1992 lines
323 KiB
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1992 lines
323 KiB
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<mods:title id="DFCB64283BF4F7615991C1E8049F36B8">Review of Grosphus Simon 1880 with description of Teruelius gen n a new buthid genus from Madagascar (Scorpiones Buthidae)</mods:title>
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<mods:namePart id="70875B91004973FFE2D32089E49B0298">Lowe, Graeme</mods:namePart>
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<mods:namePart id="C8B473A419FC74E0C440DB3BDE46DE3D">Kovařík, František</mods:namePart>
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<mods:title id="843239FF4650518304F6437140D4366E">Euscorpius</mods:title>
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Genus
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<taxonomicName id="4C124D1CFFBCFFAA2368D341FB06FE13" ID-CoL="B7Z35" authority="Lowe & Kovařík, 2019" authorityName="Lowe & Kovařík" authorityYear="2019" box="[1081,1204,568,597]" class="Arachnida" family="Buthidae" genus="Teruelius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="13" pageNumber="12" phylum="Arthropoda" rank="genus" status="gen. nov.">Teruelius</taxonomicName>
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.
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<emphasis id="B966EA8DFFBCFFAA23ACD341FABDFE12" bold="true" box="[1277,1295,568,596]" pageId="13" pageNumber="12">n</emphasis>
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.
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(
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<figureCitation id="13292A1AFFBCFFAA2402D325FC08FE32" box="[851,954,604,628]" captionStart="Figures 5–8" captionStartId="7.[113,190,1980,2002]" captionTargetBox="[138,1459,268,1917]" captionTargetId="figure-41@7.[113,1476,244,1929]" captionTargetPageId="7" captionText="Figures 5–8. Teruelius limbatus. Type species of Teruelius gen. n. Habitus. Male (5–6) and female (7–8), dorsal (5, 7) and ventral (6, 8) views. Scale bars: 10 mm." figureDoi="http://doi.org/10.5281/zenodo.7143614" httpUri="https://zenodo.org/record/7143614/files/figure.png" pageId="13" pageNumber="12">Figs. 5–8</figureCitation>
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,
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<figureCitation id="13292A1AFFBCFFAA2495D325FBB8FE32" box="[964,1034,604,628]" captionStart="Figures 9–20" captionStartId="11.[116,190,1913,1935]" captionTargetBox="[129,1461,231,1903]" captionTargetId="figure-144@11.[129,1461,231,1903]" captionTargetPageId="11" captionText="Figures 9–20. Position of femur trichobothrium d 2in Grosphus and Teruelius gen. n. Dorsal surfaces of proximal pedipalp femur of adult females (♀, left panels) and adult males (♂, right panels), underUV fluorescence to highlightgranulation,carinae and trichobothrial areolae.Whitearrows indicate positions of trichobothrium d in each image. G. madagascariensis (9, 10; 2 samples from Anjiro (9) and Andasibe (10) show consistency of d 2 2 position),G. hirtus(11), G. voahangyae (12), T. ankarafantsika (13, 15; 2 samples fromAmpijoroa show consistency ofd 2position),T. ankarana (14), T. flavopiceus (16), T. grandidieri (17), T. limbatus (18), T. mahafaliensis (19), and T. olgae (20). Scale bars: 1 mm (9–20♀) or 500 μm (20 ♂)." figureDoi="http://doi.org/10.5281/zenodo.7143618" httpUri="https://zenodo.org/record/7143618/files/figure.png" pageId="13" pageNumber="12">13–20</figureCitation>
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,
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<figureCitation id="13292A1AFFBCFFAA2345D325FBEBFE32" box="[1044,1113,604,628]" captionStart-0="Figures 21–27" captionStart-1="Figures 28–29" captionStart-2="Figures 30–35" captionStart-3="Figures 36–39" captionStartId-0="14.[113,188,1859,1881]" captionStartId-1="18.[113,190,1953,1975]" captionStartId-2="19.[113,190,1773,1795]" captionStartId-3="20.[113,190,1388,1410]" captionTargetBox-0="[114,1474,227,1840]" captionTargetBox-2="[118,1455,257,1710]" captionTargetBox-3="[121,1471,246,1361]" captionTargetId-0="figure-206@14.[113,1474,222,1851]" captionTargetId-1="figure-131@18.[113,1474,255,1915]" captionTargetId-2="figure-204@19.[113,1474,251,1714]" captionTargetId-3="figure-425@20.[113,1474,234,1383]" captionTargetPageId-0="14" captionTargetPageId-1="18" captionTargetPageId-2="19" captionTargetPageId-3="20" captionText-0="Figures 21–27. Positions of trichobothria Eb, Eb and petite Eb on manus of pedipalp chela in Grosphus, Neogrosphus and Teruelius gen. 1 2 3 n. Figures 21–24. External views of pedipalp chela, shown under UV fluorescence to highlight trichobothrial areolae. G. hirtus ♂ (21), G. madagascariensis ♀ (22), T. limbatus ♂ (23), T. ankarafantsika ♀ (24). Positions of Eb 1, Eb 2 and Eb 3 and lines joining them shown as white overlays. Scale bars: 2 mm. Figures 25–26. Horizontal histograms comparing ratios, R = d(Eb, Eb)/ d(Eb, Eb), of Eb -Eb distance, to Eb - 123 2 3 1 2 2 3 1 Eb distance, in males (25), and females (26) of Grosphus, Neogrosphus and Teruelius gen. n. Error bars indicate ranges (minimum, maximum), 2 histogram bars mid-range values. Figure 27. Scatter plot of male vs. female ratios R. Ratio is larger in males if points fall above the diagonal 123 (gray) line, larger in females if they fall below it." captionText-1="Figures 28–29. Variation in pectinal tooth count (PTC) in Grosphus, Neogrosphus and Teruelius gen. n. Horizontal histograms comparing PTC in males (28), and females (29) of Grosphus, Neogrosphus and Teruelius gen. n. Error bars indicate ranges (minimum, maximum), bars mid-range values." captionText-2="Figures 30–35. Variation in pectinal tooth count (PTC) in Grosphus, Neogrosphus and Teruelius gen. n. Figures 30–31. Logarithmic scatter plots showing scaling of PTC vs. carapace length in males (30), and females (31) of Grosphus, Neogrosphus and Teruelius gen. n. Color lines are least squares linear regression fits of scaling trends for the different genera. Figures 32–33. Logarithmic scatter plots showing scaling of PTC vs. carapace length in Figs. 30–31 superposed upon overall scaling trends for a larger sample of male (32) and female (33) buthids including both α and β trichobothrial configurations (gray symbols; 1055♂, 1046♀). Dark lines: least squares linear regression fits of scaling trends for larger sample of buthids. Figures 34–35. Logarithmic scatter plots showing scaling of PTC vs. carapace length in Figs. 30–31 superposed upon overall scaling trends for a larger sample of male (34) and female (35) α-buthids restricted to α trichobothrial configurations (gray symbols; 445♂, 371♀). Dark lines: least squares linear regression fits of scaling trends for α-buthids. Numbers: slopes of regression lines (= allometric exponents). Purple symbols and lines: outgroup Pseudolychas." captionText-3="Figures 36–39. Comparative biometrics of female basal pectinal tooth and hemispermatophore trunk of Grosphus, Neogrosphus and Teruelius gen. n. Figures 36–37. Logarithmic scatter plots showing scaling of length of female basal pectinal tooth vs. carapace length (36), and hemispermatophore trunk length vs. carapace length (37). Color lines: least squares linear regression fits of scaling trends for the different genera. Figures 38–39. Logarithmic scatter plots of female basal pectinal tooth length vs. hemispermatophore trunk length (38), and the ratio of basal pectinal tooth length/ hemispermatophore trunk length (%) vs. hemispermatophore trunk length, showing fraction of trunk spanned by the female tooth (39). Female basal pectinal tooth length measured as chord length, from more proximal corner (along comb axis) at base of tooth, to more distal corner (along comb axis) at tip of tooth. Trunk length measured from proximal edge of basal lobe to pedicel (foot). Color lines: least squares linear regression fits of scaling trends for the different genera." figureDoi-0="http://doi.org/10.5281/zenodo.7143620" figureDoi-1="http://doi.org/10.5281/zenodo.7143622" figureDoi-2="http://doi.org/10.5281/zenodo.7143626" figureDoi-3="http://doi.org/10.5281/zenodo.7143628" httpUri-0="https://zenodo.org/record/7143620/files/figure.png" httpUri-1="https://zenodo.org/record/7143622/files/figure.png" httpUri-2="https://zenodo.org/record/7143626/files/figure.png" httpUri-3="https://zenodo.org/record/7143628/files/figure.png" pageId="13" pageNumber="12">23–39</figureCitation>
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,
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<figureCitation id="13292A1AFFBCFFAA2335D325FB18FE32" box="[1124,1194,604,628]" captionStart="Figures 40–51" captionStartId="22.[113,188,1930,1952]" captionTargetBox="[113,1474,234,1920]" captionTargetId="figure-124@22.[113,1474,234,1922]" captionTargetPageId="22" captionText="Figures 40–51. Female basal pectinal teeth in Grosphus and Teruelius gen. n. Ventral views of proximal left pectine of females shown under UV fluorescence to highlight cuticular surface texture, setation and absence of peg sensillae on basal tooth vs. their presence on other teeth. G. sp. nr hirtus (40), G. madagascariensis (41), G. hirtus (42), G. voahangyae (43), T. ankarafantsika (44–45; 2 samples fromAmpijoroa show variation in tooth shape), T. ankarana (46), T. flavopiceus (47), T. grandidieri (48), T. limbatus (49), T. mahafaliensis (50), and T. olgae (51). Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7143634" httpUri="https://zenodo.org/record/7143634/files/figure.png" pageId="13" pageNumber="12">44–51</figureCitation>
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,
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<figureCitation id="13292A1AFFBCFFAA23E5D325FB48FE32" box="[1204,1274,604,628]" captionStart="Figures 71–85" captionStartId="26.[113,190,1701,1723]" captionTargetBox="[149,1434,231,1678]" captionTargetId="figure-287@26.[146,1446,225,1690]" captionTargetPageId="26" captionText="Figures 71–85. Hemispermatophores and capsule regions of Teruelius gen n. Multi-panel figures show: whole hemispermatophore; whole hemispermatophore and/or capsule with flagellum; capsule region in convex (or convex compressed), anterior and posterior views (panels in left to right sequence). Right hemispermatophores. Figure 71. T. ankarana, whole hemispermatophore. Scale bar: 4 mm. Figure 72. T. ankarana, capsule, left mirrored, Forêt d’Ankavanana, FMNH. Scale bar: 500 μm. Figure 73. T. grandidieri, whole hemispermatophore (flagellum truncated). Scale bar: 4 mm. Figure 74. T. grandidieri, capsule, Antsakabe River, FMNH. Scale bar: 500 μm. Figure 75. T. flavopiceus, whole hemispermatophore. Scale bar: 2 mm. Figure 76. T. flavopiceus, capsule, Madagascar, GLPC, FKCP. Scale bar: 500 μm. Figure 77. T. annulatus, capsule, Tsimanampetsotsa National Park, GLPC, FKCP. Scale bar: 500 μm. Figure 78. T. ankarafantsika, capsule and flagellum. Scale bar: 400 μm. Figure 79. T. ankarafantsika, capsule, Forêt d’Ankavanana, FMNH. Scale bar: 200 μm. Figure 80. T. ankarafantsika, capsule, Réserve Forestière de l’Ankarafantsika, FMNH. Scale bar: 200 μm. Figure 81. T. olgae, capsule, Itampolo village, FMNH. Scale bar: 500 μm. Figure 82. T. limbatus, whole hemispermatophore (flagellum truncated). Scale bar: 2 mm. Figure 83. T. limbatus, capsule, Forêt d’Ianasana, FMNH. Scale bar: 500 μm. Figure 84. T. mahafaliensis, capsule views, Zombitse-Vohibasia National Park, GLPC, FKCP. Scale bar: 500 μm. Figure 85. T. intertidalis, capsule, Madagascar, GLPC, FKCP. Scale bar: 500 μm." figureDoi="http://doi.org/10.5281/zenodo.7143641" httpUri="https://zenodo.org/record/7143641/files/figure.png" pageId="13" pageNumber="12">71–85</figureCitation>
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,
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<figureCitation id="13292A1AFFBCFFAA2255D325FAF8FE32" box="[1284,1354,604,628]" captionStart="Figures 90–93" captionStartId="28.[113,190,1598,1620]" captionTargetBox="[119,1454,254,1587]" captionTargetId="figure-271@28.[113,1474,234,1597]" captionTargetPageId="28" captionText="Figures 90–93. Hemispermatophores and capsule regions of Teruelius gen. n. Cross stereoscopic convex views. Figure 90. T. ankarana, left mirrored, Forêt d’Ankavanana, FMNH. Figure 91. T. limbatus, Forêt d’Ianasana, FMNH. Figure 92. T. flavopiceus, Madagascar, GLPC, FKCP. Figure 93. T. ankarafantsika, Forêt d’Ankavanana, FMNH. Scale bars: 200 μm." figureDoi="http://doi.org/10.5281/zenodo.7143647" httpUri="https://zenodo.org/record/7143647/files/figure.png" pageId="13" pageNumber="12">90–93</figureCitation>
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,
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<figureCitation id="13292A1AFFBCFFAA2205D325FC2CFED2" captionStart-0="Figures 95–110" captionStart-1="Figures 110–113" captionStart-2="Figures 114–116" captionStart-3="Figures 117–120" captionStart-4="Figure 121" captionStartId-0="18.[113,190,1408,1430]" captionStartId-1="32.[113,190,1684,1706]" captionStartId-2="33.[113,190,1610,1632]" captionStartId-3="35.[113,190,1782,1804]" captionStartId-4="36.[113,181,1915,1937]" captionTargetBox-0="[113,1475,255,1894]" captionTargetBox-1="[113,1474,276,1623]" captionTargetBox-2="[128,1470,242,1585]" captionTargetBox-3="[132,1464,278,1712]" captionTargetBox-4="[120,1469,286,1859]" captionTargetId-0="figure-131@18.[113,1474,255,1915]" captionTargetId-1="figure-16@32.[113,1474,266,1629]" captionTargetId-2="figure-262@33.[113,1474,234,1604]" captionTargetId-3="figure-194@35.[113,1474,274,1732]" captionTargetId-4="figure-94@36.[113,1474,282,1865]" captionTargetPageId-0="18" captionTargetPageId-1="32" captionTargetPageId-2="33" captionTargetPageId-3="35" captionTargetPageId-4="36" captionText-0="Figures 95–110: Orthochirus mesopotamicus comb. n., from Iraq, Basra (ca. 30°48’N 47°34’E). Figures 95, 97, 100–110. Male, carapace and tergites I–III (95), sternopectinal region and sternites (97). Pedipalp chela, dorsal (100), external (101), and ventrointernal (102) views. Pedipalp patella, dorsal (103), external (104), and ventral (105) views. Pedipalp femur and trochanter, internodorsal (106) and dorsoexternal (107) views. Pedipalp chela, movable finger dentate margin (108). Distal segments of right legs III–IV, retrolateral views (109–110). Figures 96, 98–99. Female, carapace and tergites I–III (96), sternopectinal region and sternites (98), and pedipalp chela dorsoexternal (99). The trichobothrial pattern is indicated in Figures 101–104, 106–107 (white circles)." captionText-1="Figures 110–113. Biometric scaling of spiracles of sternite IV in Grosphus, Neogrosphus and Teruelius gen. n. Figures 110–111. Logarithmic scatter plots showing scaling of single spiracle area/ (carapace length)2 vs. carapace length in males (110), and females (111) of Grosphus, Neogrosphus and Teruelius gen. n. Spiracle length was largest diameter (approximately along mediolateral axis), width was smallest diameter (approximately along rostrocaudal axis). Area was from closed boundary curve: anterior edge of opening sharply delineated, posterior edge sometimes less sharp and fitted with spline curve extrapolating lateral edge curvature and tracing shadow of raised posterior margin. Brown line is least squares linear regression fit of scaling trends for Teruelius. R = Pearson correlation coefficient, and P value as indicated. Plotted points are means, error bars SE. Figures 112–113. Logarithmic scatter plots showing thermal (terrestrial temperature) scaling of spiracle area/ (carapace length)2 in males (112), and females (113) of Grosphus, Neogrosphus and Teruelius gen. n. Gray lines are least squares linear regression fits of scaling trends for all points. R = Pearson correlation coefficient, P value as indicated. Plotted points are means, error bars SE. Land surface temperatures at known collection sites were averaged over all sites for each species. Collection sites were extracted from published data and FMNH records. Temperatures were extracted from published model data of Chabot-Couture et al. (2014), who estimated annual mean daytime temperature by statistical processing and Kriging of remote sensing spectral radiance by AQUA satellite." captionText-2="Figures 114–116. Biometrics of spiracles of sternite IV in Grosphus, Neogrosphus and Teruelius gen. n. Logarithmic scatter plots showing sexual dimorphism of spiracle area/ (carapace length)2 (114), spiracle width/ carapace length (115), and spiracle length/ width (116) in Grosphus, Neogrosphus and Teruelius gen. n. Plotted points are means, error bars SE. Ratios are larger in males if points fall above the diagonal (gray) line, larger in females if they fall below it." captionText-3="Figures 117–120. Distributions of Grosphus, Neogrosphus and Teruelius gen. n. superposed on climate maps. Figure 117. Collection sites superposed on average rainfall map. Figure 118. Collection sites superposed on land temperature map. Figure 119. Distribution of each genus according to average rainfall. Figure 120. Distribution of each genus according to land temperature. Vertical histogram bars in Figs 119–120 are means, error bars SE. P-values and asterisks indicate significant differences of Neogrosphus and Teruelius gen. n. means from Grosphus means. Symbols in bar charts are collection sites collapsed along rainfall axis (119), or temperature axis (120). Inset horizontal histograms show relative (normalized) densities of points vs. rainfall or temperature (vertical axes). Collection sites were extracted from published data and FMNH records. Rainfall and temperatures were extracted from published model data of Chabot-Couture et al. (2014), who estimated January and July rainfalls by interpolation of RFE 2.0 data, and annual mean daytime temperature by statistical processing and Kriging of remote sensing spectral radiance by AQUA satellite. We averaged January and July rainfalls to generate a single map." captionText-4="Figure 121. Bioclimatic zone distributions of Grosphus, Neogrosphus and Teruelius gen. n. in Madagascar. Collection sites were extracted from published data and FMNH records, and their bioclimatic classification visualized by superposition on zone boundaries defined by Cornet (1974). Colored regions show zones as delineated by Cornet, and symbols for genera are placed on the map and on zone keys of Cornet (upper left inset) to show ranges of humidity and temperature tolerance of each genus." figureDoi-1="http://doi.org/10.5281/zenodo.7143657" figureDoi-2="http://doi.org/10.5281/zenodo.7143663" figureDoi-3="http://doi.org/10.5281/zenodo.7143665" figureDoi-4="http://doi.org/10.5281/zenodo.7143669" httpUri-1="https://zenodo.org/record/7143657/files/figure.png" httpUri-2="https://zenodo.org/record/7143663/files/figure.png" httpUri-3="https://zenodo.org/record/7143665/files/figure.png" httpUri-4="https://zenodo.org/record/7143669/files/figure.png" pageId="13" pageNumber="12">99–105, 106–121</figureCitation>
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<figureCitation id="13292A1AFFBCFFAA24F8D305FBBBFED2" box="[937,1033,636,660]" captionStart="Figures 133–144" captionStartId="40.[113,190,1293,1315]" captionTargetBox="[118,1474,249,1273]" captionTargetId="figure-523@40.[113,1474,227,1273]" captionTargetPageId="40" captionText="Figures 133–144. Ventral setation of telotarsus III in Grosphus and Teruelius gen. n. Ventral surfaces of right telotarsus III of adult males (♂) or females (♀), shown under UV fluorescence to highlight setation. Macrosetae appear dark with strongly fluorescent cuticular sockets at their base. Putative chemosensory microsetae appear bright. G. hirtus (= G. h. garciai) (133), G. hirtus (134), G. madagascariensis (135, from Anjiro, 136 from Andasibe), G. voahangyae (137), T. ankarafantsika (138), T. limbatus (139), T. mahafaliensis (140), T. ankarana (141), T. grandidieri (142), T. flavopiceus (143), and T. olgae (144). Scale bars: 500 μm (133–140, 144) and 1 mm (141–143)." figureDoi="http://doi.org/10.5281/zenodo.7143677" httpUri="https://zenodo.org/record/7143677/files/figure.png" pageId="13" pageNumber="12">137–144</figureCitation>
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<figureCitation id="13292A1AFFBCFFAA2345D305FBC6FED2" box="[1044,1140,636,660]" captionStart-0="Figures 145–157" captionStart-1="Figures 158-160" captionStartId-0="41.[113,190,1502,1524]" captionStartId-1="42.[113,190,695,717]" captionTargetBox-0="[113,1474,234,1490]" captionTargetBox-1="[114,1470,251,674]" captionTargetId-0="figure-392@41.[113,1474,234,1490]" captionTargetId-1="figure-866@42.[113,1474,234,683]" captionTargetPageId-0="41" captionTargetPageId-1="42" captionText-0="Figures 145–157. Comparative intensity of UV fluorescence in Grosphus and Teruelius gen. n. Photographic comparison of fluorescence emission intensities of representative species of each genus, including adult males (♂) or females (♀). G. madagascariensis (145 from Anjiro, 148 from Andasibe), G. hirtus (146), G. voahangyae (147), G. hirtus (= G. h. garciai) (149), T. ankarafantsika (150), T. grandidieri (151), T. olgae (152), T. flavopiceus (153), T. ankarana (154–155), T. mahafaliensis (156), and T. limbatus (157). Images acquired under identical intensities of UV excitation (395 nm LED source, 0.35 A current) and camera exposure (Canon EOS 7D Mark II, 100 mm f/13, 0.5 s, ISO 320), with 475 nm longpass filter to block excitation wavelengths. Scale bars: 10 mm (145–150, 152–153, 156–157), 20 mm (151, 154–155)." captionText-1="Figures 158-160. Comparative intensity of UV fluorescence in Grosphus and Teruelius gen. n. Figure 158. Horizontal histograms comparing relative fluorescence emission intensities measured by photodiode current, for Grosphus and Teruelius species.Histogram bars are means, error bars are SE, species averages over all samples, male and female (sexes not significantly different, P> 0.05). Sample sizes: G. hirtus, 1♂, 1♀; G. sp. nr hirtus, 6♀; G. madagascariensis, 9♂, 1♀; G. voahangyae, 4♂, 1♀; T. ankarafantsika, 2♂, 2♀; T. ankarana, 1♂, 1♀; T. flavopiceus, 1♂, 1♀; T. grandidieri, 1♂, 1♀; T. limbatus, 6♂, 5♀; T. mahafaliensis, 2♂, 2♀; T. olgae, 1♂, 1♀. Sample means may not reflect single specimen intensities recorded in Figs. 145–157. Figure 159. Time course of photodiode current in UV fluorescence emission measurement for T. limbatus (upper curve), and two specimens of G. madagascariensis (middle and lower curves) over 620 s. Arrow indicates time of shutter opening to initiate UV excitation.160. Photobleaching time course for UV fluorescence emission of T. limbatus (middle curve) and G. madagascariensis (upper and lower curves) over 600 s following shutter opening (see Fig. 159). Ordinate is percentage bleaching relative to initial fluorescence after shutter opening. Blue curves are mono-exponential fits with decay time constants: T. limbatus, 640.2 s; G. madagascariensis, #1 654.4 s and #2 347.5 s." figureDoi-0="http://doi.org/10.5281/zenodo.7143681" figureDoi-1="http://doi.org/10.5281/zenodo.7143685" httpUri-0="https://zenodo.org/record/7143681/files/figure.png" httpUri-1="https://zenodo.org/record/7143685/files/figure.png" pageId="13" pageNumber="12">150–160</figureCitation>
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<figureCitation id="13292A1AFFBCFFAA232FD305FB6DFED2" box="[1150,1247,636,660]" captionStart="Figures 165–180" captionStartId="63.[113,190,1958,1980]" captionTargetBox="[113,1473,312,1921]" captionTargetId="figure-73@63.[113,1473,312,1921]" captionTargetPageId="63" captionText="Figures 165–180. Carapace of representative Grosphus and Teruelius gen. n. G. madagascariensis (165, 168), G. hirtus (166–167), G. voahangyae (169), T. ankarafantsika (170–171), T. ankarana (172), T. flavopiceus (173–174), T. grandidieri (175), T. limbatus (176), T.mahafaliensis (177–178) andT. olgae (179–180).UV fluorescence,♂male, ♀female.Scale bars:2mm (165–171, 176–180),4 mm (172–175)." figureDoi="http://doi.org/10.5281/zenodo.7143694" httpUri="https://zenodo.org/record/7143694/files/figure.png" pageId="13" pageNumber="12">170–180</figureCitation>
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<figureCitation id="13292A1AFFBCFFAA23B8D305FAFBFED2" box="[1257,1353,636,660]" captionStart="Figures 181–195" captionStartId="64.[113,190,1953,1975]" captionTargetId="figure-71@64.[113,1474,242,1670]" captionTargetPageId="64" captionText="Figures 181–195. Telson of representativeGrosphus andTeruelius gen. n. G. hirtus (181–182), G. madagascariensis (183–184), G. voahangyae (185), T. ankarafantsika (186–187), T. ankarana (188), T. flavopiceus (189), T. grandidieri (190–191), T. limbatus (192), T. mahafaliensis (193) and T. olgae (194–195). UV fluorescence, ♂ male, ♀ female. Scale bars: 2 mm (181–187, 192–195), 4 mm (188–191)." figureDoi="http://doi.org/10.5281/zenodo.7143700" httpUri="https://zenodo.org/record/7143700/files/figure.png" pageId="13" pageNumber="12">186–195</figureCitation>
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<figureCitation id="13292A1AFFBCFFAA2205D305FA06FED2" box="[1364,1460,636,660]" captionStart="Figures 196–210" captionStartId="65.[113,190,1950,1972]" captionTargetId="figure-77@65.[113,1474,234,1589]" captionTargetPageId="65" captionText="Figures 196–210. Female basal pectinal teeth of representative Grosphus and Teruelius gen. n. G. hirtus (196–198), G. madagascariensis (199), G. madagascariensis, paratype of G. mandena(200), T. ankarafantsika (201),T. ankarana (202), T. annulatus (203),T. bistriatus (204), T. flavopiceus (205), T. grandidieri (206), T. intertidalis (207), T. limbatus (208), T. mahafaliensis (209), T. feti (210)." figureDoi="http://doi.org/10.5281/zenodo.7143706" httpUri="https://zenodo.org/record/7143706/files/figure.png" pageId="13" pageNumber="12">201–210</figureCitation>
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<figureCitation id="13292A1AFFBCFFAA2468D3E5FC2BFEF2" box="[825,921,668,692]" captionStart="Figures 211–226" captionStartId="66.[113,190,1920,1942]" captionTargetBox="[117,1466,234,1743]" captionTargetId="figure-88@66.[113,1474,234,1743]" captionTargetPageId="66" captionText="Figures 211–226. Ventral tarsal setation of legs III or IV in Grosphus and Teruelius gen. n. G. goudoti (211), G. hirtus (212), G. hirtus (G. garciai) (213), G. madagascariensis (214), G. madagascariensis (G. mandena) (215), Teruelius ankarana (216), T. flavopiceus (217), T. annulatus (218), T. ankarafantsika (219–220), T. bistriatus (221), T. intertidalis (222), T. grandidieri (223), T. mahafaliensis (224), T. limbatus (225), T. feti (226)." figureDoi="http://doi.org/10.5281/zenodo.7143708" httpUri="https://zenodo.org/record/7143708/files/figure.png" pageId="13" pageNumber="12">216–226</figureCitation>
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<figureCitation id="13292A1AFFBCFFAA24F2D3E5FBB1FEF2" box="[931,1027,668,692]" captionStart="Figures 227–230" captionStartId="67.[113,190,1951,1973]" captionTargetBox="[113,1474,234,1902]" captionTargetId="figure-2@67.[113,1474,234,1902]" captionTargetPageId="67" captionText="Figures 227–230. Right lateral eyes of Grosphus and Teruelius gen. n. G. madagascariensis (227), G. voahangyae (228), T. limbatus (229) and T. flavopiceus (230). All species comply with the 5-eye buthid pattern with series of 3 larger ocelli in lower position, and two smaller ocelli in posterior and upper positions. UV fluorescence, males. Scale bars: 500 μm." figureDoi="http://doi.org/10.5281/zenodo.7143712" httpUri="https://zenodo.org/record/7143712/files/figure.png" pageId="13" pageNumber="12">229–230</figureCitation>
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<figureCitation id="13292A1AFFBCFFAA235FD3E5FBDCFEF2" box="[1038,1134,668,692]" captionStart="Figures 231–238" captionStartId="68.[112,189,1958,1980]" captionTargetId="figure-61@68.[113,1474,234,1421]" captionTargetPageId="68" captionText="Figures 231–238. Males. Right chelicera. Figures 231–234. Grosphus madagascariensis, dorsal (231, 233) and ventral (232, 234) views, under white light (231–232) and UV fluorescence (233–234). Figures 235–238. Teruelius limbatus, dorsal (235, 237) and ventral (236, 238) views, under white light (235–236) and UV fluorescence (237–238). Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7143714" httpUri="https://zenodo.org/record/7143714/files/figure.png" pageId="13" pageNumber="12">235–238</figureCitation>
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<figureCitation id="13292A1AFFBCFFAA2328D3E5FB6AFEF2" box="[1145,1240,668,692]" captionStart-0="Figures 387–390" captionStart-1="Figures 391–408" captionStart-10="Figure 459" captionStart-11="Figures 460–472" captionStart-12="Figures 473–476" captionStart-13="Figures 477–490" captionStart-14="Figures 491–494" captionStart-15="Figures 495–499" captionStart-16="Figures 500–511" captionStart-17="Figures 512–515" captionStart-18="Figures 516–521" captionStart-19="Figures 522–525" captionStart-2="Figures 409–417" captionStart-20="Figures 526–537" captionStart-21="Figures 538–541" captionStart-22="Figures 542–546" captionStart-23="Figures 547–551" captionStart-24="Figures 552–563" captionStart-25="Figures 564–575" captionStart-26="Figures 576–579" captionStart-3="Figures 418–419" captionStart-4="Figures 420–421" captionStart-5="Figures 422–425" captionStart-6="Figures 426–435" captionStart-7="Figures 436–438" captionStart-8="Figures 439–446" captionStart-9="Figures 447–458" captionStartId-0="87.[113,190,1981,2003]" captionStartId-1="88.[113,190,1919,1941]" captionStartId-10="97.[113,181,1982,2004]" captionStartId-11="98.[113,190,1930,1952]" captionStartId-12="99.[113,190,1981,2003]" captionStartId-13="100.[113,190,1928,1950]" captionStartId-14="101.[113,190,1981,2003]" captionStartId-15="102.[113,190,1981,2003]" captionStartId-16="103.[113,190,1928,1950]" captionStartId-17="104.[117,194,1981,2003]" captionStartId-18="105.[113,190,1958,1980]" captionStartId-19="106.[113,190,1981,2003]" captionStartId-2="89.[113,190,1958,1980]" captionStartId-20="107.[113,190,1928,1950]" captionStartId-21="108.[113,190,1981,2003]" captionStartId-22="109.[113,190,1981,2003]" captionStartId-23="110.[113,190,1981,2003]" captionStartId-24="111.[113,190,1924,1946]" captionStartId-25="112.[113,190,1950,1972]" captionStartId-26="113.[113,190,1981,2003]" captionStartId-3="90.[113,190,1992,2014]" captionStartId-4="91.[113,190,2011,2033]" captionStartId-5="92.[115,192,1981,2003]" captionStartId-6="93.[113,190,1928,1950]" captionStartId-7="94.[113,190,1981,2003]" captionStartId-8="95.[113,190,1981,2003]" captionStartId-9="96.[113,190,1958,1980]" captionTargetBox-0="[130,1452,241,1959]" captionTargetBox-1="[125,1462,242,1860]" captionTargetBox-10="[113,1474,234,1946]" captionTargetBox-11="[113,1474,232,1920]" captionTargetBox-12="[125,1454,245,1949]" captionTargetBox-13="[110,1468,256,1846]" captionTargetBox-14="[124,1467,247,1915]" captionTargetBox-15="[123,1465,254,1921]" captionTargetBox-16="[113,1465,240,1881]" captionTargetBox-17="[146,1452,271,1937]" captionTargetBox-18="[146,1426,242,1905]" captionTargetBox-19="[150,1435,241,1931]" captionTargetBox-2="[113,1474,234,1814]" captionTargetBox-20="[113,1458,234,1912]" captionTargetBox-21="[129,1451,241,1964]" captionTargetBox-22="[126,1474,251,1952]" captionTargetBox-23="[117,1471,261,1846]" captionTargetBox-24="[113,1464,245,1912]" captionTargetBox-25="[113,1438,237,1916]" captionTargetBox-26="[122,1462,264,1906]" captionTargetBox-5="[131,1456,264,1907]" captionTargetBox-6="[113,1474,234,1874]" captionTargetBox-7="[130,1447,241,1866]" captionTargetBox-8="[141,1445,264,1884]" captionTargetBox-9="[133,1446,260,1921]" captionTargetId-0="figure-29@87.[115,1476,234,1964]" captionTargetId-1="figure-93@88.[113,1474,234,1872]" captionTargetId-10="figure-36@97.[113,1474,234,1949]" captionTargetId-11="figure-86@98.[113,1474,231,1923]" captionTargetId-12="figure-28@99.[113,1474,242,1957]" captionTargetId-13="figure-92@100.[110,1471,244,1880]" captionTargetId-14="figure-27@101.[115,1475,242,1920]" captionTargetId-15="figure-48@102.[113,1474,248,1928]" captionTargetId-16="figure-73@103.[113,1474,234,1881]" captionTargetId-17="figure-34@104.[135,1459,234,1949]" captionTargetId-18="figure-51@105.[136,1452,232,1918]" captionTargetId-19="figure-39@106.[127,1460,232,1934]" captionTargetId-2="figure-58@89.[113,1474,234,1838]" captionTargetId-20="figure-80@107.[113,1468,234,1919]" captionTargetId-21="figure-37@108.[123,1459,234,1977]" captionTargetId-22="figure-50@109.[113,1474,232,1967]" captionTargetId-23="figure-45@110.[113,1474,246,1883]" captionTargetId-24="figure-71@111.[113,1474,236,1915]" captionTargetId-25="figure-53@112.[113,1474,225,1917]" captionTargetId-26="figure-33@113.[110,1474,245,1928]" captionTargetId-3="figure-22@90.[113,1474,265,1687]" captionTargetId-4="figure-22@91.[117,1478,268,1690]" captionTargetId-5="figure-27@92.[115,1476,249,1911]" captionTargetId-6="figure-72@93.[113,1474,234,1874]" captionTargetId-7="figure-29@94.[113,1474,234,1876]" captionTargetId-8="figure-37@95.[113,1474,236,1904]" captionTargetId-9="figure-61@96.[113,1474,236,1929]" captionTargetPageId-0="87" captionTargetPageId-1="88" captionTargetPageId-10="97" captionTargetPageId-11="98" captionTargetPageId-12="99" captionTargetPageId-13="100" captionTargetPageId-14="101" captionTargetPageId-15="102" captionTargetPageId-16="103" captionTargetPageId-17="104" captionTargetPageId-18="105" captionTargetPageId-19="106" captionTargetPageId-2="89" captionTargetPageId-20="107" captionTargetPageId-21="108" captionTargetPageId-22="109" captionTargetPageId-23="110" captionTargetPageId-24="111" captionTargetPageId-25="112" captionTargetPageId-26="113" captionTargetPageId-3="90" captionTargetPageId-4="91" captionTargetPageId-5="92" captionTargetPageId-6="93" captionTargetPageId-7="94" captionTargetPageId-8="95" captionTargetPageId-9="96" captionText-0="Figures 387–390. Teruelius ankarafantsika. Habitus. Male paratype (387–388) and female holotype (389–390), dorsal (387, 389) and ventral (388, 390) views. Scale bar: 10 mm." captionText-1="Figures 391–408. Teruelius ankarafantsika. Male paratype (391–393, 404–405) and female holotype (394–403, 406–408). Figures 391–402. Pedipalp.Chela in dorsal (391, 394), external (392, 395) and ventral (396) views. Patella in dorsal (397), external (398) and ventral (399) views. Femur and trochanter in internal (393, 400) and dorsal (401) views. Pedipalp chela, movable finger dentate margin (402). Figures 403–408. Telson in lateral (403, 404) views. Metasoma and telson in lateral (405, 406), dorsal (407) and ventral (408) views.Scale bar: 10 mm (406–408)." captionText-10="Figure 459. Teruelius feti. Holotype juvenile male. Habitus, dorsal view, with specimen labels (FMNH). Scale bar: 5 mm (holotype). Figure 459a. Adult female from Makay Mts., carapace and tergites I–II." captionText-11="Figures 460–472. Teruelius feti. Holotype male. Metasoma and pedipalp. Figures 460–462. Metasoma in dorsal (460), lateral (461) and ventral (462) views. Figures 463–472. Pedipalp. Chela in dorsal (463), external (464) and ventral (465) views. Patella in dorsal (466), external (467) and ventral (468) views. Femur and trochanter in internal (469), dorsal (470), ventral (471) and external (472) views. Right femur shows developmental anomaly (Figs. 469, 472). Scale bars: 2 mm (460–462, 463–472)." captionText-12="Figures 473–476. Teruelius flavopiceus. Habitus. Male (473–474) and female (475–476), dorsal (473, 475) and ventral (474, 476) views. Scale bar: 10 mm." captionText-13="Figures 477–490. Teruelius flavopiceus, male. Figures 477–486. Pedipalp chela, dorsal (477), external (478) and ventrointernal (479) views; pedipalp patella, dorsal (480), external (481) and ventral (482) views; pedipalp femur and trochanter, internal (483) and dorsal (484) views; pedipalp chela, movable (485) and fixed (486) finger dentate margin. The trichobothrial pattern is indicated by white circles in Figures 477a–484a. Figures 487–490. Right legs tibia, basitarsus and telotarsus, retrolateral views. Leg IV (487), leg III (488), leg. II (489), leg I (490)." captionText-14="Figures 491–494. Teruelius grandidieri. Habitus. Male (491–492) and female (493–494), dorsal (491, 493) and ventral (492, 494) views. Scale bar: 10 mm." captionText-15="Figures 495–499. Teruelius intertidalis, female holotype. Figures 495–496. Habitus in dorsal (495) and ventral (496) views, and original labels. Figures 497–499. Metasoma in lateral (497), ventral (498) and dorsal (499) views. Scale bars: 10 mm." captionText-16="Figures 500–511. Teruelius intertidalis, female holotype. Figures 500–508. Pedipalp chela, dorsal (500), external (501) and ventrointernal (502) views; pedipalp patella, dorsal (503), external (504) and ventral (505) views; pedipalp femur and trochanter, internal (506) and dorsal (507) views; pedipalp chela, movable finger dentate margin (508). Figure 509. Carapace and tergites I–III. Figure 510. Sternopectinal region. Figure 511. Telson lateral view." captionText-17="Figures 512–515. Teruelius intertidalis, female holotype. Left legs tibia, basitarsus and telotarsus, retrolateral views. Leg I (512), leg II (513), leg III (514), leg IV (515)." captionText-18="Figures 516–521. Teruelius limbatus. Figures 516–519. Habitus. Male (516–517) and female (518–519), dorsal (516, 518) and ventral (517, 519) views. Figure 520. Pedipalp chela external, male. Figure 521. Pedipalp chela, movable finger dentate margin, female. Scale bar: 10 mm (516–519)." captionText-19="Figures 522–525. Teruelius mahafaliensis. Habitus. Male holotype of G. rossii (522–523) and male from near the type locality of T. mahafaliensis (524–525), dorsal (522, 524) and ventral (523, 525) views. Scale bar: 10 mm." captionText-2="Figures 409–417. Teruelius ankarafantsika. Male paratype (409, 416–417) and female holotype (410–415). Figures 409–411. Carapace and tergites I–III (409–410) and sternopectinal region (411). Figures 412–417. Left legs tibia, basitarsus and telotarsus, retrolateral views. Leg I (412), leg II (413), leg III (414, 416), leg IV (415, 417)." captionText-20="Figures 526–537. Teruelius mahafaliensis, female (526, 529–530, 535), male (527–528, 531–534) and male holotype of G. rossii (536–537). Sternopectinal region (526–527). Carapace and tergites I–II (528). Pedipalp chela, movable finger dentate margin (529). Pedipalp chela dorsal (530–531) and external (532) views. Pedipalp patella dorsal (533) view. Pedipalp femur and trochanter dorsal (534) view. Metasoma V and telson lateral (535) view. Metasoma and telson dorsal (536) and ventral (537) views. Scale bar: 10 mm (536–537)." captionText-21="Figures 538–541. Teruelius mahafaliensis, male holotype of G. rossii. Left legs tibia, basitarsus and telotarsus, retrolateral views. Leg I (538), leg II (539), leg. III (540), leg IV (541)." captionText-22="Figures 542–546. Teruelius olgae, male paratype.Figures 542–543. Habitus, dorsal (542) and ventral (543) views, and original labels. Figures 544–546. Metasoma and telson in lateral (544), ventral (545) and dorsal (546) views. Scale bars: 10 mm (542–543, 544–546)." captionText-23="Figures 547–551. Teruelius olgae, female. Figures 547–548. Habitus, dorsal (547) and ventral (548) views. Scale bar: 10 mm. Figures 549–551. Metasoma and telson. lateral (549), ventral (550) and dorsal (551) views. Scale bar: 5 mm." captionText-24="Figures 552–563. Teruelius olgae, male paratype. Figures 552–560. Pedipalp chela, dorsal (552), external (553) and ventrointernal (554) views; pedipalp patella, dorsal (555), external (556) and ventral (557) views; pedipalp femur and trochanter, internal (558) and dorsal (559) views; pedipalp chela, movable finger dentate margin (560). Figure 561. Carapace and tergites I–II. Figure 562. Sternopectinal region. Figure 563. Telson lateral view." captionText-25="Figures 564–575. Teruelius olgae. Female pedipalp. Chela in dorsal (564), external (565), ventral (566) and internal (567) views. Patella in dorsal (568), external (569), ventral (570) and internal (571) views. Femur and trochanter in dorsal (572), internal (573), external (574) and ventral (575) views. Scale bar: 2 mm." captionText-26="Figures 576–579. Teruelius olgae, male paratype. Right legs tibia, basitarsus and telotarsus, retrolateral and ventral views. Leg I (576), leg II (577), leg III (578), leg IV (579)." captionText-3="Figures 418–419. Teruelius ankarana. Habitus. Male in dorsal (418) and ventral (419) views. Scale bar: 10 mm." captionText-4="Figures 420–421. Teruelius ankarana. Habitus. Female in dorsal (420) and ventral (421) views. Scale bar: 10 mm." captionText-5="Figures 422–425. Teruelius annulatus. Habitus. Male (422–423) and female (424–425), dorsal (422, 424) and ventral (423, 425) views. Scale bar: 10 mm." captionText-6="Figures 426–435. Figures 426–432. Teruelius annulatus. Male (426, 428, 430–431) and female (427, 429). Sternopectinal region (426–427). Carapace and tergites I–III (428). Pedipalp chela dorsal (429–430). Pedipalp chela, movable finger dentate margin (431). Metasoma V and telson lateroventral view (432). Figures 433–435. Teruelius bistriatus. Female (433–434) and male (435) syntypes. Sternopectinal region (433, 435). Carapace and tergites I–III (434)." captionText-7="Figures 436–438. Teruelius bistriatus. Habitus. Female syntype in dorsal (436) and ventral (437) views, and original labels (438). Scale bar: 10 mm." captionText-8="Figures 439–446. Teruelius bistriatus. Metasoma and telson. Male (439, 441–443) and female syntypes (440, 444–446), lateral (439–441, 444), ventral (442, 445) and dorsal (443, 446) views. Scale bars: 10 mm (441–443, 444–446)." captionText-9="Figures 447–458. Teruelius bistriatus. Pedipalp. Female (447–452) and male (453–458) syntypes. Pedipalp chela, dorsal (447, 453), external (448, 454) and ventrointernal (449, 455) views; pedipalp patella, dorsal (450, 456) and ventral (457) views; pedipalp femur and trochanter in dorsal (451, 458) views; pedipalp chela, movable finger dentate margin (452). Pedipalp of male is mirrored." figureDoi-0="http://doi.org/10.5281/zenodo.7143772" figureDoi-1="http://doi.org/10.5281/zenodo.7143774" figureDoi-10="http://doi.org/10.5281/zenodo.7143796" figureDoi-11="http://doi.org/10.5281/zenodo.7143798" figureDoi-12="http://doi.org/10.5281/zenodo.7143801" figureDoi-13="http://doi.org/10.5281/zenodo.7143803" figureDoi-14="http://doi.org/10.5281/zenodo.7143805" figureDoi-15="http://doi.org/10.5281/zenodo.7143809" figureDoi-16="http://doi.org/10.5281/zenodo.7143811" figureDoi-17="http://doi.org/10.5281/zenodo.7143813" figureDoi-18="http://doi.org/10.5281/zenodo.7143815" figureDoi-19="http://doi.org/10.5281/zenodo.7143817" figureDoi-2="http://doi.org/10.5281/zenodo.7143778" figureDoi-20="http://doi.org/10.5281/zenodo.7143819" figureDoi-21="http://doi.org/10.5281/zenodo.7143821" figureDoi-22="http://doi.org/10.5281/zenodo.7143823" figureDoi-23="http://doi.org/10.5281/zenodo.7143825" figureDoi-24="http://doi.org/10.5281/zenodo.7143827" figureDoi-25="http://doi.org/10.5281/zenodo.7143829" figureDoi-26="http://doi.org/10.5281/zenodo.7143831" figureDoi-3="http://doi.org/10.5281/zenodo.7143780" figureDoi-4="http://doi.org/10.5281/zenodo.7143782" figureDoi-5="http://doi.org/10.5281/zenodo.7143784" figureDoi-6="http://doi.org/10.5281/zenodo.7143788" figureDoi-7="http://doi.org/10.5281/zenodo.7143790" figureDoi-8="http://doi.org/10.5281/zenodo.7143792" figureDoi-9="http://doi.org/10.5281/zenodo.7143794" httpUri-0="https://zenodo.org/record/7143772/files/figure.png" httpUri-1="https://zenodo.org/record/7143774/files/figure.png" httpUri-10="https://zenodo.org/record/7143796/files/figure.png" httpUri-11="https://zenodo.org/record/7143798/files/figure.png" httpUri-12="https://zenodo.org/record/7143801/files/figure.png" httpUri-13="https://zenodo.org/record/7143803/files/figure.png" httpUri-14="https://zenodo.org/record/7143805/files/figure.png" httpUri-15="https://zenodo.org/record/7143809/files/figure.png" httpUri-16="https://zenodo.org/record/7143811/files/figure.png" httpUri-17="https://zenodo.org/record/7143813/files/figure.png" httpUri-18="https://zenodo.org/record/7143815/files/figure.png" httpUri-19="https://zenodo.org/record/7143817/files/figure.png" httpUri-2="https://zenodo.org/record/7143778/files/figure.png" httpUri-20="https://zenodo.org/record/7143819/files/figure.png" httpUri-21="https://zenodo.org/record/7143821/files/figure.png" httpUri-22="https://zenodo.org/record/7143823/files/figure.png" httpUri-23="https://zenodo.org/record/7143825/files/figure.png" httpUri-24="https://zenodo.org/record/7143827/files/figure.png" httpUri-25="https://zenodo.org/record/7143829/files/figure.png" httpUri-26="https://zenodo.org/record/7143831/files/figure.png" httpUri-3="https://zenodo.org/record/7143780/files/figure.png" httpUri-4="https://zenodo.org/record/7143782/files/figure.png" httpUri-5="https://zenodo.org/record/7143784/files/figure.png" httpUri-6="https://zenodo.org/record/7143788/files/figure.png" httpUri-7="https://zenodo.org/record/7143790/files/figure.png" httpUri-8="https://zenodo.org/record/7143792/files/figure.png" httpUri-9="https://zenodo.org/record/7143794/files/figure.png" pageId="13" pageNumber="12">387–579</figureCitation>
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||
,
|
||
<figureCitation id="13292A1AFFBCFFAA23B2D3E5FAF1FEF2" box="[1251,1347,668,692]" captionStart-0="Figures 584–587" captionStart-1="Figures 588–589" captionStart-2="Figures 590–592" captionStart-3="Figures 593–594" captionStart-4="Figures 595–596" captionStart-5="Figures 597–599" captionStart-6="Figures 600–601" captionStart-7="Figures 602–603" captionStart-8="Figures 604–605" captionStartId-0="116.[113,190,1984,2006]" captionStartId-1="117.[113,190,2007,2029]" captionStartId-2="118.[113,190,2005,2027]" captionStartId-3="119.[113,190,2011,2033]" captionStartId-4="120.[113,190,2011,2033]" captionStartId-5="121.[113,190,2001,2023]" captionStartId-6="122.[113,190,2003,2025]" captionStartId-7="123.[113,190,2001,2023]" captionStartId-8="124.[113,190,1999,2021]" captionTargetBox-0="[113,1473,234,1973]" captionTargetBox-1="[113,1474,234,1993]" captionTargetBox-2="[113,1474,234,1993]" captionTargetBox-3="[113,1473,234,1993]" captionTargetBox-4="[113,1474,234,1993]" captionTargetBox-5="[113,1474,236,1988]" captionTargetBox-6="[113,1474,236,1987]" captionTargetBox-7="[113,1474,232,1975]" captionTargetBox-8="[113,1474,234,1984]" captionTargetId-0="figure-26@116.[113,1473,234,1975]" captionTargetId-1="figure-17@117.[110,1476,234,1995]" captionTargetId-2="figure-19@118.[111,1476,234,1996]" captionTargetId-3="figure-17@119.[113,1473,234,1996]" captionTargetId-4="figure-20@120.[113,1474,231,1995]" captionTargetId-5="figure-21@121.[113,1474,233,1990]" captionTargetId-6="figure-17@122.[113,1474,236,1990]" captionTargetId-7="figure-19@123.[113,1474,232,1975]" captionTargetId-8="figure-21@124.[109,1474,234,1986]" captionTargetPageId-0="116" captionTargetPageId-1="117" captionTargetPageId-2="118" captionTargetPageId-3="119" captionTargetPageId-4="120" captionTargetPageId-5="121" captionTargetPageId-6="122" captionTargetPageId-7="123" captionTargetPageId-8="124" captionText-0="Figures 584–587. Teruelius ankarana. Juveniles before second (584 top), after second (584 below and 585), third (586) and fourth (587) ecdysis." captionText-1="Figures 588–589. Teruelius ankarana. Male (588) and female (589), in vivo habitus." captionText-2="Figures 590–592. Teruelius flavopiceus. Juveniles after second (590), third (591) and fourth (592) ecdysis." captionText-3="Figures 593–594. Teruelius flavopiceus. Male (593) and female (594), in vivo habitus." captionText-4="Figures 595–596. Teruelius flavopiceus. Female with newborns (595) and with juveniles after first ecdysis (596)." captionText-5="Figures 597–599. Teruelius flavopiceus. Juvenile with duplicated metasoma (597, 599) and its unsuccesful second ecdysis (598)." captionText-6="Figures 600–601. Teruelius grandidieri. Male (600) and female (601) in vivo habitus." captionText-7="Figures 602–603. Teruelius grandidieri. Female with newborns (602) and with juveniles after first ecdysis (603)." captionText-8="Figures 604–605. Teruelius limbatus. Male (in the middle) and females (604) and female with newborns (605)." figureDoi-0="http://doi.org/10.5281/zenodo.7143839" figureDoi-1="http://doi.org/10.5281/zenodo.7143843" figureDoi-2="http://doi.org/10.5281/zenodo.7143845" figureDoi-3="http://doi.org/10.5281/zenodo.7143847" figureDoi-4="http://doi.org/10.5281/zenodo.7143849" figureDoi-5="http://doi.org/10.5281/zenodo.7143851" figureDoi-6="http://doi.org/10.5281/zenodo.7143853" figureDoi-7="http://doi.org/10.5281/zenodo.7143857" figureDoi-8="http://doi.org/10.5281/zenodo.7143859" httpUri-0="https://zenodo.org/record/7143839/files/figure.png" httpUri-1="https://zenodo.org/record/7143843/files/figure.png" httpUri-2="https://zenodo.org/record/7143845/files/figure.png" httpUri-3="https://zenodo.org/record/7143847/files/figure.png" httpUri-4="https://zenodo.org/record/7143849/files/figure.png" httpUri-5="https://zenodo.org/record/7143851/files/figure.png" httpUri-6="https://zenodo.org/record/7143853/files/figure.png" httpUri-7="https://zenodo.org/record/7143857/files/figure.png" httpUri-8="https://zenodo.org/record/7143859/files/figure.png" pageId="13" pageNumber="12">584–605</figureCitation>
|
||
,
|
||
<tableCitation id="C6900324FFBCFFAA221CD3E5FA07FEF2" box="[1357,1461,668,692]" captionStart-0="Table 1" captionStart-1="Table 2" captionStart-2="Table 3" captionStart-3="Table 4" captionStartId-0="9.[115,171,1146,1167]" captionStartId-1="10.[113,169,1291,1312]" captionStartId-2="16.[110,166,1277,1298]" captionStartId-3="17.[113,169,1270,1291]" captionTargetId-0="graphics-417@9.[119,1468,237,1129]" captionTargetId-1="graphics-433@10.[121,1460,237,1263]" captionTargetId-2="graphics-479@16.[118,1471,242,1254]" captionTargetId-3="graphics-446@17.[119,1470,248,1242]" captionTargetPageId-0="9" captionTargetPageId-1="10" captionTargetPageId-2="16" captionTargetPageId-3="17" captionText-0="Table 1. Character states for species examined. States of 5 characters for Grosphus (4 spp., blue panels), Neogrosphus (1 sp., magenta panels) and Teruelius gen. n. (11 spp., yellow panels) examined in this study: position of trichobothrium d 2 on pedipalp femur; ratio R123 = d(Eb 2, Eb 3)/ d(Eb 1, Eb 2) of distances between pedipalp manus trichobothria Eb 1, Eb 2 and Eb 3; ranges of pectinal tooth counts (PTC) of males (♂) and females (♀); and size and shape of female basal pectinal tooth." captionText-1="Table 2. Character states for species examined. States of 6 characters for Grosphus (4 spp., blue panels), Neogrosphus (1 sp., magenta panels) and Teruelius gen. n. (11 spp., yellow panels) examined in this study: position of hemispermatophore basal lobe relative to base of flagellum; size of hemispermatophore posterior lobe; shape of spiracles; condition of metasoma I ventromedian carinae; telotarsal setation; and intensity of UV fluorescence." captionText-2="Table 3. Character states for species not examined. States of 6 characters for Grosphus (6 spp., blue panels), Neogrosphus (2 spp., magenta panels) and Teruelius gen. n. (7 spp., yellow panels) not examined in this study: position of trichobothrium d on pedipalp femur; ratio R = 2 123 d(Eb 2, Eb 3)/ d(Eb 1, Eb 2) of distances between pedipalp manus trichobothria Eb 1, Eb 2 and Eb 3; ranges of pectinal tooth counts (PTC) of males (♂) and females (♀);size and shape of female basal pectinal tooth; shape of spiracles; and telotarsal setation. Character states inferred from published descriptions and figures. PTC and basal pectinal tooth characters not indicated in cases where either males or females are not known or described. Other characters not indicated in cases where published descriptions did not yield clear information." captionText-3="Table 4. Character states for ‘Grosphus’ group genera and outgroup genus Pseudolychas. States of 11 characters for Grosphus, Neogrosphus, Teruelius gen. n. and Pseudolychas, summarizing data of Tables 1–3. Pseudolychas character states determined by study of materials, and from published descriptions. Primitive states indicated by green panels, derived states by red panels, with polarities determined by comparison to Pseudolychas which is assumed primitive. Unpolarized characters indicated by dark green panels." httpUri-0="http://table.plazi.org/id/DF6D6617FFB8FFAE2722D503FCADF8A1" httpUri-2="http://table.plazi.org/id/DF6D6617FFA1FFB7273FD584FB00F9E5" httpUri-3="http://table.plazi.org/id/DF6D6617FFA0FFB62720D58FFB7EF924" pageId="13" pageNumber="12" tableUuid-0="DF6D6617FFB8FFAE2722D503FCADF8A1" tableUuid-2="DF6D6617FFA1FFB7273FD584FB00F9E5" tableUuid-3="DF6D6617FFA0FFB62720D58FFB7EF924">Tabs. 1–4</tableCitation>
|
||
)
|
||
</paragraph>
|
||
<paragraph id="8BAD369FFFBCFFAA246DD3A5FAF5FF52" blockId="13.[828,1466,732,788]" pageId="13" pageNumber="12">
|
||
<emphasis id="B966EA8DFFBCFFAA246DD3A5FAF5FF52" bold="true" pageId="13" pageNumber="12">http://zoobank.org/urn:lsid:zoobank.org:act:54CB8128- BCFD-4B1F-A947-153C7CDD5B83</emphasis>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3086514FFBCFFAA2463D245FADEFFD2" pageId="13" pageNumber="12" type="reference_group">
|
||
<paragraph id="8BAD369FFFBCFFAA2463D245FADEFFD2" blockId="13.[818,1477,828,916]" pageId="13" pageNumber="12">
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||
<treatmentCitationGroup id="AB0211B1FFBCFFAA2463D245FADEFFD2" pageId="13" pageNumber="12">
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||
<treatmentCitation id="0AB3108EFFBCFFAA2463D245FBC5FF12" author="VACHON" box="[818,1143,828,852]" page="254" pageId="13" pageNumber="12" year="1940">
|
||
<taxonomicName id="4C124D1CFFBCFFAA2463D245FBC5FF12" ID-CoL="4R7C" authority="Vachon, 1940: 254" authorityName="Vachon" authorityPageNumber="254" authorityYear="1940" box="[818,1143,828,852]" class="Arachnida" family="Buthidae" genus="Grosphus" kingdom="Animalia" order="Scorpiones" pageId="13" pageNumber="12" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B966EA8DFFBCFFAA2463D245FC2BFF12" box="[818,921,828,852]" italics="true" pageId="13" pageNumber="12">Grosphus</emphasis>
|
||
<bibRefCitation id="EF834B6EFFBCFFAA24F5D245FBC5FF12" author="Vachon" box="[932,1143,828,852]" firstAuthor="Vachon" page="254" pageId="13" pageNumber="12" pagination="241 - 260" refId="ref43710" refString="VACHON, M. 1940. Sur la systematique des Scorpions. Memoires du Museum national d'Histoire Naturelle, Paris, 13 (2): 241 - 260." type="journal article" year="1940">Vachon, 1940: 254</bibRefCitation>
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||
</taxonomicName>
|
||
</treatmentCitation>
|
||
, 256;
|
||
<treatmentCitation id="0AB3108EFFBCFFAA2391D245FC3FFF32" author="LOURENCO, W. R. & V. SOARIMALALA & S. M. GOODMAN" page="375" pageId="13" pageNumber="12" year="2007">
|
||
<bibRefCitation id="EF834B6EFFBCFFAA2391D245FC3FFF32" author="LOURENCO, W. R. & V. SOARIMALALA & S. M. GOODMAN" journalOrPublisher="Boletin de la Sociedad Entomologica Aragonesa" pageId="13" pageNumber="12" pagination="369 - 375" part="41" refId="ref39953" refString="LOURENCO, W. R., V. SOARIMALALA & S. M. GOODMAN. 2007 b. Scorpions of south-west Madagascar. II. The species of Grosphus Simon (Scorpiones, Buthidae). Boletin de la Sociedad Entomologica Aragonesa, 41: 369 - 375." title="Scorpions of south-west Madagascar. II. The species of Grosphus Simon (Scorpiones, Buthidae)" type="journal article" year="2007">Lourenço et al., 2007b: 375</bibRefCitation>
|
||
</treatmentCitation>
|
||
;
|
||
<treatmentCitation id="0AB3108EFFBCFFAA24F3D225FAE3FF32" author="PRENDINI, L. & L. ESPOSITO" box="[930,1361,860,884]" page="675" pageId="13" pageNumber="12" year="2010">
|
||
<bibRefCitation id="EF834B6EFFBCFFAA24F3D225FAE3FF32" author="PRENDINI, L. & L. ESPOSITO" box="[930,1361,860,884]" journalOrPublisher="Zoological Journal of the Linnean Society" pageId="13" pageNumber="12" pagination="673 - 710" part="159" refId="ref42182" refString="PRENDINI, L. & L. ESPOSITO. 2010. A reanalysis of Parabuthus (Scorpiones: Buthidae) phylogeny with descriptions of two new Parabuthus species endemic to the Central Namib gravel plains, Namibia. Zoological Journal of the Linnean Society, 159: 673 - 710." title="A reanalysis of Parabuthus (Scorpiones: Buthidae) phylogeny with descriptions of two new Parabuthus species endemic to the Central Namib gravel plains, Namibia" type="journal article" year="2010">Prendini & Esposito, 2010: 675–676</bibRefCitation>
|
||
</treatmentCitation>
|
||
;
|
||
<treatmentCitation id="0AB3108EFFBCFFAA2234D225FB99FFD2" author="LORIA, S. F. & L. PRENDINI" page="25" pageId="13" pageNumber="12" year="2014">
|
||
<bibRefCitation id="EF834B6EFFBCFFAA2234D225FB99FFD2" author="LORIA, S. F. & L. PRENDINI" journalOrPublisher="PLoS ONE" pageId="13" pageNumber="12" pagination="112913" part="9" refId="ref37170" refString="LORIA, S. F. & L. PRENDINI. 2014. Homology of the lateral eyes of Scorpiones: a six-ocellus model. PLoS ONE 9 (12): e 112913. doi: 10.1371 / journal. pone. 0112913." title="Homology of the lateral eyes of Scorpiones: a six-ocellus model" type="journal article" year="2014">Loria & Prendini, 2014: 25</bibRefCitation>
|
||
</treatmentCitation>
|
||
;
|
||
<treatmentCitation id="0AB3108EFFBCFFAA2367D205FADAFFD2" author="LORIA, S. F. & L. PRENDINI" box="[1078,1384,892,916]" page="184" pageId="13" pageNumber="12" year="2018">
|
||
<bibRefCitation id="EF834B6EFFBCFFAA2367D205FADAFFD2" author="LORIA, S. F. & L. PRENDINI" box="[1078,1384,892,916]" journalOrPublisher="Zoologischer Anzeiger" pageId="13" pageNumber="12" pagination="183 - 191" part="273" refId="ref37215" refString="LORIA, S. F. & L. PRENDINI. 2018. Ultrastructural comparison of the eyespot and ocelli of scorpions, and implications for the systematics of Chaerilidae Pocock, 1893. Zoologischer Anzeiger, 273: 183 - 191." title="Ultrastructural comparison of the eyespot and ocelli of scorpions, and implications for the systematics of Chaerilidae Pocock, 1893" type="journal article" year="2018">Loria & Prendini, 2018: 184</bibRefCitation>
|
||
</treatmentCitation>
|
||
.
|
||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3086514FFBCFFAA2463D2C5FAA7FF92" box="[818,1301,956,980]" pageId="13" pageNumber="12" type="type_taxon">
|
||
<paragraph id="8BAD369FFFBCFFAA2463D2C5FAA7FF92" blockId="13.[818,1301,956,980]" box="[818,1301,956,980]" pageId="13" pageNumber="12">
|
||
<typeStatus id="54A9883DFFBCFFAA2463D2C5FCDAFF95" box="[818,872,956,979]" pageId="13" pageNumber="12">TYPE</typeStatus>
|
||
SPECIES.
|
||
<taxonomicName id="4C124D1CFFBCFFAA2498D2C5FAA3FF92" authority="Pocock, 1889" authorityName=", Pocock" authorityYear="1889" box="[969,1297,956,980]" class="Arachnida" family="Buthidae" genus="Buthus" kingdom="Animalia" order="Scorpiones" pageId="13" pageNumber="12" phylum="Arthropoda" rank="species" species="limbatus">
|
||
<emphasis id="B966EA8DFFBCFFAA2498D2C5FBC4FF92" box="[969,1142,956,980]" italics="true" pageId="13" pageNumber="12">Buthus limbatus</emphasis>
|
||
<bibRefCitation id="EF834B6EFFBCFFAA232DD2C5FAA3FF92" author="POCOCK, R. I." box="[1148,1297,956,980]" journalOrPublisher="Annals and Magazine of Natural History" pageId="13" pageNumber="12" pagination="334 - 351" part="3" refId="ref41689" refString="POCOCK, R. I. 1889 a. Notes on some Buthidae, new and old. Annals and Magazine of Natural History, 3: 334 - 351." title="Notes on some Buthidae, new and old" type="journal article" year="1889">Pocock, 1889</bibRefCitation>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3086514FFBCFFAA2463D285FC12F832" pageId="13" pageNumber="12" type="etymology">
|
||
<paragraph id="8BAD369FFFBCFFAA2463D285FC12F832" blockId="13.[818,1476,1020,1140]" pageId="13" pageNumber="12">
|
||
ETYMOLOGY. The generic epithet
|
||
<taxonomicName id="4C124D1CFFBCFFAA23F4D285FAB5F852" authorityName="Lowe & Kovařík" authorityYear="2019" box="[1189,1287,1020,1044]" class="Arachnida" family="Buthidae" genus="Teruelius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="13" pageNumber="12" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B966EA8DFFBCFFAA23F4D285FAB5F852" box="[1189,1287,1020,1044]" italics="true" pageId="13" pageNumber="12">Teruelius</emphasis>
|
||
</taxonomicName>
|
||
(masculine) is a patronym honoring Rolando Teruel from
|
||
<collectingCountry id="F305760FFFBCFFAA23BED565FA95F872" box="[1263,1319,1052,1076]" name="Cuba" pageId="13" pageNumber="12">Cuba</collectingCountry>
|
||
in recognition of his many important contributions to the knowledge of scorpions.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3086514FFBCFFAA2463D5E5FC56FB92" pageId="13" pageNumber="12" type="diagnosis">
|
||
<paragraph id="8BAD369FFFBCFFAA2463D5E5FC56FB92" blockId="13.[818,1476,1180,2004]" pageId="13" pageNumber="12">
|
||
DIAGNOSIS. A member of the ‘
|
||
<taxonomicName id="4C124D1CFFBCFFAA232BD5E5FB53F8F2" authorityName="Vachon" authorityYear="1940" box="[1146,1249,1180,1204]" class="Arachnida" family="Buthidae" genus="Grosphus" kingdom="Animalia" order="Scorpiones" pageId="13" pageNumber="12" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B966EA8DFFBCFFAA232BD5E5FB53F8F2" box="[1146,1249,1180,1204]" italics="true" pageId="13" pageNumber="12">Grosphus</emphasis>
|
||
</taxonomicName>
|
||
’ group differentiated as follows: medium-sized to large-sized scorpions, adults ca.
|
||
<quantity id="4CEA9B7AFFBCFFAA2463D5A5FC04F8B5" box="[818,950,1244,1268]" metricMagnitude="-2" metricUnit="m" metricValue="7.75" metricValueMax="12.0" metricValueMin="3.5" pageId="13" pageNumber="12" unit="mm" value="77.5" valueMax="120.0" valueMin="35.0">35–120 mm</quantity>
|
||
in length; pedipalp finger granule rows 10–15 (
|
||
<figureCitation id="13292A1AFFBCFFAA246BD585FC10F952" box="[826,930,1276,1300]" captionStart="Figures 391–408" captionStartId="88.[113,190,1919,1941]" captionTargetBox="[125,1462,242,1860]" captionTargetId="figure-93@88.[113,1474,234,1872]" captionTargetPageId="88" captionText="Figures 391–408. Teruelius ankarafantsika. Male paratype (391–393, 404–405) and female holotype (394–403, 406–408). Figures 391–402. Pedipalp.Chela in dorsal (391, 394), external (392, 395) and ventral (396) views. Patella in dorsal (397), external (398) and ventral (399) views. Femur and trochanter in internal (393, 400) and dorsal (401) views. Pedipalp chela, movable finger dentate margin (402). Figures 403–408. Telson in lateral (403, 404) views. Metasoma and telson in lateral (405, 406), dorsal (407) and ventral (408) views.Scale bar: 10 mm (406–408)." figureDoi="http://doi.org/10.5281/zenodo.7143774" httpUri="https://zenodo.org/record/7143774/files/figure.png" pageId="13" pageNumber="12">Figs. 402</figureCitation>
|
||
,
|
||
<figureCitation id="13292A1AFFBCFFAA24FFD585FC6AF952" box="[942,984,1276,1300]" captionStart="Figures 426–435" captionStartId="93.[113,190,1928,1950]" captionTargetBox="[113,1474,234,1874]" captionTargetId="figure-72@93.[113,1474,234,1874]" captionTargetPageId="93" captionText="Figures 426–435. Figures 426–432. Teruelius annulatus. Male (426, 428, 430–431) and female (427, 429). Sternopectinal region (426–427). Carapace and tergites I–III (428). Pedipalp chela dorsal (429–430). Pedipalp chela, movable finger dentate margin (431). Metasoma V and telson lateroventral view (432). Figures 433–435. Teruelius bistriatus. Female (433–434) and male (435) syntypes. Sternopectinal region (433, 435). Carapace and tergites I–III (434)." figureDoi="http://doi.org/10.5281/zenodo.7143788" httpUri="https://zenodo.org/record/7143788/files/figure.png" pageId="13" pageNumber="12">431</figureCitation>
|
||
,
|
||
<figureCitation id="13292A1AFFBCFFAA24B5D585FBBDF952" box="[996,1039,1276,1300]" captionStart="Figures 447–458" captionStartId="96.[113,190,1958,1980]" captionTargetBox="[133,1446,260,1921]" captionTargetId="figure-61@96.[113,1474,236,1929]" captionTargetPageId="96" captionText="Figures 447–458. Teruelius bistriatus. Pedipalp. Female (447–452) and male (453–458) syntypes. Pedipalp chela, dorsal (447, 453), external (448, 454) and ventrointernal (449, 455) views; pedipalp patella, dorsal (450, 456) and ventral (457) views; pedipalp femur and trochanter in dorsal (451, 458) views; pedipalp chela, movable finger dentate margin (452). Pedipalp of male is mirrored." figureDoi="http://doi.org/10.5281/zenodo.7143794" httpUri="https://zenodo.org/record/7143794/files/figure.png" pageId="13" pageNumber="12">452</figureCitation>
|
||
,
|
||
<figureCitation id="13292A1AFFBCFFAA234BD585FBC8F952" box="[1050,1146,1276,1300]" captionStart="Figures 477–490" captionStartId="100.[113,190,1928,1950]" captionTargetBox="[110,1468,256,1846]" captionTargetId="figure-92@100.[110,1471,244,1880]" captionTargetPageId="100" captionText="Figures 477–490. Teruelius flavopiceus, male. Figures 477–486. Pedipalp chela, dorsal (477), external (478) and ventrointernal (479) views; pedipalp patella, dorsal (480), external (481) and ventral (482) views; pedipalp femur and trochanter, internal (483) and dorsal (484) views; pedipalp chela, movable (485) and fixed (486) finger dentate margin. The trichobothrial pattern is indicated by white circles in Figures 477a–484a. Figures 487–490. Right legs tibia, basitarsus and telotarsus, retrolateral views. Leg IV (487), leg III (488), leg. II (489), leg I (490)." figureDoi="http://doi.org/10.5281/zenodo.7143803" httpUri="https://zenodo.org/record/7143803/files/figure.png" pageId="13" pageNumber="12">485–486</figureCitation>
|
||
,
|
||
<figureCitation id="13292A1AFFBCFFAA23D7D585FB02F952" box="[1158,1200,1276,1300]" captionStart="Figures 500–511" captionStartId="103.[113,190,1928,1950]" captionTargetBox="[113,1465,240,1881]" captionTargetId="figure-73@103.[113,1474,234,1881]" captionTargetPageId="103" captionText="Figures 500–511. Teruelius intertidalis, female holotype. Figures 500–508. Pedipalp chela, dorsal (500), external (501) and ventrointernal (502) views; pedipalp patella, dorsal (503), external (504) and ventral (505) views; pedipalp femur and trochanter, internal (506) and dorsal (507) views; pedipalp chela, movable finger dentate margin (508). Figure 509. Carapace and tergites I–III. Figure 510. Sternopectinal region. Figure 511. Telson lateral view." figureDoi="http://doi.org/10.5281/zenodo.7143811" httpUri="https://zenodo.org/record/7143811/files/figure.png" pageId="13" pageNumber="12">508</figureCitation>
|
||
,
|
||
<figureCitation id="13292A1AFFBCFFAA23EDD585FB54F952" box="[1212,1254,1276,1300]" captionStart="Figures 516–521" captionStartId="105.[113,190,1958,1980]" captionTargetBox="[146,1426,242,1905]" captionTargetId="figure-51@105.[136,1452,232,1918]" captionTargetPageId="105" captionText="Figures 516–521. Teruelius limbatus. Figures 516–519. Habitus. Male (516–517) and female (518–519), dorsal (516, 518) and ventral (517, 519) views. Figure 520. Pedipalp chela external, male. Figure 521. Pedipalp chela, movable finger dentate margin, female. Scale bar: 10 mm (516–519)." figureDoi="http://doi.org/10.5281/zenodo.7143815" httpUri="https://zenodo.org/record/7143815/files/figure.png" pageId="13" pageNumber="12">521</figureCitation>
|
||
,
|
||
<figureCitation id="13292A1AFFBCFFAA23A2D585FAACF952" box="[1267,1310,1276,1300]" captionStart="Figures 526–537" captionStartId="107.[113,190,1928,1950]" captionTargetBox="[113,1458,234,1912]" captionTargetId="figure-80@107.[113,1468,234,1919]" captionTargetPageId="107" captionText="Figures 526–537. Teruelius mahafaliensis, female (526, 529–530, 535), male (527–528, 531–534) and male holotype of G. rossii (536–537). Sternopectinal region (526–527). Carapace and tergites I–II (528). Pedipalp chela, movable finger dentate margin (529). Pedipalp chela dorsal (530–531) and external (532) views. Pedipalp patella dorsal (533) view. Pedipalp femur and trochanter dorsal (534) view. Metasoma V and telson lateral (535) view. Metasoma and telson dorsal (536) and ventral (537) views. Scale bar: 10 mm (536–537)." figureDoi="http://doi.org/10.5281/zenodo.7143819" httpUri="https://zenodo.org/record/7143819/files/figure.png" pageId="13" pageNumber="12">529</figureCitation>
|
||
,
|
||
<figureCitation id="13292A1AFFBCFFAA2278D585FAE7F952" box="[1321,1365,1276,1300]" captionStart="Figures 552–563" captionStartId="111.[113,190,1924,1946]" captionTargetBox="[113,1464,245,1912]" captionTargetId="figure-71@111.[113,1474,236,1915]" captionTargetPageId="111" captionText="Figures 552–563. Teruelius olgae, male paratype. Figures 552–560. Pedipalp chela, dorsal (552), external (553) and ventrointernal (554) views; pedipalp patella, dorsal (555), external (556) and ventral (557) views; pedipalp femur and trochanter, internal (558) and dorsal (559) views; pedipalp chela, movable finger dentate margin (560). Figure 561. Carapace and tergites I–II. Figure 562. Sternopectinal region. Figure 563. Telson lateral view." figureDoi="http://doi.org/10.5281/zenodo.7143827" httpUri="https://zenodo.org/record/7143827/files/figure.png" pageId="13" pageNumber="12">560</figureCitation>
|
||
), movable finger typically with 4 external subdistal granules; femur trichobothrium
|
||
<emphasis id="B966EA8DFFBCFFAA248DD445FC5BF912" box="[988,1001,1340,1364]" italics="true" pageId="13" pageNumber="12">d</emphasis>
|
||
<subScript id="179634DAFFBCFFAA24B8D435FC43F91C" attach="left" box="[1001,1009,1356,1370]" fontSize="6" pageId="13" pageNumber="12">2</subScript>
|
||
straddling dorsointernal carina, or located on dorsal surface (
|
||
<figureCitation id="13292A1AFFBCFFAA235CD425FB2AF932" box="[1037,1176,1372,1396]" captionStart="Figures 9–20" captionStartId="11.[116,190,1913,1935]" captionTargetBox="[129,1461,231,1903]" captionTargetId="figure-144@11.[129,1461,231,1903]" captionTargetPageId="11" captionText="Figures 9–20. Position of femur trichobothrium d 2in Grosphus and Teruelius gen. n. Dorsal surfaces of proximal pedipalp femur of adult females (♀, left panels) and adult males (♂, right panels), underUV fluorescence to highlightgranulation,carinae and trichobothrial areolae.Whitearrows indicate positions of trichobothrium d in each image. G. madagascariensis (9, 10; 2 samples from Anjiro (9) and Andasibe (10) show consistency of d 2 2 position),G. hirtus(11), G. voahangyae (12), T. ankarafantsika (13, 15; 2 samples fromAmpijoroa show consistency ofd 2position),T. ankarana (14), T. flavopiceus (16), T. grandidieri (17), T. limbatus (18), T. mahafaliensis (19), and T. olgae (20). Scale bars: 1 mm (9–20♀) or 500 μm (20 ♂)." figureDoi="http://doi.org/10.5281/zenodo.7143618" httpUri="https://zenodo.org/record/7143618/files/figure.png" pageId="13" pageNumber="12">Figs. 13–20</figureCitation>
|
||
); chela manus with petite trichobothrium
|
||
<emphasis id="B966EA8DFFBCFFAA248ED405FC4FF9D2" box="[991,1021,1404,1428]" italics="true" pageId="13" pageNumber="12">Eb</emphasis>
|
||
<subScript id="179634DAFFBCFFAA24ADD4F5FBB6F9DC" attach="left" box="[1020,1028,1420,1434]" fontSize="6" pageId="13" pageNumber="12">3</subScript>
|
||
near
|
||
<emphasis id="B966EA8DFFBCFFAA231AD405FBDBF9D2" box="[1099,1129,1404,1428]" italics="true" pageId="13" pageNumber="12">Eb</emphasis>
|
||
<subScript id="179634DAFFBCFFAA2339D4F5FBC2F9DC" attach="left" box="[1128,1136,1420,1434]" fontSize="6" pageId="13" pageNumber="12">2</subScript>
|
||
, closer than half the distance between
|
||
<emphasis id="B966EA8DFFBCFFAA24C6D4E5FC07F9F2" box="[919,949,1436,1460]" italics="true" pageId="13" pageNumber="12">Eb</emphasis>
|
||
<subScript id="179634DAFFBCFFAA24E5D4D5FC0EF9FC" attach="left" box="[948,956,1452,1466]" fontSize="6" pageId="13" pageNumber="12">1</subScript>
|
||
and
|
||
<emphasis id="B966EA8DFFBCFFAA24ABD4E5FBAAF9F2" box="[1018,1048,1436,1460]" italics="true" pageId="13" pageNumber="12">Eb</emphasis>
|
||
<subScript id="179634DAFFBCFFAA2349D4D5FB92F9FC" attach="left" box="[1048,1056,1452,1466]" fontSize="6" pageId="13" pageNumber="12">2</subScript>
|
||
(
|
||
<figureCitation id="13292A1AFFBCFFAA2362D4E5FB09F9F2" box="[1075,1211,1436,1460]" captionStart="Figures 21–27" captionStartId="14.[113,188,1859,1881]" captionTargetBox="[114,1474,227,1840]" captionTargetId="figure-206@14.[113,1474,222,1851]" captionTargetPageId="14" captionText="Figures 21–27. Positions of trichobothria Eb, Eb and petite Eb on manus of pedipalp chela in Grosphus, Neogrosphus and Teruelius gen. 1 2 3 n. Figures 21–24. External views of pedipalp chela, shown under UV fluorescence to highlight trichobothrial areolae. G. hirtus ♂ (21), G. madagascariensis ♀ (22), T. limbatus ♂ (23), T. ankarafantsika ♀ (24). Positions of Eb 1, Eb 2 and Eb 3 and lines joining them shown as white overlays. Scale bars: 2 mm. Figures 25–26. Horizontal histograms comparing ratios, R = d(Eb, Eb)/ d(Eb, Eb), of Eb -Eb distance, to Eb - 123 2 3 1 2 2 3 1 Eb distance, in males (25), and females (26) of Grosphus, Neogrosphus and Teruelius gen. n. Error bars indicate ranges (minimum, maximum), 2 histogram bars mid-range values. Figure 27. Scatter plot of male vs. female ratios R. Ratio is larger in males if points fall above the diagonal 123 (gray) line, larger in females if they fall below it." figureDoi="http://doi.org/10.5281/zenodo.7143620" httpUri="https://zenodo.org/record/7143620/files/figure.png" pageId="13" pageNumber="12">Figs. 23–24</figureCitation>
|
||
); manus trichobothrium
|
||
<emphasis id="B966EA8DFFBCFFAA2463D4C4FCF0F992" box="[818,834,1469,1492]" italics="true" pageId="13" pageNumber="12">V</emphasis>
|
||
<subScript id="179634DAFFBCFFAA2412D4B5FCF9F99C" attach="left" box="[835,843,1484,1498]" fontSize="6" pageId="13" pageNumber="12">2</subScript>
|
||
roughly collinear with
|
||
<emphasis id="B966EA8DFFBCFFAA2333D4C4FBC0F992" box="[1122,1138,1469,1492]" italics="true" pageId="13" pageNumber="12">V</emphasis>
|
||
<subScript id="179634DAFFBCFFAA2323D4B5FBC8F99C" attach="left" box="[1138,1146,1484,1498]" fontSize="6" pageId="13" pageNumber="12">1</subScript>
|
||
along chela axis or slightly displaced internally; higher pectinal tooth counts:
|
||
<specimenCount id="9D14FD16FFBCFFAA2204D4A5FADBF9B5" box="[1365,1385,1500,1523]" pageId="13" pageNumber="12" type="male">♂</specimenCount>
|
||
25– 41,
|
||
<specimenCount id="9D14FD16FFBCFFAA2463D485FCF4FA55" box="[818,838,1532,1555]" pageId="13" pageNumber="12" type="female">♀</specimenCount>
|
||
24–35 (
|
||
<figureCitation id="13292A1AFFBCFFAA24F1D485FB96FA52" box="[928,1060,1532,1556]" captionStart-0="Figures 28–29" captionStart-1="Figures 30–35" captionStartId-0="18.[113,190,1953,1975]" captionStartId-1="19.[113,190,1773,1795]" captionTargetBox-1="[118,1455,257,1710]" captionTargetId-0="figure-131@18.[113,1474,255,1915]" captionTargetId-1="figure-204@19.[113,1474,251,1714]" captionTargetPageId-0="18" captionTargetPageId-1="19" captionText-0="Figures 28–29. Variation in pectinal tooth count (PTC) in Grosphus, Neogrosphus and Teruelius gen. n. Horizontal histograms comparing PTC in males (28), and females (29) of Grosphus, Neogrosphus and Teruelius gen. n. Error bars indicate ranges (minimum, maximum), bars mid-range values." captionText-1="Figures 30–35. Variation in pectinal tooth count (PTC) in Grosphus, Neogrosphus and Teruelius gen. n. Figures 30–31. Logarithmic scatter plots showing scaling of PTC vs. carapace length in males (30), and females (31) of Grosphus, Neogrosphus and Teruelius gen. n. Color lines are least squares linear regression fits of scaling trends for the different genera. Figures 32–33. Logarithmic scatter plots showing scaling of PTC vs. carapace length in Figs. 30–31 superposed upon overall scaling trends for a larger sample of male (32) and female (33) buthids including both α and β trichobothrial configurations (gray symbols; 1055♂, 1046♀). Dark lines: least squares linear regression fits of scaling trends for larger sample of buthids. Figures 34–35. Logarithmic scatter plots showing scaling of PTC vs. carapace length in Figs. 30–31 superposed upon overall scaling trends for a larger sample of male (34) and female (35) α-buthids restricted to α trichobothrial configurations (gray symbols; 445♂, 371♀). Dark lines: least squares linear regression fits of scaling trends for α-buthids. Numbers: slopes of regression lines (= allometric exponents). Purple symbols and lines: outgroup Pseudolychas." figureDoi-0="http://doi.org/10.5281/zenodo.7143622" figureDoi-1="http://doi.org/10.5281/zenodo.7143626" httpUri-0="https://zenodo.org/record/7143622/files/figure.png" httpUri-1="https://zenodo.org/record/7143626/files/figure.png" pageId="13" pageNumber="12">Figs. 28–31</figureCitation>
|
||
); basal pectinal tooth of females wide, with elongate, tapering distal extension, distinctly longer than other teeth (
|
||
<figureCitation id="13292A1AFFBCFFAA2351D745FB39FA12" box="[1024,1163,1596,1620]" captionStart="Figures 40–51" captionStartId="22.[113,188,1930,1952]" captionTargetBox="[113,1474,234,1920]" captionTargetId="figure-124@22.[113,1474,234,1922]" captionTargetPageId="22" captionText="Figures 40–51. Female basal pectinal teeth in Grosphus and Teruelius gen. n. Ventral views of proximal left pectine of females shown under UV fluorescence to highlight cuticular surface texture, setation and absence of peg sensillae on basal tooth vs. their presence on other teeth. G. sp. nr hirtus (40), G. madagascariensis (41), G. hirtus (42), G. voahangyae (43), T. ankarafantsika (44–45; 2 samples fromAmpijoroa show variation in tooth shape), T. ankarana (46), T. flavopiceus (47), T. grandidieri (48), T. limbatus (49), T. mahafaliensis (50), and T. olgae (51). Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.7143634" httpUri="https://zenodo.org/record/7143634/files/figure.png" pageId="13" pageNumber="12">Figs. 44–51</figureCitation>
|
||
,
|
||
<figureCitation id="13292A1AFFBCFFAA23CFD745FB4CFA12" box="[1182,1278,1596,1620]" captionStart="Figures 196–210" captionStartId="65.[113,190,1950,1972]" captionTargetId="figure-77@65.[113,1474,234,1589]" captionTargetPageId="65" captionText="Figures 196–210. Female basal pectinal teeth of representative Grosphus and Teruelius gen. n. G. hirtus (196–198), G. madagascariensis (199), G. madagascariensis, paratype of G. mandena(200), T. ankarafantsika (201),T. ankarana (202), T. annulatus (203),T. bistriatus (204), T. flavopiceus (205), T. grandidieri (206), T. intertidalis (207), T. limbatus (208), T. mahafaliensis (209), T. feti (210)." figureDoi="http://doi.org/10.5281/zenodo.7143706" httpUri="https://zenodo.org/record/7143706/files/figure.png" pageId="13" pageNumber="12">201–210</figureCitation>
|
||
,
|
||
<figureCitation id="13292A1AFFBCFFAA2240D745FA89FA12" box="[1297,1339,1596,1620]" captionStart="Figures 409–417" captionStartId="89.[113,190,1958,1980]" captionTargetBox="[113,1474,234,1814]" captionTargetId="figure-58@89.[113,1474,234,1838]" captionTargetPageId="89" captionText="Figures 409–417. Teruelius ankarafantsika. Male paratype (409, 416–417) and female holotype (410–415). Figures 409–411. Carapace and tergites I–III (409–410) and sternopectinal region (411). Figures 412–417. Left legs tibia, basitarsus and telotarsus, retrolateral views. Leg I (412), leg II (413), leg III (414, 416), leg IV (415, 417)." figureDoi="http://doi.org/10.5281/zenodo.7143778" httpUri="https://zenodo.org/record/7143778/files/figure.png" pageId="13" pageNumber="12">411</figureCitation>
|
||
,
|
||
<figureCitation id="13292A1AFFBCFFAA221FD745FACBFA12" box="[1358,1401,1596,1620]" captionStart="Figures 500–511" captionStartId="103.[113,190,1928,1950]" captionTargetBox="[113,1465,240,1881]" captionTargetId="figure-73@103.[113,1474,234,1881]" captionTargetPageId="103" captionText="Figures 500–511. Teruelius intertidalis, female holotype. Figures 500–508. Pedipalp chela, dorsal (500), external (501) and ventrointernal (502) views; pedipalp patella, dorsal (503), external (504) and ventral (505) views; pedipalp femur and trochanter, internal (506) and dorsal (507) views; pedipalp chela, movable finger dentate margin (508). Figure 509. Carapace and tergites I–III. Figure 510. Sternopectinal region. Figure 511. Telson lateral view." figureDoi="http://doi.org/10.5281/zenodo.7143811" httpUri="https://zenodo.org/record/7143811/files/figure.png" pageId="13" pageNumber="12">510</figureCitation>
|
||
,
|
||
<figureCitation id="13292A1AFFBCFFAA22DDD745FA0AFA12" box="[1420,1464,1596,1620]" captionStart="Figures 526–537" captionStartId="107.[113,190,1928,1950]" captionTargetBox="[113,1458,234,1912]" captionTargetId="figure-80@107.[113,1468,234,1919]" captionTargetPageId="107" captionText="Figures 526–537. Teruelius mahafaliensis, female (526, 529–530, 535), male (527–528, 531–534) and male holotype of G. rossii (536–537). Sternopectinal region (526–527). Carapace and tergites I–II (528). Pedipalp chela, movable finger dentate margin (529). Pedipalp chela dorsal (530–531) and external (532) views. Pedipalp patella dorsal (533) view. Pedipalp femur and trochanter dorsal (534) view. Metasoma V and telson lateral (535) view. Metasoma and telson dorsal (536) and ventral (537) views. Scale bar: 10 mm (536–537)." figureDoi="http://doi.org/10.5281/zenodo.7143819" httpUri="https://zenodo.org/record/7143819/files/figure.png" pageId="13" pageNumber="12">526</figureCitation>
|
||
); hemispermatophore capsule short, carinate, posterior lobe rounded, without lanceolate extension (
|
||
<figureCitation id="13292A1AFFBCFFAA2387D705FAEBFAD2" box="[1238,1369,1660,1684]" captionStart="Figures 71–85" captionStartId="26.[113,190,1701,1723]" captionTargetBox="[149,1434,231,1678]" captionTargetId="figure-287@26.[146,1446,225,1690]" captionTargetPageId="26" captionText="Figures 71–85. Hemispermatophores and capsule regions of Teruelius gen n. Multi-panel figures show: whole hemispermatophore; whole hemispermatophore and/or capsule with flagellum; capsule region in convex (or convex compressed), anterior and posterior views (panels in left to right sequence). Right hemispermatophores. Figure 71. T. ankarana, whole hemispermatophore. Scale bar: 4 mm. Figure 72. T. ankarana, capsule, left mirrored, Forêt d’Ankavanana, FMNH. Scale bar: 500 μm. Figure 73. T. grandidieri, whole hemispermatophore (flagellum truncated). Scale bar: 4 mm. Figure 74. T. grandidieri, capsule, Antsakabe River, FMNH. Scale bar: 500 μm. Figure 75. T. flavopiceus, whole hemispermatophore. Scale bar: 2 mm. Figure 76. T. flavopiceus, capsule, Madagascar, GLPC, FKCP. Scale bar: 500 μm. Figure 77. T. annulatus, capsule, Tsimanampetsotsa National Park, GLPC, FKCP. Scale bar: 500 μm. Figure 78. T. ankarafantsika, capsule and flagellum. Scale bar: 400 μm. Figure 79. T. ankarafantsika, capsule, Forêt d’Ankavanana, FMNH. Scale bar: 200 μm. Figure 80. T. ankarafantsika, capsule, Réserve Forestière de l’Ankarafantsika, FMNH. Scale bar: 200 μm. Figure 81. T. olgae, capsule, Itampolo village, FMNH. Scale bar: 500 μm. Figure 82. T. limbatus, whole hemispermatophore (flagellum truncated). Scale bar: 2 mm. Figure 83. T. limbatus, capsule, Forêt d’Ianasana, FMNH. Scale bar: 500 μm. Figure 84. T. mahafaliensis, capsule views, Zombitse-Vohibasia National Park, GLPC, FKCP. Scale bar: 500 μm. Figure 85. T. intertidalis, capsule, Madagascar, GLPC, FKCP. Scale bar: 500 μm." figureDoi="http://doi.org/10.5281/zenodo.7143641" httpUri="https://zenodo.org/record/7143641/files/figure.png" pageId="13" pageNumber="12">Figs. 71–85</figureCitation>
|
||
); sternites with narrow, slit-like spiracles (
|
||
<figureCitation id="13292A1AFFBCFFAA23C6D7E5FA9EFAF2" box="[1175,1324,1692,1716]" captionStart="Figures 95–110" captionStartId="18.[113,190,1408,1430]" captionTargetBox="[113,1475,255,1894]" captionTargetId="figure-131@18.[113,1474,255,1915]" captionTargetPageId="18" captionText="Figures 95–110: Orthochirus mesopotamicus comb. n., from Iraq, Basra (ca. 30°48’N 47°34’E). Figures 95, 97, 100–110. Male, carapace and tergites I–III (95), sternopectinal region and sternites (97). Pedipalp chela, dorsal (100), external (101), and ventrointernal (102) views. Pedipalp patella, dorsal (103), external (104), and ventral (105) views. Pedipalp femur and trochanter, internodorsal (106) and dorsoexternal (107) views. Pedipalp chela, movable finger dentate margin (108). Distal segments of right legs III–IV, retrolateral views (109–110). Figures 96, 98–99. Female, carapace and tergites I–III (96), sternopectinal region and sternites (98), and pedipalp chela dorsoexternal (99). The trichobothrial pattern is indicated in Figures 101–104, 106–107 (white circles)." pageId="13" pageNumber="12">Figs. 99–105</figureCitation>
|
||
); metasoma I with ventromedian carinae moderately to weakly crenulate or smooth to obsolete (
|
||
<figureCitation id="13292A1AFFBCFFAA2369D7A5FB6BFAB2" box="[1080,1241,1756,1780]" captionStart-0="Figures 122–126" captionStart-1="Figures 127–132" captionStartId-0="38.[113,190,1956,1978]" captionStartId-1="39.[113,190,1954,1976]" captionTargetBox-0="[180,1420,247,1929]" captionTargetBox-1="[132,1462,256,1937]" captionTargetId-0="figure-75@38.[126,1459,234,1942]" captionTargetId-1="figure-68@39.[113,1474,231,1947]" captionTargetPageId-0="38" captionTargetPageId-1="39" captionText-0="Figures 122–126. Ventral aspect of metasoma and telson of adult males (♂) or females (♀) of representative Grosphus spp. and Teruelius limbatus under UV fluorescence to reveal carination and granulation. G. hirtus (122), G. madagascariensis (123 from Anjiro, 124 from Andasibe), G. voahangyae (125), and T. limbatus (126). Scale bars: 4 mm." captionText-1="Figures 127–132. Ventral aspect of metasoma and telson of adult males (♂) or females (♀) of representative Teruelius gen. n under UV fluorescence to reveal carination and granulation. T. ankarafantsika (127), T. ankarana (128), T. flavopiceus (129), T. grandidieri (130), T. mahafaliensis (131), and T. olgae (132). Scale bars: 4 mm." figureDoi-0="http://doi.org/10.5281/zenodo.7143671" figureDoi-1="http://doi.org/10.5281/zenodo.7143675" httpUri-0="https://zenodo.org/record/7143671/files/figure.png" httpUri-1="https://zenodo.org/record/7143675/files/figure.png" pageId="13" pageNumber="12">Figs. 126–132</figureCitation>
|
||
); telson with oval or bulbous vesicle, without subaculear tubercle in adults (
|
||
<figureCitation id="13292A1AFFBCFFAA22DFD785FC26FB72" captionStart="Figures 181–195" captionStartId="64.[113,190,1953,1975]" captionTargetId="figure-71@64.[113,1474,242,1670]" captionTargetPageId="64" captionText="Figures 181–195. Telson of representativeGrosphus andTeruelius gen. n. G. hirtus (181–182), G. madagascariensis (183–184), G. voahangyae (185), T. ankarafantsika (186–187), T. ankarana (188), T. flavopiceus (189), T. grandidieri (190–191), T. limbatus (192), T. mahafaliensis (193) and T. olgae (194–195). UV fluorescence, ♂ male, ♀ female. Scale bars: 2 mm (181–187, 192–195), 4 mm (188–191)." figureDoi="http://doi.org/10.5281/zenodo.7143700" httpUri="https://zenodo.org/record/7143700/files/figure.png" pageId="13" pageNumber="12">Figs. 186–195</figureCitation>
|
||
); legs with ventral surface of telotarsus densely setose or scopulate, with broad, brush-like strips of> 20 long filiform macrosetae (
|
||
<figureCitation id="13292A1AFFBCFFAA2490D625FBD4FB32" box="[961,1126,1884,1908]" captionStart="Figures 133–144" captionStartId="40.[113,190,1293,1315]" captionTargetBox="[118,1474,249,1273]" captionTargetId="figure-523@40.[113,1474,227,1273]" captionTargetPageId="40" captionText="Figures 133–144. Ventral setation of telotarsus III in Grosphus and Teruelius gen. n. Ventral surfaces of right telotarsus III of adult males (♂) or females (♀), shown under UV fluorescence to highlight setation. Macrosetae appear dark with strongly fluorescent cuticular sockets at their base. Putative chemosensory microsetae appear bright. G. hirtus (= G. h. garciai) (133), G. hirtus (134), G. madagascariensis (135, from Anjiro, 136 from Andasibe), G. voahangyae (137), T. ankarafantsika (138), T. limbatus (139), T. mahafaliensis (140), T. ankarana (141), T. grandidieri (142), T. flavopiceus (143), and T. olgae (144). Scale bars: 500 μm (133–140, 144) and 1 mm (141–143)." figureDoi="http://doi.org/10.5281/zenodo.7143677" httpUri="https://zenodo.org/record/7143677/files/figure.png" pageId="13" pageNumber="12">Figs. 138–144</figureCitation>
|
||
,
|
||
<figureCitation id="13292A1AFFBCFFAA2328D625FB68FB32" box="[1145,1242,1884,1908]" captionStart="Figures 211–226" captionStartId="66.[113,190,1920,1942]" captionTargetBox="[117,1466,234,1743]" captionTargetId="figure-88@66.[113,1474,234,1743]" captionTargetPageId="66" captionText="Figures 211–226. Ventral tarsal setation of legs III or IV in Grosphus and Teruelius gen. n. G. goudoti (211), G. hirtus (212), G. hirtus (G. garciai) (213), G. madagascariensis (214), G. madagascariensis (G. mandena) (215), Teruelius ankarana (216), T. flavopiceus (217), T. annulatus (218), T. ankarafantsika (219–220), T. bistriatus (221), T. intertidalis (222), T. grandidieri (223), T. mahafaliensis (224), T. limbatus (225), T. feti (226)." figureDoi="http://doi.org/10.5281/zenodo.7143708" httpUri="https://zenodo.org/record/7143708/files/figure.png" pageId="13" pageNumber="12">216–226</figureCitation>
|
||
,
|
||
<figureCitation id="13292A1AFFBCFFAA23BCD625FAFFFB32" box="[1261,1357,1884,1908]" captionStart="Figures 409–417" captionStartId="89.[113,190,1958,1980]" captionTargetBox="[113,1474,234,1814]" captionTargetId="figure-58@89.[113,1474,234,1838]" captionTargetPageId="89" captionText="Figures 409–417. Teruelius ankarafantsika. Male paratype (409, 416–417) and female holotype (410–415). Figures 409–411. Carapace and tergites I–III (409–410) and sternopectinal region (411). Figures 412–417. Left legs tibia, basitarsus and telotarsus, retrolateral views. Leg I (412), leg II (413), leg III (414, 416), leg IV (415, 417)." figureDoi="http://doi.org/10.5281/zenodo.7143778" httpUri="https://zenodo.org/record/7143778/files/figure.png" pageId="13" pageNumber="12">409–417</figureCitation>
|
||
,
|
||
<figureCitation id="13292A1AFFBCFFAA2231D625FA72FB32" box="[1376,1472,1884,1908]" captionStart="Figures 477–490" captionStartId="100.[113,190,1928,1950]" captionTargetBox="[110,1468,256,1846]" captionTargetId="figure-92@100.[110,1471,244,1880]" captionTargetPageId="100" captionText="Figures 477–490. Teruelius flavopiceus, male. Figures 477–486. Pedipalp chela, dorsal (477), external (478) and ventrointernal (479) views; pedipalp patella, dorsal (480), external (481) and ventral (482) views; pedipalp femur and trochanter, internal (483) and dorsal (484) views; pedipalp chela, movable (485) and fixed (486) finger dentate margin. The trichobothrial pattern is indicated by white circles in Figures 477a–484a. Figures 487–490. Right legs tibia, basitarsus and telotarsus, retrolateral views. Leg IV (487), leg III (488), leg. II (489), leg I (490)." figureDoi="http://doi.org/10.5281/zenodo.7143803" httpUri="https://zenodo.org/record/7143803/files/figure.png" pageId="13" pageNumber="12">487–490</figureCitation>
|
||
,
|
||
<figureCitation id="13292A1AFFBCFFAA2463D605FC20FBD2" box="[818,914,1916,1940]" captionStart="Figures 512–515" captionStartId="104.[117,194,1981,2003]" captionTargetBox="[146,1452,271,1937]" captionTargetId="figure-34@104.[135,1459,234,1949]" captionTargetPageId="104" captionText="Figures 512–515. Teruelius intertidalis, female holotype. Left legs tibia, basitarsus and telotarsus, retrolateral views. Leg I (512), leg II (513), leg III (514), leg IV (515)." figureDoi="http://doi.org/10.5281/zenodo.7143813" httpUri="https://zenodo.org/record/7143813/files/figure.png" pageId="13" pageNumber="12">512–515</figureCitation>
|
||
,
|
||
<figureCitation id="13292A1AFFBCFFAA24CFD605FC4CFBD2" box="[926,1022,1916,1940]" captionStart="Figures 538–541" captionStartId="108.[113,190,1981,2003]" captionTargetBox="[129,1451,241,1964]" captionTargetId="figure-37@108.[123,1459,234,1977]" captionTargetPageId="108" captionText="Figures 538–541. Teruelius mahafaliensis, male holotype of G. rossii. Left legs tibia, basitarsus and telotarsus, retrolateral views. Leg I (538), leg II (539), leg. III (540), leg IV (541)." figureDoi="http://doi.org/10.5281/zenodo.7143821" httpUri="https://zenodo.org/record/7143821/files/figure.png" pageId="13" pageNumber="12">538–541</figureCitation>
|
||
,
|
||
<figureCitation id="13292A1AFFBCFFAA2358D605FBDEFBD2" box="[1033,1132,1916,1940]" captionStart="Figures 576–579" captionStartId="113.[113,190,1981,2003]" captionTargetBox="[122,1462,264,1906]" captionTargetId="figure-33@113.[110,1474,245,1928]" captionTargetPageId="113" captionText="Figures 576–579. Teruelius olgae, male paratype. Right legs tibia, basitarsus and telotarsus, retrolateral and ventral views. Leg I (576), leg II (577), leg III (578), leg IV (579)." figureDoi="http://doi.org/10.5281/zenodo.7143831" httpUri="https://zenodo.org/record/7143831/files/figure.png" pageId="13" pageNumber="12">576–579</figureCitation>
|
||
); telotarsus with dorsal terminal process of normal size; cuticle with strong UV fluorescence (
|
||
<figureCitation id="13292A1AFFBCFFAA246BD6C5FC6AFB92" box="[826,984,1980,2004]" captionStart="Figures 145–157" captionStartId="41.[113,190,1502,1524]" captionTargetBox="[113,1474,234,1490]" captionTargetId="figure-392@41.[113,1474,234,1490]" captionTargetPageId="41" captionText="Figures 145–157. Comparative intensity of UV fluorescence in Grosphus and Teruelius gen. n. Photographic comparison of fluorescence emission intensities of representative species of each genus, including adult males (♂) or females (♀). G. madagascariensis (145 from Anjiro, 148 from Andasibe), G. hirtus (146), G. voahangyae (147), G. hirtus (= G. h. garciai) (149), T. ankarafantsika (150), T. grandidieri (151), T. olgae (152), T. flavopiceus (153), T. ankarana (154–155), T. mahafaliensis (156), and T. limbatus (157). Images acquired under identical intensities of UV excitation (395 nm LED source, 0.35 A current) and camera exposure (Canon EOS 7D Mark II, 100 mm f/13, 0.5 s, ISO 320), with 475 nm longpass filter to block excitation wavelengths. Scale bars: 10 mm (145–150, 152–153, 156–157), 20 mm (151, 154–155)." figureDoi="http://doi.org/10.5281/zenodo.7143681" httpUri="https://zenodo.org/record/7143681/files/figure.png" pageId="13" pageNumber="12">Figs. 150–157</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="DF6D6617FFBFFFA92720D63AFD89F440" ID-DOI="http://doi.org/10.5281/zenodo.7143620" ID-Zenodo-Dep="7143620" httpUri="https://zenodo.org/record/7143620/files/figure.png" pageId="14" pageNumber="13" startId="14.[113,188,1859,1881]" targetBox="[114,1474,227,1840]" targetPageId="14">
|
||
<paragraph id="8BAD369FFFBFFFA92720D63AFD89F440" blockId="14.[113,1475,1859,2054]" pageId="14" pageNumber="13">
|
||
<emphasis id="B966EA8DFFBFFFA92720D63AFEB3FB1E" bold="true" box="[113,257,1859,1881]" pageId="14" pageNumber="13">Figures 21–27</emphasis>
|
||
. Positions of trichobothria
|
||
<emphasis id="B966EA8DFFBFFFA9255DD63AFDE7FB1E" box="[524,597,1859,1881]" italics="true" pageId="14" pageNumber="13">Eb, Eb</emphasis>
|
||
and petite
|
||
<emphasis id="B966EA8DFFBFFFA92598D63AFD51FB1E" box="[713,739,1859,1880]" italics="true" pageId="14" pageNumber="13">Eb</emphasis>
|
||
on manus of pedipalp chela in
|
||
<taxonomicName id="4C124D1CFFBFFFA92370D63AFBCEFB1E" authorityName="Vachon" authorityYear="1940" box="[1057,1148,1859,1880]" class="Arachnida" family="Buthidae" genus="Grosphus" kingdom="Animalia" order="Scorpiones" pageId="14" pageNumber="13" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B966EA8DFFBFFFA92370D63AFBCEFB1E" box="[1057,1148,1859,1880]" italics="true" pageId="14" pageNumber="13">Grosphus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C124D1CFFBFFFA923DAD63AFAB4FB1E" authorityName="Lourenco" authorityYear="1995" box="[1163,1286,1859,1880]" class="Arachnida" family="Buthidae" genus="Neogrosphus" kingdom="Animalia" order="Scorpiones" pageId="14" pageNumber="13" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B966EA8DFFBFFFA923DAD63AFAB4FB1E" box="[1163,1286,1859,1880]" italics="true" pageId="14" pageNumber="13">Neogrosphus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C124D1CFFBFFFA92268D63AFA22FB1E" authorityName="Lowe & Kovařík" authorityYear="2019" box="[1337,1424,1859,1880]" class="Arachnida" family="Buthidae" genus="Teruelius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="14" pageNumber="13" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B966EA8DFFBFFFA92268D63AFA22FB1E" box="[1337,1424,1859,1880]" italics="true" pageId="14" pageNumber="13">Teruelius</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B966EA8DFFBFFFA922C8D63AFA0EFB1E" bold="true" box="[1433,1468,1859,1880]" pageId="14" pageNumber="13">gen</emphasis>
|
||
. 1 2 3
|
||
<emphasis id="B966EA8DFFBFFFA92720D619FEAFFB33" bold="true" box="[113,285,1888,1910]" pageId="14" pageNumber="13">n. Figures 21–24</emphasis>
|
||
. External views of pedipalp chela, shown under UV fluorescence to highlight trichobothrial areolae.
|
||
<taxonomicName id="4C124D1CFFBFFFA923AAD619FAE0FB33" baseAuthorityName="Lourenco & Wilme" baseAuthorityYear="2016" box="[1275,1362,1888,1909]" class="Arachnida" family="Buthidae" genus="Grosphus" kingdom="Animalia" order="Scorpiones" pageId="14" pageNumber="13" phylum="Arthropoda" rank="species" species="hirtus">
|
||
<emphasis id="B966EA8DFFBFFFA923AAD619FABEFB33" box="[1275,1292,1888,1909]" italics="true" pageId="14" pageNumber="13">G</emphasis>
|
||
.
|
||
<emphasis id="B966EA8DFFBFFFA9224AD619FAE0FB33" box="[1307,1362,1888,1909]" italics="true" pageId="14" pageNumber="13">hirtus</emphasis>
|
||
</taxonomicName>
|
||
♂ (21),
|
||
<taxonomicName id="4C124D1CFFBFFFA922FAD619FEA9FBD4" class="Arachnida" family="Buthidae" genus="Grosphus" kingdom="Animalia" order="Scorpiones" pageId="14" pageNumber="13" phylum="Arthropoda" rank="species" species="madagascariensis">
|
||
<emphasis id="B966EA8DFFBFFFA922FAD619FA0EFB33" box="[1451,1468,1888,1909]" italics="true" pageId="14" pageNumber="13">G</emphasis>
|
||
.
|
||
<emphasis id="B966EA8DFFBFFFA92720D604FEA9FBD4" box="[113,283,1917,1938]" italics="true" pageId="14" pageNumber="13">madagascariensis</emphasis>
|
||
</taxonomicName>
|
||
♀ (22),
|
||
<taxonomicName id="4C124D1CFFBFFFA92621D604FE6EFBD4" authorityName=", Pocock" authorityYear="1889" box="[368,476,1917,1938]" class="Arachnida" family="Buthidae" genus="Buthus" kingdom="Animalia" order="Scorpiones" pageId="14" pageNumber="13" phylum="Arthropoda" rank="species" species="limbatus">
|
||
<emphasis id="B966EA8DFFBFFFA92621D604FECFFBD4" box="[368,381,1917,1938]" italics="true" pageId="14" pageNumber="13">T</emphasis>
|
||
.
|
||
<emphasis id="B966EA8DFFBFFFA926DAD604FE6EFBD4" box="[395,476,1917,1938]" italics="true" pageId="14" pageNumber="13">limbatus</emphasis>
|
||
</taxonomicName>
|
||
♂ (23),
|
||
<taxonomicName id="4C124D1CFFBFFFA92563D604FD68FBD4" authorityName="Lowe & Kovařík" authorityYear="2019" baseAuthorityName="Lourenco" baseAuthorityYear="2003" box="[562,730,1917,1938]" class="Arachnida" family="Buthidae" genus="Teruelius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="14" pageNumber="13" phylum="Arthropoda" rank="species" species="ankarafantsika">
|
||
<emphasis id="B966EA8DFFBFFFA92563D604FD8DFBD4" box="[562,575,1917,1938]" italics="true" pageId="14" pageNumber="13">T</emphasis>
|
||
.
|
||
<emphasis id="B966EA8DFFBFFFA9251CD604FD68FBD4" box="[589,730,1917,1938]" italics="true" pageId="14" pageNumber="13">ankarafantsika</emphasis>
|
||
</taxonomicName>
|
||
♀ (24). Positions of
|
||
<emphasis id="B966EA8DFFBFFFA924F9D604FC70FBD4" box="[936,962,1917,1938]" italics="true" pageId="14" pageNumber="13">Eb</emphasis>
|
||
<subScript id="179634DAFFBFFFA92493D6F5FC7BFBDE" attach="left" box="[962,969,1932,1944]" fontSize="5" pageId="14" pageNumber="13">1</subScript>
|
||
,
|
||
<emphasis id="B966EA8DFFBFFFA92486D604FC43FBD4" box="[983,1009,1917,1938]" italics="true" pageId="14" pageNumber="13">Eb</emphasis>
|
||
<subScript id="179634DAFFBFFFA924A0D6F5FC4AFBDE" attach="left" box="[1009,1016,1932,1944]" fontSize="5" pageId="14" pageNumber="13">2</subScript>
|
||
and
|
||
<emphasis id="B966EA8DFFBFFFA92377D604FBF2FBD4" box="[1062,1088,1917,1938]" italics="true" pageId="14" pageNumber="13">Eb</emphasis>
|
||
<subScript id="179634DAFFBFFFA92311D6F5FBF5FBDE" attach="left" box="[1088,1095,1932,1944]" fontSize="5" pageId="14" pageNumber="13">3</subScript>
|
||
and lines joining them shown as white overlays. Scale bars: 2 mm.
|
||
<emphasis id="B966EA8DFFBFFFA9262DD6E0FDBBFBE8" bold="true" box="[380,521,1945,1967]" pageId="14" pageNumber="13">Figures 25–26</emphasis>
|
||
. Horizontal histograms comparing ratios,
|
||
<emphasis id="B966EA8DFFBFFFA924C7D6E3FC17FBE9" box="[918,933,1946,1967]" italics="true" pageId="14" pageNumber="13">R</emphasis>
|
||
= d(
|
||
<emphasis id="B966EA8DFFBFFFA924B6D6E3FBB3FBE9" box="[999,1025,1946,1967]" italics="true" pageId="14" pageNumber="13">Eb</emphasis>
|
||
,
|
||
<emphasis id="B966EA8DFFBFFFA92345D6E3FB9CFBE9" box="[1044,1070,1946,1967]" italics="true" pageId="14" pageNumber="13">Eb</emphasis>
|
||
)/ d(
|
||
<emphasis id="B966EA8DFFBFFFA9230DD6E3FBC4FBE9" box="[1116,1142,1946,1967]" italics="true" pageId="14" pageNumber="13">Eb</emphasis>
|
||
,
|
||
<emphasis id="B966EA8DFFBFFFA923D8D6E3FB11FBE9" box="[1161,1187,1946,1967]" italics="true" pageId="14" pageNumber="13">Eb</emphasis>
|
||
), of
|
||
<emphasis id="B966EA8DFFBFFFA92389D6E3FAA9FBE9" box="[1240,1307,1946,1967]" italics="true" pageId="14" pageNumber="13">Eb -Eb</emphasis>
|
||
distance, to
|
||
<emphasis id="B966EA8DFFBFFFA922C8D6E3FA01FBE9" box="[1433,1459,1946,1967]" italics="true" pageId="14" pageNumber="13">Eb</emphasis>
|
||
-
|
||
<subScript id="179634DAFFBFFFA924F4D6D0FB87FBF3" attach="left" box="[933,1077,1961,1973]" fontSize="5" pageId="14" pageNumber="13">123 2 3</subScript>
|
||
<subScript id="179634DAFFBFFFA92326D6D0FB19FBF3" attach="right" box="[1143,1195,1961,1973]" fontSize="5" pageId="14" pageNumber="13">1 2</subScript>
|
||
<subScript id="179634DAFFBFFFA923A3D6D0FA08FBF3" attach="left" box="[1266,1466,1961,1973]" fontSize="5" pageId="14" pageNumber="13">2 3 1</subScript>
|
||
<emphasis id="B966EA8DFFBFFFA92720D6CFFF39FB8D" box="[113,139,1974,1995]" italics="true" pageId="14" pageNumber="13">Eb</emphasis>
|
||
distance, in males (25), and females (26) of
|
||
<taxonomicName id="4C124D1CFFBFFFA92569D6CFFD21FB8D" authorityName="Vachon" authorityYear="1940" box="[568,659,1974,1995]" class="Arachnida" family="Buthidae" genus="Grosphus" kingdom="Animalia" order="Scorpiones" pageId="14" pageNumber="13" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B966EA8DFFBFFFA92569D6CFFD21FB8D" box="[568,659,1974,1995]" italics="true" pageId="14" pageNumber="13">Grosphus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C124D1CFFBFFFA925CFD6CFFCABFB8D" authorityName="Lourenco" authorityYear="1995" box="[670,793,1974,1995]" class="Arachnida" family="Buthidae" genus="Neogrosphus" kingdom="Animalia" order="Scorpiones" pageId="14" pageNumber="13" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B966EA8DFFBFFFA925CFD6CFFCABFB8D" box="[670,793,1974,1995]" italics="true" pageId="14" pageNumber="13">Neogrosphus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C124D1CFFBFFFA92419D6CFFC2DFB8D" authority="Lowe & Kovařík, 2019" authorityName="Lowe & Kovařík" authorityYear="2019" box="[840,927,1974,1995]" class="Arachnida" family="Buthidae" genus="Teruelius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="14" pageNumber="13" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="B966EA8DFFBFFFA92419D6CFFC2DFB8D" box="[840,927,1974,1995]" italics="true" pageId="14" pageNumber="13">Teruelius</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A25557F6FFBFFFA924F5D6CEFC55FB8A" box="[932,999,1975,1996]" pageId="14" pageNumber="13" rank="genus">
|
||
<emphasis id="B966EA8DFFBFFFA924F5D6CEFC75FB8A" bold="true" box="[932,967,1975,1996]" pageId="14" pageNumber="13">gen</emphasis>
|
||
.
|
||
<emphasis id="B966EA8DFFBFFFA92485D6CEFC53FB8A" bold="true" box="[980,993,1975,1996]" pageId="14" pageNumber="13">n</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
Error bars indicate ranges (minimum, maximum), 2 histogram bars mid-range values.
|
||
<emphasis id="B966EA8DFFBFFFA926E2D6AAFDA6FBAE" bold="true" box="[435,532,2003,2025]" pageId="14" pageNumber="13">Figure 27</emphasis>
|
||
. Scatter plot of male vs. female ratios
|
||
<emphasis id="B966EA8DFFBFFFA924D2D6ADFC20FBAF" box="[899,914,2004,2025]" italics="true" pageId="14" pageNumber="13">R</emphasis>
|
||
. Ratio is larger in males if points fall above the diagonal 123 (gray) line, larger in females if they fall below it.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C3086514FFBEFFA82722D192FDB7F864" pageId="15" pageNumber="14" type="discussion">
|
||
<paragraph id="8BAD369FFFBEFFA82722D192FEF5FD45" blockId="15.[115,759,235,963]" box="[115,327,235,259]" pageId="15" pageNumber="14">SUBORDINATE TAXA.</paragraph>
|
||
<paragraph id="8BAD369FFFBEFFA82722D072FD8EFD24" blockId="15.[115,759,235,963]" pageId="15" pageNumber="14">
|
||
<taxonomicName id="4C124D1CFFBEFFA82722D072FD8FFD65" authority="(Lourenco, 2003)" authorityName="Lowe & Kovařík" authorityYear="2019" baseAuthorityName="Lourenco" baseAuthorityYear="2003" box="[115,573,267,291]" class="Arachnida" family="Buthidae" genus="Teruelius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="15" pageNumber="14" phylum="Arthropoda" rank="species" species="ankarafantsika" status="comb. nov.">
|
||
<emphasis id="B966EA8DFFBEFFA82722D072FECEFD65" box="[115,380,267,291]" italics="true" pageId="15" pageNumber="14">Teruelius ankarafantsika</emphasis>
|
||
(
|
||
<bibRefCitation id="EF834B6EFFBEFFA826DBD072FD84FD65" author="LOURENCO, W. R." box="[394,566,267,291]" editor="GOODMAN, S. & BENSTEAD, J." journalOrPublisher="The University of Chicago Press, Chicago, Illinois" pageId="15" pageNumber="14" pagination="575 - 579" refId="ref38188" refString="LOURENCO, W. R. 2003 a. Scorpiones, scorpions. Pp. 575 - 579 in GOODMAN, S., BENSTEAD, J. (Eds.). The Natural History of Madagascar. The University of Chicago Press, Chicago, Illinois." title="Scorpiones, scorpions." type="book chapter" volumeTitle="The Natural History of Madagascar" year="2003">Lourenço, 2003</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A25557F6FFBEFFA82515D072FD14FD64" box="[580,678,267,291]" pageId="15" pageNumber="14" rank="species">
|
||
<emphasis id="B966EA8DFFBEFFA82515D072FD30FD65" bold="true" box="[580,642,267,291]" pageId="15" pageNumber="14">comb</emphasis>
|
||
.
|
||
<emphasis id="B966EA8DFFBEFFA825C1D072FD2DFD65" bold="true" box="[656,671,267,291]" pageId="15" pageNumber="14">n</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
<taxonomicName id="4C124D1CFFBEFFA82722D052FD3DFD05" authority="(Lourenco & Goodman, 2003)" authorityName="Lowe & Kovařík" authorityYear="2019" baseAuthorityName="Lourenco & Goodman" baseAuthorityYear="2003" box="[115,655,299,323]" class="Arachnida" family="Buthidae" genus="Teruelius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="15" pageNumber="14" phylum="Arthropoda" rank="species" species="ankarana" status="comb. nov.">
|
||
<emphasis id="B966EA8DFFBEFFA82722D052FEF0FD05" box="[115,322,299,323]" italics="true" pageId="15" pageNumber="14">Teruelius ankarana</emphasis>
|
||
(
|
||
<bibRefCitation id="EF834B6EFFBEFFA82601D052FD3AFD05" author="LOURENCO, W. R. & S. M. GOODMAN" box="[336,648,299,323]" journalOrPublisher="Revista Iberica de Aracnologia" pageId="15" pageNumber="14" pagination="19 - 28" part="7" refId="ref39272" refString="LOURENCO, W. R. & S. M. GOODMAN. 2003 a. Description of a new species of Grosphus Simon (Scorpiones, Buthidae) from the Ankarana Massif, Madagascar. Revista Iberica de Aracnologia, 7: 19 - 28." title="Description of a new species of Grosphus Simon (Scorpiones, Buthidae) from the Ankarana Massif, Madagascar" type="journal article" year="2003">Lourenço & Goodman, 2003</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A25557F6FFBEFFA825C7D052FD45FD04" box="[662,759,299,323]" pageId="15" pageNumber="14" rank="species">
|
||
<emphasis id="B966EA8DFFBEFFA825C7D052FD66FD05" bold="true" box="[662,724,299,323]" pageId="15" pageNumber="14">comb</emphasis>
|
||
.
|
||
<emphasis id="B966EA8DFFBEFFA825B0D052FD42FD05" bold="true" box="[737,752,299,323]" pageId="15" pageNumber="14">n</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
<taxonomicName id="4C124D1CFFBEFFA82722D032FE67FD25" authority="(Fage, 1929)" authorityName="Lowe & Kovařík" authorityYear="2019" baseAuthorityName="Fage" baseAuthorityYear="1929" box="[115,469,331,355]" class="Arachnida" family="Buthidae" genus="Teruelius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="15" pageNumber="14" phylum="Arthropoda" rank="species" species="annulatus" status="comb. nov.">
|
||
<emphasis id="B966EA8DFFBEFFA82722D032FEF7FD25" box="[115,325,331,355]" italics="true" pageId="15" pageNumber="14">Teruelius annulatus</emphasis>
|
||
(
|
||
<bibRefCitation id="EF834B6EFFBEFFA82605D032FE7FFD25" author="FAGE, L." box="[340,461,331,355]" journalOrPublisher="Societe d'Editions Geographiques Maritimes et Coloniales, Paris" pageId="15" pageNumber="14" pagination="637 - 693" refId="ref33536" refString="FAGE, L. 1929. Les scorpions de Madagascar, leur affinites, leur distribution geographique. Pp. 637 - 693 in: Faune des Colonies Francaises. Societe d'Editions Geographiques Maritimes et Coloniales, Paris." title="Les scorpions de Madagascar, leur affinites, leur distribution geographique" type="book chapter" volumeTitle="Faune des Colonies Francaises" year="1929">Fage, 1929</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A25557F6FFBEFFA8268AD032FD8EFD24" box="[475,572,331,355]" pageId="15" pageNumber="14" rank="species">
|
||
<emphasis id="B966EA8DFFBEFFA8268AD032FDABFD25" bold="true" box="[475,537,331,355]" pageId="15" pageNumber="14">comb</emphasis>
|
||
.
|
||
<emphasis id="B966EA8DFFBEFFA82576D032FD84FD25" bold="true" box="[551,566,331,355]" pageId="15" pageNumber="14">n</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
</paragraph>
|
||
<paragraph id="8BAD369FFFBEFFA82722D012FF67FDE4" blockId="15.[115,759,235,963]" pageId="15" pageNumber="14">
|
||
<taxonomicName id="4C124D1CFFBEFFA82722D012FD7DFDC5" authority="(Lourenco, Wilme & Waeber, 2018)" authorityName="Lowe & Kovařík" authorityYear="2019" baseAuthorityName="Lourenco, Wilme & Waeber" baseAuthorityYear="2018" box="[115,719,363,387]" class="Arachnida" family="Buthidae" genus="Teruelius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="15" pageNumber="14" phylum="Arthropoda" rank="species" species="bemaraha" status="comb. nov.">
|
||
<emphasis id="B966EA8DFFBEFFA82722D012FEFAFDC5" box="[115,328,363,387]" italics="true" pageId="15" pageNumber="14">Teruelius bemaraha</emphasis>
|
||
(
|
||
<bibRefCitation id="EF834B6EFFBEFFA82607D012FD7AFDC5" author="LOURENCO, W. R. & L. WILME & P. O. WAEBER" box="[342,712,363,387]" journalOrPublisher="Comptes Rendus Biologies" pageId="15" pageNumber="14" pagination="131 - 143" part="341" refId="ref40549" refString="LOURENCO, W. R., L. WILME & P. O. WAEBER. 2018 a. The genus Opisthacanthus Peters, 1861 (Scorpiones: Hormuridae), a remarkable Gondwanian group of scorpions. Comptes Rendus Biologies, 341: 131 - 143." title="The genus Opisthacanthus Peters, 1861 (Scorpiones: Hormuridae), a remarkable Gondwanian group of scorpions" type="journal article" year="2018">Lourenço, Wilmé & Waeber, 2018</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A25557F6FFBEFFA82722D0F2FF67FDE4" box="[115,213,395,419]" pageId="15" pageNumber="14" rank="species">
|
||
<emphasis id="B966EA8DFFBEFFA82722D0F2FF03FDE5" bold="true" box="[115,177,395,419]" pageId="15" pageNumber="14">comb</emphasis>
|
||
.
|
||
<emphasis id="B966EA8DFFBEFFA827EED0F2FF7CFDE5" bold="true" box="[191,206,395,419]" pageId="15" pageNumber="14">n</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
</paragraph>
|
||
<paragraph id="8BAD369FFFBEFFA82722D0D2FDE1FD84" blockId="15.[115,759,235,963]" box="[115,595,427,451]" pageId="15" pageNumber="14">
|
||
<taxonomicName id="4C124D1CFFBEFFA82722D0D2FE58FD85" authority="(Lourenco, 2012)" authorityName="Lowe & Kovařík" authorityYear="2019" baseAuthorityName="Lourenco" baseAuthorityYear="2012" box="[115,490,427,451]" class="Arachnida" family="Buthidae" genus="Teruelius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="15" pageNumber="14" phylum="Arthropoda" rank="species" species="bicolor" status="comb. nov.">
|
||
<emphasis id="B966EA8DFFBEFFA82722D0D2FE9BFD85" box="[115,297,427,451]" italics="true" pageId="15" pageNumber="14">Teruelius bicolor</emphasis>
|
||
(
|
||
<bibRefCitation id="EF834B6EFFBEFFA82666D0D2FE51FD85" author="LOURENCO, W. R." box="[311,483,427,451]" journalOrPublisher="Boletin de la Sociedad Entomologica Aragonesa" pageId="15" pageNumber="14" pagination="187 - 192" part="50" refId="ref38648" refString="LOURENCO, W. R. 2012 a. Further taxonomic considerations on the genus Buthoscorpio W erner, 1936 (Scorpiones, Buthidae), with description of a new species from India. Boletin de la Sociedad Entomologica Aragonesa, 50: 187 - 192." title="Further taxonomic considerations on the genus Buthoscorpio W erner, 1936 (Scorpiones, Buthidae), with description of a new species from India" type="journal article" year="2012">Lourenço, 2012</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A25557F6FFBEFFA826A0D0D2FDE1FD84" box="[497,595,427,451]" pageId="15" pageNumber="14" rank="species">
|
||
<emphasis id="B966EA8DFFBEFFA826A0D0D2FD9DFD85" bold="true" box="[497,559,427,451]" pageId="15" pageNumber="14">comb</emphasis>
|
||
.
|
||
<emphasis id="B966EA8DFFBEFFA8256CD0D2FDFEFD85" bold="true" box="[573,588,427,451]" pageId="15" pageNumber="14">n</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
</paragraph>
|
||
<paragraph id="8BAD369FFFBEFFA82722D0B2FDDFFDA4" blockId="15.[115,759,235,963]" box="[115,621,459,483]" pageId="15" pageNumber="14">
|
||
<taxonomicName id="4C124D1CFFBEFFA82722D0B2FDB4FDA5" authority="(Kraepelin, 1900)" authorityName="Lowe & Kovařík" authorityYear="2019" baseAuthorityName="Kraepelin" baseAuthorityYear="1900" box="[115,518,459,483]" class="Arachnida" family="Buthidae" genus="Teruelius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="15" pageNumber="14" phylum="Arthropoda" rank="species" species="bistriatus" status="comb. nov.">
|
||
<emphasis id="B966EA8DFFBEFFA82722D0B2FEF3FDA5" box="[115,321,459,483]" italics="true" pageId="15" pageNumber="14">Teruelius bistriatus</emphasis>
|
||
(
|
||
<bibRefCitation id="EF834B6EFFBEFFA8261ED0B2FE4CFDA5" author="KRAEPELIN, K." box="[335,510,459,483]" journalOrPublisher="Abhandlungen aus dem Gebiete der Naturwissenschaften. Herausgegeben vom Naturwissenschaftlichen Verein in Hamburg" pageId="15" pageNumber="14" pagination="1 - 17" part="16" refId="ref36651" refString="KRAEPELIN, K. 1900. Ueber einige neue Gliederspinnen. Abhandlungen aus dem Gebiete der Naturwissenschaften. Herausgegeben vom Naturwissenschaftlichen Verein in Hamburg, 16, 1 (4): 1 - 17." title="Ueber einige neue Gliederspinnen" type="journal article" year="1900">Kraepelin, 1900</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A25557F6FFBEFFA8255DD0B2FDDFFDA4" box="[524,621,459,483]" pageId="15" pageNumber="14" rank="species">
|
||
<emphasis id="B966EA8DFFBEFFA8255DD0B2FDF8FDA5" bold="true" box="[524,586,459,483]" pageId="15" pageNumber="14">comb</emphasis>
|
||
.
|
||
<emphasis id="B966EA8DFFBEFFA82506D0B2FDD4FDA5" bold="true" box="[599,614,459,483]" pageId="15" pageNumber="14">n</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
</paragraph>
|
||
<paragraph id="8BAD369FFFBEFFA82722D092FD9AFE64" blockId="15.[115,759,235,963]" pageId="15" pageNumber="14">
|
||
<taxonomicName id="4C124D1CFFBEFFA82722D092FD36FE45" authority="(Lourenco & Wilme, 2016)" authorityName="Lowe & Kovařík" authorityYear="2019" baseAuthorityName="Lourenco & Wilme" baseAuthorityYear="2016" box="[115,644,491,515]" class="Arachnida" family="Buthidae" genus="Teruelius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="15" pageNumber="14" phylum="Arthropoda" rank="species" species="eliseanneae" status="comb. nov.">
|
||
<emphasis id="B966EA8DFFBEFFA82722D092FEE8FE45" box="[115,346,491,515]" italics="true" pageId="15" pageNumber="14">Teruelius eliseanneae</emphasis>
|
||
(
|
||
<bibRefCitation id="EF834B6EFFBEFFA82639D092FDCFFE45" author="LOURENCO, W. R. & L. WILME" box="[360,637,491,515]" journalOrPublisher="Madagascar Conservation & Development" pageId="15" pageNumber="14" pagination="1 - 14" part="11" refId="ref40275" refString="LOURENCO, W. R. & L. WILME. 2016. Three new species of Grosphus Simon 1880, (Scorpiones: Buthidae) from Madagascar; possible vicariant cases within the Grosphus bistriatus group of species. Madagascar Conservation & Development, 11 (2): 1 - 14." title="Three new species of Grosphus Simon 1880, (Scorpiones: Buthidae) from Madagascar; possible vicariant cases within the Grosphus bistriatus group of species" type="journal article" year="2016">Lourenço & Wilmé, 2016</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A25557F6FFBEFFA825DAD092FD5FFE44" box="[651,749,491,515]" pageId="15" pageNumber="14" rank="species">
|
||
<emphasis id="B966EA8DFFBEFFA825DAD092FD7BFE45" bold="true" box="[651,713,491,515]" pageId="15" pageNumber="14">comb</emphasis>
|
||
.
|
||
<emphasis id="B966EA8DFFBEFFA82586D092FD54FE45" bold="true" box="[727,742,491,515]" pageId="15" pageNumber="14">n</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
<taxonomicName id="4C124D1CFFBEFFA82722D372FE72FE65" authority="(Lourenco, 1996)" authorityName="Lowe & Kovařík" authorityYear="2019" baseAuthorityName="Lourenco" baseAuthorityYear="1996" box="[115,448,523,547]" class="Arachnida" family="Buthidae" genus="Teruelius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="15" pageNumber="14" phylum="Arthropoda" rank="species" species="feti" status="comb. nov.">
|
||
<emphasis id="B966EA8DFFBEFFA82722D372FF4CFE65" box="[115,254,523,547]" italics="true" pageId="15" pageNumber="14">Teruelius feti</emphasis>
|
||
(
|
||
<bibRefCitation id="EF834B6EFFBEFFA8265DD372FE0AFE65" author="LOURENCO, W. R." box="[268,440,523,547]" journalOrPublisher="Biogeographie de Madagascar. ORSTOM, Paris" pageId="15" pageNumber="14" pagination="441 - 455" refId="ref37587" refString="LOURENCO, W. R. 1996 a. Origins and affinities of the scorpion fauna of Madagascar. Pp. 441 - 455 in: Lourenco, W. R. (Ed.). Biogeographie de Madagascar. ORSTOM, Paris." title="Origins and affinities of the scorpion fauna of Madagascar" type="book chapter" volumeTitle="Lourenco, W. R" year="1996">Lourenço, 1996</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A25557F6FFBEFFA82697D372FD9AFE64" box="[454,552,523,547]" pageId="15" pageNumber="14" rank="species">
|
||
<emphasis id="B966EA8DFFBEFFA82697D372FDB6FE65" bold="true" box="[454,516,523,547]" pageId="15" pageNumber="14">comb</emphasis>
|
||
.
|
||
<emphasis id="B966EA8DFFBEFFA82543D372FD93FE65" bold="true" box="[530,545,523,547]" pageId="15" pageNumber="14">n</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
</paragraph>
|
||
<paragraph id="8BAD369FFFBEFFA82722D352FD32FE04" blockId="15.[115,759,235,963]" box="[115,640,555,579]" pageId="15" pageNumber="14">
|
||
<taxonomicName id="4C124D1CFFBEFFA82722D352FDABFE05" authority="(Kraepelin, 1900)" authorityName="Lowe & Kovařík" authorityYear="2019" baseAuthorityName="Kraepelin" baseAuthorityYear="1900" box="[115,537,555,579]" class="Arachnida" family="Buthidae" genus="Teruelius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="15" pageNumber="14" phylum="Arthropoda" rank="species" species="flavopiceus" status="comb. nov.">
|
||
<emphasis id="B966EA8DFFBEFFA82722D352FEE6FE05" box="[115,340,555,579]" italics="true" pageId="15" pageNumber="14">Teruelius flavopiceus</emphasis>
|
||
(
|
||
<bibRefCitation id="EF834B6EFFBEFFA82633D352FDA3FE05" author="KRAEPELIN, K." box="[354,529,555,579]" journalOrPublisher="Abhandlungen aus dem Gebiete der Naturwissenschaften. Herausgegeben vom Naturwissenschaftlichen Verein in Hamburg" pageId="15" pageNumber="14" pagination="1 - 17" part="16" refId="ref36651" refString="KRAEPELIN, K. 1900. Ueber einige neue Gliederspinnen. Abhandlungen aus dem Gebiete der Naturwissenschaften. Herausgegeben vom Naturwissenschaftlichen Verein in Hamburg, 16, 1 (4): 1 - 17." title="Ueber einige neue Gliederspinnen" type="journal article" year="1900">Kraepelin, 1900</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A25557F6FFBEFFA8254ED352FD32FE04" box="[543,640,555,579]" pageId="15" pageNumber="14" rank="species">
|
||
<emphasis id="B966EA8DFFBEFFA8254ED352FDEFFE05" bold="true" box="[543,605,555,579]" pageId="15" pageNumber="14">comb</emphasis>
|
||
.
|
||
<emphasis id="B966EA8DFFBEFFA8253AD352FDC8FE05" bold="true" box="[619,634,555,579]" pageId="15" pageNumber="14">n</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
</paragraph>
|
||
<paragraph id="8BAD369FFFBEFFA82722D332FF67FEC4" blockId="15.[115,759,235,963]" pageId="15" pageNumber="14">
|
||
<taxonomicName id="4C124D1CFFBEFFA82722D332FD7DFE25" authority="(Lourenco, Wilme & Waeber, 2016)" authorityName="Lowe & Kovařík" authorityYear="2019" baseAuthorityName="Lourenco, Wilme & Waeber" baseAuthorityYear="2016" box="[115,719,587,611]" class="Arachnida" family="Buthidae" genus="Teruelius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="15" pageNumber="14" phylum="Arthropoda" rank="species" species="ganzhorni" status="comb. nov.">
|
||
<emphasis id="B966EA8DFFBEFFA82722D332FEFAFE25" box="[115,328,587,611]" italics="true" pageId="15" pageNumber="14">Teruelius ganzhorni</emphasis>
|
||
(
|
||
<bibRefCitation id="EF834B6EFFBEFFA82607D332FD7AFE25" author="LOURENCO, W. R. & P. O. WAEBER & L. WILME" box="[342,712,587,611]" journalOrPublisher="Comptes Rendus Biologies" pageId="15" pageNumber="14" pagination="427 - 436" part="339" refId="ref40105" refString="LOURENCO, W. R., P. O. WAEBER & L. WILME. 2016 a. The geographical pattern of distribution of the genus Tityobuthus Pocock, 1890, a typical Ananterinae element endemic to Madagascar (Scorpiones: Buthidae). Comptes Rendus Biologies, 339: 427 - 436." title="The geographical pattern of distribution of the genus Tityobuthus Pocock, 1890, a typical Ananterinae element endemic to Madagascar (Scorpiones: Buthidae)" type="journal article" year="2016">Lourenço, Wilmé & Waeber, 2016</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A25557F6FFBEFFA82722D312FF67FEC4" box="[115,213,619,643]" pageId="15" pageNumber="14" rank="species">
|
||
<emphasis id="B966EA8DFFBEFFA82722D312FF03FEC5" bold="true" box="[115,177,619,643]" pageId="15" pageNumber="14">comb</emphasis>
|
||
.
|
||
<emphasis id="B966EA8DFFBEFFA827EED312FF7CFEC5" bold="true" box="[191,206,619,643]" pageId="15" pageNumber="14">n</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
</paragraph>
|
||
<paragraph id="8BAD369FFFBEFFA82722D3F2FD30FEE4" blockId="15.[115,759,235,963]" box="[115,642,651,675]" pageId="15" pageNumber="14">
|
||
<taxonomicName id="4C124D1CFFBEFFA82722D3F2FDA8FEE5" authority="(Kraepelin, 1900)" authorityName="Lowe & Kovařík" authorityYear="2019" baseAuthorityName="Kraepelin" baseAuthorityYear="1900" box="[115,538,651,675]" class="Arachnida" family="Buthidae" genus="Teruelius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="15" pageNumber="14" phylum="Arthropoda" rank="species" species="grandidieri" status="comb. nov.">
|
||
<emphasis id="B966EA8DFFBEFFA82722D3F2FEE7FEE5" box="[115,341,651,675]" italics="true" pageId="15" pageNumber="14">Teruelius grandidieri</emphasis>
|
||
(
|
||
<bibRefCitation id="EF834B6EFFBEFFA82635D3F2FDA1FEE5" author="KRAEPELIN, K." box="[356,531,651,675]" journalOrPublisher="Abhandlungen aus dem Gebiete der Naturwissenschaften. Herausgegeben vom Naturwissenschaftlichen Verein in Hamburg" pageId="15" pageNumber="14" pagination="1 - 17" part="16" refId="ref36651" refString="KRAEPELIN, K. 1900. Ueber einige neue Gliederspinnen. Abhandlungen aus dem Gebiete der Naturwissenschaften. Herausgegeben vom Naturwissenschaftlichen Verein in Hamburg, 16, 1 (4): 1 - 17." title="Ueber einige neue Gliederspinnen" type="journal article" year="1900">Kraepelin, 1900</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A25557F6FFBEFFA82570D3F2FD30FEE4" box="[545,642,651,675]" pageId="15" pageNumber="14" rank="species">
|
||
<emphasis id="B966EA8DFFBEFFA82570D3F2FDEDFEE5" bold="true" box="[545,607,651,675]" pageId="15" pageNumber="14">comb</emphasis>
|
||
.
|
||
<emphasis id="B966EA8DFFBEFFA8253DD3F2FDC9FEE5" bold="true" box="[620,635,651,675]" pageId="15" pageNumber="14">n</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
</paragraph>
|
||
<paragraph id="8BAD369FFFBEFFA82722D3D2FDC9FE84" blockId="15.[115,759,235,963]" box="[115,635,683,707]" pageId="15" pageNumber="14">
|
||
<taxonomicName id="4C124D1CFFBEFFA82722D3D2FDA1FE85" authority="(Lourenco, 1999)" authorityName="Lowe & Kovařík" authorityYear="2019" baseAuthorityName="Lourenco" baseAuthorityYear="1999" box="[115,531,683,707]" class="Arachnida" family="Buthidae" genus="Teruelius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="15" pageNumber="14" phylum="Arthropoda" rank="species" species="intertidalis" status="comb. nov.">
|
||
<emphasis id="B966EA8DFFBEFFA82722D3D2FEE3FE85" box="[115,337,683,707]" italics="true" pageId="15" pageNumber="14">Teruelius intertidalis</emphasis>
|
||
(
|
||
<bibRefCitation id="EF834B6EFFBEFFA8260ED3D2FDB9FE85" author="LOURENCO, W. R." box="[351,523,683,707]" journalOrPublisher="Entomologische Mitteilungen aus dem Zoologischen Museum Hamburg" pageId="15" pageNumber="14" pagination="183 - 188" part="12" refId="ref37834" refString="LOURENCO, W. R. 1999. A new species of Grosphus Simon (Scorpiones, Buthidae), the first record of an intertidal scorpion from Madagascar. Entomologische Mitteilungen aus dem Zoologischen Museum Hamburg, 12 (156): 183 - 188." title="A new species of Grosphus Simon (Scorpiones, Buthidae), the first record of an intertidal scorpion from Madagascar" type="journal article" year="1999">Lourenço, 1999</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A25557F6FFBEFFA82548D3D2FDC9FE84" box="[537,635,683,707]" pageId="15" pageNumber="14" rank="species">
|
||
<emphasis id="B966EA8DFFBEFFA82548D3D2FDE5FE85" bold="true" box="[537,599,683,707]" pageId="15" pageNumber="14">comb</emphasis>
|
||
.
|
||
<emphasis id="B966EA8DFFBEFFA82534D3D2FDC6FE85" bold="true" box="[613,628,683,707]" pageId="15" pageNumber="14">n</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
</paragraph>
|
||
<paragraph id="8BAD369FFFBEFFA82722D3B2FDF8FEA4" blockId="15.[115,759,235,963]" box="[115,586,715,739]" pageId="15" pageNumber="14">
|
||
<taxonomicName id="4C124D1CFFBEFFA82722D3B2FE50FEA5" authority="(Pocock, 1889)" authorityName="Lowe & Kovařík" authorityYear="2019" baseAuthorityName="Pocock" baseAuthorityYear="1889" box="[115,482,715,739]" class="Arachnida" family="Buthidae" genus="Teruelius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="15" pageNumber="14" phylum="Arthropoda" rank="species" species="limbatus" status="comb. nov.">
|
||
<emphasis id="B966EA8DFFBEFFA82722D3B2FE8AFEA5" box="[115,312,715,739]" italics="true" pageId="15" pageNumber="14">Teruelius limbatus</emphasis>
|
||
(
|
||
<bibRefCitation id="EF834B6EFFBEFFA82617D3B2FE68FEA5" author="POCOCK, R. I." box="[326,474,715,739]" journalOrPublisher="Annals and Magazine of Natural History" pageId="15" pageNumber="14" pagination="334 - 351" part="3" refId="ref41689" refString="POCOCK, R. I. 1889 a. Notes on some Buthidae, new and old. Annals and Magazine of Natural History, 3: 334 - 351." title="Notes on some Buthidae, new and old" type="journal article" year="1889">Pocock, 1889</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A25557F6FFBEFFA826B9D3B2FDF8FEA4" box="[488,586,715,739]" pageId="15" pageNumber="14" rank="species">
|
||
<emphasis id="B966EA8DFFBEFFA826B9D3B2FD94FEA5" bold="true" box="[488,550,715,739]" pageId="15" pageNumber="14">comb</emphasis>
|
||
.
|
||
<emphasis id="B966EA8DFFBEFFA82565D3B2FDF1FEA5" bold="true" box="[564,579,715,739]" pageId="15" pageNumber="14">n</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
</paragraph>
|
||
<paragraph id="8BAD369FFFBEFFA82722D392FDC7FF44" blockId="15.[115,759,235,963]" box="[115,629,747,771]" pageId="15" pageNumber="14">
|
||
<taxonomicName id="4C124D1CFFBEFFA82722D392FDBEFF45" authority="(Lourenco, 2014)" authorityName="Lowe & Kovařík" authorityYear="2019" baseAuthorityName="Lourenco" baseAuthorityYear="2014" box="[115,524,747,771]" class="Arachnida" family="Buthidae" genus="Teruelius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="15" pageNumber="14" phylum="Arthropoda" rank="species" species="magalieae" status="comb. nov.">
|
||
<emphasis id="B966EA8DFFBEFFA82722D392FEF9FF45" box="[115,331,747,771]" italics="true" pageId="15" pageNumber="14">Teruelius magalieae</emphasis>
|
||
(
|
||
<bibRefCitation id="EF834B6EFFBEFFA82608D392FDB7FF45" author="LOURENCO, W. R." box="[345,517,747,771]" journalOrPublisher="Zoosystema" pageId="15" pageNumber="14" pagination="631 - 645" part="36" refId="ref38903" refString="LOURENCO, W. R. 2014. The genus Grosphus Simon, 1880 in South-Western Madagascar, with the description of a new species (Scorpiones, Buthidae). Zoosystema, 36 (3): 631 - 645." title="The genus Grosphus Simon, 1880 in South-Western Madagascar, with the description of a new species (Scorpiones, Buthidae)" type="journal article" year="2014">Lourenço, 2014</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A25557F6FFBEFFA82542D392FDC7FF44" box="[531,629,747,771]" pageId="15" pageNumber="14" rank="species">
|
||
<emphasis id="B966EA8DFFBEFFA82542D392FDE3FF45" bold="true" box="[531,593,747,771]" pageId="15" pageNumber="14">comb</emphasis>
|
||
.
|
||
<emphasis id="B966EA8DFFBEFFA8250ED392FDDCFF45" bold="true" box="[607,622,747,771]" pageId="15" pageNumber="14">n</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
</paragraph>
|
||
<paragraph id="8BAD369FFFBEFFA82722D272FEECFF65" blockId="15.[115,759,235,963]" box="[115,350,779,803]" pageId="15" pageNumber="14">
|
||
(=
|
||
<taxonomicName id="4C124D1CFFBEFFA827C3D272FEF0FF65" box="[146,322,779,803]" class="Arachnida" family="Buthidae" genus="Teruelius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="15" pageNumber="14" phylum="Arthropoda" rank="species" species="mahafaliensis">
|
||
<emphasis id="B966EA8DFFBEFFA827C3D272FF13FF64" box="[146,161,779,802]" italics="true" pageId="15" pageNumber="14">T</emphasis>
|
||
.
|
||
<emphasis id="B966EA8DFFBEFFA827FFD272FEF0FF65" box="[174,322,779,803]" italics="true" pageId="15" pageNumber="14">mahafaliensis</emphasis>
|
||
</taxonomicName>
|
||
?)
|
||
</paragraph>
|
||
<paragraph id="8BAD369FFFBEFFA82722D252FE1BFF24" blockId="15.[115,759,235,963]" pageId="15" pageNumber="14">
|
||
<taxonomicName id="4C124D1CFFBEFFA82722D252FEF3FF25" authority="(Lourenco, Goodman & Ramilijaona, 2004)" authorityName="Lowe & Kovařík" authorityYear="2019" baseAuthorityName="Lourenco, Goodman & Ramilijaona" baseAuthorityYear="2004" class="Arachnida" family="Buthidae" genus="Teruelius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="15" pageNumber="14" phylum="Arthropoda" rank="species" species="mahafaliensis" status="comb. nov.">
|
||
<emphasis id="B966EA8DFFBEFFA82722D252FEC2FF05" box="[115,368,811,835]" italics="true" pageId="15" pageNumber="14">Teruelius mahafaliensis</emphasis>
|
||
(
|
||
<bibRefCitation id="EF834B6EFFBEFFA8262FD252FE88FF25" author="LOURENCO, W. R. & S. M. GOODMAN & O. RAMILIJAONA" journalOrPublisher="Revista Iberica de Aracnologia" pageId="15" pageNumber="14" pagination="225 - 234" part="9" refId="ref39550" refString="LOURENCO, W. R., S. M. GOODMAN & O. RAMILIJAONA. 2004. Three new species of Grosphus Simon from Madagascar (Scorpiones, Buthidae). Revista Iberica de Aracnologia, 9: 225 - 234." title="Three new species of Grosphus Simon from Madagascar (Scorpiones, Buthidae)" type="journal article" year="2004">Lourenço, Goodman & Ramilijaona, 2004</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A25557F6FFBEFFA82619D232FE1BFF24" box="[328,425,843,867]" pageId="15" pageNumber="14" rank="species">
|
||
<emphasis id="B966EA8DFFBEFFA82619D232FE34FF25" bold="true" box="[328,390,843,867]" pageId="15" pageNumber="14">comb</emphasis>
|
||
.
|
||
<emphasis id="B966EA8DFFBEFFA826C2D232FE10FF25" bold="true" box="[403,418,843,867]" pageId="15" pageNumber="14">n</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
</paragraph>
|
||
<paragraph id="8BAD369FFFBEFFA82722D212FDF3FFC4" blockId="15.[115,759,235,963]" box="[115,577,875,899]" pageId="15" pageNumber="14">
|
||
<taxonomicName id="4C124D1CFFBEFFA82722D212FE6BFFC5" authority="(Lourenco, 2004)" authorityName="Lowe & Kovařík" authorityYear="2019" baseAuthorityName="Lourenco" baseAuthorityYear="2004" box="[115,473,875,899]" class="Arachnida" family="Buthidae" genus="Teruelius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="15" pageNumber="14" phylum="Arthropoda" rank="species" species="olgae" status="comb. nov.">
|
||
<emphasis id="B966EA8DFFBEFFA82722D212FEA5FFC5" box="[115,279,875,899]" italics="true" pageId="15" pageNumber="14">Teruelius olgae</emphasis>
|
||
(
|
||
<bibRefCitation id="EF834B6EFFBEFFA82674D212FE60FFC5" author="LOURENCO, W. R." bookContentInfo="105 pp." box="[293,466,875,899]" editor="Raselimanana A. P. & Goodman S. M." journalOrPublisher="Recherches pour le developpement, Serie Sciences Biologiques" pageId="15" pageNumber="14" pagination="25 - 35" part="21" refId="ref38323" refString="LOURENCO, W. R. 2004 a. Scorpions du sud-ouest de Madagascar et en particulier de la foret des Mikea. Pp. 25 - 35, In: Raselimanana A. P. & Goodman S. M. (eds), Inventaire floristique et faunistique de la foret de Mikea: Paysage ecologique et diversite biologique d'une preoccupation majeure pour la conservation, Recherches pour le developpement, Serie Sciences Biologiques, 21: 105 pp." title="Scorpions du sud-ouest de Madagascar et en particulier de la foret des Mikea" type="book chapter" volumeTitle="Inventaire floristique et faunistique de la foret de Mikea: Paysage ecologique et diversite biologique d'une preoccupation majeure pour la conservation" year="2004">Lourenço, 2004</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A25557F6FFBEFFA8268ED212FDF3FFC4" box="[479,577,875,899]" pageId="15" pageNumber="14" rank="species">
|
||
<emphasis id="B966EA8DFFBEFFA8268ED212FDAFFFC5" bold="true" box="[479,541,875,899]" pageId="15" pageNumber="14">comb</emphasis>
|
||
.
|
||
<emphasis id="B966EA8DFFBEFFA8257AD212FD88FFC5" bold="true" box="[555,570,875,899]" pageId="15" pageNumber="14">n</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
</paragraph>
|
||
<paragraph id="8BAD369FFFBEFFA82722D2F2FD77FF84" blockId="15.[115,759,235,963]" pageId="15" pageNumber="14">
|
||
<taxonomicName id="4C124D1CFFBEFFA82722D2F2FDD7FFE5" authority="(Lourenco & Wilme, 2016)" authorityName="Lowe & Kovařík" authorityYear="2019" baseAuthorityName="Lourenco & Wilme" baseAuthorityYear="2016" box="[115,613,907,931]" class="Arachnida" family="Buthidae" genus="Teruelius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="15" pageNumber="14" phylum="Arthropoda" rank="species" species="sabineae" status="comb. nov.">
|
||
<emphasis id="B966EA8DFFBEFFA82722D2F2FE89FFE5" box="[115,315,907,931]" italics="true" pageId="15" pageNumber="14">Teruelius sabineae</emphasis>
|
||
(
|
||
<bibRefCitation id="EF834B6EFFBEFFA82618D2F2FDECFFE5" author="LOURENCO, W. R. & L. WILME" box="[329,606,907,931]" journalOrPublisher="Madagascar Conservation & Development" pageId="15" pageNumber="14" pagination="1 - 14" part="11" refId="ref40275" refString="LOURENCO, W. R. & L. WILME. 2016. Three new species of Grosphus Simon 1880, (Scorpiones: Buthidae) from Madagascar; possible vicariant cases within the Grosphus bistriatus group of species. Madagascar Conservation & Development, 11 (2): 1 - 14." title="Three new species of Grosphus Simon 1880, (Scorpiones: Buthidae) from Madagascar; possible vicariant cases within the Grosphus bistriatus group of species" type="journal article" year="2016">Lourenço & Wilmé, 2016</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A25557F6FFBEFFA8253DD2F2FD7FFFE4" box="[620,717,907,931]" pageId="15" pageNumber="14" rank="species">
|
||
<emphasis id="B966EA8DFFBEFFA8253DD2F2FD18FFE5" bold="true" box="[620,682,907,931]" pageId="15" pageNumber="14">comb</emphasis>
|
||
.
|
||
<emphasis id="B966EA8DFFBEFFA825E9D2F2FD75FFE5" bold="true" box="[696,711,907,931]" pageId="15" pageNumber="14">n</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
<taxonomicName id="4C124D1CFFBEFFA82722D2D2FDEEFF85" authority="(Lourenco & Wilme, 2016)" authorityName="Lowe & Kovařík" authorityYear="2019" baseAuthorityName="Lourenco & Wilme" baseAuthorityYear="2016" box="[115,604,939,963]" class="Arachnida" family="Buthidae" genus="Teruelius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="15" pageNumber="14" phylum="Arthropoda" rank="species" species="waeberi" status="comb. nov.">
|
||
<emphasis id="B966EA8DFFBEFFA82722D2D2FE80FF85" box="[115,306,939,963]" italics="true" pageId="15" pageNumber="14">Teruelius waeberi</emphasis>
|
||
(
|
||
<bibRefCitation id="EF834B6EFFBEFFA82611D2D2FDE7FF85" author="LOURENCO, W. R. & L. WILME" box="[320,597,939,963]" journalOrPublisher="Madagascar Conservation & Development" pageId="15" pageNumber="14" pagination="1 - 14" part="11" refId="ref40275" refString="LOURENCO, W. R. & L. WILME. 2016. Three new species of Grosphus Simon 1880, (Scorpiones: Buthidae) from Madagascar; possible vicariant cases within the Grosphus bistriatus group of species. Madagascar Conservation & Development, 11 (2): 1 - 14." title="Three new species of Grosphus Simon 1880, (Scorpiones: Buthidae) from Madagascar; possible vicariant cases within the Grosphus bistriatus group of species" type="journal article" year="2016">Lourenço & Wilmé, 2016</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A25557F6FFBEFFA82532D2D2FD77FF84" box="[611,709,939,963]" pageId="15" pageNumber="14" rank="species">
|
||
<emphasis id="B966EA8DFFBEFFA82532D2D2FD13FF85" bold="true" box="[611,673,939,963]" pageId="15" pageNumber="14">comb</emphasis>
|
||
.
|
||
<emphasis id="B966EA8DFFBEFFA825FED2D2FD0CFF85" bold="true" box="[687,702,939,963]" pageId="15" pageNumber="14">n</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
</paragraph>
|
||
<paragraph id="8BAD369FFFBEFFA82722D292FDB7F864" blockId="15.[115,773,1003,1059]" pageId="15" pageNumber="14">
|
||
See
|
||
<tableCitation id="C6900324FFBEFFA827F4D292FEAFF845" box="[165,285,1003,1027]" captionStart-0="Table 1" captionStart-1="Table 2" captionStart-2="Table 3" captionStartId-0="9.[115,171,1146,1167]" captionStartId-1="10.[113,169,1291,1312]" captionStartId-2="16.[110,166,1277,1298]" captionTargetId-0="graphics-417@9.[119,1468,237,1129]" captionTargetId-1="graphics-433@10.[121,1460,237,1263]" captionTargetId-2="graphics-479@16.[118,1471,242,1254]" captionTargetPageId-0="9" captionTargetPageId-1="10" captionTargetPageId-2="16" captionText-0="Table 1. Character states for species examined. States of 5 characters for Grosphus (4 spp., blue panels), Neogrosphus (1 sp., magenta panels) and Teruelius gen. n. (11 spp., yellow panels) examined in this study: position of trichobothrium d 2 on pedipalp femur; ratio R123 = d(Eb 2, Eb 3)/ d(Eb 1, Eb 2) of distances between pedipalp manus trichobothria Eb 1, Eb 2 and Eb 3; ranges of pectinal tooth counts (PTC) of males (♂) and females (♀); and size and shape of female basal pectinal tooth." captionText-1="Table 2. Character states for species examined. States of 6 characters for Grosphus (4 spp., blue panels), Neogrosphus (1 sp., magenta panels) and Teruelius gen. n. (11 spp., yellow panels) examined in this study: position of hemispermatophore basal lobe relative to base of flagellum; size of hemispermatophore posterior lobe; shape of spiracles; condition of metasoma I ventromedian carinae; telotarsal setation; and intensity of UV fluorescence." captionText-2="Table 3. Character states for species not examined. States of 6 characters for Grosphus (6 spp., blue panels), Neogrosphus (2 spp., magenta panels) and Teruelius gen. n. (7 spp., yellow panels) not examined in this study: position of trichobothrium d on pedipalp femur; ratio R = 2 123 d(Eb 2, Eb 3)/ d(Eb 1, Eb 2) of distances between pedipalp manus trichobothria Eb 1, Eb 2 and Eb 3; ranges of pectinal tooth counts (PTC) of males (♂) and females (♀);size and shape of female basal pectinal tooth; shape of spiracles; and telotarsal setation. Character states inferred from published descriptions and figures. PTC and basal pectinal tooth characters not indicated in cases where either males or females are not known or described. Other characters not indicated in cases where published descriptions did not yield clear information." httpUri-0="http://table.plazi.org/id/DF6D6617FFB8FFAE2722D503FCADF8A1" httpUri-2="http://table.plazi.org/id/DF6D6617FFA1FFB7273FD584FB00F9E5" pageId="15" pageNumber="14" tableUuid-0="DF6D6617FFB8FFAE2722D503FCADF8A1" tableUuid-2="DF6D6617FFA1FFB7273FD584FB00F9E5">Tables 1–3</tableCitation>
|
||
for diagnostic characters used to place the above species under
|
||
<taxonomicName id="4C124D1CFFBEFFA82600D572FE01F865" authority="Lowe & Kovařík, 2019" authorityName="Lowe & Kovařík" authorityYear="2019" box="[337,435,1035,1059]" class="Arachnida" family="Buthidae" genus="Teruelius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="15" pageNumber="14" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="B966EA8DFFBEFFA82600D572FE01F865" box="[337,435,1035,1059]" italics="true" pageId="15" pageNumber="14">Teruelius</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A25557F6FFBEFFA826EBD572FDB7F864" box="[442,517,1035,1059]" pageId="15" pageNumber="14" rank="genus">
|
||
<emphasis id="B966EA8DFFBEFFA826EBD572FE50F865" bold="true" box="[442,482,1035,1059]" pageId="15" pageNumber="14">gen</emphasis>
|
||
.
|
||
<emphasis id="B966EA8DFFBEFFA826BED572FE4CF865" bold="true" box="[495,510,1035,1059]" pageId="15" pageNumber="14">n</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3086514FFBEFFB22722D535FDFCF845" lastPageId="21" lastPageNumber="20" pageId="15" pageNumber="14" type="discussion">
|
||
<paragraph id="8BAD369FFFBEFFA82722D535FE16FBE0" blockId="15.[115,774,1099,2022]" pageId="15" pageNumber="14">
|
||
REMARKS. Recognition of
|
||
<taxonomicName id="4C124D1CFFBEFFA826F9D537FDB8F820" authority="Lowe & Kovařík, 2019" authorityName="Lowe & Kovařík" authorityYear="2019" box="[424,522,1102,1126]" class="Arachnida" family="Buthidae" genus="Teruelius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="15" pageNumber="14" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="B966EA8DFFBEFFA826F9D537FDB8F820" box="[424,522,1102,1126]" italics="true" pageId="15" pageNumber="14">Teruelius</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A25557F6FFBEFFA82547D537FDD5F823" box="[534,615,1102,1126]" pageId="15" pageNumber="14" rank="genus">
|
||
<emphasis id="B966EA8DFFBEFFA82547D537FD8CF820" bold="true" box="[534,574,1102,1126]" pageId="15" pageNumber="14">gen</emphasis>
|
||
.
|
||
<emphasis id="B966EA8DFFBEFFA82500D537FDD2F820" bold="true" box="[593,608,1102,1126]" pageId="15" pageNumber="14">n</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
as a separate genus, distinct from
|
||
<taxonomicName id="4C124D1CFFBEFFA82605D517FE09F8C0" authorityName="Vachon" authorityYear="1940" box="[340,443,1134,1158]" class="Arachnida" family="Buthidae" genus="Grosphus" kingdom="Animalia" order="Scorpiones" pageId="15" pageNumber="14" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B966EA8DFFBEFFA82605D517FE09F8C0" box="[340,443,1134,1158]" italics="true" pageId="15" pageNumber="14">Grosphus</emphasis>
|
||
</taxonomicName>
|
||
, necessitates revision of some previous concepts about taxonomy and biogeography of
|
||
<taxonomicName id="4C124D1CFFBEFFA82722D5D7FF68F880" authorityName="Vachon" authorityYear="1940" box="[115,218,1198,1222]" class="Arachnida" family="Buthidae" genus="Grosphus" kingdom="Animalia" order="Scorpiones" pageId="15" pageNumber="14" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B966EA8DFFBEFFA82722D5D7FF68F880" box="[115,218,1198,1222]" italics="true" pageId="15" pageNumber="14">Grosphus</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
<bibRefCitation id="EF834B6EFFBEFFA827B9D5D7FE64F880" author="LOURENCO, W. R. & L. WILME L & V. SOARIMALALA & P. O. WAEBER" box="[232,470,1198,1222]" journalOrPublisher="Revista Iberica de Arachnologia" pageId="15" pageNumber="14" pagination="61 - 69" part="30" refId="ref40436" refString="LOURENCO, W. R., L. WILME L, V. SOARIMALALA & P. O. WAEBER. 2017. Species of Grosphus Simon, 1880 associated to Grosphus simoni Lourenco, Goodman & Ramilijaona, 2004 with description of two new species (Scorpiones: Buthidae). Revista Iberica de Arachnologia, 30: 61 - 69." title="Species of Grosphus Simon, 1880 associated to Grosphus simoni Lourenco, Goodman & Ramilijaona, 2004 with description of two new species (Scorpiones: Buthidae)" type="journal article" year="2017">Lourenço et al. (2017)</bibRefCitation>
|
||
associated
|
||
<emphasis id="B966EA8DFFBEFFA82503D5D7FDD7F883" box="[594,613,1198,1221]" italics="true" pageId="15" pageNumber="14">G</emphasis>
|
||
. ‘
|
||
<taxonomicName id="4C124D1CFFBEFFA8252DD5D6FD71F880" authorityName="Lourenco, Goodman & Ramilijaona" authorityYear="2004" box="[636,707,1199,1222]" class="Arachnida" family="Buthidae" genus="Grosphus" kingdom="Animalia" order="Scorpiones" pageId="15" pageNumber="14" phylum="Arthropoda" rank="species" species="simoni">
|
||
<emphasis id="B966EA8DFFBEFFA8252DD5D6FD71F880" box="[636,707,1199,1222]" italics="true" pageId="15" pageNumber="14">simoni</emphasis>
|
||
</taxonomicName>
|
||
’(=
|
||
<taxonomicName id="4C124D1CFFBEFFA825BAD5D7FE86F8A0" class="Arachnida" family="Buthidae" genus="Grosphus" kingdom="Animalia" order="Scorpiones" pageId="15" pageNumber="14" phylum="Arthropoda" rank="species" species="madagascariensis">
|
||
<emphasis id="B966EA8DFFBEFFA825BAD5D7FD4CF883" box="[747,766,1198,1221]" italics="true" pageId="15" pageNumber="14">G</emphasis>
|
||
.
|
||
<emphasis id="B966EA8DFFBEFFA82722D5B7FE86F8A0" box="[115,308,1230,1254]" italics="true" pageId="15" pageNumber="14">madagascariensis</emphasis>
|
||
</taxonomicName>
|
||
) with
|
||
<taxonomicName id="4C124D1CFFBEFFA8262ED5B7FD89F8A0" authorityName="Lourenco, Wilme, Soarimalala & Waeber" authorityYear="2017" box="[383,571,1230,1254]" class="Arachnida" family="Buthidae" genus="Grosphus" kingdom="Animalia" order="Scorpiones" pageId="15" pageNumber="14" phylum="Arthropoda" rank="species" species="rakotoariveloi">
|
||
<emphasis id="B966EA8DFFBEFFA8262ED5B7FE20F8A3" box="[383,402,1230,1253]" italics="true" pageId="15" pageNumber="14">G</emphasis>
|
||
.
|
||
<emphasis id="B966EA8DFFBEFFA826F3D5B7FD89F8A0" box="[418,571,1230,1254]" italics="true" pageId="15" pageNumber="14">rakotoariveloi</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<emphasis id="B966EA8DFFBEFFA82522D5B7FD34F8A3" box="[627,646,1230,1253]" italics="true" pageId="15" pageNumber="14">G</emphasis>
|
||
. ‘
|
||
<taxonomicName id="4C124D1CFFBEFFA825CED5B7FD47F8A0" authorityName="Lourenco, Wilme, Soarimalala & Waeber, Previous" authorityYear="2017" box="[671,757,1230,1254]" class="Arachnida" family="Buthidae" genus="Grosphus" kingdom="Animalia" order="Scorpiones" pageId="15" pageNumber="14" phylum="Arthropoda" rank="species" species="halleuxi">
|
||
<emphasis id="B966EA8DFFBEFFA825CED5B7FD47F8A0" box="[671,757,1230,1254]" italics="true" pageId="15" pageNumber="14">halleuxi</emphasis>
|
||
</taxonomicName>
|
||
’, and subsequently
|
||
<bibRefCitation id="EF834B6EFFBEFFA8266FD597FDF8F940" author="LOURENCO, W. R. & L. WILME & P. O. WAEBER" box="[318,586,1262,1286]" journalOrPublisher="Revista Iberica de Arachnologia" pageId="15" pageNumber="14" pagination="73 - 80" part="32" refId="ref40598" refString="LOURENCO, W. R., L. WILME & P. O. WAEBER. 2018 b. Two more new species of Grosphus Simon, 1880, associated to the ' Grosphus simoni group' (Scorpiones: Buthidae) from the regions of the Tsingy de Bemaraha and Montagne d'Ambre (Madagascar). Revista Iberica de Arachnologia, 32: 73 - 80." title="Two more new species of Grosphus Simon, 1880, associated to the ' Grosphus simoni group' (Scorpiones: Buthidae) from the regions of the Tsingy de Bemaraha and Montagne d'Ambre (Madagascar)" type="journal article" year="2018">Lourenço et al. (2018b)</bibRefCitation>
|
||
elaborated on a ‘
|
||
<taxonomicName id="4C124D1CFFBEFFA8272DD477FE82F960" authorityName="Lourenco, Goodman & Ramilijaona" authorityYear="2004" box="[124,304,1294,1318]" class="Arachnida" family="Buthidae" genus="Grosphus" kingdom="Animalia" order="Scorpiones" pageId="15" pageNumber="14" phylum="Arthropoda" rank="species" species="simoni">
|
||
<emphasis id="B966EA8DFFBEFFA8272DD477FE82F960" box="[124,304,1294,1318]" italics="true" pageId="15" pageNumber="14">Grosphus simoni</emphasis>
|
||
</taxonomicName>
|
||
’ group, treating it as a monophyletic unit of closely related “sister” species including the aforementioned three, plus
|
||
<taxonomicName id="4C124D1CFFBEFFA827A4D437FEECF920" authorityName="Lourenco, Wilme & Waeber" authorityYear="2018" box="[245,350,1358,1382]" class="Arachnida" family="Buthidae" genus="Grosphus" kingdom="Animalia" order="Scorpiones" pageId="15" pageNumber="14" phylum="Arthropoda" rank="species" species="ambre">
|
||
<emphasis id="B966EA8DFFBEFFA827A4D437FEBAF923" box="[245,264,1358,1381]" italics="true" pageId="15" pageNumber="14">G</emphasis>
|
||
.
|
||
<emphasis id="B966EA8DFFBEFFA8264AD437FEECF920" box="[283,350,1358,1382]" italics="true" pageId="15" pageNumber="14">ambre</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C124D1CFFBEFFA82623D437FDB6F920" authorityName="Lowe & Kovařík" authorityYear="2019" baseAuthorityName="Lourenco, Wilme & Waeber" baseAuthorityYear="2018" box="[370,516,1358,1382]" class="Arachnida" family="Buthidae" genus="Teruelius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="15" pageNumber="14" phylum="Arthropoda" rank="species" species="bemaraha">
|
||
<emphasis id="B966EA8DFFBEFFA82623D437FE37F923" box="[370,389,1358,1381]" italics="true" pageId="15" pageNumber="14">G</emphasis>
|
||
.
|
||
<emphasis id="B966EA8DFFBEFFA826C9D437FDB6F920" box="[408,516,1358,1382]" italics="true" pageId="15" pageNumber="14">bemaraha</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C124D1CFFBEFFA82515D437FD4CF920" authorityName="Lowe & Kovařík" authorityYear="2019" baseAuthorityName="Lourenco, Goodman & Ramilijaona" baseAuthorityYear="2004" box="[580,766,1358,1382]" class="Arachnida" family="Buthidae" genus="Teruelius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="15" pageNumber="14" phylum="Arthropoda" rank="species" species="mahafaliensis">
|
||
<emphasis id="B966EA8DFFBEFFA82515D437FDE5F923" box="[580,599,1358,1381]" italics="true" pageId="15" pageNumber="14">G</emphasis>
|
||
.
|
||
<emphasis id="B966EA8DFFBEFFA8253BD437FD4CF920" box="[618,766,1358,1382]" italics="true" pageId="15" pageNumber="14">mahafaliensis</emphasis>
|
||
</taxonomicName>
|
||
. A group diagnosis was not provided but, for
|
||
<taxonomicName id="4C124D1CFFBEFFA8253FD417FD4CF9C0" authorityName="Lowe & Kovařík" authorityYear="2019" baseAuthorityName="Lourenco, Wilme & Waeber" baseAuthorityYear="2018" box="[622,766,1390,1414]" class="Arachnida" family="Buthidae" genus="Teruelius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="15" pageNumber="14" phylum="Arthropoda" rank="species" species="bemaraha">
|
||
<emphasis id="B966EA8DFFBEFFA8253FD417FD33F9C3" box="[622,641,1390,1413]" italics="true" pageId="15" pageNumber="14">G</emphasis>
|
||
.
|
||
<emphasis id="B966EA8DFFBEFFA825C3D417FD4CF9C0" box="[658,766,1390,1414]" italics="true" pageId="15" pageNumber="14">bemaraha</emphasis>
|
||
</taxonomicName>
|
||
, mention was made of “a number of features such as spiniform granules on the dorsal carinae of metasomal segments II-IV and on internal carinae of pedipalp femur and patella”. We place
|
||
<taxonomicName id="4C124D1CFFBEFFA827E7D497FEF5FA40" authorityName="Lowe & Kovařík" authorityYear="2019" baseAuthorityName="Lourenco, Wilme & Waeber" baseAuthorityYear="2018" box="[182,327,1518,1542]" class="Arachnida" family="Buthidae" genus="Teruelius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="15" pageNumber="14" phylum="Arthropoda" rank="species" species="bemaraha">
|
||
<emphasis id="B966EA8DFFBEFFA827E7D497FF7BFA43" box="[182,201,1518,1541]" italics="true" pageId="15" pageNumber="14">G</emphasis>
|
||
.
|
||
<emphasis id="B966EA8DFFBEFFA8278AD497FEF5FA40" box="[219,327,1518,1542]" italics="true" pageId="15" pageNumber="14">bemaraha</emphasis>
|
||
</taxonomicName>
|
||
under
|
||
<taxonomicName id="4C124D1CFFBEFFA826C9D497FE48FA40" authorityName="Lowe & Kovařík" authorityYear="2019" box="[408,506,1518,1542]" class="Arachnida" family="Buthidae" genus="Teruelius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="15" pageNumber="14" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B966EA8DFFBEFFA826C9D497FE48FA40" box="[408,506,1518,1542]" italics="true" pageId="15" pageNumber="14">Teruelius</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B966EA8DFFBEFFA82554D497FDF5FA40" bold="true" box="[517,583,1518,1542]" pageId="15" pageNumber="14">gen n</emphasis>
|
||
., on the basis of narrow spiracles, dense tarsal setation and high pectinal tooth count (
|
||
<tableCitation id="C6900324FFBEFFA827ECD757FEA2FA00" box="[189,272,1582,1606]" captionStart="Table 3" captionStartId="16.[110,166,1277,1298]" captionTargetId="graphics-479@16.[118,1471,242,1254]" captionTargetPageId="16" captionText="Table 3. Character states for species not examined. States of 6 characters for Grosphus (6 spp., blue panels), Neogrosphus (2 spp., magenta panels) and Teruelius gen. n. (7 spp., yellow panels) not examined in this study: position of trichobothrium d on pedipalp femur; ratio R = 2 123 d(Eb 2, Eb 3)/ d(Eb 1, Eb 2) of distances between pedipalp manus trichobothria Eb 1, Eb 2 and Eb 3; ranges of pectinal tooth counts (PTC) of males (♂) and females (♀);size and shape of female basal pectinal tooth; shape of spiracles; and telotarsal setation. Character states inferred from published descriptions and figures. PTC and basal pectinal tooth characters not indicated in cases where either males or females are not known or described. Other characters not indicated in cases where published descriptions did not yield clear information." httpUri="http://table.plazi.org/id/DF6D6617FFA1FFB7273FD584FB00F9E5" pageId="15" pageNumber="14" tableUuid="DF6D6617FFA1FFB7273FD584FB00F9E5">Table 3</tableCitation>
|
||
), and consider spiniform granules shared with
|
||
<taxonomicName id="4C124D1CFFBEFFA82722D737FF6CFA20" authorityName="Lourenco, Goodman & Ramilijaona" authorityYear="2004" box="[115,222,1614,1638]" class="Arachnida" family="Buthidae" genus="Grosphus" kingdom="Animalia" order="Scorpiones" pageId="15" pageNumber="14" phylum="Arthropoda" rank="species" species="simoni">
|
||
<emphasis id="B966EA8DFFBEFFA82722D737FF34FA23" box="[115,134,1614,1637]" italics="true" pageId="15" pageNumber="14">G</emphasis>
|
||
.
|
||
<emphasis id="B966EA8DFFBEFFA827C6D736FF6CFA20" box="[151,222,1615,1638]" italics="true" pageId="15" pageNumber="14">simoni</emphasis>
|
||
</taxonomicName>
|
||
to be a homoplasy.
|
||
<taxonomicName id="4C124D1CFFBEFFA82695D737FDE6FA20" authorityName="Lowe & Kovařík" authorityYear="2019" baseAuthorityName="Lourenco, Wilme & Waeber" baseAuthorityYear="2018" box="[452,596,1614,1638]" class="Arachnida" family="Buthidae" genus="Teruelius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="15" pageNumber="14" phylum="Arthropoda" rank="species" species="bemaraha">
|
||
<emphasis id="B966EA8DFFBEFFA82695D737FE65FA23" box="[452,471,1614,1637]" italics="true" pageId="15" pageNumber="14">G</emphasis>
|
||
.
|
||
<emphasis id="B966EA8DFFBEFFA826B9D737FDE6FA20" box="[488,596,1614,1638]" italics="true" pageId="15" pageNumber="14">bemaraha</emphasis>
|
||
</taxonomicName>
|
||
was claimed to be closer to
|
||
<taxonomicName id="4C124D1CFFBEFFA82652D717FE70FAC0" authorityName="Lourenco, Wilme, Soarimalala & Waeber" authorityYear="2017" box="[259,450,1646,1670]" class="Arachnida" family="Buthidae" genus="Grosphus" kingdom="Animalia" order="Scorpiones" pageId="15" pageNumber="14" phylum="Arthropoda" rank="species" species="rakotoariveloi">
|
||
<emphasis id="B966EA8DFFBEFFA82652D717FEA4FAC3" box="[259,278,1646,1669]" italics="true" pageId="15" pageNumber="14">G</emphasis>
|
||
.
|
||
<emphasis id="B966EA8DFFBEFFA82678D717FE70FAC0" box="[297,450,1646,1670]" italics="true" pageId="15" pageNumber="14">rakotoariveloi</emphasis>
|
||
</taxonomicName>
|
||
, but these species belong to different genera.
|
||
<bibRefCitation id="EF834B6EFFBEFFA82676D7F7FDF3FAE0" author="LOURENCO, W. R. & L. WILME & P. O. WAEBER" box="[295,577,1678,1702]" journalOrPublisher="Revista Iberica de Arachnologia" pageId="15" pageNumber="14" pagination="73 - 80" part="32" refId="ref40598" refString="LOURENCO, W. R., L. WILME & P. O. WAEBER. 2018 b. Two more new species of Grosphus Simon, 1880, associated to the ' Grosphus simoni group' (Scorpiones: Buthidae) from the regions of the Tsingy de Bemaraha and Montagne d'Ambre (Madagascar). Revista Iberica de Arachnologia, 32: 73 - 80." title="Two more new species of Grosphus Simon, 1880, associated to the ' Grosphus simoni group' (Scorpiones: Buthidae) from the regions of the Tsingy de Bemaraha and Montagne d'Ambre (Madagascar)" type="journal article" year="2018">Lourenço et al. (2018b: 74</bibRefCitation>
|
||
, fig.1) included
|
||
<taxonomicName id="4C124D1CFFBEFFA825BAD7F7FEB5FA80" authorityName="Lowe & Kovařík" authorityYear="2019" baseAuthorityName="Lourenco, Goodman & Ramilijaona" baseAuthorityYear="2004" class="Arachnida" family="Buthidae" genus="Teruelius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="15" pageNumber="14" phylum="Arthropoda" rank="species" species="mahafaliensis">
|
||
<emphasis id="B966EA8DFFBEFFA825BAD7F7FD4CFAE3" box="[747,766,1678,1701]" italics="true" pageId="15" pageNumber="14">G</emphasis>
|
||
.
|
||
<emphasis id="B966EA8DFFBEFFA82722D7D7FEB5FA80" box="[115,263,1710,1734]" italics="true" pageId="15" pageNumber="14">mahafaliensis</emphasis>
|
||
</taxonomicName>
|
||
in the ‘
|
||
<taxonomicName id="4C124D1CFFBEFFA82607D7D6FE2FFA80" authorityName="Lourenco, Goodman & Ramilijaona" authorityYear="2004" box="[342,413,1711,1734]" class="Arachnida" family="Buthidae" genus="Grosphus" kingdom="Animalia" order="Scorpiones" pageId="15" pageNumber="14" phylum="Arthropoda" rank="species" species="simoni">
|
||
<emphasis id="B966EA8DFFBEFFA82607D7D6FE2FFA80" box="[342,413,1711,1734]" italics="true" pageId="15" pageNumber="14">simoni</emphasis>
|
||
</taxonomicName>
|
||
’group, perhaps due to similarities to
|
||
<taxonomicName id="4C124D1CFFBEFFA827DED7B7FEA9FAA0" authorityName="Lowe & Kovařík" authorityYear="2019" baseAuthorityName="Lourenco, Wilme & Waeber" baseAuthorityYear="2018" box="[143,283,1742,1766]" class="Arachnida" family="Buthidae" genus="Teruelius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="15" pageNumber="14" phylum="Arthropoda" rank="species" species="bemaraha">
|
||
<emphasis id="B966EA8DFFBEFFA827DED7B7FF10FAA3" box="[143,162,1742,1765]" italics="true" pageId="15" pageNumber="14">G</emphasis>
|
||
.
|
||
<emphasis id="B966EA8DFFBEFFA827FED7B7FEA9FAA0" box="[175,283,1742,1766]" italics="true" pageId="15" pageNumber="14">bemaraha</emphasis>
|
||
</taxonomicName>
|
||
, noting in particular a high number of pectine teeth (but it lacks spiniform granules on metasomal carinae). In contrast, we find that
|
||
<taxonomicName id="4C124D1CFFBEFFA82629D676FD9BFB60" authorityName="Lowe & Kovařík" authorityYear="2019" baseAuthorityName="Lourenco, Goodman & Ramilijaona" baseAuthorityYear="2004" box="[376,553,1806,1830]" class="Arachnida" family="Buthidae" genus="Teruelius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="15" pageNumber="14" phylum="Arthropoda" rank="species" species="mahafaliensis" status="comb. nov.">
|
||
<emphasis id="B966EA8DFFBEFFA82629D676FE35FB60" box="[376,391,1807,1830]" italics="true" pageId="15" pageNumber="14">T</emphasis>
|
||
.
|
||
<emphasis id="B966EA8DFFBEFFA826C4D677FD9BFB60" box="[405,553,1806,1830]" italics="true" pageId="15" pageNumber="14">mahafaliensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A25557F6FFBEFFA82561D677FD23FB63" box="[560,657,1806,1830]" pageId="15" pageNumber="14" rank="species">
|
||
<emphasis id="B966EA8DFFBEFFA82561D677FDDCFB60" bold="true" box="[560,622,1806,1830]" pageId="15" pageNumber="14">comb</emphasis>
|
||
.
|
||
<emphasis id="B966EA8DFFBEFFA8252DD677FD39FB60" bold="true" box="[636,651,1806,1830]" pageId="15" pageNumber="14">n</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
is very far removed from
|
||
<emphasis id="B966EA8DFFBEFFA82642D657FE94FB03" box="[275,294,1838,1861]" italics="true" pageId="15" pageNumber="14">G</emphasis>
|
||
. ‘
|
||
<taxonomicName id="4C124D1CFFBEFFA8266CD656FE36FB00" authorityName="Lourenco, Goodman & Ramilijaona" authorityYear="2004" box="[317,388,1839,1862]" class="Arachnida" family="Buthidae" genus="Grosphus" kingdom="Animalia" order="Scorpiones" pageId="15" pageNumber="14" phylum="Arthropoda" rank="species" species="simoni">
|
||
<emphasis id="B966EA8DFFBEFFA8266CD656FE36FB00" box="[317,388,1839,1862]" italics="true" pageId="15" pageNumber="14">simoni</emphasis>
|
||
</taxonomicName>
|
||
’ (=
|
||
<taxonomicName id="4C124D1CFFBEFFA826FED657FD21FB00" box="[431,659,1838,1862]" class="Arachnida" family="Buthidae" genus="Grosphus" kingdom="Animalia" order="Scorpiones" pageId="15" pageNumber="14" phylum="Arthropoda" rank="species" species="madagascariensis">
|
||
<emphasis id="B966EA8DFFBEFFA826FED657FE70FB03" box="[431,450,1838,1861]" italics="true" pageId="15" pageNumber="14">G</emphasis>
|
||
.
|
||
<emphasis id="B966EA8DFFBEFFA82683D657FD21FB00" box="[466,659,1838,1862]" italics="true" pageId="15" pageNumber="14">madagascariensis</emphasis>
|
||
</taxonomicName>
|
||
), differing in all nine genus-level diagnostic characters, and in the fundamental architecture of the hemispermatophore capsule (cf.
|
||
<figureCitation id="13292A1AFFBEFFA827CFD6F7FF58FBE0" box="[158,234,1934,1958]" captionStart="Figures 71–85" captionStartId="26.[113,190,1701,1723]" captionTargetBox="[149,1434,231,1678]" captionTargetId="figure-287@26.[146,1446,225,1690]" captionTargetPageId="26" captionText="Figures 71–85. Hemispermatophores and capsule regions of Teruelius gen n. Multi-panel figures show: whole hemispermatophore; whole hemispermatophore and/or capsule with flagellum; capsule region in convex (or convex compressed), anterior and posterior views (panels in left to right sequence). Right hemispermatophores. Figure 71. T. ankarana, whole hemispermatophore. Scale bar: 4 mm. Figure 72. T. ankarana, capsule, left mirrored, Forêt d’Ankavanana, FMNH. Scale bar: 500 μm. Figure 73. T. grandidieri, whole hemispermatophore (flagellum truncated). Scale bar: 4 mm. Figure 74. T. grandidieri, capsule, Antsakabe River, FMNH. Scale bar: 500 μm. Figure 75. T. flavopiceus, whole hemispermatophore. Scale bar: 2 mm. Figure 76. T. flavopiceus, capsule, Madagascar, GLPC, FKCP. Scale bar: 500 μm. Figure 77. T. annulatus, capsule, Tsimanampetsotsa National Park, GLPC, FKCP. Scale bar: 500 μm. Figure 78. T. ankarafantsika, capsule and flagellum. Scale bar: 400 μm. Figure 79. T. ankarafantsika, capsule, Forêt d’Ankavanana, FMNH. Scale bar: 200 μm. Figure 80. T. ankarafantsika, capsule, Réserve Forestière de l’Ankarafantsika, FMNH. Scale bar: 200 μm. Figure 81. T. olgae, capsule, Itampolo village, FMNH. Scale bar: 500 μm. Figure 82. T. limbatus, whole hemispermatophore (flagellum truncated). Scale bar: 2 mm. Figure 83. T. limbatus, capsule, Forêt d’Ianasana, FMNH. Scale bar: 500 μm. Figure 84. T. mahafaliensis, capsule views, Zombitse-Vohibasia National Park, GLPC, FKCP. Scale bar: 500 μm. Figure 85. T. intertidalis, capsule, Madagascar, GLPC, FKCP. Scale bar: 500 μm." figureDoi="http://doi.org/10.5281/zenodo.7143641" httpUri="https://zenodo.org/record/7143641/files/figure.png" pageId="15" pageNumber="14">Fig. 84</figureCitation>
|
||
vs.
|
||
<figureCitation id="13292A1AFFBEFFA82644D6F7FE2AFBE0" box="[277,408,1934,1958]" captionStart="Figures 52–70" captionStartId="24.[113,190,1591,1613]" captionTargetBox="[116,1470,248,1526]" captionTargetId="figure-361@24.[113,1474,248,1527]" captionTargetPageId="24" captionText="Figures 52–70. Hemispermatophores and capsule regions of Grosphus and Neogrosphus. Multi-panel figures show: whole hemispermatophore; whole hemispermatophore and capsule with flagellum; capsule region in convex (and/ or convex compressed), anterior and posterior views (panels in left to right sequence). Right hemispermatophores unless indicated as mirrored left images. Figure 52. G. madagascariensis, whole hemispermatophore (scale bar: 2 mm), capsule and flagellum (scale bar: 1 mm). Figure 53. G. madagascariensis, capsule, Sc1197, Andasibe, GLPC, FKCP. Scale bar: 500 μm. Figure 54. G. madagascariensis, capsule, Anjiro, G. halleuxi nr topotype, GLPC. Scale bar: 500 μm. Figure 55. G. madagascariensis, capsule, Mandena- Fort Dauphin, G. mandena paratype, MHNG.Scale bar: 500 μm.Figure 56. G. madagascariensis, capsule, Madagascar, det. Vachon, MHNG. Scale bar: 500 μm. Figure 57. G. madagascariensis, capsule, Andasibe, GLPC, FKCP. Scale bar: 500 μm. Figure 58. G. goudoti, whole hemispermatophore. Scale bar: 2 mm. Figure 59. G. goudoti, capsule, Forêt de Bobankota, holotype, MHNG. Scale bar: 500 μm. Figure 60. G. hirtus, whole hemispermatophore (scale bar: 2 mm), capsule and flagellum (scale bar: 1 mm), Antsiranana, Ramena vill., mirrored left, GLPC, FKCP. Figure 61. G. hirtus, capsule, Mahajamba River, GLPC, FKCP. Scale bar: 500 μm. Figures 62–65. G. hirtus, capsules in convex view.Antsiranana, Ramena vill., mirrored left, GLPC, FKCP (62), Jardin Botanique, MHNG (63), Ranohira-Llakaka, ZMUH, mirrored left (64), Forêt de Vohitaly, MHNG (65). Scale bars: 500 μm. Figure 66. G. hirtus, capsule, Forest Station Ampijoroa, G. garciai holotype, MHNG. Scale bar: 500 μm. Figure 67. G. voahangyae, whole hemispermatophore. Scale bar: 2 mm. Figure 68. G. voahangyae, capsule, Analamy Forest, FMNH. Scale bar: 500 μm. Figure 69. N. griveaudi, whole hemispermatophore (flagellum truncated). Scale bar: 2 mm. Figure 70. N. griveaudi, capsule, mirrored left, Tsimanampetsotsa National Park, GLPC, FKCP." figureDoi="http://doi.org/10.5281/zenodo.7143636" httpUri="https://zenodo.org/record/7143636/files/figure.png" pageId="15" pageNumber="14">Figs. 52–57</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BAD369FFFBEFFA827F2D6D7FB51FFC5" blockId="15.[115,774,1099,2022]" lastBlockId="15.[818,1476,235,1347]" pageId="15" pageNumber="14">
|
||
<bibRefCitation id="EF834B6EFFBEFFA827F2D6D7FE66FB80" author="LOURENCO, W. R. & L. WILME & P. O. WAEBER" box="[163,468,1966,1990]" journalOrPublisher="Revista Iberica de Arachnologia" pageId="15" pageNumber="14" pagination="73 - 80" part="32" refId="ref40598" refString="LOURENCO, W. R., L. WILME & P. O. WAEBER. 2018 b. Two more new species of Grosphus Simon, 1880, associated to the ' Grosphus simoni group' (Scorpiones: Buthidae) from the regions of the Tsingy de Bemaraha and Montagne d'Ambre (Madagascar). Revista Iberica de Arachnologia, 32: 73 - 80." title="Two more new species of Grosphus Simon, 1880, associated to the ' Grosphus simoni group' (Scorpiones: Buthidae) from the regions of the Tsingy de Bemaraha and Montagne d'Ambre (Madagascar)" type="journal article" year="2018">Lourenço et al. (2018b)</bibRefCitation>
|
||
discussed biogeographic hypotheses attempting to explain the distribution of the incongruous, polyphyletic ‘
|
||
<taxonomicName id="4C124D1CFFBEFFA82309D192FABEFD44" authorityName="Lourenco, Goodman & Ramilijaona" authorityYear="2004" box="[1112,1292,235,259]" class="Arachnida" family="Buthidae" genus="Grosphus" kingdom="Animalia" order="Scorpiones" pageId="15" pageNumber="14" phylum="Arthropoda" rank="species" species="simoni">
|
||
<emphasis id="B966EA8DFFBEFFA82309D192FABEFD44" box="[1112,1292,235,259]" italics="true" pageId="15" pageNumber="14">Grosphus simoni</emphasis>
|
||
</taxonomicName>
|
||
’ group. The very wide distribution of the group meant that it was “adapted to humid, dry and subarid environments”, and the two most disjunct species,
|
||
<taxonomicName id="4C124D1CFFBEFFA824A5D032FB35FD25" authorityName="Lowe & Kovařík" authorityYear="2019" baseAuthorityName="Lourenco, Wilme & Waeber" baseAuthorityYear="2018" box="[1012,1159,331,355]" class="Arachnida" family="Buthidae" genus="Teruelius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="15" pageNumber="14" phylum="Arthropoda" rank="species" species="bemaraha">
|
||
<emphasis id="B966EA8DFFBEFFA824A5D032FBB5FD24" box="[1012,1031,331,354]" italics="true" pageId="15" pageNumber="14">G</emphasis>
|
||
.
|
||
<emphasis id="B966EA8DFFBEFFA8234AD032FB35FD25" box="[1051,1159,331,355]" italics="true" pageId="15" pageNumber="14">bemaraha</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C124D1CFFBEFFA82399D032FA31FD25" authorityName="Lowe & Kovařík" authorityYear="2019" baseAuthorityName="Lourenco, Goodman & Ramilijaona" baseAuthorityYear="2004" box="[1224,1411,331,355]" class="Arachnida" family="Buthidae" genus="Teruelius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="15" pageNumber="14" phylum="Arthropoda" rank="species" species="mahafaliensis">
|
||
<emphasis id="B966EA8DFFBEFFA82399D032FB69FD24" box="[1224,1243,331,354]" italics="true" pageId="15" pageNumber="14">G</emphasis>
|
||
.
|
||
<emphasis id="B966EA8DFFBEFFA823BED032FA31FD25" box="[1263,1411,331,355]" italics="true" pageId="15" pageNumber="14">mahafaliensis</emphasis>
|
||
</taxonomicName>
|
||
from western and southern localities, were speculated to “belong to relict populations, which may have survived in humid refugia encountered in the sedimentary basins during the dry episodes of the paleoclimate oscillations”. However, these two species are not closely related to the other four group members, but belong to
|
||
<taxonomicName id="4C124D1CFFBEFFA82314D372FB15FE65" authority="Lowe & Kovařík, 2019" authorityName="Lowe & Kovařík" authorityYear="2019" box="[1093,1191,523,547]" class="Arachnida" family="Buthidae" genus="Teruelius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="15" pageNumber="14" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="B966EA8DFFBEFFA82314D372FB15FE65" box="[1093,1191,523,547]" italics="true" pageId="15" pageNumber="14">Teruelius</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A25557F6FFBEFFA823E3D372FAB2FE64" box="[1202,1280,523,547]" pageId="15" pageNumber="14" rank="genus">
|
||
<emphasis id="B966EA8DFFBEFFA823E3D372FB68FE65" bold="true" box="[1202,1242,523,547]" pageId="15" pageNumber="14">gen</emphasis>
|
||
.
|
||
<emphasis id="B966EA8DFFBEFFA823BAD372FB48FE65" bold="true" box="[1259,1274,523,547]" pageId="15" pageNumber="14">n</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
, whose ancestors may have already been adapted to dry environments. Although Pleistocene climatic fluctuations could be relevant for recent speciation events in
|
||
<taxonomicName id="4C124D1CFFBEFFA82358D312FBC2FEC5" authorityName="Vachon" authorityYear="1940" box="[1033,1136,619,643]" class="Arachnida" family="Buthidae" genus="Grosphus" kingdom="Animalia" order="Scorpiones" pageId="15" pageNumber="14" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B966EA8DFFBEFFA82358D312FBC2FEC5" box="[1033,1136,619,643]" italics="true" pageId="15" pageNumber="14">Grosphus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C124D1CFFBEFFA823F2D312FAB7FEC5" authority="Lowe & Kovařík, 2019" authorityName="Lowe & Kovařík" authorityYear="2019" box="[1187,1285,619,643]" class="Arachnida" family="Buthidae" genus="Teruelius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="15" pageNumber="14" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="B966EA8DFFBEFFA823F2D312FAB7FEC5" box="[1187,1285,619,643]" italics="true" pageId="15" pageNumber="14">Teruelius</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A25557F6FFBEFFA8225AD312FAE7FEC4" box="[1291,1365,619,643]" pageId="15" pageNumber="14" rank="genus">
|
||
<emphasis id="B966EA8DFFBEFFA8225AD312FA81FEC5" bold="true" box="[1291,1331,619,643]" pageId="15" pageNumber="14">gen</emphasis>
|
||
.
|
||
<emphasis id="B966EA8DFFBEFFA82211D312FAFDFEC5" bold="true" box="[1344,1359,619,643]" pageId="15" pageNumber="14">n</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
, the many correlated characters separating these two genera suggest a far earlier split, as in other taxa. For example, dated molecular phylogenies of
|
||
<collectingCountry id="F305760FFFBEFFA824B2D3B2FBD6FEA5" box="[995,1124,715,739]" name="Madagascar" pageId="15" pageNumber="14">Madagascar</collectingCountry>
|
||
archaeid spiders (Wood et al., 2015),
|
||
<taxonomicName id="4C124D1CFFBEFFA824D1D392FC59FF45" box="[896,1003,747,771]" class="Reptilia" family="Chamaeleonidae" genus="Brookesia" kingdom="Animalia" order="Squamata" pageId="15" pageNumber="14" phylum="Chordata" rank="genus">
|
||
<emphasis id="B966EA8DFFBEFFA824D1D392FC59FF45" box="[896,1003,747,771]" italics="true" pageId="15" pageNumber="14">Brookesia</emphasis>
|
||
</taxonomicName>
|
||
and other chameleons (
|
||
<bibRefCitation id="EF834B6EFFBEFFA823A3D392FA72FF45" author="Tolley" box="[1266,1472,747,771]" etAl="et al." firstAuthor="Tolley" pageId="15" pageNumber="14" pagination="20130184" refId="ref43585" refString="TOLLEY, K. T., TOWNSEND & M. VENCES. 2013. Largescale phylogeny of chameleons suggests African origins and Eocene diversification. Proceedings of the Royal Society B, 280: 20130184." type="journal article" year="2013">Tolley et al., 2013</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EF834B6EFFBEFFA82463D272FB97FF65" author="Townsend" box="[818,1061,779,803]" etAl="et al." firstAuthor="Townsend" pageId="15" pageNumber="14" pagination="641 - 656" refId="ref43622" refString="TOWNSEND, T. M., D. R. VIEITES, F. GLAW & M. VENCES. 2009. Testing species-level diversification hypotheses in Madagascar: the case of microendemic Brookesia leaf chameleons. Systematic Biology, 58 (6): 641 - 656." type="journal article" year="2009">Townsend et al., 2009</bibRefCitation>
|
||
) and
|
||
<taxonomicName id="4C124D1CFFBEFFA82334D272FB51FF64" authorityName="Boulenger" authorityYear="1887" box="[1125,1251,779,802]" class="Reptilia" family="Gerrhosauridae" genus="Zonosaurus" kingdom="Animalia" order="Squamata" pageId="15" pageNumber="14" phylum="Chordata" rank="genus">
|
||
<emphasis id="B966EA8DFFBEFFA82334D272FB51FF64" box="[1125,1251,779,802]" italics="true" pageId="15" pageNumber="14">Zonosaurus</emphasis>
|
||
</taxonomicName>
|
||
plated lizards (
|
||
<bibRefCitation id="EF834B6EFFBEFFA822DCD272FC19FF05" author="BLAIR, C. & B. P. NOONAN & J. L. BROWN & A. P. RASELIMANANA & M. VENCES & A. D. YODER" journalOrPublisher="Journal of Evolutionary Biology" pageId="15" pageNumber="14" pagination="481 - 492" part="28" refId="ref32794" refString="BLAIR, C., B. P. NOONAN, J. L. BROWN, A. P. RASELIMANANA, M. VENCES & A. D. YODER. 2015. Multilocus phylogenetic and geospatial analyses illuminate diversification patterns and the biogeographic history of Malagasy endemic plated lizards (Gerrhosauridae: Zonosaurinae). Journal of Evolutionary Biology, 28: 481 - 492." title="Multilocus phylogenetic and geospatial analyses illuminate diversification patterns and the biogeographic history of Malagasy endemic plated lizards (Gerrhosauridae: Zonosaurinae)" type="journal article" year="2015">Blair et al., 2015</bibRefCitation>
|
||
) have revealed that the majority of divergences in these other endemic taxa are quite deep, occurring long before the advent of Pleistocene climate cycles.
|
||
</paragraph>
|
||
<paragraph id="8BAD369FFFBEFFA82433D2F2FAEFF905" blockId="15.[818,1476,235,1347]" pageId="15" pageNumber="14">
|
||
Species of
|
||
<taxonomicName id="4C124D1CFFBEFFA82487D2F2FB8AFFE5" authority="Lowe & Kovařík, 2019" authorityName="Lowe & Kovařík" authorityYear="2019" box="[982,1080,907,931]" class="Arachnida" family="Buthidae" genus="Teruelius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="15" pageNumber="14" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="B966EA8DFFBEFFA82487D2F2FB8AFFE5" box="[982,1080,907,931]" italics="true" pageId="15" pageNumber="14">Teruelius</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A25557F6FFBEFFA8236ED2F2FB3BFFE4" box="[1087,1161,907,931]" pageId="15" pageNumber="14" rank="genus">
|
||
<emphasis id="B966EA8DFFBEFFA8236ED2F2FBD5FFE5" bold="true" box="[1087,1127,907,931]" pageId="15" pageNumber="14">gen</emphasis>
|
||
.
|
||
<emphasis id="B966EA8DFFBEFFA82325D2F2FB31FFE5" bold="true" box="[1140,1155,907,931]" pageId="15" pageNumber="14">n</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
can be loosely subdivided by size and coloration: large species,
|
||
<taxonomicName id="4C124D1CFFBEFFA823CDD2D2FA9DFF85" authorityName="Lowe & Kovařík" authorityYear="2019" baseAuthorityName="Kraepelin" baseAuthorityYear="1900" box="[1180,1327,939,963]" class="Arachnida" family="Buthidae" genus="Teruelius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="15" pageNumber="14" phylum="Arthropoda" rank="species" species="flavopiceus">
|
||
<emphasis id="B966EA8DFFBEFFA823CDD2D2FB19FF84" box="[1180,1195,939,962]" italics="true" pageId="15" pageNumber="14">T</emphasis>
|
||
.
|
||
<emphasis id="B966EA8DFFBEFFA823E6D2D2FA9DFF85" box="[1207,1327,939,963]" italics="true" pageId="15" pageNumber="14">flavopiceus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C124D1CFFBEFFA8226DD2D2FA0FFF85" authorityName="Lowe & Kovařík" authorityYear="2019" baseAuthorityName="Lourenco & Goodman" baseAuthorityYear="2003" box="[1340,1469,939,963]" class="Arachnida" family="Buthidae" genus="Teruelius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="15" pageNumber="14" phylum="Arthropoda" rank="species" species="ankarana">
|
||
<emphasis id="B966EA8DFFBEFFA8226DD2D2FAF9FF84" box="[1340,1355,939,962]" italics="true" pageId="15" pageNumber="14">T</emphasis>
|
||
.
|
||
<emphasis id="B966EA8DFFBEFFA82206D2D2FA0FFF85" box="[1367,1469,939,963]" italics="true" pageId="15" pageNumber="14">ankarana</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C124D1CFFBEFFA82463D2B2FC79FFA5" authorityName="Lowe & Kovařík" authorityYear="2019" baseAuthorityName="Kraepelin" baseAuthorityYear="1900" box="[818,971,971,995]" class="Arachnida" family="Buthidae" genus="Teruelius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="15" pageNumber="14" phylum="Arthropoda" rank="species" species="grandidieri">
|
||
<emphasis id="B966EA8DFFBEFFA82463D2B2FCF3FFA4" box="[818,833,971,994]" italics="true" pageId="15" pageNumber="14">T</emphasis>
|
||
.
|
||
<emphasis id="B966EA8DFFBEFFA82403D2B2FC79FFA5" box="[850,971,971,995]" italics="true" pageId="15" pageNumber="14">grandidieri</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C124D1CFFBEFFA82357D2B2FBC1FFA5" authorityName="Lowe & Kovařík" authorityYear="2019" baseAuthorityName="Lourenco" baseAuthorityYear="2012" box="[1030,1139,971,995]" class="Arachnida" family="Buthidae" genus="Teruelius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="15" pageNumber="14" phylum="Arthropoda" rank="species" species="bicolor">
|
||
<emphasis id="B966EA8DFFBEFFA82357D2B2FBA7FFA4" box="[1030,1045,971,994]" italics="true" pageId="15" pageNumber="14">T</emphasis>
|
||
.
|
||
<emphasis id="B966EA8DFFBEFFA82377D2B2FBC1FFA5" box="[1062,1139,971,995]" italics="true" pageId="15" pageNumber="14">bicolor</emphasis>
|
||
</taxonomicName>
|
||
; species with patterns of dark stripes on tergites (‘
|
||
<taxonomicName id="4C124D1CFFBEFFA8235FD292FBC1F845" authorityName="Lowe & Kovařík" authorityYear="2019" baseAuthorityName="Kraepelin" baseAuthorityYear="1900" box="[1038,1139,1003,1027]" class="Arachnida" family="Buthidae" genus="Teruelius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="15" pageNumber="14" phylum="Arthropoda" rank="species" species="bistriatus">
|
||
<emphasis id="B966EA8DFFBEFFA8235FD292FBC1F845" box="[1038,1139,1003,1027]" italics="true" pageId="15" pageNumber="14">bistriatus</emphasis>
|
||
</taxonomicName>
|
||
’ group of
|
||
<bibRefCitation id="EF834B6EFFBEFFA823B8D292FCDFF865" author="LOURENCO, W. R. & L. WILME" journalOrPublisher="Madagascar Conservation & Development" pageId="15" pageNumber="14" pagination="1 - 14" part="11" refId="ref40275" refString="LOURENCO, W. R. & L. WILME. 2016. Three new species of Grosphus Simon 1880, (Scorpiones: Buthidae) from Madagascar; possible vicariant cases within the Grosphus bistriatus group of species. Madagascar Conservation & Development, 11 (2): 1 - 14." title="Three new species of Grosphus Simon 1880, (Scorpiones: Buthidae) from Madagascar; possible vicariant cases within the Grosphus bistriatus group of species" type="journal article" year="2016">Lourenço & Wilmé, 2016</bibRefCitation>
|
||
):
|
||
<taxonomicName id="4C124D1CFFBEFFA824D7D572FBFBF865" authorityName="Lowe & Kovařík" authorityYear="2019" baseAuthorityName="Lourenco" baseAuthorityYear="2003" box="[902,1097,1035,1059]" class="Arachnida" family="Buthidae" genus="Teruelius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="15" pageNumber="14" phylum="Arthropoda" rank="species" species="ankarafantsika">
|
||
<emphasis id="B966EA8DFFBEFFA824D7D572FC27F864" box="[902,917,1035,1058]" italics="true" pageId="15" pageNumber="14">T</emphasis>
|
||
.
|
||
<emphasis id="B966EA8DFFBEFFA824F8D572FBFBF865" box="[937,1097,1035,1059]" italics="true" pageId="15" pageNumber="14">ankarafantsika</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C124D1CFFBEFFA8230CD572FB57F865" authorityName="Lowe & Kovařík" authorityYear="2019" baseAuthorityName="Kraepelin" baseAuthorityYear="1900" box="[1117,1253,1035,1059]" class="Arachnida" family="Buthidae" genus="Teruelius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="15" pageNumber="14" phylum="Arthropoda" rank="species" species="bistriatus">
|
||
<emphasis id="B966EA8DFFBEFFA8230CD572FBDEF864" box="[1117,1132,1035,1058]" italics="true" pageId="15" pageNumber="14">T</emphasis>
|
||
.
|
||
<emphasis id="B966EA8DFFBEFFA823D1D572FB57F865" box="[1152,1253,1035,1059]" italics="true" pageId="15" pageNumber="14">bistriatus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C124D1CFFBEFFA823A8D572FA28F865" authorityName="Lowe & Kovařík" authorityYear="2019" baseAuthorityName="Lourenco & Wilme" baseAuthorityYear="2016" box="[1273,1434,1035,1059]" class="Arachnida" family="Buthidae" genus="Teruelius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="15" pageNumber="14" phylum="Arthropoda" rank="species" species="eliseanneae">
|
||
<emphasis id="B966EA8DFFBEFFA823A8D572FABAF864" box="[1273,1288,1035,1058]" italics="true" pageId="15" pageNumber="14">T</emphasis>
|
||
.
|
||
<emphasis id="B966EA8DFFBEFFA8224DD572FA28F865" box="[1308,1434,1035,1059]" italics="true" pageId="15" pageNumber="14">eliseanneae</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C124D1CFFBEFFA822FFD572FCE6F805" authorityName="Lowe & Kovařík" authorityYear="2019" baseAuthorityName="Lourenco" baseAuthorityYear="1996" class="Arachnida" family="Buthidae" genus="Teruelius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="15" pageNumber="14" phylum="Arthropoda" rank="species" species="feti">
|
||
<emphasis id="B966EA8DFFBEFFA822FFD572FA0FF864" box="[1454,1469,1035,1058]" italics="true" pageId="15" pageNumber="14">T</emphasis>
|
||
.
|
||
<emphasis id="B966EA8DFFBEFFA82463D552FCE6F805" box="[818,852,1067,1091]" italics="true" pageId="15" pageNumber="14">feti</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C124D1CFFBEFFA82437D552FC51F805" authorityName="Lowe & Kovařík" authorityYear="2019" baseAuthorityName="Pocock" baseAuthorityYear="1889" box="[870,995,1067,1091]" class="Arachnida" family="Buthidae" genus="Teruelius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="15" pageNumber="14" phylum="Arthropoda" rank="species" species="limbatus">
|
||
<emphasis id="B966EA8DFFBEFFA82437D552FCC7F804" box="[870,885,1067,1090]" italics="true" pageId="15" pageNumber="14">T</emphasis>
|
||
.
|
||
<emphasis id="B966EA8DFFBEFFA824D6D552FC51F805" box="[903,995,1067,1091]" italics="true" pageId="15" pageNumber="14">limbatus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C124D1CFFBEFFA824A4D552FBC7F805" authorityName="Lowe & Kovařík" authorityYear="2019" baseAuthorityName="Lourenco & Wilme" baseAuthorityYear="2016" box="[1013,1141,1067,1091]" class="Arachnida" family="Buthidae" genus="Teruelius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="15" pageNumber="14" phylum="Arthropoda" rank="species" species="sabineae">
|
||
<emphasis id="B966EA8DFFBEFFA824A4D552FBB6F804" box="[1013,1028,1067,1090]" italics="true" pageId="15" pageNumber="14">T</emphasis>
|
||
.
|
||
<emphasis id="B966EA8DFFBEFFA82347D552FBC7F805" box="[1046,1141,1067,1091]" italics="true" pageId="15" pageNumber="14">sabineae</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C124D1CFFBEFFA823E3D552FA9BF805" authorityName="Lowe & Kovařík" authorityYear="2019" baseAuthorityName="Lourenco & Wilme" baseAuthorityYear="2016" box="[1202,1321,1067,1091]" class="Arachnida" family="Buthidae" genus="Teruelius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="15" pageNumber="14" phylum="Arthropoda" rank="species" species="waeberi">
|
||
<emphasis id="B966EA8DFFBEFFA823E3D552FB73F804" box="[1202,1217,1067,1090]" italics="true" pageId="15" pageNumber="14">T</emphasis>
|
||
.
|
||
<emphasis id="B966EA8DFFBEFFA82382D552FA9BF805" box="[1235,1321,1067,1091]" italics="true" pageId="15" pageNumber="14">waeberi</emphasis>
|
||
</taxonomicName>
|
||
; species with almost uniform yellow, orange or brown tergites, and maybe darker metasoma IV or V:
|
||
<taxonomicName id="4C124D1CFFBEFFA8233CD512FB4BF8C5" authorityName="Lowe & Kovařík" authorityYear="2019" baseAuthorityName="Fage" baseAuthorityYear="1929" box="[1133,1273,1131,1155]" class="Arachnida" family="Buthidae" genus="Teruelius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="15" pageNumber="14" phylum="Arthropoda" rank="species" species="annulatus">
|
||
<emphasis id="B966EA8DFFBEFFA8233CD512FBCEF8C4" box="[1133,1148,1131,1154]" italics="true" pageId="15" pageNumber="14">T</emphasis>
|
||
.
|
||
<emphasis id="B966EA8DFFBEFFA823C1D512FB4BF8C5" box="[1168,1273,1131,1155]" italics="true" pageId="15" pageNumber="14">annulatus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C124D1CFFBEFFA8225DD512FA29F8C5" authorityName="Lowe & Kovařík" authorityYear="2019" baseAuthorityName="Lourenco, Wilme & Waeber" baseAuthorityYear="2018" box="[1292,1435,1131,1155]" class="Arachnida" family="Buthidae" genus="Teruelius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="15" pageNumber="14" phylum="Arthropoda" rank="species" species="bemaraha">
|
||
<emphasis id="B966EA8DFFBEFFA8225DD512FAA9F8C4" box="[1292,1307,1131,1154]" italics="true" pageId="15" pageNumber="14">T</emphasis>
|
||
.
|
||
<emphasis id="B966EA8DFFBEFFA8227ED512FA29F8C5" box="[1327,1435,1131,1155]" italics="true" pageId="15" pageNumber="14">bemaraha</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C124D1CFFBEFFA822FFD512FC2CF8E5" authorityName="Lowe & Kovařík" authorityYear="2019" baseAuthorityName="Lourenco, Wilme & Waeber" baseAuthorityYear="2016" class="Arachnida" family="Buthidae" genus="Teruelius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="15" pageNumber="14" phylum="Arthropoda" rank="species" species="ganzhorni">
|
||
<emphasis id="B966EA8DFFBEFFA822FFD512FA0FF8C4" box="[1454,1469,1131,1154]" italics="true" pageId="15" pageNumber="14">T</emphasis>
|
||
.
|
||
<emphasis id="B966EA8DFFBEFFA82463D5F2FC2CF8E5" box="[818,926,1163,1187]" italics="true" pageId="15" pageNumber="14">ganzhorni</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C124D1CFFBEFFA824FDD5F2FB8CF8E5" authorityName="Lowe & Kovařík" authorityYear="2019" baseAuthorityName="Lourenco" baseAuthorityYear="1999" box="[940,1086,1163,1187]" class="Arachnida" family="Buthidae" genus="Teruelius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="15" pageNumber="14" phylum="Arthropoda" rank="species" species="intertidalis">
|
||
<emphasis id="B966EA8DFFBEFFA824FDD5F2FC09F8E4" box="[940,955,1163,1186]" italics="true" pageId="15" pageNumber="14">T</emphasis>
|
||
.
|
||
<emphasis id="B966EA8DFFBEFFA82498D5F2FB8CF8E5" box="[969,1086,1163,1187]" italics="true" pageId="15" pageNumber="14">intertidalis</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C124D1CFFBEFFA8231DD5F2FB65F8E5" authorityName="Lowe & Kovařík" authorityYear="2019" baseAuthorityName="Lourenco" baseAuthorityYear="2014" box="[1100,1239,1163,1187]" class="Arachnida" family="Buthidae" genus="Teruelius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="15" pageNumber="14" phylum="Arthropoda" rank="species" species="magalieae">
|
||
<emphasis id="B966EA8DFFBEFFA8231DD5F2FBE9F8E4" box="[1100,1115,1163,1186]" italics="true" pageId="15" pageNumber="14">T</emphasis>
|
||
.
|
||
<emphasis id="B966EA8DFFBEFFA82339D5F2FB65F8E5" box="[1128,1239,1163,1187]" italics="true" pageId="15" pageNumber="14">magalieae</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C124D1CFFBEFFA823B4D5F2FA24F8E5" authorityName="Lowe & Kovařík" authorityYear="2019" baseAuthorityName="Lourenco, Goodman & Ramilijaona" baseAuthorityYear="2004" box="[1253,1430,1163,1187]" class="Arachnida" family="Buthidae" genus="Teruelius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="15" pageNumber="14" phylum="Arthropoda" rank="species" species="mahafaliensis">
|
||
<emphasis id="B966EA8DFFBEFFA823B4D5F2FB46F8E4" box="[1253,1268,1163,1186]" italics="true" pageId="15" pageNumber="14">T</emphasis>
|
||
.
|
||
<emphasis id="B966EA8DFFBEFFA82253D5F2FA24F8E5" box="[1282,1430,1163,1187]" italics="true" pageId="15" pageNumber="14">mahafaliensis</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C124D1CFFBEFFA82463D5D2FC3BF885" authorityName="Lowe & Kovařík" authorityYear="2019" baseAuthorityName="Lourenco" baseAuthorityYear="2004" box="[818,905,1195,1219]" class="Arachnida" family="Buthidae" genus="Teruelius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="15" pageNumber="14" phylum="Arthropoda" rank="species" species="olgae">
|
||
<emphasis id="B966EA8DFFBEFFA82463D5D2FCF3F884" box="[818,833,1195,1218]" italics="true" pageId="15" pageNumber="14">T</emphasis>
|
||
.
|
||
<emphasis id="B966EA8DFFBEFFA8241FD5D2FC3BF885" box="[846,905,1195,1219]" italics="true" pageId="15" pageNumber="14">olgae</emphasis>
|
||
</taxonomicName>
|
||
. These groupings have been used to construct species keys, in conjunction with some other characters including shapes of female basal pectine teeth (
|
||
<bibRefCitation id="EF834B6EFFBEFFA8239CD592FAFBF945" author="FAGE, L." box="[1229,1353,1259,1283]" journalOrPublisher="Societe d'Editions Geographiques Maritimes et Coloniales, Paris" pageId="15" pageNumber="14" pagination="637 - 693" refId="ref33536" refString="FAGE, L. 1929. Les scorpions de Madagascar, leur affinites, leur distribution geographique. Pp. 637 - 693 in: Faune des Colonies Francaises. Societe d'Editions Geographiques Maritimes et Coloniales, Paris." title="Les scorpions de Madagascar, leur affinites, leur distribution geographique" type="book chapter" volumeTitle="Faune des Colonies Francaises" year="1929">Fage, 1929</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EF834B6EFFBEFFA82206D592FCC4F965" author="LOURENCO, W. R." journalOrPublisher="Revue Suisse de Zoologie" pageId="15" pageNumber="14" pagination="141 - 154" part="110" refId="ref38276" refString="LOURENCO, W. R. 2003 c. New taxonomic considerations on some species of the genus Grosphus Simon, with description of a new species (Scorpiones, Buthidae). Revue Suisse de Zoologie, 110 (1): 141 - 154." title="New taxonomic considerations on some species of the genus Grosphus Simon, with description of a new species (Scorpiones, Buthidae)" type="journal article" year="2003">Lourenço, 2003c</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="EF834B6EFFBEFFA824D2D472FC75F965" author="LOURENCO, W. R." bookContentInfo="105 pp." box="[899,967,1291,1315]" editor="Raselimanana A. P. & Goodman S. M." journalOrPublisher="Recherches pour le developpement, Serie Sciences Biologiques" pageId="15" pageNumber="14" pagination="25 - 35" part="21" refId="ref38323" refString="LOURENCO, W. R. 2004 a. Scorpions du sud-ouest de Madagascar et en particulier de la foret des Mikea. Pp. 25 - 35, In: Raselimanana A. P. & Goodman S. M. (eds), Inventaire floristique et faunistique de la foret de Mikea: Paysage ecologique et diversite biologique d'une preoccupation majeure pour la conservation, Recherches pour le developpement, Serie Sciences Biologiques, 21: 105 pp." title="Scorpions du sud-ouest de Madagascar et en particulier de la foret des Mikea" type="book chapter" volumeTitle="Inventaire floristique et faunistique de la foret de Mikea: Paysage ecologique et diversite biologique d'une preoccupation majeure pour la conservation" year="2004">2004a</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="EF834B6EFFBEFFA82482D472FBBEF965" author="LOURENCO, W. R." box="[979,1036,1291,1315]" journalOrPublisher="Zoosystema" pageId="15" pageNumber="14" pagination="631 - 645" part="36" refId="ref38903" refString="LOURENCO, W. R. 2014. The genus Grosphus Simon, 1880 in South-Western Madagascar, with the description of a new species (Scorpiones, Buthidae). Zoosystema, 36 (3): 631 - 645." title="The genus Grosphus Simon, 1880 in South-Western Madagascar, with the description of a new species (Scorpiones, Buthidae)" type="journal article" year="2014">2014</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EF834B6EFFBEFFA82348D472FAA6F965" author="LOURENCO, W. R. & V. SOARIMALALA & S. M. GOODMAN" box="[1049,1300,1291,1315]" journalOrPublisher="Boletin de la Sociedad Entomologica Aragonesa" pageId="15" pageNumber="14" pagination="369 - 375" part="41" refId="ref39953" refString="LOURENCO, W. R., V. SOARIMALALA & S. M. GOODMAN. 2007 b. Scorpions of south-west Madagascar. II. The species of Grosphus Simon (Scorpiones, Buthidae). Boletin de la Sociedad Entomologica Aragonesa, 41: 369 - 375." title="Scorpions of south-west Madagascar. II. The species of Grosphus Simon (Scorpiones, Buthidae)" type="journal article" year="2007">Lourenço et al., 2007b</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EF834B6EFFBEFFA82271D472FA0AF965" author="Vachon" box="[1312,1464,1291,1315]" firstAuthor="Vachon" pageId="15" pageNumber="14" pagination="476 - 483" refId="ref44004" refString="VACHON, M. 1969. Grosphus griveaudi, nouvelle espece de Scorpion Buthidae malgache. Bulletin du Museum national d'Histoire naturelle, Paris, (2), 41: 476 - 483." type="journal article" year="1969">Vachon, 1969</bibRefCitation>
|
||
). Monophyly of these groupings remains to be tested.
|
||
</paragraph>
|
||
<paragraph id="8BAD369FFFBEFFA82463D412FC44F9C5" blockId="15.[818,1477,1387,2019]" box="[818,1014,1387,1411]" pageId="15" pageNumber="14">NEW SYNONYMIES.</paragraph>
|
||
<paragraph id="8BAD369FFFBEFFA82463D4F2FB1EF984" blockId="15.[818,1477,1387,2019]" pageId="15" pageNumber="14">
|
||
<taxonomicName id="4C124D1CFFBEFFA82463D4F2FAA3F9E5" authority="Lourenco & Wilme, 2015" authorityName="Lourenco & Wilme" authorityYear="2015" box="[818,1297,1419,1443]" class="Arachnida" family="Buthidae" genus="Grosphus" kingdom="Animalia" order="Scorpiones" pageId="15" pageNumber="14" phylum="Arthropoda" rank="species" species="makay">
|
||
<emphasis id="B966EA8DFFBEFFA82463D4F2FC5BF9E5" box="[818,1001,1419,1443]" italics="true" pageId="15" pageNumber="14">Grosphus makay</emphasis>
|
||
<bibRefCitation id="EF834B6EFFBEFFA824A5D4F2FAA3F9E5" author="LOURENCO, W. R. & L. WILME" box="[1012,1297,1419,1443]" journalOrPublisher="Entomologische Mitteilungen aus dem Zoologischen Museum Hamburg" pageId="15" pageNumber="14" pagination="207 - 223" part="17" refId="ref40161" refString="LOURENCO, W. R. & L. WILME. 2015 a. Species of Grosphus Simon, 1880, associated to the group madagascariensis / hirtus (Scorpiones: Buthidae); description of a peculiar new species from the humid eastern forests of Madagascar. Entomologische Mitteilungen aus dem Zoologischen Museum Hamburg, 17 (194): 207 - 223." title="Species of Grosphus Simon, 1880, associated to the group madagascariensis / hirtus (Scorpiones: Buthidae); description of a peculiar new species from the humid eastern forests of Madagascar" type="journal article" year="2015">Lourenço & Wilmé, 2015</bibRefCitation>
|
||
</taxonomicName>
|
||
<emphasis id="B966EA8DFFBEFFA8224AD4F2FA98F9E5" bold="true" box="[1307,1322,1419,1443]" pageId="15" pageNumber="14">=</emphasis>
|
||
<taxonomicName id="4C124D1CFFBEFFA82264D4F2FC5FF985" authority="(Lourenco, 1996)" authorityName="Lowe & Kovařík" authorityYear="2019" baseAuthorityName="Lourenco" baseAuthorityYear="1996" class="Arachnida" family="Buthidae" genus="Teruelius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="15" pageNumber="14" phylum="Arthropoda" rank="species" species="feti" status="comb. n., syn. n.">
|
||
<emphasis id="B966EA8DFFBEFFA82264D4F2FA76F9E5" box="[1333,1476,1419,1443]" italics="true" pageId="15" pageNumber="14">Teruelius feti</emphasis>
|
||
(
|
||
<bibRefCitation id="EF834B6EFFBEFFA8246BD4D2FC54F985" author="LOURENCO, W. R." box="[826,998,1451,1475]" journalOrPublisher="Biogeographie de Madagascar. ORSTOM, Paris" pageId="15" pageNumber="14" pagination="441 - 455" refId="ref37587" refString="LOURENCO, W. R. 1996 a. Origins and affinities of the scorpion fauna of Madagascar. Pp. 441 - 455 in: Lourenco, W. R. (Ed.). Biogeographie de Madagascar. ORSTOM, Paris." title="Origins and affinities of the scorpion fauna of Madagascar" type="book chapter" volumeTitle="Lourenco, W. R" year="1996">Lourenço, 1996</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A25557F6FFBEFFA824A5D4D2FB1EF984" box="[1012,1196,1451,1475]" pageId="15" pageNumber="14">
|
||
<emphasis id="B966EA8DFFBEFFA824A5D4D2FB80F985" bold="true" box="[1012,1074,1451,1475]" pageId="15" pageNumber="14">comb</emphasis>
|
||
.
|
||
<emphasis id="B966EA8DFFBEFFA8236ED4D2FBFCF985" bold="true" box="[1087,1102,1451,1475]" pageId="15" pageNumber="14">n</emphasis>
|
||
.,
|
||
<emphasis id="B966EA8DFFBEFFA82333D4D2FB3BF985" bold="true" box="[1122,1161,1451,1475]" pageId="15" pageNumber="14">syn</emphasis>
|
||
.
|
||
<emphasis id="B966EA8DFFBEFFA823C7D4D2FB17F985" bold="true" box="[1174,1189,1451,1475]" pageId="15" pageNumber="14">n</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
</paragraph>
|
||
<paragraph id="8BAD369FFFBEFFB72433D4B2FD04FA60" blockId="15.[818,1477,1387,2019]" lastBlockId="16.[113,772,1486,2022]" lastPageId="16" lastPageNumber="15" pageId="15" pageNumber="14">
|
||
<taxonomicName id="4C124D1CFFBEFFA82433D4B2FC46F9A5" authorityName="Lourenco" authorityYear="1996" box="[866,1012,1483,1507]" class="Arachnida" family="Buthidae" genus="Grosphus" kingdom="Animalia" order="Scorpiones" pageId="15" pageNumber="14" phylum="Arthropoda" rank="species" species="feti">
|
||
<emphasis id="B966EA8DFFBEFFA82433D4B2FC46F9A5" box="[866,1012,1483,1507]" italics="true" pageId="15" pageNumber="14">Grosphus feti</emphasis>
|
||
</taxonomicName>
|
||
was described by
|
||
<bibRefCitation id="EF834B6EFFBEFFA82392D4B2FA35F9A5" author="LOURENCO, W. R." bookContentInfo="102 pp." box="[1219,1415,1483,1507]" journalOrPublisher="Museum National d'Histoire Naturelle, Paris" pageId="15" pageNumber="14" refId="ref37632" refString="LOURENCO, W. R. 1996 b. Faune de Madagascar. 87. Scorpions (Chelicerata, Scorpiones). Museum National d'Histoire Naturelle, Paris. 102 pp." title="Faune de Madagascar. 87. Scorpions (Chelicerata, Scorpiones)" type="book" year="1996">Lourenço (1996b)</bibRefCitation>
|
||
from a juvenile male
|
||
<typeStatus id="54A9883DFFBEFFA824BDD492FBFEFA45" box="[1004,1100,1515,1539]" pageId="15" pageNumber="14" type="holotype">holotype</typeStatus>
|
||
, ostensibly collected from “Prov. Tulear, Tanjon’ I Vohimena [= Cap Sainte Marie] Réserve spéciale,
|
||
<date id="FFAC105FFFBEFFA824C8D752FC58FA05" box="[921,1002,1579,1603]" pageId="15" pageNumber="14" value="1995-10">
|
||
<collectingDate id="EFE8E9B7FFBEFFA824C8D752FC58FA05" box="[921,1002,1579,1603]" pageId="15" pageNumber="14" value="1995-10">X.1995</collectingDate>
|
||
</date>
|
||
” and deposited in FMNH.
|
||
<collectingCounty id="62CC4E13FFBEFFA8224FD752FAF3FA04" box="[1310,1345,1579,1602]" pageId="15" pageNumber="14">We</collectingCounty>
|
||
loaned and studied the
|
||
<typeStatus id="54A9883DFFBEFFA824ECD732FBA8FA25" box="[957,1050,1611,1635]" pageId="15" pageNumber="14" type="holotype">holotype</typeStatus>
|
||
and a second juvenile male labeled as “
|
||
<specimenCount id="9D14FD16FFBEFFA8240ED712FCC6FAC5" box="[863,884,1643,1667]" pageId="15" pageNumber="14" type="male">♂</specimenCount>
|
||
<typeStatus id="54A9883DFFBEFFA824D1D712FC6CFAC5" box="[896,990,1643,1667]" pageId="15" pageNumber="14" type="paratype">paratype</typeStatus>
|
||
” (which we concur is conspecific with the
|
||
<typeStatus id="54A9883DFFBEFFA82463D7F2FC26FAE5" box="[818,916,1675,1699]" pageId="15" pageNumber="14" type="holotype">holotype</typeStatus>
|
||
).
|
||
<collectingMunicipality id="6BC9ACE5FFBEFFA824FCD7F2FC56FAE5" box="[941,996,1675,1699]" pageId="15" pageNumber="14">Until</collectingMunicipality>
|
||
now, this species was only known from these
|
||
<specimenCount id="9D14FD16FFBEFFA82423D7D2FC6DFA85" box="[882,991,1707,1731]" pageId="15" pageNumber="14" type="generic">two types</specimenCount>
|
||
.
|
||
<location id="8ECD6044FFBEFFA824B8D7D2FBECFA85" LSID="urn:lsid:plazi:treatment:03BB8789FFBCFFB224B1D341FDFCF845:8ECD6044FFBEFFA824B8D7D2FBECFA85" box="[1001,1118,1707,1731]" country="Madagascar" county="We" latitude="-22.683332" longLatPrecision="125" longitude="44.83" municipality="Until" name="Associated" pageId="15" pageNumber="14" stateProvince="Toliara">Associated</location>
|
||
with the type, we found locality labels (
|
||
<figureCitation id="13292A1AFFBEFFA824D0D7B2FC6DFAA5" box="[897,991,1739,1763]" captionStart="Figure 459" captionStartId="97.[113,181,1982,2004]" captionTargetBox="[113,1474,234,1946]" captionTargetId="figure-36@97.[113,1474,234,1949]" captionTargetPageId="97" captionText="Figure 459. Teruelius feti. Holotype juvenile male. Habitus, dorsal view, with specimen labels (FMNH). Scale bar: 5 mm (holotype). Figure 459a. Adult female from Makay Mts., carapace and tergites I–II." figureDoi="http://doi.org/10.5281/zenodo.7143796" httpUri="https://zenodo.org/record/7143796/files/figure.png" pageId="15" pageNumber="14">Fig. 459</figureCitation>
|
||
) that differ from the published type locality: “
|
||
<collectingCountry id="F305760FFFBEFFA8246DD792FBB2FB45" box="[828,1024,1771,1795]" name="Madagascar" pageId="15" pageNumber="14">MADAGASCAR</collectingCountry>
|
||
:
|
||
<collectingRegion id="49D6F87DFFBEFFA8235DD792FB51FB45" box="[1036,1251,1771,1795]" country="Madagascar" name="Toliara" pageId="15" pageNumber="14">Province de Toliara</collectingRegion>
|
||
, Fôret de Vohimena,
|
||
<locationDeviation id="9A1F2FBDFFBEFFA82463D672FBA0FB65" box="[818,1042,1803,1827]" location="Sakaraha" pageId="15" pageNumber="14">
|
||
<quantity id="4CEA9B7AFFBEFFA82463D672FCCBFB65" box="[818,889,1803,1827]" metricMagnitude="4" metricUnit="m" metricValue="3.5" pageId="15" pageNumber="14" unit="km" value="35.0">35 km</quantity>
|
||
SE Sakaraha
|
||
</locationDeviation>
|
||
,
|
||
<date id="FFAC105FFFBEFFA82371D672FB18FB65" box="[1056,1194,1803,1827]" pageId="15" pageNumber="14" value="1996-01-17" valueMax="1996-01-24" valueMin="1996-01-17">
|
||
<collectingDate id="EFE8E9B7FFBEFFA82371D672FB18FB65" box="[1056,1194,1803,1827]" pageId="15" pageNumber="14" value="1996-01-17" valueMax="1996-01-24" valueMin="1996-01-17">17-24.i.1996</collectingDate>
|
||
</date>
|
||
, MyrCE-7541,
|
||
<geoCoordinate id="EE265058FFBEFFA82209D672FA76FB65" box="[1368,1476,1803,1827]" degrees="22" direction="south" minutes="41.0" orientation="latitude" pageId="15" pageNumber="14" precision="92" value="-22.683332">22°41.0’S</geoCoordinate>
|
||
<geoCoordinate id="EE265058FFBEFFA82463D652FC10FB05" box="[818,930,1835,1859]" degrees="44" direction="east" minutes="49.8" orientation="longitude" pageId="15" pageNumber="14" precision="92" value="44.83">44°49.8’E</geoCoordinate>
|
||
,
|
||
<quantity id="4CEA9B7AFFBEFFA824E0D652FC49FB04" box="[945,1019,1835,1859]" metricMagnitude="2" metricUnit="m" metricValue="7.8" pageId="15" pageNumber="14" unit="m" value="780.0">
|
||
<elevation id="003FD1ACFFBEFFA824E0D652FC49FB04" box="[945,1019,1835,1859]" metricMagnitude="2" metricUnit="m" metricValue="7.8" pageId="15" pageNumber="14" unit="m" value="780.0">780 m</elevation>
|
||
</quantity>
|
||
,
|
||
<collectorName id="26E75349FFBEFFA8235AD652FB0FFB05" box="[1035,1213,1835,1859]" pageId="15" pageNumber="14">S. M. Goodman</collectorName>
|
||
0000 011 031 FMNH- INS
|
||
<taxonomicName id="4C124D1CFFBEFFA82437D632FBD4FB24" authority="Lourenco" authorityName="Lourenco" authorityYear="1996" box="[870,1126,1867,1891]" class="Arachnida" family="Buthidae" genus="Grosphus" kingdom="Animalia" order="Scorpiones" pageId="15" pageNumber="14" phylum="Arthropoda" rank="species" species="feti">
|
||
<emphasis id="B966EA8DFFBEFFA82437D632FC45FB25" box="[870,1015,1867,1891]" italics="true" pageId="15" pageNumber="14">Grosphus feti</emphasis>
|
||
Lourenço
|
||
</taxonomicName>
|
||
<typeStatus id="54A9883DFFBEFFA8233ED632FAB2FB25" box="[1135,1280,1867,1891]" pageId="15" pageNumber="14" type="holotype">HOLOTYPE</typeStatus>
|
||
: det. 1996 ”. This locality is ca.
|
||
<quantity id="4CEA9B7AFFBEFFA82485D612FB9BFBC5" box="[980,1065,1899,1923]" metricMagnitude="5" metricUnit="m" metricValue="3.25" pageId="15" pageNumber="14" unit="km" value="325.0">325 km</quantity>
|
||
roughly north of the published type locality at Cap Sainte Marie and the collection date of
|
||
<date id="FFAC105FFFBEFFA82220D6F2FCD5FB85" pageId="15" pageNumber="14" value="1996-01">
|
||
<collectingDate id="EFE8E9B7FFBEFFA82220D6F2FCD5FB85" pageId="15" pageNumber="14" value="1996-01">January 1996</collectingDate>
|
||
</date>
|
||
is several months later. Either a labeling error occurred after description, or the published type locality is incorrect. Cap Sainte Marie is a biological study area and frequent source of scorpion materials (e.g.,
|
||
<bibRefCitation id="EF834B6EFFA1FFB726B7D497FD4EFA40" author="LOURENCO, W. R. & L. WILME" box="[486,764,1518,1542]" journalOrPublisher="Madagascar Conservation & Development" pageId="16" pageNumber="15" pagination="1 - 14" part="11" refId="ref40275" refString="LOURENCO, W. R. & L. WILME. 2016. Three new species of Grosphus Simon 1880, (Scorpiones: Buthidae) from Madagascar; possible vicariant cases within the Grosphus bistriatus group of species. Madagascar Conservation & Development, 11 (2): 1 - 14." title="Three new species of Grosphus Simon 1880, (Scorpiones: Buthidae) from Madagascar; possible vicariant cases within the Grosphus bistriatus group of species" type="journal article" year="2016">Lourenço & Wilmé, 2016</bibRefCitation>
|
||
) so data labels of specimens could have been confused.
|
||
</paragraph>
|
||
<paragraph id="8BAD369FFFA1FFB7278AD060FA2AF892" pageId="16" pageNumber="15">
|
||
<table id="F912C43FFFA10058272AD071FA01F892" box="[123,1459,264,1236]" gridcols="8" gridrows="9" pageId="16" pageNumber="15">
|
||
<tr id="352234DDFFA10058272AD071FA01FD02" box="[123,1459,264,324]" gridrow="0" pageId="16" pageNumber="15">
|
||
<th id="76F35DA1FFA10058272AD071FE23FD02" box="[123,401,264,324]" gridcol="0" gridrow="0" pageId="16" pageNumber="15">
|
||
<emphasis id="B966EA8DFFA1FFB7278AD060FEFDFD77" bold="true" box="[219,335,281,305]" pageId="16" pageNumber="15">SPECIES</emphasis>
|
||
</th>
|
||
<th id="76F35DA1FFA1005826EFD071FD9BFD02" box="[446,553,264,324]" gridcol="1" gridrow="0" pageId="16" pageNumber="15">
|
||
<emphasis id="B966EA8DFFA1FFB726EFD061FDA9FD76" bold="true" box="[446,539,280,304]" pageId="16" pageNumber="15">
|
||
femur
|
||
<emphasis id="B966EA8DFFA1FFB7255CD061FDA9FD76" bold="true" box="[525,539,280,304]" italics="true" pageId="16" pageNumber="15">d</emphasis>
|
||
</emphasis>
|
||
<emphasis id="B966EA8DFFA1FFB72571D05AFD9BFD72" bold="true" box="[544,553,291,308]" pageId="16" pageNumber="15">2</emphasis>
|
||
</th>
|
||
<th id="76F35DA1FFA100582514D071FD39FD02" box="[581,651,264,324]" gridcol="2" gridrow="0" pageId="16" pageNumber="15">
|
||
<emphasis id="B966EA8DFFA1FFB7251FD061FDD2FD69" bold="true" box="[590,608,280,303]" italics="true" pageId="16" pageNumber="15">R</emphasis>
|
||
<emphasis id="B966EA8DFFA1FFB72537D05AFD30FD72" bold="true" box="[614,642,291,308]" pageId="16" pageNumber="15">123</emphasis>
|
||
</th>
|
||
<th id="76F35DA1FFA100582592D071FCA5FD02" box="[707,791,264,324]" gridcol="3" gridrow="0" pageId="16" pageNumber="15">
|
||
<emphasis id="B966EA8DFFA1FFB72592D060FCA5FD77" bold="true" box="[707,791,279,305]" pageId="16" pageNumber="15">PTC ♂</emphasis>
|
||
</th>
|
||
<th id="76F35DA1FFA100582432D071FC04FD02" box="[867,950,264,324]" gridcol="4" gridrow="0" pageId="16" pageNumber="15">
|
||
<emphasis id="B966EA8DFFA1FFB72432D060FC04FD77" bold="true" box="[867,950,279,305]" pageId="16" pageNumber="15">PTC ♀</emphasis>
|
||
</th>
|
||
<th id="76F35DA1FFA1005824BCD071FB34FD02" box="[1005,1158,264,324]" gridcol="5" gridrow="0" pageId="16" pageNumber="15">
|
||
<emphasis id="B966EA8DFFA1FFB724BCD071FBD5FD05" bold="true" pageId="16" pageNumber="15">basal pectine tooth ♀</emphasis>
|
||
</th>
|
||
<th id="76F35DA1FFA1005823E2D071FAA8FD02" box="[1203,1306,264,324]" gridcol="6" gridrow="0" pageId="16" pageNumber="15">
|
||
<emphasis id="B966EA8DFFA1FFB723E2D060FAA8FD77" bold="true" box="[1203,1306,281,305]" pageId="16" pageNumber="15">spiracles</emphasis>
|
||
</th>
|
||
<th id="76F35DA1FFA10058226ED071FA01FD02" box="[1343,1459,264,324]" gridcol="7" gridrow="0" pageId="16" pageNumber="15">
|
||
<emphasis id="B966EA8DFFA1FFB7226ED071FA15FD05" bold="true" box="[1343,1459,264,323]" pageId="16" pageNumber="15">telotarsus setation</emphasis>
|
||
</th>
|
||
</tr>
|
||
<tr id="352234DDFFA10058272AD018FA01FEBA" box="[123,1459,353,764]" gridrow="1" pageId="16" pageNumber="15">
|
||
<th id="76F35DA1FFA10058272AD018FE23FEBA" box="[123,401,353,764]" gridcol="0" gridrow="1" pageId="16" pageNumber="15">
|
||
<emphasis id="B966EA8DFFA1FFB7272AD018FEEEFEBD" italics="true" pageId="16" pageNumber="15">
|
||
<taxonomicName id="4C124D1CFFA1FFB7272AD018FE84FD3F" authorityName="Lourenco, Wilme & Waeber" authorityYear="2018" box="[123,310,353,377]" class="Arachnida" family="Buthidae" genus="Grosphus" kingdom="Animalia" order="Scorpiones" pageId="16" pageNumber="15" phylum="Arthropoda" rank="species" species="ambre">Grosphus ambre</taxonomicName>
|
||
<taxonomicName id="4C124D1CFFA1FFB7272AD0EDFEC3FDEA" authorityName="Lourenco, Goodman & Ramilijaona" authorityYear="2004" box="[123,369,404,428]" class="Arachnida" family="Buthidae" genus="Grosphus" kingdom="Animalia" order="Scorpiones" pageId="16" pageNumber="15" phylum="Arthropoda" rank="species" species="darainensis">Grosphus darainensis</taxonomicName>
|
||
<taxonomicName id="4C124D1CFFA1FFB7272AD0BEFEC0FD99" authorityName="Lourenco & Goodman" authorityYear="2009" box="[123,370,455,479]" class="Arachnida" family="Buthidae" genus="Grosphus" kingdom="Animalia" order="Scorpiones" pageId="16" pageNumber="15" phylum="Arthropoda" rank="species" species="mayottensis">Grosphus mayottensis</taxonomicName>
|
||
<taxonomicName id="4C124D1CFFA1FFB7272AD083FE8CFE54" authorityName="Lourenco, Qi & Goodman" authorityYear="2007" box="[123,318,506,530]" class="Arachnida" family="Buthidae" genus="Grosphus" kingdom="Animalia" order="Scorpiones" pageId="16" pageNumber="15" phylum="Arthropoda" rank="species" species="polskyi">Grosphus polskyi</taxonomicName>
|
||
<taxonomicName id="4C124D1CFFA1FFB7272AD354FE3CFE03" authorityName="Lourenco, Wilme, Soarimalala & Waeber" authorityYear="2017" box="[123,398,557,581]" class="Arachnida" family="Buthidae" genus="Grosphus" kingdom="Animalia" order="Scorpiones" pageId="16" pageNumber="15" phylum="Arthropoda" rank="species" species="rakotoariveloi">Grosphus rakotoariveloi</taxonomicName>
|
||
<taxonomicName id="4C124D1CFFA1FFB7272AD319FEEAFE3E" authorityName="Lourenco, Soarimalala & Goodman" authorityYear="2009" box="[123,344,608,632]" class="Arachnida" family="Buthidae" genus="Grosphus" kingdom="Animalia" order="Scorpiones" pageId="16" pageNumber="15" phylum="Arthropoda" rank="species" species="tavaratra">Grosphus tavaratra</taxonomicName>
|
||
<taxonomicName id="4C124D1CFFA1FFB7272AD3DBFE23FEFC" authorityName="Lourenco, Wilme & Waeber" authorityYear="2015" box="[123,401,674,698]" class="Arachnida" family="Buthidae" genus="Neogrosphus" kingdom="Animalia" order="Scorpiones" pageId="16" pageNumber="15" phylum="Arthropoda" rank="species" species="andrafiabe">Neogrosphus andrafiabe</taxonomicName>
|
||
<taxonomicName id="4C124D1CFFA1FFB7272AD39AFEEEFEBD" authorityName="Lourenco" authorityYear="1996" box="[123,348,739,763]" class="Arachnida" family="Buthidae" genus="Neogrosphus" kingdom="Animalia" order="Scorpiones" pageId="16" pageNumber="15" phylum="Arthropoda" rank="species" species="blanci">Neogrosphus blanci</taxonomicName>
|
||
</emphasis>
|
||
</th>
|
||
<td id="76F35DA1FFA1005826EFD018FD9BFEBA" box="[446,553,353,764]" gridcol="1" gridrow="1" pageId="16" pageNumber="15">internal? internal internal internal internal dorsal/ carinal dorsal</td>
|
||
<td id="76F35DA1FFA100582514D018FD39FEBA" box="[581,651,353,764]" gridcol="2" gridrow="1" pageId="16" pageNumber="15">> 0.5? ~ 0.5 <0.5> 0.5> 0.5 – –</td>
|
||
<td id="76F35DA1FFA100582592D018FCA5FEBA" box="[707,791,353,764]" gridcol="3" gridrow="1" pageId="16" pageNumber="15">17 17–18 – 18–19 18–19 22–23 27–28 27</td>
|
||
<td id="76F35DA1FFA100582432D018FC04FEBA" box="[867,950,353,764]" gridcol="4" gridrow="1" pageId="16" pageNumber="15">– – 17–18 – – – – –</td>
|
||
<td id="76F35DA1FFA1005824BCD018FB34FEBA" box="[1005,1158,353,764]" gridcol="5" gridrow="1" pageId="16" pageNumber="15">– – short, wide – – – – –</td>
|
||
<td id="76F35DA1FFA1005823E2D018FAA8FEBA" box="[1203,1306,353,764]" gridcol="6" gridrow="1" pageId="16" pageNumber="15">elliptic – – – – elliptic – –</td>
|
||
<td id="76F35DA1FFA10058226ED018FA01FEBA" box="[1343,1459,353,764]" gridcol="7" gridrow="1" pageId="16" pageNumber="15">sparse – – – sparse sparse – –</td>
|
||
</tr>
|
||
<tr id="352234DDFFA10058272AD26FFA01FF69" box="[123,1459,790,815]" gridrow="2" pageId="16" pageNumber="15">
|
||
<th id="76F35DA1FFA10058272AD26FFE23FF69" box="[123,401,790,815]" gridcol="0" gridrow="2" pageId="16" pageNumber="15">
|
||
<taxonomicName id="4C124D1CFFA1FFB7272AD26FFEEFFF68" authorityName="Lowe & Kovařík" authorityYear="2019" baseAuthorityName="Lourenco, Wilme & Waeber" baseAuthorityYear="2018" box="[123,349,790,814]" class="Arachnida" family="Buthidae" genus="Teruelius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="16" pageNumber="15" phylum="Arthropoda" rank="species" species="bemaraha">
|
||
<emphasis id="B966EA8DFFA1FFB7272AD26FFEEFFF68" box="[123,349,790,814]" italics="true" pageId="16" pageNumber="15">Teruelius bemaraha</emphasis>
|
||
</taxonomicName>
|
||
</th>
|
||
<td id="76F35DA1FFA1005826EFD26FFD9BFF69" box="[446,553,790,815]" gridcol="1" gridrow="2" pageId="16" pageNumber="15">carinal</td>
|
||
<td id="76F35DA1FFA100582514D26FFD39FF69" box="[581,651,790,815]" gridcol="2" gridrow="2" pageId="16" pageNumber="15">?</td>
|
||
<td id="76F35DA1FFA100582592D26FFCA5FF69" box="[707,791,790,815]" gridcol="3" gridrow="2" pageId="16" pageNumber="15">29–30</td>
|
||
<td id="76F35DA1FFA100582432D26FFC04FF69" box="[867,950,790,815]" gridcol="4" gridrow="2" pageId="16" pageNumber="15">–</td>
|
||
<td id="76F35DA1FFA1005824BCD26FFB34FF69" box="[1005,1158,790,815]" gridcol="5" gridrow="2" pageId="16" pageNumber="15">–</td>
|
||
<td id="76F35DA1FFA1005823E2D26FFAA8FF69" box="[1203,1306,790,815]" gridcol="6" gridrow="2" pageId="16" pageNumber="15">narrow</td>
|
||
<td id="76F35DA1FFA10058226ED26FFA01FF69" box="[1343,1459,790,815]" gridcol="7" gridrow="2" pageId="16" pageNumber="15">dense</td>
|
||
</tr>
|
||
<tr id="352234DDFFA10058272AD22FFA01FF28" box="[123,1459,854,878]" gridrow="3" pageId="16" pageNumber="15">
|
||
<th id="76F35DA1FFA10058272AD22FFE23FF28" box="[123,401,854,878]" gridcol="0" gridrow="3" pageId="16" pageNumber="15">
|
||
<taxonomicName id="4C124D1CFFA1FFB7272AD22FFE8EFF28" authorityName="Lowe & Kovařík" authorityYear="2019" baseAuthorityName="Lourenco" baseAuthorityYear="2012" box="[123,316,854,878]" class="Arachnida" family="Buthidae" genus="Teruelius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="16" pageNumber="15" phylum="Arthropoda" rank="species" species="bicolor">
|
||
<emphasis id="B966EA8DFFA1FFB7272AD22FFE8EFF28" box="[123,316,854,878]" italics="true" pageId="16" pageNumber="15">Teruelius bicolor</emphasis>
|
||
</taxonomicName>
|
||
</th>
|
||
<td id="76F35DA1FFA1005826EFD22FFD9BFF28" box="[446,553,854,878]" gridcol="1" gridrow="3" pageId="16" pageNumber="15">–</td>
|
||
<td id="76F35DA1FFA100582514D22FFD39FF28" box="[581,651,854,878]" gridcol="2" gridrow="3" pageId="16" pageNumber="15">?</td>
|
||
<td id="76F35DA1FFA100582592D22FFCA5FF28" box="[707,791,854,878]" gridcol="3" gridrow="3" pageId="16" pageNumber="15">36</td>
|
||
<td id="76F35DA1FFA100582432D22FFC04FF28" box="[867,950,854,878]" gridcol="4" gridrow="3" pageId="16" pageNumber="15">–</td>
|
||
<td id="76F35DA1FFA1005824BCD22FFB34FF28" box="[1005,1158,854,878]" gridcol="5" gridrow="3" pageId="16" pageNumber="15">–</td>
|
||
<td id="76F35DA1FFA1005823E2D22FFAA8FF28" box="[1203,1306,854,878]" gridcol="6" gridrow="3" pageId="16" pageNumber="15">–</td>
|
||
<td id="76F35DA1FFA10058226ED22FFA01FF28" box="[1343,1459,854,878]" gridcol="7" gridrow="3" pageId="16" pageNumber="15">–</td>
|
||
</tr>
|
||
<tr id="352234DDFFA10058272AD2DBFA01FFFC" box="[123,1459,930,954]" gridrow="4" pageId="16" pageNumber="15">
|
||
<th id="76F35DA1FFA10058272AD2DBFE23FFFC" box="[123,401,930,954]" gridcol="0" gridrow="4" pageId="16" pageNumber="15">
|
||
<taxonomicName id="4C124D1CFFA1FFB7272AD2DBFEDDFFFC" authorityName="Lowe & Kovařík" authorityYear="2019" baseAuthorityName="Lourenco & Wilme" baseAuthorityYear="2016" box="[123,367,930,954]" class="Arachnida" family="Buthidae" genus="Teruelius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="16" pageNumber="15" phylum="Arthropoda" rank="species" species="eliseanneae">
|
||
<emphasis id="B966EA8DFFA1FFB7272AD2DBFEDDFFFC" box="[123,367,930,954]" italics="true" pageId="16" pageNumber="15">Teruelius eliseanneae</emphasis>
|
||
</taxonomicName>
|
||
</th>
|
||
<td id="76F35DA1FFA1005826EFD2DBFD9BFFFC" box="[446,553,930,954]" gridcol="1" gridrow="4" pageId="16" pageNumber="15">internal</td>
|
||
<td id="76F35DA1FFA100582514D2DBFD39FFFC" box="[581,651,930,954]" gridcol="2" gridrow="4" pageId="16" pageNumber="15"><0.5</td>
|
||
<td id="76F35DA1FFA100582592D2DBFCA5FFFC" box="[707,791,930,954]" gridcol="3" gridrow="4" pageId="16" pageNumber="15">–</td>
|
||
<td id="76F35DA1FFA100582432D2DBFC04FFFC" box="[867,950,930,954]" gridcol="4" gridrow="4" pageId="16" pageNumber="15">24</td>
|
||
<td id="76F35DA1FFA1005824BCD2DBFB34FFFC" box="[1005,1158,930,954]" gridcol="5" gridrow="4" pageId="16" pageNumber="15">long, narrow</td>
|
||
<td id="76F35DA1FFA1005823E2D2DBFAA8FFFC" box="[1203,1306,930,954]" gridcol="6" gridrow="4" pageId="16" pageNumber="15">–</td>
|
||
<td id="76F35DA1FFA10058226ED2DBFA01FFFC" box="[1343,1459,930,954]" gridcol="7" gridrow="4" pageId="16" pageNumber="15">–</td>
|
||
</tr>
|
||
<tr id="352234DDFFA10058272AD298FA01FFBF" box="[123,1459,993,1017]" gridrow="5" pageId="16" pageNumber="15">
|
||
<th id="76F35DA1FFA10058272AD298FE23FFBF" box="[123,401,993,1017]" gridcol="0" gridrow="5" pageId="16" pageNumber="15">
|
||
<taxonomicName id="4C124D1CFFA1FFB7272AD298FEEFFFBF" authorityName="Lowe & Kovařík" authorityYear="2019" baseAuthorityName="Lourenco, Wilme & Waeber" baseAuthorityYear="2016" box="[123,349,993,1017]" class="Arachnida" family="Buthidae" genus="Teruelius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="16" pageNumber="15" phylum="Arthropoda" rank="species" species="ganzhorni">
|
||
<emphasis id="B966EA8DFFA1FFB7272AD298FEEFFFBF" box="[123,349,993,1017]" italics="true" pageId="16" pageNumber="15">Teruelius ganzhorni</emphasis>
|
||
</taxonomicName>
|
||
</th>
|
||
<td id="76F35DA1FFA1005826EFD298FD9BFFBF" box="[446,553,993,1017]" gridcol="1" gridrow="5" pageId="16" pageNumber="15">dorsal</td>
|
||
<td id="76F35DA1FFA100582514D298FD39FFBF" box="[581,651,993,1017]" gridcol="2" gridrow="5" pageId="16" pageNumber="15">> 0.5?</td>
|
||
<td id="76F35DA1FFA100582592D298FCA5FFBF" box="[707,791,993,1017]" gridcol="3" gridrow="5" pageId="16" pageNumber="15">–</td>
|
||
<td id="76F35DA1FFA100582432D298FC04FFBF" box="[867,950,993,1017]" gridcol="4" gridrow="5" pageId="16" pageNumber="15">25–28</td>
|
||
<td id="76F35DA1FFA1005824BCD298FB34FFBF" box="[1005,1158,993,1017]" gridcol="5" gridrow="5" pageId="16" pageNumber="15">long, narrow</td>
|
||
<td id="76F35DA1FFA1005823E2D298FAA8FFBF" box="[1203,1306,993,1017]" gridcol="6" gridrow="5" pageId="16" pageNumber="15">narrow</td>
|
||
<td id="76F35DA1FFA10058226ED298FA01FFBF" box="[1343,1459,993,1017]" gridcol="7" gridrow="5" pageId="16" pageNumber="15">dense</td>
|
||
</tr>
|
||
<tr id="352234DDFFA10058272AD55AFA01F87D" box="[123,1459,1059,1083]" gridrow="6" pageId="16" pageNumber="15">
|
||
<th id="76F35DA1FFA10058272AD55AFE23F87D" box="[123,401,1059,1083]" gridcol="0" gridrow="6" pageId="16" pageNumber="15">
|
||
<taxonomicName id="4C124D1CFFA1FFB7272AD55AFED2F87D" authorityName="Lowe & Kovařík" authorityYear="2019" baseAuthorityName="Lourenco" baseAuthorityYear="2014" box="[123,352,1059,1083]" class="Arachnida" family="Buthidae" genus="Teruelius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="16" pageNumber="15" phylum="Arthropoda" rank="species" species="magalieae">
|
||
<emphasis id="B966EA8DFFA1FFB7272AD55AFED2F87D" box="[123,352,1059,1083]" italics="true" pageId="16" pageNumber="15">Teruelius magalieae</emphasis>
|
||
</taxonomicName>
|
||
</th>
|
||
<td id="76F35DA1FFA1005826EFD55AFD9BF87D" box="[446,553,1059,1083]" gridcol="1" gridrow="6" pageId="16" pageNumber="15">dorsal</td>
|
||
<td id="76F35DA1FFA100582514D55AFD39F87D" box="[581,651,1059,1083]" gridcol="2" gridrow="6" pageId="16" pageNumber="15"><0.5</td>
|
||
<td id="76F35DA1FFA100582592D55AFCA5F87D" box="[707,791,1059,1083]" gridcol="3" gridrow="6" pageId="16" pageNumber="15">36</td>
|
||
<td id="76F35DA1FFA100582432D55AFC04F87D" box="[867,950,1059,1083]" gridcol="4" gridrow="6" pageId="16" pageNumber="15">–</td>
|
||
<td id="76F35DA1FFA1005824BCD55AFB34F87D" box="[1005,1158,1059,1083]" gridcol="5" gridrow="6" pageId="16" pageNumber="15">–</td>
|
||
<td id="76F35DA1FFA1005823E2D55AFAA8F87D" box="[1203,1306,1059,1083]" gridcol="6" gridrow="6" pageId="16" pageNumber="15">narrow</td>
|
||
<td id="76F35DA1FFA10058226ED55AFA01F87D" box="[1343,1459,1059,1083]" gridcol="7" gridrow="6" pageId="16" pageNumber="15">dense</td>
|
||
</tr>
|
||
<tr id="352234DDFFA10058272AD50AFA01F8CD" box="[123,1459,1139,1163]" gridrow="7" pageId="16" pageNumber="15">
|
||
<th id="76F35DA1FFA10058272AD50AFE23F8CD" box="[123,401,1139,1163]" gridcol="0" gridrow="7" pageId="16" pageNumber="15">
|
||
<taxonomicName id="4C124D1CFFA1FFB7272AD50AFEFDF8CD" authorityName="Lowe & Kovařík" authorityYear="2019" baseAuthorityName="Lourenco & Wilme" baseAuthorityYear="2016" box="[123,335,1139,1163]" class="Arachnida" family="Buthidae" genus="Teruelius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="16" pageNumber="15" phylum="Arthropoda" rank="species" species="sabineae">
|
||
<emphasis id="B966EA8DFFA1FFB7272AD50AFEFDF8CD" box="[123,335,1139,1163]" italics="true" pageId="16" pageNumber="15">Teruelius sabineae</emphasis>
|
||
</taxonomicName>
|
||
</th>
|
||
<td id="76F35DA1FFA1005826EFD50AFD9BF8CD" box="[446,553,1139,1163]" gridcol="1" gridrow="7" pageId="16" pageNumber="15">dorsal</td>
|
||
<td id="76F35DA1FFA100582514D50AFD39F8CD" box="[581,651,1139,1163]" gridcol="2" gridrow="7" pageId="16" pageNumber="15"><0.5</td>
|
||
<td id="76F35DA1FFA100582592D50AFCA5F8CD" box="[707,791,1139,1163]" gridcol="3" gridrow="7" pageId="16" pageNumber="15">–</td>
|
||
<td id="76F35DA1FFA100582432D50AFC04F8CD" box="[867,950,1139,1163]" gridcol="4" gridrow="7" pageId="16" pageNumber="15">25</td>
|
||
<td id="76F35DA1FFA1005824BCD50AFB34F8CD" box="[1005,1158,1139,1163]" gridcol="5" gridrow="7" pageId="16" pageNumber="15">long, narrow</td>
|
||
<td id="76F35DA1FFA1005823E2D50AFAA8F8CD" box="[1203,1306,1139,1163]" gridcol="6" gridrow="7" pageId="16" pageNumber="15">narrow</td>
|
||
<td id="76F35DA1FFA10058226ED50AFA01F8CD" box="[1343,1459,1139,1163]" gridcol="7" gridrow="7" pageId="16" pageNumber="15">dense</td>
|
||
</tr>
|
||
<tr id="352234DDFFA10058272AD5C5FA01F892" box="[123,1459,1212,1236]" gridrow="8" pageId="16" pageNumber="15">
|
||
<th id="76F35DA1FFA10058272AD5C5FE23F892" box="[123,401,1212,1236]" gridcol="0" gridrow="8" pageId="16" pageNumber="15">
|
||
<taxonomicName id="4C124D1CFFA1FFB7272AD5C5FEF4F892" authorityName="Lowe & Kovařík" authorityYear="2019" baseAuthorityName="Lourenco & Wilme" baseAuthorityYear="2016" box="[123,326,1212,1236]" class="Arachnida" family="Buthidae" genus="Teruelius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="16" pageNumber="15" phylum="Arthropoda" rank="species" species="waeberi">
|
||
<emphasis id="B966EA8DFFA1FFB7272AD5C5FEF4F892" box="[123,326,1212,1236]" italics="true" pageId="16" pageNumber="15">Teruelius waeberi</emphasis>
|
||
</taxonomicName>
|
||
</th>
|
||
<td id="76F35DA1FFA1005826EFD5C5FD9BF892" box="[446,553,1212,1236]" gridcol="1" gridrow="8" pageId="16" pageNumber="15">dorsal</td>
|
||
<td id="76F35DA1FFA100582514D5C5FD39F892" box="[581,651,1212,1236]" gridcol="2" gridrow="8" pageId="16" pageNumber="15">~ 0.5</td>
|
||
<td id="76F35DA1FFA100582592D5C5FCA5F892" box="[707,791,1212,1236]" gridcol="3" gridrow="8" pageId="16" pageNumber="15">26–30</td>
|
||
<td id="76F35DA1FFA100582432D5C5FC04F892" box="[867,950,1212,1236]" gridcol="4" gridrow="8" pageId="16" pageNumber="15">–</td>
|
||
<td id="76F35DA1FFA1005824BCD5C5FB34F892" box="[1005,1158,1212,1236]" gridcol="5" gridrow="8" pageId="16" pageNumber="15">–</td>
|
||
<td id="76F35DA1FFA1005823E2D5C5FAA8F892" box="[1203,1306,1212,1236]" gridcol="6" gridrow="8" pageId="16" pageNumber="15">narrow</td>
|
||
<td id="76F35DA1FFA10058226ED5C5FA01F892" box="[1343,1459,1212,1236]" gridcol="7" gridrow="8" pageId="16" pageNumber="15">dense</td>
|
||
</tr>
|
||
</table>
|
||
</paragraph>
|
||
<caption id="DF6D6617FFA1FFB7273FD584FB00F9E5" ID-Table-UUID="DF6D6617FFA1FFB7273FD584FB00F9E5" httpUri="http://table.plazi.org/id/DF6D6617FFA1FFB7273FD584FB00F9E5" pageId="16" pageNumber="15" startId="16.[110,166,1277,1298]" targetBox="[123,1459,264,1236]" targetIsTable="true" targetPageId="16">
|
||
<paragraph id="8BAD369FFFA1FFB7273FD584FB00F9E5" blockId="16.[110,1471,1276,1443]" pageId="16" pageNumber="15">
|
||
<emphasis id="B966EA8DFFA1FFB7273FD584FF0AF957" bold="true" box="[110,184,1276,1298]" pageId="16" pageNumber="15">Table 3</emphasis>
|
||
.
|
||
<emphasis id="B966EA8DFFA1FFB72795D585FDDFF954" bold="true" box="[196,621,1276,1298]" pageId="16" pageNumber="15">Character states for species not examined</emphasis>
|
||
. States of 6 characters for
|
||
<taxonomicName id="4C124D1CFFA1FFB7243DD584FC7AF954" authorityName="Vachon" authorityYear="1940" box="[876,968,1277,1298]" class="Arachnida" family="Buthidae" genus="Grosphus" kingdom="Animalia" order="Scorpiones" pageId="16" pageNumber="15" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B966EA8DFFA1FFB7243DD584FC7AF954" box="[876,968,1277,1298]" italics="true" pageId="16" pageNumber="15">Grosphus</emphasis>
|
||
</taxonomicName>
|
||
(6 spp., blue panels),
|
||
<taxonomicName id="4C124D1CFFA1FFB723CDD584FAA8F954" authorityName="Lourenco" authorityYear="1995" box="[1180,1306,1277,1298]" class="Arachnida" family="Buthidae" genus="Neogrosphus" kingdom="Animalia" order="Scorpiones" pageId="16" pageNumber="15" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B966EA8DFFA1FFB723CDD584FAA8F954" box="[1180,1306,1277,1298]" italics="true" pageId="16" pageNumber="15">Neogrosphus</emphasis>
|
||
</taxonomicName>
|
||
(2 spp., magenta panels) and
|
||
<taxonomicName id="4C124D1CFFA1FFB727B3D460FE88F968" authority="Lowe & Kovařík, 2019" authorityName="Lowe & Kovařík" authorityYear="2019" box="[226,314,1305,1326]" class="Arachnida" family="Buthidae" genus="Teruelius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="16" pageNumber="15" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="B966EA8DFFA1FFB727B3D460FE88F968" box="[226,314,1305,1326]" italics="true" pageId="16" pageNumber="15">Teruelius</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A25557F6FFA1FFB72610D460FE37F969" box="[321,389,1305,1327]" pageId="16" pageNumber="15" rank="genus">
|
||
<emphasis id="B966EA8DFFA1FFB72610D460FED7F968" bold="true" box="[321,357,1305,1326]" pageId="16" pageNumber="15">gen</emphasis>
|
||
.
|
||
<emphasis id="B966EA8DFFA1FFB72623D460FECDF968" bold="true" box="[370,383,1305,1326]" pageId="16" pageNumber="15">n</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
(7 spp., yellow panels) not examined in this study: position of trichobothrium
|
||
<emphasis id="B966EA8DFFA1FFB7232CD460FB3BF968" box="[1149,1161,1305,1326]" italics="true" pageId="16" pageNumber="15">d</emphasis>
|
||
on pedipalp femur; ratio
|
||
<emphasis id="B966EA8DFFA1FFB722D6D463FA24F969" box="[1415,1430,1306,1327]" italics="true" pageId="16" pageNumber="15">R</emphasis>
|
||
= 2 123 d(
|
||
<emphasis id="B966EA8DFFA1FFB727D3D44FFF2FF90D" box="[130,157,1334,1355]" italics="true" pageId="16" pageNumber="15">Eb</emphasis>
|
||
<subScript id="179634DAFFA1FFB727CDD43CFF11F917" attach="left" box="[156,163,1349,1361]" fontSize="5" pageId="16" pageNumber="15">2</subScript>
|
||
,
|
||
<emphasis id="B966EA8DFFA1FFB727E1D44FFF79F90D" box="[176,203,1334,1355]" italics="true" pageId="16" pageNumber="15">Eb</emphasis>
|
||
<subScript id="179634DAFFA1FFB7279AD43CFF60F917" attach="left" box="[203,210,1349,1361]" fontSize="5" pageId="16" pageNumber="15">3</subScript>
|
||
)/ d(
|
||
<emphasis id="B966EA8DFFA1FFB727AAD44FFEA4F90D" box="[251,278,1334,1355]" italics="true" pageId="16" pageNumber="15">Eb</emphasis>
|
||
<subScript id="179634DAFFA1FFB72644D43CFEAEF917" attach="left" box="[277,284,1349,1361]" fontSize="5" pageId="16" pageNumber="15">1</subScript>
|
||
,
|
||
<emphasis id="B966EA8DFFA1FFB72678D44FFEF6F90D" box="[297,324,1334,1355]" italics="true" pageId="16" pageNumber="15">Eb</emphasis>
|
||
<subScript id="179634DAFFA1FFB72615D43CFEF9F917" attach="left" box="[324,331,1349,1361]" fontSize="5" pageId="16" pageNumber="15">2</subScript>
|
||
) of distances between pedipalp manus trichobothria
|
||
<emphasis id="B966EA8DFFA1FFB7241BD44FFCD7F90D" box="[842,869,1334,1355]" italics="true" pageId="16" pageNumber="15">Eb</emphasis>
|
||
<subScript id="179634DAFFA1FFB72434D43CFCDEF917" attach="left" box="[869,876,1349,1361]" fontSize="5" pageId="16" pageNumber="15">1</subScript>
|
||
,
|
||
<emphasis id="B966EA8DFFA1FFB72428D44FFC26F90D" box="[889,916,1334,1355]" italics="true" pageId="16" pageNumber="15">Eb</emphasis>
|
||
<subScript id="179634DAFFA1FFB724C2D43CFC28F917" attach="left" box="[915,922,1349,1361]" fontSize="5" pageId="16" pageNumber="15">2</subScript>
|
||
and
|
||
<emphasis id="B966EA8DFFA1FFB7249BD44FFC57F90D" box="[970,997,1334,1355]" italics="true" pageId="16" pageNumber="15">Eb</emphasis>
|
||
<subScript id="179634DAFFA1FFB724B4D43CFC5EF917" attach="left" box="[997,1004,1349,1361]" fontSize="5" pageId="16" pageNumber="15">3</subScript>
|
||
; ranges of pectinal tooth counts (PTC) of males (♂) and females (♀);size and shape of female basal pectinal tooth; shape of spiracles; and telotarsal setation. Character states inferred from published descriptions and figures. PTC and basal pectinal tooth characters not indicated in cases where either males or females are not known or described. Other characters not indicated in cases where published descriptions did not yield clear information.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BAD369FFFA1FFB627F0D757FC2FFA48" blockId="16.[113,772,1486,2022]" lastBlockId="17.[816,1474,1429,1550]" lastPageId="17" lastPageNumber="16" pageId="16" pageNumber="15">
|
||
The FMNH label site is ca.
|
||
<quantity id="4CEA9B7AFFA1FFB7268ED757FD81FA00" box="[479,563,1582,1606]" metricMagnitude="5" metricUnit="m" metricValue="1.45" pageId="16" pageNumber="15" unit="km" value="145.0">145 km</quantity>
|
||
south and slightly west of the type locality of
|
||
<taxonomicName id="4C124D1CFFA1FFB726E0D737FE92FAC0" authority="(Lourenco & Wilme, 2015 b)" baseAuthorityName="Lourenco & Wilme" baseAuthorityYear="2015" class="Arachnida" family="Buthidae" genus="Grosphus" kingdom="Animalia" order="Scorpiones" pageId="16" pageNumber="15" phylum="Arthropoda" rank="species" species="makay">
|
||
<emphasis id="B966EA8DFFA1FFB726E0D737FDDBFA20" box="[433,617,1614,1638]" italics="true" pageId="16" pageNumber="15">Grosphus makay</emphasis>
|
||
(
|
||
<bibRefCitation id="EF834B6EFFA1FFB7252AD737FEAAFAC0" author="LOURENCO, W. R. & L. WILME" journalOrPublisher="Revista Iberica de Arachnologia" pageId="16" pageNumber="15" pagination="55 - 61" part="26" refId="ref40226" refString="LOURENCO, W. R. & L. WILME. 2015 b. Scorpions collected in the Makay mountain range, Madagascar (Scorpiones: Hormuridae, Buthidae) and with description of a new species. Revista Iberica de Arachnologia, 26: 55 - 61." title="Scorpions collected in the Makay mountain range, Madagascar (Scorpiones: Hormuridae, Buthidae) and with description of a new species" type="journal article" year="2015">Lourenço & Wilmé, 2015b</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
: “Region Atsimo-Andrefana, ex Province of
|
||
<collectingRegion id="49D6F87DFFA1FFB727C2D7F7FF53FAE0" box="[147,225,1678,1702]" country="Madagascar" name="Toliara" pageId="16" pageNumber="15">Toliara</collectingRegion>
|
||
,
|
||
<taxonomicName id="4C124D1CFFA1FFB727A0D7F7FE6DFAE0" authority="Mts., General" authorityName="Mts., General" box="[241,479,1678,1702]" class="Arachnida" family="Buthidae" genus="Grosphus" kingdom="Animalia" order="Scorpiones" pageId="16" pageNumber="15" phylum="Arthropoda" rank="species" species="makay">Makay Mts., General</taxonomicName>
|
||
Collection, dry-Forest on sandy soil,
|
||
<date id="FFAC105FFFA1FFB727ADD7D7FEC7FA80" box="[252,373,1710,1734]" pageId="16" pageNumber="15" value="2010-03-12">12/III/2010</date>
|
||
(B. L. Fisher et al.). BLF25549. Female
|
||
<typeStatus id="54A9883DFFA1FFB72797D7B7FE91FAA0" box="[198,291,1742,1766]" pageId="16" pageNumber="15" type="holotype">holotype</typeStatus>
|
||
(CAS)”. These two localities have sandstone substrates, similar elevation and are located in the same general bioclimatic region. Comparison of the published habitus of the
|
||
<typeStatus id="54A9883DFFA1FFB727ECD657FEA8FB00" box="[189,282,1838,1862]" pageId="16" pageNumber="15" type="holotype">holotype</typeStatus>
|
||
adult female of
|
||
<taxonomicName id="4C124D1CFFA1FFB72688D657FD8DFB00" box="[473,575,1838,1862]" class="Arachnida" family="Buthidae" genus="Grosphus" kingdom="Animalia" order="Scorpiones" pageId="16" pageNumber="15" phylum="Arthropoda" rank="species" species="makay">
|
||
<emphasis id="B966EA8DFFA1FFB72688D657FE5AFB03" box="[473,488,1838,1861]" italics="true" pageId="16" pageNumber="15">T</emphasis>
|
||
.
|
||
<emphasis id="B966EA8DFFA1FFB726A8D657FD8DFB00" box="[505,575,1838,1862]" italics="true" pageId="16" pageNumber="15">makay</emphasis>
|
||
</taxonomicName>
|
||
and the
|
||
<typeStatus id="54A9883DFFA1FFB725F7D657FCB1FB00" box="[678,771,1838,1862]" pageId="16" pageNumber="15" type="holotype">holotype</typeStatus>
|
||
juvenile male of
|
||
<taxonomicName id="4C124D1CFFA1FFB72677D637FED6FB20" authorityName="Lourenco" authorityYear="1996" box="[294,356,1870,1894]" class="Arachnida" family="Buthidae" genus="Grosphus" kingdom="Animalia" order="Scorpiones" pageId="16" pageNumber="15" phylum="Arthropoda" rank="species" species="feti">
|
||
<emphasis id="B966EA8DFFA1FFB72677D637FE87FB23" box="[294,309,1870,1893]" italics="true" pageId="16" pageNumber="15">T</emphasis>
|
||
.
|
||
<emphasis id="B966EA8DFFA1FFB72613D637FED6FB20" box="[322,356,1870,1894]" italics="true" pageId="16" pageNumber="15">feti</emphasis>
|
||
</taxonomicName>
|
||
revealed very similar morphology and morphometrics even though sex and age differ. Most notably, their color patterns are identical in all details including: pattern of fuscosity on interocular triangle of carapace with pale cut-out behind lateral eyes; thin median line, precise fuscous banding patterns and transverse lateral striping on all tergites; darkly marked ventrolateral and ventromedian carinae on metasoma I–IV; fuscous patterns on metasoma V and telson; short fuscous strip on interno-proximal margin of pedipalp patella; leg femora with distal short, pale cut-outs on distal dark areas of prolateral surfaces and pale narrow lines on dorsal margins; and leg patellae with fuscous ventral margins on prolateral surfaces (compare
|
||
<figureCitation id="13292A1AFFA1FFB723B4D7D7FA8CFA80" box="[1253,1342,1710,1734]" captionStart="Figure 459" captionStartId="97.[113,181,1982,2004]" captionTargetBox="[113,1474,234,1946]" captionTargetId="figure-36@97.[113,1474,234,1949]" captionTargetPageId="97" captionText="Figure 459. Teruelius feti. Holotype juvenile male. Habitus, dorsal view, with specimen labels (FMNH). Scale bar: 5 mm (holotype). Figure 459a. Adult female from Makay Mts., carapace and tergites I–II." figureDoi="http://doi.org/10.5281/zenodo.7143796" httpUri="https://zenodo.org/record/7143796/files/figure.png" pageId="16" pageNumber="15">Fig. 459</figureCitation>
|
||
to fig. 13 of
|
||
<bibRefCitation id="EF834B6EFFA1FFB72461D7B7FBE9FAA0" author="LOURENCO, W. R. & L. WILME" box="[816,1115,1742,1766]" journalOrPublisher="Revista Iberica de Arachnologia" pageId="16" pageNumber="15" pagination="55 - 61" part="26" refId="ref40226" refString="LOURENCO, W. R. & L. WILME. 2015 b. Scorpions collected in the Makay mountain range, Madagascar (Scorpiones: Hormuridae, Buthidae) and with description of a new species. Revista Iberica de Arachnologia, 26: 55 - 61." title="Scorpions collected in the Makay mountain range, Madagascar (Scorpiones: Hormuridae, Buthidae) and with description of a new species" type="journal article" year="2015">Lourenço & Wilmé, 2015b</bibRefCitation>
|
||
). These very particular details of pigmentation pattern are not found in
|
||
<taxonomicName id="4C124D1CFFA1FFB7239FD797FAF5FB40" authorityName="Lowe & Kovařík" authorityYear="2019" baseAuthorityName="Pocock" baseAuthorityYear="1889" box="[1230,1351,1774,1798]" class="Arachnida" family="Buthidae" genus="Teruelius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="16" pageNumber="15" phylum="Arthropoda" rank="species" species="limbatus">
|
||
<emphasis id="B966EA8DFFA1FFB7239FD797FB6FFB43" box="[1230,1245,1774,1797]" italics="true" pageId="16" pageNumber="15">T</emphasis>
|
||
.
|
||
<emphasis id="B966EA8DFFA1FFB723BAD797FAF5FB40" box="[1259,1351,1774,1798]" italics="true" pageId="16" pageNumber="15">limbatus</emphasis>
|
||
</taxonomicName>
|
||
which was regarded as a closely related species (
|
||
<figureCitation id="13292A1AFFA1FFB7238AD677FAF4FB60" box="[1243,1350,1806,1830]" captionStart="Figures 5–8" captionStartId="7.[113,190,1980,2002]" captionTargetBox="[138,1459,268,1917]" captionTargetId="figure-41@7.[113,1476,244,1929]" captionTargetPageId="7" captionText="Figures 5–8. Teruelius limbatus. Type species of Teruelius gen. n. Habitus. Male (5–6) and female (7–8), dorsal (5, 7) and ventral (6, 8) views. Scale bars: 10 mm." figureDoi="http://doi.org/10.5281/zenodo.7143614" httpUri="https://zenodo.org/record/7143614/files/figure.png" pageId="16" pageNumber="15">Figs. 5–8</figureCitation>
|
||
,
|
||
<figureCitation id="13292A1AFFA1FFB72204D677FA05FB60" box="[1365,1463,1806,1830]" captionStart="Figures 516–521" captionStartId="105.[113,190,1958,1980]" captionTargetBox="[146,1426,242,1905]" captionTargetId="figure-51@105.[136,1452,232,1918]" captionTargetPageId="105" captionText="Figures 516–521. Teruelius limbatus. Figures 516–519. Habitus. Male (516–517) and female (518–519), dorsal (516, 518) and ventral (517, 519) views. Figure 520. Pedipalp chela external, male. Figure 521. Pedipalp chela, movable finger dentate margin, female. Scale bar: 10 mm (516–519)." figureDoi="http://doi.org/10.5281/zenodo.7143815" httpUri="https://zenodo.org/record/7143815/files/figure.png" pageId="16" pageNumber="15">516–521</figureCitation>
|
||
). We also examined
|
||
<specimenCount id="9D14FD16FFA1FFB72357D657FBDFFB00" box="[1030,1133,1838,1862]" pageId="16" pageNumber="15" type="adult">two adult</specimenCount>
|
||
females, near topotypes of
|
||
<taxonomicName id="4C124D1CFFA1FFB722F9D657FCC4FB20" class="Arachnida" family="Buthidae" genus="Grosphus" kingdom="Animalia" order="Scorpiones" pageId="16" pageNumber="15" phylum="Arthropoda" rank="species" species="makay">
|
||
<emphasis id="B966EA8DFFA1FFB722F9D657FA09FB03" box="[1448,1467,1838,1861]" italics="true" pageId="16" pageNumber="15">G</emphasis>
|
||
.
|
||
<emphasis id="B966EA8DFFA1FFB72461D637FCC4FB20" box="[816,886,1870,1894]" italics="true" pageId="16" pageNumber="15">makay</emphasis>
|
||
</taxonomicName>
|
||
from the
|
||
<taxonomicName id="4C124D1CFFA1FFB724B8D637FB01FB20" authority="Mountains" authorityName="Mountains" box="[1001,1203,1870,1894]" class="Arachnida" family="Buthidae" genus="Grosphus" kingdom="Animalia" order="Scorpiones" pageId="16" pageNumber="15" phylum="Arthropoda" rank="species" species="makay">Makay Mountains</taxonomicName>
|
||
, that exhibited the same coloration patterns and morphometrics as
|
||
<taxonomicName id="4C124D1CFFA1FFB723ABD617FA88FBC0" authorityName="Lourenco" authorityYear="1996" box="[1274,1338,1902,1926]" class="Arachnida" family="Buthidae" genus="Grosphus" kingdom="Animalia" order="Scorpiones" pageId="16" pageNumber="15" phylum="Arthropoda" rank="species" species="feti">
|
||
<emphasis id="B966EA8DFFA1FFB723ABD617FABBFBC3" box="[1274,1289,1902,1925]" italics="true" pageId="16" pageNumber="15">T</emphasis>
|
||
.
|
||
<emphasis id="B966EA8DFFA1FFB72249D617FA88FBC0" box="[1304,1338,1902,1926]" italics="true" pageId="16" pageNumber="15">feti</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="13292A1AFFA1FFB7221AD617FA05FBC0" box="[1355,1463,1902,1926]" captionStart="Figure 459" captionStartId="97.[113,181,1982,2004]" captionTargetBox="[113,1474,234,1946]" captionTargetId="figure-36@97.[113,1474,234,1949]" captionTargetPageId="97" captionText="Figure 459. Teruelius feti. Holotype juvenile male. Habitus, dorsal view, with specimen labels (FMNH). Scale bar: 5 mm (holotype). Figure 459a. Adult female from Makay Mts., carapace and tergites I–II." figureDoi="http://doi.org/10.5281/zenodo.7143796" httpUri="https://zenodo.org/record/7143796/files/figure.png" pageId="16" pageNumber="15">Fig. 459a</figureCitation>
|
||
). We allowed for the fact that juveniles of pigmented scorpions usually display darker, more intense markings, and that in adults these color patterns are somewhat faded. Details of coloration pattern have been given high priority as characters for species-level taxonomy of
|
||
<taxonomicName id="4C124D1CFFA0FFB62680D4CFFD8AF988" authorityName="Vachon" authorityYear="1940" box="[465,568,1462,1486]" class="Arachnida" family="Buthidae" genus="Grosphus" kingdom="Animalia" order="Scorpiones" pageId="17" pageNumber="16" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B966EA8DFFA0FFB62680D4CFFD8AF988" box="[465,568,1462,1486]" italics="true" pageId="17" pageNumber="16">Grosphus</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<bibRefCitation id="EF834B6EFFA0FFB6251DD4CFFF0AF9A8" author="LOURENCO, W. R. & V. SOARIMALALA & S. M. GOODMAN" journalOrPublisher="Malagasy Nature" pageId="17" pageNumber="16" pagination="144 - 153" part="2" refId="ref40004" refString="LOURENCO, W. R., V. SOARIMALALA & S. M. GOODMAN. 2009 b. The species of Grosphus Simon (Scorpiones, Buthidae) distributed in the northern and eastern regions of Madagascar with the description of a new species. Malagasy Nature, 2: 144 - 153." title="The species of Grosphus Simon (Scorpiones, Buthidae) distributed in the northern and eastern regions of Madagascar with the description of a new species" type="journal article" year="2009">Lourenço et al., 2009b</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EF834B6EFFA0FFB62798D4AFFECCF9A8" author="LOURENCO, W. R." box="[201,382,1494,1518]" journalOrPublisher="Zoosystema" pageId="17" pageNumber="16" pagination="631 - 645" part="36" refId="ref38903" refString="LOURENCO, W. R. 2014. The genus Grosphus Simon, 1880 in South-Western Madagascar, with the description of a new species (Scorpiones, Buthidae). Zoosystema, 36 (3): 631 - 645." title="The genus Grosphus Simon, 1880 in South-Western Madagascar, with the description of a new species (Scorpiones, Buthidae)" type="journal article" year="2014">Lourenço, 2014</bibRefCitation>
|
||
). We therefore consider
|
||
<taxonomicName id="4C124D1CFFA0FFB625CDD4AFFF05FA48" class="Arachnida" family="Buthidae" genus="Grosphus" kingdom="Animalia" order="Scorpiones" pageId="17" pageNumber="16" phylum="Arthropoda" rank="species" species="makay">
|
||
<emphasis id="B966EA8DFFA0FFB625CDD4AFFF05FA48" italics="true" pageId="17" pageNumber="16">Grosphus makay</emphasis>
|
||
</taxonomicName>
|
||
to be a junior synonym of
|
||
<taxonomicName id="4C124D1CFFA0FFB6268ED48FFDACFA48" authorityName="Lourenco" authorityYear="1996" box="[479,542,1526,1550]" class="Arachnida" family="Buthidae" genus="Grosphus" kingdom="Animalia" order="Scorpiones" pageId="17" pageNumber="16" phylum="Arthropoda" rank="species" species="feti">
|
||
<emphasis id="B966EA8DFFA0FFB6268ED48FFE5CFA4B" box="[479,494,1526,1549]" italics="true" pageId="17" pageNumber="16">T</emphasis>
|
||
.
|
||
<emphasis id="B966EA8DFFA0FFB626ADD48FFDACFA48" box="[508,542,1526,1550]" italics="true" pageId="17" pageNumber="16">feti</emphasis>
|
||
</taxonomicName>
|
||
, and the correct type locality of the latter to be that indicated on FMNH data labels.
|
||
<taxonomicName id="4C124D1CFFA0FFB62720D74FFF01FA08" authorityName="Lourenco" authorityYear="1996" box="[113,179,1590,1614]" class="Arachnida" family="Buthidae" genus="Grosphus" kingdom="Animalia" order="Scorpiones" pageId="17" pageNumber="16" phylum="Arthropoda" rank="species" species="feti">
|
||
<emphasis id="B966EA8DFFA0FFB62720D74FFF32FA0B" box="[113,128,1590,1613]" italics="true" pageId="17" pageNumber="16">T</emphasis>
|
||
.
|
||
<emphasis id="B966EA8DFFA0FFB627C0D74FFF01FA08" box="[145,179,1590,1614]" italics="true" pageId="17" pageNumber="16">feti</emphasis>
|
||
</taxonomicName>
|
||
was never again collected from Cap Sainte Marie in over two decades of fieldwork since its description, although other scorpion species (e.g.,
|
||
<taxonomicName id="4C124D1CFFA0FFB626E8D70FFD88FAC8" authorityName="Lowe & Kovařík" authorityYear="2019" baseAuthorityName="Lourenco & Wilme" baseAuthorityYear="2016" box="[441,570,1654,1678]" class="Arachnida" family="Buthidae" genus="Teruelius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="17" pageNumber="16" phylum="Arthropoda" rank="species" species="sabineae">
|
||
<emphasis id="B966EA8DFFA0FFB626E8D70FFE7AFACB" box="[441,456,1654,1677]" italics="true" pageId="17" pageNumber="16">T</emphasis>
|
||
.
|
||
<emphasis id="B966EA8DFFA0FFB6268AD70FFD88FAC8" box="[475,570,1654,1678]" italics="true" pageId="17" pageNumber="16">sabineae</emphasis>
|
||
</taxonomicName>
|
||
) were discovered there. The adult male remains unknown. Our opinion could be verified by collection and analysis of topotypic adult and juvenile specimens from the FMNH locality. The description of
|
||
<taxonomicName id="4C124D1CFFA0FFB627C0D78FFE62FB48" authority="(Lourenco, 1996 b: 14)" baseAuthorityName="Lourenco" baseAuthorityPageNumber="14" baseAuthorityYear="1996" box="[145,464,1782,1806]" class="Arachnida" family="Buthidae" genus="Grosphus" kingdom="Animalia" order="Scorpiones" pageId="17" pageNumber="16" phylum="Arthropoda" rank="species" species="feti">
|
||
<emphasis id="B966EA8DFFA0FFB627C0D78FFF12FB4B" box="[145,160,1782,1805]" italics="true" pageId="17" pageNumber="16">T</emphasis>
|
||
.
|
||
<emphasis id="B966EA8DFFA0FFB627FED78FFF63FB48" box="[175,209,1782,1806]" italics="true" pageId="17" pageNumber="16">feti</emphasis>
|
||
(
|
||
<bibRefCitation id="EF834B6EFFA0FFB627B3D78FFE7AFB48" author="LOURENCO, W. R." bookContentInfo="102 pp." box="[226,456,1782,1806]" journalOrPublisher="Museum National d'Histoire Naturelle, Paris" pageId="17" pageNumber="16" refId="ref37632" refString="LOURENCO, W. R. 1996 b. Faune de Madagascar. 87. Scorpions (Chelicerata, Scorpiones). Museum National d'Histoire Naturelle, Paris. 102 pp." title="Faune de Madagascar. 87. Scorpions (Chelicerata, Scorpiones)" type="book" year="1996">Lourenço, 1996b: 14</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
noted the juvenile status of the
|
||
<typeStatus id="54A9883DFFA0FFB627CAD66FFF4AFB68" box="[155,248,1814,1838]" pageId="17" pageNumber="16" type="holotype">holotype</typeStatus>
|
||
which is ca.
|
||
<quantity id="4CEA9B7AFFA0FFB626DCD66FFE6FFB6B" box="[397,477,1814,1838]" metricMagnitude="-2" metricUnit="m" metricValue="3.0" pageId="17" pageNumber="16" unit="mm" value="30.0">30 mm</quantity>
|
||
, but
|
||
<bibRefCitation id="EF834B6EFFA0FFB62544D66FFCB1FB68" author="LOURENCO, W. R." box="[533,771,1814,1838]" journalOrPublisher="Zoosystema" pageId="17" pageNumber="16" pagination="631 - 645" part="36" refId="ref38903" refString="LOURENCO, W. R. 2014. The genus Grosphus Simon, 1880 in South-Western Madagascar, with the description of a new species (Scorpiones, Buthidae). Zoosystema, 36 (3): 631 - 645." title="The genus Grosphus Simon, 1880 in South-Western Madagascar, with the description of a new species (Scorpiones, Buthidae)" type="journal article" year="2014">Lourenço (2014: 636)</bibRefCitation>
|
||
diagnosed the species as “of small size with a total length of
|
||
<quantity id="4CEA9B7AFFA0FFB627C0D62FFE97FB28" box="[145,293,1878,1902]" metricMagnitude="-2" metricUnit="m" metricValue="3.5" metricValueMax="4.0" metricValueMin="3.0" pageId="17" pageNumber="16" unit="mm" value="35.0" valueMax="40.0" valueMin="30.0">30 to 40 mm</quantity>
|
||
”. Small size cannot be a species diagnostic character if it is a property of the juvenile, and adult males are probably medium-sized, comparable to an adult female of
|
||
<taxonomicName id="4C124D1CFFA0FFB625B8D6EFFF05FB88" class="Arachnida" family="Buthidae" genus="Grosphus" kingdom="Animalia" order="Scorpiones" pageId="17" pageNumber="16" phylum="Arthropoda" rank="species" species="makay">
|
||
<emphasis id="B966EA8DFFA0FFB625B8D6EFFD4EFBEB" box="[745,764,1942,1965]" italics="true" pageId="17" pageNumber="16">G</emphasis>
|
||
.
|
||
<emphasis id="B966EA8DFFA0FFB62720D6CFFF05FB88" box="[113,183,1974,1998]" italics="true" pageId="17" pageNumber="16">makay</emphasis>
|
||
</taxonomicName>
|
||
, ca.
|
||
<quantity id="4CEA9B7AFFA0FFB627B9D6CCFE80FB8B" box="[232,306,1973,1997]" metricMagnitude="-2" metricUnit="m" metricValue="5.6" pageId="17" pageNumber="16" unit="mm" value="56.0">56 mm</quantity>
|
||
in body length. Our synonymy also implies that this is not a microendemic species of the upper Central Menabe, as suggested of
|
||
<bibRefCitation id="EF834B6EFFA0FFB62310D4EFFAC3F9EB" author="LOURENCO, W. R. & L. WILME" box="[1089,1393,1429,1454]" journalOrPublisher="Revista Iberica de Arachnologia" pageId="17" pageNumber="16" pagination="55 - 61" part="26" refId="ref40226" refString="LOURENCO, W. R. & L. WILME. 2015 b. Scorpions collected in the Makay mountain range, Madagascar (Scorpiones: Hormuridae, Buthidae) and with description of a new species. Revista Iberica de Arachnologia, 26: 55 - 61." title="Scorpions collected in the Makay mountain range, Madagascar (Scorpiones: Hormuridae, Buthidae) and with description of a new species" type="journal article" year="2015">Lourenço & Wilmé (2015b)</bibRefCitation>
|
||
. The
|
||
<taxonomicName id="4C124D1CFFA0FFB622FCD4EFFCE0F988" baseAuthorityName="Lourenco" baseAuthorityYear="1996" class="Arachnida" family="Buthidae" genus="Grosphus" kingdom="Animalia" order="Scorpiones" pageId="17" pageNumber="16" phylum="Arthropoda" rank="species" species="feti">
|
||
<emphasis id="B966EA8DFFA0FFB622FCD4EFFA0EF9EB" box="[1453,1468,1430,1453]" italics="true" pageId="17" pageNumber="16">T</emphasis>
|
||
.
|
||
<emphasis id="B966EA8DFFA0FFB62461D4CFFCE0F988" box="[816,850,1462,1486]" italics="true" pageId="17" pageNumber="16">feti</emphasis>
|
||
</taxonomicName>
|
||
emended type locality lies a significant distance south of the
|
||
<taxonomicName id="4C124D1CFFA0FFB6240BD4AFFBADF9A8" authority="mountains" authorityName="Mountains" box="[858,1055,1494,1518]" class="Arachnida" family="Buthidae" genus="Grosphus" kingdom="Animalia" order="Scorpiones" pageId="17" pageNumber="15" phylum="Arthropoda" rank="species" species="makay">Makay mountains</taxonomicName>
|
||
, in the Mangoky watershed (
|
||
<bibRefCitation id="EF834B6EFFA0FFB6220CD4AFFC20FA48" author="Wilme" etAl="et al." firstAuthor="Wilme" pageId="17" pageNumber="16" pagination="1063 - 1065" refId="ref44604" refString="WILME, L., S. M. GOODMAN & J. U. GANZHORN. 2006. Biogeographic evolution of Madagascar's microendemic biota. Science, 312: 1063 - 1065." type="journal article" year="2006">Wilmé et al., 2006</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BAD369FFFA0FFB627B8D06AFAF6F886" pageId="17" pageNumber="16">
|
||
<table id="F912C43FFFA0005827C4D06AFA20F886" box="[149,1426,275,1216]" gridcols="5" gridrows="12" pageId="17" pageNumber="16">
|
||
<tr id="352234DDFFA0005827C4D06AFA20FD68" box="[149,1426,275,302]" gridrow="0" pageId="17" pageNumber="16">
|
||
<th id="76F35DA1FFA0005827C4D06AFE11FD68" box="[149,419,275,302]" gridcol="0" gridrow="0" pageId="17" pageNumber="16">
|
||
<emphasis id="B966EA8DFFA0FFB627B8D06AFEFCFD6B" bold="true" box="[233,334,275,301]" pageId="17" pageNumber="16">GENUS</emphasis>
|
||
</th>
|
||
<th id="76F35DA1FFA00058268BD06AFD2CFD68" box="[474,670,275,302]" gridcol="1" gridrow="0" pageId="17" pageNumber="16">
|
||
<taxonomicName id="4C124D1CFFA0FFB626B9D06AFD3FFD68" authorityName="Kraepelin" authorityYear="1911" box="[488,653,275,302]" class="Arachnida" family="Buthidae" genus="Pseudolychas" kingdom="Animalia" order="Scorpiones" pageId="17" pageNumber="16" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B966EA8DFFA0FFB626B9D06AFD3FFD68" bold="true" box="[488,653,275,302]" italics="true" pageId="17" pageNumber="16">Pseudolychas</emphasis>
|
||
</taxonomicName>
|
||
</th>
|
||
<th id="76F35DA1FFA000582590D06AFC3EFD68" box="[705,908,275,302]" gridcol="2" gridrow="0" pageId="17" pageNumber="16">
|
||
<taxonomicName id="4C124D1CFFA0FFB625BBD06AFCD2FD68" authorityName="Vachon" authorityYear="1940" box="[746,864,275,302]" class="Arachnida" family="Buthidae" genus="Grosphus" kingdom="Animalia" order="Scorpiones" pageId="17" pageNumber="16" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B966EA8DFFA0FFB625BBD06AFCD2FD68" bold="true" box="[746,864,275,302]" italics="true" pageId="17" pageNumber="16">Grosphus</emphasis>
|
||
</taxonomicName>
|
||
</th>
|
||
<th id="76F35DA1FFA0005824F4D06AFBC2FD68" box="[933,1136,275,302]" gridcol="3" gridrow="0" pageId="17" pageNumber="16">
|
||
<taxonomicName id="4C124D1CFFA0FFB624E8D06AFBEBFD68" authorityName="Lourenco" authorityYear="1995" box="[953,1113,275,302]" class="Arachnida" family="Buthidae" genus="Neogrosphus" kingdom="Animalia" order="Scorpiones" pageId="17" pageNumber="16" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B966EA8DFFA0FFB624E8D06AFBEBFD68" bold="true" box="[953,1113,275,302]" italics="true" pageId="17" pageNumber="16">Neogrosphus</emphasis>
|
||
</taxonomicName>
|
||
</th>
|
||
<th id="76F35DA1FFA0005823FCD06AFA20FD68" box="[1197,1426,275,302]" gridcol="4" gridrow="0" pageId="17" pageNumber="16">
|
||
<emphasis id="B966EA8DFFA0FFB623E6D06AFA38FD68" bold="true" box="[1207,1418,275,302]" pageId="17" pageNumber="16">
|
||
<taxonomicName id="4C124D1CFFA0FFB623E6D06AFA98FD68" authority="Lowe & Kovařík, 2019" authorityName="Lowe & Kovařík" authorityYear="2019" box="[1207,1322,275,302]" class="Arachnida" family="Buthidae" genus="Teruelius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="17" pageNumber="16" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="B966EA8DFFA0FFB623E6D06AFA98FD68" bold="true" box="[1207,1322,275,302]" italics="true" pageId="17" pageNumber="16">Teruelius</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A25557F6FFA0FFB62266D06DFA38FD68" box="[1335,1418,276,302]" pageId="17" pageNumber="16" rank="genus">gen. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
</th>
|
||
</tr>
|
||
<tr id="352234DDFFA0005827C4D037FA20FDCB" box="[149,1426,334,397]" gridrow="1" pageId="17" pageNumber="16">
|
||
<th id="76F35DA1FFA0005827C4D037FE11FDCB" box="[149,419,334,397]" gridcol="0" gridrow="1" pageId="17" pageNumber="16">
|
||
<emphasis id="B966EA8DFFA0FFB627B2D026FEF7FD3C" bold="true" box="[227,325,351,378]" pageId="17" pageNumber="16">
|
||
femur
|
||
<emphasis id="B966EA8DFFA0FFB62667D026FEF7FD3C" bold="true" box="[310,325,351,378]" italics="true" pageId="17" pageNumber="16">d</emphasis>
|
||
</emphasis>
|
||
<emphasis id="B966EA8DFFA0FFB6261BD015FEE6FD3B" bold="true" box="[330,340,364,381]" pageId="17" pageNumber="16">2</emphasis>
|
||
</th>
|
||
<td id="76F35DA1FFA00058268BD037FD2CFDCB" box="[474,670,334,397]" gridcol="1" gridrow="1" pageId="17" pageNumber="16">dorsal</td>
|
||
<td id="76F35DA1FFA000582590D037FC3EFDCB" box="[705,908,334,397]" gridcol="2" gridrow="1" pageId="17" pageNumber="16">carinal/ internal</td>
|
||
<td id="76F35DA1FFA0005824F4D037FBC2FDCB" box="[933,1136,334,397]" gridcol="3" gridrow="1" pageId="17" pageNumber="16">internal/carinal/ external</td>
|
||
<td id="76F35DA1FFA0005823FCD037FA20FDCB" box="[1197,1426,334,397]" gridcol="4" gridrow="1" pageId="17" pageNumber="16">carinal/ dorsal</td>
|
||
</tr>
|
||
<tr id="352234DDFFA0005827C4D0CBFA20FD96" box="[149,1426,434,464]" gridrow="2" pageId="17" pageNumber="16">
|
||
<th id="76F35DA1FFA0005827C4D0CBFE11FD96" box="[149,419,434,464]" gridcol="0" gridrow="2" pageId="17" pageNumber="16">
|
||
<emphasis id="B966EA8DFFA0FFB62653D0CBFEA5FD8A" bold="true" box="[258,279,434,460]" pageId="17" pageNumber="16">R</emphasis>
|
||
<emphasis id="B966EA8DFFA0FFB62646D0C6FE87FD96" bold="true" box="[279,309,447,464]" pageId="17" pageNumber="16">123</emphasis>
|
||
</th>
|
||
<td id="76F35DA1FFA00058268BD0CBFD2CFD96" box="[474,670,434,464]" gridcol="1" gridrow="2" pageId="17" pageNumber="16">2/3 spp. ≥ 0.50</td>
|
||
<td id="76F35DA1FFA000582590D0CBFC3EFD96" box="[705,908,434,464]" gridcol="2" gridrow="2" pageId="17" pageNumber="16">0.32–1.04</td>
|
||
<td id="76F35DA1FFA0005824F4D0CBFBC2FD96" box="[933,1136,434,464]" gridcol="3" gridrow="2" pageId="17" pageNumber="16">0.54</td>
|
||
<td id="76F35DA1FFA0005823FCD0CBFA20FD96" box="[1197,1426,434,464]" gridcol="4" gridrow="2" pageId="17" pageNumber="16"><0.50</td>
|
||
</tr>
|
||
<tr id="352234DDFFA0005827C4D37AFA20FE66" box="[149,1426,515,544]" gridrow="3" pageId="17" pageNumber="16">
|
||
<th id="76F35DA1FFA0005827C4D37AFE11FE66" box="[149,419,515,544]" gridcol="0" gridrow="3" pageId="17" pageNumber="16">
|
||
<emphasis id="B966EA8DFFA0FFB627A1D37FFEFAFE66" bold="true" box="[240,328,515,544]" pageId="17" pageNumber="16">PTC ♂</emphasis>
|
||
</th>
|
||
<td id="76F35DA1FFA00058268BD37AFD2CFE66" box="[474,670,515,544]" gridcol="1" gridrow="3" pageId="17" pageNumber="16">11–17</td>
|
||
<td id="76F35DA1FFA000582590D37AFC3EFE66" box="[705,908,515,544]" gridcol="2" gridrow="3" pageId="17" pageNumber="16">15–23</td>
|
||
<td id="76F35DA1FFA0005824F4D37AFBC2FE66" box="[933,1136,515,544]" gridcol="3" gridrow="3" pageId="17" pageNumber="16">27–31</td>
|
||
<td id="76F35DA1FFA0005823FCD37AFA20FE66" box="[1197,1426,515,544]" gridcol="4" gridrow="3" pageId="17" pageNumber="16">25–41</td>
|
||
</tr>
|
||
<tr id="352234DDFFA0005827C4D32FFA20FE35" box="[149,1426,598,627]" gridrow="4" pageId="17" pageNumber="16">
|
||
<th id="76F35DA1FFA0005827C4D32FFE11FE35" box="[149,419,598,627]" gridcol="0" gridrow="4" pageId="17" pageNumber="16">
|
||
<emphasis id="B966EA8DFFA0FFB627A1D321FEFAFE35" bold="true" box="[240,328,598,627]" pageId="17" pageNumber="16">PTC ♀</emphasis>
|
||
</th>
|
||
<td id="76F35DA1FFA00058268BD32FFD2CFE35" box="[474,670,598,627]" gridcol="1" gridrow="4" pageId="17" pageNumber="16">9–14</td>
|
||
<td id="76F35DA1FFA000582590D32FFC3EFE35" box="[705,908,598,627]" gridcol="2" gridrow="4" pageId="17" pageNumber="16">12–19</td>
|
||
<td id="76F35DA1FFA0005824F4D32FFBC2FE35" box="[933,1136,598,627]" gridcol="3" gridrow="4" pageId="17" pageNumber="16">27–28</td>
|
||
<td id="76F35DA1FFA0005823FCD32FFA20FE35" box="[1197,1426,598,627]" gridcol="4" gridrow="4" pageId="17" pageNumber="16">24–35</td>
|
||
</tr>
|
||
<tr id="352234DDFFA0005827C4D3E1FA20FE9E" box="[149,1426,664,728]" gridrow="5" pageId="17" pageNumber="16">
|
||
<th id="76F35DA1FFA0005827C4D3E1FE11FE9E" box="[149,419,664,728]" gridcol="0" gridrow="5" pageId="17" pageNumber="16">
|
||
<emphasis id="B966EA8DFFA0FFB627C9D3D2FE12FE83" bold="true" box="[152,416,680,709]" pageId="17" pageNumber="16">basal pectine tooth ♀</emphasis>
|
||
</th>
|
||
<td id="76F35DA1FFA00058268BD3E1FD2CFE9E" box="[474,670,664,728]" gridcol="1" gridrow="5" pageId="17" pageNumber="16">short, ovoid (2/3 spp.)</td>
|
||
<td id="76F35DA1FFA000582590D3E1FC3EFE9E" box="[705,908,664,728]" gridcol="2" gridrow="5" pageId="17" pageNumber="16">short, ovoid</td>
|
||
<td id="76F35DA1FFA0005824F4D3E1FBC2FE9E" box="[933,1136,664,728]" gridcol="3" gridrow="5" pageId="17" pageNumber="16">short, ovoid</td>
|
||
<td id="76F35DA1FFA0005823FCD3E1FA20FE9E" box="[1197,1426,664,728]" gridcol="4" gridrow="5" pageId="17" pageNumber="16">elongate, falcate</td>
|
||
</tr>
|
||
<tr id="352234DDFFA0005827C4D394FA20FF6B" box="[149,1426,749,813]" gridrow="6" pageId="17" pageNumber="16">
|
||
<th id="76F35DA1FFA0005827C4D394FE11FF6B" box="[149,419,749,813]" gridcol="0" gridrow="6" pageId="17" pageNumber="16">
|
||
<emphasis id="B966EA8DFFA0FFB627CFD394FEE8FF6B" bold="true" box="[158,410,749,813]" pageId="17" pageNumber="16">hemispermatophore basal lobe</emphasis>
|
||
</th>
|
||
<td id="76F35DA1FFA00058268BD394FD2CFF6B" box="[474,670,749,813]" gridcol="1" gridrow="6" pageId="17" pageNumber="16">distal</td>
|
||
<td id="76F35DA1FFA000582590D394FC3EFF6B" box="[705,908,749,813]" gridcol="2" gridrow="6" pageId="17" pageNumber="16">distal/ proximal</td>
|
||
<td id="76F35DA1FFA0005824F4D394FBC2FF6B" box="[933,1136,749,813]" gridcol="3" gridrow="6" pageId="17" pageNumber="16">distal</td>
|
||
<td id="76F35DA1FFA0005823FCD394FA20FF6B" box="[1197,1426,749,813]" gridcol="4" gridrow="6" pageId="17" pageNumber="16">distal</td>
|
||
</tr>
|
||
<tr id="352234DDFFA0005827C4D23BFA20FFC4" box="[149,1426,834,898]" gridrow="7" pageId="17" pageNumber="16">
|
||
<th id="76F35DA1FFA0005827C4D23BFE11FFC4" box="[149,419,834,898]" gridcol="0" gridrow="7" pageId="17" pageNumber="16">
|
||
<emphasis id="B966EA8DFFA0FFB627CFD23BFEC0FFC4" bold="true" box="[158,410,834,898]" pageId="17" pageNumber="16">hemispermatophore posterior lobe</emphasis>
|
||
</th>
|
||
<td id="76F35DA1FFA00058268BD23BFD2CFFC4" box="[474,670,834,898]" gridcol="1" gridrow="7" pageId="17" pageNumber="16">long</td>
|
||
<td id="76F35DA1FFA000582590D23BFC3EFFC4" box="[705,908,834,898]" gridcol="2" gridrow="7" pageId="17" pageNumber="16">long</td>
|
||
<td id="76F35DA1FFA0005824F4D23BFBC2FFC4" box="[933,1136,834,898]" gridcol="3" gridrow="7" pageId="17" pageNumber="16">short</td>
|
||
<td id="76F35DA1FFA0005823FCD23BFA20FFC4" box="[1197,1426,834,898]" gridcol="4" gridrow="7" pageId="17" pageNumber="16">short</td>
|
||
</tr>
|
||
<tr id="352234DDFFA0005827C4D2DEFA20FF84" box="[149,1426,935,962]" gridrow="8" pageId="17" pageNumber="16">
|
||
<th id="76F35DA1FFA0005827C4D2DEFE11FF84" box="[149,419,935,962]" gridcol="0" gridrow="8" pageId="17" pageNumber="16">
|
||
<emphasis id="B966EA8DFFA0FFB627B4D2DEFEE0FF87" bold="true" box="[229,338,935,961]" pageId="17" pageNumber="16">spiracles</emphasis>
|
||
</th>
|
||
<td id="76F35DA1FFA00058268BD2DEFD2CFF84" box="[474,670,935,962]" gridcol="1" gridrow="8" pageId="17" pageNumber="16">elliptic, L/W <5</td>
|
||
<td id="76F35DA1FFA000582590D2DEFC3EFF84" box="[705,908,935,962]" gridcol="2" gridrow="8" pageId="17" pageNumber="16">elliptic, L/W <5</td>
|
||
<td id="76F35DA1FFA0005824F4D2DEFBC2FF84" box="[933,1136,935,962]" gridcol="3" gridrow="8" pageId="17" pageNumber="16">elliptic, L/W <5</td>
|
||
<td id="76F35DA1FFA0005823FCD2DEFA20FF84" box="[1197,1426,935,962]" gridcol="4" gridrow="8" pageId="17" pageNumber="16">narrow, L/W> 5</td>
|
||
</tr>
|
||
<tr id="352234DDFFA0005827C4D293FA20F86F" box="[149,1426,1002,1065]" gridrow="9" pageId="17" pageNumber="16">
|
||
<th id="76F35DA1FFA0005827C4D293FE11F86F" box="[149,419,1002,1065]" gridcol="0" gridrow="9" pageId="17" pageNumber="16">
|
||
<emphasis id="B966EA8DFFA0FFB627C4D284FE11F851" bold="true" box="[149,419,1021,1047]" pageId="17" pageNumber="16">met. I ventral carinae</emphasis>
|
||
</th>
|
||
<td id="76F35DA1FFA00058268BD293FD2CF86F" box="[474,670,1002,1065]" gridcol="1" gridrow="9" pageId="17" pageNumber="16">granulate</td>
|
||
<td id="76F35DA1FFA000582590D293FC3EF86F" box="[705,908,1002,1065]" gridcol="2" gridrow="9" pageId="17" pageNumber="16">crenulate/ granulate</td>
|
||
<td id="76F35DA1FFA0005824F4D293FBC2F86F" box="[933,1136,1002,1065]" gridcol="3" gridrow="9" pageId="17" pageNumber="16">granulate</td>
|
||
<td id="76F35DA1FFA0005823FCD293FA20F86F" box="[1197,1426,1002,1065]" gridcol="4" gridrow="9" pageId="17" pageNumber="16">weakly granulate to smooth</td>
|
||
</tr>
|
||
<tr id="352234DDFFA0005827C4D52AFA20F82B" box="[149,1426,1107,1133]" gridrow="10" pageId="17" pageNumber="16">
|
||
<th id="76F35DA1FFA0005827C4D52AFE11F82B" box="[149,419,1107,1133]" gridcol="0" gridrow="10" pageId="17" pageNumber="16">
|
||
<emphasis id="B966EA8DFFA0FFB627FBD52AFE3FF82B" bold="true" box="[170,397,1107,1133]" pageId="17" pageNumber="16">telotarsus setation</emphasis>
|
||
</th>
|
||
<td id="76F35DA1FFA00058268BD52AFD2CF82B" box="[474,670,1107,1133]" gridcol="1" gridrow="10" pageId="17" pageNumber="16">sparse, 2 rows</td>
|
||
<td id="76F35DA1FFA000582590D52AFC3EF82B" box="[705,908,1107,1133]" gridcol="2" gridrow="10" pageId="17" pageNumber="16">sparse, 2 rows</td>
|
||
<td id="76F35DA1FFA0005824F4D52AFBC2F82B" box="[933,1136,1107,1133]" gridcol="3" gridrow="10" pageId="17" pageNumber="16">dense, irregular</td>
|
||
<td id="76F35DA1FFA0005823FCD52AFA20F82B" box="[1197,1426,1107,1133]" gridcol="4" gridrow="10" pageId="17" pageNumber="16">dense, irregular</td>
|
||
</tr>
|
||
<tr id="352234DDFFA0005827C4D5DCFA20F886" box="[149,1426,1189,1216]" gridrow="11" pageId="17" pageNumber="16">
|
||
<th id="76F35DA1FFA0005827C4D5DCFE11F886" box="[149,419,1189,1216]" gridcol="0" gridrow="11" pageId="17" pageNumber="16">
|
||
<emphasis id="B966EA8DFFA0FFB627E4D5DFFE30F8F9" bold="true" box="[181,386,1189,1216]" pageId="17" pageNumber="16">UV fluorescence</emphasis>
|
||
</th>
|
||
<td id="76F35DA1FFA00058268BD5DCFD2CF886" box="[474,670,1189,1216]" gridcol="1" gridrow="11" pageId="17" pageNumber="16">weak</td>
|
||
<td id="76F35DA1FFA000582590D5DCFC3EF886" box="[705,908,1189,1216]" gridcol="2" gridrow="11" pageId="17" pageNumber="16">weak</td>
|
||
<td id="76F35DA1FFA0005824F4D5DCFBC2F886" box="[933,1136,1189,1216]" gridcol="3" gridrow="11" pageId="17" pageNumber="16">strong</td>
|
||
<td id="76F35DA1FFA0005823FCD5DCFA20F886" box="[1197,1426,1189,1216]" gridcol="4" gridrow="11" pageId="17" pageNumber="16">strong</td>
|
||
</tr>
|
||
</table>
|
||
</paragraph>
|
||
<caption id="DF6D6617FFA0FFB62720D58FFB7EF924" ID-Table-UUID="DF6D6617FFA0FFB62720D58FFB7EF924" httpUri="http://table.plazi.org/id/DF6D6617FFA0FFB62720D58FFB7EF924" pageId="17" pageNumber="16" startId="17.[113,169,1270,1291]" targetBox="[149,1426,275,1216]" targetIsTable="true" targetPageId="17">
|
||
<paragraph id="8BAD369FFFA0FFB62720D58FFB7EF924" blockId="17.[113,1474,1269,1378]" pageId="17" pageNumber="16">
|
||
<emphasis id="B966EA8DFFA0FFB62720D58FFF73F94C" bold="true" box="[113,193,1269,1291]" pageId="17" pageNumber="16">Table 4</emphasis>
|
||
.
|
||
<emphasis id="B966EA8DFFA0FFB62782D58CFB81F94D" bold="true" box="[211,1075,1269,1291]" pageId="17" pageNumber="16">
|
||
Character states for ‘
|
||
<taxonomicName id="4C124D1CFFA0FFB626EFD58CFDACF94D" authorityName="Vachon" authorityYear="1940" box="[446,542,1269,1291]" class="Arachnida" family="Buthidae" genus="Grosphus" kingdom="Animalia" order="Scorpiones" pageId="17" pageNumber="16" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B966EA8DFFA0FFB626EFD58CFDACF94D" bold="true" box="[446,542,1269,1291]" italics="true" pageId="17" pageNumber="16">Grosphus</emphasis>
|
||
</taxonomicName>
|
||
’ group genera and outgroup genus
|
||
<taxonomicName id="4C124D1CFFA0FFB624FDD58CFB81F94D" authorityName="Kraepelin" authorityYear="1911" box="[940,1075,1269,1291]" class="Arachnida" family="Buthidae" genus="Pseudolychas" kingdom="Animalia" order="Scorpiones" pageId="17" pageNumber="16" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B966EA8DFFA0FFB624FDD58CFB81F94D" bold="true" box="[940,1075,1269,1291]" italics="true" pageId="17" pageNumber="16">Pseudolychas</emphasis>
|
||
</taxonomicName>
|
||
</emphasis>
|
||
. States of 11 characters for
|
||
<taxonomicName id="4C124D1CFFA0FFB62231D58FFA0EF94D" authorityName="Vachon" authorityYear="1940" box="[1376,1468,1270,1291]" class="Arachnida" family="Buthidae" genus="Grosphus" kingdom="Animalia" order="Scorpiones" pageId="17" pageNumber="16" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B966EA8DFFA0FFB62231D58FFA0EF94D" box="[1376,1468,1270,1291]" italics="true" pageId="17" pageNumber="16">Grosphus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C124D1CFFA0FFB62720D46BFF5DF961" authorityName="Lourenco" authorityYear="1995" box="[113,239,1298,1319]" class="Arachnida" family="Buthidae" genus="Neogrosphus" kingdom="Animalia" order="Scorpiones" pageId="17" pageNumber="16" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B966EA8DFFA0FFB62720D46BFF5DF961" box="[113,239,1298,1319]" italics="true" pageId="17" pageNumber="16">Neogrosphus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C124D1CFFA0FFB627ACD46BFEE7F961" authority="Lowe & Kovařík, 2019" authorityName="Lowe & Kovařík" authorityYear="2019" box="[253,341,1298,1319]" class="Arachnida" family="Buthidae" genus="Teruelius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="17" pageNumber="16" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="B966EA8DFFA0FFB627ACD46BFEE7F961" box="[253,341,1298,1319]" italics="true" pageId="17" pageNumber="16">Teruelius</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A25557F6FFA0FFB6260FD46AFE16F96E" box="[350,420,1299,1320]" pageId="17" pageNumber="16" rank="genus">
|
||
<emphasis id="B966EA8DFFA0FFB6260FD46AFE30F96E" bold="true" box="[350,386,1299,1320]" pageId="17" pageNumber="16">gen</emphasis>
|
||
.
|
||
<emphasis id="B966EA8DFFA0FFB626C1D46AFE2FF96E" bold="true" box="[400,413,1299,1320]" pageId="17" pageNumber="16">n</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
and
|
||
<taxonomicName id="4C124D1CFFA0FFB62686D46BFDE9F961" authorityName="Kraepelin" authorityYear="1911" box="[471,603,1298,1319]" class="Arachnida" family="Buthidae" genus="Pseudolychas" kingdom="Animalia" order="Scorpiones" pageId="17" pageNumber="16" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B966EA8DFFA0FFB62686D46BFDE9F961" box="[471,603,1298,1319]" italics="true" pageId="17" pageNumber="16">Pseudolychas</emphasis>
|
||
</taxonomicName>
|
||
, summarizing data of Tables 1–3.
|
||
<taxonomicName id="4C124D1CFFA0FFB624E2D46BFB85F961" authorityName="Kraepelin" authorityYear="1911" box="[947,1079,1298,1319]" class="Arachnida" family="Buthidae" genus="Pseudolychas" kingdom="Animalia" order="Scorpiones" pageId="17" pageNumber="16" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B966EA8DFFA0FFB624E2D46BFB85F961" box="[947,1079,1298,1319]" italics="true" pageId="17" pageNumber="16">Pseudolychas</emphasis>
|
||
</taxonomicName>
|
||
character states determined by study of materials, and from published descriptions. Primitive states indicated by green panels, derived states by red panels, with polarities determined by comparison to
|
||
<taxonomicName id="4C124D1CFFA0FFB6264FD435FE10F927" authorityName="Kraepelin" authorityYear="1911" box="[286,418,1356,1377]" class="Arachnida" family="Buthidae" genus="Pseudolychas" kingdom="Animalia" order="Scorpiones" pageId="17" pageNumber="16" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B966EA8DFFA0FFB6264FD435FE10F927" box="[286,418,1356,1377]" italics="true" pageId="17" pageNumber="16">Pseudolychas</emphasis>
|
||
</taxonomicName>
|
||
which is assumed primitive. Unpolarized characters indicated by dark green panels.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BAD369FFFA0FFB62461D74FFA70FA2B" blockId="17.[816,1475,1590,2030]" pageId="17" pageNumber="16">
|
||
<taxonomicName id="4C124D1CFFA0FFB62461D74FFB2AFA08" authority="Lourenco, 2013" authorityName="Lourenco" authorityYear="2013" box="[816,1176,1590,1614]" class="Arachnida" family="Buthidae" genus="Grosphus" kingdom="Animalia" order="Scorpiones" pageId="17" pageNumber="16" phylum="Arthropoda" rank="species" species="rossii">
|
||
<emphasis id="B966EA8DFFA0FFB62461D74FFC6FFA0B" box="[816,989,1590,1614]" italics="true" pageId="17" pageNumber="16">Grosphus rossii</emphasis>
|
||
<bibRefCitation id="EF834B6EFFA0FFB624BBD74FFB2AFA08" author="LOURENCO, W. R." box="[1002,1176,1590,1614]" journalOrPublisher="Comptes Rendus Biologies" pageId="17" pageNumber="16" pagination="515 - 520" part="336" refId="ref38813" refString="LOURENCO, W. R. 2013 a. The remarkable peri-Saharan distribution of the genus Butheoloides Hirst (Scorpiones, Buthidae), with the description of a new species from Cameroon. Comptes Rendus Biologies, 336 (10): 515 - 520." title="The remarkable peri-Saharan distribution of the genus Butheoloides Hirst (Scorpiones, Buthidae), with the description of a new species from Cameroon" type="journal article" year="2013">Lourenço, 2013</bibRefCitation>
|
||
</taxonomicName>
|
||
=
|
||
<taxonomicName id="4C124D1CFFA0FFB623EED74FFAB6FA28" authority="(Lourenco, Goodman & Ramilijaona, 2004)" authorityName="Lowe & Kovařík" authorityYear="2019" baseAuthorityName="Lourenco, Goodman & Ramilijaona" baseAuthorityYear="2004" class="Arachnida" family="Buthidae" genus="Teruelius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="17" pageNumber="16" phylum="Arthropoda" rank="species" species="mahafaliensis" status="comb. n., syn. n.">
|
||
<emphasis id="B966EA8DFFA0FFB623EED74FFA70FA08" box="[1215,1474,1590,1614]" italics="true" pageId="17" pageNumber="16">Teruelius mahafaliensis</emphasis>
|
||
(
|
||
<bibRefCitation id="EF834B6EFFA0FFB62469D72FFB4CFA28" author="LOURENCO, W. R. & S. M. GOODMAN & O. RAMILIJAONA" box="[824,1278,1622,1646]" journalOrPublisher="Revista Iberica de Aracnologia" pageId="17" pageNumber="16" pagination="225 - 234" part="9" refId="ref39550" refString="LOURENCO, W. R., S. M. GOODMAN & O. RAMILIJAONA. 2004. Three new species of Grosphus Simon from Madagascar (Scorpiones, Buthidae). Revista Iberica de Aracnologia, 9: 225 - 234." title="Three new species of Grosphus Simon from Madagascar (Scorpiones, Buthidae)" type="journal article" year="2004">Lourenço, Goodman & Ramilijaona, 2004</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A25557F6FFA0FFB6225AD72FFA70FA2B" box="[1291,1474,1622,1646]" pageId="17" pageNumber="16">
|
||
<emphasis id="B966EA8DFFA0FFB6225AD72FFAFBFA28" bold="true" box="[1291,1353,1622,1646]" pageId="17" pageNumber="16">comb</emphasis>
|
||
.
|
||
<emphasis id="B966EA8DFFA0FFB62207D72FFAD7FA28" bold="true" box="[1366,1381,1622,1646]" pageId="17" pageNumber="16">n</emphasis>
|
||
.,
|
||
<emphasis id="B966EA8DFFA0FFB62228D72FFA12FA28" bold="true" box="[1401,1440,1622,1646]" pageId="17" pageNumber="16">syn</emphasis>
|
||
.
|
||
<emphasis id="B966EA8DFFA0FFB622FCD72FFA0EFA28" bold="true" box="[1453,1468,1622,1646]" pageId="17" pageNumber="16">n</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
</paragraph>
|
||
<paragraph id="8BAD369FFFA0FFB22431D70FFEF3FD24" blockId="17.[816,1475,1590,2030]" lastBlockId="21.[113,771,235,1027]" lastPageId="21" lastPageNumber="20" pageId="17" pageNumber="16">
|
||
<taxonomicName id="4C124D1CFFA0FFB62431D70FFBB4FACB" authorityName="Lourenco" authorityYear="2013" box="[864,1030,1654,1678]" class="Arachnida" family="Buthidae" genus="Grosphus" kingdom="Animalia" order="Scorpiones" pageId="17" pageNumber="16" phylum="Arthropoda" rank="species" species="rossii">
|
||
<emphasis id="B966EA8DFFA0FFB62431D70FFBB4FACB" box="[864,1030,1654,1678]" italics="true" pageId="17" pageNumber="16">Grosphus rossii</emphasis>
|
||
</taxonomicName>
|
||
was described by
|
||
<bibRefCitation id="EF834B6EFFA0FFB62399D70FFA3BFAC8" author="LOURENCO, W. R." box="[1224,1417,1654,1678]" journalOrPublisher="Entomologische Mitteilungen aus dem Zoologischen Museum Hamburg" pageId="17" pageNumber="16" pagination="57 - 62" part="16" refId="ref38860" refString="LOURENCO, W. R. 2013 b. A new species of Grosphus Simon, 1880 (Scorpiones, Buthidae) from Central Madagascar. Entomologische Mitteilungen aus dem Zoologischen Museum Hamburg, 16 (189): 57 - 62." title="A new species of Grosphus Simon, 1880 (Scorpiones, Buthidae) from Central Madagascar" type="journal article" year="2013">Lourenço (2013b)</bibRefCitation>
|
||
from a single adult male
|
||
<typeStatus id="54A9883DFFA0FFB62359D7EFFBD7FAE8" box="[1032,1125,1686,1710]" pageId="17" pageNumber="16" type="holotype">holotype</typeStatus>
|
||
collected from “
|
||
<collectingRegion id="49D6F87DFFA0FFB6224FD7EFFA0CFAE8" box="[1310,1470,1686,1710]" country="Ghana" name="Central" pageId="17" pageNumber="16">Central region</collectingRegion>
|
||
, NE Manandona, S of Antsirabe, in secondary growth forest, under log,
|
||
<date id="FFAC105FFFA0FFB624F6D7AFFBFBFAA8" box="[935,1097,1750,1774]" pageId="17" pageNumber="16" value="2004-08-08">8 August 2004</date>
|
||
, W. R Lourenço” and deposited in ZMUH. We loaned and studied the
|
||
<typeStatus id="54A9883DFFA0FFB623FCD78FFAB8FB48" box="[1197,1290,1782,1806]" pageId="17" pageNumber="16" type="holotype">holotype</typeStatus>
|
||
and found that it was virtually identical in coloration, external morphological characters and morphometrics to
|
||
<taxonomicName id="4C124D1CFFA0FFB623EAD64FFAA8FB28" authority="(Lourenco, Goodman & Ramilijaona, 2004)" authorityName="Lowe & Kovařík" authorityYear="2019" baseAuthorityName="Lourenco, Goodman & Ramilijaona" baseAuthorityYear="2004" class="Arachnida" family="Buthidae" genus="Teruelius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="17" pageNumber="16" phylum="Arthropoda" rank="species" species="mahafaliensis" status="comb. nov.">
|
||
<emphasis id="B966EA8DFFA0FFB623EAD64FFA70FB08" box="[1211,1474,1846,1870]" italics="true" pageId="17" pageNumber="16">Teruelius mahafaliensis</emphasis>
|
||
(
|
||
<bibRefCitation id="EF834B6EFFA0FFB62469D62FFAA1FB28" author="LOURENCO, W. R. & S. M. GOODMAN & O. RAMILIJAONA" box="[824,1299,1878,1902]" journalOrPublisher="Revista Iberica de Aracnologia" pageId="17" pageNumber="16" pagination="225 - 234" part="9" refId="ref39550" refString="LOURENCO, W. R., S. M. GOODMAN & O. RAMILIJAONA. 2004. Three new species of Grosphus Simon from Madagascar (Scorpiones, Buthidae). Revista Iberica de Aracnologia, 9: 225 - 234." title="Three new species of Grosphus Simon from Madagascar (Scorpiones, Buthidae)" type="journal article" year="2004">Lourenço, Goodman & Ramilijaona, 2004</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A25557F6FFA0FFB62277D62FFA3FFB2B" box="[1318,1421,1878,1902]" pageId="17" pageNumber="16" rank="species">
|
||
<emphasis id="B966EA8DFFA0FFB62277D62FFAD6FB28" bold="true" box="[1318,1380,1878,1902]" pageId="17" pageNumber="16">comb</emphasis>
|
||
.
|
||
<emphasis id="B966EA8DFFA0FFB62226D62FFA34FB28" bold="true" box="[1399,1414,1878,1902]" pageId="17" pageNumber="16">n</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
Our comparative materials of the latter included a male collected near the type locality of that species (
|
||
<figureCitation id="13292A1AFFA0FFB623B3D6EFFA36FBEB" box="[1250,1412,1941,1966]" captionStart="Figures 522–525" captionStartId="106.[113,190,1981,2003]" captionTargetBox="[150,1435,241,1931]" captionTargetId="figure-39@106.[127,1460,232,1934]" captionTargetPageId="106" captionText="Figures 522–525. Teruelius mahafaliensis. Habitus. Male holotype of G. rossii (522–523) and male from near the type locality of T. mahafaliensis (524–525), dorsal (522, 524) and ventral (523, 525) views. Scale bar: 10 mm." figureDoi="http://doi.org/10.5281/zenodo.7143817" httpUri="https://zenodo.org/record/7143817/files/figure.png" pageId="17" pageNumber="16">Figs. 522–525</figureCitation>
|
||
), and determined material loaned from FMNH including
|
||
<specimenCount id="9D14FD16FFA0FFB6220CD6CFFA70FB88" box="[1373,1474,1974,1998]" pageId="17" pageNumber="16" type="adult">two adult</specimenCount>
|
||
males. In his description,
|
||
<bibRefCitation id="EF834B6EFFA0FFB6236ED6AFFA95FBAB" author="LOURENCO, W. R." box="[1087,1319,2005,2030]" journalOrPublisher="Entomologische Mitteilungen aus dem Zoologischen Museum Hamburg" pageId="17" pageNumber="16" pagination="57 - 62" part="16" refId="ref38860" refString="LOURENCO, W. R. 2013 b. A new species of Grosphus Simon, 1880 (Scorpiones, Buthidae) from Central Madagascar. Entomologische Mitteilungen aus dem Zoologischen Museum Hamburg, 16 (189): 57 - 62." title="A new species of Grosphus Simon, 1880 (Scorpiones, Buthidae) from Central Madagascar" type="journal article" year="2013">Lourenço (2013b: 59)</bibRefCitation>
|
||
compared and contrasted
|
||
<taxonomicName id="4C124D1CFFA5FFB327B4D708FE8CFACE" authorityName="Lourenco" authorityYear="2013" box="[229,318,1649,1672]" class="Arachnida" family="Buthidae" genus="Grosphus" kingdom="Animalia" order="Scorpiones" pageId="20" pageNumber="19" phylum="Arthropoda" rank="species" species="rossii">
|
||
<emphasis id="B966EA8DFFA5FFB327B4D708FF4AFACE" box="[229,248,1649,1672]" italics="true" pageId="20" pageNumber="19">G</emphasis>
|
||
.
|
||
<emphasis id="B966EA8DFFA5FFB32655D708FE8CFACE" box="[260,318,1649,1672]" italics="true" pageId="20" pageNumber="19">rossii</emphasis>
|
||
</taxonomicName>
|
||
to
|
||
<taxonomicName id="4C124D1CFFA5FFB3260ED708FE64FACE" authorityName="Lowe & Kovařík" authorityYear="2019" baseAuthorityName="Pocock" baseAuthorityYear="1889" box="[351,470,1648,1672]" class="Arachnida" family="Buthidae" genus="Teruelius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="20" pageNumber="19" phylum="Arthropoda" rank="species" species="limbatus">
|
||
<emphasis id="B966EA8DFFA5FFB3260ED708FEDCFACE" box="[351,366,1649,1672]" italics="true" pageId="20" pageNumber="19">T</emphasis>
|
||
.
|
||
<emphasis id="B966EA8DFFA5FFB3262BD709FE64FACE" box="[378,470,1648,1672]" italics="true" pageId="20" pageNumber="19">limbatus</emphasis>
|
||
</taxonomicName>
|
||
, but not to
|
||
<taxonomicName id="4C124D1CFFA5FFB3251CD708FD4EFACE" authorityName="Lowe & Kovařík" authorityYear="2019" baseAuthorityName="Lourenco, Goodman & Ramilijaona" baseAuthorityYear="2004" box="[589,764,1648,1672]" class="Arachnida" family="Buthidae" genus="Teruelius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="20" pageNumber="19" phylum="Arthropoda" rank="species" species="mahafaliensis">
|
||
<emphasis id="B966EA8DFFA5FFB3251CD708FDEEFACE" box="[589,604,1649,1672]" italics="true" pageId="20" pageNumber="19">T</emphasis>
|
||
.
|
||
<emphasis id="B966EA8DFFA5FFB32539D709FD4EFACE" box="[616,764,1648,1672]" italics="true" pageId="20" pageNumber="19">mahafaliensis</emphasis>
|
||
</taxonomicName>
|
||
. A potential diagnostic difference is the pectinal tooth count (= 28) in
|
||
<taxonomicName id="4C124D1CFFA5FFB327EAD7C8FEA5FA8E" authorityName="Lourenco" authorityYear="2013" box="[187,279,1713,1736]" class="Arachnida" family="Buthidae" genus="Grosphus" kingdom="Animalia" order="Scorpiones" pageId="20" pageNumber="19" phylum="Arthropoda" rank="species" species="rossii">
|
||
<emphasis id="B966EA8DFFA5FFB327EAD7C8FF7CFA8E" box="[187,206,1713,1736]" italics="true" pageId="20" pageNumber="19">G</emphasis>
|
||
.
|
||
<emphasis id="B966EA8DFFA5FFB3278CD7C8FEA5FA8E" box="[221,279,1713,1736]" italics="true" pageId="20" pageNumber="19">rossii</emphasis>
|
||
</taxonomicName>
|
||
being lower than the range (35–40) reported for male
|
||
<taxonomicName id="4C124D1CFFA5FFB3279ED7A8FECCFAAE" authorityName="Lowe & Kovařík" authorityYear="2019" baseAuthorityName="Lourenco, Goodman & Ramilijaona" baseAuthorityYear="2004" box="[207,382,1744,1768]" class="Arachnida" family="Buthidae" genus="Teruelius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="20" pageNumber="19" phylum="Arthropoda" rank="species" species="mahafaliensis">
|
||
<emphasis id="B966EA8DFFA5FFB3279ED7A8FF6CFAAE" box="[207,222,1745,1768]" italics="true" pageId="20" pageNumber="19">T</emphasis>
|
||
.
|
||
<emphasis id="B966EA8DFFA5FFB327BBD7A9FECCFAAE" box="[234,382,1744,1768]" italics="true" pageId="20" pageNumber="19">mahafaliensis</emphasis>
|
||
</taxonomicName>
|
||
by Lourenço et al., (2007b: 373, tab. III). However, we examined a male
|
||
<taxonomicName id="4C124D1CFFA5FFB326BCD788FD2FFB4E" authorityName="Lowe & Kovařík" authorityYear="2019" baseAuthorityName="Lourenco, Goodman & Ramilijaona" baseAuthorityYear="2004" box="[493,669,1776,1800]" class="Arachnida" family="Buthidae" genus="Teruelius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="20" pageNumber="19" phylum="Arthropoda" rank="species" species="mahafaliensis">
|
||
<emphasis id="B966EA8DFFA5FFB326BCD788FE4EFB4E" box="[493,508,1777,1800]" italics="true" pageId="20" pageNumber="19">T</emphasis>
|
||
.
|
||
<emphasis id="B966EA8DFFA5FFB32558D789FD2FFB4E" box="[521,669,1776,1800]" italics="true" pageId="20" pageNumber="19">mahafaliensis</emphasis>
|
||
</taxonomicName>
|
||
collected near the species type locality with PTC of 29–33 (
|
||
<figureCitation id="13292A1AFFA5FFB325C0D669FF2FFB0E" captionStart="Figures 522–525" captionStartId="106.[113,190,1981,2003]" captionTargetBox="[150,1435,241,1931]" captionTargetId="figure-39@106.[127,1460,232,1934]" captionTargetPageId="106" captionText="Figures 522–525. Teruelius mahafaliensis. Habitus. Male holotype of G. rossii (522–523) and male from near the type locality of T. mahafaliensis (524–525), dorsal (522, 524) and ventral (523, 525) views. Scale bar: 10 mm." figureDoi="http://doi.org/10.5281/zenodo.7143817" httpUri="https://zenodo.org/record/7143817/files/figure.png" pageId="20" pageNumber="19">Figs. 524– 525</figureCitation>
|
||
). The type locality of
|
||
<taxonomicName id="4C124D1CFFA5FFB326C5D648FE43FB0E" authorityName="Lourenco" authorityYear="2013" box="[404,497,1841,1864]" class="Arachnida" family="Buthidae" genus="Grosphus" kingdom="Animalia" order="Scorpiones" pageId="20" pageNumber="19" phylum="Arthropoda" rank="species" species="rossii">
|
||
<emphasis id="B966EA8DFFA5FFB326C5D648FE15FB0E" box="[404,423,1841,1864]" italics="true" pageId="20" pageNumber="19">G</emphasis>
|
||
.
|
||
<emphasis id="B966EA8DFFA5FFB326E6D648FE43FB0E" box="[439,497,1841,1864]" italics="true" pageId="20" pageNumber="19">rossii</emphasis>
|
||
</taxonomicName>
|
||
on the central plateau at ca.
|
||
<quantity id="4CEA9B7AFFA5FFB327C9D629FF58FB2E" box="[152,234,1872,1896]" metricMagnitude="3" metricUnit="m" metricValue="1.4" pageId="20" pageNumber="19" unit="m" value="1400.0">1400 m</quantity>
|
||
a. s. l., is in a cooler, more humid zone, quite far from the other records of
|
||
<taxonomicName id="4C124D1CFFA5FFB326C6D608FDF9FBCE" authorityName="Lowe & Kovařík" authorityYear="2019" baseAuthorityName="Lourenco, Goodman & Ramilijaona" baseAuthorityYear="2004" box="[407,587,1904,1928]" class="Arachnida" family="Buthidae" genus="Teruelius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="20" pageNumber="19" phylum="Arthropoda" rank="species" species="mahafaliensis">
|
||
<emphasis id="B966EA8DFFA5FFB326C6D608FE14FBCE" box="[407,422,1905,1928]" italics="true" pageId="20" pageNumber="19">T</emphasis>
|
||
.
|
||
<emphasis id="B966EA8DFFA5FFB326E6D609FDF9FBCE" box="[439,587,1904,1928]" italics="true" pageId="20" pageNumber="19">mahafaliensis</emphasis>
|
||
</taxonomicName>
|
||
concentrated on the Mahafaly Plateau, a region of subarid thorn scrub along the southwest coast (ca.
|
||
<quantity id="4CEA9B7AFFA5FFB326D0D6C9FE75FB8E" box="[385,455,1968,1992]" metricMagnitude="2" metricUnit="m" metricValue="1.2" pageId="20" pageNumber="19" unit="m" value="120.0">120 m</quantity>
|
||
a.s.l.). It was suggested that
|
||
<taxonomicName id="4C124D1CFFA5FFB32720D6A8FF7DFBAE" authorityName="Lourenco" authorityYear="2013" box="[113,207,2001,2024]" class="Arachnida" family="Buthidae" genus="Grosphus" kingdom="Animalia" order="Scorpiones" pageId="20" pageNumber="19" phylum="Arthropoda" rank="species" species="rossii">
|
||
<emphasis id="B966EA8DFFA5FFB32720D6A8FF36FBAE" box="[113,132,2001,2024]" italics="true" pageId="20" pageNumber="19">G</emphasis>
|
||
.
|
||
<emphasis id="B966EA8DFFA5FFB327C4D6A8FF7DFBAE" box="[149,207,2001,2024]" italics="true" pageId="20" pageNumber="19">rossii</emphasis>
|
||
</taxonomicName>
|
||
was evidence of microendemism. We take a more conservative position and interpret the very close morphologies of
|
||
<taxonomicName id="4C124D1CFFA5FFB3241ED7E8FC19FAEE" authorityName="Lourenco" authorityYear="2013" box="[847,939,1681,1704]" class="Arachnida" family="Buthidae" genus="Grosphus" kingdom="Animalia" order="Scorpiones" pageId="20" pageNumber="19" phylum="Arthropoda" rank="species" species="rossii">
|
||
<emphasis id="B966EA8DFFA5FFB3241ED7E8FCD0FAEE" box="[847,866,1681,1704]" italics="true" pageId="20" pageNumber="19">G</emphasis>
|
||
.
|
||
<emphasis id="B966EA8DFFA5FFB32420D7E8FC19FAEE" box="[881,939,1681,1704]" italics="true" pageId="20" pageNumber="19">rossii</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C124D1CFFA5FFB324B3D7E8FB21FAEE" authorityName="Lowe & Kovařík" authorityYear="2019" baseAuthorityName="Lourenco, Goodman & Ramilijaona" baseAuthorityYear="2004" box="[994,1171,1680,1704]" class="Arachnida" family="Buthidae" genus="Teruelius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="20" pageNumber="19" phylum="Arthropoda" rank="species" species="mahafaliensis">
|
||
<emphasis id="B966EA8DFFA5FFB324B3D7E8FC43FAEE" box="[994,1009,1681,1704]" italics="true" pageId="20" pageNumber="19">T</emphasis>
|
||
.
|
||
<emphasis id="B966EA8DFFA5FFB324AED7E9FB21FAEE" box="[1023,1171,1680,1704]" italics="true" pageId="20" pageNumber="19">mahafaliensis</emphasis>
|
||
</taxonomicName>
|
||
as indicative of a eurytopic species with wider distribution. Broad elevation ranges of>
|
||
<quantity id="4CEA9B7AFFA5FFB32418D7A9FC2FFAAE" box="[841,925,1744,1768]" metricMagnitude="3" metricUnit="m" metricValue="1.4" pageId="20" pageNumber="19" unit="m" value="1400.0">1400 m</quantity>
|
||
are known for some widely distributed scorpions that inhabit varied bioclimatic zones (e.g.,
|
||
<taxonomicName id="4C124D1CFFA5FFB3226AD788FB32FB6E" authority="Soleglad & Fet, 2004" authorityName="Soleglad & Fet" authorityYear="2004" class="Arachnida" family="Chactidae" genus="Anuroctonus" kingdom="Animalia" order="Scorpiones" pageId="20" pageNumber="19" phylum="Arthropoda" rank="species" species="pococki">
|
||
<emphasis id="B966EA8DFFA5FFB3226AD788FC31FB6E" italics="true" pageId="20" pageNumber="19">Anuroctonus pococki</emphasis>
|
||
<bibRefCitation id="EF834B6EFFA5FFB324DFD669FB32FB6E" author="SOLEGLAD, M. E. & V. FET" box="[910,1152,1808,1832]" journalOrPublisher="Revista Iberica de Aracnologia" pageId="20" pageNumber="19" pagination="81 - 128" part="10" refId="ref43142" refString="SOLEGLAD, M. E. & V. FET. 2004. The systematics of the scorpion subfamily Uroctoninae (Scorpiones: Chactidae). Revista Iberica de Aracnologia, 10: 81 - 128." title="The systematics of the scorpion subfamily Uroctoninae (Scorpiones: Chactidae)" type="journal article" year="2004">Soleglad & Fet, 2004</bibRefCitation>
|
||
</taxonomicName>
|
||
, 300–
|
||
<quantity id="4CEA9B7AFFA5FFB32395D669FAAAFB6E" box="[1220,1304,1808,1832]" metricMagnitude="3" metricUnit="m" metricValue="1.85" pageId="20" pageNumber="19" unit="m" value="1850.0">1850 m</quantity>
|
||
a.s.l.,
|
||
<bibRefCitation id="EF834B6EFFA5FFB32234D669FC73FB0E" author="SOLEGLAD, M. E. & V. FET" journalOrPublisher="Revista Iberica de Aracnologia" pageId="20" pageNumber="19" pagination="81 - 128" part="10" refId="ref43142" refString="SOLEGLAD, M. E. & V. FET. 2004. The systematics of the scorpion subfamily Uroctoninae (Scorpiones: Chactidae). Revista Iberica de Aracnologia, 10: 81 - 128." title="The systematics of the scorpion subfamily Uroctoninae (Scorpiones: Chactidae)" type="journal article" year="2004">Soleglad & Fet, 2004</bibRefCitation>
|
||
;
|
||
<taxonomicName id="4C124D1CFFA5FFB32482D649FA3CFB0E" authority="Kraepelin, 1894" authorityName="Kraepelin" authorityYear="1894" box="[979,1422,1840,1864]" class="Arachnida" family="Bothriuridae" genus="Bothriurus" kingdom="Animalia" order="Scorpiones" pageId="20" pageNumber="19" phylum="Arthropoda" rank="species" species="burmeisteri">
|
||
<emphasis id="B966EA8DFFA5FFB32482D649FB7CFB0E" box="[979,1230,1840,1864]" italics="true" pageId="20" pageNumber="19">Bothriurus burmeisteri</emphasis>
|
||
Kraepelin, 1894
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C124D1CFFA5FFB32461D629FACCFB2E" authority="(Thorell, 1876)" baseAuthorityName="Thorell" baseAuthorityYear="1876" box="[816,1406,1872,1896]" class="Arachnida" family="Bothriuridae" genus="Brachistosternus" kingdom="Animalia" order="Scorpiones" pageId="20" pageNumber="19" phylum="Arthropoda" rank="species" species="weijenberghi">
|
||
<emphasis id="B966EA8DFFA5FFB32461D629FBCFFB2E" box="[816,1149,1872,1896]" italics="true" pageId="20" pageNumber="19">Brachistosternus weijenberghi</emphasis>
|
||
(Thorell, 1876),
|
||
<quantity id="4CEA9B7AFFA5FFB3221BD629FACCFB2E" box="[1354,1406,1872,1896]" metricMagnitude="3" metricUnit="m" metricValue="2.0" metricValueMax="3.0" metricValueMin="1.0" pageId="20" pageNumber="19" unit="m" value="2000.0" valueMax="3000.0" valueMin="1000.0">1000</quantity>
|
||
</taxonomicName>
|
||
–3000 m a.s.l.,
|
||
<bibRefCitation id="EF834B6EFFA5FFB324CCD609FB20FBCE" author="CAMPON, F. FERNANDEZ & S. LAGOS SILNIK & L. A. FEDELI" box="[925,1170,1904,1928]" journalOrPublisher="Journal of Arachnology" pageId="20" pageNumber="19" pagination="163 - 169" part="42" refId="ref32963" refString="CAMPON, F. FERNANDEZ, S. LAGOS SILNIK & L. A. FEDELI. 2014. Scorpion diversity of the Central Andes in Argentina. Journal of Arachnology, 42: 163 - 169." title="Scorpion diversity of the Central Andes in Argentina" type="journal article" year="2014">Campón et al., 2014</bibRefCitation>
|
||
;
|
||
<taxonomicName id="4C124D1CFFA5FFB323F7D609FC4EFBEE" authority="(Kraepelin, 1901)" baseAuthorityName="Kraepelin" baseAuthorityYear="1901" class="Arachnida" family="Buthidae" genus="Compsobuthus" kingdom="Animalia" order="Scorpiones" pageId="20" pageNumber="19" phylum="Arthropoda" rank="species" species="maindroni">
|
||
<emphasis id="B966EA8DFFA5FFB323F7D609FA70FBCE" box="[1190,1474,1904,1928]" italics="true" pageId="20" pageNumber="19">Compsobuthus maindroni</emphasis>
|
||
(Kraepelin, 1901)
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C124D1CFFA5FFB32345D6E8FA0CFBEE" authority="(Pocock, 1895)" baseAuthorityName="Pocock" baseAuthorityYear="1895" box="[1044,1470,1936,1960]" class="Arachnida" family="Buthidae" genus="Hottentotta" kingdom="Animalia" order="Scorpiones" pageId="20" pageNumber="19" phylum="Arthropoda" rank="species" species="jayakari">
|
||
<emphasis id="B966EA8DFFA5FFB32345D6E8FB4BFBEE" box="[1044,1273,1936,1960]" italics="true" pageId="20" pageNumber="19">Hottentotta jayakari</emphasis>
|
||
(Pocock, 1895)
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C124D1CFFA5FFB32461D6C9FB34FB8E" authority="Francke, 1980" authorityName="Francke" authorityYear="1980" box="[816,1158,1968,1992]" class="Arachnida" family="Diplocentridae" genus="Nebo" kingdom="Animalia" order="Scorpiones" pageId="20" pageNumber="19" phylum="Arthropoda" rank="species" species="omanensis">
|
||
<emphasis id="B966EA8DFFA5FFB32461D6C9FC51FB8E" box="[816,995,1968,1992]" italics="true" pageId="20" pageNumber="19">Nebo omanensis</emphasis>
|
||
Francke, 1980
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C124D1CFFA5FFB323EED6C9FC43FBAE" authority="(Kraepelin, 1903)" baseAuthorityName="Kraepelin" baseAuthorityYear="1903" class="Arachnida" family="Buthidae" genus="Orthochirus" kingdom="Animalia" order="Scorpiones" pageId="20" pageNumber="19" phylum="Arthropoda" rank="species" species="glabrifrons">
|
||
<emphasis id="B966EA8DFFA5FFB323EED6C9FA70FB8E" box="[1215,1474,1968,1992]" italics="true" pageId="20" pageNumber="19">Orthochirus glabrifrons</emphasis>
|
||
(Kraepelin, 1903)
|
||
</taxonomicName>
|
||
,
|
||
<quantity id="4CEA9B7AFFA5FFB324ACD6A9FBD8FBAE" box="[1021,1130,2000,2024]" metricMagnitude="2" metricUnit="m" metricValue="9.25" metricValueMax="18.5" metricValueMin="0.0" pageId="20" pageNumber="19" unit="m" value="925.0" valueMax="1850.0" valueMin="0.0">0–1850 m</quantity>
|
||
a.s.l.,
|
||
<bibRefCitation id="EF834B6EFFA5FFB323E0D6A8FAF0FBAE" author="LOWE, G." box="[1201,1346,2000,2024]" journalOrPublisher="Euscorpius" pageId="20" pageNumber="19" pagination="1 - 23" part="103" refId="ref40812" refString="LOWE, G. 2010 c. Two new species of Hottentotta Birula, 1908 (Scorpiones: Buthidae) from northern Oman. Euscorpius, 103: 1 - 23." title="Two new species of Hottentotta Birula, 1908 (Scorpiones: Buthidae) from northern Oman" type="journal article" year="2010">Lowe, 2010c</bibRefCitation>
|
||
). The status of
|
||
<taxonomicName id="4C124D1CFFA4FFB227C0D192FF5CFD44" authorityName="Lourenco" authorityYear="2013" box="[145,238,235,258]" class="Arachnida" family="Buthidae" genus="Grosphus" kingdom="Animalia" order="Scorpiones" pageId="21" pageNumber="20" phylum="Arthropoda" rank="species" species="rossii">
|
||
<emphasis id="B966EA8DFFA4FFB227C0D192FF16FD44" box="[145,164,235,258]" italics="true" pageId="21" pageNumber="20">G</emphasis>
|
||
.
|
||
<emphasis id="B966EA8DFFA4FFB227E5D192FF5CFD44" box="[180,238,235,258]" italics="true" pageId="21" pageNumber="20">rossii</emphasis>
|
||
</taxonomicName>
|
||
should be reviewed when topotypic females are collected and their basal pectinal tooth compared to that of
|
||
<taxonomicName id="4C124D1CFFA4FFB225BFD072FEB7FD05" authorityName="Lowe & Kovařík" authorityYear="2019" baseAuthorityName="Lourenco, Goodman & Ramilijaona" baseAuthorityYear="2004" class="Arachnida" family="Buthidae" genus="Teruelius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="21" pageNumber="20" phylum="Arthropoda" rank="species" species="mahafaliensis">
|
||
<emphasis id="B966EA8DFFA4FFB225BFD072FD4FFD64" box="[750,765,267,290]" italics="true" pageId="21" pageNumber="20">T</emphasis>
|
||
.
|
||
<emphasis id="B966EA8DFFA4FFB22720D052FEB7FD05" box="[113,261,299,323]" italics="true" pageId="21" pageNumber="20">mahafaliensis</emphasis>
|
||
</taxonomicName>
|
||
, as this is a more reliable diagnostic character in
|
||
<taxonomicName id="4C124D1CFFA4FFB227DCD032FF5DFD25" authority="Lowe & Kovařík, 2019" authorityName="Lowe & Kovařík" authorityYear="2019" box="[141,239,331,355]" class="Arachnida" family="Buthidae" genus="Teruelius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="21" pageNumber="20" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="B966EA8DFFA4FFB227DCD032FF5DFD25" box="[141,239,331,355]" italics="true" pageId="21" pageNumber="20">Teruelius</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A25557F6FFA4FFB227A7D032FEF3FD24" box="[246,321,331,355]" pageId="21" pageNumber="20" rank="genus">
|
||
<emphasis id="B966EA8DFFA4FFB227A7D032FEACFD25" bold="true" box="[246,286,331,355]" pageId="21" pageNumber="20">gen</emphasis>
|
||
.
|
||
<emphasis id="B966EA8DFFA4FFB2267AD032FE88FD25" bold="true" box="[299,314,331,355]" pageId="21" pageNumber="20">n</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
</paragraph>
|
||
<caption id="DF6D6617FFA3FFB52720D6D8FEACFBB6" ID-DOI="http://doi.org/10.5281/zenodo.7143622" ID-Zenodo-Dep="7143622" httpUri="https://zenodo.org/record/7143622/files/figure.png" pageId="18" pageNumber="17" startId="18.[113,190,1953,1975]" targetBox="[113,1475,255,1894]" targetPageId="18">
|
||
<paragraph id="8BAD369FFFA3FFB52720D6D8FEACFBB6" blockId="18.[113,1474,1953,2032]" pageId="18" pageNumber="17">
|
||
<emphasis id="B966EA8DFFA3FFB52720D6D8FEB0FBF0" bold="true" box="[113,258,1953,1975]" pageId="18" pageNumber="17">Figures 28–29</emphasis>
|
||
. Variation in pectinal tooth count (PTC) in
|
||
<taxonomicName id="4C124D1CFFA3FFB525E2D6D8FCBDFBF0" authorityName="Vachon" authorityYear="1940" box="[691,783,1953,1974]" class="Arachnida" family="Buthidae" genus="Grosphus" kingdom="Animalia" order="Scorpiones" pageId="18" pageNumber="17" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B966EA8DFFA3FFB525E2D6D8FCBDFBF0" box="[691,783,1953,1974]" italics="true" pageId="18" pageNumber="17">Grosphus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C124D1CFFA3FFB5244FD6D8FC2EFBF0" authorityName="Lourenco" authorityYear="1995" box="[798,924,1953,1974]" class="Arachnida" family="Buthidae" genus="Neogrosphus" kingdom="Animalia" order="Scorpiones" pageId="18" pageNumber="17" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B966EA8DFFA3FFB5244FD6D8FC2EFBF0" box="[798,924,1953,1974]" italics="true" pageId="18" pageNumber="17">Neogrosphus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C124D1CFFA3FFB5249FD6D8FB94FBF0" authority="Lowe & Kovařík, 2019" authorityName="Lowe & Kovařík" authorityYear="2019" box="[974,1062,1953,1974]" class="Arachnida" family="Buthidae" genus="Teruelius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="18" pageNumber="17" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="B966EA8DFFA3FFB5249FD6D8FB94FBF0" box="[974,1062,1953,1974]" italics="true" pageId="18" pageNumber="17">Teruelius</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A25557F6FFA3FFB5237FD6D8FBC6FBF0" box="[1070,1140,1953,1974]" pageId="18" pageNumber="17" rank="genus">
|
||
<emphasis id="B966EA8DFFA3FFB5237FD6D8FBE0FBF0" bold="true" box="[1070,1106,1953,1974]" pageId="18" pageNumber="17">gen</emphasis>
|
||
.
|
||
<emphasis id="B966EA8DFFA3FFB52331D6D8FBDFFBF0" bold="true" box="[1120,1133,1953,1974]" pageId="18" pageNumber="17">n</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
Horizontal histograms comparing PTC in males (28), and females (29) of
|
||
<taxonomicName id="4C124D1CFFA3FFB526A2D6C7FDFDFB95" authorityName="Vachon" authorityYear="1940" box="[499,591,1982,2003]" class="Arachnida" family="Buthidae" genus="Grosphus" kingdom="Animalia" order="Scorpiones" pageId="18" pageNumber="17" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B966EA8DFFA3FFB526A2D6C7FDFDFB95" box="[499,591,1982,2003]" italics="true" pageId="18" pageNumber="17">Grosphus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C124D1CFFA3FFB5250DD6C7FD68FB95" authorityName="Lourenco" authorityYear="1995" box="[604,730,1982,2003]" class="Arachnida" family="Buthidae" genus="Neogrosphus" kingdom="Animalia" order="Scorpiones" pageId="18" pageNumber="17" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B966EA8DFFA3FFB5250DD6C7FD68FB95" box="[604,730,1982,2003]" italics="true" pageId="18" pageNumber="17">Neogrosphus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C124D1CFFA3FFB5245BD6C7FCD0FB95" authority="Lowe & Kovařík, 2019" authorityName="Lowe & Kovařík" authorityYear="2019" box="[778,866,1982,2003]" class="Arachnida" family="Buthidae" genus="Teruelius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="18" pageNumber="17" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="B966EA8DFFA3FFB5245BD6C7FCD0FB95" box="[778,866,1982,2003]" italics="true" pageId="18" pageNumber="17">Teruelius</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A25557F6FFA3FFB52438D6C7FC1FFB95" box="[873,941,1982,2003]" pageId="18" pageNumber="17" rank="genus">
|
||
<emphasis id="B966EA8DFFA3FFB52438D6C7FC3FFB95" bold="true" box="[873,909,1982,2003]" pageId="18" pageNumber="17">gen</emphasis>
|
||
.
|
||
<emphasis id="B966EA8DFFA3FFB524C8D6C7FC14FB95" bold="true" box="[921,934,1982,2003]" pageId="18" pageNumber="17">n</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
Error bars indicate ranges (minimum, maximum), bars mid-range values.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF6D6617FFA2FFB42720D794FCFFFBAF" ID-DOI="http://doi.org/10.5281/zenodo.7143626" ID-Zenodo-Dep="7143626" httpUri="https://zenodo.org/record/7143626/files/figure.png" pageId="19" pageNumber="18" startId="19.[113,190,1773,1795]" targetBox="[118,1455,257,1710]" targetPageId="19">
|
||
<paragraph id="8BAD369FFFA2FFB42720D794FCFFFBAF" blockId="19.[113,1474,1773,2025]" pageId="19" pageNumber="18">
|
||
<emphasis id="B966EA8DFFA2FFB42720D794FEB3FB44" bold="true" box="[113,257,1773,1795]" pageId="19" pageNumber="18">Figures 30–35</emphasis>
|
||
. Variation in pectinal tooth count (PTC) in
|
||
<taxonomicName id="4C124D1CFFA2FFB425F8D794FCB7FB44" authorityName="Vachon" authorityYear="1940" box="[681,773,1773,1794]" class="Arachnida" family="Buthidae" genus="Grosphus" kingdom="Animalia" order="Scorpiones" pageId="19" pageNumber="18" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B966EA8DFFA2FFB425F8D794FCB7FB44" box="[681,773,1773,1794]" italics="true" pageId="19" pageNumber="18">Grosphus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C124D1CFFA2FFB42443D794FC22FB44" authorityName="Lourenco" authorityYear="1995" box="[786,912,1773,1794]" class="Arachnida" family="Buthidae" genus="Neogrosphus" kingdom="Animalia" order="Scorpiones" pageId="19" pageNumber="18" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B966EA8DFFA2FFB42443D794FC22FB44" box="[786,912,1773,1794]" italics="true" pageId="19" pageNumber="18">Neogrosphus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C124D1CFFA2FFB42491D794FBAAFB44" authority="Lowe & Kovařík, 2019" authorityName="Lowe & Kovařík" authorityYear="2019" box="[960,1048,1773,1794]" class="Arachnida" family="Buthidae" genus="Teruelius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="19" pageNumber="18" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="B966EA8DFFA2FFB42491D794FBAAFB44" box="[960,1048,1773,1794]" italics="true" pageId="19" pageNumber="18">Teruelius</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A25557F6FFA2FFB4234ED794FBD1FB44" box="[1055,1123,1773,1794]" pageId="19" pageNumber="18" rank="genus">
|
||
<emphasis id="B966EA8DFFA2FFB4234ED794FBF1FB44" bold="true" box="[1055,1091,1773,1794]" pageId="19" pageNumber="18">gen</emphasis>
|
||
.
|
||
<emphasis id="B966EA8DFFA2FFB42301D794FBEFFB44" bold="true" box="[1104,1117,1773,1794]" pageId="19" pageNumber="18">n</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
<emphasis id="B966EA8DFFA2FFB4233BD794FB4BFB44" bold="true" box="[1130,1273,1773,1795]" pageId="19" pageNumber="18">Figures 30–31</emphasis>
|
||
. Logarithmic scatter plots showing scaling of PTC vs. carapace length in males (30), and females (31) of
|
||
<taxonomicName id="4C124D1CFFA2FFB424CDD673FC4AFB59" authorityName="Vachon" authorityYear="1940" box="[924,1016,1802,1823]" class="Arachnida" family="Buthidae" genus="Grosphus" kingdom="Animalia" order="Scorpiones" pageId="19" pageNumber="18" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B966EA8DFFA2FFB424CDD673FC4AFB59" box="[924,1016,1802,1823]" italics="true" pageId="19" pageNumber="18">Grosphus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C124D1CFFA2FFB42355D673FB30FB59" authorityName="Lourenco" authorityYear="1995" box="[1028,1154,1802,1823]" class="Arachnida" family="Buthidae" genus="Neogrosphus" kingdom="Animalia" order="Scorpiones" pageId="19" pageNumber="18" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B966EA8DFFA2FFB42355D673FB30FB59" box="[1028,1154,1802,1823]" italics="true" pageId="19" pageNumber="18">Neogrosphus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C124D1CFFA2FFB423E1D673FABAFB59" authority="Lowe & Kovařík, 2019" authorityName="Lowe & Kovařík" authorityYear="2019" box="[1200,1288,1802,1823]" class="Arachnida" family="Buthidae" genus="Teruelius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="19" pageNumber="18" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="B966EA8DFFA2FFB423E1D673FABAFB59" box="[1200,1288,1802,1823]" italics="true" pageId="19" pageNumber="18">Teruelius</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A25557F6FFA2FFB4225ED673FAE0FB59" box="[1295,1362,1802,1823]" pageId="19" pageNumber="18" rank="genus">
|
||
<emphasis id="B966EA8DFFA2FFB4225ED673FA81FB59" bold="true" box="[1295,1331,1802,1823]" pageId="19" pageNumber="18">gen</emphasis>
|
||
.
|
||
<emphasis id="B966EA8DFFA2FFB4226ED673FAFEFB59" bold="true" box="[1343,1356,1802,1823]" pageId="19" pageNumber="18">n</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
Color lines are least squares linear regression fits of scaling trends for the different genera.
|
||
<emphasis id="B966EA8DFFA2FFB424D9D65FFBABFB7D" bold="true" box="[904,1049,1830,1852]" pageId="19" pageNumber="18">Figures 32–33</emphasis>
|
||
. Logarithmic scatter plots showing scaling of PTC vs. carapace length in Figs. 30–31 superposed upon overall scaling trends for a larger sample of male (32) and female (33) buthids including both α and β trichobothrial configurations (gray symbols; 1055♂, 1046♀). Dark lines: least squares linear regression fits of scaling trends for larger sample of buthids.
|
||
<emphasis id="B966EA8DFFA2FFB42689D604FDDBFBD4" bold="true" box="[472,617,1917,1939]" pageId="19" pageNumber="18">Figures 34–35</emphasis>
|
||
. Logarithmic scatter plots showing scaling of PTC vs. carapace length in Figs. 30–31 superposed upon overall scaling trends for a larger sample of male (34) and female (35) α-buthids restricted to α trichobothrial configurations (gray symbols; 445♂, 371♀). Dark lines: least squares linear regression fits of scaling trends for α-buthids. Numbers: slopes of regression lines (= allometric exponents). Purple symbols and lines: outgroup
|
||
<taxonomicName id="4C124D1CFFA2FFB42592D6AAFCF5FBAE" authorityName="Kraepelin" authorityYear="1911" box="[707,839,2003,2024]" class="Arachnida" family="Buthidae" genus="Pseudolychas" kingdom="Animalia" order="Scorpiones" pageId="19" pageNumber="18" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B966EA8DFFA2FFB42592D6AAFCF5FBAE" box="[707,839,2003,2024]" italics="true" pageId="19" pageNumber="18">Pseudolychas</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF6D6617FFA5FFB32720D415FCCEFA0A" ID-DOI="http://doi.org/10.5281/zenodo.7143628" ID-Zenodo-Dep="7143628" httpUri="https://zenodo.org/record/7143628/files/figure.png" pageId="20" pageNumber="19" startId="20.[113,190,1388,1410]" targetBox="[121,1471,246,1361]" targetPageId="20">
|
||
<paragraph id="8BAD369FFFA5FFB32720D415FCCEFA0A" blockId="20.[113,1474,1388,1612]" pageId="20" pageNumber="19">
|
||
<emphasis id="B966EA8DFFA5FFB32720D415FF48F9C4" bold="true" box="[113,250,1388,1410]" pageId="20" pageNumber="19">Figures 36–39</emphasis>
|
||
. Comparative biometrics of female basal pectinal tooth and hemispermatophore trunk of
|
||
<taxonomicName id="4C124D1CFFA5FFB32307D414FB00F9C4" authorityName="Vachon" authorityYear="1940" box="[1110,1202,1389,1410]" class="Arachnida" family="Buthidae" genus="Grosphus" kingdom="Animalia" order="Scorpiones" pageId="20" pageNumber="19" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B966EA8DFFA5FFB32307D414FB00F9C4" box="[1110,1202,1389,1410]" italics="true" pageId="20" pageNumber="19">Grosphus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C124D1CFFA5FFB323EFD414FA8EF9C4" authorityName="Lourenco" authorityYear="1995" box="[1214,1340,1389,1410]" class="Arachnida" family="Buthidae" genus="Neogrosphus" kingdom="Animalia" order="Scorpiones" pageId="20" pageNumber="19" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B966EA8DFFA5FFB323EFD414FA8EF9C4" box="[1214,1340,1389,1410]" italics="true" pageId="20" pageNumber="19">Neogrosphus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C124D1CFFA5FFB3223BD414FA70F9C4" authority="Lowe & Kovařík, 2019" authorityName="Lowe & Kovařík" authorityYear="2019" box="[1386,1474,1389,1410]" class="Arachnida" family="Buthidae" genus="Teruelius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="20" pageNumber="19" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="B966EA8DFFA5FFB3223BD414FA70F9C4" box="[1386,1474,1389,1410]" italics="true" pageId="20" pageNumber="19">Teruelius</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A25557F6FFA5FFB32720D4F3FF0AF9D9" box="[113,184,1418,1439]" pageId="20" pageNumber="19" rank="genus">
|
||
<emphasis id="B966EA8DFFA5FFB32720D4F3FF27F9D9" bold="true" box="[113,149,1418,1439]" pageId="20" pageNumber="19">gen</emphasis>
|
||
.
|
||
<emphasis id="B966EA8DFFA5FFB327F4D4F3FF00F9D9" bold="true" box="[165,178,1418,1439]" pageId="20" pageNumber="19">n</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
<emphasis id="B966EA8DFFA5FFB32790D4F0FEE6F9D8" bold="true" box="[193,340,1417,1439]" pageId="20" pageNumber="19">Figures 36–37</emphasis>
|
||
. Logarithmic scatter plots showing scaling of length of female basal pectinal tooth vs. carapace length (36), and hemispermatophore trunk length vs. carapace length (37). Color lines: least squares linear regression fits of scaling trends for the different genera.
|
||
<emphasis id="B966EA8DFFA5FFB327EFD4BAFEFFF99E" bold="true" box="[190,333,1475,1497]" pageId="20" pageNumber="19">Figures 38–39</emphasis>
|
||
. Logarithmic scatter plots of female basal pectinal tooth length vs. hemispermatophore trunk length (38), and the ratio of basal pectinal tooth length/ hemispermatophore trunk length (%) vs. hemispermatophore trunk length, showing fraction of trunk spanned by the female tooth (39). Female basal pectinal tooth length measured as chord length, from more proximal corner (along comb axis) at base of tooth, to more distal corner (along comb axis) at tip of tooth. Trunk length measured from proximal edge of basal lobe to pedicel (foot). Color lines: least squares linear regression fits of scaling trends for the different genera.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BAD369FFFA4FFB227F0D012FDFCF845" blockId="21.[113,771,235,1027]" pageId="21" pageNumber="20">
|
||
Another potential synonym of
|
||
<taxonomicName id="4C124D1CFFA4FFB22556D012FD0DFDC5" authorityName="Lowe & Kovařík" authorityYear="2019" baseAuthorityName="Lourenco, Goodman & Ramilijaona" baseAuthorityYear="2004" box="[519,703,363,387]" class="Arachnida" family="Buthidae" genus="Teruelius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="21" pageNumber="20" phylum="Arthropoda" rank="species" species="mahafaliensis">
|
||
<emphasis id="B966EA8DFFA4FFB22556D012FDA4FDC4" box="[519,534,363,386]" italics="true" pageId="21" pageNumber="20">T</emphasis>
|
||
.
|
||
<emphasis id="B966EA8DFFA4FFB2257AD012FD0DFDC5" box="[555,703,363,387]" italics="true" pageId="21" pageNumber="20">mahafaliensis</emphasis>
|
||
</taxonomicName>
|
||
is
|
||
<taxonomicName id="4C124D1CFFA4FFB225BFD012FE00FDE5" authority="(Lourenco, 2014)" authorityName="Lowe & Kovařík" authorityYear="2019" baseAuthorityName="Lourenco" baseAuthorityYear="2014" class="Arachnida" family="Buthidae" genus="Teruelius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="21" pageNumber="20" phylum="Arthropoda" rank="species" species="magalieae">
|
||
<emphasis id="B966EA8DFFA4FFB225BFD012FD4FFDC4" box="[750,765,363,386]" italics="true" pageId="21" pageNumber="20">T</emphasis>
|
||
.
|
||
<emphasis id="B966EA8DFFA4FFB22720D0F2FF52FDE5" box="[113,224,395,419]" italics="true" pageId="21" pageNumber="20">magalieae</emphasis>
|
||
(
|
||
<bibRefCitation id="EF834B6EFFA4FFB227A4D0F2FE18FDE5" author="LOURENCO, W. R." box="[245,426,395,419]" journalOrPublisher="Zoosystema" pageId="21" pageNumber="20" pagination="631 - 645" part="36" refId="ref38903" refString="LOURENCO, W. R. 2014. The genus Grosphus Simon, 1880 in South-Western Madagascar, with the description of a new species (Scorpiones, Buthidae). Zoosystema, 36 (3): 631 - 645." title="The genus Grosphus Simon, 1880 in South-Western Madagascar, with the description of a new species (Scorpiones, Buthidae)" type="journal article" year="2014">Lourenço, 2014</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
. According to its description, the morphometrics, coloration, and meristics of
|
||
<taxonomicName id="4C124D1CFFA4FFB22526D0D2FCB1FD85" authorityName="Lowe & Kovařík" authorityYear="2019" baseAuthorityName="Lourenco" baseAuthorityYear="2014" box="[631,771,427,451]" class="Arachnida" family="Buthidae" genus="Teruelius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="21" pageNumber="20" phylum="Arthropoda" rank="species" species="magalieae">
|
||
<emphasis id="B966EA8DFFA4FFB22526D0D2FD34FD84" box="[631,646,427,450]" italics="true" pageId="21" pageNumber="20">T</emphasis>
|
||
.
|
||
<emphasis id="B966EA8DFFA4FFB225C5D0D2FCB1FD85" box="[660,771,427,451]" italics="true" pageId="21" pageNumber="20">magalieae</emphasis>
|
||
</taxonomicName>
|
||
are very close to those of
|
||
<taxonomicName id="4C124D1CFFA4FFB226DDD0B2FD8FFDA5" authorityName="Lowe & Kovařík" authorityYear="2019" baseAuthorityName="Lourenco, Goodman & Ramilijaona" baseAuthorityYear="2004" box="[396,573,459,483]" class="Arachnida" family="Buthidae" genus="Teruelius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="21" pageNumber="20" phylum="Arthropoda" rank="species" species="mahafaliensis">
|
||
<emphasis id="B966EA8DFFA4FFB226DDD0B2FE29FDA4" box="[396,411,459,482]" italics="true" pageId="21" pageNumber="20">T</emphasis>
|
||
.
|
||
<emphasis id="B966EA8DFFA4FFB226F8D0B2FD8FFDA5" box="[425,573,459,483]" italics="true" pageId="21" pageNumber="20">mahafaliensis</emphasis>
|
||
</taxonomicName>
|
||
. The type locality of Cap Saint Marie (
|
||
<typeStatus id="54A9883DFFA4FFB2261ED092FE1CFE45" box="[335,430,491,515]" pageId="21" pageNumber="20" type="holotype">holotype</typeStatus>
|
||
male as only known specimen) lies on the southwestern coast in the same bioclimatic region as the latter species.
|
||
<bibRefCitation id="EF834B6EFFA4FFB22609D352FDFAFE05" author="LOURENCO, W. R." box="[344,584,555,579]" journalOrPublisher="Zoosystema" pageId="21" pageNumber="20" pagination="631 - 645" part="36" refId="ref38903" refString="LOURENCO, W. R. 2014. The genus Grosphus Simon, 1880 in South-Western Madagascar, with the description of a new species (Scorpiones, Buthidae). Zoosystema, 36 (3): 631 - 645." title="The genus Grosphus Simon, 1880 in South-Western Madagascar, with the description of a new species (Scorpiones, Buthidae)" type="journal article" year="2014">Lourenço (2014: 633)</bibRefCitation>
|
||
did not compare
|
||
<taxonomicName id="4C124D1CFFA4FFB22720D332FEB5FE25" authorityName="Lowe & Kovařík" authorityYear="2019" baseAuthorityName="Lourenco" baseAuthorityYear="2014" box="[113,263,587,611]" class="Arachnida" family="Buthidae" genus="Teruelius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="21" pageNumber="20" phylum="Arthropoda" rank="species" species="magalieae">
|
||
<emphasis id="B966EA8DFFA4FFB22720D332FF32FE24" box="[113,128,587,610]" italics="true" pageId="21" pageNumber="20">T</emphasis>
|
||
.
|
||
<emphasis id="B966EA8DFFA4FFB227C9D332FEB5FE25" box="[152,263,587,611]" italics="true" pageId="21" pageNumber="20">magalieae</emphasis>
|
||
</taxonomicName>
|
||
to
|
||
<taxonomicName id="4C124D1CFFA4FFB2266FD332FE4BFE25" authorityName="Lowe & Kovařík" authorityYear="2019" baseAuthorityName="Lourenco, Goodman & Ramilijaona" baseAuthorityYear="2004" box="[318,505,587,611]" class="Arachnida" family="Buthidae" genus="Teruelius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="21" pageNumber="20" phylum="Arthropoda" rank="species" species="mahafaliensis">
|
||
<emphasis id="B966EA8DFFA4FFB2266FD332FEFFFE24" box="[318,333,587,610]" italics="true" pageId="21" pageNumber="20">T</emphasis>
|
||
.
|
||
<emphasis id="B966EA8DFFA4FFB22634D332FE4BFE25" box="[357,505,587,611]" italics="true" pageId="21" pageNumber="20">mahafaliensis</emphasis>
|
||
</taxonomicName>
|
||
, but claimed that the most closely related species was
|
||
<taxonomicName id="4C124D1CFFA4FFB226BAD312FDF9FEC4" authorityName="Lourenco" authorityYear="2013" box="[491,587,619,642]" class="Arachnida" family="Buthidae" genus="Grosphus" kingdom="Animalia" order="Scorpiones" pageId="21" pageNumber="20" phylum="Arthropoda" rank="species" species="rossii">
|
||
<emphasis id="B966EA8DFFA4FFB226BAD312FE4CFEC4" box="[491,510,619,642]" italics="true" pageId="21" pageNumber="20">G</emphasis>
|
||
.
|
||
<emphasis id="B966EA8DFFA4FFB22540D312FDF9FEC4" box="[529,587,619,642]" italics="true" pageId="21" pageNumber="20">rossii</emphasis>
|
||
</taxonomicName>
|
||
, which we here synonymize under
|
||
<taxonomicName id="4C124D1CFFA4FFB22666D3F2FE57FEE5" authorityName="Lowe & Kovařík" authorityYear="2019" baseAuthorityName="Lourenco, Goodman & Ramilijaona" baseAuthorityYear="2004" box="[311,485,651,675]" class="Arachnida" family="Buthidae" genus="Teruelius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="21" pageNumber="20" phylum="Arthropoda" rank="species" species="mahafaliensis">
|
||
<emphasis id="B966EA8DFFA4FFB22666D3F2FEF4FEE4" box="[311,326,651,674]" italics="true" pageId="21" pageNumber="20">T</emphasis>
|
||
.
|
||
<emphasis id="B966EA8DFFA4FFB22600D3F2FE57FEE5" box="[337,485,651,675]" italics="true" pageId="21" pageNumber="20">mahafaliensis</emphasis>
|
||
</taxonomicName>
|
||
. The diagnostic differences between
|
||
<taxonomicName id="4C124D1CFFA4FFB22784D3D2FED6FE85" authorityName="Lowe & Kovařík" authorityYear="2019" baseAuthorityName="Lourenco" baseAuthorityYear="2014" box="[213,356,683,707]" class="Arachnida" family="Buthidae" genus="Teruelius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="21" pageNumber="20" phylum="Arthropoda" rank="species" species="magalieae">
|
||
<emphasis id="B966EA8DFFA4FFB22784D3D2FF56FE84" box="[213,228,683,706]" italics="true" pageId="21" pageNumber="20">T</emphasis>
|
||
.
|
||
<emphasis id="B966EA8DFFA4FFB227A4D3D2FED6FE85" box="[245,356,683,707]" italics="true" pageId="21" pageNumber="20">magalieae</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C124D1CFFA4FFB226CED3D2FE4CFE84" authorityName="Lourenco" authorityYear="2013" box="[415,510,683,706]" class="Arachnida" family="Buthidae" genus="Grosphus" kingdom="Animalia" order="Scorpiones" pageId="21" pageNumber="20" phylum="Arthropoda" rank="species" species="rossii">
|
||
<emphasis id="B966EA8DFFA4FFB226CED3D2FE00FE84" box="[415,434,683,706]" italics="true" pageId="21" pageNumber="20">G</emphasis>
|
||
.
|
||
<emphasis id="B966EA8DFFA4FFB22695D3D2FE4CFE84" box="[452,510,683,706]" italics="true" pageId="21" pageNumber="20">rossii</emphasis>
|
||
</taxonomicName>
|
||
are not compelling: (i) pectines with 36 vs. 28 teeth (a range of variation allowed here for
|
||
<taxonomicName id="4C124D1CFFA4FFB22782D392FE35FF45" authorityName="Lowe & Kovařík" authorityYear="2019" baseAuthorityName="Lourenco, Goodman & Ramilijaona" baseAuthorityYear="2004" box="[211,391,747,771]" class="Arachnida" family="Buthidae" genus="Teruelius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="21" pageNumber="20" phylum="Arthropoda" rank="species" species="mahafaliensis">
|
||
<emphasis id="B966EA8DFFA4FFB22782D392FF50FF44" box="[211,226,747,770]" italics="true" pageId="21" pageNumber="20">T</emphasis>
|
||
.
|
||
<emphasis id="B966EA8DFFA4FFB227A2D392FE35FF45" box="[243,391,747,771]" italics="true" pageId="21" pageNumber="20">mahafaliensis</emphasis>
|
||
</taxonomicName>
|
||
); (ii) pedipalp fingers with 12–13 vs. 12–12 granule rows (overlapping counts); and (iii) overall paler coloration (differences in color shade are not uncommon for different populations of a species inhabiting areas with different substrates). We provisionally list this species, until it can be critically evaluated by study of more material and analysis of variation. The female of
|
||
<taxonomicName id="4C124D1CFFA4FFB226A1D2D2FDC9FF85" authorityName="Lowe & Kovařík" authorityYear="2019" baseAuthorityName="Lourenco" baseAuthorityYear="2014" box="[496,635,939,963]" class="Arachnida" family="Buthidae" genus="Teruelius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="21" pageNumber="20" phylum="Arthropoda" rank="species" species="magalieae">
|
||
<emphasis id="B966EA8DFFA4FFB226A1D2D2FE4DFF84" box="[496,511,939,962]" italics="true" pageId="21" pageNumber="20">T</emphasis>
|
||
.
|
||
<emphasis id="B966EA8DFFA4FFB2255DD2D2FDC9FF85" box="[524,635,939,963]" italics="true" pageId="21" pageNumber="20">magalieae</emphasis>
|
||
</taxonomicName>
|
||
is unknown, and the species might be better diagnosed if the female basal pectinal tooth were determined to be unique.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |