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<mods:title id="E1FC377C5162F228C1AF7E69BF5ADE74">Gilpinia hakonensis and similar species in Japan and ovipositors of five European Gilpinia species (Hymenoptera, Diprionidae)</mods:title>
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<taxonomicName id="4C589DFAFFCA694FFBB5FC26FDACF9FF" authority="Okutani, 1965" authorityName="Okutani" authorityYear="1965" box="[151,572,906,933]" class="Insecta" family="Diprionidae" genus="Gilpinia" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="478" phylum="Arthropoda" rank="species" species="amamiana">
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<emphasis id="B92C3A6BFFCA694FFBB5FC26FE13F9FE" bold="true" box="[151,387,906,933]" italics="true" pageId="7" pageNumber="478">Gilpinia amamiana</emphasis>
<bibRefCitation id="EFC99B88FFCA694FFAA8FC26FDACF9FF" author="Okutani, T." box="[394,572,906,932]" pageId="7" pageNumber="478" pagination="73 - 84" refId="ref14145" refString="Okutani, T. (1965) Sawflies and horntails from the Ryukyus. Kontyu, 33, 73 - 84." type="journal article" year="1965">Okutani, 1965</bibRefCitation>
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(
<figureCitation id="1363FAFCFFCA694FFBBDFC03FEC1F992" box="[159,337,943,969]" captionStart="FIGURES 2" captionStartId="4.[151,264,1746,1770]" captionTargetBox="[166,1435,215,1720]" captionTargetId="figure-19@4.[151,1435,181,1723]" captionTargetPageId="4" captionText="FIGURES 2AH. AF, IK, Gilpinia amamiana: A, B, holotype, female; C, D, female, Tokunoshima Isl.; E, F, male, Amami Oshima Isl. G, H, L, Gilpinia okinawa, holotype, female. A, C, E, G, Dorsal view; B, D, F, H, ventrolateral view; IL, head, frontal view." figureDoi="http://doi.org/10.5281/zenodo.5057391" httpUri="https://zenodo.org/record/5057391/files/figure.png" pageId="7" pageNumber="478">Figs 2AF, IK</figureCitation>
,
<figureCitation id="1363FAFCFFCA694FFA7FFC03FDBEF992" box="[349,558,943,969]" captionStart="FIGURES 3" captionStartId="9.[151,264,1786,1810]" captionTargetBox="[151,1435,181,1762]" captionTargetId="figure-19@9.[151,1435,181,1762]" captionTargetPageId="9" captionText="FIGURES 3AX. AH, Head, dorsal view; IP, dorsal part of head, anterolateral view; Q, head, lateral view; RX, antenna, inner view (RT, V, W) or outer view (U, X). AD, IL, QU, Gilpinia hakonensis: A, I, Q, R, lectotype of Lophyrus hakonensis, female; B, J, S, holotype of G. hakonensis var. laticincta, female; C, K, T, paratype of Diprion fukudai, female; D, L, U, male, Honshu. EG, MO, V, W, Gilpinia amamiana: E, M, V, holotype, female; F, N, W, female, Tokunoshima Isl.; G, O, male, Amami Oshima Isl. H, P, X, Gilpinia okinawa, holotype, female. T, Reversed." figureDoi="http://doi.org/10.5281/zenodo.5057395" httpUri="https://zenodo.org/record/5057395/files/figure.png" pageId="7" pageNumber="478">3EG, MO, V, W</figureCitation>
, 4IL, Q, 5GJ, S, 6FH, 7DF)
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<taxonomicName id="4C589DFAFFCA694FFBB5FC5FFD8DFE50" authority="Okutani, 1965: 76" authorityName="Okutani" authorityPageNumber="76" authorityYear="1965" box="[151,541,1011,1035]" class="Insecta" family="Diprionidae" genus="Gilpinia" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="478" phylum="Arthropoda" rank="species" species="amamiana">
<emphasis id="B92C3A6BFFCA694FFBB5FC5FFECDFE50" box="[151,349,1011,1035]" italics="true" pageId="7" pageNumber="478">Gilpinia amamiana</emphasis>
<bibRefCitation id="EFC99B88FFCA694FFA41FC5FFD8DFE50" author="Okutani, T." box="[355,541,1011,1035]" pageId="7" pageNumber="478" pagination="73 - 84" refId="ref14145" refString="Okutani, T. (1965) Sawflies and horntails from the Ryukyus. Kontyu, 33, 73 - 84." type="journal article" year="1965">Okutani, 1965: 76</bibRefCitation>
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<bibRefCitation id="EFC99B88FFCA694FF90AFC5FFD72FE50" author="Okutani, T." box="[552,738,1011,1035]" pageId="7" pageNumber="478" pagination="23 - 26" refId="ref14216" refString="Okutani, T. (1984) [Symphyta of Japan IX]. Nature and Insects, 19 (1), 23 - 26. [in Japanese]" type="journal article" year="1984">Okutani, 1984: 26</bibRefCitation>
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<bibRefCitation id="EFC99B88FFCA694FF9C9FC5FFC6AFE50" author="Abe, M. &amp; Togashi, I." box="[747,1018,1011,1035]" pageId="7" pageNumber="478" pagination="541 - 560" refId="ref12897" refString="Abe, M. &amp; Togashi, I. (1989) Symphyta. In: Hirashima, Y. (Ed.), A Check List of Japanese Insects. Entomological Laboratory, Faculty of Agriculture, Kyushu University, Fukuoka, pp. 541 - 560. [in Japanese]" type="book chapter" year="1989">Abe &amp; Togashi, 1989: 545</bibRefCitation>
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Azuma
<emphasis id="B92C3A6BFFCA694FFF71FC58FBEFFE50" box="[1107,1151,1011,1035]" italics="true" pageId="7" pageNumber="478">et al</emphasis>
., 2002: 301
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;
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<bibRefCitation id="EFC99B88FFCA694FFBCCFBBEFE18FE71" author="Hara, H." box="[238,392,1042,1066]" pageId="7" pageNumber="478" pagination="42 - 46" refId="ref13152" refString="Hara, H. (2019) Family Diprionidae. In: Editorial Committee of Catalogue of the Insects of Japan (Ed.), Catalogue of the Insects of Japan. Vol. 9. Hymenoptera. Part 1. Symphyta. Entomological Society of Japan, Kyoto, pp. 42 - 46. [in Japanese]" type="book chapter" year="2019">Hara, 2019: 42</bibRefCitation>
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<bibRefCitation id="EFC99B88FFCA694FFAB1FBBEFDA9FE71" author="Hara, H." box="[403,569,1042,1066]" pageId="7" pageNumber="478" pagination="286 - 294" refId="ref13213" refString="Hara, H. (2020) Family Diprionidae. In: Naito, T., Shinohara, A., Hara, H. &amp; Ito, F., Sawflies and Woodwasps of Japan. Hokkaido University Press, Sapporo, pp. 48 - 53 + 286 - 294. [in Japanese]" type="book chapter" year="2020">Hara, 2020: 291</bibRefCitation>
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<paragraph id="8BE7E679FFCA694FFBB5FBF4FAE6FC8D" blockId="7.[151,1437,1112,2039]" pageId="7" pageNumber="478">
<emphasis id="B92C3A6BFFCA694FFBB5FBF4FF7EFE29" bold="true" box="[151,238,1112,1138]" pageId="7" pageNumber="478">Female</emphasis>
(
<figureCitation id="1363FAFCFFCA694FFBDDFBF4FE13FE29" box="[255,387,1112,1138]" captionStart="FIGURES 2" captionStartId="4.[151,264,1746,1770]" captionTargetBox="[166,1435,215,1720]" captionTargetId="figure-19@4.[151,1435,181,1723]" captionTargetPageId="4" captionText="FIGURES 2AH. AF, IK, Gilpinia amamiana: A, B, holotype, female; C, D, female, Tokunoshima Isl.; E, F, male, Amami Oshima Isl. G, H, L, Gilpinia okinawa, holotype, female. A, C, E, G, Dorsal view; B, D, F, H, ventrolateral view; IL, head, frontal view." figureDoi="http://doi.org/10.5281/zenodo.5057391" httpUri="https://zenodo.org/record/5057391/files/figure.png" pageId="7" pageNumber="478">Figs 2AD</figureCitation>
). Length 7.59.0 mm. Head black, yellow on supraclypeal area, narrow area around torulus and clypeus except for narrow lateral part (
<figureCitation id="1363FAFCFFCA694FF95BFBD0FD74FECC" box="[633,740,1148,1175]" captionStart="FIGURES 2" captionStartId="4.[151,264,1746,1770]" captionTargetBox="[166,1435,215,1720]" captionTargetId="figure-19@4.[151,1435,181,1723]" captionTargetPageId="4" captionText="FIGURES 2AH. AF, IK, Gilpinia amamiana: A, B, holotype, female; C, D, female, Tokunoshima Isl.; E, F, male, Amami Oshima Isl. G, H, L, Gilpinia okinawa, holotype, female. A, C, E, G, Dorsal view; B, D, F, H, ventrolateral view; IL, head, frontal view." figureDoi="http://doi.org/10.5281/zenodo.5057391" httpUri="https://zenodo.org/record/5057391/files/figure.png" pageId="7" pageNumber="478">Figs 2I, J</figureCitation>
); postocellar area narrowly or mostly yellow to brown; vertex sometimes narrowly or widely yellow or brown (
<figureCitation id="1363FAFCFFCA694FF9EEFB0CFCC7FEE0" box="[716,855,1184,1211]" captionStart="FIGURES 3" captionStartId="9.[151,264,1786,1810]" captionTargetBox="[151,1435,181,1762]" captionTargetId="figure-19@9.[151,1435,181,1762]" captionTargetPageId="9" captionText="FIGURES 3AX. AH, Head, dorsal view; IP, dorsal part of head, anterolateral view; Q, head, lateral view; RX, antenna, inner view (RT, V, W) or outer view (U, X). AD, IL, QU, Gilpinia hakonensis: A, I, Q, R, lectotype of Lophyrus hakonensis, female; B, J, S, holotype of G. hakonensis var. laticincta, female; C, K, T, paratype of Diprion fukudai, female; D, L, U, male, Honshu. EG, MO, V, W, Gilpinia amamiana: E, M, V, holotype, female; F, N, W, female, Tokunoshima Isl.; G, O, male, Amami Oshima Isl. H, P, X, Gilpinia okinawa, holotype, female. T, Reversed." figureDoi="http://doi.org/10.5281/zenodo.5057395" httpUri="https://zenodo.org/record/5057395/files/figure.png" pageId="7" pageNumber="478">Figs 3M, N</figureCitation>
); gena widely or mostly yellow to brown; antenna black, basally narrowly yellow (
<figureCitation id="1363FAFCFFCA694FF922FB68FD15FE84" box="[512,645,1220,1247]" captionStart="FIGURES 3" captionStartId="9.[151,264,1786,1810]" captionTargetBox="[151,1435,181,1762]" captionTargetId="figure-19@9.[151,1435,181,1762]" captionTargetPageId="9" captionText="FIGURES 3AX. AH, Head, dorsal view; IP, dorsal part of head, anterolateral view; Q, head, lateral view; RX, antenna, inner view (RT, V, W) or outer view (U, X). AD, IL, QU, Gilpinia hakonensis: A, I, Q, R, lectotype of Lophyrus hakonensis, female; B, J, S, holotype of G. hakonensis var. laticincta, female; C, K, T, paratype of Diprion fukudai, female; D, L, U, male, Honshu. EG, MO, V, W, Gilpinia amamiana: E, M, V, holotype, female; F, N, W, female, Tokunoshima Isl.; G, O, male, Amami Oshima Isl. H, P, X, Gilpinia okinawa, holotype, female. T, Reversed." figureDoi="http://doi.org/10.5281/zenodo.5057395" httpUri="https://zenodo.org/record/5057395/files/figure.png" pageId="7" pageNumber="478">Figs 3V, W</figureCitation>
); labrum dark brown (
<figureCitation id="1363FAFCFFCA694FF85DFB68FC7BFE84" box="[895,1003,1220,1247]" captionStart="FIGURES 2" captionStartId="4.[151,264,1746,1770]" captionTargetBox="[166,1435,215,1720]" captionTargetId="figure-19@4.[151,1435,181,1723]" captionTargetPageId="4" captionText="FIGURES 2AH. AF, IK, Gilpinia amamiana: A, B, holotype, female; C, D, female, Tokunoshima Isl.; E, F, male, Amami Oshima Isl. G, H, L, Gilpinia okinawa, holotype, female. A, C, E, G, Dorsal view; B, D, F, H, ventrolateral view; IL, head, frontal view." figureDoi="http://doi.org/10.5281/zenodo.5057391" httpUri="https://zenodo.org/record/5057391/files/figure.png" pageId="7" pageNumber="478">Figs 2I, J</figureCitation>
), sometimes yellow; mandible reddish brown, basally darkened, with yellow spot on outer basal part; palpi yellow, often basally brownish. Thorax yellow (
<figureCitation id="1363FAFCFFCA694FFBBDFAA0FEB3FF7D" box="[159,291,1292,1318]" captionStart="FIGURES 2" captionStartId="4.[151,264,1746,1770]" captionTargetBox="[166,1435,215,1720]" captionTargetId="figure-19@4.[151,1435,181,1723]" captionTargetPageId="4" captionText="FIGURES 2AH. AF, IK, Gilpinia amamiana: A, B, holotype, female; C, D, female, Tokunoshima Isl.; E, F, male, Amami Oshima Isl. G, H, L, Gilpinia okinawa, holotype, female. A, C, E, G, Dorsal view; B, D, F, H, ventrolateral view; IL, head, frontal view." figureDoi="http://doi.org/10.5281/zenodo.5057391" httpUri="https://zenodo.org/record/5057391/files/figure.png" pageId="7" pageNumber="478">Figs 2AD</figureCitation>
), dark brown to black on narrow anteromedial part of pronotum, wide anteromedial part of median mesoscutal lobe, most of lateral mesoscutal lobe, very narrow anteromedial margin and narrow posterior margin of mesoscutellum (Fig. 4I), mesoscutellar appendage, anterior or most part of metascutum, metascutellum, and wide ventral part of mesepisternum (
<figureCitation id="1363FAFCFFCA694FFAD5FAD4FDE6FFC8" box="[503,630,1400,1427]" captionStart="FIGURES 2" captionStartId="4.[151,264,1746,1770]" captionTargetBox="[166,1435,215,1720]" captionTargetId="figure-19@4.[151,1435,181,1723]" captionTargetPageId="4" captionText="FIGURES 2AH. AF, IK, Gilpinia amamiana: A, B, holotype, female; C, D, female, Tokunoshima Isl.; E, F, male, Amami Oshima Isl. G, H, L, Gilpinia okinawa, holotype, female. A, C, E, G, Dorsal view; B, D, F, H, ventrolateral view; IL, head, frontal view." figureDoi="http://doi.org/10.5281/zenodo.5057391" httpUri="https://zenodo.org/record/5057391/files/figure.png" pageId="7" pageNumber="478">Figs 2B, D</figureCitation>
); mesopostnotum medially brown; postspiracular sclerite, anteroventral part of anepimeron and katepimeron except for posterior margin sometimes brown to dark brown; metapleuron partly brown. Legs yellow to brown yellow (
<figureCitation id="1363FAFCFFCA694FF9B7FA6CFC88FF81" box="[661,792,1472,1498]" captionStart="FIGURES 2" captionStartId="4.[151,264,1746,1770]" captionTargetBox="[166,1435,215,1720]" captionTargetId="figure-19@4.[151,1435,181,1723]" captionTargetPageId="4" captionText="FIGURES 2AH. AF, IK, Gilpinia amamiana: A, B, holotype, female; C, D, female, Tokunoshima Isl.; E, F, male, Amami Oshima Isl. G, H, L, Gilpinia okinawa, holotype, female. A, C, E, G, Dorsal view; B, D, F, H, ventrolateral view; IL, head, frontal view." figureDoi="http://doi.org/10.5281/zenodo.5057391" httpUri="https://zenodo.org/record/5057391/files/figure.png" pageId="7" pageNumber="478">Figs 2AD</figureCitation>
); coxae basally brown to dark brown; femora sometimes basally narrowly darkened; hind tibia sometimes narrowly darkened apically; middle and hind tarsi often slightly darkened apically. Wings colorless transparent (
<figureCitation id="1363FAFCFFCA694FF989F9A4FCB8FC79" box="[683,808,1544,1570]" captionStart="FIGURES 2" captionStartId="4.[151,264,1746,1770]" captionTargetBox="[166,1435,215,1720]" captionTargetId="figure-19@4.[151,1435,181,1723]" captionTargetPageId="4" captionText="FIGURES 2AH. AF, IK, Gilpinia amamiana: A, B, holotype, female; C, D, female, Tokunoshima Isl.; E, F, male, Amami Oshima Isl. G, H, L, Gilpinia okinawa, holotype, female. A, C, E, G, Dorsal view; B, D, F, H, ventrolateral view; IL, head, frontal view." figureDoi="http://doi.org/10.5281/zenodo.5057391" httpUri="https://zenodo.org/record/5057391/files/figure.png" pageId="7" pageNumber="478">Figs 2A, C</figureCitation>
); veins brown to black; most of vein C and bases of vein 1A and 2A+3A yellow; vein R1 basal to stigma sometimes widely pale; stigma brown yellow or brown, black on narrow margins and basal third. Abdomen yellow (
<figureCitation id="1363FAFCFFCA694FF9F4F9FCFCC8FC31" box="[726,856,1616,1642]" captionStart="FIGURES 2" captionStartId="4.[151,264,1746,1770]" captionTargetBox="[166,1435,215,1720]" captionTargetId="figure-19@4.[151,1435,181,1723]" captionTargetPageId="4" captionText="FIGURES 2AH. AF, IK, Gilpinia amamiana: A, B, holotype, female; C, D, female, Tokunoshima Isl.; E, F, male, Amami Oshima Isl. G, H, L, Gilpinia okinawa, holotype, female. A, C, E, G, Dorsal view; B, D, F, H, ventrolateral view; IL, head, frontal view." figureDoi="http://doi.org/10.5281/zenodo.5057391" httpUri="https://zenodo.org/record/5057391/files/figure.png" pageId="7" pageNumber="478">Figs 2AD</figureCitation>
); first tergum mostly yellow or mostly dark brown; second to eighth terga each dark brown posteriorly; ninth tergum marginally dark brown narrowly or widely; tenth tergum laterally and posteriorly brown to dark brown; posterior sterna each darkened along posterior margin; cercus dark brown; ovipositor sheath with valvifer 2 yellow to brown, anteriorly darkened and valvula 3 mostly black.
</paragraph>
<paragraph id="8BE7E679FFCA694FFBE5F94CFE62FD89" blockId="7.[151,1437,1112,2039]" pageId="7" pageNumber="478">
Postocellar area slightly or distinctly swollen; lateral furrow distinct, vestigial or indistinct (
<figureCitation id="1363FAFCFFCA694FFFE1F94CFAD8FCA0" box="[1219,1352,1760,1787]" captionStart="FIGURES 3" captionStartId="9.[151,264,1786,1810]" captionTargetBox="[151,1435,181,1762]" captionTargetId="figure-19@9.[151,1435,181,1762]" captionTargetPageId="9" captionText="FIGURES 3AX. AH, Head, dorsal view; IP, dorsal part of head, anterolateral view; Q, head, lateral view; RX, antenna, inner view (RT, V, W) or outer view (U, X). AD, IL, QU, Gilpinia hakonensis: A, I, Q, R, lectotype of Lophyrus hakonensis, female; B, J, S, holotype of G. hakonensis var. laticincta, female; C, K, T, paratype of Diprion fukudai, female; D, L, U, male, Honshu. EG, MO, V, W, Gilpinia amamiana: E, M, V, holotype, female; F, N, W, female, Tokunoshima Isl.; G, O, male, Amami Oshima Isl. H, P, X, Gilpinia okinawa, holotype, female. T, Reversed." figureDoi="http://doi.org/10.5281/zenodo.5057395" httpUri="https://zenodo.org/record/5057395/files/figure.png" pageId="7" pageNumber="478">Figs 3M, N</figureCitation>
). OOL: POL:OOCL 0.91.1:1.0:0.80.9. OOCL 2.33.1 × lateral ocellus length. Inner margins of eyes convergent below (
<figureCitation id="1363FAFCFFCA694FFBBDF884FE9EFD18" box="[159,270,1832,1859]" captionStart="FIGURES 2" captionStartId="4.[151,264,1746,1770]" captionTargetBox="[166,1435,215,1720]" captionTargetId="figure-19@4.[151,1435,181,1723]" captionTargetPageId="4" captionText="FIGURES 2AH. AF, IK, Gilpinia amamiana: A, B, holotype, female; C, D, female, Tokunoshima Isl.; E, F, male, Amami Oshima Isl. G, H, L, Gilpinia okinawa, holotype, female. A, C, E, G, Dorsal view; B, D, F, H, ventrolateral view; IL, head, frontal view." figureDoi="http://doi.org/10.5281/zenodo.5057391" httpUri="https://zenodo.org/record/5057391/files/figure.png" pageId="7" pageNumber="478">Figs 2I, J</figureCitation>
). Distance between eyes at level of torulus 1.82.1 × major axis of eye. Supraclypeal area slightly or moderately swollen. Clypeus with ventral margin truncate or very slightly concave medially; median height 1.01.3 × torulus height. Malar space 0.91.2 × median ocellus width. Antenna with 2123 antennomeres (
<figureCitation id="1363FAFCFFCA694FFFF9F8DCFACDFDD0" box="[1243,1373,1904,1931]" captionStart="FIGURES 3" captionStartId="9.[151,264,1786,1810]" captionTargetBox="[151,1435,181,1762]" captionTargetId="figure-19@9.[151,1435,181,1762]" captionTargetPageId="9" captionText="FIGURES 3AX. AH, Head, dorsal view; IP, dorsal part of head, anterolateral view; Q, head, lateral view; RX, antenna, inner view (RT, V, W) or outer view (U, X). AD, IL, QU, Gilpinia hakonensis: A, I, Q, R, lectotype of Lophyrus hakonensis, female; B, J, S, holotype of G. hakonensis var. laticincta, female; C, K, T, paratype of Diprion fukudai, female; D, L, U, male, Honshu. EG, MO, V, W, Gilpinia amamiana: E, M, V, holotype, female; F, N, W, female, Tokunoshima Isl.; G, O, male, Amami Oshima Isl. H, P, X, Gilpinia okinawa, holotype, female. T, Reversed." figureDoi="http://doi.org/10.5281/zenodo.5057395" httpUri="https://zenodo.org/record/5057395/files/figure.png" pageId="7" pageNumber="478">Figs 3V, W</figureCitation>
); first flagellomere without serration, with apical breadth in lateral view 0.81.1 × dorsal length; serrations around fourth to eighth flagellomeres largest.
</paragraph>
<paragraph id="8BE7E679FFCA6940FBE5F870FAD9FAA2" blockId="7.[151,1437,1112,2039]" lastBlockId="8.[151,1437,150,1905]" lastPageId="8" lastPageNumber="479" pageId="7" pageNumber="478">Distance between cenchri 0.60.7 × major axis of cenchrus (Fig. 4I). Metascutellum in dorsal view not or very slightly angulated posterolaterally with posterior margin convex; median length 0.30.5 × major axis of cenchrus. Hind leg with posterior tibial spur with length 0.81.0 × dorsal length of first tarsomere, 0.81.1 × apical breadth of tibia in lateral view; dorsal length of second and third tarsomeres combined 1.31.6 × dorsal length of first.</paragraph>
<paragraph id="8BE7E679FFC56940FBE5FEAFFCF2F842" blockId="8.[151,1437,150,1905]" pageId="8" pageNumber="479">
Valvula
<quantity id="4CA04B9CFFC56940FA04FEAFFEC3FB46" box="[294,339,259,285]" metricMagnitude="-2" metricUnit="m" metricValue="7.62" pageId="8" pageNumber="479" unit="in" value="3.0">3 in</quantity>
dorsal view about 3 × as wide as cercus, apically with round lateral projection and small acute medial projection (
<figureCitation id="1363FAFCFFC56940FA49FE8BFE70FB1A" box="[363,480,294,321]" captionStart="FIGURES 5" captionStartId="11.[151,264,1868,1892]" captionTargetBox="[204,1384,181,1835]" captionTargetId="figure-19@11.[204,1384,181,1843]" captionTargetPageId="11" captionText="FIGURES 5AU.AP, Valvula 3, dorsal and posterior views; QU, ovipositor or lance, dorsal view (Q, S, T) or lateral view (R, U). AF, Q, R, Gilpinia hakonensis: AB, lectotype of Lophyrus hakonensis; C, D, holotype of G. hakonensis var. laticincta; E, F, P, paratype of Diprion fukudai; Q, R, Honshu. GJ, S, Gilpinia amamiana: G, H, holotype; I, J, Tokunoshima Isl.; S, Amami Oshima Isl. K, L, T, U, Gilpinia okinawa, holotype. M, N, Gilpinia frutetorum, Sweden. O, P, Gilpinia laricis, Germany. R, Reversed." figureDoi="http://doi.org/10.5281/zenodo.5057397" httpUri="https://zenodo.org/record/5057397/files/figure.png" pageId="8" pageNumber="479">Figs 5G, I</figureCitation>
); scopa about 1.52.0 × as wide as cercus, in posterior view long oval, about 2.02.5 × as high as wide (
<figureCitation id="1363FAFCFFC56940FA47FEE7FE4AFB3E" box="[357,474,330,357]" captionStart="FIGURES 5" captionStartId="11.[151,264,1868,1892]" captionTargetBox="[204,1384,181,1835]" captionTargetId="figure-19@11.[204,1384,181,1843]" captionTargetPageId="11" captionText="FIGURES 5AU.AP, Valvula 3, dorsal and posterior views; QU, ovipositor or lance, dorsal view (Q, S, T) or lateral view (R, U). AF, Q, R, Gilpinia hakonensis: AB, lectotype of Lophyrus hakonensis; C, D, holotype of G. hakonensis var. laticincta; E, F, P, paratype of Diprion fukudai; Q, R, Honshu. GJ, S, Gilpinia amamiana: G, H, holotype; I, J, Tokunoshima Isl.; S, Amami Oshima Isl. K, L, T, U, Gilpinia okinawa, holotype. M, N, Gilpinia frutetorum, Sweden. O, P, Gilpinia laricis, Germany. R, Reversed." figureDoi="http://doi.org/10.5281/zenodo.5057397" httpUri="https://zenodo.org/record/5057397/files/figure.png" pageId="8" pageNumber="479">Figs 5H, J</figureCitation>
). Lance with posterior projection on processus articularis apically extending to or near most basal annular suture of lance (
<figureCitation id="1363FAFCFFC56940F910FEC3FD19FBD2" box="[562,649,366,393]" captionStart="FIGURES 5" captionStartId="11.[151,264,1868,1892]" captionTargetBox="[204,1384,181,1835]" captionTargetId="figure-19@11.[204,1384,181,1843]" captionTargetPageId="11" captionText="FIGURES 5AU.AP, Valvula 3, dorsal and posterior views; QU, ovipositor or lance, dorsal view (Q, S, T) or lateral view (R, U). AF, Q, R, Gilpinia hakonensis: AB, lectotype of Lophyrus hakonensis; C, D, holotype of G. hakonensis var. laticincta; E, F, P, paratype of Diprion fukudai; Q, R, Honshu. GJ, S, Gilpinia amamiana: G, H, holotype; I, J, Tokunoshima Isl.; S, Amami Oshima Isl. K, L, T, U, Gilpinia okinawa, holotype. M, N, Gilpinia frutetorum, Sweden. O, P, Gilpinia laricis, Germany. R, Reversed." figureDoi="http://doi.org/10.5281/zenodo.5057397" httpUri="https://zenodo.org/record/5057397/files/figure.png" pageId="8" pageNumber="479">Fig. 5S</figureCitation>
). Lancet
<figureCitation id="1363FAFCFFC56940F9D7FEC3FCE1FBD2" box="[757,881,367,393]" captionStart="FIGURES 6" captionStartId="12.[151,264,1869,1893]" captionTargetBox="[154,1435,181,1833]" captionTargetId="figure-19@12.[151,1435,181,1845]" captionTargetPageId="12" captionText="FIGURES 6AJ. Lancet. AF, Gilpinia hakonensis: A, lectotype of Lophyrus hakonensis; B, holotype of G. hakonensis var. laticincta; C, Kyushu, D, Honshu, E, paratype of Diprion fukudai. FH, Gilpinia amamiana: F, holotype; G, Amami Oshima Isl.; H, Tokunoshima Isl. I, J, Gilpinia okinawa: I, holotype; J, Okinawa Isl. 1, 2, 5, first (most basal), second and fifth annuli. DF, H, Reversed." figureDoi="http://doi.org/10.5281/zenodo.5057401" httpUri="https://zenodo.org/record/5057401/files/figure.png" pageId="8" pageNumber="479">Figs 6FH</figureCitation>
; lamnium with 9 annuli and length from ventral end of first ctenidium to apex 2.12.3 × maximum width; ctenidia nearly straight except second arched and third sometimes slightly arched; sclerite before first annulus narrow; serrulae of second and third annuli each with distinct precalcar (precalcar sometimes absent, probably due to severe wear,
<figureCitation id="1363FAFCFFC56940F8AFFE77FC75FBAD" box="[909,997,475,502]" captionStart="FIGURES 6" captionStartId="12.[151,264,1869,1893]" captionTargetBox="[154,1435,181,1833]" captionTargetId="figure-19@12.[151,1435,181,1845]" captionTargetPageId="12" captionText="FIGURES 6AJ. Lancet. AF, Gilpinia hakonensis: A, lectotype of Lophyrus hakonensis; B, holotype of G. hakonensis var. laticincta; C, Kyushu, D, Honshu, E, paratype of Diprion fukudai. FH, Gilpinia amamiana: F, holotype; G, Amami Oshima Isl.; H, Tokunoshima Isl. I, J, Gilpinia okinawa: I, holotype; J, Okinawa Isl. 1, 2, 5, first (most basal), second and fifth annuli. DF, H, Reversed." figureDoi="http://doi.org/10.5281/zenodo.5057401" httpUri="https://zenodo.org/record/5057401/files/figure.png" pageId="8" pageNumber="479">Fig. 6H</figureCitation>
); serrula of second annulus much larger than other serrulae, with precalcar relatively close to postcalcar.
</paragraph>
<paragraph id="8BE7E679FFC56940FBE5FD8FFDF3F8DE" blockId="8.[151,1437,150,1905]" pageId="8" pageNumber="479">
Head and thorax mostly covered with dense punctures (
<figureCitation id="1363FAFCFFC56940F860FD8FFC59F866" box="[834,969,547,573]" captionStart="FIGURES 3" captionStartId="9.[151,264,1786,1810]" captionTargetBox="[151,1435,181,1762]" captionTargetId="figure-19@9.[151,1435,181,1762]" captionTargetPageId="9" captionText="FIGURES 3AX. AH, Head, dorsal view; IP, dorsal part of head, anterolateral view; Q, head, lateral view; RX, antenna, inner view (RT, V, W) or outer view (U, X). AD, IL, QU, Gilpinia hakonensis: A, I, Q, R, lectotype of Lophyrus hakonensis, female; B, J, S, holotype of G. hakonensis var. laticincta, female; C, K, T, paratype of Diprion fukudai, female; D, L, U, male, Honshu. EG, MO, V, W, Gilpinia amamiana: E, M, V, holotype, female; F, N, W, female, Tokunoshima Isl.; G, O, male, Amami Oshima Isl. H, P, X, Gilpinia okinawa, holotype, female. T, Reversed." figureDoi="http://doi.org/10.5281/zenodo.5057395" httpUri="https://zenodo.org/record/5057395/files/figure.png" pageId="8" pageNumber="479">Figs 3M, N</figureCitation>
, 4I, J); punctures very often fused; inter- spaces generally narrower than diameters of punctures. Punctures on mesoscutellum much larger than those of mesoscutum. Abdomen microsculptured.
</paragraph>
<paragraph id="8BE7E679FFC56940FBE5FD23FAA7F962" blockId="8.[151,1437,150,1905]" pageId="8" pageNumber="479">
<emphasis id="B92C3A6BFFC56940FBE5FD23FE94F8F2" bold="true" box="[199,260,655,681]" pageId="8" pageNumber="479">Male</emphasis>
(
<figureCitation id="1363FAFCFFC56940FA36FD23FE00F8F2" box="[276,400,655,681]" captionStart="FIGURES 2" captionStartId="4.[151,264,1746,1770]" captionTargetBox="[166,1435,215,1720]" captionTargetId="figure-19@4.[151,1435,181,1723]" captionTargetPageId="4" captionText="FIGURES 2AH. AF, IK, Gilpinia amamiana: A, B, holotype, female; C, D, female, Tokunoshima Isl.; E, F, male, Amami Oshima Isl. G, H, L, Gilpinia okinawa, holotype, female. A, C, E, G, Dorsal view; B, D, F, H, ventrolateral view; IL, head, frontal view." figureDoi="http://doi.org/10.5281/zenodo.5057391" httpUri="https://zenodo.org/record/5057391/files/figure.png" pageId="8" pageNumber="479">Figs 2E, F</figureCitation>
). As in female, but differing as follows except for usual sexual differences. Length 7.08.0 mm. Black, but antenna sometimes mostly dark brown, labrum dark brown, mandible reddish brown except for narrow base, pronotum posterolaterally widely yellow, tegula yellow, medially dark brown, abdomen ventrally yellow, sometimes with laterotergites each narrowly brown along posterior margin and subgenital plate dark brown. Legs yellow; coxae black except for narrow apices; trochanters basally black; hind tibia apically brown yellow.
</paragraph>
<paragraph id="8BE7E679FFC56940FBE5FCEFFB4DFEFA" blockId="8.[151,1437,150,1905]" pageId="8" pageNumber="479">
OOL:POL:OOCL 0.80.9:1.0:0.6. OOCL 1.62.0 × lateral ocellus length. Distance between eyes at level of torulus 2.1 × major axis of eye. Clypeus with median height 0.9 × torulus height. Malar space 1.11.2 × median ocellus width. Antenna with 2425 antennomeres; flagellomeres each with two rami, but most basal and preapical flagellomeres with one ramus and apical most flagellomere without ramus. Distance between cenchri 0.71.0 × major axis of cenchrus (Fig. 4K). Metascutellum in dorsal view posterolaterally slightly angulated, with posterior margin slightly convex; median length 0.50.6 × major axis of cenchrus. Posterior hind tibial spur.with length 0.81.0 × dorsal length of hind first tarsomere, 1.11.2 × apical breadth of tibia in lateral view; dorsal length of second and third tarsomeres combined 1.31.4× as that of first. Subgenital plate apically nearly truncate in dorsal or ventral view (
<figureCitation id="1363FAFCFFC56940FA6CFBCFFE34FE26" box="[334,420,1123,1149]" captionStart="FIGURES 2" captionStartId="4.[151,264,1746,1770]" captionTargetBox="[166,1435,215,1720]" captionTargetId="figure-19@4.[151,1435,181,1723]" captionTargetPageId="4" captionText="FIGURES 2AH. AF, IK, Gilpinia amamiana: A, B, holotype, female; C, D, female, Tokunoshima Isl.; E, F, male, Amami Oshima Isl. G, H, L, Gilpinia okinawa, holotype, female. A, C, E, G, Dorsal view; B, D, F, H, ventrolateral view; IL, head, frontal view." figureDoi="http://doi.org/10.5281/zenodo.5057391" httpUri="https://zenodo.org/record/5057391/files/figure.png" pageId="8" pageNumber="479">Fig. 2E</figureCitation>
). Genitalia
<figureCitation id="1363FAFCFFC56940F906FBCFFD0DFE26" box="[548,669,1123,1149]" captionStart="FIGURES 7" captionStartId="13.[151,264,1915,1939]" captionTargetBox="[151,1423,960,1891]" captionTargetId="figure-347@13.[151,1435,957,1891]" captionTargetPageId="13" captionText="FIGURES 7AF. A, D, Male genitalia, dorsal view; B, E, male genitalia (penis valve removed), ventral view; C, F, penis valve, lateral view (left dorsal). AC, Gilpinia hakonensis, Honshu. DF, Gilpinia amamiana, Amami Oshima Isl." figureDoi="http://doi.org/10.5281/zenodo.5057405" httpUri="https://zenodo.org/record/5057405/files/figure.png" pageId="8" pageNumber="479">Figs 7DF</figureCitation>
; in ventral view, parapenis apically truncate, with lateral margin very short, (
<figureCitation id="1363FAFCFFC56940FBC4FB2BFEADFEFA" box="[230,317,1159,1185]" captionStart="FIGURES 7" captionStartId="13.[151,264,1915,1939]" captionTargetBox="[151,1423,960,1891]" captionTargetId="figure-347@13.[151,1435,957,1891]" captionTargetPageId="13" captionText="FIGURES 7AF. A, D, Male genitalia, dorsal view; B, E, male genitalia (penis valve removed), ventral view; C, F, penis valve, lateral view (left dorsal). AC, Gilpinia hakonensis, Honshu. DF, Gilpinia amamiana, Amami Oshima Isl." figureDoi="http://doi.org/10.5281/zenodo.5057405" httpUri="https://zenodo.org/record/5057405/files/figure.png" pageId="8" pageNumber="479">Fig. 7E</figureCitation>
); penis valve with valviceps exceedingly convex ventrally at apical third (
<figureCitation id="1363FAFCFFC56940FF58FB2BFB40FEFA" box="[1146,1232,1159,1185]" captionStart="FIGURES 7" captionStartId="13.[151,264,1915,1939]" captionTargetBox="[151,1423,960,1891]" captionTargetId="figure-347@13.[151,1435,957,1891]" captionTargetPageId="13" captionText="FIGURES 7AF. A, D, Male genitalia, dorsal view; B, E, male genitalia (penis valve removed), ventral view; C, F, penis valve, lateral view (left dorsal). AC, Gilpinia hakonensis, Honshu. DF, Gilpinia amamiana, Amami Oshima Isl." figureDoi="http://doi.org/10.5281/zenodo.5057405" httpUri="https://zenodo.org/record/5057405/files/figure.png" pageId="8" pageNumber="479">Fig. 7F</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C342B5F2FFC56940FBE5FB06FCB8FFBE" pageId="8" pageNumber="479" type="materials_examined">
<paragraph id="8BE7E679FFC56940FBE5FB06FEBFFF6A" blockId="8.[151,1437,150,1905]" pageId="8" pageNumber="479">
<emphasis id="B92C3A6BFFC56940FBE5FB06FE3AFE9F" bold="true" box="[199,426,1194,1220]" pageId="8" pageNumber="479">Material examined</emphasis>
.
<typeStatus id="54E358DBFFC56940FA9BFB07FDB1FE9E" box="[441,545,1195,1221]" pageId="8" pageNumber="479" type="holotype">Holotype</typeStatus>
(
<figureCitation id="1363FAFCFFC56940F912FB07FD57FE9E" box="[560,711,1195,1221]" captionStart="FIGURES 2" captionStartId="4.[151,264,1746,1770]" captionTargetBox="[166,1435,215,1720]" captionTargetId="figure-19@4.[151,1435,181,1723]" captionTargetPageId="4" captionText="FIGURES 2AH. AF, IK, Gilpinia amamiana: A, B, holotype, female; C, D, female, Tokunoshima Isl.; E, F, male, Amami Oshima Isl. G, H, L, Gilpinia okinawa, holotype, female. A, C, E, G, Dorsal view; B, D, F, H, ventrolateral view; IL, head, frontal view." figureDoi="http://doi.org/10.5281/zenodo.5057391" httpUri="https://zenodo.org/record/5057391/files/figure.png" pageId="8" pageNumber="479">Figs 2A, B, I</figureCitation>
,
<figureCitation id="1363FAFCFFC56940F9F6FB07FCACFE9E" box="[724,828,1195,1221]" captionStart="FIGURES 3" captionStartId="9.[151,264,1786,1810]" captionTargetBox="[151,1435,181,1762]" captionTargetId="figure-19@9.[151,1435,181,1762]" captionTargetPageId="9" captionText="FIGURES 3AX. AH, Head, dorsal view; IP, dorsal part of head, anterolateral view; Q, head, lateral view; RX, antenna, inner view (RT, V, W) or outer view (U, X). AD, IL, QU, Gilpinia hakonensis: A, I, Q, R, lectotype of Lophyrus hakonensis, female; B, J, S, holotype of G. hakonensis var. laticincta, female; C, K, T, paratype of Diprion fukudai, female; D, L, U, male, Honshu. EG, MO, V, W, Gilpinia amamiana: E, M, V, holotype, female; F, N, W, female, Tokunoshima Isl.; G, O, male, Amami Oshima Isl. H, P, X, Gilpinia okinawa, holotype, female. T, Reversed." figureDoi="http://doi.org/10.5281/zenodo.5057395" httpUri="https://zenodo.org/record/5057395/files/figure.png" pageId="8" pageNumber="479">3E, M, V</figureCitation>
, 4I, J, 5G, H, 6F):
<specimenCount id="9D5E2DF0FFC56940FF33FB07FBB6FE9E" box="[1041,1062,1195,1221]" pageId="8" pageNumber="479" type="female"></specimenCount>
, “Hatsuno Amami 2. April. 1963 Y. Arita”, “
<typeStatus id="54E358DBFFC56940FA3AFB63FE13FEB2" box="[280,387,1231,1257]" pageId="8" pageNumber="479" type="holotype">Holotype</typeStatus>
<taxonomicName id="4C589DFAFFC56940FAACFB63FC89FEB2" authority="Okutani, 1965" authorityName="Okutani" authorityYear="1965" box="[398,793,1230,1257]" class="Insecta" family="Diprionidae" genus="Gilpinia" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="479" phylum="Arthropoda" rank="species" species="amamiana">
Gilpinia amamiana
<bibRefCitation id="EFC99B88FFC56940F953FB63FC89FEB2" author="Okutani, T." box="[625,793,1230,1257]" pageId="8" pageNumber="479" pagination="73 - 84" refId="ref14145" refString="Okutani, T. (1965) Sawflies and horntails from the Ryukyus. Kontyu, 33, 73 - 84." type="journal article" year="1965">Okutani, 1965</bibRefCitation>
</taxonomicName>
”, “lancet mounted”, “2”, “
<typeStatus id="54E358DBFFC56940FF6CFB63FB29FEB2" box="[1102,1209,1231,1257]" pageId="8" pageNumber="479" type="holotype">Holotype</typeStatus>
<taxonomicName id="4C589DFAFFC56940FFE1FB63FF67FF56" authority="Okutani" authorityName="Okutani" authorityYear="1965" class="Insecta" family="Diprionidae" genus="Gilpinia" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="479" phylum="Arthropoda" rank="species" species="amamiana">Gilpinia amamiana Okutani</taxonomicName>
”. The photo of the lancet (
<figureCitation id="1363FAFCFFC56940F93DFB5FFDE4FF55" box="[543,628,1267,1294]" captionStart="FIGURES 6" captionStartId="12.[151,264,1869,1893]" captionTargetBox="[154,1435,181,1833]" captionTargetId="figure-19@12.[151,1435,181,1845]" captionTargetPageId="12" captionText="FIGURES 6AJ. Lancet. AF, Gilpinia hakonensis: A, lectotype of Lophyrus hakonensis; B, holotype of G. hakonensis var. laticincta; C, Kyushu, D, Honshu, E, paratype of Diprion fukudai. FH, Gilpinia amamiana: F, holotype; G, Amami Oshima Isl.; H, Tokunoshima Isl. I, J, Gilpinia okinawa: I, holotype; J, Okinawa Isl. 1, 2, 5, first (most basal), second and fifth annuli. DF, H, Reversed." figureDoi="http://doi.org/10.5281/zenodo.5057401" httpUri="https://zenodo.org/record/5057401/files/figure.png" pageId="8" pageNumber="479">Fig. 6E</figureCitation>
) was taken by Smith in 1979. After that, the slide of the lancet probably has been lost.
</paragraph>
<paragraph id="8BE7E679FFC56940FBE5FA97FCB8FFBE" blockId="8.[151,1437,150,1905]" pageId="8" pageNumber="479">
Other material examined:
<materialsCitation id="3B30EC24FFC56940FACDFA97FC91FF0E" box="[495,769,1339,1365]" country="Japan" location="Amami Islands" pageId="8" pageNumber="479" specimenCount="1">
<collectingCountry id="F34FA6E9FFC56940FACDFA97FDD8FF0E" box="[495,584,1339,1365]" name="Japan" pageId="8" pageNumber="479">JAPAN</collectingCountry>
:
<location id="8E87B0A2FFC56940F973FA97FC91FF0E" LSID="urn:lsid:plazi:treatment:03F1576FFFCA6942FBB5FC26FD1CFD8E:8E87B0A2FFC56940F973FA97FC91FF0E" box="[593,769,1339,1365]" country="Japan" name="Amami Islands" pageId="8" pageNumber="479">Amami Islands</location>
</materialsCitation>
:
<materialsCitation id="3B30EC24FFC56940F82EFA97FC8DFF22" collectingDate="2009-06-23" collectionCode="VI" collectorName="M. Owada" country="Japan" county="Nagakumo-toge" location="Amami Oshima Island" municipality="Tatsugo" pageId="8" pageNumber="479" specimenCount="1" specimenCount-male="1">
<specimenCount id="9D5E2DF0FFC56940F82EFA97FCBFFF0E" box="[780,815,1339,1365]" pageId="8" pageNumber="479" type="male">1♂</specimenCount>
(
<figureCitation id="1363FAFCFFC56940F81DFA97FC72FF0E" box="[831,994,1339,1365]" captionStart="FIGURES 2" captionStartId="4.[151,264,1746,1770]" captionTargetBox="[166,1435,215,1720]" captionTargetId="figure-19@4.[151,1435,181,1723]" captionTargetPageId="4" captionText="FIGURES 2AH. AF, IK, Gilpinia amamiana: A, B, holotype, female; C, D, female, Tokunoshima Isl.; E, F, male, Amami Oshima Isl. G, H, L, Gilpinia okinawa, holotype, female. A, C, E, G, Dorsal view; B, D, F, H, ventrolateral view; IL, head, frontal view." figureDoi="http://doi.org/10.5281/zenodo.5057391" httpUri="https://zenodo.org/record/5057391/files/figure.png" pageId="8" pageNumber="479">Figs 2E, F, K</figureCitation>
,
<figureCitation id="1363FAFCFFC56940F8CCFA97FBA5FF0E" box="[1006,1077,1339,1365]" captionStart="FIGURES 3" captionStartId="9.[151,264,1786,1810]" captionTargetBox="[151,1435,181,1762]" captionTargetId="figure-19@9.[151,1435,181,1762]" captionTargetPageId="9" captionText="FIGURES 3AX. AH, Head, dorsal view; IP, dorsal part of head, anterolateral view; Q, head, lateral view; RX, antenna, inner view (RT, V, W) or outer view (U, X). AD, IL, QU, Gilpinia hakonensis: A, I, Q, R, lectotype of Lophyrus hakonensis, female; B, J, S, holotype of G. hakonensis var. laticincta, female; C, K, T, paratype of Diprion fukudai, female; D, L, U, male, Honshu. EG, MO, V, W, Gilpinia amamiana: E, M, V, holotype, female; F, N, W, female, Tokunoshima Isl.; G, O, male, Amami Oshima Isl. H, P, X, Gilpinia okinawa, holotype, female. T, Reversed." figureDoi="http://doi.org/10.5281/zenodo.5057395" httpUri="https://zenodo.org/record/5057395/files/figure.png" pageId="8" pageNumber="479">3G, O</figureCitation>
, 4K, L, 7DF),
<location id="8E87B0A2FFC56940FFCBFA97FF4EFF22" LSID="urn:lsid:plazi:treatment:03F1576FFFCA6942FBB5FC26FD1CFD8E:8E87B0A2FFC56940FFCBFA97FF4EFF22" country="Japan" county="Nagakumo-toge" municipality="Tatsugo" name="Amami Oshima Island" pageId="8" pageNumber="479">Amami Oshima Island</location>
,
<collectingMunicipality id="6B837C03FFC56940FBCAFAF3FED3FF22" box="[232,323,1375,1401]" pageId="8" pageNumber="479">Tatsugo</collectingMunicipality>
,
<collectingCounty id="62869EF5FFC56940FA6FFAF3FD94FF22" box="[333,516,1375,1401]" pageId="8" pageNumber="479">Nagakumo-toge</collectingCounty>
,
<date id="FFE6C0B9FFC56940F92CFAF3FD0BFF22" box="[526,667,1375,1401]" pageId="8" pageNumber="479" value="2009-06-23">
<collectingDate id="EFA23951FFC56940F92CFAF3FD0BFF22" box="[526,667,1375,1401]" pageId="8" pageNumber="479" value="2009-06-23">
23.
<collectionCode id="ED497EBCFFC56940F914FAF3FDC7FF22" box="[566,599,1375,1401]" country="Norway" name="Mykotektet, National Veterinary Institute" pageId="8" pageNumber="479">VI</collectionCode>
. 2009
</collectingDate>
</date>
,
<collectorName id="26AD83AFFFC56940F987FAF3FC8DFF22" box="[677,797,1375,1401]" pageId="8" pageNumber="479">M. Owada</collectorName>
</materialsCitation>
;
<materialsCitation id="3B30EC24FFC56940F805FAF3FE82FFC6" collectingDate="1982-09-28" collectorName="S. Yamane" country="Japan" location="Amami Oshima Island" municipality="Naze" pageId="8" pageNumber="479" specimenCount="1" specimenCount-female="1">
<specimenCount id="9D5E2DF0FFC56940F805FAF3FCDAFF22" box="[807,842,1375,1401]" pageId="8" pageNumber="479" type="female">1♀</specimenCount>
(
<figureCitation id="1363FAFCFFC56940F87AFAF3FC3AFF22" box="[856,938,1374,1401]" captionStart="FIGURES 5" captionStartId="11.[151,264,1868,1892]" captionTargetBox="[204,1384,181,1835]" captionTargetId="figure-19@11.[204,1384,181,1843]" captionTargetPageId="11" captionText="FIGURES 5AU.AP, Valvula 3, dorsal and posterior views; QU, ovipositor or lance, dorsal view (Q, S, T) or lateral view (R, U). AF, Q, R, Gilpinia hakonensis: AB, lectotype of Lophyrus hakonensis; C, D, holotype of G. hakonensis var. laticincta; E, F, P, paratype of Diprion fukudai; Q, R, Honshu. GJ, S, Gilpinia amamiana: G, H, holotype; I, J, Tokunoshima Isl.; S, Amami Oshima Isl. K, L, T, U, Gilpinia okinawa, holotype. M, N, Gilpinia frutetorum, Sweden. O, P, Gilpinia laricis, Germany. R, Reversed." figureDoi="http://doi.org/10.5281/zenodo.5057397" httpUri="https://zenodo.org/record/5057397/files/figure.png" pageId="8" pageNumber="479">Fig. 5S</figureCitation>
),
<location id="8E87B0A2FFC56940F89FFAF3FB2AFF22" LSID="urn:lsid:plazi:treatment:03F1576FFFCA6942FBB5FC26FD1CFD8E:8E87B0A2FFC56940F89FFAF3FB2AFF22" box="[957,1210,1375,1401]" country="Japan" municipality="Naze" name="Amami Oshima Island" pageId="8" pageNumber="479">Amami Oshima Island</location>
,
<collectingMunicipality id="6B837C03FFC56940FFE6FAF3FA90FF22" box="[1220,1280,1375,1401]" pageId="8" pageNumber="479">Naze</collectingMunicipality>
,
<date id="FFE6C0B9FFC56940FE28FAF3FA08FF22" box="[1290,1432,1375,1401]" pageId="8" pageNumber="479" value="1982-09-28">
<collectingDate id="EFA23951FFC56940FE28FAF3FA08FF22" box="[1290,1432,1375,1401]" pageId="8" pageNumber="479" value="1982-09-28">28. IX. 1982</collectingDate>
</date>
,
<collectorName id="26AD83AFFFC56940FBB5FA2FFE82FFC6" box="[151,274,1411,1437]" pageId="8" pageNumber="479">S. Yamane</collectorName>
</materialsCitation>
;
<materialsCitation id="3B30EC24FFC56940FA3FFA2FFB1BFFC6" box="[285,1163,1411,1438]" collectingDate="2012-03-23" collectorName="H. Ono" country="Japan" county="Kinsakubaru" location="Figs" municipality="Amami Oshima Island" pageId="8" pageNumber="479" specimenCount="1" specimenCount-female="1">
<specimenCount id="9D5E2DF0FFC56940FA3FFA2FFED0FFC6" box="[285,320,1411,1437]" pageId="8" pageNumber="479" type="female">1♀</specimenCount>
(
<location id="8E87B0A2FFC56940FA6DFA2FFE11FFC6" LSID="urn:lsid:plazi:treatment:03F1576FFFCA6942FBB5FC26FD1CFD8E:8E87B0A2FFC56940FA6DFA2FFE11FFC6" box="[335,385,1411,1437]" country="Japan" county="Kinsakubaru" municipality="Amami Oshima Island" name="Figs" pageId="8" pageNumber="479">Figs</location>
4Q, 6G),
<collectingMunicipality id="6B837C03FFC56940FAD3FA2FFD61FFC6" box="[497,753,1411,1437]" pageId="8" pageNumber="479">Amami Oshima Island</collectingMunicipality>
,
<collectingCounty id="62869EF5FFC56940F9DFFA2FFC1FFFC6" box="[765,911,1411,1437]" pageId="8" pageNumber="479">Kinsakubaru</collectingCounty>
,
<date id="FFE6C0B9FFC56940F8B8FA2FFBB9FFC6" box="[922,1065,1411,1437]" pageId="8" pageNumber="479" value="2012-03-23">
<collectingDate id="EFA23951FFC56940F8B8FA2FFBB9FFC6" box="[922,1065,1411,1437]" pageId="8" pageNumber="479" value="2012-03-23">23. III. 2012</collectingDate>
</date>
,
<collectorName id="26AD83AFFFC56940FF16FA2FFB1BFFC6" box="[1076,1163,1411,1437]" pageId="8" pageNumber="479">H. Ono</collectorName>
</materialsCitation>
;
<materialsCitation id="3B30EC24FFC56940FFB4FA2FFB82FF9A" collectingDate="2021-03-26" collectorName="H. Hara" country="Japan" latitude="28.35" location="Amami Oshima" longLatPrecision="925" longitude="129.46584" municipality="Amami" pageId="8" pageNumber="479" specimenCount="2" specimenCount-male="2">
<specimenCount id="9D5E2DF0FFC56940FFB4FA2FFB2BFFC6" box="[1174,1211,1411,1437]" pageId="8" pageNumber="479" type="male">2♂</specimenCount>
,
<location id="8E87B0A2FFC56940FFE7FA2FFAE7FFC6" LSID="urn:lsid:plazi:treatment:03F1576FFFCA6942FBB5FC26FD1CFD8E:8E87B0A2FFC56940FFE7FA2FFAE7FFC6" box="[1221,1399,1411,1437]" country="Japan" latitude="28.35" longLatPrecision="925" longitude="129.46584" municipality="Amami" name="Amami Oshima" pageId="8" pageNumber="479">Amami Oshima</location>
Is- land,
<collectingMunicipality id="6B837C03FFC56940FBF6FA0BFEBBFF9A" box="[212,299,1447,1473]" pageId="8" pageNumber="479">Amami</collectingMunicipality>
,
<geoCoordinate id="EE6C80BEFFC56940FA14FA0BFE03FF9A" box="[310,403,1447,1473]" degrees="28" direction="north" minutes="21" orientation="latitude" pageId="8" pageNumber="479" precision="925" value="28.35">28°21'N</geoCoordinate>
<geoCoordinate id="EE6C80BEFFC56940FAB8FA0AFDB0FF9A" box="[410,544,1446,1473]" degrees="129" direction="east" minutes="27" orientation="longitude" pageId="8" pageNumber="479" precision="15" seconds="57" value="129.46584">129°27'57E</geoCoordinate>
,
<collectedFrom id="3844A584FFC56940F90EFA0BFC9CFF9A" box="[556,780,1447,1473]" pageId="8" pageNumber="479">
on
<taxonomicName id="4C589DFAFFC56940F96DFA0BFC9CFF9A" box="[591,780,1447,1473]" class="Pinopsida" family="Pinaceae" genus="Pinus" kingdom="Plantae" order="Pinales" pageId="8" pageNumber="479" phylum="Tracheophyta" rank="species" species="luchuensis">
<emphasis id="B92C3A6BFFC56940F96DFA0BFC9CFF9A" box="[591,780,1447,1473]" italics="true" pageId="8" pageNumber="479">Pinus luchuensis</emphasis>
</taxonomicName>
</collectedFrom>
,
<date id="FFE6C0B9FFC56940F839FA0BFC3AFF9A" box="[795,938,1447,1473]" pageId="8" pageNumber="479" value="2021-03-26">
<collectingDate id="EFA23951FFC56940F839FA0BFC3AFF9A" box="[795,938,1447,1473]" pageId="8" pageNumber="479" value="2021-03-26">26. III. 2021</collectingDate>
</date>
,
<collectorName id="26AD83AFFFC56940F897FA0BFB82FF9A" box="[949,1042,1447,1473]" pageId="8" pageNumber="479">H. Hara</collectorName>
</materialsCitation>
;
<materialsCitation id="3B30EC24FFC56940FF3FFA0BFCB4FFBE" collectingDate="1998-04-09" collectorName="M. Kimura" country="Japan" location="Tokunoshima Island" municipality="Mikyo" pageId="8" pageNumber="479" specimenCount="1" specimenCount-female="1">
<specimenCount id="9D5E2DF0FFC56940FF3FFA0BFBD0FF9A" box="[1053,1088,1447,1473]" pageId="8" pageNumber="479" type="female">1♀</specimenCount>
(
<figureCitation id="1363FAFCFFC56940FF6DFA0BFB77FF9A" box="[1103,1255,1447,1473]" captionStart="FIGURES 2" captionStartId="4.[151,264,1746,1770]" captionTargetBox="[166,1435,215,1720]" captionTargetId="figure-19@4.[151,1435,181,1723]" captionTargetPageId="4" captionText="FIGURES 2AH. AF, IK, Gilpinia amamiana: A, B, holotype, female; C, D, female, Tokunoshima Isl.; E, F, male, Amami Oshima Isl. G, H, L, Gilpinia okinawa, holotype, female. A, C, E, G, Dorsal view; B, D, F, H, ventrolateral view; IL, head, frontal view." figureDoi="http://doi.org/10.5281/zenodo.5057391" httpUri="https://zenodo.org/record/5057391/files/figure.png" pageId="8" pageNumber="479">Figs 2C, D, J</figureCitation>
,
<figureCitation id="1363FAFCFFC56940FFD1FA0BFACAFF9A" box="[1267,1370,1447,1473]" captionStart="FIGURES 3" captionStartId="9.[151,264,1786,1810]" captionTargetBox="[151,1435,181,1762]" captionTargetId="figure-19@9.[151,1435,181,1762]" captionTargetPageId="9" captionText="FIGURES 3AX. AH, Head, dorsal view; IP, dorsal part of head, anterolateral view; Q, head, lateral view; RX, antenna, inner view (RT, V, W) or outer view (U, X). AD, IL, QU, Gilpinia hakonensis: A, I, Q, R, lectotype of Lophyrus hakonensis, female; B, J, S, holotype of G. hakonensis var. laticincta, female; C, K, T, paratype of Diprion fukudai, female; D, L, U, male, Honshu. EG, MO, V, W, Gilpinia amamiana: E, M, V, holotype, female; F, N, W, female, Tokunoshima Isl.; G, O, male, Amami Oshima Isl. H, P, X, Gilpinia okinawa, holotype, female. T, Reversed." figureDoi="http://doi.org/10.5281/zenodo.5057395" httpUri="https://zenodo.org/record/5057395/files/figure.png" pageId="8" pageNumber="479">3F, N, W</figureCitation>
,
<figureCitation id="1363FAFCFFC56940FE47FA0AFA07FF9A" box="[1381,1431,1446,1473]" captionStart="FIGURES 5" captionStartId="11.[151,264,1868,1892]" captionTargetBox="[204,1384,181,1835]" captionTargetId="figure-19@11.[204,1384,181,1843]" captionTargetPageId="11" captionText="FIGURES 5AU.AP, Valvula 3, dorsal and posterior views; QU, ovipositor or lance, dorsal view (Q, S, T) or lateral view (R, U). AF, Q, R, Gilpinia hakonensis: AB, lectotype of Lophyrus hakonensis; C, D, holotype of G. hakonensis var. laticincta; E, F, P, paratype of Diprion fukudai; Q, R, Honshu. GJ, S, Gilpinia amamiana: G, H, holotype; I, J, Tokunoshima Isl.; S, Amami Oshima Isl. K, L, T, U, Gilpinia okinawa, holotype. M, N, Gilpinia frutetorum, Sweden. O, P, Gilpinia laricis, Germany. R, Reversed." figureDoi="http://doi.org/10.5281/zenodo.5057397" httpUri="https://zenodo.org/record/5057397/files/figure.png" pageId="8" pageNumber="479">5I, J</figureCitation>
,
<figureCitation id="1363FAFCFFC56940FBB5FA67FF2DFFBD" box="[151,189,1483,1510]" captionStart="FIGURES 6" captionStartId="12.[151,264,1869,1893]" captionTargetBox="[154,1435,181,1833]" captionTargetId="figure-19@12.[151,1435,181,1845]" captionTargetPageId="12" captionText="FIGURES 6AJ. Lancet. AF, Gilpinia hakonensis: A, lectotype of Lophyrus hakonensis; B, holotype of G. hakonensis var. laticincta; C, Kyushu, D, Honshu, E, paratype of Diprion fukudai. FH, Gilpinia amamiana: F, holotype; G, Amami Oshima Isl.; H, Tokunoshima Isl. I, J, Gilpinia okinawa: I, holotype; J, Okinawa Isl. 1, 2, 5, first (most basal), second and fifth annuli. DF, H, Reversed." figureDoi="http://doi.org/10.5281/zenodo.5057401" httpUri="https://zenodo.org/record/5057401/files/figure.png" pageId="8" pageNumber="479">6H</figureCitation>
),
<location id="8E87B0A2FFC56940FBF2FA67FE26FFBE" LSID="urn:lsid:plazi:treatment:03F1576FFFCA6942FBB5FC26FD1CFD8E:8E87B0A2FFC56940FBF2FA67FE26FFBE" box="[208,438,1483,1509]" country="Japan" municipality="Mikyo" name="Tokunoshima Island" pageId="8" pageNumber="479">Tokunoshima Island</location>
,
<collectingMunicipality id="6B837C03FFC56940FAE0FA67FD80FFBE" box="[450,528,1483,1509]" pageId="8" pageNumber="479">Mikyo</collectingMunicipality>
,
<date id="FFE6C0B9FFC56940F938FA67FD09FFBE" box="[538,665,1483,1509]" pageId="8" pageNumber="479" value="1998-04-09">
<collectingDate id="EFA23951FFC56940F938FA67FD09FFBE" box="[538,665,1483,1509]" pageId="8" pageNumber="479" value="1998-04-09">9. IV. 1998</collectingDate>
</date>
,
<collectorName id="26AD83AFFFC56940F986FA67FCB4FFBE" box="[676,804,1483,1509]" pageId="8" pageNumber="479">M. Kimura</collectorName>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C342B5F2FFC56940FBE5FA42FE9CFC2E" pageId="8" pageNumber="479" type="distribution">
<paragraph id="8BE7E679FFC56940FBE5FA42FB64FC51" blockId="8.[151,1437,150,1905]" box="[199,1268,1518,1546]" pageId="8" pageNumber="479">
<emphasis id="B92C3A6BFFC56940FBE5FA42FEC9FC53" bold="true" box="[199,345,1518,1544]" pageId="8" pageNumber="479">Distribution</emphasis>
.
<collectingCountry id="F34FA6E9FFC56940FA45FA43FE3BFC52" box="[359,427,1519,1545]" name="Japan" pageId="8" pageNumber="479">Japan</collectingCountry>
: Amami Oshima Island (
<bibRefCitation id="EFC99B88FFC56940F9E2FA43FCCEFC52" author="Okutani, T." box="[704,862,1518,1545]" pageId="8" pageNumber="479" pagination="73 - 84" refId="ref14145" refString="Okutani, T. (1965) Sawflies and horntails from the Ryukyus. Kontyu, 33, 73 - 84." type="journal article" year="1965">Okutani 1965</bibRefCitation>
), Tokunoshima Island (new record).
</paragraph>
<paragraph id="8BE7E679FFC56940FBE5F9BEFE9CFC2E" blockId="8.[151,1437,150,1905]" pageId="8" pageNumber="479">
<emphasis id="B92C3A6BFFC56940FBE5F9BEFEC2FC77" bold="true" box="[199,338,1554,1580]" pageId="8" pageNumber="479">Life history</emphasis>
. The adults were collected in March, April, June and September. This sawfly probably has several generations a year. Hara collected
<specimenCount id="9D5E2DF0FFC56940F93DF99BFD02FC0A" box="[543,658,1591,1617]" pageId="8" pageNumber="479" type="male">two males</specimenCount>
on a tree of
<taxonomicName id="4C589DFAFFC56940F807F99BFBF1FC0A" authority="Mayr. The" authorityName="Mayr. The" box="[805,1121,1591,1617]" class="Pinopsida" family="Pinaceae" genus="Pinus" kingdom="Plantae" order="Pinales" pageId="8" pageNumber="479" phylum="Tracheophyta" rank="species" species="luchuensis">
<emphasis id="B92C3A6BFFC56940F807F99BFC73FC0A" box="[805,995,1591,1617]" italics="true" pageId="8" pageNumber="479">Pinus luchuensis</emphasis>
Mayr. The
</taxonomicName>
tree species is probably the host plant.
</paragraph>
</subSubSection>
<subSubSection id="C342B5F2FFC56942FBE5F9D3FD1CFD8E" lastPageId="10" lastPageNumber="481" pageId="8" pageNumber="479" type="discussion">
<paragraph id="8BE7E679FFC56940FBE5F9D3FF61FD2A" blockId="8.[151,1437,150,1905]" pageId="8" pageNumber="479">
<emphasis id="B92C3A6BFFC56940FBE5F9D3FEA4FCC2" bold="true" box="[199,308,1663,1689]" pageId="8" pageNumber="479">Remarks</emphasis>
.
<taxonomicName id="4C589DFAFFC56940FA60F9D3FD8BFCC3" authorityName="Okutani" authorityYear="1965" box="[322,539,1663,1689]" class="Insecta" family="Diprionidae" genus="Gilpinia" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="479" phylum="Arthropoda" rank="species" species="amamiana">
<emphasis id="B92C3A6BFFC56940FA60F9D3FD8BFCC3" box="[322,539,1663,1689]" italics="true" pageId="8" pageNumber="479">Gilpinia amamiana</emphasis>
</taxonomicName>
is distinguished from other congeners as stated in the section of the
<taxonomicName id="4C589DFAFFC56940FE30F9D3FF25FCE7" authorityName=": Benson" authorityYear="1939" class="Insecta" family="Diprionidae" genus="Gilpinia" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="479" phylum="Arthropoda" rank="species" species="hakonensis">
<emphasis id="B92C3A6BFFC56940FE30F9D3FF25FCE7" italics="true" pageId="8" pageNumber="479">G. hakonensis</emphasis>
</taxonomicName>
and similar species and the key above.
<bibRefCitation id="EFC99B88FFC56940F95DF90FFCBCFCE6" author="Okutani, T." box="[639,812,1698,1725]" pageId="8" pageNumber="479" pagination="73 - 84" refId="ref14145" refString="Okutani, T. (1965) Sawflies and horntails from the Ryukyus. Kontyu, 33, 73 - 84." type="journal article" year="1965">Okutani (1965)</bibRefCitation>
wrote about the female
<typeStatus id="54E358DBFFC56940FF66F90FFB36FCE6" box="[1092,1190,1699,1725]" pageId="8" pageNumber="479" type="holotype">holotype</typeStatus>
of
<taxonomicName id="4C589DFAFFC56940FFEDF90FFAF9FCE7" authorityName="Okutani" authorityYear="1965" box="[1231,1385,1699,1725]" class="Insecta" family="Diprionidae" genus="Gilpinia" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="479" phylum="Arthropoda" rank="species" species="amamiana">
<emphasis id="B92C3A6BFFC56940FFEDF90FFAF9FCE7" box="[1231,1385,1699,1725]" italics="true" pageId="8" pageNumber="479">G. amamiana</emphasis>
</taxonomicName>
that “this species is closely allied to
<taxonomicName id="4C589DFAFFC56940FAD5F96BFCE4FCBA" authority="(Matsumura, 1912)" baseAuthorityName="Matsumura" baseAuthorityYear="1912" box="[503,884,1735,1761]" class="Insecta" family="Diprionidae" genus="Gilpinia" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="479" phylum="Arthropoda" rank="species" species="hakonensis">
<emphasis id="B92C3A6BFFC56940FAD5F96BFD04FCBA" box="[503,660,1735,1761]" italics="true" pageId="8" pageNumber="479">G. hakonensis</emphasis>
(
<bibRefCitation id="EFC99B88FFC56940F980F96BFCFCFCBA" author="Matsumura, S." box="[674,876,1735,1761]" pageId="8" pageNumber="479" refId="ref13717" refString="Matsumura, S. (1912) Thousand Insects of Japan. Supplement IV. Keiseisha-shoten, Tokyo, 247 pp., 55 pls. [in Japanese and English]" type="book" year="1912">Matsumura, 1912</bibRefCitation>
)
</taxonomicName>
, but easily distinguished by the larger metascutellum, truncate clypeus, hardly defined postocellar area, and yellow legs”. Actually,
<taxonomicName id="4C589DFAFFC56940FF08F947FB51FD5F" authorityName="Okutani" authorityYear="1965" box="[1066,1217,1771,1797]" class="Insecta" family="Diprionidae" genus="Gilpinia" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="479" phylum="Arthropoda" rank="species" species="amamiana">
<emphasis id="B92C3A6BFFC56940FF08F947FB51FD5F" box="[1066,1217,1771,1797]" italics="true" pageId="8" pageNumber="479">G. amamiana</emphasis>
</taxonomicName>
has the metascutellum not larger than that of the latter (compare Fig. 4I with Figs 4A, C, E), the clypeus not different from that of the latter, and the postocellar area with a distinct anterior furrow and a distinct, vestigial or indistinct lateral furrow (
<figureCitation id="1363FAFCFFC56940FE49F89FFF74FD2A" captionStart="FIGURES 3" captionStartId="9.[151,264,1786,1810]" captionTargetBox="[151,1435,181,1762]" captionTargetId="figure-19@9.[151,1435,181,1762]" captionTargetPageId="9" captionText="FIGURES 3AX. AH, Head, dorsal view; IP, dorsal part of head, anterolateral view; Q, head, lateral view; RX, antenna, inner view (RT, V, W) or outer view (U, X). AD, IL, QU, Gilpinia hakonensis: A, I, Q, R, lectotype of Lophyrus hakonensis, female; B, J, S, holotype of G. hakonensis var. laticincta, female; C, K, T, paratype of Diprion fukudai, female; D, L, U, male, Honshu. EG, MO, V, W, Gilpinia amamiana: E, M, V, holotype, female; F, N, W, female, Tokunoshima Isl.; G, O, male, Amami Oshima Isl. H, P, X, Gilpinia okinawa, holotype, female. T, Reversed." figureDoi="http://doi.org/10.5281/zenodo.5057395" httpUri="https://zenodo.org/record/5057395/files/figure.png" pageId="8" pageNumber="479">Figs 3M, N</figureCitation>
).
</paragraph>
<caption id="DF27B6F1FFC46941FBB5F956FC3EFDF9" ID-DOI="http://doi.org/10.5281/zenodo.5057395" ID-Zenodo-Dep="5057395" httpUri="https://zenodo.org/record/5057395/files/figure.png" pageId="9" pageNumber="480" startId="9.[151,264,1786,1810]" targetBox="[151,1435,181,1762]" targetPageId="9">
<paragraph id="8BE7E679FFC46941FBB5F956FC3EFDF9" blockId="9.[151,1436,1786,1954]" pageId="9" pageNumber="480">
<emphasis id="B92C3A6BFFC46941FBB5F956FEC4FD49" bold="true" box="[151,340,1786,1810]" pageId="9" pageNumber="480">FIGURES 3AX.</emphasis>
AH, Head, dorsal view; IP, dorsal part of head, anterolateral view; Q, head, lateral view; RX, antenna, inner view (RT, V, W) or outer view (U, X). AD, IL, QU,
<taxonomicName id="4C589DFAFFC46941F802F8B2FC7DFD6D" baseAuthorityName="Matsumura" baseAuthorityYear="1912" box="[800,1005,1822,1846]" class="Insecta" family="Diprionidae" genus="Gilpinia" kingdom="Animalia" order="Hymenoptera" pageId="9" pageNumber="480" phylum="Arthropoda" rank="species" species="hakonensis">
<emphasis id="B92C3A6BFFC46941F802F8B2FC7DFD6D" box="[800,1005,1822,1846]" italics="true" pageId="9" pageNumber="480">Gilpinia hakonensis</emphasis>
</taxonomicName>
: A, I, Q, R, lectotype of
<taxonomicName id="4C589DFAFFC46941FFCDF8B2FF5AFD01" authorityName="Matsumura" authorityYear="1912" class="Insecta" family="Miridae" genus="Lophyrus" higherTaxonomySource="CoL" kingdom="Animalia" order="Hemiptera" pageId="9" pageNumber="480" phylum="Arthropoda" rank="species" species="hakonensis">
<emphasis id="B92C3A6BFFC46941FFCDF8B2FF5AFD01" italics="true" pageId="9" pageNumber="480">Lophyrus hakonensis</emphasis>
</taxonomicName>
, female; B, J, S, holotype of
<taxonomicName id="4C589DFAFFC46941FAD6F8EEFC86FD01" box="[500,790,1858,1882]" class="Insecta" family="Miridae" genus="Lophyrus" higherTaxonomySource="CoL" kingdom="Animalia" order="Hemiptera" pageId="9" pageNumber="480" phylum="Arthropoda" rank="variety" species="hakonensis" variety="laticincta">
<emphasis id="B92C3A6BFFC46941FAD6F8EEFD14FD01" box="[500,644,1858,1882]" italics="true" pageId="9" pageNumber="480">G. hakonensis</emphasis>
var.
<emphasis id="B92C3A6BFFC46941F994F8EEFC86FD01" box="[694,790,1858,1882]" italics="true" pageId="9" pageNumber="480">laticincta</emphasis>
</taxonomicName>
, female; C, K, T, paratype of
<taxonomicName id="4C589DFAFFC46941FF64F8EFFB79FD01" baseAuthorityName="Togashi" baseAuthorityYear="1964" box="[1094,1257,1858,1882]" class="Insecta" family="Diprionidae" genus="Diprion" kingdom="Animalia" order="Hymenoptera" pageId="9" pageNumber="480" phylum="Arthropoda" rank="species" species="fukudai">
<emphasis id="B92C3A6BFFC46941FF64F8EFFB79FD01" box="[1094,1257,1858,1882]" italics="true" pageId="9" pageNumber="480">Diprion fukudai</emphasis>
</taxonomicName>
, female; D, L, U, male, Honshu. EG, MO, V, W,
<taxonomicName id="4C589DFAFFC46941FAC4F8CAFD3BFD25" authorityName="Okutani" authorityYear="1965" box="[486,683,1894,1918]" class="Insecta" family="Diprionidae" genus="Gilpinia" kingdom="Animalia" order="Hymenoptera" pageId="9" pageNumber="480" phylum="Arthropoda" rank="species" species="amamiana">
<emphasis id="B92C3A6BFFC46941FAC4F8CAFD3BFD25" box="[486,683,1894,1918]" italics="true" pageId="9" pageNumber="480">Gilpinia amamiana</emphasis>
</taxonomicName>
: E, M, V, holotype, female; F, N, W, female, Tokunoshima Isl.; G, O, male, Amami Oshima Isl. H, P, X,
<taxonomicName id="4C589DFAFFC46941FA99F826FDFBFDF9" authorityName="Hara &amp; Smith &amp; Shinohara" authorityYear="2021" box="[443,619,1930,1954]" class="Insecta" family="Diprionidae" genus="Gilpinia" kingdom="Animalia" order="Hymenoptera" pageId="9" pageNumber="480" phylum="Arthropoda" rank="species" species="okinawa">
<emphasis id="B92C3A6BFFC46941FA99F826FDFBFDF9" box="[443,619,1930,1954]" italics="true" pageId="9" pageNumber="480">Gilpinia okinawa</emphasis>
</taxonomicName>
, holotype, female. T, Reversed.
</paragraph>
</caption>
<paragraph id="8BE7E679FFC76942FBB5F881FD1CFD8E" blockId="10.[151,1436,1837,2005]" pageId="10" pageNumber="481">
<emphasis id="B92C3A6BFFC76942FBB5F881FEC4FD1E" bold="true" box="[151,340,1837,1861]" pageId="10" pageNumber="481">FIGURES 4AR.</emphasis>
A, C, E, G, I, K, M, Posteromedial part of thorax, dorsal view; B, D, F, H, J, L, N, mesepisternum, lateral view; OR, female seventh abdominal sternum, ventrolateral view. AH, O, P,
<taxonomicName id="4C589DFAFFC76942F889F8FDFBE5FD32" baseAuthorityName="Matsumura" baseAuthorityYear="1912" box="[939,1141,1873,1897]" class="Insecta" family="Diprionidae" genus="Gilpinia" kingdom="Animalia" order="Hymenoptera" pageId="10" pageNumber="481" phylum="Arthropoda" rank="species" species="hakonensis">
<emphasis id="B92C3A6BFFC76942F889F8FDFBE5FD32" box="[939,1141,1873,1897]" italics="true" pageId="10" pageNumber="481">Gilpinia hakonensis</emphasis>
</taxonomicName>
: A, B, O,
<typeStatus id="54E358DBFFC76942FFFAF8FDFAA5FD32" box="[1240,1333,1873,1897]" pageId="10" pageNumber="481" type="lectotype">lectotype</typeStatus>
of
<taxonomicName id="4C589DFAFFC76942FE76F8FDFEBEFDD6" authorityName="Matsumura" authorityYear="1912" class="Insecta" family="Miridae" genus="Lophyrus" higherTaxonomySource="CoL" kingdom="Animalia" order="Hemiptera" pageId="10" pageNumber="481" phylum="Arthropoda" rank="species" species="hakonensis">
<emphasis id="B92C3A6BFFC76942FE76F8FDFEBEFDD6" italics="true" pageId="10" pageNumber="481">Lophyrus hakonensis</emphasis>
</taxonomicName>
, female; C, D,
<typeStatus id="54E358DBFFC76942FAEAF8D9FDB1FDD6" box="[456,545,1909,1933]" pageId="10" pageNumber="481" type="holotype">holotype</typeStatus>
of
<taxonomicName id="4C589DFAFFC76942F960F8D9FCF2FDD6" box="[578,866,1909,1933]" class="Insecta" family="Miridae" genus="Lophyrus" higherTaxonomySource="CoL" kingdom="Animalia" order="Hemiptera" pageId="10" pageNumber="481" phylum="Arthropoda" rank="variety" species="hakonensis" variety="laticincta">
<emphasis id="B92C3A6BFFC76942F960F8D9FD41FDD6" box="[578,721,1909,1933]" italics="true" pageId="10" pageNumber="481">G. hakonensis</emphasis>
var.
<emphasis id="B92C3A6BFFC76942F820F8D9FCF2FDD6" box="[770,866,1909,1933]" italics="true" pageId="10" pageNumber="481">laticincta</emphasis>
</taxonomicName>
, female; E, F, P,
<typeStatus id="54E358DBFFC76942FF29F8DAFBF2FDD5" box="[1035,1122,1910,1934]" pageId="10" pageNumber="481" type="paratype">paratype</typeStatus>
of
<taxonomicName id="4C589DFAFFC76942FFA1F8DAFAB5FDD6" baseAuthorityName="Togashi" baseAuthorityYear="1964" box="[1155,1317,1909,1933]" class="Insecta" family="Diprionidae" genus="Diprion" kingdom="Animalia" order="Hymenoptera" pageId="10" pageNumber="481" phylum="Arthropoda" rank="species" species="fukudai">
<emphasis id="B92C3A6BFFC76942FFA1F8DAFAB5FDD6" box="[1155,1317,1909,1933]" italics="true" pageId="10" pageNumber="481">Diprion fukudai</emphasis>
</taxonomicName>
, female; G, H, male, Honshu. IL, Q,
<taxonomicName id="4C589DFAFFC76942FAB8F835FDCFFDEA" authorityName="Okutani" authorityYear="1965" box="[410,607,1945,1969]" class="Insecta" family="Diprionidae" genus="Gilpinia" kingdom="Animalia" order="Hymenoptera" pageId="10" pageNumber="481" phylum="Arthropoda" rank="species" species="amamiana">
<emphasis id="B92C3A6BFFC76942FAB8F835FDCFFDEA" box="[410,607,1945,1969]" italics="true" pageId="10" pageNumber="481">Gilpinia amamiana</emphasis>
</taxonomicName>
: I, J,
<typeStatus id="54E358DBFFC76942F9B6F835FD7FFDEA" box="[660,751,1945,1969]" pageId="10" pageNumber="481" type="holotype">holotype</typeStatus>
, female; K, L, male, Amami Oshima Isl.; Q, female, Amami Oshima Isl. M, N, R,
<taxonomicName id="4C589DFAFFC76942FA3FF811FE5EFD8E" authorityName="Hara &amp; Smith &amp; Shinohara" authorityYear="2021" box="[285,462,1981,2005]" class="Insecta" family="Diprionidae" genus="Gilpinia" kingdom="Animalia" order="Hymenoptera" pageId="10" pageNumber="481" phylum="Arthropoda" rank="species" species="okinawa">
<emphasis id="B92C3A6BFFC76942FA3FF811FE5EFD8E" box="[285,462,1981,2005]" italics="true" pageId="10" pageNumber="481">Gilpinia okinawa</emphasis>
</taxonomicName>
,
<typeStatus id="54E358DBFFC76942FAF8F811FDA5FD8E" box="[474,565,1981,2005]" pageId="10" pageNumber="481" type="holotype">holotype</typeStatus>
, female.
</paragraph>
</subSubSection>
</treatment>
</document>