231 lines
45 KiB
XML
231 lines
45 KiB
XML
<document id="69611D5AC29E43DE38F63F5479B91AED" ID-CLB-Dataset="25859" ID-DOI="10.11646/zootaxa.4661.3.9" ID-GBIF-Dataset="05e4bcce-91c1-4172-84cd-384cb981987c" ID-ISSN="1175-5326" ID-Zenodo-Dep="3380981" ID-ZooBank="64DCCEC6-AD9D-4546-983B-8919E7D9F501" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1567061293248" checkinUser="plazi" docAuthor="Lukashevich, Elena D." docDate="2019" docId="03BB879EFF89E97E46C942AE4476333A" docLanguage="en" docName="zootaxa.4661.3.9.pdf" docOrigin="Zootaxa 4661 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Zhiganka Lukashevich 1995" docType="treatment" docVersion="10" lastPageNumber="568" masterDocId="FF82FFE6FF88E97C465E447B46553761" masterDocTitle="Review of the fossil record of Bittacomorphinae (Diptera: Ptychopteridae)" masterLastPageNumber="578" masterPageNumber="566" pageNumber="567" updateTime="1698753122357" updateUser="ExternalLinkService">
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<mods:title id="30F42CE35DC38308DA6C1478C35A424E">Review of the fossil record of Bittacomorphinae (Diptera: Ptychopteridae)</mods:title>
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<treatment id="03BB879EFF89E97E46C942AE4476333A" ID-DOI="http://doi.org/10.5281/zenodo.5936279" ID-GBIF-Taxon="159404833" ID-Zenodo-Dep="5936279" LSID="urn:lsid:plazi:treatment:03BB879EFF89E97E46C942AE4476333A" httpUri="http://treatment.plazi.org/id/03BB879EFF89E97E46C942AE4476333A" lastPageId="2" lastPageNumber="568" pageId="1" pageNumber="567">
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<subSubSection id="C3086503FF89E97D46C942AE440B318E" box="[151,606,1749,1776]" pageId="1" pageNumber="567" type="nomenclature">
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<paragraph id="8BAD3688FF89E97D46C942AE440B318E" blockId="1.[151,606,1749,1812]" box="[151,606,1749,1776]" pageId="1" pageNumber="567">
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<heading id="D0E581E4FF89E97D46C942AE440B318E" bold="true" box="[151,606,1749,1776]" fontSize="11" level="1" pageId="1" pageNumber="567" reason="1">
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<emphasis id="B966EA9AFF89E97D46C942AE440B318E" bold="true" box="[151,606,1749,1776]" pageId="1" pageNumber="567">
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Genus:
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<taxonomicName id="4C124D0BFF89E97D46A642AE440B318E" ID-CoL="89RRR" authority="Lukashevich, 1995" authorityName="Lukashevich" authorityYear="1995" box="[248,606,1749,1776]" class="Insecta" family="Ptychopteridae" genus="Zhiganka" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="567" phylum="Arthropoda" rank="genus">
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<emphasis id="B966EA9AFF89E97D46A642AE47383191" bold="true" box="[248,365,1749,1776]" italics="true" pageId="1" pageNumber="567">Zhiganka</emphasis>
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<bibRefCitation id="EF834B79FF89E97D472B42AE440B318E" author="Lukashevich, E. D." box="[373,606,1749,1775]" pageId="1" pageNumber="567" pagination="164 - 170" refId="ref7214" refString="Lukashevich, E. D. (1995) First pupae of the Eoptychopteridae and Ptychopteridae from the Mesozoic of Siberia (Insecta: Diptera). Paleontological Journal, 29 (4), 164 - 170." type="journal article" year="1995">Lukashevich, 1995</bibRefCitation>
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</taxonomicName>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C3086503FF89E97D46C9428247E73075" box="[151,434,1785,1812]" pageId="1" pageNumber="567" type="description">
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<paragraph id="8BAD3688FF89E97D46C9428247E73075" blockId="1.[151,606,1749,1812]" box="[151,434,1785,1812]" pageId="1" pageNumber="567">
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(
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<figureCitation id="13292A0DFF89E97D46C1428247723072" box="[159,295,1785,1811]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStart-3="FIGURE 4" captionStartId-0="3.[151,250,1902,1927]" captionStartId-1="4.[151,250,1869,1894]" captionStartId-2="5.[151,250,1787,1812]" captionStartId-3="6.[151,250,1833,1858]" captionTargetBox-0="[201,1387,166,1882]" captionTargetBox-1="[187,1399,166,1849]" captionTargetBox-2="[214,1364,173,1767]" captionTargetBox-3="[227,1361,166,1813]" captionTargetId-0="figure@3.[201,1387,166,1882]" captionTargetId-1="figure@4.[187,1399,166,1849]" captionTargetId-2="figure@5.[214,1372,166,1767]" captionTargetId-3="figure@6.[227,1361,166,1813]" captionTargetPageId-0="3" captionTargetPageId-1="4" captionTargetPageId-2="5" captionTargetPageId-3="6" captionText-0="FIGURE 1. Zhiganka longialata sp. nov.: A–E. Male PIN 5340/953, holotype:A. Head and thorax. B. Habitus. C. Two spurs on hind tibia. D, E. Two spurs on left and right midtibia. F. Spur on fore tibia, PIN 5340/1494. G. Habitus of female PIN 5340/951, paratype. H. Terminalia of female PIN 5026/1595. Abbreviations: cl, clypeus; lbl, labellum; lbr, labrum; mp, maxillary palp; s, scutal suture; sl, scutellum." captionText-1="FIGURE 2. Zhiganka longialata sp. nov., male PIN 5026/1599, paratype: A. Anterior part of body. B. Wing. C. Proximal flagellomeres of antenna, inset—distal flagellomeres, arrows mark setae. D. Head. E, F. Terminalia, part and counterpart. G, Distal part of wing. Abbreviations: apn, antepronotum; cl, clypeus; ep, epandrium; gcx, gonocoxite; gst, gonostylus; hyp, hypandrium; lbl, labellum; lbr, labrum; mp, maxillary palp; ss, surstylus." captionText-2="FIGURE 3. Zhiganka longialata sp. nov.: A–C. Paratype PIN 5026/1599: A. Anterior part of body. B. Distal and proximal flagellomeres of antenna. C. Wing. D. Abdomen of female PIN 5026/1597, dorsal view. E. Abdomen of male PIN 5340/807, lateral view (flipped horizontally). F–H. Genitalia of male. F. Paratype PIN 5026/1599. G. Paratype PIN 5026/1594. H. Paratype PIN 5340/807. Scale bars equal 2 mm (A, C, D, E), 200 μm (B, F, G, H). Abbreviations: apn, antepronotum; b, transepimeral suture; cl, clypeus; cx, coxa; ep, epandrium; gcx, gonocoxite; gst, gonostylus; hyp, hypandrium; lbl, labellum; lbr, labrum; mp, maxillary palp; ps, prescutal suture; s, scutal suture; ss, surstylus." captionText-3="FIGURE 4. Zhiganka longialata sp. nov.: A, B. Anterior and basal part of wing, PIN 5340/1494, part and counterpart. C. Part of wing, female PIN 5340/951. D. Genitalia of male PIN 5026/1594. E. Genitalia of male PIN 5340/953. F. Thorax and head, female PIN 5340/952. Abbreviations: aem, anepimerum or pteropleurite; apn, antepronotum; b, transepimeral suture; cx, coxa; ep, epandrium; gcx, gonocoxite; gst, gonostylus; hyp, hypandrium; lc, laterocervicale; mt, mediotergum, mpl, meropleurum; ps, prescutal suture; pt, pleurotergum; s, scutal suture; sl, scutellum; ss, surstylus." httpUri-0="https://zenodo.org/record/3380983/files/figure.png" httpUri-1="https://zenodo.org/record/3380985/files/figure.png" httpUri-2="https://zenodo.org/record/3380987/files/figure.png" httpUri-3="https://zenodo.org/record/3380989/files/figure.png" pageId="1" pageNumber="567">Figures 1–4</figureCitation>
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,
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<figureCitation id="13292A0DFF89E97D476A4282470D3075" box="[308,344,1785,1812]" captionStart="FIGURE 5" captionStartId="7.[151,250,1936,1961]" captionTargetBox="[151,1417,526,1917]" captionTargetId="figure@7.[151,1417,526,1917]" captionTargetPageId="7" captionText="FIGURE 5. SEM images:A. Zhiganka longialata sp. nov., holotype PIN 5340/953, wing; Khasurty, K 1. B. Zhiganka comitans, holotype PIN 4502/13, Zhigansk, K 1. C. Crenoptychoptera antica, PIN 1255/2123, wing; Kubekovo, J 2. Arrows mark spurious vein. Scale bars equal 2 mm." httpUri="https://zenodo.org/record/3380991/files/figure.png" pageId="1" pageNumber="567">5A</figureCitation>
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,
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<figureCitation id="13292A0DFF89E97D473A428247243075" box="[356,369,1785,1812]" captionStart="FIGURE 6" captionStartId="8.[151,250,1675,1700]" captionTargetBox="[151,1436,174,1652]" captionTargetId="figure@8.[151,1436,174,1655]" captionTargetPageId="8" captionText="FIGURE 6. SEM images: A–D. Zhiganka longialata sp. nov., wing of holotype PIN 5340/953, Zhigansk, K 1: A. Distal part. B. Part of wing. C, D. Fore and hind margin of wing. E, F. Crenoptychoptera antica, apical and middle part of wing PIN 1255/2123, Kubekovo, J 2. Black arrows mark spurious vein, white—setae on veins and membrane." httpUri="https://zenodo.org/record/3380993/files/figure.png" pageId="1" pageNumber="567">6</figureCitation>
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A–D)
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</paragraph>
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</subSubSection>
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<subSubSection id="C3086503FF89E97D46C9434545183037" box="[151,845,1853,1878]" pageId="1" pageNumber="567" type="type_taxon">
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<paragraph id="8BAD3688FF89E97D46C9434545183037" blockId="1.[151,845,1853,1878]" box="[151,845,1853,1878]" pageId="1" pageNumber="567">
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<emphasis id="B966EA9AFF89E97D46C9434547763037" bold="true" box="[151,291,1853,1878]" pageId="1" pageNumber="567">
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<typeStatus id="54A9882AFF89E97D46C9434546993037" box="[151,204,1854,1878]" pageId="1" pageNumber="567">Type</typeStatus>
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species.
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</emphasis>
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<taxonomicName id="4C124D0BFF89E97D4774434544BD3037" ID-CoL="8FP3G" authority="Lukashevich, 1995: 170" authorityName="Lukashevich" authorityPageNumber="170" authorityYear="1995" box="[298,744,1853,1878]" class="Insecta" family="Ptychopteridae" genus="Zhiganka" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="567" phylum="Arthropoda" rank="species" species="comitans">
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<emphasis id="B966EA9AFF89E97D4774434547BE3034" box="[298,491,1854,1878]" italics="true" pageId="1" pageNumber="567">Zhiganka comitans</emphasis>
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<bibRefCitation id="EF834B79FF89E97D47AC434644BD3037" author="Lukashevich, E. D." box="[498,744,1853,1878]" pageId="1" pageNumber="567" pagination="164 - 170" refId="ref7214" refString="Lukashevich, E. D. (1995) First pupae of the Eoptychopteridae and Ptychopteridae from the Mesozoic of Siberia (Insecta: Diptera). Paleontological Journal, 29 (4), 164 - 170." type="journal article" year="1995">Lukashevich, 1995: 170</bibRefCitation>
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</taxonomicName>
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, fig. 1e,g.
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</paragraph>
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</subSubSection>
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||
<subSubSection id="C3086503FF89E97E46C943F8446035CA" lastPageId="2" lastPageNumber="568" pageId="1" pageNumber="567" type="diagnosis">
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<paragraph id="8BAD3688FF89E97E46C943F842EE357A" blockId="1.[151,1437,1923,2023]" lastBlockId="2.[151,1437,153,1115]" lastPageId="2" lastPageNumber="568" pageId="1" pageNumber="567">
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<emphasis id="B966EA9AFF89E97D46C943F847C330FF" bold="true" box="[151,406,1923,1950]" pageId="1" pageNumber="567">Diagnosis (emended).</emphasis>
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Adult. Medium-size gracile flies,
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<quantity id="4CEA9B6DFF89E97D454B43FF45D530FE" box="[789,896,1924,1951]" metricMagnitude="-2" metricUnit="m" metricValue="1.1" metricValueMax="1.3" metricValueMin="0.9" pageId="1" pageNumber="567" unit="mm" value="11.0" valueMax="13.0" valueMin="9.0">9–13 mm</quantity>
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long (
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<figureCitation id="13292A0DFF89E97D459643FF421730FF" box="[968,1090,1924,1950]" captionStart="FIGURE 1" captionStartId="3.[151,250,1902,1927]" captionTargetBox="[201,1387,166,1882]" captionTargetId="figure@3.[201,1387,166,1882]" captionTargetPageId="3" captionText="FIGURE 1. Zhiganka longialata sp. nov.: A–E. Male PIN 5340/953, holotype:A. Head and thorax. B. Habitus. C. Two spurs on hind tibia. D, E. Two spurs on left and right midtibia. F. Spur on fore tibia, PIN 5340/1494. G. Habitus of female PIN 5340/951, paratype. H. Terminalia of female PIN 5026/1595. Abbreviations: cl, clypeus; lbl, labellum; lbr, labrum; mp, maxillary palp; s, scutal suture; sl, scutellum." httpUri="https://zenodo.org/record/3380983/files/figure.png" pageId="1" pageNumber="567">Fig. 1B, G</figureCitation>
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). Head not appressed to thorax (
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<figureCitation id="13292A0DFF89E97D46C143D3474A30A2" box="[159,287,1960,1987]" captionStart="FIGURE 2" captionStartId="4.[151,250,1869,1894]" captionTargetBox="[187,1399,166,1849]" captionTargetId="figure@4.[187,1399,166,1849]" captionTargetPageId="4" captionText="FIGURE 2. Zhiganka longialata sp. nov., male PIN 5026/1599, paratype: A. Anterior part of body. B. Wing. C. Proximal flagellomeres of antenna, inset—distal flagellomeres, arrows mark setae. D. Head. E, F. Terminalia, part and counterpart. G, Distal part of wing. Abbreviations: apn, antepronotum; cl, clypeus; ep, epandrium; gcx, gonocoxite; gst, gonostylus; hyp, hypandrium; lbl, labellum; lbr, labrum; mp, maxillary palp; ss, surstylus." httpUri="https://zenodo.org/record/3380985/files/figure.png" pageId="1" pageNumber="567">Figs 2A, D</figureCitation>
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,
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<figureCitation id="13292A0DFF89E97D477543D3470530A3" box="[299,336,1960,1986]" captionStart="FIGURE 3" captionStartId="5.[151,250,1787,1812]" captionTargetBox="[214,1364,173,1767]" captionTargetId="figure@5.[214,1372,166,1767]" captionTargetPageId="5" captionText="FIGURE 3. Zhiganka longialata sp. nov.: A–C. Paratype PIN 5026/1599: A. Anterior part of body. B. Distal and proximal flagellomeres of antenna. C. Wing. D. Abdomen of female PIN 5026/1597, dorsal view. E. Abdomen of male PIN 5340/807, lateral view (flipped horizontally). F–H. Genitalia of male. F. Paratype PIN 5026/1599. G. Paratype PIN 5026/1594. H. Paratype PIN 5340/807. Scale bars equal 2 mm (A, C, D, E), 200 μm (B, F, G, H). Abbreviations: apn, antepronotum; b, transepimeral suture; cl, clypeus; cx, coxa; ep, epandrium; gcx, gonocoxite; gst, gonostylus; hyp, hypandrium; lbl, labellum; lbr, labrum; mp, maxillary palp; ps, prescutal suture; s, scutal suture; ss, surstylus." httpUri="https://zenodo.org/record/3380987/files/figure.png" pageId="1" pageNumber="567">3A</figureCitation>
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). Antenna with 19 flagellomeres, each with distinct scattered (4?) macrosetae and short pubescence, apical flagellomere reduced to button-like structure (
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<figureCitation id="13292A0DFF89E97D455243B745DF3087" box="[780,906,1996,2022]" captionStart="FIGURE 2" captionStartId="4.[151,250,1869,1894]" captionTargetBox="[187,1399,166,1849]" captionTargetId="figure@4.[187,1399,166,1849]" captionTargetPageId="4" captionText="FIGURE 2. Zhiganka longialata sp. nov., male PIN 5026/1599, paratype: A. Anterior part of body. B. Wing. C. Proximal flagellomeres of antenna, inset—distal flagellomeres, arrows mark setae. D. Head. E, F. Terminalia, part and counterpart. G, Distal part of wing. Abbreviations: apn, antepronotum; cl, clypeus; ep, epandrium; gcx, gonocoxite; gst, gonostylus; hyp, hypandrium; lbl, labellum; lbr, labrum; mp, maxillary palp; ss, surstylus." httpUri="https://zenodo.org/record/3380985/files/figure.png" pageId="1" pageNumber="567">Figs 2A, C</figureCitation>
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,
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<figureCitation id="13292A0DFF89E97D45C843B745EC3087" box="[918,953,1996,2022]" captionStart="FIGURE 3" captionStartId="5.[151,250,1787,1812]" captionTargetBox="[214,1364,173,1767]" captionTargetId="figure@5.[214,1372,166,1767]" captionTargetPageId="5" captionText="FIGURE 3. Zhiganka longialata sp. nov.: A–C. Paratype PIN 5026/1599: A. Anterior part of body. B. Distal and proximal flagellomeres of antenna. C. Wing. D. Abdomen of female PIN 5026/1597, dorsal view. E. Abdomen of male PIN 5340/807, lateral view (flipped horizontally). F–H. Genitalia of male. F. Paratype PIN 5026/1599. G. Paratype PIN 5026/1594. H. Paratype PIN 5340/807. Scale bars equal 2 mm (A, C, D, E), 200 μm (B, F, G, H). Abbreviations: apn, antepronotum; b, transepimeral suture; cl, clypeus; cx, coxa; ep, epandrium; gcx, gonocoxite; gst, gonostylus; hyp, hypandrium; lbl, labellum; lbr, labrum; mp, maxillary palp; ps, prescutal suture; s, scutal suture; ss, surstylus." httpUri="https://zenodo.org/record/3380987/files/figure.png" pageId="1" pageNumber="567">3B</figureCitation>
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). Clypeus acutely triangular, labrum small, sclerotized, labella large. Maxillary palpus elongate, with apical and preapical segments subequal (
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<figureCitation id="13292A0DFF8AE97E435B44E2433037D2" box="[1285,1381,153,179]" captionStart="FIGURE 1" captionStartId="3.[151,250,1902,1927]" captionTargetBox="[201,1387,166,1882]" captionTargetId="figure@3.[201,1387,166,1882]" captionTargetPageId="3" captionText="FIGURE 1. Zhiganka longialata sp. nov.: A–E. Male PIN 5340/953, holotype:A. Head and thorax. B. Habitus. C. Two spurs on hind tibia. D, E. Two spurs on left and right midtibia. F. Spur on fore tibia, PIN 5340/1494. G. Habitus of female PIN 5340/951, paratype. H. Terminalia of female PIN 5026/1595. Abbreviations: cl, clypeus; lbl, labellum; lbr, labrum; mp, maxillary palp; s, scutal suture; sl, scutellum." httpUri="https://zenodo.org/record/3380983/files/figure.png" pageId="2" pageNumber="568">Figs 1A</figureCitation>
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,
|
||
<figureCitation id="13292A0DFF8AE97E432D44E243CD37D2" box="[1395,1432,153,179]" captionStart="FIGURE 2" captionStartId="4.[151,250,1869,1894]" captionTargetBox="[187,1399,166,1849]" captionTargetId="figure@4.[187,1399,166,1849]" captionTargetPageId="4" captionText="FIGURE 2. Zhiganka longialata sp. nov., male PIN 5026/1599, paratype: A. Anterior part of body. B. Wing. C. Proximal flagellomeres of antenna, inset—distal flagellomeres, arrows mark setae. D. Head. E, F. Terminalia, part and counterpart. G, Distal part of wing. Abbreviations: apn, antepronotum; cl, clypeus; ep, epandrium; gcx, gonocoxite; gst, gonostylus; hyp, hypandrium; lbl, labellum; lbr, labrum; mp, maxillary palp; ss, surstylus." httpUri="https://zenodo.org/record/3380985/files/figure.png" pageId="2" pageNumber="568">2D</figureCitation>
|
||
,
|
||
<figureCitation id="13292A0DFF8AE97E46C944C646E837B6" box="[151,189,189,215]" captionStart="FIGURE 3" captionStartId="5.[151,250,1787,1812]" captionTargetBox="[214,1364,173,1767]" captionTargetId="figure@5.[214,1372,166,1767]" captionTargetPageId="5" captionText="FIGURE 3. Zhiganka longialata sp. nov.: A–C. Paratype PIN 5026/1599: A. Anterior part of body. B. Distal and proximal flagellomeres of antenna. C. Wing. D. Abdomen of female PIN 5026/1597, dorsal view. E. Abdomen of male PIN 5340/807, lateral view (flipped horizontally). F–H. Genitalia of male. F. Paratype PIN 5026/1599. G. Paratype PIN 5026/1594. H. Paratype PIN 5340/807. Scale bars equal 2 mm (A, C, D, E), 200 μm (B, F, G, H). Abbreviations: apn, antepronotum; b, transepimeral suture; cl, clypeus; cx, coxa; ep, epandrium; gcx, gonocoxite; gst, gonostylus; hyp, hypandrium; lbl, labellum; lbr, labrum; mp, maxillary palp; ps, prescutal suture; s, scutal suture; ss, surstylus." httpUri="https://zenodo.org/record/3380987/files/figure.png" pageId="2" pageNumber="568">3A</figureCitation>
|
||
). Antepronotum well developed, prescutal suture distinct, scutum with incomplete transverse suture, all pleural sclerites without setae, transepimeral suture present (
|
||
<figureCitation id="13292A0DFF8AE97E44AB449A4501379A" box="[757,852,225,251]" captionStart="FIGURE 2" captionStartId="4.[151,250,1869,1894]" captionTargetBox="[187,1399,166,1849]" captionTargetId="figure@4.[187,1399,166,1849]" captionTargetPageId="4" captionText="FIGURE 2. Zhiganka longialata sp. nov., male PIN 5026/1599, paratype: A. Anterior part of body. B. Wing. C. Proximal flagellomeres of antenna, inset—distal flagellomeres, arrows mark setae. D. Head. E, F. Terminalia, part and counterpart. G, Distal part of wing. Abbreviations: apn, antepronotum; cl, clypeus; ep, epandrium; gcx, gonocoxite; gst, gonostylus; hyp, hypandrium; lbl, labellum; lbr, labrum; mp, maxillary palp; ss, surstylus." httpUri="https://zenodo.org/record/3380985/files/figure.png" pageId="2" pageNumber="568">Figs 2D</figureCitation>
|
||
,
|
||
<figureCitation id="13292A0DFF8AE97E453C449A45D2379A" box="[866,903,225,251]" captionStart="FIGURE 3" captionStartId="5.[151,250,1787,1812]" captionTargetBox="[214,1364,173,1767]" captionTargetId="figure@5.[214,1372,166,1767]" captionTargetPageId="5" captionText="FIGURE 3. Zhiganka longialata sp. nov.: A–C. Paratype PIN 5026/1599: A. Anterior part of body. B. Distal and proximal flagellomeres of antenna. C. Wing. D. Abdomen of female PIN 5026/1597, dorsal view. E. Abdomen of male PIN 5340/807, lateral view (flipped horizontally). F–H. Genitalia of male. F. Paratype PIN 5026/1599. G. Paratype PIN 5026/1594. H. Paratype PIN 5340/807. Scale bars equal 2 mm (A, C, D, E), 200 μm (B, F, G, H). Abbreviations: apn, antepronotum; b, transepimeral suture; cl, clypeus; cx, coxa; ep, epandrium; gcx, gonocoxite; gst, gonostylus; hyp, hypandrium; lbl, labellum; lbr, labrum; mp, maxillary palp; ps, prescutal suture; s, scutal suture; ss, surstylus." httpUri="https://zenodo.org/record/3380987/files/figure.png" pageId="2" pageNumber="568">3A</figureCitation>
|
||
,
|
||
<figureCitation id="13292A0DFF8AE97E45CB449A45E3379A" box="[917,950,225,251]" captionStart="FIGURE 4" captionStartId="6.[151,250,1833,1858]" captionTargetBox="[227,1361,166,1813]" captionTargetId="figure@6.[227,1361,166,1813]" captionTargetPageId="6" captionText="FIGURE 4. Zhiganka longialata sp. nov.: A, B. Anterior and basal part of wing, PIN 5340/1494, part and counterpart. C. Part of wing, female PIN 5340/951. D. Genitalia of male PIN 5026/1594. E. Genitalia of male PIN 5340/953. F. Thorax and head, female PIN 5340/952. Abbreviations: aem, anepimerum or pteropleurite; apn, antepronotum; b, transepimeral suture; cx, coxa; ep, epandrium; gcx, gonocoxite; gst, gonostylus; hyp, hypandrium; lc, laterocervicale; mt, mediotergum, mpl, meropleurum; ps, prescutal suture; pt, pleurotergum; s, scutal suture; sl, scutellum; ss, surstylus." httpUri="https://zenodo.org/record/3380989/files/figure.png" pageId="2" pageNumber="568">4F</figureCitation>
|
||
). Wings rather broad, slightly shorter than abdomen, with macrotrichia confined to apex adjacent to costa; C ending at wing apex, Rs stem long, aligned with straight R
|
||
<subScript id="179634CDFF8AE97E475B4541474A3629" attach="left" box="[261,287,314,328]" fontSize="6" pageId="2" pageNumber="568">4+5</subScript>
|
||
stem, fork R
|
||
<subScript id="179634CDFF8AE97E47F3454147923629" attach="left" box="[429,455,314,328]" fontSize="6" pageId="2" pageNumber="568">4+5</subScript>
|
||
longer than R
|
||
<subScript id="179634CDFF8AE97E443B4541442A3629" attach="left" box="[613,639,314,328]" fontSize="6" pageId="2" pageNumber="568">4+5</subScript>
|
||
stem; crossvein
|
||
<emphasis id="B966EA9AFF8AE97E4564455145373622" box="[826,866,298,323]" italics="true" pageId="2" pageNumber="568">r-m</emphasis>
|
||
under Rs furcation; spurious vein between veins R and M long; M forks before
|
||
<emphasis id="B966EA9AFF8AE97E4788453547AB3606" box="[470,510,334,359]" italics="true" pageId="2" pageNumber="568">r-m</emphasis>
|
||
; M
|
||
<subScript id="179634CDFF8AE97E447B4525446A360D" attach="left" box="[549,575,350,364]" fontSize="6" pageId="2" pageNumber="568">1+2</subScript>
|
||
unforked, straight, and extremely thin; CuA slightly sigmoidal beyond
|
||
<emphasis id="B966EA9AFF8AE97E4302453543C13606" box="[1372,1428,334,359]" italics="true" pageId="2" pageNumber="568">m-cu</emphasis>
|
||
; anal lobe well developed (
|
||
<figureCitation id="13292A0DFF8AE97E47E1450A446B36EA" box="[447,574,369,395]" captionStart="FIGURE 1" captionStartId="3.[151,250,1902,1927]" captionTargetBox="[201,1387,166,1882]" captionTargetId="figure@3.[201,1387,166,1882]" captionTargetPageId="3" captionText="FIGURE 1. Zhiganka longialata sp. nov.: A–E. Male PIN 5340/953, holotype:A. Head and thorax. B. Habitus. C. Two spurs on hind tibia. D, E. Two spurs on left and right midtibia. F. Spur on fore tibia, PIN 5340/1494. G. Habitus of female PIN 5340/951, paratype. H. Terminalia of female PIN 5026/1595. Abbreviations: cl, clypeus; lbl, labellum; lbr, labrum; mp, maxillary palp; s, scutal suture; sl, scutellum." httpUri="https://zenodo.org/record/3380983/files/figure.png" pageId="2" pageNumber="568">Figs 1B, G</figureCitation>
|
||
,
|
||
<figureCitation id="13292A0DFF8AE97E4414450A44C536EA" box="[586,656,369,395]" captionStart="FIGURE 2" captionStartId="4.[151,250,1869,1894]" captionTargetBox="[187,1399,166,1849]" captionTargetId="figure@4.[187,1399,166,1849]" captionTargetPageId="4" captionText="FIGURE 2. Zhiganka longialata sp. nov., male PIN 5026/1599, paratype: A. Anterior part of body. B. Wing. C. Proximal flagellomeres of antenna, inset—distal flagellomeres, arrows mark setae. D. Head. E, F. Terminalia, part and counterpart. G, Distal part of wing. Abbreviations: apn, antepronotum; cl, clypeus; ep, epandrium; gcx, gonocoxite; gst, gonostylus; hyp, hypandrium; lbl, labellum; lbr, labrum; mp, maxillary palp; ss, surstylus." httpUri="https://zenodo.org/record/3380985/files/figure.png" pageId="2" pageNumber="568">2B, G</figureCitation>
|
||
,
|
||
<figureCitation id="13292A0DFF8AE97E44C2450A44EB36EA" box="[668,702,369,395]" captionStart="FIGURE 3" captionStartId="5.[151,250,1787,1812]" captionTargetBox="[214,1364,173,1767]" captionTargetId="figure@5.[214,1372,166,1767]" captionTargetPageId="5" captionText="FIGURE 3. Zhiganka longialata sp. nov.: A–C. Paratype PIN 5026/1599: A. Anterior part of body. B. Distal and proximal flagellomeres of antenna. C. Wing. D. Abdomen of female PIN 5026/1597, dorsal view. E. Abdomen of male PIN 5340/807, lateral view (flipped horizontally). F–H. Genitalia of male. F. Paratype PIN 5026/1599. G. Paratype PIN 5026/1594. H. Paratype PIN 5340/807. Scale bars equal 2 mm (A, C, D, E), 200 μm (B, F, G, H). Abbreviations: apn, antepronotum; b, transepimeral suture; cl, clypeus; cx, coxa; ep, epandrium; gcx, gonocoxite; gst, gonostylus; hyp, hypandrium; lbl, labellum; lbr, labrum; mp, maxillary palp; ps, prescutal suture; s, scutal suture; ss, surstylus." httpUri="https://zenodo.org/record/3380987/files/figure.png" pageId="2" pageNumber="568">3C</figureCitation>
|
||
,
|
||
<figureCitation id="13292A0DFF8AE97E4495450A448C36EA" box="[715,729,369,395]" captionStart="FIGURE 4" captionStartId="6.[151,250,1833,1858]" captionTargetBox="[227,1361,166,1813]" captionTargetId="figure@6.[227,1361,166,1813]" captionTargetPageId="6" captionText="FIGURE 4. Zhiganka longialata sp. nov.: A, B. Anterior and basal part of wing, PIN 5340/1494, part and counterpart. C. Part of wing, female PIN 5340/951. D. Genitalia of male PIN 5026/1594. E. Genitalia of male PIN 5340/953. F. Thorax and head, female PIN 5340/952. Abbreviations: aem, anepimerum or pteropleurite; apn, antepronotum; b, transepimeral suture; cx, coxa; ep, epandrium; gcx, gonocoxite; gst, gonostylus; hyp, hypandrium; lc, laterocervicale; mt, mediotergum, mpl, meropleurum; ps, prescutal suture; pt, pleurotergum; s, scutal suture; sl, scutellum; ss, surstylus." httpUri="https://zenodo.org/record/3380989/files/figure.png" pageId="2" pageNumber="568">4</figureCitation>
|
||
A–C, 5A, 6A–C). Legs uniformly dark, without banding, tibial spur formula 1:2:2, foreleg spur spine-like, unmodified (
|
||
<figureCitation id="13292A0DFF8AE97E454945EE450336CE" box="[791,854,405,431]" captionStart="FIGURE 1" captionStartId="3.[151,250,1902,1927]" captionTargetBox="[201,1387,166,1882]" captionTargetId="figure@3.[201,1387,166,1882]" captionTargetPageId="3" captionText="FIGURE 1. Zhiganka longialata sp. nov.: A–E. Male PIN 5340/953, holotype:A. Head and thorax. B. Habitus. C. Two spurs on hind tibia. D, E. Two spurs on left and right midtibia. F. Spur on fore tibia, PIN 5340/1494. G. Habitus of female PIN 5340/951, paratype. H. Terminalia of female PIN 5026/1595. Abbreviations: cl, clypeus; lbl, labellum; lbr, labrum; mp, maxillary palp; s, scutal suture; sl, scutellum." httpUri="https://zenodo.org/record/3380983/files/figure.png" pageId="2" pageNumber="568">Fig. 1</figureCitation>
|
||
C–F). Abdomen narrow in both sexes (
|
||
<figureCitation id="13292A0DFF8AE97E434D45EE43C536CE" box="[1299,1424,405,431]" captionStart="FIGURE 1" captionStartId="3.[151,250,1902,1927]" captionTargetBox="[201,1387,166,1882]" captionTargetId="figure@3.[201,1387,166,1882]" captionTargetPageId="3" captionText="FIGURE 1. Zhiganka longialata sp. nov.: A–E. Male PIN 5340/953, holotype:A. Head and thorax. B. Habitus. C. Two spurs on hind tibia. D, E. Two spurs on left and right midtibia. F. Spur on fore tibia, PIN 5340/1494. G. Habitus of female PIN 5340/951, paratype. H. Terminalia of female PIN 5026/1595. Abbreviations: cl, clypeus; lbl, labellum; lbr, labrum; mp, maxillary palp; s, scutal suture; sl, scutellum." httpUri="https://zenodo.org/record/3380983/files/figure.png" pageId="2" pageNumber="568">Fig. 1B, G</figureCitation>
|
||
). Male: epandrium divided into epandrial lobes, surstyli distinct. Gonocoxite without secondary lobe. Gonostylus simple lobe-like, pubescent. Hypandrium divided into basal and terminal divisions (
|
||
<figureCitation id="13292A0DFF8AE97E421545A642973696" box="[1099,1218,477,503]" captionStart="FIGURE 2" captionStartId="4.[151,250,1869,1894]" captionTargetBox="[187,1399,166,1849]" captionTargetId="figure@4.[187,1399,166,1849]" captionTargetPageId="4" captionText="FIGURE 2. Zhiganka longialata sp. nov., male PIN 5026/1599, paratype: A. Anterior part of body. B. Wing. C. Proximal flagellomeres of antenna, inset—distal flagellomeres, arrows mark setae. D. Head. E, F. Terminalia, part and counterpart. G, Distal part of wing. Abbreviations: apn, antepronotum; cl, clypeus; ep, epandrium; gcx, gonocoxite; gst, gonostylus; hyp, hypandrium; lbl, labellum; lbr, labrum; mp, maxillary palp; ss, surstylus." httpUri="https://zenodo.org/record/3380985/files/figure.png" pageId="2" pageNumber="568">Figs 2E, F</figureCitation>
|
||
,
|
||
<figureCitation id="13292A0DFF8AE97E429045A6428F3696" box="[1230,1242,477,503]" captionStart="FIGURE 3" captionStartId="5.[151,250,1787,1812]" captionTargetBox="[214,1364,173,1767]" captionTargetId="figure@5.[214,1372,166,1767]" captionTargetPageId="5" captionText="FIGURE 3. Zhiganka longialata sp. nov.: A–C. Paratype PIN 5026/1599: A. Anterior part of body. B. Distal and proximal flagellomeres of antenna. C. Wing. D. Abdomen of female PIN 5026/1597, dorsal view. E. Abdomen of male PIN 5340/807, lateral view (flipped horizontally). F–H. Genitalia of male. F. Paratype PIN 5026/1599. G. Paratype PIN 5026/1594. H. Paratype PIN 5340/807. Scale bars equal 2 mm (A, C, D, E), 200 μm (B, F, G, H). Abbreviations: apn, antepronotum; b, transepimeral suture; cl, clypeus; cx, coxa; ep, epandrium; gcx, gonocoxite; gst, gonostylus; hyp, hypandrium; lbl, labellum; lbr, labrum; mp, maxillary palp; ps, prescutal suture; s, scutal suture; ss, surstylus." httpUri="https://zenodo.org/record/3380987/files/figure.png" pageId="2" pageNumber="568">3</figureCitation>
|
||
F–H, 4D, E). Female: cerci tablike, setose, lightly sclerotized (
|
||
<figureCitation id="13292A0DFF8AE97E44C1467A44AD357A" box="[671,760,513,539]" captionStart="FIGURE 1" captionStartId="3.[151,250,1902,1927]" captionTargetBox="[201,1387,166,1882]" captionTargetId="figure@3.[201,1387,166,1882]" captionTargetPageId="3" captionText="FIGURE 1. Zhiganka longialata sp. nov.: A–E. Male PIN 5340/953, holotype:A. Head and thorax. B. Habitus. C. Two spurs on hind tibia. D, E. Two spurs on left and right midtibia. F. Spur on fore tibia, PIN 5340/1494. G. Habitus of female PIN 5340/951, paratype. H. Terminalia of female PIN 5026/1595. Abbreviations: cl, clypeus; lbl, labellum; lbr, labrum; mp, maxillary palp; s, scutal suture; sl, scutellum." httpUri="https://zenodo.org/record/3380983/files/figure.png" pageId="2" pageNumber="568">Fig. 1H</figureCitation>
|
||
). Spermathecae absent or unsclerotized.
|
||
</paragraph>
|
||
<paragraph id="8BAD3688FF8AE97E4699465F446035CA" blockId="2.[151,1437,153,1115]" pageId="2" pageNumber="568">
|
||
<emphasis id="B966EA9AFF8AE97E4699465F47C1355E" bold="true" box="[199,404,548,575]" pageId="2" pageNumber="568">Species included.</emphasis>
|
||
Besides the
|
||
<typeStatus id="54A9882AFF8AE97E447A465E4401355E" box="[548,596,549,575]" pageId="2" pageNumber="568">type</typeStatus>
|
||
species from Zhigansk (Batylykh Formation, Yakutia,
|
||
<collectingCountry id="F3057618FF8AE97E429E465E435B355E" box="[1216,1294,549,575]" name="Russia" pageId="2" pageNumber="568">Russia</collectingCountry>
|
||
; Lower Cretaceous),
|
||
<taxonomicName id="4C124D0BFF8AE97E475D463145383502" authority="Lukashevich, Coram, Jarzembowski, 2001" authorityName="Lukashevich, Coram, Jarzembowski" authorityYear="2001" box="[259,877,585,611]" class="Insecta" family="Ptychopteridae" genus="Zhiganka" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="568" phylum="Arthropoda" rank="species" species="woolgari">
|
||
<emphasis id="B966EA9AFF8AE97E475D463147D33502" box="[259,390,585,611]" italics="true" pageId="2" pageNumber="568">Z. woolgari</emphasis>
|
||
<bibRefCitation id="EF834B79FF8AE97E47D0463245383502" author="Lukashevich, E. D. & Coram, R. A. & Jarzembowski, E. A." box="[398,877,585,611]" pageId="2" pageNumber="568" pagination="451 - 460" refId="ref7530" refString="Lukashevich, E. D. Coram, R. A. & Jarzembowski, E. A. (2001) New true flies (Insecta: Diptera) from the Lower Cretaceous of Southern England. Cretaceous Research, 22, 451 - 460. https: // doi. org / 10.1006 / cres. 2001.026 5" type="journal article" year="2001">Lukashevich, Coram, Jarzembowski, 2001</bibRefCitation>
|
||
</taxonomicName>
|
||
from Clockhouse (Wealden Group,
|
||
<collectingCountry id="F3057618FF8AE97E43594632469935E6" name="United Kingdom" pageId="2" pageNumber="568">United Kingdom</collectingCountry>
|
||
; Upper Hauterivian) and
|
||
<taxonomicName id="4C124D0BFF8AE97E47B94615442D35E6" authority="Lukashevich, 2019" authorityName="Lukashevich" authorityYear="2019" box="[487,632,621,647]" class="Insecta" family="Ptychopteridae" genus="Zhiganka" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="568" phylum="Arthropoda" rank="species" species="longialata" status="sp. nov.">
|
||
<emphasis id="B966EA9AFF8AE97E47B94615442D35E6" box="[487,632,621,647]" italics="true" pageId="2" pageNumber="568">Z. longialata</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A25557E1FF8AE97E44204616448C35E6" box="[638,729,621,647]" pageId="2" pageNumber="568" rank="species">
|
||
<emphasis id="B966EA9AFF8AE97E44204616448735E6" bold="true" box="[638,722,621,647]" pageId="2" pageNumber="568">sp. nov</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
described below from Khasurty (Gusinoe Ozero Group, Transbaikalia,
|
||
<collectingCountry id="F3057618FF8AE97E46A046EA471935CA" box="[254,332,657,683]" name="Russia" pageId="2" pageNumber="568">Russia</collectingCountry>
|
||
; Lower Cretaceous).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3086503FF8AE97E469946CE4476333A" pageId="2" pageNumber="568" type="discussion">
|
||
<paragraph id="8BAD3688FF8AE97E469946CE4476333A" blockId="2.[151,1437,153,1115]" pageId="2" pageNumber="568">
|
||
<emphasis id="B966EA9AFF8AE97E469946CE476E35AE" bold="true" box="[199,315,693,719]" pageId="2" pageNumber="568">Remarks.</emphasis>
|
||
The genus was established based on isolated wings and pupal fragments (the only immature stage known so far). For adults of each previously known species only an incompletly preserved isolated wing was described, so the diagnosis was limited to wing venation. The additional crossvein
|
||
<emphasis id="B966EA9AFF8AE97E42794685421A3476" box="[1063,1103,766,791]" italics="true" pageId="2" pageNumber="568">r-m</emphasis>
|
||
was cautiously shown with a dotted line in
|
||
<taxonomicName id="4C124D0BFF8AE97E4769475947E9345A" authorityName="Lukashevich" authorityYear="1995" box="[311,444,802,827]" class="Insecta" family="Ptychopteridae" genus="Zhiganka" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="568" phylum="Arthropoda" rank="species" species="comitans">
|
||
<emphasis id="B966EA9AFF8AE97E4769475947E9345A" box="[311,444,802,827]" italics="true" pageId="2" pageNumber="568">Z. comitans</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<bibRefCitation id="EF834B79FF8AE97E4792475A44FC345A" author="Lukashevich, E. D." box="[460,681,800,827]" pageId="2" pageNumber="568" pagination="164 - 170" refId="ref7214" refString="Lukashevich, E. D. (1995) First pupae of the Eoptychopteridae and Ptychopteridae from the Mesozoic of Siberia (Insecta: Diptera). Paleontological Journal, 29 (4), 164 - 170." type="journal article" year="1995">Lukashevich, 1995</bibRefCitation>
|
||
: fig.
|
||
<quantity id="4CEA9B6DFF8AE97E44BB475A4555345A" box="[741,768,801,827]" metricMagnitude="-3" metricUnit="kg" metricValue="1.0" pageId="2" pageNumber="568" unit="g" value="1.0">1g</quantity>
|
||
) but was not observed in
|
||
<taxonomicName id="4C124D0BFF8AE97E4273475942E4345A" authorityName="Lukashevich, Coram, Jarzembowski" authorityYear="2001" box="[1069,1201,801,827]" class="Insecta" family="Ptychopteridae" genus="Zhiganka" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="568" phylum="Arthropoda" rank="species" species="woolgari">
|
||
<emphasis id="B966EA9AFF8AE97E4273475942E4345A" box="[1069,1201,801,827]" italics="true" pageId="2" pageNumber="568">Z. woolgari</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<bibRefCitation id="EF834B79FF8AE97E429C475A4686343E" author="Lukashevich, E. D. & Coram, R. A. & Jarzembowski, E. A." pageId="2" pageNumber="568" pagination="451 - 460" refId="ref7530" refString="Lukashevich, E. D. Coram, R. A. & Jarzembowski, E. A. (2001) New true flies (Insecta: Diptera) from the Lower Cretaceous of Southern England. Cretaceous Research, 22, 451 - 460. https: // doi. org / 10.1006 / cres. 2001.026 5" type="journal article" year="2001">
|
||
Lukashevich
|
||
<emphasis id="B966EA9AFF8AE97E4305475943DB345A" box="[1371,1422,801,827]" italics="true" pageId="2" pageNumber="568">et al</emphasis>
|
||
., 2001
|
||
</bibRefCitation>
|
||
: fig.10). Reexamination of the
|
||
<typeStatus id="54A9882AFF8AE97E4472473E44DB343E" box="[556,654,837,863]" pageId="2" pageNumber="568" type="holotype">holotype</typeStatus>
|
||
of
|
||
<taxonomicName id="4C124D0BFF8AE97E44EC473D4561343E" authorityName="Lukashevich" authorityYear="1995" box="[690,820,838,863]" class="Insecta" family="Ptychopteridae" genus="Zhiganka" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="568" phylum="Arthropoda" rank="species" species="comitans">
|
||
<emphasis id="B966EA9AFF8AE97E44EC473D4561343E" box="[690,820,838,863]" italics="true" pageId="2" pageNumber="568">Z. comitans</emphasis>
|
||
</taxonomicName>
|
||
by SEM (
|
||
<figureCitation id="13292A0DFF8AE97E45FB473E45AE343E" box="[933,1019,836,863]" captionStart="FIGURE 5" captionStartId="7.[151,250,1936,1961]" captionTargetBox="[151,1417,526,1917]" captionTargetId="figure@7.[151,1417,526,1917]" captionTargetPageId="7" captionText="FIGURE 5. SEM images:A. Zhiganka longialata sp. nov., holotype PIN 5340/953, wing; Khasurty, K 1. B. Zhiganka comitans, holotype PIN 4502/13, Zhigansk, K 1. C. Crenoptychoptera antica, PIN 1255/2123, wing; Kubekovo, J 2. Arrows mark spurious vein. Scale bars equal 2 mm." httpUri="https://zenodo.org/record/3380991/files/figure.png" pageId="2" pageNumber="568">Fig. 5B</figureCitation>
|
||
) made it clear that this additional
|
||
<emphasis id="B966EA9AFF8AE97E432A473D43C9343E" box="[1396,1436,838,863]" italics="true" pageId="2" pageNumber="568">r-m</emphasis>
|
||
is a preservation artifact, the vein M
|
||
<subScript id="179634CDFF8AE97E446E4701441C34E9" attach="left" box="[560,585,890,904]" fontSize="6" pageId="2" pageNumber="568">1+2</subScript>
|
||
is the thinnest and a long spurious vein (a longitudinal vein-like thickening in the membrane between veins R and M, bisecting crossvein
|
||
<emphasis id="B966EA9AFF8AE97E450447F545D734C6" box="[858,898,910,935]" italics="true" pageId="2" pageNumber="568">r-m</emphasis>
|
||
and ending in cell r
|
||
<subScript id="179634CDFF8AE97E423647E5422534CD" attach="left" box="[1128,1136,926,940]" fontSize="6" pageId="2" pageNumber="568">5</subScript>
|
||
) is present. Therefore, the venation patterns of the type species and the new species described herein (
|
||
<figureCitation id="13292A0DFF8AE97E45B647CA421534AA" box="[1000,1088,944,971]" captionStart="FIGURE 5" captionStartId="7.[151,250,1936,1961]" captionTargetBox="[151,1417,526,1917]" captionTargetId="figure@7.[151,1417,526,1917]" captionTargetPageId="7" captionText="FIGURE 5. SEM images:A. Zhiganka longialata sp. nov., holotype PIN 5340/953, wing; Khasurty, K 1. B. Zhiganka comitans, holotype PIN 4502/13, Zhigansk, K 1. C. Crenoptychoptera antica, PIN 1255/2123, wing; Kubekovo, J 2. Arrows mark spurious vein. Scale bars equal 2 mm." httpUri="https://zenodo.org/record/3380991/files/figure.png" pageId="2" pageNumber="568">Fig. 5A</figureCitation>
|
||
) are identical. The
|
||
<typeStatus id="54A9882AFF8AE97E434547CA432834AA" box="[1307,1405,945,971]" pageId="2" pageNumber="568" type="holotype">holotype</typeStatus>
|
||
of
|
||
<taxonomicName id="4C124D0BFF8AE97E46C947AD474F348E" authorityName="Lukashevich, Coram, Jarzembowski" authorityYear="2001" box="[151,282,981,1007]" class="Insecta" family="Ptychopteridae" genus="Zhiganka" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="568" phylum="Arthropoda" rank="species" species="woolgari">
|
||
<emphasis id="B966EA9AFF8AE97E46C947AD474F348E" box="[151,282,981,1007]" italics="true" pageId="2" pageNumber="568">Z. woolgari</emphasis>
|
||
</taxonomicName>
|
||
was not available for the study. Macrotrichia on the apical part of the wing are conspicuous in the new species (
|
||
<figureCitation id="13292A0DFF8AE97E46A9478247013372" box="[247,340,1017,1043]" captionStart="FIGURE 2" captionStartId="4.[151,250,1869,1894]" captionTargetBox="[187,1399,166,1849]" captionTargetId="figure@4.[187,1399,166,1849]" captionTargetPageId="4" captionText="FIGURE 2. Zhiganka longialata sp. nov., male PIN 5026/1599, paratype: A. Anterior part of body. B. Wing. C. Proximal flagellomeres of antenna, inset—distal flagellomeres, arrows mark setae. D. Head. E, F. Terminalia, part and counterpart. G, Distal part of wing. Abbreviations: apn, antepronotum; cl, clypeus; ep, epandrium; gcx, gonocoxite; gst, gonostylus; hyp, hypandrium; lbl, labellum; lbr, labrum; mp, maxillary palp; ss, surstylus." httpUri="https://zenodo.org/record/3380985/files/figure.png" pageId="2" pageNumber="568">Figs 2G</figureCitation>
|
||
,
|
||
<figureCitation id="13292A0DFF8AE97E4701478247F63372" box="[351,419,1017,1043]" captionStart="FIGURE 6" captionStartId="8.[151,250,1675,1700]" captionTargetBox="[151,1436,174,1652]" captionTargetId="figure@8.[151,1436,174,1655]" captionTargetPageId="8" captionText="FIGURE 6. SEM images: A–D. Zhiganka longialata sp. nov., wing of holotype PIN 5340/953, Zhigansk, K 1: A. Distal part. B. Part of wing. C, D. Fore and hind margin of wing. E, F. Crenoptychoptera antica, apical and middle part of wing PIN 1255/2123, Kubekovo, J 2. Black arrows mark spurious vein, white—setae on veins and membrane." httpUri="https://zenodo.org/record/3380993/files/figure.png" pageId="2" pageNumber="568">6B, C</figureCitation>
|
||
), but are barely discernible, without certainty, in the type species; however, this can be due to a poor state of preservation of
|
||
<emphasis id="B966EA9AFF8AE97E4459406544C53356" box="[519,656,1054,1079]" italics="true" pageId="2" pageNumber="568">
|
||
<taxonomicName id="4C124D0BFF8AE97E4459406544D83356" authorityName="Lukashevich" authorityYear="1995" box="[519,653,1054,1079]" class="Insecta" family="Ptychopteridae" genus="Zhiganka" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="568" phylum="Arthropoda" rank="species" species="comitans">Z. comitans</taxonomicName>
|
||
.
|
||
</emphasis>
|
||
The new species is, therefore, included in
|
||
<taxonomicName id="4C124D0BFF8AE97E4234406642813356" authorityName="Lukashevich" authorityYear="1995" box="[1130,1236,1053,1079]" class="Insecta" family="Ptychopteridae" genus="Zhiganka" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="568" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B966EA9AFF8AE97E4234406642813356" box="[1130,1236,1053,1079]" italics="true" pageId="2" pageNumber="568">Zhiganka</emphasis>
|
||
</taxonomicName>
|
||
and the diagnosis of the genus is revised accordingly.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |