCode id="ED03AE27FFA6FF83E025256FBBFEF8CC" box="[333,387,1815,1840]" country="Germany" httpUri="http://grbio.org/cool/jp8y-i5nv" name="Museum f<>r Naturkunde" pageId="10" pageNumber="211">MfN</collectionCode>
collection, bears three labels separated by // and transcribed verbatim:
<collectorName id="26E75334FFA6FF83E5F12560BEB0F8D3" box="[1177,1229,1816,1839]" pageId="10" pageNumber="211">Faro</collectorName>
86
<collectorName id="26E75334FFA6FF83E59D256FBF62F8CC" box="[1269,1311,1815,1840]" pageId="10" pageNumber="211">Hnl</collectorName>
// genitalia vial M-9047 [male symbol]
<collectorName id="26E75334FFA6FF83E0B82544B861F8AF" box="[464,540,1852,1875]" pageId="10" pageNumber="211">Lee D.</collectorName>
Miller //
<taxonomicName id="4C124D61FFA6FF83E3EC2543B973F8A8" box="[644,782,1851,1876]" class="Insecta" family="Nymphalidae" genus="Bia" kingdom="Animalia" order="Lepidoptera" pageId="10" pageNumber="211" phylum="Arthropoda" rank="species" species="actorion">Bia actorion</taxonomicName>
form
<taxonomicName id="4C124D61FFA6FF83E2382543BE66F8A8" authority="Weym." authorityName="Weym." box="[848,1051,1851,1876]" class="Insecta" family="Nymphalidae" genus="Bia" kingdom="Animalia" order="Lepidoptera" pageId="10" pageNumber="211" phylum="Arthropoda" rank="species" species="decaerulea">decaerulea Weym</taxonomicName>
</materialsCitation>
. LECTOTYPE [male symbol] det. R.I. Vane-Wright, 1969 [the latter is printed across the center of the label] // Lectotypus.
</paragraph>
</subSubSection>
<subSubSection id="C3086569FFA6FF83E1AF25FBBB94F83C" pageId="10" pageNumber="211" type="discussion">
<paragraph id="8BAD36E2FFA6FF83E1AF25FBBB94F83C" blockId="10.[151,1437,582,1984]" pageId="10" pageNumber="211">
<emphasis id="B966EAF0FFA6FF83E1AF25FBBB46F860" bold="true" box="[199,315,1923,1948]" pageId="10" pageNumber="211">Remarks.</emphasis>
Geographical variation in some diagnostic characters serves as the basis for the subspecific classification proposed below.
</paragraph>
</subSubSection>
</treatment>
</document>