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<document id="1F455FFDFA7A2CEE288885874F3EA72A" ID-CLB-Dataset="24019" ID-DOI="10.1080/00222933.2018.1524032" ID-GBIF-Dataset="034ebcd9-efa7-416a-822c-b6aee0e57a1c" ID-ISSN="1464-5262" ID-Zenodo-Dep="3670229" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1581941790231" checkinUser="valdenar" docAuthor="DeVries, Thomas J." docDate="2019" docId="03BB2338E341E234FE16A20D59B4FE3F" docLanguage="en" docName="JNATHIST.53.25–26.1533–1584.pdf.imf" docOrigin="Journal of Natural History 53 (25)" docStyle="DocumentStyle:485A7191CD41D6261F41521A3380DC73.9:JNatHist.2017-.journal_article.1cover" docStyleId="485A7191CD41D6261F41521A3380DC73" docStyleName="JNatHist.2017-.journal_article.1cover" docStyleVersion="9" docTitle="Ambigostrea sechura" docType="treatment" docVersion="23" lastPageNumber="1547" masterDocId="FF825B40E348E23BFFA1A1435A0CFFCC" masterDocTitle="Early Paleogene brackish-water molluscs from the Caballas Formation of the East Pisco Basin (Southern Peru)" masterLastPageNumber="1584" masterPageNumber="1533" pageNumber="1541" updateTime="1698783606523" updateUser="ExternalLinkService">
<mods:mods id="4DFEE128E0E9BB1CDBF36E3F499FC32B" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="E627394555CBBEF3C739AD134770721A">
<mods:title id="9531546A25566DBF247BD8FD0DD1A63C">Early Paleogene brackish-water molluscs from the Caballas Formation of the East Pisco Basin (Southern Peru)</mods:title>
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<mods:namePart id="1A2EF1AB28C3300134A2BD560FA39ECD">DeVries, Thomas J.</mods:namePart>
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<mods:title id="8B477C45C9EFFC049166CC703D4AAD76">Journal of Natural History</mods:title>
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<mods:part id="32E555B33E292458D2A7BAB241E48C1D">
<mods:date id="645C1A64CE3783B6DBE632B49FD57F22">2019</mods:date>
<mods:detail id="CD037CFCC4C1BA11BA3A531E5ED8FAE5" type="pubDate">
<mods:number id="FEF1B3D3E495300419AE5CAE8E2FA359">2018-12-17</mods:number>
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<mods:number id="952FE8F2CB666B7929D9C10FEB9C7984">53</mods:number>
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<mods:number id="CD7D5972854F8B94E58C79609DC5318B">25</mods:number>
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<mods:classification id="5DB7A95DD8BEC07D02772370235A6615">journal article</mods:classification>
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<mods:identifier id="AAC01611011503F7D5DC68730CB96DDB" type="DOI">10.1080/00222933.2018.1524032</mods:identifier>
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<treatment id="03BB2338E341E234FE16A20D59B4FE3F" ID-DOI="http://doi.org/10.5281/zenodo.3671184" ID-GBIF-Taxon="162146538" ID-Zenodo-Dep="3671184" LSID="urn:lsid:plazi:treatment:03BB2338E341E234FE16A20D59B4FE3F" httpUri="http://treatment.plazi.org/id/03BB2338E341E234FE16A20D59B4FE3F" lastPageId="15" lastPageNumber="1547" pageId="9" pageNumber="1541">
<subSubSection id="C308C1A5E341E232FE16A20D5960FCAB" box="[439,876,846,871]" pageId="9" pageNumber="1541" type="nomenclature">
<paragraph id="8BAD922EE341E232FE16A20D5960FCAB" blockId="9.[439,876,846,906]" box="[439,876,846,871]" pageId="9" pageNumber="1541">
<heading id="D0E52542E341E232FE16A20D5960FCAB" box="[439,876,846,871]" centered="true" fontSize="11" level="2" pageId="9" pageNumber="1541" reason="2">
<taxonomicName id="4C12E9ADE341E232FE16A20D5960FCAB" authority="(Olssoni 1944)" baseAuthorityName="Olssoni" baseAuthorityYear="1944" box="[439,876,846,871]" class="Bivalvia" family="Ostreidae" genus="Ambigostrea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ostreida" pageId="9" pageNumber="1541" phylum="Mollusca" rank="species" species="sechura">
<emphasis id="B9664E3CE341E232FE16A20D58B7FCAB" bold="true" box="[439,699,846,871]" italics="true" pageId="9" pageNumber="1541">Ambigostrea sechura</emphasis>
(Olsson, 1944)
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C308C1A5E341E232FDE1A23258EFFC46" box="[576,739,880,906]" pageId="9" pageNumber="1541" type="description">
<paragraph id="8BAD922EE341E232FDE1A23258EFFC46" blockId="9.[439,876,846,906]" box="[576,739,880,906]" pageId="9" pageNumber="1541">
<heading id="D0E52542E341E232FDE1A23258EFFC46" box="[576,739,880,906]" centered="true" fontSize="11" level="2" pageId="9" pageNumber="1541" reason="5">
(
<figureCitation id="13298EABE341E232FDE9A23258A0FC46" box="[584,684,881,906]" captionStart="Figure 2" captionStartId="7.[160,225,1097,1120]" captionTargetBox="[165,1151,144,1064]" captionTargetId="figure@7.[163,1153,142,1066]" captionTargetPageId="7" captionText="Figure 2. Bivalves from the Caballas Formation. (a–c) Incanopsis fastigiata sp. nov. (a) UWBM 107752, holotype, B8772, L 17.4 mm. Left valve, exterior. (b) UWBM 107752, cardinal area. (c) UWBM 107753, paratype, B8769, L 14.7 mm. Right valve, broken. (d, e) Brachidontes euglyphus (Woods, 1922). UWBM 107551, B8772, L 30.0 mm. Right valve, mould in part. (d) Exterior. (e) Exterior, anterior view. (f–s) Ambigostrea sechura (Olsson, 1944). (f) MUSM INV 235, B8769, L 55.6 mm. Left valve, exterior. (g) UWBM 107558, B8769, L 40.6 mm. Paired valves, posterior view. (h) MUSM INV 235, interior. (i) UWBM 107562, B8770, L 16.5 mm. Left valve, exterior. (j) UWBM 107562, right valve, exterior. (k) UWBM 107563, B8770, L 16.7 mm. Right valve, exterior. (l) UWBM 107568, B8772, L 26.0 mm. Paired valves, right valve resting on left valve. (m) UWBM 107554, B8769, L 76.8 mm. Left valve, interior. (n) MUSM INV 241, B8772, L 15.2 mm. Paired valves, right valve resting on left valve; with attached oyster spat. (o) UWBM 107565, B8772, L 24.6 mm. Left valve, interior. (p) UWBM 107555, B8769, L 52.2 mm. Right valve, exterior. (q) UWBM 107555, interior. (r) MUSM INV 238, B8769, L 29.7 mm. Left valve, interior. (s) MUSM INV 236, B8769, H 29.1 mm. Left valve, exterior, with twig mould in attachment scar. (t, u) undescribed gryphaeid oyster, Los Choros Member of the Paracas Formation, B8768. (t) UWBM 107642, H 49.0 mm. Right valve, interior showing vermiculate dorsoposterior margin. (u) UWBM 107641, H 65.2 mm (left to right in photo). Left valve, anterior view." figureDoi="http://doi.org/10.5281/zenodo.3670233" httpUri="https://zenodo.org/record/3670233/files/figure.png" pageId="9" pageNumber="1541">Figure 2</figureCitation>
(f
<emphasis id="B9664E3CE341E232FD1BA23358C4FC45" box="[698,712,880,905]" italics="true" pageId="9" pageNumber="1541">–</emphasis>
s))
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C308C1A5E341E232FF01A2E55968FC2C" pageId="9" pageNumber="1541" type="reference_group">
<paragraph id="8BAD922EE341E232FF01A2E5591CFC72" blockId="9.[160,1147,933,1027]" box="[160,784,933,958]" pageId="9" pageNumber="1541">
<taxonomicName id="4C12E9ADE341E232FF01A2E5583FFC72" authority="Olsson 1944" authorityName="Olssoni" authorityYear="1944" box="[160,563,934,958]" class="Bivalvia" family="Ostreidae" genus="Ostrea" kingdom="Animalia" order="Ostreoida" pageId="9" pageNumber="1541" phylum="Mollusca" rank="species" species="sechura" subGenus="Lopha">
<emphasis id="B9664E3CE341E232FF01A2E55AEBFC72" box="[160,231,934,958]" italics="true" pageId="9" pageNumber="1541">Ostrea</emphasis>
(
<emphasis id="B9664E3CE341E232FF56A2E55B37FC72" box="[247,315,934,958]" italics="true" pageId="9" pageNumber="1541">Lopha</emphasis>
)
<emphasis id="B9664E3CE341E232FEEAA2E55B93FC72" box="[331,415,934,958]" italics="true" pageId="9" pageNumber="1541">sechura</emphasis>
<bibRefCitation id="EF83EFDFE341E232FE09A2E5583FFC72" author="Olsson AA" box="[424,563,934,958]" pageId="9" pageNumber="1541" pagination="1 - 147" refId="ref25769" refString="Olsson AA. 1944. Contributions to the Tertiary paleontology of northern Peru. Part VII. The Cretaceous of the Paita region. Bulletins Am Paleontol. 19 (68): 1 - 147." type="journal article" year="1944">Olsson 1944</bibRefCitation>
</taxonomicName>
, 41, pl. 1, figs. 7, 8.
</paragraph>
<paragraph id="8BAD922EE341E232FF01A28B5968FC2C" blockId="9.[160,1147,933,1027]" box="[160,868,968,992]" pageId="9" pageNumber="1541">
<emphasis id="B9664E3CE341E232FF01A28B5AFEFC2C" box="[160,242,968,992]" italics="true" pageId="9" pageNumber="1541">‘Ostrea’</emphasis>
<taxonomicName id="4C12E9ADE341E232FF5BA28B5810FC2C" authority="Olssoni 1944" authorityName="Olssoni" authorityYear="1944" box="[250,540,968,992]" class="Bivalvia" family="Ostreidae" genus="Ostrea" kingdom="Animalia" order="Ostreoida" pageId="9" pageNumber="1541" phylum="Mollusca" rank="species" species="sechura">
[sic]
<emphasis id="B9664E3CE341E232FE8EA28B5B8FFC2C" box="[303,387,968,992]" italics="true" pageId="9" pageNumber="1541">sechura</emphasis>
Olsson, 1944
</taxonomicName>
.
<bibRefCitation id="EF83EFDFE341E232FD8AA28B58CBFC2C" author="Alleman V" box="[555,711,968,992]" pageId="9" pageNumber="1541" pagination="67 - 71" refId="ref21624" refString="Alleman V. 1995. Lamelibranquios tipicos des Senoniano superior de la zona Baculitesi de Tortuga (Paitai Depto. de Piurai Peru). Biotempo. 2: 67 - 71." type="journal article" year="1995">
Alleman
<date id="FFACB4EEE341E232FD31A28B58CBFC2C" box="[656,711,968,992]" pageId="9" pageNumber="1541" value="1995-01">1995</date>
</bibRefCitation>
, 68, figs. 5, 6.
</paragraph>
</subSubSection>
<subSubSection id="C308C1A5E341E232FF01A2A85E77FBCF" box="[160,1147,1003,1027]" pageId="9" pageNumber="1541" type="description">
<paragraph id="8BAD922EE341E232FF01A2A85E77FBCF" blockId="9.[160,1147,933,1027]" box="[160,1147,1003,1027]" pageId="9" pageNumber="1541">
? [sic]
<taxonomicName id="4C12E9ADE341E232FF49A2A85864FBCF" authority="Olssoni 1944" authorityName="Olssoni" authorityYear="1944" box="[232,616,1003,1027]" class="Bivalvia" family="Ostreidae" genus="Ambigostrea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ostreida" pageId="9" pageNumber="1541" phylum="Mollusca" rank="species" species="sechura">
<emphasis id="B9664E3CE341E232FF49A2A85BC3FBCF" box="[232,463,1003,1027]" italics="true" pageId="9" pageNumber="1541">Ambigostrea sechura</emphasis>
Olsson, 1944
</taxonomicName>
. Dhondt and Jaillard
<date id="FFACB4EEE341E232FCFEA2A85990FBCF" box="[863,924,1003,1027]" pageId="9" pageNumber="1541" value="2005-01">2005,</date>
339, pl. 1, figs 1, 2.
</paragraph>
</subSubSection>
<subSubSection id="C308C1A5E341E236FF01A5765BB4FE12" lastPageId="13" lastPageNumber="1545" pageId="9" pageNumber="1541" type="discussion">
<paragraph id="8BAD922EE341E232FF01A5765B0AFB83" blockId="9.[160,1156,1077,1727]" box="[160,262,1077,1103]" pageId="9" pageNumber="1541">
<emphasis id="B9664E3CE341E232FF01A5765B0AFB83" bold="true" box="[160,262,1077,1103]" italics="true" pageId="9" pageNumber="1541">Remarks</emphasis>
</paragraph>
<paragraph id="8BAD922EE341E232FF01A519593FFB31" blockId="9.[160,1156,1077,1727]" pageId="9" pageNumber="1541">
<taxonomicName id="4C12E9ADE341E232FF01A5195848FBBE" authority="Olssoni 1944" authorityName="Olssoni" authorityYear="1944" box="[160,580,1114,1138]" class="Bivalvia" family="Ostreidae" genus="Ostrea" kingdom="Animalia" order="Ostreoida" pageId="9" pageNumber="1541" phylum="Mollusca" rank="species" species="sechura" subGenus="Lopha">
<emphasis id="B9664E3CE341E232FF01A5195AEBFBBE" box="[160,231,1114,1138]" italics="true" pageId="9" pageNumber="1541">Ostrea</emphasis>
(
<emphasis id="B9664E3CE341E232FF5BA5195B32FBBE" box="[250,318,1114,1138]" italics="true" pageId="9" pageNumber="1541">Lopha</emphasis>
)
<emphasis id="B9664E3CE341E232FEF0A5195BA9FBBE" box="[337,421,1114,1138]" italics="true" pageId="9" pageNumber="1541">sechura</emphasis>
Olsson, 1944
</taxonomicName>
; was assigned to
<taxonomicName id="4C12E9ADE341E232FCBCA5195E5BFBBE" authority="Malchusi 1990" authorityName="Malchusi" authorityYear="1990" box="[797,1111,1114,1138]" class="Bivalvia" family="Ostreidae" genus="Ambigostrea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ostreida" pageId="9" pageNumber="1541" phylum="Mollusca" rank="genus">
<emphasis id="B9664E3CE341E232FCBCA51959AAFBBE" box="[797,934,1114,1138]" italics="true" pageId="9" pageNumber="1541">Ambigostrea</emphasis>
Malchus, 1990
</taxonomicName>
; by Dhondt and Jaillard (2005); but with some hesitation for want of a study of shell microstructure of the Peruvian specimens. An examination of nearly 100 Caballas Formation specimens supports the assignment to
<taxonomicName id="4C12E9ADE341E232FD7DA5815969FB16" authorityName="Malchusi" authorityYear="1990" box="[732,869,1218,1242]" class="Bivalvia" family="Ostreidae" genus="Ambigostrea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ostreida" pageId="9" pageNumber="1541" phylum="Mollusca" rank="genus">
<emphasis id="B9664E3CE341E232FD7DA5815969FB16" box="[732,869,1218,1242]" italics="true" pageId="9" pageNumber="1541">Ambigostrea</emphasis>
</taxonomicName>
based on shell morphology and the presence of a simple foliated microstructure.
</paragraph>
<paragraph id="8BAD922EE341E232FF61A4445824F99B" blockId="9.[160,1156,1077,1727]" pageId="9" pageNumber="1541">
Full-sized left valves of Caballas Formation oysters exhibit the distinctive characters seen on left valves of
<taxonomicName id="4C12E9ADE341E232FE0DA4695899FA8E" authorityName="Olssoni" authorityYear="1944" box="[428,661,1322,1346]" class="Bivalvia" family="Ostreidae" genus="Ambigostrea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ostreida" pageId="9" pageNumber="1541" phylum="Mollusca" rank="species" species="sechura">
<emphasis id="B9664E3CE341E232FE0DA4695899FA8E" box="[428,661,1322,1346]" italics="true" pageId="9" pageNumber="1541">Ambigostrea sechura</emphasis>
</taxonomicName>
: a sickle shape; strong radiating chevronshaped ribs, which are few in number; and posterior and anterior margins crossed transversely away from the commissural plane by closely spaced corrugations, usually near the umbones but in large specimens extending as far as the hooked ventral margin. The corrugations are produced by the stacking of closely spaced marginal crenulations in successively older shell layers. The marginal crenulations are produced near the umbones by the overlap of chomata with the margin and farther from the umbones by marginal crenulations that are several times smaller and at oblique or right angles to the radiating chevron-shaped ribs.
</paragraph>
<paragraph id="8BAD922EE341E231FF61A7215BBAFEFD" blockId="9.[160,1156,1077,1727]" lastBlockId="10.[160,1156,142,1726]" lastPageId="10" lastPageNumber="1542" pageId="9" pageNumber="1541">
With more material available than was known to
<bibRefCitation id="EF83EFDFE341E232FD4CA721598BF9B6" author="Olsson AA" box="[749,903,1634,1658]" pageId="9" pageNumber="1541" pagination="1 - 147" refId="ref25769" refString="Olsson AA. 1944. Contributions to the Tertiary paleontology of northern Peru. Part VII. The Cretaceous of the Paita region. Bulletins Am Paleontol. 19 (68): 1 - 147." type="journal article" year="1944">Olsson (1944)</bibRefCitation>
, further description of the left valve is possible. The adductor muscle scar has a skewed U-shape, with the posterior edge extending farther towards the hinge than the anterior edge. The entire ventral edge of the adductor scar is raised above the floor of the valve and bordered by a myophoric buttress. The chevron-shaped radial ribs begin directly beyond the ventral edge of the attachment scar. The transverse corrugations appear on juveniles, but only close to the umbones. Chomata extend for variable distances ventrally on the posterior and/or anterior margins.
</paragraph>
<paragraph id="8BAD922EE342E231FF61A07F5E05FE12" blockId="10.[160,1156,142,1726]" pageId="10" pageNumber="1542">
The right valve of
<taxonomicName id="4C12E9ADE342E231FE26A07F586BFE98" authorityName="Olssoni" authorityYear="1944" box="[391,615,316,340]" class="Bivalvia" family="Ostreidae" genus="Ambigostrea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ostreida" pageId="10" pageNumber="1542" phylum="Mollusca" rank="species" species="sechura">
<emphasis id="B9664E3CE342E231FE26A07F586BFE98" box="[391,615,316,340]" italics="true" pageId="10" pageNumber="1542">Ambigostrea sechura</emphasis>
</taxonomicName>
is usually but not invariably less convex than the left valve and usually but not invariably has weaker radiating ribs. The adductor scar is U-shaped and swollen above the floor of the valve. Transverse corrugations are developed to the extent permitted by the shell thickness and appear on juvenile specimens near the umbones and on larger specimens at least as far ventrally as the adductor scar.
</paragraph>
<paragraph id="8BAD922EE342E231FF61A0AA599FFD1D" blockId="10.[160,1156,142,1726]" pageId="10" pageNumber="1542">
Specimens of the Argentinian Maastrichtian oyster,
<taxonomicName id="4C12E9ADE342E231FCA2A0AA5E73FDCD" authority="(Iheringi 1907)" baseAuthorityName="Iheringi" baseAuthorityYear="1907" box="[771,1151,489,513]" class="Bivalvia" family="Ostreidae" genus="Ambigostrea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ostreida" pageId="10" pageNumber="1542" phylum="Mollusca" rank="species" species="clarae">
<emphasis id="B9664E3CE342E231FCA2A0AA59D9FDCD" box="[771,981,489,513]" italics="true" pageId="10" pageNumber="1542">Ambigostrea clarae</emphasis>
(Ihering, 1907)
</taxonomicName>
, exhibit the sickle shape, comma-shaped posterior adductor muscle scar, transverse corrugations, and bifurcating radial chevron-shaped ribs on right and left valves seen on specimens of
<taxonomicName id="4C12E9ADE342E231FED1A3125892FDA5" authority="(Casadio 1998)" baseAuthorityName="Casadio" baseAuthorityYear="1998" box="[368,670,593,617]" class="Bivalvia" family="Ostreidae" genus="Ambigostrea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ostreida" pageId="10" pageNumber="1542" phylum="Mollusca" rank="species" species="sechura">
<emphasis id="B9664E3CE342E231FED1A3125BEAFDA5" box="[368,486,593,617]" italics="true" pageId="10" pageNumber="1542">A. sechura</emphasis>
(Casadío 1998)
</taxonomicName>
. The chomata of
<taxonomicName id="4C12E9ADE342E231FCD6A31259D7FDA5" baseAuthorityName="Iheringi" baseAuthorityYear="1907" box="[887,987,593,617]" class="Bivalvia" family="Ostreidae" genus="Ambigostrea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ostreida" pageId="10" pageNumber="1542" phylum="Mollusca" rank="species" species="clarae">
<emphasis id="B9664E3CE342E231FCD6A31259D7FDA5" box="[887,987,593,617]" italics="true" pageId="10" pageNumber="1542">A. clarae</emphasis>
</taxonomicName>
, however, are always restricted to the dorsal half of the posterior and anterior margins, which on some specimens of
<taxonomicName id="4C12E9ADE342E231FEE6A3D55BB1FD62" baseAuthorityName="Casadio" baseAuthorityYear="1998" box="[327,445,662,686]" class="Bivalvia" family="Ostreidae" genus="Ambigostrea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ostreida" pageId="10" pageNumber="1542" phylum="Mollusca" rank="species" species="sechura">
<emphasis id="B9664E3CE342E231FEE6A3D55BB1FD62" box="[327,445,662,686]" italics="true" pageId="10" pageNumber="1542">A. sechura</emphasis>
</taxonomicName>
(
<figureCitation id="13298EABE342E231FE73A3D55839FD62" box="[466,565,662,686]" captionStart="Figure 2" captionStartId="7.[160,225,1097,1120]" captionTargetBox="[165,1151,144,1064]" captionTargetId="figure@7.[163,1153,142,1066]" captionTargetPageId="7" captionText="Figure 2. Bivalves from the Caballas Formation. (a–c) Incanopsis fastigiata sp. nov. (a) UWBM 107752, holotype, B8772, L 17.4 mm. Left valve, exterior. (b) UWBM 107752, cardinal area. (c) UWBM 107753, paratype, B8769, L 14.7 mm. Right valve, broken. (d, e) Brachidontes euglyphus (Woods, 1922). UWBM 107551, B8772, L 30.0 mm. Right valve, mould in part. (d) Exterior. (e) Exterior, anterior view. (f–s) Ambigostrea sechura (Olsson, 1944). (f) MUSM INV 235, B8769, L 55.6 mm. Left valve, exterior. (g) UWBM 107558, B8769, L 40.6 mm. Paired valves, posterior view. (h) MUSM INV 235, interior. (i) UWBM 107562, B8770, L 16.5 mm. Left valve, exterior. (j) UWBM 107562, right valve, exterior. (k) UWBM 107563, B8770, L 16.7 mm. Right valve, exterior. (l) UWBM 107568, B8772, L 26.0 mm. Paired valves, right valve resting on left valve. (m) UWBM 107554, B8769, L 76.8 mm. Left valve, interior. (n) MUSM INV 241, B8772, L 15.2 mm. Paired valves, right valve resting on left valve; with attached oyster spat. (o) UWBM 107565, B8772, L 24.6 mm. Left valve, interior. (p) UWBM 107555, B8769, L 52.2 mm. Right valve, exterior. (q) UWBM 107555, interior. (r) MUSM INV 238, B8769, L 29.7 mm. Left valve, interior. (s) MUSM INV 236, B8769, H 29.1 mm. Left valve, exterior, with twig mould in attachment scar. (t, u) undescribed gryphaeid oyster, Los Choros Member of the Paracas Formation, B8768. (t) UWBM 107642, H 49.0 mm. Right valve, interior showing vermiculate dorsoposterior margin. (u) UWBM 107641, H 65.2 mm (left to right in photo). Left valve, anterior view." figureDoi="http://doi.org/10.5281/zenodo.3670233" httpUri="https://zenodo.org/record/3670233/files/figure.png" pageId="10" pageNumber="1542">Figure 2</figureCitation>
(h)) is not the case, and the radial ribs are more numerous (8
<emphasis id="B9664E3CE342E231FE91A3FA5B31FD1D" box="[304,317,697,721]" italics="true" pageId="10" pageNumber="1542">–</emphasis>
13 ribs) than on specimens of
<taxonomicName id="4C12E9ADE342E231FD3AA3FA5900FD1D" baseAuthorityName="Casadio" baseAuthorityYear="1998" box="[667,780,697,721]" class="Bivalvia" family="Ostreidae" genus="Ambigostrea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ostreida" pageId="10" pageNumber="1542" phylum="Mollusca" rank="species" species="sechura">
<emphasis id="B9664E3CE342E231FD3AA3FA5900FD1D" box="[667,780,697,721]" italics="true" pageId="10" pageNumber="1542">A. sechura</emphasis>
</taxonomicName>
(5
<emphasis id="B9664E3CE342E231FC8BA3FA593BFD1D" box="[810,823,697,721]" italics="true" pageId="10" pageNumber="1542">–</emphasis>
10 ribs).
</paragraph>
<paragraph id="8BAD922EE342E231FF61A3985B3BFC6D" blockId="10.[160,1156,142,1726]" pageId="10" pageNumber="1542">
A few oyster specimens from the
<collectingCountry id="F305D2BEE342E231FDE3A3985896FD3F" box="[578,666,731,755]" name="Haiti" pageId="10" pageNumber="1542">Tortuga</collectingCountry>
and La Mesa formations were assigned by Dhondt and Jaillard (2005) to the finely ribbed
<taxonomicName id="4C12E9ADE342E231FD77A3BD5E6AFCDA" authority="(Coquandi 1862)" baseAuthorityName="Coquandi" baseAuthorityYear="1862" box="[726,1126,766,790]" class="Bivalvia" family="Ostreidae" genus="Ambigostrea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ostreida" pageId="10" pageNumber="1542" phylum="Mollusca" rank="species" species="villei">
<emphasis id="B9664E3CE342E231FD77A3BD5990FCDA" box="[726,924,766,790]" italics="true" pageId="10" pageNumber="1542">Ambigostrea villei</emphasis>
(Coquand, 1862)
</taxonomicName>
, a Campanian
<emphasis id="B9664E3CE342E231FEBEA2625B20FCF5" box="[287,300,801,825]" italics="true" pageId="10" pageNumber="1542">–</emphasis>
Maastrichtian species known throughout North Africa and the Middle East (
<bibRefCitation id="EF83EFDFE342E231FF07A2005B4FFC97" author="Malchus N" box="[166,323,835,859]" pageId="10" pageNumber="1542" pagination="1 - 231" refId="ref25063" refString="Malchus N. 1990. Revision der Kreide-Austern (Bivalvia: Pteriomorphia) Agyptens (Biostratigraphiei Systematik). Berliner Geowissenschaftliche Abhandlungen (A). 125: 1 - 231." type="journal article" year="1990">Malchus 1990</bibRefCitation>
). The specimens from the Caballas Formation, however, closely resemble the coarsely ribbed specimens from the
<collectingCountry id="F305D2BEE342E231FDCBA22558CEFCB2" box="[618,706,870,894]" name="Haiti" pageId="10" pageNumber="1542">Tortuga</collectingCountry>
and La Mesa formations, i.e. specimens of
<taxonomicName id="4C12E9ADE342E231FF1EA2CA5B3DFC6D" baseAuthorityName="Casadio" baseAuthorityYear="1998" box="[191,305,905,929]" class="Bivalvia" family="Ostreidae" genus="Ambigostrea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ostreida" pageId="10" pageNumber="1542" phylum="Mollusca" rank="species" species="sechura">
<emphasis id="B9664E3CE342E231FF1EA2CA5B3DFC6D" box="[191,305,905,929]" italics="true" pageId="10" pageNumber="1542">A. sechura</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8BAD922EE342E231FF61A2E85AF4FB7A" blockId="10.[160,1156,142,1726]" pageId="10" pageNumber="1542">
Species of
<taxonomicName id="4C12E9ADE342E231FE98A2E85832FC0F" authority="Saccoi 1897" authorityName="Saccoi" authorityYear="1897" box="[313,574,939,963]" class="Bivalvia" family="Ostreidae" genus="Cubitostrea" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Ostreida" pageId="10" pageNumber="1542" phylum="Mollusca" rank="genus">
<emphasis id="B9664E3CE342E231FE98A2E85BBFFC0F" box="[313,435,939,963]" italics="true" pageId="10" pageNumber="1542">Cubitostrea</emphasis>
Sacco, 1897
</taxonomicName>
, a genus that ranges from the Maastrichtian to late Miocene in South America (
<bibRefCitation id="EF83EFDFE342E231FE69A28D5873FC2A" author="Griffin Mi Casadio Si Parras A" box="[456,639,974,998]" pageId="10" pageNumber="1542" pagination="257 - 276" refId="ref24097" refString="Griffin Mi Casadio Si Parras A. 2005. Maastrichtian and Danian species of Turkostreini (Ostreidaei Crassostreinae) from the Neuquen Basini Argentina. Ameghiniana. 42 (2): 257 - 276." type="journal article" year="2005">Griffin et al. 2005</bibRefCitation>
), may exhibit the sickle-shaped, strongly ribbed form of
<taxonomicName id="4C12E9ADE342E231FF5BA2B25BD6FBC5" baseAuthorityName="Casadio" baseAuthorityYear="1998" box="[250,474,1009,1033]" class="Bivalvia" family="Ostreidae" genus="Ambigostrea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ostreida" pageId="10" pageNumber="1542" phylum="Mollusca" rank="species" species="sechura">
<emphasis id="B9664E3CE342E231FF5BA2B25BD6FBC5" box="[250,474,1009,1033]" italics="true" pageId="10" pageNumber="1542">Ambigostrea sechura</emphasis>
</taxonomicName>
, and some species, e.g.
<taxonomicName id="4C12E9ADE342E231FD46A2B25942FBC5" baseAuthorityName="Deshayes" baseAuthorityYear="1832" box="[743,846,1009,1033]" class="Bivalvia" family="Ostreidae" genus="Cubitostrea" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Ostreida" pageId="10" pageNumber="1542" phylum="Mollusca" rank="species" species="cubitus">
<emphasis id="B9664E3CE342E231FD46A2B25942FBC5" box="[743,846,1009,1033]" italics="true" pageId="10" pageNumber="1542">C. cubitus</emphasis>
</taxonomicName>
(Deshayes,
<date id="FFACB4EEE342E231FC73A2B25E47FBC5" box="[978,1099,1009,1033]" pageId="10" pageNumber="1542" value="1824" valueMax="1837">
1824
<emphasis id="B9664E3CE342E231FBA9A2B25E19FBC5" box="[1032,1045,1009,1033]" italics="true" pageId="10" pageNumber="1542">–</emphasis>
1837
</date>
), the
<typeStatus id="54A92C8CE342E231FF01A5505ADDFBE7" box="[160,209,1043,1067]" pageId="10" pageNumber="1542">type</typeStatus>
species of
<taxonomicName id="4C12E9ADE342E231FEC5A5505BD1FBE7" authorityName="Saccoi" authorityYear="1897" box="[356,477,1043,1067]" class="Bivalvia" family="Ostreidae" genus="Cubitostrea" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Ostreida" pageId="10" pageNumber="1542" phylum="Mollusca" rank="genus">
<emphasis id="B9664E3CE342E231FEC5A5505BD1FBE7" box="[356,477,1043,1067]" italics="true" pageId="10" pageNumber="1542">Cubitostrea</emphasis>
</taxonomicName>
, and
<taxonomicName id="4C12E9ADE342E231FD8BA5505E6EFBE7" authority="Griffini Casadioi and Parrasi 2005" authorityName="Griffini Casadioi and Parrasi" authorityYear="2005" box="[554,1122,1043,1067]" class="Bivalvia" family="Ostreidae" genus="Cubitostrea" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Ostreida" pageId="10" pageNumber="1542" phylum="Mollusca" rank="species" species="primordialis">
<emphasis id="B9664E3CE342E231FD8BA55058C0FBE7" box="[554,716,1043,1067]" italics="true" pageId="10" pageNumber="1542">C. primordialis</emphasis>
Griffin, Casadío, and Parras, 2005
</taxonomicName>
, a Maastrichtian species from Argentina, also exhibit closely spaced transverse corrugations like those of
<taxonomicName id="4C12E9ADE342E231FE97A51A5BABFBBD" baseAuthorityName="Casadio" baseAuthorityYear="1998" box="[310,423,1113,1137]" class="Bivalvia" family="Ostreidae" genus="Ambigostrea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ostreida" pageId="10" pageNumber="1542" phylum="Mollusca" rank="species" species="sechura">
<emphasis id="B9664E3CE342E231FE97A51A5BABFBBD" box="[310,423,1113,1137]" italics="true" pageId="10" pageNumber="1542">A. sechura</emphasis>
</taxonomicName>
. The right valve of
<taxonomicName id="4C12E9ADE342E231FD2BA51A5908FBBD" authorityName="Saccoi" authorityYear="1897" box="[650,772,1113,1137]" class="Bivalvia" family="Ostreidae" genus="Cubitostrea" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Ostreida" pageId="10" pageNumber="1542" phylum="Mollusca" rank="genus">
<emphasis id="B9664E3CE342E231FD2BA51A5908FBBD" box="[650,772,1113,1137]" italics="true" pageId="10" pageNumber="1542">Cubitostrea</emphasis>
</taxonomicName>
taxa, however, is entirely smooth (
<bibRefCitation id="EF83EFDFE342E231FF07A5385B60FB5F" author="Griffin Mi Casadio Si Parras A" box="[166,364,1147,1171]" pageId="10" pageNumber="1542" pagination="257 - 276" refId="ref24097" refString="Griffin Mi Casadio Si Parras A. 2005. Maastrichtian and Danian species of Turkostreini (Ostreidaei Crassostreinae) from the Neuquen Basini Argentina. Ameghiniana. 42 (2): 257 - 276." type="journal article" year="2005">Griffin et al. 2005</bibRefCitation>
), lacking the chevron-shaped radiating ribs seen on right valves of
<taxonomicName id="4C12E9ADE342E231FBCEA5385AFEFB7A" baseAuthorityName="Casadio" baseAuthorityYear="1998" class="Bivalvia" family="Ostreidae" genus="Ambigostrea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ostreida" pageId="10" pageNumber="1542" phylum="Mollusca" rank="species" species="sechura">
<emphasis id="B9664E3CE342E231FBCEA5385AFEFB7A" italics="true" pageId="10" pageNumber="1542">A. sechura</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8BAD922EE342E231FF61A5825B71FA22" blockId="10.[160,1156,142,1726]" pageId="10" pageNumber="1542">
Specimens of
<taxonomicName id="4C12E9ADE342E231FEC3A5825845FB15" baseAuthorityName="Casadio" baseAuthorityYear="1998" box="[354,585,1217,1241]" class="Bivalvia" family="Ostreidae" genus="Ambigostrea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ostreida" pageId="10" pageNumber="1542" phylum="Mollusca" rank="species" species="sechura">
<emphasis id="B9664E3CE342E231FEC3A5825845FB15" box="[354,585,1217,1241]" italics="true" pageId="10" pageNumber="1542">Ambigostrea sechura</emphasis>
</taxonomicName>
differ from equally large specimens of the upper lower Eocene
<taxonomicName id="4C12E9ADE342E231FE9EA5A05852FB37" authority="Woodsi 1922" authorityName="Woodsi" authorityYear="1922" box="[319,606,1251,1275]" class="Bivalvia" family="Ostreidae" genus="Ostrea" kingdom="Animalia" order="Ostreoida" pageId="10" pageNumber="1542" phylum="Mollusca" rank="species" species="buski">
<emphasis id="B9664E3CE342E231FE9EA5A05BC9FB37" box="[319,453,1251,1275]" italics="true" pageId="10" pageNumber="1542">Ostrea buski</emphasis>
Woods, 1922
</taxonomicName>
, from northern
<collectingCountry id="F305D2BEE342E231FCB3A5A05949FB37" box="[786,837,1251,1275]" name="Peru" pageId="10" pageNumber="1542">Peru</collectingCountry>
(
<bibRefCitation id="EF83EFDFE342E231FCF3A5A059ECFB37" author="Woods H" box="[850,992,1251,1275]" pageId="10" pageNumber="1542" pagination="51 - 113" refId="ref27434" refString="Woods H. 1922. Mollusca from Eocene and Miocene deposits of Peru. In: Bosworth TOi editor. Geology of the Tertiary and Quaternary periods in the northern part of Peru. London: Macmillan; p. 51 - 113." type="book chapter" year="1922">
Woods
<date id="FFACB4EEE342E231FC08A5A059ECFB37" box="[937,992,1251,1275]" pageId="10" pageNumber="1542" value="1922-01">1922</date>
</bibRefCitation>
, figs. 3
<emphasis id="B9664E3CE342E231FB8FA5A05E37FB37" box="[1070,1083,1251,1275]" italics="true" pageId="10" pageNumber="1542">–</emphasis>
5) and other specimens of
<taxonomicName id="4C12E9ADE342E231FE01A445580EFAD2" authorityName="Woodsi" authorityYear="1922" box="[416,514,1286,1310]" class="Bivalvia" family="Ostreidae" genus="Ostrea" kingdom="Animalia" order="Ostreoida" pageId="10" pageNumber="1542" phylum="Mollusca" rank="species" species="buski">
<emphasis id="B9664E3CE342E231FE01A445580EFAD2" box="[416,514,1286,1310]" italics="true" pageId="10" pageNumber="1542">O. buski</emphasis>
</taxonomicName>
examined in Olsson
<emphasis id="B9664E3CE342E231FCA9A4455901FAD2" box="[776,781,1286,1310]" italics="true" pageId="10" pageNumber="1542">’</emphasis>
s undescribed material at the Paleontological Research Institution and the Smithsonian National Museum of Natural History. Large specimens of
<taxonomicName id="4C12E9ADE342E231FE43A4085836FAAF" authorityName="Woodsi" authorityYear="1922" box="[482,570,1355,1379]" class="Bivalvia" family="Ostreidae" genus="Ostrea" kingdom="Animalia" order="Ostreoida" pageId="10" pageNumber="1542" phylum="Mollusca" rank="species" species="buski">
<emphasis id="B9664E3CE342E231FE43A4085836FAAF" box="[482,570,1355,1379]" italics="true" pageId="10" pageNumber="1542">O. buski</emphasis>
</taxonomicName>
are fan-shaped; usually have smaller, more numerous chevron-shaped radiating ribs (15
<emphasis id="B9664E3CE342E231FDE8A42D585AFA4A" box="[585,598,1390,1414]" italics="true" pageId="10" pageNumber="1542">–</emphasis>
20 ribs, compared with 5
<emphasis id="B9664E3CE342E231FCCDA42D5975FA4A" box="[876,889,1390,1414]" italics="true" pageId="10" pageNumber="1542">–</emphasis>
10 ribs on specimens of
<taxonomicName id="4C12E9ADE342E231FF01A4D25B1CFA65" baseAuthorityName="Casadio" baseAuthorityYear="1998" box="[160,272,1425,1449]" class="Bivalvia" family="Ostreidae" genus="Ambigostrea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ostreida" pageId="10" pageNumber="1542" phylum="Mollusca" rank="species" species="sechura">
<emphasis id="B9664E3CE342E231FF01A4D25B1CFA65" box="[160,272,1425,1449]" italics="true" pageId="10" pageNumber="1542">A. sechura</emphasis>
</taxonomicName>
); have anterior and posterior margins near the umbones of both valves scored by deeply scooped chomata; and lack the transverse corrugations seen on most specimens of
<taxonomicName id="4C12E9ADE342E231FEA4A4955B7BFA22" baseAuthorityName="Casadio" baseAuthorityYear="1998" box="[261,375,1494,1518]" class="Bivalvia" family="Ostreidae" genus="Ambigostrea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ostreida" pageId="10" pageNumber="1542" phylum="Mollusca" rank="species" species="sechura">
<emphasis id="B9664E3CE342E231FEA4A4955B7BFA22" box="[261,375,1494,1518]" italics="true" pageId="10" pageNumber="1542">A. sechura</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8BAD922EE342E236FF61A4BA5BDFFE98" blockId="10.[160,1156,142,1726]" lastBlockId="13.[160,1156,142,478]" lastPageId="13" lastPageNumber="1545" pageId="10" pageNumber="1542">
Inland from Puerto Caballas, in the lower part of the Los Choros Member of the Paracas Formation (DeVries 2017), specimens are found of an undescribed oyster equally large and similarly sculpted as the more ancient
<taxonomicName id="4C12E9ADE342E231FD70A77D59B4F99A" baseAuthorityName="Casadio" baseAuthorityYear="1998" box="[721,952,1598,1622]" class="Bivalvia" family="Ostreidae" genus="Ambigostrea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ostreida" pageId="10" pageNumber="1542" phylum="Mollusca" rank="species" species="sechura">
<emphasis id="B9664E3CE342E231FD70A77D59B4F99A" box="[721,952,1598,1622]" italics="true" pageId="10" pageNumber="1542">Ambigostrea sechura</emphasis>
</taxonomicName>
(
<figureCitation id="13298EABE342E231FC69A77D5E2BF99A" box="[968,1063,1598,1622]" captionStart="Figure 2" captionStartId="7.[160,225,1097,1120]" captionTargetBox="[165,1151,144,1064]" captionTargetId="figure@7.[163,1153,142,1066]" captionTargetPageId="7" captionText="Figure 2. Bivalves from the Caballas Formation. (a–c) Incanopsis fastigiata sp. nov. (a) UWBM 107752, holotype, B8772, L 17.4 mm. Left valve, exterior. (b) UWBM 107752, cardinal area. (c) UWBM 107753, paratype, B8769, L 14.7 mm. Right valve, broken. (d, e) Brachidontes euglyphus (Woods, 1922). UWBM 107551, B8772, L 30.0 mm. Right valve, mould in part. (d) Exterior. (e) Exterior, anterior view. (f–s) Ambigostrea sechura (Olsson, 1944). (f) MUSM INV 235, B8769, L 55.6 mm. Left valve, exterior. (g) UWBM 107558, B8769, L 40.6 mm. Paired valves, posterior view. (h) MUSM INV 235, interior. (i) UWBM 107562, B8770, L 16.5 mm. Left valve, exterior. (j) UWBM 107562, right valve, exterior. (k) UWBM 107563, B8770, L 16.7 mm. Right valve, exterior. (l) UWBM 107568, B8772, L 26.0 mm. Paired valves, right valve resting on left valve. (m) UWBM 107554, B8769, L 76.8 mm. Left valve, interior. (n) MUSM INV 241, B8772, L 15.2 mm. Paired valves, right valve resting on left valve; with attached oyster spat. (o) UWBM 107565, B8772, L 24.6 mm. Left valve, interior. (p) UWBM 107555, B8769, L 52.2 mm. Right valve, exterior. (q) UWBM 107555, interior. (r) MUSM INV 238, B8769, L 29.7 mm. Left valve, interior. (s) MUSM INV 236, B8769, H 29.1 mm. Left valve, exterior, with twig mould in attachment scar. (t, u) undescribed gryphaeid oyster, Los Choros Member of the Paracas Formation, B8768. (t) UWBM 107642, H 49.0 mm. Right valve, interior showing vermiculate dorsoposterior margin. (u) UWBM 107641, H 65.2 mm (left to right in photo). Left valve, anterior view." figureDoi="http://doi.org/10.5281/zenodo.3670233" httpUri="https://zenodo.org/record/3670233/files/figure.png" pageId="10" pageNumber="1542">Figure 2</figureCitation>
(u)). The left valve of the Los Choros Member species, however, is deeply arched and the attachment scar is several centimetres long, rather than less than two centimetres long, which is the case even for large specimens of
<taxonomicName id="4C12E9ADE342E231FD5AA7E55961F972" baseAuthorityName="Casadio" baseAuthorityYear="1998" box="[763,877,1702,1726]" class="Bivalvia" family="Ostreidae" genus="Ambigostrea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ostreida" pageId="10" pageNumber="1542" phylum="Mollusca" rank="species" species="sechura">
<emphasis id="B9664E3CE342E231FD5AA7E55961F972" box="[763,877,1702,1726]" italics="true" pageId="10" pageNumber="1542">A. sechura</emphasis>
</taxonomicName>
. The posterior adductor muscle scar is circular and barely raised above the floor of the valve. The posterodorsal margin is lined with vermiculate chomata. A right valve (
<figureCitation id="13298EABE345E236FCF4A1F259B4FF05" box="[853,952,177,201]" captionStart="Figure 2" captionStartId="7.[160,225,1097,1120]" captionTargetBox="[165,1151,144,1064]" captionTargetId="figure@7.[163,1153,142,1066]" captionTargetPageId="7" captionText="Figure 2. Bivalves from the Caballas Formation. (a–c) Incanopsis fastigiata sp. nov. (a) UWBM 107752, holotype, B8772, L 17.4 mm. Left valve, exterior. (b) UWBM 107752, cardinal area. (c) UWBM 107753, paratype, B8769, L 14.7 mm. Right valve, broken. (d, e) Brachidontes euglyphus (Woods, 1922). UWBM 107551, B8772, L 30.0 mm. Right valve, mould in part. (d) Exterior. (e) Exterior, anterior view. (f–s) Ambigostrea sechura (Olsson, 1944). (f) MUSM INV 235, B8769, L 55.6 mm. Left valve, exterior. (g) UWBM 107558, B8769, L 40.6 mm. Paired valves, posterior view. (h) MUSM INV 235, interior. (i) UWBM 107562, B8770, L 16.5 mm. Left valve, exterior. (j) UWBM 107562, right valve, exterior. (k) UWBM 107563, B8770, L 16.7 mm. Right valve, exterior. (l) UWBM 107568, B8772, L 26.0 mm. Paired valves, right valve resting on left valve. (m) UWBM 107554, B8769, L 76.8 mm. Left valve, interior. (n) MUSM INV 241, B8772, L 15.2 mm. Paired valves, right valve resting on left valve; with attached oyster spat. (o) UWBM 107565, B8772, L 24.6 mm. Left valve, interior. (p) UWBM 107555, B8769, L 52.2 mm. Right valve, exterior. (q) UWBM 107555, interior. (r) MUSM INV 238, B8769, L 29.7 mm. Left valve, interior. (s) MUSM INV 236, B8769, H 29.1 mm. Left valve, exterior, with twig mould in attachment scar. (t, u) undescribed gryphaeid oyster, Los Choros Member of the Paracas Formation, B8768. (t) UWBM 107642, H 49.0 mm. Right valve, interior showing vermiculate dorsoposterior margin. (u) UWBM 107641, H 65.2 mm (left to right in photo). Left valve, anterior view." figureDoi="http://doi.org/10.5281/zenodo.3670233" httpUri="https://zenodo.org/record/3670233/files/figure.png" pageId="13" pageNumber="1545">Figure 2</figureCitation>
(t)) has a circular adductor scar that is not lifted above the floor of the valve and a posterodorsal margin fringed with vermiculate chomata. These features indicate that the Los Choros Member species should be assigned to Gryphaeidae, rather than
<taxonomicName id="4C12E9ADE345E236FC83A05A5981FEFD" authorityName="Rafinesquei" authorityYear="1815" box="[802,909,281,305]" class="Bivalvia" family="Ostreidae" kingdom="Animalia" order="Ostreoida" pageId="13" pageNumber="1545" phylum="Mollusca" rank="family">Ostreidae</taxonomicName>
(Coan and Valentich- Scott 2012;
<bibRefCitation id="EF83EFDFE345E236FE86A07F5BCAFE98" author="Kosenko IN" box="[295,454,316,340]" pageId="13" pageNumber="1545" pagination="759 - 778" refId="ref24467" refString="Kosenko IN. 2017. Late Jurassic-Early Cretaceous oysters from Siberia: a systematic review. Acta Palaeontol Pol. 62: 759 - 778." type="journal article" year="2017">Kosenko 2017</bibRefCitation>
).
</paragraph>
<caption id="DF6DC2A6E343E230FF01A5AE5977F916" ID-DOI="http://doi.org/10.5281/zenodo.3670235" ID-Zenodo-Dep="3670235" httpUri="https://zenodo.org/record/3670235/files/figure.png" pageId="11" pageNumber="1543" startId="11.[160,225,1261,1284]" targetBox="[200,1113,144,1228]" targetPageId="11">
<paragraph id="8BAD922EE343E230FF01A5AE5977F916" blockId="11.[160,1157,1261,1754]" pageId="11" pageNumber="1543">
<emphasis id="B9664E3CE343E230FF01A5AE5B0EFAC8" bold="true" box="[160,258,1261,1284]" pageId="11" pageNumber="1543">Figure 3.</emphasis>
Bivalves from the Caballas Formation. (a, b) Twig moulds on attachment scars of
<taxonomicName id="4C12E9ADE343E230FF01A4495B66FAED" baseAuthorityName="Casadio" baseAuthorityYear="1998" box="[160,362,1290,1313]" class="Bivalvia" family="Ostreidae" genus="Ambigostrea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ostreida" pageId="11" pageNumber="1543" phylum="Mollusca" rank="species" species="sechura">
<emphasis id="B9664E3CE343E230FF01A4495B66FAED" box="[160,362,1290,1313]" italics="true" pageId="11" pageNumber="1543">Ambigostrea sechura</emphasis>
</taxonomicName>
. (a) UWBM 107557, latex cast. (b) UWBM 107564, oyster length 11 mm. (c–e)
<taxonomicName id="4C12E9ADE343E230FF01A46B5844FAF3" authority="Olsson, 1928" authorityName="Olssoni" authorityYear="1928" box="[160,584,1320,1343]" class="Bivalvia" family="Anomiidae" genus="Carolia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ostreida" pageId="11" pageNumber="1543" phylum="Mollusca" rank="species" species="parinensis" subGenus="Parinomya">
<emphasis id="B9664E3CE343E230FF01A46B5AEFFAF3" box="[160,227,1320,1343]" italics="true" pageId="11" pageNumber="1543">Carolia</emphasis>
(
<emphasis id="B9664E3CE343E230FF50A46B5B54FAF3" box="[241,344,1320,1343]" italics="true" pageId="11" pageNumber="1543">Parinomya</emphasis>
)
<emphasis id="B9664E3CE343E230FEC4A46B5BC9FAF3" box="[357,453,1320,1343]" italics="true" pageId="11" pageNumber="1543">parinensis</emphasis>
Olsson, 1928
</taxonomicName>
, on oyster attachment scars. (c) MUSM INV 243, latex cast of external mould on
<taxonomicName id="4C12E9ADE343E230FEDAA4065849FA90" baseAuthorityName="Casadio" baseAuthorityYear="1998" box="[379,581,1349,1372]" class="Bivalvia" family="Ostreidae" genus="Ambigostrea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ostreida" pageId="11" pageNumber="1543" phylum="Mollusca" rank="species" species="sechura">
<emphasis id="B9664E3CE343E230FEDAA4065849FA90" box="[379,581,1349,1372]" italics="true" pageId="11" pageNumber="1543">Ambigostrea sechura</emphasis>
</taxonomicName>
(MUSM INV 237),
<taxonomicName id="4C12E9ADE343E230FD5CA406594CFA90" authorityName="Cantrainei" authorityYear="1838" box="[765,832,1349,1372]" class="Bivalvia" family="Anomiidae" genus="Carolia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ostreida" pageId="11" pageNumber="1543" phylum="Mollusca" rank="genus">
<emphasis id="B9664E3CE343E230FD5CA406594CFA90" box="[765,832,1349,1372]" italics="true" pageId="11" pageNumber="1543">Carolia</emphasis>
</taxonomicName>
H 11.7 mm (top to bottom). (d) UWBM 107569, external mould on
<taxonomicName id="4C12E9ADE343E230FE5EA42158C6FAB5" baseAuthorityName="Casadio" baseAuthorityYear="1998" box="[511,714,1378,1401]" class="Bivalvia" family="Ostreidae" genus="Ambigostrea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ostreida" pageId="11" pageNumber="1543" phylum="Mollusca" rank="species" species="sechura">
<emphasis id="B9664E3CE343E230FE5EA42158C6FAB5" box="[511,714,1378,1401]" italics="true" pageId="11" pageNumber="1543">Ambigostrea sechura</emphasis>
</taxonomicName>
(UWBM 107559),
<taxonomicName id="4C12E9ADE343E230FCDFA42159CDFAB5" authorityName="Cantrainei" authorityYear="1838" box="[894,961,1378,1401]" class="Bivalvia" family="Anomiidae" genus="Carolia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ostreida" pageId="11" pageNumber="1543" phylum="Mollusca" rank="genus">
<emphasis id="B9664E3CE343E230FCDFA42159CDFAB5" box="[894,961,1378,1401]" italics="true" pageId="11" pageNumber="1543">Carolia</emphasis>
</taxonomicName>
H 22.9 mm (left to right). (e) UWBM 107570, external mould on
<taxonomicName id="4C12E9ADE343E230FD2CA4C35951FA5B" baseAuthorityName="Casadio" baseAuthorityYear="1998" box="[653,861,1408,1431]" class="Bivalvia" family="Ostreidae" genus="Ambigostrea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ostreida" pageId="11" pageNumber="1543" phylum="Mollusca" rank="species" species="sechura">
<emphasis id="B9664E3CE343E230FD2CA4C35951FA5B" box="[653,861,1408,1431]" italics="true" pageId="11" pageNumber="1543">Ambigostrea sechura</emphasis>
</taxonomicName>
(UWBM 107560),
<taxonomicName id="4C12E9ADE343E230FB82A4C35E6AFA5B" authorityName="Cantrainei" authorityYear="1838" box="[1059,1126,1408,1431]" class="Bivalvia" family="Anomiidae" genus="Carolia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ostreida" pageId="11" pageNumber="1543" phylum="Mollusca" rank="genus">
<emphasis id="B9664E3CE343E230FB82A4C35E6AFA5B" box="[1059,1126,1408,1431]" italics="true" pageId="11" pageNumber="1543">Carolia</emphasis>
</taxonomicName>
H 15.3 mm (left to right). (f–k)
<taxonomicName id="4C12E9ADE343E230FE7CA4DE5884FA78" authority="DeVries, 2019" authorityName="DeVries" authorityYear="2019" box="[477,648,1437,1460]" class="Bivalvia" family="Crassatellidae" genus="Incatina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Carditoida" pageId="11" pageNumber="1543" phylum="Mollusca" rank="species" species="collinsae" status="sp. nov.">
<emphasis id="B9664E3CE343E230FE7CA4DE5884FA78" box="[477,648,1437,1460]" italics="true" pageId="11" pageNumber="1543">Incatina collinsae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A255F347E343E230FD34A4DE58E6FA78" box="[661,746,1437,1460]" pageId="11" pageNumber="1543" rank="species">sp. nov.</taxonomicNameLabel>
, B8772. (f) UWBM 107637, holotype, L 13.7 mm. Right valve of pair, exterior. (g) UWBM 107637, left valve of pair, exterior. (h) UWBM 107637, paired valves, dorsal view. (i) UWBM 107638, paratype, L 10.0 mm. Right valve of pair. (j) UWBM 107639, paratype, L 16.4 mm. Right valve of pair. (k) UWBM 107638, left valve of pair. (l–q)
<taxonomicName id="4C12E9ADE343E230FF01A7515802F9E5" authority="DeVries, 2019" authorityName="DeVries" authorityYear="2019" box="[160,526,1554,1577]" class="Bivalvia" family="Cardiidae" genus="Pleuriocardia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Venerida" pageId="11" pageNumber="1543" phylum="Mollusca" rank="species" species="schneideri" status="sp. nov." subGenus="Incacardium">
<emphasis id="B9664E3CE343E230FF01A7515B11F9E5" box="[160,285,1554,1577]" italics="true" pageId="11" pageNumber="1543">Pleuriocardia</emphasis>
(
<emphasis id="B9664E3CE343E230FE8AA7515BADF9E5" box="[299,417,1554,1577]" italics="true" pageId="11" pageNumber="1543">Incacardium</emphasis>
)
<emphasis id="B9664E3CE343E230FE0EA7515802F9E5" box="[431,526,1554,1577]" italics="true" pageId="11" pageNumber="1543">schneideri</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A255F347E343E230FDB7A751586FF9E5" box="[534,611,1554,1577]" pageId="11" pageNumber="1543" rank="species">sp. nov.</taxonomicNameLabel>
(l) UWBM 107575, paratype, B8769, L 13.7 mm. Paired valves, dorsal view, anterior to right. (m) UWBM 107574, paratype, B8770, L 25.0 mm. Right valve, exterior. (n) MUSM INV 247, paratype, B8772, L 21.1 mm. Right valve, exterior. (o) UWBM 107576, paratype, B8769, L 25.5 mm. External mould of left valve, oblique view. (p) MUSM INV 246, paratype, B8772, L 20.4 mm. External mould of right valve. (q) 107573, holotype, B8772, L 21.5 mm. Left valve, exterior. (r–t)
<taxonomicName id="4C12E9ADE343E230FE8AA7E6581BF970" authority="DeVries, 2019" authorityName="DeVries" authorityYear="2019" box="[299,535,1701,1724]" class="Bivalvia" family="Mactridae" genus="Mulinoides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Venerida" pageId="11" pageNumber="1543" phylum="Mollusca" rank="species" species="plenicostatus" status="sp. nov.">
<emphasis id="B9664E3CE343E230FE8AA7E6581BF970" box="[299,535,1701,1724]" italics="true" pageId="11" pageNumber="1543">Mulinoides plenicostatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A255F347E343E230FDBFA7E65863F970" box="[542,623,1701,1724]" pageId="11" pageNumber="1543" rank="species">sp. nov.</taxonomicNameLabel>
, UWBM 107571, holotype, B8769, L 28.1 mm. (r) Left valve, exterior. (s) Dorsal view, anterior on right. (t) Right valve, exterior.
</paragraph>
</caption>
<paragraph id="8BAD922EE345E236FF61A01D5BB4FE12" blockId="13.[160,1156,142,478]" pageId="13" pageNumber="1545">
Moulds on the small attachment scar on left valves of Caballas Formation specimens of
<taxonomicName id="4C12E9ADE345E236FF61A0C25BABFE55" baseAuthorityName="Casadio" baseAuthorityYear="1998" box="[192,423,385,409]" class="Bivalvia" family="Ostreidae" genus="Ambigostrea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ostreida" pageId="13" pageNumber="1545" phylum="Mollusca" rank="species" species="sechura">
<emphasis id="B9664E3CE345E236FF61A0C25BABFE55" box="[192,423,385,409]" italics="true" pageId="13" pageNumber="1545">Ambigostrea sechura</emphasis>
</taxonomicName>
include specimens of
<emphasis id="B9664E3CE345E236FD0AA0C2592FFE55" box="[683,803,385,409]" italics="true" pageId="13" pageNumber="1545">
<taxonomicName id="4C12E9ADE345E236FD0AA0C25913FE55" baseAuthorityName="Casadio" baseAuthorityYear="1998" box="[683,799,385,409]" class="Bivalvia" family="Ostreidae" genus="Ambigostrea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ostreida" pageId="13" pageNumber="1545" phylum="Mollusca" rank="species" species="sechura">A. sechura</taxonomicName>
,
</emphasis>
<taxonomicName id="4C12E9ADE345E236FC8DA0C25E72FE55" authorityName="Olssoni" authorityYear="1928" box="[812,1150,385,409]" class="Bivalvia" family="Anomiidae" genus="Carolia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ostreida" pageId="13" pageNumber="1545" phylum="Mollusca" rank="species" species="parinensis" subGenus="Parinomya">
<emphasis id="B9664E3CE345E236FC8DA0C25975FE55" box="[812,889,385,409]" italics="true" pageId="13" pageNumber="1545">Carolia</emphasis>
(
<emphasis id="B9664E3CE345E236FC2BA0C259F3FE55" box="[906,1023,385,409]" italics="true" pageId="13" pageNumber="1545">Parinomya</emphasis>
)
<emphasis id="B9664E3CE345E236FBB0A0C25E72FE55" box="[1041,1150,385,409]" italics="true" pageId="13" pageNumber="1545">parinensis</emphasis>
</taxonomicName>
, venerid bivalves of indeterminate genera,
<taxonomicName id="4C12E9ADE345E236FD2CA0E7596CFE70" authority="DeVries, 2019" authorityName="DeVries" authorityYear="2019" box="[653,864,420,444]" class="Gastropoda" family="Pachychilidae" genus="Nodifaunus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="13" pageNumber="1545" phylum="Mollusca" rank="species" species="gainesi" status="sp. nov.">
<emphasis id="B9664E3CE345E236FD2CA0E7596CFE70" box="[653,864,420,444]" italics="true" pageId="13" pageNumber="1545">Nodifaunus gainesi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A255F347E345E236FCCAA0E759C4FE70" box="[875,968,420,444]" pageId="13" pageNumber="1545" rank="species">sp. nov.</taxonomicNameLabel>
, and plant fragments (
<figureCitation id="13298EABE345E236FF57A0855B58FE12" box="[246,340,454,478]" captionStart="Figure 3" captionStartId="11.[160,225,1261,1284]" captionTargetBox="[200,1113,144,1228]" captionTargetId="figure@11.[199,1117,142,1230]" captionTargetPageId="11" captionText="Figure 3. Bivalves from the Caballas Formation. (a, b) Twig moulds on attachment scars of Ambigostrea sechura. (a) UWBM 107557, latex cast. (b) UWBM 107564, oyster length 11 mm. (c–e) Carolia (Parinomya) parinensis Olsson, 1928, on oyster attachment scars. (c) MUSM INV 243, latex cast of external mould on Ambigostrea sechura (MUSM INV 237), Carolia H 11.7 mm (top to bottom). (d) UWBM 107569, external mould on Ambigostrea sechura (UWBM 107559), Carolia H 22.9 mm (left to right). (e) UWBM 107570, external mould on Ambigostrea sechura (UWBM 107560), Carolia H 15.3 mm (left to right). (f–k) Incatina collinsae sp. nov., B8772. (f) UWBM 107637, holotype, L 13.7 mm. Right valve of pair, exterior. (g) UWBM 107637, left valve of pair, exterior. (h) UWBM 107637, paired valves, dorsal view. (i) UWBM 107638, paratype, L 10.0 mm. Right valve of pair. (j) UWBM 107639, paratype, L 16.4 mm. Right valve of pair. (k) UWBM 107638, left valve of pair. (l–q) Pleuriocardia (Incacardium) schneideri sp. nov. (l) UWBM 107575, paratype, B8769, L 13.7 mm. Paired valves, dorsal view, anterior to right. (m) UWBM 107574, paratype, B8770, L 25.0 mm. Right valve, exterior. (n) MUSM INV 247, paratype, B8772, L 21.1 mm. Right valve, exterior. (o) UWBM 107576, paratype, B8769, L 25.5 mm. External mould of left valve, oblique view. (p) MUSM INV 246, paratype, B8772, L 20.4 mm. External mould of right valve. (q) 107573, holotype, B8772, L 21.5 mm. Left valve, exterior. (r–t) Mulinoides plenicostatus sp. nov., UWBM 107571, holotype, B8769, L 28.1 mm. (r) Left valve, exterior. (s) Dorsal view, anterior on right. (t) Right valve, exterior." figureDoi="http://doi.org/10.5281/zenodo.3670235" httpUri="https://zenodo.org/record/3670235/files/figure.png" pageId="13" pageNumber="1545">Figure 3</figureCitation>
(a), 3(b)).
</paragraph>
</subSubSection>
<subSubSection id="C308C1A5E345E234FF01A35C59B4FE3F" lastPageId="15" lastPageNumber="1547" pageId="13" pageNumber="1545" type="materials_examined">
<paragraph id="8BAD922EE345E236FF01A35C5B0FFDF5" blockId="13.[160,1156,543,846]" box="[160,259,543,569]" pageId="13" pageNumber="1545">
<emphasis id="B9664E3CE345E236FF01A35C5B0FFDF5" bold="true" box="[160,259,543,569]" italics="true" pageId="13" pageNumber="1545">Material</emphasis>
</paragraph>
<paragraph id="8BAD922EE345E234FF01A3005E44FEFD" blockId="13.[160,1156,543,846]" lastBlockId="15.[160,1157,142,305]" lastPageId="15" lastPageNumber="1547" pageId="13" pageNumber="1545">
<collectionCode id="ED030AEBE345E236FF01A3005AE7FD97" box="[160,235,579,603]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34878" lsid="urn:lsid:biocol.org:col:34878" name="University of Washington, Burke Museum" pageId="13" pageNumber="1545" type="Museum">UWBM</collectionCode>
107554, left valve, B8769, L 76.8, H (44), W (22);
<collectionCode id="ED030AEBE345E236FD58A3005948FD97" box="[761,836,579,603]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34878" lsid="urn:lsid:biocol.org:col:34878" name="University of Washington, Burke Museum" pageId="13" pageNumber="1545" type="Museum">UWBM</collectionCode>
107555, right valve, B8769, L (52.2), H (65.4), W (18);
<collectionCode id="ED030AEBE345E236FE05A3255BE3FDB2" box="[420,495,614,638]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34878" lsid="urn:lsid:biocol.org:col:34878" name="University of Washington, Burke Museum" pageId="13" pageNumber="1545" type="Museum">UWBM</collectionCode>
107556, left valve, B8769, L (32.0), H (57.4),
<geoCoordinate id="EE26F4E9E345E236FC7FA3255E21FDB2" box="[990,1069,614,638]" degrees="10.0" direction="west" orientation="longitude" pageId="13" pageNumber="1545" precision="5555" value="-10.0">W 10.0</geoCoordinate>
;
<collectionCode id="ED030AEBE345E236FB99A3255E8FFDB2" box="[1080,1155,614,638]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34878" lsid="urn:lsid:biocol.org:col:34878" name="University of Washington, Burke Museum" pageId="13" pageNumber="1545" type="Museum">UWBM</collectionCode>
107557, left valve, B8769, L 22.2, H 33.3,
<geoCoordinate id="EE26F4E9E345E236FDEFA3CA5881FD6D" box="[590,653,649,673]" degrees="7.7" direction="west" orientation="longitude" pageId="13" pageNumber="1545" precision="5555" value="-7.7">W 7.7</geoCoordinate>
;
<collectionCode id="ED030AEBE345E236FD34A3CA58ECFD6D" box="[661,736,649,673]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34878" lsid="urn:lsid:biocol.org:col:34878" name="University of Washington, Burke Museum" pageId="13" pageNumber="1545" type="Museum">UWBM</collectionCode>
107558, paired valves, B8769, L 40.6, H 44.3,
<geoCoordinate id="EE26F4E9E345E236FF79A3E85B2EFD0F" box="[216,290,683,707]" degrees="21.9" direction="west" orientation="longitude" pageId="13" pageNumber="1545" precision="5555" value="-21.9">W 21.9</geoCoordinate>
(pair);
<collectionCode id="ED030AEBE345E236FECBA3E85BB9FD0F" box="[362,437,683,707]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34878" lsid="urn:lsid:biocol.org:col:34878" name="University of Washington, Burke Museum" pageId="13" pageNumber="1545" type="Museum">UWBM</collectionCode>
107559, left valve, B8769, L (25), H 37.8, W (15);
<collectionCode id="ED030AEBE345E236FC15A3E859F3FD0F" box="[948,1023,683,707]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34878" lsid="urn:lsid:biocol.org:col:34878" name="University of Washington, Burke Museum" pageId="13" pageNumber="1545" type="Museum">UWBM</collectionCode>
107560, left valve, B8769, L (19.5), H (25.1),
<geoCoordinate id="EE26F4E9E345E236FE55A38D5839FD2A" box="[500,565,718,742]" degrees="9.9" direction="west" orientation="longitude" pageId="13" pageNumber="1545" precision="5555" value="-9.9">W 9.9</geoCoordinate>
;
<collectionCode id="ED030AEBE345E236FD9EA38D5886FD2A" box="[575,650,718,742]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34878" lsid="urn:lsid:biocol.org:col:34878" name="University of Washington, Burke Museum" pageId="13" pageNumber="1545" type="Museum">UWBM</collectionCode>
107561, left valve, B8769, L (21.9), H (29.3),
<geoCoordinate id="EE26F4E9E345E236FBCCA38D5ACFFCC5" degrees="9.0" direction="west" orientation="longitude" pageId="13" pageNumber="1545" precision="5555" value="-9.0">W 9.0</geoCoordinate>
;
<collectionCode id="ED030AEBE345E236FF6CA3B25B14FCC5" box="[205,280,753,777]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34878" lsid="urn:lsid:biocol.org:col:34878" name="University of Washington, Burke Museum" pageId="13" pageNumber="1545" type="Museum">UWBM</collectionCode>
107562, paired valves, B8770, L 16.5, H 24.1,
<geoCoordinate id="EE26F4E9E345E236FCA9A3B25958FCC5" box="[776,852,753,777]" degrees="12.7" direction="west" orientation="longitude" pageId="13" pageNumber="1545" precision="5555" value="-12.7">W 12.7</geoCoordinate>
(pair);
<collectionCode id="ED030AEBE345E236FC01A3B259E7FCC5" box="[928,1003,753,777]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34878" lsid="urn:lsid:biocol.org:col:34878" name="University of Washington, Burke Museum" pageId="13" pageNumber="1545" type="Museum">UWBM</collectionCode>
107563, right valve, B8770, L 16.7, H 21.1,
<geoCoordinate id="EE26F4E9E345E236FE74A250581AFCE7" box="[469,534,787,811]" degrees="4.4" direction="west" orientation="longitude" pageId="13" pageNumber="1545" precision="5555" value="-4.4">W 4.4</geoCoordinate>
;
<collectionCode id="ED030AEBE345E236FDBEA2505866FCE7" box="[543,618,787,811]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34878" lsid="urn:lsid:biocol.org:col:34878" name="University of Washington, Burke Museum" pageId="13" pageNumber="1545" type="Museum">UWBM</collectionCode>
107564, left valve, B8770, L (11), H (15.4);
<collectionCode id="ED030AEBE345E236FB99A2505E8FFCE7" box="[1080,1155,787,811]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34878" lsid="urn:lsid:biocol.org:col:34878" name="University of Washington, Burke Museum" pageId="13" pageNumber="1545" type="Museum">UWBM</collectionCode>
107565, left valve, B8772, L 24.6, H 32.4,
<geoCoordinate id="EE26F4E9E345E236FDC5A27558A9FC82" box="[612,677,822,846]" degrees="7.8" direction="west" orientation="longitude" pageId="13" pageNumber="1545" precision="5555" value="-7.8">W 7.8</geoCoordinate>
;
<collectionCode id="ED030AEBE345E236FD0EA27558F6FC82" box="[687,762,822,846]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34878" lsid="urn:lsid:biocol.org:col:34878" name="University of Washington, Burke Museum" pageId="13" pageNumber="1545" type="Museum">UWBM</collectionCode>
107566, left valve, B8772, L 18.8, H (31.7),
<geoCoordinate id="EE26F4E9E346E235FF49A7525B24F9E5" box="[232,296,1553,1577]" degrees="7.0" direction="west" orientation="longitude" pageId="14" pageNumber="1546" precision="5555" value="-7.0">W 7.0</geoCoordinate>
;
<collectionCode id="ED030AEBE346E235FE90A7525B70F9E5" box="[305,380,1553,1577]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34878" lsid="urn:lsid:biocol.org:col:34878" name="University of Washington, Burke Museum" pageId="14" pageNumber="1546" type="Museum">UWBM</collectionCode>
107567, left valve, B8772, L 23.3, H 27.2,
<geoCoordinate id="EE26F4E9E346E235FC98A752598AF9E5" box="[825,902,1553,1577]" degrees="10.2" direction="west" orientation="longitude" pageId="14" pageNumber="1546" precision="5555" value="-10.2">W 10.2</geoCoordinate>
;
<collectionCode id="ED030AEBE346E235FC31A75259D7F9E5" box="[912,987,1553,1577]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34878" lsid="urn:lsid:biocol.org:col:34878" name="University of Washington, Burke Museum" pageId="14" pageNumber="1546" type="Museum">UWBM</collectionCode>
107568, paired valves, B8772, L 26.0, H 13.9,
<geoCoordinate id="EE26F4E9E346E235FE76A770581FF987" box="[471,531,1587,1611]" degrees="6.5" direction="west" orientation="longitude" pageId="14" pageNumber="1546" precision="5555" value="-6.5">W 6.5</geoCoordinate>
(pair);
<collectionCode id="ED030AEBE346E235FDFDA77058A9F987" box="[604,677,1587,1611]" pageId="14" pageNumber="1546">MUSM</collectionCode>
INV 235, left valve, B8769, L 55.6, H (59.6),
<geoCoordinate id="EE26F4E9E346E235FBCCA7705ADCF9A2" degrees="18.1" direction="west" orientation="longitude" pageId="14" pageNumber="1546" precision="5555" value="-18.1">W 18.1</geoCoordinate>
;
<collectionCode id="ED030AEBE346E235FF79A7155B2DF9A2" box="[216,289,1622,1646]" pageId="14" pageNumber="1546">MUSM</collectionCode>
INV 236, left valve, B8769, L 21.6, H 29.1,
<geoCoordinate id="EE26F4E9E346E235FD7BA7155915F9A2" box="[730,793,1622,1646]" degrees="7.3" direction="west" orientation="longitude" pageId="14" pageNumber="1546" precision="5555" value="-7.3">W 7.3</geoCoordinate>
;
<collectionCode id="ED030AEBE346E235FC80A7155966F9A2" box="[801,874,1622,1646]" pageId="14" pageNumber="1546">MUSM</collectionCode>
INV 237, left valve, B8769, L (22.4), H (20.6),
<geoCoordinate id="EE26F4E9E346E235FEFCA73A5B91F95D" box="[349,413,1657,1681]" degrees="8.0" direction="west" orientation="longitude" pageId="14" pageNumber="1546" precision="5555" value="-8.0">W 8.0</geoCoordinate>
;
<collectionCode id="ED030AEBE346E235FE07A73A5BE3F95D" box="[422,495,1657,1681]" pageId="14" pageNumber="1546">MUSM</collectionCode>
INV 238, left valve, B8769, L 29.7, H 32.4,
<geoCoordinate id="EE26F4E9E346E235FC17A73A5E08F95D" box="[950,1028,1657,1681]" degrees="12.7" direction="west" orientation="longitude" pageId="14" pageNumber="1546" precision="5555" value="-12.7">W 12.7</geoCoordinate>
;
<collectionCode id="ED030AEBE346E235FBAFA73A5E5BF95D" box="[1038,1111,1657,1681]" pageId="14" pageNumber="1546">MUSM</collectionCode>
INV 239, paired valves, B8770, L 25.0, H 30.8,
<geoCoordinate id="EE26F4E9E346E235FDFEA7D858A7F97F" box="[607,683,1691,1715]" degrees="15.9" direction="west" orientation="longitude" pageId="14" pageNumber="1546" precision="5555" value="-15.9">W 15.9</geoCoordinate>
(pair);
<collectionCode id="ED030AEBE346E235FD57A7D85933F97F" box="[758,831,1691,1715]" pageId="14" pageNumber="1546">MUSM</collectionCode>
INV 240, right valve, B8770, L 15.9, H 17.8,
<geoCoordinate id="EE26F4E9E347E234FE8DA1CD5B60FF6A" box="[300,364,142,166]" degrees="3.9" direction="west" orientation="longitude" pageId="15" pageNumber="1547" precision="5555" value="-3.9">W 3.9</geoCoordinate>
;
<collectionCode id="ED030AEBE347E234FED4A1CD5BB2FF6A" box="[373,446,142,166]" pageId="15" pageNumber="1547">MUSM</collectionCode>
INV 241, paired valves, B8772, L 15.2, H 24.9,
<geoCoordinate id="EE26F4E9E347E234FC10A1CD59FCFF6A" box="[945,1008,142,166]" degrees="8.5" direction="west" orientation="longitude" pageId="15" pageNumber="1547" precision="5555" value="-8.5">W 8.5</geoCoordinate>
(pair);
<collectionCode id="ED030AEBE347E234FB9AA1CD5E88FF6A" box="[1083,1156,142,166]" pageId="15" pageNumber="1547">MUSM</collectionCode>
INV 242, left valve, B8772, L 22.3, H 28.2,
<geoCoordinate id="EE26F4E9E347E234FDF7A1F25899FF05" box="[598,661,177,201]" degrees="5.8" direction="west" orientation="longitude" pageId="15" pageNumber="1547" precision="5555" value="-5.8">W 5.8</geoCoordinate>
.
<collectorName id="26E7F7F8E347E234FD3FA1F2591CFF05" box="[670,784,177,201]" pageId="15" pageNumber="1547">Gryphaeid</collectorName>
oysters from
<collectorName id="26E7F7F8E347E234FC02A1F25B89FF20" pageId="15" pageNumber="1547">Los Choros Member, Paracas Formation, B</collectorName>
8768:
<specimenCode id="DBB43A55E347E234FE66A1975861FF20" box="[455,621,212,236]" collectionCode="UWBM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34878" lsid="urn:lsid:biocol.org:col:34878" name="University of Washington, Burke Museum" pageId="15" pageNumber="1547" type="Museum">UWBM 107641</specimenCode>
, left valve, L 46.6, H 65.2,
<geoCoordinate id="EE26F4E9E347E234FC25A19759DEFF20" box="[900,978,212,236]" degrees="33.6" direction="west" orientation="longitude" pageId="15" pageNumber="1547" precision="5555" value="-33.6">W 33.6</geoCoordinate>
;
<specimenCode id="DBB43A55E347E234FC7AA1975E8DFF20" box="[987,1153,212,236]" collectionCode="UWBM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34878" lsid="urn:lsid:biocol.org:col:34878" name="University of Washington, Burke Museum" pageId="15" pageNumber="1547" type="Museum">UWBM 107642</specimenCode>
, right valve, L 49.0, H 53.6,
<geoCoordinate id="EE26F4E9E347E234FE60A1B55802FEC2" box="[449,526,246,270]" degrees="10.8" direction="west" orientation="longitude" pageId="15" pageNumber="1547" precision="5555" value="-10.8">W 10.8</geoCoordinate>
;
<specimenCode id="DBB43A55E347E234FDB9A1B558B2FEC2" box="[536,702,246,270]" collectionCode="UWBM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34878" lsid="urn:lsid:biocol.org:col:34878" name="University of Washington, Burke Museum" pageId="15" pageNumber="1547" type="Museum">UWBM 107643</specimenCode>
, valve uncertain, L (47.9), H (30.4);
<specimenCode id="DBB43A55E347E234FB99A1B55AF8FEFD" collectionCode="UWBM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34878" lsid="urn:lsid:biocol.org:col:34878" name="University of Washington, Burke Museum" pageId="15" pageNumber="1547" type="Museum">UWBM 107644</specimenCode>
, right valve, L (29.0), H 42.6, W (5);
<specimenCode id="DBB43A55E347E234FDD3A05A5914FEFD" box="[626,792,281,305]" collectionCode="UWBM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34878" lsid="urn:lsid:biocol.org:col:34878" name="University of Washington, Burke Museum" pageId="15" pageNumber="1547" type="Museum">UWBM 107645</specimenCode>
, left valve, L (42.1), H (39.5).
</paragraph>
<caption id="DF6DC2A6E345E236FF01A2EB589EF976" ID-DOI="http://doi.org/10.5281/zenodo.3686966" ID-Zenodo-Dep="3686966" httpUri="https://zenodo.org/record/3686966/files/figure.png" pageId="13" pageNumber="1545" targetBox="[164,1150,144,1035]" targetPageId="14">
<paragraph id="8BAD922EE345E236FF01A2EB589EF976" blockId="13.[160,1157,936,1722]" pageId="13" pageNumber="1545">
<emphasis id="B9664E3CE345E236FF01A2EB5AF2FC73" bold="true" box="[160,254,936,959]" pageId="13" pageNumber="1545">Figure 4.</emphasis>
Gastropods from the Caballas Formation. (a–e)
<taxonomicName id="4C12E9ADE345E236FD7EA2EB597EFC73" authority="DeVries, 2019" authorityName="DeVries" authorityYear="2019" box="[735,882,936,959]" class="Gastropoda" family="Neritidae" genus="Nerita" kingdom="Animalia" order="Cycloneritimorpha" pageId="13" pageNumber="1545" phylum="Mollusca" rank="species" species="squatina" status="sp. nov.">
<emphasis id="B9664E3CE345E236FD7EA2EB597EFC73" box="[735,882,936,959]" italics="true" pageId="13" pageNumber="1545">Nerita squatina</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A255F347E345E236FCDAA2EB59C7FC73" box="[891,971,936,959]" pageId="13" pageNumber="1545" rank="species">sp. nov.</taxonomicNameLabel>
, B8771. (a) UWBM 107577, holotype, W 5.7 mm, spire. (b) UWBM 107648, paratype, W 8.5 mm, shoulder of last whorl. (c) UWBM 107649, paratype, W 10.1 mm, anterior of outer lip. (d) UWBM 107647, paratype, W 7.8 mm, inner surface of outer lip. (e) UWBM 107646, paratype, W 6.2 mm, columnar teeth. (f)
<taxonomicName id="4C12E9ADE345E236FC06A5435E07FBDB" authorityName="Lamarcki" authorityYear="1799" box="[935,1035,1024,1047]" class="Gastropoda" family="Calyptraeidae" genus="Calyptraea" kingdom="Animalia" order="Littorinimorpha" pageId="13" pageNumber="1545" phylum="Mollusca" rank="genus">
<emphasis id="B9664E3CE345E236FC06A5435E07FBDB" box="[935,1035,1024,1047]" italics="true" pageId="13" pageNumber="1545">Calyptraea</emphasis>
</taxonomicName>
cf.
<taxonomicName id="4C12E9ADE345E236FB8FA5435B22FBF9" authority="Solander, 1766" authorityName="Solanderi" authorityYear="1766" class="Gastropoda" family="Calyptraeidae" genus="Calyptraea" kingdom="Animalia" order="Littorinimorpha" pageId="13" pageNumber="1545" phylum="Mollusca" rank="species" species="aperta">
<emphasis id="B9664E3CE345E236FB8FA5435E8EFBDB" box="[1070,1154,1024,1047]" italics="true" pageId="13" pageNumber="1545">C. aperta</emphasis>
<bibRefCitation id="EF83EFDFE345E236FF01A55D5B22FBF9" author="Solander DC" box="[160,302,1054,1077]" pageId="13" pageNumber="1545" refId="ref26472" refString="Solander DC. 1766. In: Brander Gi editor. Fossilia Hantoniensia collectai et in Mus. London: Brittanico deposita." type="book" year="1766">Solander, 1766</bibRefCitation>
</taxonomicName>
, UWBM 107578, B8772, W (diameter) 9.9 mm, external mould. (g)
<taxonomicName id="4C12E9ADE345E236FC08A55D5AC3FB9E" authority="Gabb, 1870" authorityName="Gabbi" authorityYear="1870" class="Dothideomycetes" family="Didymellaceae" genus="Ampullina" higherTaxonomySource="CoL" kingdom="Fungi" order="Pleosporales" pageId="13" pageNumber="1545" phylum="Ascomycota" rank="species" species="ortoni">
<emphasis id="B9664E3CE345E236FC08A55D5E4AFBF9" box="[937,1094,1054,1077]" italics="true" pageId="13" pageNumber="1545">Ampullina ortoni</emphasis>
<bibRefCitation id="EF83EFDFE345E236FBEAA55D5AC3FB9E" author="Gabb WM" pageId="13" pageNumber="1545" pagination="25 - 32" refId="ref23664" refString="Gabb WM. 1870. Description of new species of South American fossils: no. 1 i Tertiary. Am J Conchol. 5 (1): 25 - 32." type="journal article" year="1870">Gabb, 1870</bibRefCitation>
</taxonomicName>
, UWBM 107,627, B8772, L 36.8 mm, abapertural view, last whorl mostly missing; left margin annotated. (h–l)
<taxonomicName id="4C12E9ADE345E236FEE1A51B581EFBA3" authority="DeVries, 2019" authorityName="DeVries" authorityYear="2019" box="[320,530,1112,1135]" class="Gastropoda" family="Eulimidae" genus="Melanella" higherTaxonomySource="CoL" kingdom="Animalia" order="Littorinimorpha" pageId="13" pageNumber="1545" phylum="Mollusca" rank="species" species="indicaformis" status="sp. nov.">
<emphasis id="B9664E3CE345E236FEE1A51B581EFBA3" box="[320,530,1112,1135]" italics="true" pageId="13" pageNumber="1545">Melanella indicaformis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A255F347E345E236FDB9A51B586FFBA3" box="[536,611,1112,1135]" pageId="13" pageNumber="1545" rank="species">sp. nov.</taxonomicNameLabel>
(h) UWBM 107620, paratype, B8772, L 8.2 mm, apertural view. (i) UWBM 107619, paratype, B8769, L 9.4 mm, apertural/lateral view. (j) UWBM 107623, paratype, B8769, L 8.2 mm, apertural view. (k) UWBM 107618, paratype, B8772, L 10.6, abapertural view. (l) UWBM 107617, holotype, B8769, L 11.1 mm, abapertural view. (m–q)
<taxonomicName id="4C12E9ADE345E236FCB5A5F359B4FB0B" authority="DeVries, 2019" authorityName="DeVries" authorityYear="2019" box="[788,952,1200,1223]" class="Gastropoda" family="Potamididae" genus="Potamides" kingdom="Animalia" pageId="13" pageNumber="1545" phylum="Mollusca" rank="species" species="henryi" status="sp. nov.">
<emphasis id="B9664E3CE345E236FCB5A5F359B4FB0B" box="[788,952,1200,1223]" italics="true" pageId="13" pageNumber="1545">Potamides henryi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A255F347E345E236FC60A5F35E1CFB0B" box="[961,1040,1200,1223]" pageId="13" pageNumber="1545" rank="species">sp. nov.</taxonomicNameLabel>
(m) UWBM 107582, holotype, B8772, L 17.8 mm, lateral view. (n) UWBM 107582, W 6.9 mm, basal view. (o) UWBM 107586, paratype, B8771, L 11.2 mm, lateral view. (p) UWBM 107587, paratype, B8769, L 9.5 mm, abapertural view. (q) UWBM 107650, paratype, B8770, L 15.4 mm, lateral view, shell compressed. (r–t)
<taxonomicName id="4C12E9ADE345E236FF01A4655B49FAF1" authority="DeVries, 2019" authorityName="DeVries" authorityYear="2019" box="[160,325,1318,1341]" class="Gastropoda" family="Potamididae" genus="Potamides" kingdom="Animalia" pageId="13" pageNumber="1545" phylum="Mollusca" rank="species" species="janeae" status="sp. nov.">
<emphasis id="B9664E3CE345E236FF01A4655B49FAF1" box="[160,325,1318,1341]" italics="true" pageId="13" pageNumber="1545">Potamides janeae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A255F347E345E236FEEBA4655B9BFAF1" box="[330,407,1318,1341]" pageId="13" pageNumber="1545" rank="species">sp. nov.</taxonomicNameLabel>
, B8771. (r) UWBM 107579, holotype, L 9.0 mm, lateral view. (s) MUSM INV 249, L 8.8 mm, lateral and columellar view. (t) UWBM 107580, L 9.5 mm, layeral and columellar view. (u, v, x)
<taxonomicName id="4C12E9ADE345E236FF01A4235B62FABB" authority="DeVries, 2019" authorityName="DeVries" authorityYear="2019" box="[160,366,1376,1399]" class="Gastropoda" family="Potamididae" genus="Papposilenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neotaenioglossa" pageId="13" pageNumber="1545" phylum="Mollusca" rank="species" species="utriculus" status="gen. nov., sp. nov.">
<emphasis id="B9664E3CE345E236FF01A4235B62FABB" box="[160,366,1376,1399]" italics="true" pageId="13" pageNumber="1545">Papposilenus utriculus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A255F347E345E236FED2A423582CFABB" box="[371,544,1376,1399]" pageId="13" pageNumber="1545" rank="species">gen. nov., sp. nov.</taxonomicNameLabel>
(u) UWBM 107592, paratype, B8772, L 18.7 mm, oblique view of base. (v) UWBM 107589, holotype, B8772, L 25.5 mm, lateral view. (x) UWBM 107590, B8770, L 17.6 mm, lateral view. (w, y, z, jj)
<taxonomicName id="4C12E9ADE345E236FED9A4D85865FA7E" authority="DeVries, 2019" authorityName="DeVries" authorityYear="2019" box="[376,617,1435,1458]" class="Gastropoda" family="Potamididae" genus="Rhinotamides" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="13" pageNumber="1545" phylum="Mollusca" rank="species" species="everriculum" status="sp. nov.">
<emphasis id="B9664E3CE345E236FED9A4D85865FA7E" box="[376,617,1435,1458]" italics="true" pageId="13" pageNumber="1545">Rhinotamides everriculum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A255F347E345E236FDCCA4D858BBFA7E" box="[621,695,1435,1458]" pageId="13" pageNumber="1545" rank="species">sp. nov.</taxonomicNameLabel>
(w) UWBM 107598, paratype, B8772, L 23.5 mm, abapertural view, juvenile. (y) UWBM 107593, holotype, B8769, L 21.1 mm, abapertural view. (z) UWBM 107594, paratype, B8772, L 19.4 mm, apertural view, posterior and anterior missing. (jj) UWBM 107593, apertural view. (aa, bb)
<taxonomicName id="4C12E9ADE345E236FEDFA4B05803F9C6" authority="DeVries, 2019" authorityName="DeVries" authorityYear="2019" box="[382,527,1523,1546]" class="Gastropoda" family="Potamididae" genus="Terebralia" kingdom="Animalia" pageId="13" pageNumber="1545" phylum="Mollusca" rank="species" species="pauli" status="gen. nov.">
<emphasis id="B9664E3CE345E236FEDFA4B05803F9C6" box="[382,527,1523,1546]" italics="true" pageId="13" pageNumber="1545">Terebralia pauli</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A255F347E345E236FDB2A4B05860F9C6" box="[531,620,1523,1546]" pageId="13" pageNumber="1545" rank="genus">gen. nov.</taxonomicNameLabel>
sp. nov., UWBM 107599, holotype, B8771, L 9.2 mm. (aa) Abapertural view. (bb) Apertural view, outer lip missing. (cc)
<taxonomicName id="4C12E9ADE345E236FD4DA7535989F9EB" authority="DeVries, 2019" authorityName="DeVries" authorityYear="2019" box="[748,901,1552,1575]" class="Gastropoda" family="Potamididae" genus="Terebralia" kingdom="Animalia" pageId="13" pageNumber="1545" phylum="Mollusca" rank="species" species="marki" status="gen. nov.">
<emphasis id="B9664E3CE345E236FD4DA7535989F9EB" box="[748,901,1552,1575]" italics="true" pageId="13" pageNumber="1545">Terebralia marki</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A255F347E345E236FC2AA75359EAF9EB" box="[907,998,1552,1575]" pageId="13" pageNumber="1545" rank="genus">gen. nov.</taxonomicNameLabel>
sp. nov., UWBM 107600, holotype, B8771, L 9.8, lateral view. (dd–ii).
<taxonomicName id="4C12E9ADE345E236FD0BA76D5952F989" authority="DeVries, 2019" authorityName="DeVries" authorityYear="2019" box="[682,862,1582,1605]" class="Gastropoda" family="Pachychilidae" genus="Nodifaunus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="13" pageNumber="1545" phylum="Mollusca" rank="species" species="gainesi" status="sp. nov.">
<emphasis id="B9664E3CE345E236FD0BA76D5952F989" box="[682,862,1582,1605]" italics="true" pageId="13" pageNumber="1545">Nodifaunus gainesi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A255F347E345E236FCC6A76D59BAF989" box="[871,950,1582,1605]" pageId="13" pageNumber="1545" rank="species">sp. nov.</taxonomicNameLabel>
(dd) UWBM 107609, holotype, B8769, L 42.4 mm, apertural view. (ee) MUSM INV 264, paratype, B8772, L 31.0 mm, apertural view, anterior missing. (ff) UWBM 107609, abapertural view. (gg) UWBM 107612, paratype, B8770, L 14.2 mm, abapertural view, juvenile. (hh) UWBM 107612, apertural view. (ii) UWBM 107610, paratype, B8769, L 25.9 mm, apertural view, anterior missing.
</paragraph>
</caption>
<caption id="DF6DC2A6E346E235FF01A56F5BDDFA0C" ID-DOI="http://doi.org/10.5281/zenodo.3670237" ID-Zenodo-Dep="3670237" httpUri="https://zenodo.org/record/3670237/files/figure.png" pageId="14" pageNumber="1546" startId="14.[160,225,1068,1091]" targetBox="[164,1150,144,1035]" targetPageId="14">
<paragraph id="8BAD922EE346E235FF01A56F5BDDFA0C" blockId="14.[160,1157,1068,1472]" pageId="14" pageNumber="1546">
<emphasis id="B9664E3CE346E235FF01A56F5AF0FB8F" bold="true" box="[160,252,1068,1091]" pageId="14" pageNumber="1546">Figure 5.</emphasis>
Gastropods from the Caballas Formation. (a–e)
<taxonomicName id="4C12E9ADE346E235FD43A56F59CEFB8F" authority="DeVries, 2019" authorityName="DeVries" authorityYear="2019" box="[738,962,1068,1091]" class="Gastropoda" family="Potamididae" genus="Caballasea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neotaenioglossa" pageId="14" pageNumber="1546" phylum="Mollusca" rank="species" species="segmentata" status="gen. nov., sp. nov.">
<emphasis id="B9664E3CE346E235FD43A56F59CEFB8F" box="[738,962,1068,1091]" italics="true" pageId="14" pageNumber="1546">Caballasea segmentata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A255F347E346E235FC6AA56F5E88FB8F" box="[971,1156,1068,1091]" pageId="14" pageNumber="1546" rank="species">gen. nov., sp. nov.</taxonomicNameLabel>
B8769. (a) UWBM 107607, paratype, L 12.4 mm, apertural view. (b) UWBM 107607, abapertural view. (c) UWBM 107606, holotype, L 12.6 mm, abapertural view. (d) MUSM INV 262, paratype, L 8.3 mm, oblique view of parietal callus and columella. (e) UWBM 107608, paratype, L 15.3 mm, abapertural view. (f, g)
<taxonomicName id="4C12E9ADE346E235FEB0A5E25BC0FB74" authority="DeVries, 2019" authorityName="DeVries" authorityYear="2019" box="[273,460,1185,1208]" class="Gastropoda" family="Turritellidae" genus="Turritella" higherTaxonomySource="CoL" kingdom="Animalia" order="Littorinimorpha" pageId="14" pageNumber="1546" phylum="Mollusca" rank="species" species="capistrata" status="sp. nov.">
<emphasis id="B9664E3CE346E235FEB0A5E25BC0FB74" box="[273,460,1185,1208]" italics="true" pageId="14" pageNumber="1546">Turritella capistrata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A255F347E346E235FE75A5E2582EFB74" box="[468,546,1185,1208]" pageId="14" pageNumber="1546" rank="species">sp. nov.</taxonomicNameLabel>
(f) MUSM INV 259, paratype, B8772, L 16.3 mm, lateral view. (g) UWBM 107601, holotype, B8770, L 14.4 mm, apertural view. (h)
<taxonomicName id="4C12E9ADE346E235FCF4A5FC5E72FB1A" authority="Olsson, 1944" authorityName="Olssoni" authorityYear="1944" box="[853,1150,1215,1238]" class="Gastropoda" family="Turritellidae" genus="Turritella" higherTaxonomySource="CoL" kingdom="Animalia" order="Littorinimorpha" pageId="14" pageNumber="1546" phylum="Mollusca" rank="species" species="saposa">
<emphasis id="B9664E3CE346E235FCF4A5FC59F9FB1A" box="[853,1013,1215,1238]" italics="true" pageId="14" pageNumber="1546">Turritella saposa</emphasis>
<bibRefCitation id="EF83EFDFE346E235FC5FA5FC5E72FB1A" author="Olsson AA" box="[1022,1150,1215,1238]" pageId="14" pageNumber="1546" pagination="1 - 147" refId="ref25769" refString="Olsson AA. 1944. Contributions to the Tertiary paleontology of northern Peru. Part VII. The Cretaceous of the Paita region. Bulletins Am Paleontol. 19 (68): 1 - 147." type="journal article" year="1944">Olsson, 1944</bibRefCitation>
</taxonomicName>
, UWBM 107602, B8770, L 15.5 mm. (i)
<taxonomicName id="4C12E9ADE346E235FD93A59F591AFB3F" authority="DeVries, 2019" authorityName="DeVries" authorityYear="2019" box="[562,790,1244,1267]" class="Spirochaetes" family="Spirochaetaceae" genus="Cristispira" higherTaxonomySource="CoL" kingdom="Bacteria" order="Spirochaetales" pageId="14" pageNumber="1546" phylum="Spirochaetes" rank="species" species="caballasensis" status="sp. nov.">
<emphasis id="B9664E3CE346E235FD93A59F591AFB3F" box="[562,790,1244,1267]" italics="true" pageId="14" pageNumber="1546">Cristispira caballasensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A255F347E346E235FC80A59F5978FB3F" box="[801,884,1244,1267]" pageId="14" pageNumber="1546" rank="species">sp. nov.</taxonomicNameLabel>
, UWBM 107605, holotype, B8769, L 5.1 mm, fragment. (j, k, m, n)
<taxonomicName id="4C12E9ADE346E235FD8AA5BA591CFADC" authority="DeVries, 2019" authorityName="DeVries" authorityYear="2019" box="[555,784,1273,1296]" class="Gastropoda" family="Melongenidae" genus="Rhombopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neogastropoda" pageId="14" pageNumber="1546" phylum="Mollusca" rank="species" species="longinquus" status="sp. nov.">
<emphasis id="B9664E3CE346E235FD8AA5BA591CFADC" box="[555,784,1273,1296]" italics="true" pageId="14" pageNumber="1546">Rhombopsis longinquus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A255F347E346E235FCB6A5BA5969FADC" box="[791,869,1273,1296]" pageId="14" pageNumber="1546" rank="species">sp. nov.</taxonomicNameLabel>
(j) UWBM 107630, holotype, B8772, L 18.3 mm, apertural view, outer lip missing. (k) MUSM INV 269, paratype, B8769, L 19.9 mm, abapertural view. (m) UWBM 107631, paratype, B8772, L 22.5 mm, abapertural view. (n) UWBM 107632, paratype, B8769, L 16.4 mm, lateral view of spire and columella. (l, o–q)
<taxonomicName id="4C12E9ADE346E235FC5DA4125B2FFA4A" authority="DeVries, 2019" authorityName="DeVries" authorityYear="2019" class="Gastropoda" family="Fasciolariidae" genus="Tristeirostoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neogastropoda" pageId="14" pageNumber="1546" phylum="Mollusca" rank="species" species="saxaquadrata" status="sp. nov.">
<emphasis id="B9664E3CE346E235FC5DA4125B2FFA4A" italics="true" pageId="14" pageNumber="1546">Tristeirostoma saxaquadrata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A255F347E346E235FE8FA42C5B73FA4A" box="[302,383,1391,1414]" pageId="14" pageNumber="1546" rank="species">sp. nov.</taxonomicNameLabel>
B8771. (l) UWBM 107636, paratype, L 10.0 mm, spire. (o) UWBM 107,635, holotype, L 7.6 mm, abapertural view. (p) UWBM 107635, lateral view of columella. (q) UWBM 107635, apertural/lateral view.
</paragraph>
</caption>
<paragraph id="8BAD922EE347E234FF01A0315B28FE40" blockId="15.[160,1155,370,499]" box="[160,292,370,396]" pageId="15" pageNumber="1547">
<emphasis id="B9664E3CE347E234FF01A0315B28FE40" bold="true" box="[160,292,370,396]" italics="true" pageId="15" pageNumber="1547">Occurrence</emphasis>
</paragraph>
<paragraph id="8BAD922EE347E234FF01A0D559B4FE3F" blockId="15.[160,1155,370,499]" pageId="15" pageNumber="1547">
Upper Campanian to Maastrichtian,
<collectingCountry id="F305D2BEE347E234FDEBA0D558AEFE62" box="[586,674,406,430]" name="Haiti" pageId="15" pageNumber="1547">Tortuga</collectingCountry>
and La Mesa formations, Sechura Basin, northern
<collectingCountry id="F305D2BEE347E234FEACA0FA5B4EFE1D" box="[269,322,441,465]" name="Peru" pageId="15" pageNumber="1547">Peru</collectingCountry>
; Upper Cretaceous, eastern
<collectingCountry id="F305D2BEE347E234FD2BA0FA58B1FE1D" box="[650,701,441,465]" name="Peru" pageId="15" pageNumber="1547">Peru</collectingCountry>
(
<bibRefCitation id="EF83EFDFE347E234FD6FA0FA59ADFE1D" author="Cerron Fi Galloso Ai Chumpitaz M" box="[718,929,441,465]" pageId="15" pageNumber="1547" pagination="1 - 212" refId="ref22273" refString="Cerron Fi Galloso Ai Chumpitaz M. 1998. Geologia de los cuadrangulos Uracusa y Cachiyacu. Inst Geologico Minero y Metalurgico Serie A Boletin. 117: 1 - 212." type="journal article" year="1998">Cerron et al. 1998</bibRefCitation>
); Lower Paleogene, Cuenca Member, Caballas Formation, East Pisco Basin, southern
<collectingCountry id="F305D2BEE347E234FCDEA09859B9FE3F" box="[895,949,475,499]" name="Peru" pageId="15" pageNumber="1547">Peru</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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