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<document id="AF467677B896A9AED7AFCB20B5838579" ID-CLB-Dataset="24019" ID-DOI="10.1080/00222933.2018.1524032" ID-GBIF-Dataset="034ebcd9-efa7-416a-822c-b6aee0e57a1c" ID-ISSN="1464-5262" ID-Zenodo-Dep="3670229" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1581941790231" checkinUser="valdenar" docAuthor="DeVries, Thomas J." docDate="2019" docId="03BB2338E354E226FE75A3245E88FEB4" docLanguage="en" docName="JNATHIST.53.25–26.1533–1584.pdf.imf" docOrigin="Journal of Natural History 53 (25)" docStyle="DocumentStyle:485A7191CD41D6261F41521A3380DC73.9:JNatHist.2017-.journal_article.1cover" docStyleId="485A7191CD41D6261F41521A3380DC73" docStyleName="JNatHist.2017-.journal_article.1cover" docStyleVersion="9" docTitle="Papposilenus utriculus DeVries 2019, sp. nov." docType="treatment" docVersion="23" lastPageNumber="1561" masterDocId="FF825B40E348E23BFFA1A1435A0CFFCC" masterDocTitle="Early Paleogene brackish-water molluscs from the Caballas Formation of the East Pisco Basin (Southern Peru)" masterLastPageNumber="1584" masterPageNumber="1533" pageNumber="1560" updateTime="1698783606523" updateUser="ExternalLinkService">
<mods:mods id="90D16E65E4CFB4246CDDB924C2F62E09" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="E26408A493AEAF2B263C6D298316B8D2">
<mods:title id="33CC4CC7B5573A81D1540F4F5534B4E6">Early Paleogene brackish-water molluscs from the Caballas Formation of the East Pisco Basin (Southern Peru)</mods:title>
</mods:titleInfo>
<mods:name id="AED11F7EF211D54D7F7C108718196F49" type="personal">
<mods:role id="5105FE3BB5572F396D6505273B7B1901">
<mods:roleTerm id="DE1238145B21BD5C9EA2C4AF02304349">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="B2BC12DDF624D53D7CCFDCFBEC54786D">DeVries, Thomas J.</mods:namePart>
</mods:name>
<mods:typeOfResource id="C9D65BDD90E8BA9A47603B3B1FF14617">text</mods:typeOfResource>
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<mods:title id="16661E834BB7F11E612F3384978961E3">Journal of Natural History</mods:title>
</mods:titleInfo>
<mods:part id="8A74583A9EF0E67EFA4675B4B43CBB34">
<mods:date id="B4E9F477E451A5674CAB6A37C411C7B8">2019</mods:date>
<mods:detail id="F2AAEDE81F5EC1D648F1314F9ED5FFA5" type="pubDate">
<mods:number id="117F8B4BCDDD698246B726EF2D4E7EA1">2018-12-17</mods:number>
</mods:detail>
<mods:detail id="BBA4A2579A946C8E92BC6071277082B9" type="volume">
<mods:number id="B9DA83521E569FE80EA90D6DE23C00DA">53</mods:number>
</mods:detail>
<mods:detail id="D1AAA73759CB6482EC036090D43E69FF" type="issue">
<mods:number id="270DE279B59FA0C880FE91B38F0422B5">25</mods:number>
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<mods:start id="482D55A86199FD00393FEC4C8B1877AE">1533</mods:start>
<mods:end id="19CD491C276869C1A2682E21EA7A1090">1584</mods:end>
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<mods:classification id="36C4B957722DA3EBA6EE0628E27B64A3">journal article</mods:classification>
<mods:identifier id="285F4EE77B4A3D34031763E670814669" type="CLB-Dataset">24019</mods:identifier>
<mods:identifier id="321C4274A2486D27828D44F68F4E702D" type="DOI">10.1080/00222933.2018.1524032</mods:identifier>
<mods:identifier id="CAB5AEB3F961EDE52FB3DE41A4312285" type="GBIF-Dataset">034ebcd9-efa7-416a-822c-b6aee0e57a1c</mods:identifier>
<mods:identifier id="3E5BE0877ACBC72ACFB960FC587BA226" type="ISSN">1464-5262</mods:identifier>
<mods:identifier id="5471799CF99C51429153E2E8F6BF96CE" type="Zenodo-Dep">3670229</mods:identifier>
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<treatment id="03BB2338E354E226FE75A3245E88FEB4" ID-DOI="http://doi.org/10.5281/zenodo.3671204" ID-GBIF-Taxon="162146509" ID-Zenodo-Dep="3671204" LSID="urn:lsid:plazi:treatment:03BB2338E354E226FE75A3245E88FEB4" httpUri="http://treatment.plazi.org/id/03BB2338E354E226FE75A3245E88FEB4" lastPageId="29" lastPageNumber="1561" pageId="28" pageNumber="1560">
<subSubSection id="C308C1A5E354E227FE75A3245943FD4C" box="[468,847,615,640]" pageId="28" pageNumber="1560" type="nomenclature">
<paragraph id="8BAD922EE354E227FE75A3245943FD4C" blockId="28.[468,847,615,675]" box="[468,847,615,640]" pageId="28" pageNumber="1560">
<heading id="D0E52542E354E227FE75A3245943FD4C" box="[468,847,615,640]" centered="true" fontSize="11" level="2" pageId="28" pageNumber="1560" reason="2">
<taxonomicName id="4C12E9ADE354E227FE75A32458E5FD4C" authority="DeVries, 2019" authorityName="DeVries" authorityYear="2019" box="[468,745,615,640]" class="Gastropoda" family="Potamididae" genus="Papposilenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neotaenioglossa" pageId="28" pageNumber="1560" phylum="Mollusca" rank="species" species="utriculus" status="sp. nov.">
<emphasis id="B9664E3CE354E227FE75A32458E5FD4C" bold="true" box="[468,745,615,640]" italics="true" pageId="28" pageNumber="1560">Papposilenus utriculus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A255F347E354E227FD52A3245943FD4C" box="[755,847,615,640]" pageId="28" pageNumber="1560" rank="species">sp. nov.</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C308C1A5E354E227FD8CA3C958FBFD6F" box="[557,759,650,675]" pageId="28" pageNumber="1560" type="description">
<paragraph id="8BAD922EE354E227FD8CA3C958FBFD6F" blockId="28.[468,847,615,675]" box="[557,759,650,675]" pageId="28" pageNumber="1560">
<heading id="D0E52542E354E227FD8CA3C958FBFD6F" box="[557,759,650,675]" centered="true" fontSize="11" level="2" pageId="28" pageNumber="1560" reason="5">
(
<figureCitation id="13298EABE354E227FD94A3C95894FD6F" box="[565,664,650,675]" captionStart="Figure 4" captionStartId="13.[160,225,936,959]" captionTargetId="figure@14.[162,1153,142,1037]" captionTargetPageId="14" captionText="Figure 4. Gastropods from the Caballas Formation. (a–e) Nerita squatina sp. nov., B8771. (a) UWBM 107577, holotype, W 5.7 mm, spire. (b) UWBM 107648, paratype, W 8.5 mm, shoulder of last whorl. (c) UWBM 107649,paratype, W 10.1 mm, anterior of outer lip.(d) UWBM 107647,paratype,W 7.8 mm,inner surface of outer lip. (e) UWBM 107646, paratype, W 6.2 mm, columnar teeth. (f) Calyptraea cf. C. aperta Solander, 1766, UWBM 107578, B8772,W (diameter) 9.9 mm, external mould. (g) Ampullina ortoni Gabb, 1870, UWBM 107,627, B8772, L 36.8 mm, abapertural view, last whorl mostly missing; left margin annotated. (h–l) Melanella indicaformis sp. nov. (h) UWBM 107620, paratype, B8772, L 8.2 mm, apertural view. (i) UWBM 107619, paratype, B8769, L 9.4 mm, apertural/lateral view. (j) UWBM 107623, paratype, B8769,L 8.2 mm, apertural view. (k) UWBM 107618, paratype, B8772, L 10.6, abapertural view. (l) UWBM 107617, holotype, B8769, L 11.1 mm, abapertural view. (m–q) Potamides henryi sp. nov. (m) UWBM 107582, holotype, B8772, L 17.8 mm, lateral view. (n) UWBM 107582, W 6.9 mm, basal view. (o) UWBM 107586, paratype, B8771, L 11.2 mm, lateral view. (p) UWBM 107587, paratype, B8769, L 9.5 mm, abapertural view. (q) UWBM 107650, paratype, B8770, L 15.4 mm, lateral view, shell compressed. (r–t) Potamides janeae sp. nov., B8771.(r) UWBM 107579, holotype, L 9.0 mm, lateral view. (s) MUSM INV 249, L 8.8 mm, lateral and columellar view. (t) UWBM 107580, L 9.5 mm, layeral and columellar view. (u, v, x) Papposilenus utriculus gen.nov., sp. nov. (u) UWBM 107592,paratype, B8772, L 18.7 mm, oblique view of base. (v) UWBM 107589, holotype, B8772, L 25.5 mm, lateral view. (x) UWBM 107590, B8770, L 17.6 mm, lateral view.(w, y,z,jj) Rhinotamides everriculum sp.nov. (w) UWBM 107598,paratype, B8772,L 23.5 mm, abapertural view, juvenile. (y) UWBM 107593, holotype, B8769, L 21.1 mm, abapertural view. (z) UWBM 107594, paratype, B8772, L 19.4 mm, apertural view, posterior and anterior missing. (jj) UWBM 107593, apertural view.(aa,bb) Terebralia pauli gen. nov.sp.nov., UWBM 107599,holotype,B8771, L 9.2 mm.(aa) Abapertural view. (bb) Apertural view, outer lip missing. (cc) Terebralia marki gen. nov. sp. nov., UWBM 107600, holotype, B8771, L 9.8, lateral view. (dd–ii). Nodifaunus gainesi sp. nov. (dd) UWBM 107609, holotype, B8769, L 42.4 mm, apertural view. (ee) MUSM INV 264, paratype, B8772, L 31.0 mm, apertural view, anterior missing. (ff) UWBM 107609, abapertural view. (gg) UWBM 107612, paratype, B8770, L 14.2 mm, abapertural view, juvenile. (hh) UWBM 107612, apertural view. (ii) UWBM 107610, paratype, B8769, L 25.9 mm, apertural view, anterior missing." figureDoi="http://doi.org/10.5281/zenodo.3686966" httpUri="https://zenodo.org/record/3686966/files/figure.png" pageId="28" pageNumber="1560">Figure 4</figureCitation>
(u, v, x))
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C308C1A5E354E227FF01A3A05B15FC8E" pageId="28" pageNumber="1560" type="diagnosis">
<paragraph id="8BAD922EE354E227FF01A3A05B18FD31" blockId="28.[160,1155,739,834]" box="[160,276,739,765]" pageId="28" pageNumber="1560">
<emphasis id="B9664E3CE354E227FF01A3A05B18FD31" bold="true" box="[160,276,739,765]" italics="true" pageId="28" pageNumber="1560">Diagnosis</emphasis>
</paragraph>
<paragraph id="8BAD922EE354E227FF01A2445B15FC8E" blockId="28.[160,1155,739,834]" pageId="28" pageNumber="1560">Characters as for genus; two anterior spiral cords on earliest teleoconch whorls unbeaded.</paragraph>
</subSubSection>
<subSubSection id="C308C1A5E354E227FF01A2C15838FAF0" pageId="28" pageNumber="1560" type="description">
<paragraph id="8BAD922EE354E227FF01A2C15B2AFC50" blockId="28.[160,1156,898,1340]" box="[160,294,898,924]" pageId="28" pageNumber="1560">
<emphasis id="B9664E3CE354E227FF01A2C15B2AFC50" bold="true" box="[160,294,898,924]" italics="true" pageId="28" pageNumber="1560">Description</emphasis>
</paragraph>
<paragraph id="8BAD922EE354E227FF01A2E55838FAF0" blockId="28.[160,1156,898,1340]" pageId="28" pageNumber="1560">Estimated shell length about 30 mm, stoutly conical, spire angle about 35°. Protoconch unknown. Spire with about nine whorls. Whorls flat-sided or slightly convex, sutures thinly incised, recessed. All preserved whorls with three spiral cords. Posteriormost spiral cord strongest, beaded on very early whorls, beads rhombohedral, flattened or nodular, with long axis parallel to shell axis and sides prosocline. Two anterior spiral cords slightly narrower, equal in size to each other, divided by thin incision on early whorls, not beaded. Last three or four whorls with all cords fully beaded; beads aligned, producing orthocline to slightly prosocline axial groove, broadly sulcate medially. Spire varices absent. Base poorly preserved; with at least four unbeaded or weakly beaded spiral cords separated by equally wide interspaces. Columella without folds. Outer lip flared from posterior attachment point but mostly not preserved. Anterior canal strongly recurved but mostly not preserved.</paragraph>
</subSubSection>
<subSubSection id="C308C1A5E354E227FF01A43F585FF9EC" pageId="28" pageNumber="1560" type="discussion">
<paragraph id="8BAD922EE354E227FF01A43F5B0AFA5A" blockId="28.[160,1155,1404,1568]" box="[160,262,1404,1430]" pageId="28" pageNumber="1560">
<emphasis id="B9664E3CE354E227FF01A43F5B0AFA5A" bold="true" box="[160,262,1404,1430]" italics="true" pageId="28" pageNumber="1560">Remarks</emphasis>
</paragraph>
<paragraph id="8BAD922EE354E227FF01A4E3585FF9EC" blockId="28.[160,1155,1404,1568]" pageId="28" pageNumber="1560">
Specimens of the early Eocene
<taxonomicName id="4C12E9ADE354E227FDA3A4E358FFFA74" authority="DeVries, 2019" authorityName="DeVries" authorityYear="2019" box="[514,755,1440,1464]" class="Gastropoda" family="Potamididae" genus="Papposilenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neotaenioglossa" pageId="28" pageNumber="1560" phylum="Mollusca" rank="species" species="utriculus" status="sp. nov.">
<emphasis id="B9664E3CE354E227FDA3A4E358FFFA74" box="[514,755,1440,1464]" italics="true" pageId="28" pageNumber="1560">Papposilenus utriculus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A255F347E354E227FD5AA4E3595FFA74" box="[763,851,1440,1464]" pageId="28" pageNumber="1560" rank="species">sp. nov.</taxonomicNameLabel>
differ in few respects from the widely distributed Miocene
<taxonomicName id="4C12E9ADE354E227FDAAA48058B6FA17" baseAuthorityName="Gabbi" baseAuthorityYear="1873" box="[523,698,1475,1499]" class="Gastropoda" family="Potamididae" genus="Potamides" kingdom="Animalia" pageId="28" pageNumber="1560" phylum="Mollusca" rank="species" species="suprasulcatus">
<emphasis id="B9664E3CE354E227FDAAA48058B6FA17" box="[523,698,1475,1499]" italics="true" pageId="28" pageNumber="1560">P. suprasulcatus</emphasis>
</taxonomicName>
. The Eocene species usually has a more strongly opisthocyrt growth line, wider interspaces between the beaded cords, and beads that are more spirally elongate.
</paragraph>
</subSubSection>
<subSubSection id="C308C1A5E354E227FF01A72358A0F973" pageId="28" pageNumber="1560" type="etymology">
<paragraph id="8BAD922EE354E227FF01A7235B11F9B6" blockId="28.[160,1155,1632,1727]" box="[160,285,1632,1658]" pageId="28" pageNumber="1560">
<emphasis id="B9664E3CE354E227FF01A7235B11F9B6" bold="true" box="[160,285,1632,1658]" italics="true" pageId="28" pageNumber="1560">Etymology</emphasis>
</paragraph>
<paragraph id="8BAD922EE354E227FF01A7C758A0F973" blockId="28.[160,1155,1632,1727]" pageId="28" pageNumber="1560">
<emphasis id="B9664E3CE354E227FF01A7C75AAAF950" box="[160,166,1668,1692]" italics="true" pageId="28" pageNumber="1560">‘</emphasis>
Utriculus
<emphasis id="B9664E3CE354E227FEA8A7C75B03F950" box="[265,271,1668,1692]" italics="true" pageId="28" pageNumber="1560">’</emphasis>
, wineskin, a leather bag made from goat hide used to dispense wine; its profile is like that of the shell of
<taxonomicName id="4C12E9ADE354E227FDB9A7E458AAF973" authority="DeVries, 2019" authorityName="DeVries" authorityYear="2019" box="[536,678,1703,1727]" class="Gastropoda" family="Potamididae" genus="Papposilenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neotaenioglossa" pageId="28" pageNumber="1576" phylum="Mollusca" rank="genus" status="gen. nov.">
<emphasis id="B9664E3CE354E227FDB9A7E458AAF973" box="[536,678,1703,1727]" italics="true" pageId="28" pageNumber="1560">Papposilenus</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C308C1A5E355E226FF01A1CE5E88FEB4" pageId="29" pageNumber="1561" type="materials_examined">
<paragraph id="8BAD922EE355E226FF01A1CE5B0FFF6B" blockId="29.[160,1156,141,271]" box="[160,259,141,167]" pageId="29" pageNumber="1561">
<emphasis id="B9664E3CE355E226FF01A1CE5B0FFF6B" bold="true" box="[160,259,141,167]" italics="true" pageId="29" pageNumber="1561">Material</emphasis>
</paragraph>
<paragraph id="8BAD922EE355E226FF01A1F15E88FF20" blockId="29.[160,1156,141,271]" pageId="29" pageNumber="1561">
<materialsCitation id="3B7A9873E355E226FF01A1F15983FF06" ID-GBIF-Occurrence="2572160118" box="[160,911,178,202]" collectionCode="UWBM" pageId="29" pageNumber="1561" specimenCode="UWBM 107589" specimenCount="1" typeStatus="holotype">
<specimenCode id="DBB43A55E355E226FF01A1F15B47FF06" box="[160,331,178,202]" pageId="29" pageNumber="1561">UWBM 107589</specimenCode>
,
<typeStatus id="54A92C8CE355E226FEF8A1F15BCCFF06" box="[345,448,178,202]" pageId="29" pageNumber="1561" type="holotype">holotype</typeStatus>
, B8772 (type locality), L (25.5), W (15.0)
</materialsCitation>
;
<materialsCitation id="3B7A9873E355E226FC3FA1F158BFFF20" ID-GBIF-Occurrence="2572160117" collectionCode="UWBM" pageId="29" pageNumber="1561" specimenCode="UWBM 107590" specimenCount="1" typeStatus="paratype">
remainder are
<typeStatus id="54A92C8CE355E226FBE8A1F15AD3FF20" pageId="29" pageNumber="1561" type="paratype">paratypes</typeStatus>
:
<specimenCode id="DBB43A55E355E226FF4BA1975B9FFF20" box="[234,403,212,236]" pageId="29" pageNumber="1561">UWBM 107590</specimenCode>
, B8770, L (17.6), W (12.5);
</materialsCitation>
<materialsCitation id="3B7A9873E355E226FD1AA1975E88FF20" ID-GBIF-Occurrence="2565890709" box="[699,1156,212,236]" pageId="29" pageNumber="1561">
<collectionCode id="ED030AEBE355E226FD1AA197590BFF20" box="[699,775,212,236]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34878" lsid="urn:lsid:biocol.org:col:34878" name="University of Washington, Burke Museum" pageId="29" pageNumber="1561" type="Museum">UWBM</collectionCode>
107592, B8772, L (18.7), W (12.0);
</materialsCitation>
</paragraph>
<paragraph id="8BAD922EE355E226FF01A1B45E47FEC3" blockId="29.[160,1156,141,271]" box="[160,1099,247,271]" pageId="29" pageNumber="1561">
<materialsCitation id="3B7A9873E355E226FF01A1B4587DFEC3" ID-GBIF-Occurrence="2565890692" box="[160,625,247,271]" pageId="29" pageNumber="1561">
<collectionCode id="ED030AEBE355E226FF01A1B45AE5FEC3" box="[160,233,247,271]" pageId="29" pageNumber="1561">MUSM</collectionCode>
INV 253, B8770, L (25.3), W (12.9);
</materialsCitation>
<materialsCitation id="3B7A9873E355E226FDD8A1B45E47FEC3" ID-GBIF-Occurrence="2565890666" box="[633,1099,247,271]" pageId="29" pageNumber="1561">
<collectionCode id="ED030AEBE355E226FDD8A1B458CEFEC3" box="[633,706,247,271]" pageId="29" pageNumber="1561">MUSM</collectionCode>
INV 254, B8769, L (22.6), W (12.9).
</materialsCitation>
</paragraph>
<paragraph id="8BAD922EE355E226FF01A0785B28FE99" blockId="29.[160,1156,315,376]" box="[160,292,315,341]" pageId="29" pageNumber="1561">
<emphasis id="B9664E3CE355E226FF01A0785B28FE99" bold="true" box="[160,292,315,341]" italics="true" pageId="29" pageNumber="1561">Occurrence</emphasis>
</paragraph>
<paragraph id="8BAD922EE355E226FF01A0235E88FEB4" blockId="29.[160,1156,315,376]" box="[160,1156,352,376]" pageId="29" pageNumber="1561">
Lower Paleogene, Cuenca Member, Caballas Formation, East Pisco Basin, southern
<collectingCountry id="F305D2BEE355E226FBEAA0235E8DFEB4" box="[1099,1153,352,376]" name="Peru" pageId="29" pageNumber="1561">Peru</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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