360 lines
54 KiB
XML
360 lines
54 KiB
XML
<document id="4CB337AB3F2C1875B30D4097AEB4C5C1" ID-CLB-Dataset="46283" ID-DOI="10.5281/zenodo.278802" ID-GBIF-Dataset="275e1f6f-69ff-4f2e-a9a9-0d1cb7d286d8" ID-ISSN="1175-5326" ID-Zenodo-Dep="278802" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460392962328" checkinUser="plazi" docAuthor="Anker, Arthur" docDate="2011" docId="03BB87D5FF9CFF83FF7F2365FB899961" docLanguage="en" docName="zt03041p038.pdf" docOrigin="Zootaxa 3041" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Leptalpheus azuero Anker, 2011, sp. nov." docType="treatment" docVersion="13" lastPageNumber="12" masterDocId="FF82FFADFF94FF88FFE82158FFA79D55" masterDocTitle="Six new species and three new records of infaunal alpheid shrimps from the genera Leptalpheus Williams, 1965 and Fenneralpheus Felder & Manning, 1986 (Crustacea, Decapoda)" masterLastPageNumber="38" masterPageNumber="1" pageNumber="9" updateTime="1698298436616" updateUser="ExternalLinkService">
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<mods:mods id="B6B0BA2BB491DCB58CBE799BA43EF5A2" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="178B697CC8103B391EAFF4046FE80442">
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<mods:title id="605AE80D1575B01A7C26FC96C88A051B">Six new species and three new records of infaunal alpheid shrimps from the genera Leptalpheus Williams, 1965 and Fenneralpheus Felder & Manning, 1986 (Crustacea, Decapoda)</mods:title>
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</mods:titleInfo>
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<mods:name id="C90375965CD243A474F0BD37EAFCB0B8" type="personal">
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<mods:role id="3F7556A0CD6EF2AC745F76E97307073E">
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<mods:roleTerm id="D459E690048D4EE8FA02EDC0B2897BB8">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="BD15D8108C909C6CA7A2B5A46FCFDD87">Anker, Arthur</mods:namePart>
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</mods:name>
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<mods:typeOfResource id="E4480FDC0BFF73BA15243B1E4B74DB4E">text</mods:typeOfResource>
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<mods:titleInfo id="AE42946D8CAD931B3414FD59BF160AE1">
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<mods:title id="FE12107E57C8CC8BCE8D8923CB443DFA">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="336C28A647FE1FF8A58E992E4C9E0C39">
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<mods:date id="FD3D976A131B99A6729FD6665BDC953E">2011</mods:date>
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<mods:detail id="8C7CE887BB485EBA57E3FEB56E9F3AB0" type="volume">
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<mods:number id="817BA3AB256C0ABC3AFBA0A2CAFC7575">3041</mods:number>
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</mods:detail>
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<mods:extent id="9D64DB2691D0DDB5A8A3658FA9D9417D" unit="page">
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<mods:start id="645DDCE8ABF5FE8A9915FBFDE1A4BC04">1</mods:start>
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<mods:end id="4DD37C64D9C22776B25AAF9ADA348629">38</mods:end>
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</mods:extent>
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</mods:relatedItem>
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<mods:classification id="79D2EC1A8D792CBB531200F519D9FC3C">journal article</mods:classification>
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<mods:identifier id="238865EDA116F000C5611B23F2EB582A" type="CLB-Dataset">46283</mods:identifier>
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<mods:identifier id="3242F18A2D0B7CEC7706D9B2B85363E2" type="DOI">10.5281/zenodo.278802</mods:identifier>
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<mods:identifier id="5B817075DF733DE5F8C82DA366F2769C" type="GBIF-Dataset">275e1f6f-69ff-4f2e-a9a9-0d1cb7d286d8</mods:identifier>
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<mods:identifier id="4207B7949443721DD4C41466E0D3165D" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="DEEFE63ED1FB50430049FFF9AE8ABD22" type="Zenodo-Dep">278802</mods:identifier>
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<treatment id="03BB87D5FF9CFF83FF7F2365FB899961" ID-DOI="http://doi.org/10.5281/zenodo.4893565" ID-GBIF-Taxon="119417252" ID-Zenodo-Dep="4893565" LSID="urn:lsid:plazi:treatment:03BB87D5FF9CFF83FF7F2365FB899961" httpUri="http://treatment.plazi.org/id/03BB87D5FF9CFF83FF7F2365FB899961" lastPageId="11" lastPageNumber="12" pageId="8" pageNumber="9">
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<subSubSection id="C3086548FF9CFF80FF7F2365FF589F22" pageId="8" pageNumber="9" type="nomenclature">
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<paragraph id="8BAD36C3FF9CFF80FF7F2365FE4F9F02" blockId="8.[151,488,573,631]" box="[151,488,573,599]" pageId="8" pageNumber="9">
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<heading id="D0E581AFFF9CFF80FF7F2365FE4F9F02" bold="true" box="[151,488,573,599]" fontSize="11" level="1" pageId="8" pageNumber="9" reason="1">
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||
<emphasis id="B966EAD1FF9CFF80FF7F2365FE4F9F02" bold="true" box="[151,488,573,599]" pageId="8" pageNumber="9">
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||
<taxonomicName id="4C124D40FF9CFF80FF7F2365FE249F02" ID-CoL="85TJC" box="[151,387,573,599]" class="Malacostraca" family="Alpheidae" genus="Leptalpheus" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="azuero" status="sp. nov.">
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<emphasis id="B966EAD1FF9CFF80FF7F2365FE249F02" bold="true" box="[151,387,573,599]" italics="true" pageId="8" pageNumber="9">Leptalpheus azuero</emphasis>
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||
</taxonomicName>
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||
<taxonomicNameLabel id="A25557AAFF9CFF80FE632365FE4F9F02" box="[395,488,573,599]" pageId="8" pageNumber="9" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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||
</heading>
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||
</paragraph>
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<paragraph id="8BAD36C3FF9CFF80FF7F2307FF589F22" blockId="8.[151,488,573,631]" box="[151,255,607,631]" pageId="8" pageNumber="9">
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<figureCitation id="13292A46FF9CFF80FF7F2307FF589F22" box="[151,255,607,631]" captionStart-0="FIGURE 5" captionStart-1="FIGURE 6" captionStart-2="FIGURE 7" captionStartId-0="9.[151,250,1808,1831]" captionStartId-1="10.[151,250,1281,1304]" captionStartId-2="10.[151,250,1734,1757]" captionTargetBox-0="[173,1407,200,1771]" captionTargetBox-1="[169,1409,208,1241]" captionTargetBox-2="[151,1435,1414,1713]" captionTargetId-0="figure@9.[151,1436,193,1787]" captionTargetId-1="figure@10.[157,1430,193,1260]" captionTargetId-2="figure@10.[151,1436,1414,1713]" captionTargetPageId-0="9" captionTargetPageId-1="10" captionTargetPageId-2="10" captionText-0="FIGURE 5. Leptalpheus azuero sp. nov.: all except I—paratype, female from Playa El Aguillito, Azuero Peninsula, Pacific coast of Panama (RMNH D 54560); I—paratype, male from the same locality (MNHN-IU- 2011 - 5240): A, frontal region, dorsal view; B, same, lateral view; C, telson, dorsal view; D, ventromesial tooth on first article of antennular peduncle, lateral view; E, scaphocerite, dorsal view; F, third maxilliped, lateral view; G, second pereiopod, lateral view; H, third pereiopod, lateral view; I, appendix masculina and appendix interna, dorsal view; J, uropod, dorsal view (setae omitted)." captionText-1="FIGURE 6. Leptalpheus azuero sp. nov., paratype, female from Playa El Aguillito, Azuero Peninsula, Pacific coast of Panama (RMNH D 54560): A, major cheliped, lateral view; B, same, mesial view; C, same, carpus and chela, lateral view; D, major chela fingers, lateral view; E, minor cheliped, lateral view; F, same, carpus and chela, mesial view." captionText-2="FIGURE 7. Leptalpheus azuero sp. nov., A, holotype, female from Playa El Aguillito, Azuero Peninsula, Pacific coast of Panama (MNHN-IU- 2011 - 5239), lateral view; B, non-type female from the same locality, dorsal view (UP)." httpUri-0="https://zenodo.org/record/278807/files/figure.png" httpUri-1="https://zenodo.org/record/278808/files/figure.png" httpUri-2="https://zenodo.org/record/278809/files/figure.png" pageId="8" pageNumber="9">Figs. 5–7</figureCitation>
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</paragraph>
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||
</subSubSection>
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<subSubSection id="C3086548FF9CFF80FF7F23F0FAF19E21" pageId="8" pageNumber="9" type="materials_examined">
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<paragraph id="8BAD36C3FF9CFF80FF7F23F0FBEE9E1A" blockId="8.[151,1436,679,1964]" pageId="8" pageNumber="9">
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<emphasis id="B966EAD1FF9CFF80FF7F23F0FEEE9F95" bold="true" box="[151,329,679,704]" pageId="8" pageNumber="9">
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||
<typeStatus id="54A98861FF9CFF80FF7F23F0FF759F95" box="[151,210,680,704]" pageId="8" pageNumber="9">Type</typeStatus>
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material.
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</emphasis>
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<typeStatus id="54A98861FF9CFF80FEBC23FFFE669FEA" box="[340,449,679,703]" pageId="8" pageNumber="9" type="holotype">Holotype</typeStatus>
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: female (cl
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<quantity id="4CEA9B26FF9CFF80FDA623FFFD0B9FEA" box="[590,684,679,703]" metricMagnitude="-3" metricUnit="m" metricValue="2.8" pageId="8" pageNumber="9" unit="mm" value="2.8">2.8 mm</quantity>
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), MNHN-IU-2011-5239,
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<collectingCountry id="F3057653FF9CFF80FC3B23F0FB889FEA" box="[979,1071,680,703]" name="Panama" pageId="8" pageNumber="9">Panama</collectingCountry>
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, Pacific coast, Chitré, Playa El Aguillito, mudflat near mangroves, yabby pump,
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<quantity id="4CEA9B26FF9CFF80FD562394FC899FB1" box="[702,814,716,740]" metricMagnitude="-1" metricUnit="m" metricValue="3.5" metricValueMax="5.0" metricValueMin="2.0" pageId="8" pageNumber="9" unit="m" value="0.35" valueMax="0.5" valueMin="0.2">0.2–0.5 m</quantity>
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at low tide, leg. A. Anker, J.A. Vera Caripe, 0
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<date id="FFAC1003FF9CFF80FAC52394FA3B9FB1" box="[1325,1436,716,740]" pageId="8" pageNumber="9" value="2006-08-11">8.11.2006</date>
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[fcn 06-536].
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<typeStatus id="54A98861FF9CFF80FEDE23A8FE009E5D" box="[310,423,752,776]" pageId="8" pageNumber="9" type="paratype">Paratypes</typeStatus>
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:
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<specimenCount id="9D14FD4AFF9CFF80FE5B23B7FDB19E52" box="[435,534,751,775]" pageId="8" pageNumber="9" type="female">1 female</specimenCount>
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(cl
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<quantity id="4CEA9B26FF9CFF80FDAF23B7FD049E52" box="[583,675,751,775]" metricMagnitude="-3" metricUnit="m" metricValue="2.7" pageId="8" pageNumber="9" unit="mm" value="2.7">2.7 mm</quantity>
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),
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<collectionCode id="ED03AE06FF9CFF80FD5023B7FC8F9E52" box="[696,808,751,775]" pageId="8" pageNumber="9">OUMNH</collectionCode>
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.ZC. 2011-06-020, same collection data [fcn 06-537];
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<specimenCount id="9D14FD4AFF9CFF80FA6623B7FF499E79" pageId="8" pageNumber="9" type="female">2 females</specimenCount>
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(cl 2.4,
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<quantity id="4CEA9B26FF9CFF80FEBA224CFE089E79" box="[338,431,788,812]" metricMagnitude="-3" metricUnit="m" metricValue="2.6" pageId="8" pageNumber="9" unit="mm" value="2.6">2.6 mm</quantity>
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),
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<collectionCode id="ED03AE06FF9CFF80FE2D224DFDBE9E79" LSID="urn:lsid:biocol.org:col:34992" box="[453,537,789,812]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="8" pageNumber="9">RMNH</collectionCode>
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D54560, same collection data [fcn 06-526A, 06-526B];
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<specimenCount id="9D14FD4AFF9CFF80FB41224CFAB09E79" box="[1193,1303,788,812]" pageId="8" pageNumber="9" type="female">3 females</specimenCount>
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(cl
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<quantity id="4CEA9B26FF9CFF80FAA0224CFF609E1A" metricMagnitude="-3" metricUnit="m" metricValue="2.8499999999999996" metricValueMax="2.9" metricValueMin="2.8" pageId="8" pageNumber="9" unit="mm" value="2.8499999999999996" valueMax="2.9" valueMin="2.8">2.8–2.9 mm</quantity>
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),
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<specimenCount id="9D14FD4AFF9CFF80FF32226FFE829E1A" box="[218,293,823,847]" pageId="8" pageNumber="9" type="male">1 male</specimenCount>
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(cl
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<quantity id="4CEA9B26FF9CFF80FEB9226FFE0C9E1A" box="[337,427,823,847]" metricMagnitude="-3" metricUnit="m" metricValue="3.7" pageId="8" pageNumber="9" unit="mm" value="3.7">3.7 mm</quantity>
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), MNHN-IU-2011-5240, same collection data [fcn 06-528].
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</paragraph>
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<paragraph id="8BAD36C3FF9CFF80FF2F2203FAF19E21" blockId="8.[151,1436,679,1964]" box="[199,1366,859,885]" pageId="8" pageNumber="9">
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<emphasis id="B966EAD1FF9CFF80FF2F2203FE1E9E21" bold="true" box="[199,441,859,884]" pageId="8" pageNumber="9">Additional material.</emphasis>
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<specimenCount id="9D14FD4AFF9CFF80FE282204FD869E21" box="[448,545,860,884]" pageId="8" pageNumber="9" type="female">1 female</specimenCount>
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||
(cl
|
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<quantity id="4CEA9B26FF9CFF80FDA52204FD009E21" box="[589,679,860,884]" metricMagnitude="-3" metricUnit="m" metricValue="2.6" pageId="8" pageNumber="9" unit="mm" value="2.6">2.6 mm</quantity>
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), UP, same collection data as for the
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<typeStatus id="54A98861FF9CFF80FBAC2205FBD39E20" box="[1092,1140,861,885]" pageId="8" pageNumber="9">type</typeStatus>
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series [fcn 06-534].
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</paragraph>
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</subSubSection>
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||
<subSubSection id="C3086548FF9CFF82FF2F2227FE399AF9" lastPageId="10" lastPageNumber="11" pageId="8" pageNumber="9" type="description">
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<paragraph id="8BAD36C3FF9CFF80FF2F2227FAFE9951" blockId="8.[151,1436,679,1964]" pageId="8" pageNumber="9">
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<emphasis id="B966EAD1FF9CFF80FF2F2227FEFE9ECD" bold="true" box="[199,345,895,920]" pageId="8" pageNumber="9">Description.</emphasis>
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Frontal margin of carapace broadly triangular, with short, blunt rostral projection, without orbital crests (
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<figureCitation id="13292A46FF9CFF80FF0E22FCFE8F9EE9" box="[230,296,932,956]" captionStart="FIGURE 5" captionStartId="9.[151,250,1808,1831]" captionTargetBox="[173,1407,200,1771]" captionTargetId="figure@9.[151,1436,193,1787]" captionTargetPageId="9" captionText="FIGURE 5. Leptalpheus azuero sp. nov.: all except I—paratype, female from Playa El Aguillito, Azuero Peninsula, Pacific coast of Panama (RMNH D 54560); I—paratype, male from the same locality (MNHN-IU- 2011 - 5240): A, frontal region, dorsal view; B, same, lateral view; C, telson, dorsal view; D, ventromesial tooth on first article of antennular peduncle, lateral view; E, scaphocerite, dorsal view; F, third maxilliped, lateral view; G, second pereiopod, lateral view; H, third pereiopod, lateral view; I, appendix masculina and appendix interna, dorsal view; J, uropod, dorsal view (setae omitted)." httpUri="https://zenodo.org/record/278807/files/figure.png" pageId="8" pageNumber="9">Fig. 5</figureCitation>
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A, B). Telson widest in proximal third, very slightly tapering distally; dorsal surface with two pairs of strong spiniform setae inserted in deep pits relatively close to lateral margin; posterior margin rounded, with two pairs of spiniform setae at posterolateral angles, lateral much more slender and shorter than mesial (
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<figureCitation id="13292A46FF9CFF80FB1C22B4FA919951" box="[1268,1334,1004,1028]" captionStart="FIGURE 5" captionStartId="9.[151,250,1808,1831]" captionTargetBox="[173,1407,200,1771]" captionTargetId="figure@9.[151,1436,193,1787]" captionTargetPageId="9" captionText="FIGURE 5. Leptalpheus azuero sp. nov.: all except I—paratype, female from Playa El Aguillito, Azuero Peninsula, Pacific coast of Panama (RMNH D 54560); I—paratype, male from the same locality (MNHN-IU- 2011 - 5240): A, frontal region, dorsal view; B, same, lateral view; C, telson, dorsal view; D, ventromesial tooth on first article of antennular peduncle, lateral view; E, scaphocerite, dorsal view; F, third maxilliped, lateral view; G, second pereiopod, lateral view; H, third pereiopod, lateral view; I, appendix masculina and appendix interna, dorsal view; J, uropod, dorsal view (setae omitted)." httpUri="https://zenodo.org/record/278807/files/figure.png" pageId="8" pageNumber="9">Fig. 5</figureCitation>
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C).
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</paragraph>
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<paragraph id="8BAD36C3FF9CFF80FF2F2548FEA69871" blockId="8.[151,1436,679,1964]" pageId="8" pageNumber="9">
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Eyestalks with anteromesial margin rounded (
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<figureCitation id="13292A46FF9CFF80FD2F2548FCAE997D" box="[711,777,1040,1064]" captionStart="FIGURE 5" captionStartId="9.[151,250,1808,1831]" captionTargetBox="[173,1407,200,1771]" captionTargetId="figure@9.[151,1436,193,1787]" captionTargetPageId="9" captionText="FIGURE 5. Leptalpheus azuero sp. nov.: all except I—paratype, female from Playa El Aguillito, Azuero Peninsula, Pacific coast of Panama (RMNH D 54560); I—paratype, male from the same locality (MNHN-IU- 2011 - 5240): A, frontal region, dorsal view; B, same, lateral view; C, telson, dorsal view; D, ventromesial tooth on first article of antennular peduncle, lateral view; E, scaphocerite, dorsal view; F, third maxilliped, lateral view; G, second pereiopod, lateral view; H, third pereiopod, lateral view; I, appendix masculina and appendix interna, dorsal view; J, uropod, dorsal view (setae omitted)." httpUri="https://zenodo.org/record/278807/files/figure.png" pageId="8" pageNumber="9">Fig. 5</figureCitation>
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A). Antennular peduncles rather stout, not greatly flattened dorsoventrally; stylocerite not appressed, exceeding distal margin of first article; ventromesial carina with strong, distally bifid tooth; second article subquadrate, slightly shorter than wide; lateral flagellum with well-developed secondary ramus (
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<figureCitation id="13292A46FF9CFF80FE8D2524FE0099C1" box="[357,423,1148,1172]" captionStart="FIGURE 5" captionStartId="9.[151,250,1808,1831]" captionTargetBox="[173,1407,200,1771]" captionTargetId="figure@9.[151,1436,193,1787]" captionTargetPageId="9" captionText="FIGURE 5. Leptalpheus azuero sp. nov.: all except I—paratype, female from Playa El Aguillito, Azuero Peninsula, Pacific coast of Panama (RMNH D 54560); I—paratype, male from the same locality (MNHN-IU- 2011 - 5240): A, frontal region, dorsal view; B, same, lateral view; C, telson, dorsal view; D, ventromesial tooth on first article of antennular peduncle, lateral view; E, scaphocerite, dorsal view; F, third maxilliped, lateral view; G, second pereiopod, lateral view; H, third pereiopod, lateral view; I, appendix masculina and appendix interna, dorsal view; J, uropod, dorsal view (setae omitted)." httpUri="https://zenodo.org/record/278807/files/figure.png" pageId="8" pageNumber="9">Fig. 5</figureCitation>
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A, B, D). Antenna with moderately stout basicerite bearing strong distoventral tooth; scaphocerite broadly oval, with small, acute distolateral tooth not reaching beyond anterior margin of blade; carpocerite stout, reaching slightly beyond scaphocerite, not reaching beyond end of antennular peduncle (
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||
<figureCitation id="13292A46FF9CFF80FAF8259CFAF19989" box="[1296,1366,1220,1244]" captionStart="FIGURE 5" captionStartId="9.[151,250,1808,1831]" captionTargetBox="[173,1407,200,1771]" captionTargetId="figure@9.[151,1436,193,1787]" captionTargetPageId="9" captionText="FIGURE 5. Leptalpheus azuero sp. nov.: all except I—paratype, female from Playa El Aguillito, Azuero Peninsula, Pacific coast of Panama (RMNH D 54560); I—paratype, male from the same locality (MNHN-IU- 2011 - 5240): A, frontal region, dorsal view; B, same, lateral view; C, telson, dorsal view; D, ventromesial tooth on first article of antennular peduncle, lateral view; E, scaphocerite, dorsal view; F, third maxilliped, lateral view; G, second pereiopod, lateral view; H, third pereiopod, lateral view; I, appendix masculina and appendix interna, dorsal view; J, uropod, dorsal view (setae omitted)." httpUri="https://zenodo.org/record/278807/files/figure.png" pageId="8" pageNumber="9">Fig. 5</figureCitation>
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B, E). Mouthparts typical for genus. Third maxilliped with moderately elongate, distally subacute lateral plate on coxa (
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<figureCitation id="13292A46FF9CFF80FF772454FF459871" box="[159,226,1292,1316]" captionStart="FIGURE 5" captionStartId="9.[151,250,1808,1831]" captionTargetBox="[173,1407,200,1771]" captionTargetId="figure@9.[151,1436,193,1787]" captionTargetPageId="9" captionText="FIGURE 5. Leptalpheus azuero sp. nov.: all except I—paratype, female from Playa El Aguillito, Azuero Peninsula, Pacific coast of Panama (RMNH D 54560); I—paratype, male from the same locality (MNHN-IU- 2011 - 5240): A, frontal region, dorsal view; B, same, lateral view; C, telson, dorsal view; D, ventromesial tooth on first article of antennular peduncle, lateral view; E, scaphocerite, dorsal view; F, third maxilliped, lateral view; G, second pereiopod, lateral view; H, third pereiopod, lateral view; I, appendix masculina and appendix interna, dorsal view; J, uropod, dorsal view (setae omitted)." httpUri="https://zenodo.org/record/278807/files/figure.png" pageId="8" pageNumber="9">Fig. 5</figureCitation>
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F).
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</paragraph>
|
||
<paragraph id="8BAD36C3FF9CFF80FF2D2468FED59B3D" blockId="8.[151,1436,679,1964]" pageId="8" pageNumber="9">
|
||
Major cheliped slender; ischium unarmed mesially; merus slender, depressed ventrally, with smooth margins; carpus very short, cup-shaped, without large processes distally; chela slender, with palm mostly smooth, depressed ventrally; small field of low tubercles and rugosites present near pollex; fingers about half palm length, not strongly twisted or curved, not gaping when closed; cutting edge of dactylus with large blunt tooth fitting into broad hiatus on pollex, cutting edge of pollex with larger, broadly triangular, proximal tooth and smaller, more narrowly triangular, subdistal tooth, dactylus tip curved upwards, subacute; adhesive disks present (
|
||
<figureCitation id="13292A46FF9CFF80FB2F24BCFAAE98A9" box="[1223,1289,1508,1532]" captionStart="FIGURE 6" captionStartId="10.[151,250,1281,1304]" captionTargetBox="[169,1409,208,1241]" captionTargetId="figure@10.[157,1430,193,1260]" captionTargetPageId="10" captionText="FIGURE 6. Leptalpheus azuero sp. nov., paratype, female from Playa El Aguillito, Azuero Peninsula, Pacific coast of Panama (RMNH D 54560): A, major cheliped, lateral view; B, same, mesial view; C, same, carpus and chela, lateral view; D, major chela fingers, lateral view; E, minor cheliped, lateral view; F, same, carpus and chela, mesial view." httpUri="https://zenodo.org/record/278808/files/figure.png" pageId="8" pageNumber="9">Fig. 6</figureCitation>
|
||
A–D). Minor cheliped with ischium unarmed; merus slender, ventrally flattened, with smooth margins; carpus short, cup-shaped; chela very slender, simple, with fingers slightly longer than palm, tips crossing distally; cutting edges minutely rugose (
|
||
<figureCitation id="13292A46FF9CFF80FF192708FE949B3D" box="[241,307,1616,1640]" captionStart="FIGURE 6" captionStartId="10.[151,250,1281,1304]" captionTargetBox="[169,1409,208,1241]" captionTargetId="figure@10.[157,1430,193,1260]" captionTargetPageId="10" captionText="FIGURE 6. Leptalpheus azuero sp. nov., paratype, female from Playa El Aguillito, Azuero Peninsula, Pacific coast of Panama (RMNH D 54560): A, major cheliped, lateral view; B, same, mesial view; C, same, carpus and chela, lateral view; D, major chela fingers, lateral view; E, minor cheliped, lateral view; F, same, carpus and chela, mesial view." httpUri="https://zenodo.org/record/278808/files/figure.png" pageId="8" pageNumber="9">Fig. 6</figureCitation>
|
||
E, F).
|
||
</paragraph>
|
||
<paragraph id="8BAD36C3FF9CFF80FF2D272CFCCA9A15" blockId="8.[151,1436,679,1964]" pageId="8" pageNumber="9">
|
||
Second pereiopod with merus slightly shorter than carpus; carpus five-articulated, with article ratio approximately equal to 2.5: 1: 1: 0.8: 2 (
|
||
<figureCitation id="13292A46FF9CFF80FDF327C0FDFB9BE5" box="[539,604,1688,1712]" captionStart="FIGURE 5" captionStartId="9.[151,250,1808,1831]" captionTargetBox="[173,1407,200,1771]" captionTargetId="figure@9.[151,1436,193,1787]" captionTargetPageId="9" captionText="FIGURE 5. Leptalpheus azuero sp. nov.: all except I—paratype, female from Playa El Aguillito, Azuero Peninsula, Pacific coast of Panama (RMNH D 54560); I—paratype, male from the same locality (MNHN-IU- 2011 - 5240): A, frontal region, dorsal view; B, same, lateral view; C, telson, dorsal view; D, ventromesial tooth on first article of antennular peduncle, lateral view; E, scaphocerite, dorsal view; F, third maxilliped, lateral view; G, second pereiopod, lateral view; H, third pereiopod, lateral view; I, appendix masculina and appendix interna, dorsal view; J, uropod, dorsal view (setae omitted)." httpUri="https://zenodo.org/record/278807/files/figure.png" pageId="8" pageNumber="9">Fig. 5</figureCitation>
|
||
G). Third and fourth pereiopods similar; third pereiopod relatively slender, compressed; ischium with small spiniform seta on ventrolateral surface; merus almost five times as long as wide; carpus about 0.4 length of merus, with distoventral spiniform seta; propodus with four spiniform setae along ventral margin; dactylus about half as long as propodus, conical, slender, acute distally (
|
||
<figureCitation id="13292A46FF9CFF80FBAD265CFB209A49" box="[1093,1159,1796,1820]" captionStart="FIGURE 5" captionStartId="9.[151,250,1808,1831]" captionTargetBox="[173,1407,200,1771]" captionTargetId="figure@9.[151,1436,193,1787]" captionTargetPageId="9" captionText="FIGURE 5. Leptalpheus azuero sp. nov.: all except I—paratype, female from Playa El Aguillito, Azuero Peninsula, Pacific coast of Panama (RMNH D 54560); I—paratype, male from the same locality (MNHN-IU- 2011 - 5240): A, frontal region, dorsal view; B, same, lateral view; C, telson, dorsal view; D, ventromesial tooth on first article of antennular peduncle, lateral view; E, scaphocerite, dorsal view; F, third maxilliped, lateral view; G, second pereiopod, lateral view; H, third pereiopod, lateral view; I, appendix masculina and appendix interna, dorsal view; J, uropod, dorsal view (setae omitted)." httpUri="https://zenodo.org/record/278807/files/figure.png" pageId="8" pageNumber="9">Fig. 5</figureCitation>
|
||
H). Fifth pereiopod slen- der, not compressed, with three distal rows of setae on propodus.
|
||
</paragraph>
|
||
<paragraph id="8BAD36C3FF9CFF80FF2D2614FA9F9AF9" blockId="8.[151,1436,679,1964]" pageId="8" pageNumber="9">
|
||
Male second pleopod with slender appendix masculina only slightly exceeding appendix interna, apex with three stiff setae (
|
||
<figureCitation id="13292A46FF9CFF80FEBA2628FE319ADD" box="[338,406,1904,1928]" captionStart="FIGURE 5" captionStartId="9.[151,250,1808,1831]" captionTargetBox="[173,1407,200,1771]" captionTargetId="figure@9.[151,1436,193,1787]" captionTargetPageId="9" captionText="FIGURE 5. Leptalpheus azuero sp. nov.: all except I—paratype, female from Playa El Aguillito, Azuero Peninsula, Pacific coast of Panama (RMNH D 54560); I—paratype, male from the same locality (MNHN-IU- 2011 - 5240): A, frontal region, dorsal view; B, same, lateral view; C, telson, dorsal view; D, ventromesial tooth on first article of antennular peduncle, lateral view; E, scaphocerite, dorsal view; F, third maxilliped, lateral view; G, second pereiopod, lateral view; H, third pereiopod, lateral view; I, appendix masculina and appendix interna, dorsal view; J, uropod, dorsal view (setae omitted)." httpUri="https://zenodo.org/record/278807/files/figure.png" pageId="8" pageNumber="9">
|
||
Fig.
|
||
<date id="FFAC1003FF9CFF80FE6F2628FE319ADD" box="[391,406,1904,1928]" pageId="8" pageNumber="9">5</date>
|
||
</figureCitation>
|
||
I). Uropod with lateral lobe of protopod ending in two subacute teeth; exopod with rounded distal margin; diaeresis straight laterally, deeply incised and with blunt tooth near mesial margin (
|
||
<figureCitation id="13292A46FF9CFF80FB3326CCFAB99AF9" box="[1243,1310,1940,1964]" captionStart="FIGURE 5" captionStartId="9.[151,250,1808,1831]" captionTargetBox="[173,1407,200,1771]" captionTargetId="figure@9.[151,1436,193,1787]" captionTargetPageId="9" captionText="FIGURE 5. Leptalpheus azuero sp. nov.: all except I—paratype, female from Playa El Aguillito, Azuero Peninsula, Pacific coast of Panama (RMNH D 54560); I—paratype, male from the same locality (MNHN-IU- 2011 - 5240): A, frontal region, dorsal view; B, same, lateral view; C, telson, dorsal view; D, ventromesial tooth on first article of antennular peduncle, lateral view; E, scaphocerite, dorsal view; F, third maxilliped, lateral view; G, second pereiopod, lateral view; H, third pereiopod, lateral view; I, appendix masculina and appendix interna, dorsal view; J, uropod, dorsal view (setae omitted)." httpUri="https://zenodo.org/record/278807/files/figure.png" pageId="8" pageNumber="9">Fig. 5</figureCitation>
|
||
J).
|
||
</paragraph>
|
||
<caption id="DF6D664BFF9DFF81FF7F2648FBC29AF4" httpUri="https://zenodo.org/record/278807/files/figure.png" pageId="9" pageNumber="10" targetBox="[173,1407,200,1771]" targetPageId="9">
|
||
<paragraph id="8BAD36C3FF9DFF81FF7F2648FBC29AF4" blockId="9.[151,1436,1808,1953]" pageId="9" pageNumber="10">
|
||
<emphasis id="B966EAD1FF9DFF81FF7F2648FEB39A73" bold="true" box="[151,276,1808,1831]" pageId="9" pageNumber="10">FIGURE 5.</emphasis>
|
||
<taxonomicName id="4C124D40FF9DFF81FEF42648FE439A72" box="[284,484,1808,1831]" class="Malacostraca" family="Alpheidae" genus="Leptalpheus" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="azuero" status="sp. nov.">
|
||
<emphasis id="B966EAD1FF9DFF81FEF42648FE439A72" box="[284,484,1808,1831]" italics="true" pageId="9" pageNumber="10">Leptalpheus azuero</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B966EAD1FF9DFF81FE052649FD989A73" bold="true" box="[493,575,1809,1830]" pageId="9" pageNumber="10">
|
||
<taxonomicNameLabel id="A25557AAFF9DFF81FE052649FD989A73" box="[493,575,1809,1830]" pageId="9" pageNumber="10" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
: all except I—paratype, female from Playa El Aguillito,
|
||
<taxonomicName id="4C124D40FF9DFF81FB7A2649FB7A9A73" box="[1170,1245,1809,1830]" class="Malacostraca" family="Alpheidae" genus="Leptalpheus" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="19" phylum="Arthropoda" rank="species" species="azuero" status="sp. nov.">Azuero</taxonomicName>
|
||
Peninsula, Pacific coast of Panama (RMNH D54560); I—paratype, male from the same locality (MNHN-IU-2011-5240): A, frontal region, dorsal view; B, same, lateral view; C, telson, dorsal view; D, ventromesial tooth on first article of antennular peduncle, lateral view; E, scaphocerite, dorsal view; F, third maxilliped, lateral view; G, second pereiopod, lateral view; H, third pereiopod, lateral view; I, appendix masculina and appendix interna, dorsal view; J, uropod, dorsal view (setae omitted).
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF6D664BFF9EFF82FF7F2459FBDA9801" httpUri="https://zenodo.org/record/278808/files/figure.png" pageId="10" pageNumber="11" targetBox="[169,1409,208,1241]" targetPageId="10">
|
||
<paragraph id="8BAD36C3FF9EFF82FF7F2459FBDA9801" blockId="10.[151,1436,1281,1364]" pageId="10" pageNumber="11">
|
||
<emphasis id="B966EAD1FF9EFF82FF7F2459FEB59843" bold="true" box="[151,274,1281,1304]" pageId="10" pageNumber="11">FIGURE 6.</emphasis>
|
||
<taxonomicName id="4C124D40FF9EFF82FEF02459FE799842" box="[280,478,1281,1303]" class="Malacostraca" family="Alpheidae" genus="Leptalpheus" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="azuero" status="sp. nov.">
|
||
<emphasis id="B966EAD1FF9EFF82FEF02459FE799842" box="[280,478,1281,1303]" italics="true" pageId="10" pageNumber="11">Leptalpheus azuero</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B966EAD1FF9EFF82FE0C245AFD929842" bold="true" box="[484,565,1282,1303]" pageId="10" pageNumber="11">
|
||
<taxonomicNameLabel id="A25557AAFF9EFF82FE0C245AFD929842" box="[484,565,1282,1303]" pageId="10" pageNumber="11" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, paratype, female from Playa El Aguillito,
|
||
<taxonomicName id="4C124D40FF9EFF82FC02245AFB939842" box="[1002,1076,1282,1303]" class="Malacostraca" family="Alpheidae" genus="Leptalpheus" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="19" phylum="Arthropoda" rank="species" species="azuero" status="sp. nov.">Azuero</taxonomicName>
|
||
Peninsula, Pacific coast of Panama (RMNH D54560): A, major cheliped, lateral view; B, same, mesial view; C, same, carpus and chela, lateral view; D, major chela fingers, lateral view; E, minor cheliped, lateral view; F, same, carpus and chela, mesial view.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF6D664BFF9EFF82FF7F279EFB1D9BA9" httpUri="https://zenodo.org/record/278809/files/figure.png" pageId="10" pageNumber="11" targetBox="[151,1435,1414,1713]" targetPageId="10">
|
||
<paragraph id="8BAD36C3FF9EFF82FF7F279EFB1D9BA9" blockId="10.[151,1435,1734,1788]" pageId="10" pageNumber="11">
|
||
<emphasis id="B966EAD1FF9EFF82FF7F279EFEB59B89" bold="true" box="[151,274,1734,1757]" pageId="10" pageNumber="11">FIGURE 7.</emphasis>
|
||
<taxonomicName id="4C124D40FF9EFF82FEF1279FFE789B88" box="[281,479,1735,1757]" class="Malacostraca" family="Alpheidae" genus="Leptalpheus" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="azuero" status="sp. nov.">
|
||
<emphasis id="B966EAD1FF9EFF82FEF1279FFE789B88" box="[281,479,1735,1757]" italics="true" pageId="10" pageNumber="11">Leptalpheus azuero</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B966EAD1FF9EFF82FE0D279FFD929B89" bold="true" box="[485,565,1735,1756]" pageId="10" pageNumber="11">
|
||
<taxonomicNameLabel id="A25557AAFF9EFF82FE0D279FFD929B89" box="[485,565,1735,1756]" pageId="10" pageNumber="11" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, A, holotype, female from Playa El Aguillito,
|
||
<taxonomicName id="4C124D40FF9EFF82FBE4279FFBF19B89" box="[1036,1110,1735,1756]" class="Malacostraca" family="Alpheidae" genus="Leptalpheus" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="19" phylum="Arthropoda" rank="species" species="azuero" status="sp. nov.">Azuero</taxonomicName>
|
||
Peninsula, Pacific coast of Panama (MNHN-IU-2011-5239), lateral view; B, non-type female from the same locality, dorsal view (UP).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BAD36C3FF9EFF82FF2F267FFE399AF9" blockId="10.[151,1436,1831,2036]" pageId="10" pageNumber="11">
|
||
<emphasis id="B966EAD1FF9EFF82FF2F267FFF599A15" bold="true" box="[199,254,1831,1856]" pageId="10" pageNumber="11">Size.</emphasis>
|
||
The
|
||
<typeStatus id="54A98861FF9EFF82FEDF2671FEC09A14" box="[311,359,1833,1857]" pageId="10" pageNumber="11">type</typeStatus>
|
||
specimens range from
|
||
<quantity id="4CEA9B26FF9EFF82FD852670FCAD9A15" box="[621,778,1832,1856]" metricMagnitude="-3" metricUnit="m" metricValue="3.05" metricValueMax="3.7" metricValueMin="2.4" pageId="10" pageNumber="11" unit="mm" value="3.05" valueMax="3.7" valueMin="2.4">2.4 to 3.7 mm</quantity>
|
||
cl, the largest being a male missing its major cheliped.
|
||
<emphasis id="B966EAD1FF9EFF82FF2F2614FECA9A30" bold="true" box="[199,365,1868,1893]" pageId="10" pageNumber="11">Colour in life.</emphasis>
|
||
Semitransparent with small red chromatophores, some forming diffuse transverse bands on the abdomen; antennular and antennal peduncles and tail fan with red chromatophores; chelipeds and walking legs mostly whitish (
|
||
<figureCitation id="13292A46FF9EFF82FEA426CCFE379AF9" box="[332,400,1940,1964]" captionStart="FIGURE 7" captionStartId="10.[151,250,1734,1757]" captionTargetBox="[151,1435,1414,1713]" captionTargetId="figure@10.[151,1436,1414,1713]" captionTargetPageId="10" captionText="FIGURE 7. Leptalpheus azuero sp. nov., A, holotype, female from Playa El Aguillito, Azuero Peninsula, Pacific coast of Panama (MNHN-IU- 2011 - 5239), lateral view; B, non-type female from the same locality, dorsal view (UP)." httpUri="https://zenodo.org/record/278809/files/figure.png" pageId="10" pageNumber="11">Fig. 7</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3086548FF9EFF82FF2F26E0FE7A9AA1" pageId="10" pageNumber="11" type="etymology">
|
||
<paragraph id="8BAD36C3FF9EFF82FF2F26E0FE7A9AA1" blockId="10.[151,1436,1831,2036]" pageId="10" pageNumber="11">
|
||
<emphasis id="B966EAD1FF9EFF82FF2F26E0FEE99A85" bold="true" box="[199,334,1976,2000]" pageId="10" pageNumber="11">Etymology.</emphasis>
|
||
Named after Panama’s
|
||
<taxonomicName id="4C124D40FF9EFF82FD8B26E1FD129A84" box="[611,693,1977,2001]" class="Malacostraca" family="Alpheidae" genus="Leptalpheus" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="19" phylum="Arthropoda" rank="species" species="azuero" status="sp. nov.">Azuero</taxonomicName>
|
||
Peninsula, on which the
|
||
<typeStatus id="54A98861FF9EFF82FC3426E1FBAB9A84" box="[988,1036,1977,2001]" pageId="10" pageNumber="11">type</typeStatus>
|
||
locality of this species is situated; used as a noun in opposition.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3086548FF9FFF83FF2F21C0FC629DFA" box="[199,965,151,176]" pageId="11" pageNumber="12" type="materials_examined">
|
||
<paragraph id="8BAD36C3FF9FFF83FF2F21C0FC629DFA" blockId="11.[151,1437,151,1076]" box="[199,965,151,176]" pageId="11" pageNumber="12">
|
||
<emphasis id="B966EAD1FF9FFF83FF2F21C0FEC29DE5" bold="true" box="[199,357,151,176]" pageId="11" pageNumber="12">
|
||
<typeStatus id="54A98861FF9FFF83FF2F21C0FEA59DE5" box="[199,258,152,176]" pageId="11" pageNumber="12">Type</typeStatus>
|
||
locality.
|
||
</emphasis>
|
||
<collectingCountry id="F3057653FF9FFF83FE8421C0FE6F9DFA" box="[364,456,152,175]" name="Panama" pageId="11" pageNumber="12">Panama</collectingCountry>
|
||
, Pacific coast, Playa El Aguillito near Chitré.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3086548FF9FFF83FF2F21E3FCDC9C31" pageId="11" pageNumber="12" type="distribution">
|
||
<paragraph id="8BAD36C3FF9FFF83FF2F21E3FF6A9DA2" blockId="11.[151,1437,151,1076]" pageId="11" pageNumber="12">
|
||
<emphasis id="B966EAD1FF9FFF83FF2F21E3FEC79D81" bold="true" box="[199,352,187,212]" pageId="11" pageNumber="12">Distribution.</emphasis>
|
||
Eastern Pacific: presently known only from the
|
||
<typeStatus id="54A98861FF9FFF83FC7821E5FC679D80" box="[912,960,189,213]" pageId="11" pageNumber="12">type</typeStatus>
|
||
locality on the
|
||
<taxonomicName id="4C124D40FF9FFF83FB9E21E4FB6F9D81" box="[1142,1224,188,212]" class="Malacostraca" family="Alpheidae" genus="Leptalpheus" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="19" phylum="Arthropoda" rank="species" species="azuero" status="sp. nov.">Azuero</taxonomicName>
|
||
Peninsula of
|
||
<collectingCountry id="F3057653FF9FFF83FA8121E5FF6E9DA2" name="Panama" pageId="11" pageNumber="12">Panama</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BAD36C3FF9FFF83FF2F205CFCDC9C31" blockId="11.[151,1437,151,1076]" pageId="11" pageNumber="12">
|
||
<emphasis id="B966EAD1FF9FFF83FF2F205CFE8C9C49" bold="true" box="[199,299,260,284]" pageId="11" pageNumber="12">Ecology.</emphasis>
|
||
Shallow sand-mud flat near mangroves, in burrows of unknown hosts; several species of
|
||
<taxonomicName id="4C124D40FF9FFF83FA89205DFE8F9C15" class="Malacostraca" family="Callianassidae" genus="Lepidophthalmus" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="12" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B966EAD1FF9FFF83FA89205DFE8F9C15" italics="true" pageId="11" pageNumber="12">Lepidophthalmus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C124D40FF9FFF83FEDD2070FE039C15" box="[309,420,296,320]" class="Malacostraca" family="Upogebiidae" genus="Upogebia" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="12" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B966EAD1FF9FFF83FEDD2070FE039C15" box="[309,420,296,320]" italics="true" pageId="11" pageNumber="12">Upogebia</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C124D40FF9FFF83FE312070FDEB9C15" box="[473,588,296,320]" class="Malacostraca" family="Laomediidae" genus="Axianassa" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="12" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B966EAD1FF9FFF83FE312070FDEB9C15" box="[473,588,296,320]" italics="true" pageId="11" pageNumber="12">Axianassa</emphasis>
|
||
</taxonomicName>
|
||
were also collected at this site, but never together with specimens of
|
||
<taxonomicName id="4C124D40FF9FFF83FAA72071FF1C9C31" class="Malacostraca" family="Alpheidae" genus="Leptalpheus" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="azuero" status="sp. nov.">
|
||
<emphasis id="B966EAD1FF9FFF83FAA72071FF1C9C31" italics="true" pageId="11" pageNumber="12">L. azuero</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B966EAD1FF9FFF83FF2A2014FEBB9C31" bold="true" box="[194,284,332,356]" pageId="11" pageNumber="12">
|
||
<taxonomicNameLabel id="A25557AAFF9FFF83FF2A2014FEBB9C31" box="[194,284,332,356]" pageId="11" pageNumber="12" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
(A. Anker, pers. obs.; Dworschak & Anker, in study).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3086548FF9FFF83FF2F2028FB899961" pageId="11" pageNumber="12" type="discussion">
|
||
<paragraph id="8BAD36C3FF9FFF83FF2F2028FD4F9FBA" blockId="11.[151,1437,151,1076]" pageId="11" pageNumber="12">
|
||
<emphasis id="B966EAD1FF9FFF83FF2F2028FE9C9CDD" bold="true" box="[199,315,368,392]" pageId="11" pageNumber="12">Remarks.</emphasis>
|
||
<taxonomicName id="4C124D40FF9FFF83FEA92028FDBC9CDD" box="[321,539,368,392]" class="Malacostraca" family="Alpheidae" genus="Leptalpheus" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="azuero" status="sp. nov.">
|
||
<emphasis id="B966EAD1FF9FFF83FEA92028FDBC9CDD" box="[321,539,368,392]" italics="true" pageId="11" pageNumber="12">Leptalpheus azuero</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B966EAD1FF9FFF83FDCA2028FDDD9CDD" bold="true" box="[546,634,368,392]" pageId="11" pageNumber="12">
|
||
<taxonomicNameLabel id="A25557AAFF9FFF83FDCA2028FDDD9CDD" box="[546,634,368,392]" pageId="11" pageNumber="12" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
is morphologically closest to the western Atlantic
|
||
<taxonomicName id="4C124D40FF9FFF83FB472029FAE09CDD" box="[1199,1351,369,392]" class="Malacostraca" family="Alpheidae" genus="Leptalpheus" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="axianassae">
|
||
<emphasis id="B966EAD1FF9FFF83FB472029FAE09CDD" box="[1199,1351,369,392]" italics="true" pageId="11" pageNumber="12">L. axianassae</emphasis>
|
||
</taxonomicName>
|
||
, differing from the latter by the more produced rostral projection; the presence of a small spiniform seta on the third and fourth pereiopods; and the armature on the major chela fingers, especially the more developed proximal tooth of the pollex and the presence of an additional subdistal tooth near the pollex tip (cf.
|
||
<figureCitation id="13292A46FF9FFF83FBA22084FB299CA1" box="[1098,1166,476,500]" captionStart="FIGURE 6" captionStartId="10.[151,250,1281,1304]" captionTargetBox="[169,1409,208,1241]" captionTargetId="figure@10.[157,1430,193,1260]" captionTargetPageId="10" captionText="FIGURE 6. Leptalpheus azuero sp. nov., paratype, female from Playa El Aguillito, Azuero Peninsula, Pacific coast of Panama (RMNH D 54560): A, major cheliped, lateral view; B, same, mesial view; C, same, carpus and chela, lateral view; D, major chela fingers, lateral view; E, minor cheliped, lateral view; F, same, carpus and chela, mesial view." httpUri="https://zenodo.org/record/278808/files/figure.png" pageId="11" pageNumber="12">Fig. 6</figureCitation>
|
||
C, D and
|
||
<bibRefCitation id="EF834B32FF9FFF83FB162084FE8D9F42" author="Dworschak" pageId="11" pageNumber="12" refString="Dworschak, P. C. & Coelho, V. R. (1999) On two alpheids from Araca (Sao Paulo, Brazil) with a description of a new species of Leptalpheus (Decapoda: Caridea: Alpheidae). Annalen des Naturhistorischen Museums in Wien, 101 B, 475 - 488." type="journal article" year="1999">Dworschak & Coelho 1999</bibRefCitation>
|
||
, figs. 18, 19).
|
||
<taxonomicName id="4C124D40FF9FFF83FE242358FD0F9F4D" box="[460,680,512,536]" class="Malacostraca" family="Alpheidae" genus="Leptalpheus" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="azuero" status="sp. nov.">
|
||
<emphasis id="B966EAD1FF9FFF83FE242358FD0F9F4D" box="[460,680,512,536]" italics="true" pageId="11" pageNumber="12">Leptalpheus azuero</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B966EAD1FF9FFF83FD472358FCAD9F4D" bold="true" box="[687,778,512,536]" pageId="11" pageNumber="12">
|
||
<taxonomicNameLabel id="A25557AAFF9FFF83FD472358FCAD9F4D" box="[687,778,512,536]" pageId="11" pageNumber="12" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
also shows some affinities to the species of the Indo-West Pacific
|
||
<taxonomicName id="4C124D40FF9FFF83FF04237DFED79F69" box="[236,368,548,572]" class="Malacostraca" family="Alpheidae" genus="Leptalpheus" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="pacificus">
|
||
<emphasis id="B966EAD1FF9FFF83FF04237DFED79F69" box="[236,368,548,572]" italics="true" pageId="11" pageNumber="12">L. pacificus</emphasis>
|
||
</taxonomicName>
|
||
complex, viz.
|
||
<taxonomicName id="4C124D40FF9FFF83FDFF237DFD3C9F69" box="[535,667,548,572]" class="Malacostraca" family="Alpheidae" genus="Leptalpheus" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="pacificus">
|
||
<emphasis id="B966EAD1FF9FFF83FDFF237DFD3C9F69" box="[535,667,548,572]" italics="true" pageId="11" pageNumber="12">L. pacificus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C124D40FF9FFF83FD41237DFCF79F69" box="[681,848,548,572]" class="Malacostraca" family="Alpheidae" genus="Leptalpheus" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="denticulatus">
|
||
<emphasis id="B966EAD1FF9FFF83FD41237DFCF79F69" box="[681,848,548,572]" italics="true" pageId="11" pageNumber="12">L. denticulatus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C124D40FF9FFF83FC60237DFB809F69" box="[904,1063,548,572]" class="Malacostraca" family="Alpheidae" genus="Leptalpheus" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="dworschaki">
|
||
<emphasis id="B966EAD1FF9FFF83FC60237DFB809F69" box="[904,1063,548,572]" italics="true" pageId="11" pageNumber="12">L. dworschaki</emphasis>
|
||
</taxonomicName>
|
||
, but can be separated from these three species by the frontal margin produced anteriorly into a broadly triangular rostral projection (rounded in the other species); the longer stylocerite, exceeding distal margin of the first article of the antennular peduncle (not reaching this margin in the other species); the second article of the antennular peduncle being wider than long (much longer than wide in the other species); and the configuration of teeth on the pollex of the major cheliped (cf.
|
||
<figureCitation id="13292A46FF9FFF83FF7F238FFF439FBA" box="[151,228,727,751]" captionStart="FIGURE 5" captionStartId="9.[151,250,1808,1831]" captionTargetBox="[173,1407,200,1771]" captionTargetId="figure@9.[151,1436,193,1787]" captionTargetPageId="9" captionText="FIGURE 5. Leptalpheus azuero sp. nov.: all except I—paratype, female from Playa El Aguillito, Azuero Peninsula, Pacific coast of Panama (RMNH D 54560); I—paratype, male from the same locality (MNHN-IU- 2011 - 5240): A, frontal region, dorsal view; B, same, lateral view; C, telson, dorsal view; D, ventromesial tooth on first article of antennular peduncle, lateral view; E, scaphocerite, dorsal view; F, third maxilliped, lateral view; G, second pereiopod, lateral view; H, third pereiopod, lateral view; I, appendix masculina and appendix interna, dorsal view; J, uropod, dorsal view (setae omitted)." httpUri="https://zenodo.org/record/278807/files/figure.png" pageId="11" pageNumber="12">Figs. 5</figureCitation>
|
||
,
|
||
<figureCitation id="13292A46FF9FFF83FF19238FFF589FBA" box="[241,255,727,751]" captionStart="FIGURE 6" captionStartId="10.[151,250,1281,1304]" captionTargetBox="[169,1409,208,1241]" captionTargetId="figure@10.[157,1430,193,1260]" captionTargetPageId="10" captionText="FIGURE 6. Leptalpheus azuero sp. nov., paratype, female from Playa El Aguillito, Azuero Peninsula, Pacific coast of Panama (RMNH D 54560): A, major cheliped, lateral view; B, same, mesial view; C, same, carpus and chela, lateral view; D, major chela fingers, lateral view; E, minor cheliped, lateral view; F, same, carpus and chela, mesial view." httpUri="https://zenodo.org/record/278808/files/figure.png" pageId="11" pageNumber="12">6</figureCitation>
|
||
and
|
||
<bibRefCitation id="EF834B32FF9FFF83FEDD238FFD819FBA" author="Anker" box="[309,550,727,751]" pageId="11" pageNumber="12" refString="Anker A, Marin, I. N. (2009) The alpheid shrimp genus Leptalpheus Williams, 1965 in the tropical western Pacific, with descriptions of two new species (Crustacea: Decapoda: Caridea). The Raffles Bulletin of Zoology, 57, 91 - 107." type="journal article" year="2009">Anker & Marin 2009</bibRefCitation>
|
||
, figs. 4, 7d, 11d).
|
||
</paragraph>
|
||
<paragraph id="8BAD36C3FF9FFF83FF2D23A4FB899961" blockId="11.[151,1437,151,1076]" pageId="11" pageNumber="12">
|
||
<taxonomicName id="4C124D40FF9FFF83FF2D23A4FE389E41" box="[197,415,764,788]" class="Malacostraca" family="Alpheidae" genus="Leptalpheus" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="azuero" status="sp. nov.">
|
||
<emphasis id="B966EAD1FF9FFF83FF2D23A4FE389E41" box="[197,415,764,788]" italics="true" pageId="11" pageNumber="12">Leptalpheus azuero</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B966EAD1FF9FFF83FE4D23A4FE599E41" bold="true" box="[421,510,764,788]" pageId="11" pageNumber="12">
|
||
<taxonomicNameLabel id="A25557AAFF9FFF83FE4D23A4FE599E41" box="[421,510,764,788]" pageId="11" pageNumber="12" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
appears to be one of the smallest species of the genus based on the size of the present specimens. The
|
||
<typeStatus id="54A98861FF9FFF83FE9F2279FE009E6C" box="[375,423,801,825]" pageId="11" pageNumber="12">type</typeStatus>
|
||
series consists of a single, apparently adult male (cl
|
||
<quantity id="4CEA9B26FF9FFF83FC142247FBF49E62" box="[1020,1107,799,823]" metricMagnitude="-3" metricUnit="m" metricValue="3.7" pageId="11" pageNumber="12" unit="mm" value="3.7">3.7 mm</quantity>
|
||
) and several, possibly young or subadult, non-ovigerous females (cl
|
||
<quantity id="4CEA9B26FF9FFF83FDB8221CFD7C9E09" box="[592,731,836,860]" metricMagnitude="-3" metricUnit="m" metricValue="2.65" metricValueMax="2.9" metricValueMin="2.4" pageId="11" pageNumber="12" unit="mm" value="2.65" valueMax="2.9" valueMin="2.4">2.4–2.9 mm</quantity>
|
||
). The male is missing its major cheliped, but has a well-developed appendix masculina (
|
||
<figureCitation id="13292A46FF9FFF83FE2E223FFDAE9E2A" box="[454,521,871,895]" captionStart="FIGURE 5" captionStartId="9.[151,250,1808,1831]" captionTargetBox="[173,1407,200,1771]" captionTargetId="figure@9.[151,1436,193,1787]" captionTargetPageId="9" captionText="FIGURE 5. Leptalpheus azuero sp. nov.: all except I—paratype, female from Playa El Aguillito, Azuero Peninsula, Pacific coast of Panama (RMNH D 54560); I—paratype, male from the same locality (MNHN-IU- 2011 - 5240): A, frontal region, dorsal view; B, same, lateral view; C, telson, dorsal view; D, ventromesial tooth on first article of antennular peduncle, lateral view; E, scaphocerite, dorsal view; F, third maxilliped, lateral view; G, second pereiopod, lateral view; H, third pereiopod, lateral view; I, appendix masculina and appendix interna, dorsal view; J, uropod, dorsal view (setae omitted)." httpUri="https://zenodo.org/record/278807/files/figure.png" pageId="11" pageNumber="12">
|
||
Fig.
|
||
<date id="FFAC1003FF9FFF83FE13223FFDAE9E2A" box="[507,521,871,895]" pageId="11" pageNumber="12">5</date>
|
||
</figureCitation>
|
||
I) and is otherwise very similar to the females. Four females, including the
|
||
<typeStatus id="54A98861FF9FFF83FA88223FFF6D9EF0" pageId="11" pageNumber="12" type="holotype">holotype</typeStatus>
|
||
, have a well-developed, species-diagnostic major cheliped, with very little variation in the armature of the chela fingers (
|
||
<figureCitation id="13292A46FF9FFF83FEDF22F7FEDC9E92" box="[311,379,943,967]" captionStart="FIGURE 6" captionStartId="10.[151,250,1281,1304]" captionTargetBox="[169,1409,208,1241]" captionTargetId="figure@10.[157,1430,193,1260]" captionTargetPageId="10" captionText="FIGURE 6. Leptalpheus azuero sp. nov., paratype, female from Playa El Aguillito, Azuero Peninsula, Pacific coast of Panama (RMNH D 54560): A, major cheliped, lateral view; B, same, mesial view; C, same, carpus and chela, lateral view; D, major chela fingers, lateral view; E, minor cheliped, lateral view; F, same, carpus and chela, mesial view." httpUri="https://zenodo.org/record/278808/files/figure.png" pageId="11" pageNumber="12">Fig. 6</figureCitation>
|
||
D). The
|
||
<typeStatus id="54A98861FF9FFF83FE3122E9FDAE9E9C" box="[473,521,945,969]" pageId="11" pageNumber="12">type</typeStatus>
|
||
specimens of another eastern Pacific species,
|
||
<taxonomicName id="4C124D40FF9FFF83FBFA22E9FB019E9D" box="[1042,1190,945,968]" class="Malacostraca" family="Alpheidae" genus="Leptalpheus" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="mexicanus">
|
||
<emphasis id="B966EAD1FF9FFF83FBFA22E9FB019E9D" box="[1042,1190,945,968]" italics="true" pageId="11" pageNumber="12">L. mexicanus</emphasis>
|
||
</taxonomicName>
|
||
, range from
|
||
<quantity id="4CEA9B26FF9FFF83FAD022F7FA3C9E9D" box="[1336,1435,943,968]" metricMagnitude="-3" metricUnit="m" metricValue="2.88" pageId="11" pageNumber="12" unit="mm" value="2.88">2.88 mm</quantity>
|
||
to
|
||
<quantity id="4CEA9B26FF9FFF83FF5C228CFEBE9EB9" box="[180,281,980,1004]" metricMagnitude="-3" metricUnit="m" metricValue="3.54" pageId="11" pageNumber="12" unit="mm" value="3.54">3.54 mm</quantity>
|
||
(
|
||
<bibRefCitation id="EF834B32FF9FFF83FEC0228CFD979EB9" author="Rios" box="[296,560,980,1004]" pageId="11" pageNumber="12" refString="Rios R. & Carvacho A. (1983) Caridean shrimps of the Gulf of California III. Leptalpheus mexicanus, new species (Crustacea, Decapoda, Alpheidae). Journal of Crustacean Biology, 3, 306 - 313." type="journal article" year="1983">Ríos & Carvacho 1983</bibRefCitation>
|
||
), i.e. size range comparable to the
|
||
<typeStatus id="54A98861FF9FFF83FC51228DFC4E9EB8" box="[953,1001,981,1005]" pageId="11" pageNumber="12">type</typeStatus>
|
||
specimens of
|
||
<taxonomicName id="4C124D40FF9FFF83FB64228DFB519EB9" box="[1164,1270,981,1004]" class="Malacostraca" family="Alpheidae" genus="Leptalpheus" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="azuero" status="sp. nov.">
|
||
<emphasis id="B966EAD1FF9FFF83FB64228DFB519EB9" box="[1164,1270,981,1004]" italics="true" pageId="11" pageNumber="12">L. azuero</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B966EAD1FF9FFF83FB15228CFAFF9EB9" bold="true" box="[1277,1368,980,1004]" pageId="11" pageNumber="12">
|
||
<taxonomicNameLabel id="A25557AAFF9FFF83FB15228CFAFF9EB9" box="[1277,1368,980,1004]" pageId="11" pageNumber="12" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
More sampling of infaunal shrimps is planned at the
|
||
<typeStatus id="54A98861FF9FFF83FD4A22A1FD759944" box="[674,722,1017,1041]" pageId="11" pageNumber="12">type</typeStatus>
|
||
locality and other areas of the Pacific coast of
|
||
<collectingCountry id="F3057653FF9FFF83FB3722A1FA9F9945" box="[1247,1336,1017,1040]" name="Panama" pageId="11" pageNumber="12">Panama</collectingCountry>
|
||
to determine the maximum size of
|
||
<taxonomicName id="4C124D40FF9FFF83FE202545FD969961" box="[456,561,1053,1076]" class="Malacostraca" family="Alpheidae" genus="Leptalpheus" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="azuero" status="sp. nov.">
|
||
<emphasis id="B966EAD1FF9FFF83FE202545FD969961" box="[456,561,1053,1076]" italics="true" pageId="11" pageNumber="12">L. azuero</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B966EAD1FF9FFF83FDD02544FD359961" bold="true" box="[568,658,1052,1076]" pageId="11" pageNumber="12">
|
||
<taxonomicNameLabel id="A25557AAFF9FFF83FDD02544FD359961" box="[568,658,1052,1076]" pageId="11" pageNumber="12" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
and also to identify its host or hosts.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |