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<mods:title id="5EC280CAC548A1BEC6296CD7A18E75D6">First records of some coral reef-associated brachyuran crabs from the Nicobar archipelago, India</mods:title>
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<mods:namePart id="3158D1925D847F01A31063E1F0B08337">Padate, Vinay P.</mods:namePart>
<mods:affiliation id="5433979A71898F8A8AE835D6214369B1">Centre for Marine Living Resources &amp; Ecology, Ministry of Earth Sciences, Government of India, Atal Bhavan, LNG Terminus Road, Puthuvype, Kochi, 682508, India.</mods:affiliation>
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<mods:namePart id="9B61F21017F5876D66A361B9EE4CD3ED">Cubelio, Sherine Sonia</mods:namePart>
<mods:affiliation id="6ED5DCEF738D8A21D1A5A188F5A1AC62">Centre for Marine Living Resources &amp; Ecology, Ministry of Earth Sciences, Government of India, Atal Bhavan, LNG Terminus Road, Puthuvype, Kochi, 682508, India.</mods:affiliation>
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<mods:namePart id="1FC4C48C25515C2A702C6D8DE29688C3">Takeda, Masatsune</mods:namePart>
<mods:affiliation id="C0C368DF009CFA6333ED91C26C5314B6">Department of Zoology, National Museum of Nature and Science, Tokyo. 4 - 1 - 1 Amakubo, Tsukuba, Ibaraki 305 - 0005, Japan.</mods:affiliation>
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<treatment id="03EA87917F2CFF81FF79FC42B5A1D4EC" LSID="urn:lsid:plazi:treatment:03EA87917F2CFF81FF79FC42B5A1D4EC" httpUri="http://treatment.plazi.org/id/03EA87917F2CFF81FF79FC42B5A1D4EC" lastPageId="9" lastPageNumber="485" pageId="7" pageNumber="483">
<subSubSection id="C359650C7F2CFF8FFF79FC42B7FFD498" box="[151,694,931,958]" pageId="7" pageNumber="483" type="nomenclature">
<paragraph id="8BFC36877F2CFF8FFF79FC42B7FFD498" blockId="7.[151,694,931,994]" box="[151,694,931,958]" pageId="7" pageNumber="483">
<heading id="D0B481EB7F2CFF8FFF79FC42B7FFD498" bold="true" box="[151,694,931,958]" fontSize="11" level="1" pageId="7" pageNumber="483" reason="1">
<taxonomicName id="4C434D047F2CFF8FFF79FC42B7FFD498" ID-CoL="6WFQ8" authority="(Calman, 1900)" baseAuthorityName="Calman" baseAuthorityYear="1900" box="[151,694,931,958]" class="Malacostraca" family="Parthenopidae" genus="Pseudolambrus" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="483" phylum="Arthropoda" rank="species" species="confragosus">
<emphasis id="B937EA957F2CFF8FFF79FC42B7FFD498" bold="true" box="[151,694,931,958]" pageId="7" pageNumber="483">
<emphasis id="B937EA957F2CFF8FFF79FC42B4A7D498" bold="true" box="[151,494,931,958]" italics="true" pageId="7" pageNumber="483">Pseudolambrus confragosus</emphasis>
(
<bibRefCitation id="EFD24B767F2CFF8FFE10FC42B7E4D498" author="Calman, W. T." box="[510,685,931,957]" pageId="7" pageNumber="483" pagination="1 - 50" refId="ref10828" refString="Calman, W. T. (1900) On a collection of Brachyura from Torres Straits. Transactions of the Linnean Society of London, Series 2, Zoology, 8, 1 - 50, pls. 1 - 3. https: // doi. org / 10.1111 / j. 1096 - 3642.1900. tb 00307. x" type="journal article" year="1900">Calman, 1900</bibRefCitation>
)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C359650C7F2CFF8FFF79FC26B4F2D4C7" box="[151,443,967,994]" pageId="7" pageNumber="483" type="description">
<paragraph id="8BFC36877F2CFF8FFF79FC26B4F2D4C7" blockId="7.[151,694,931,994]" box="[151,443,967,994]" pageId="7" pageNumber="483">
(
<figureCitation id="13782A027F2CFF8FFF71FC26B44BD4C7" box="[159,258,967,994]" captionStart="FIGURE 2" captionStartId="3.[152,255,1683,1707]" captionTargetBox="[177,1410,181,1659]" captionTargetId="figure-37@3.[177,1410,181,1659]" captionTargetPageId="3" captionText="FIGURE 2. Dorsal habitus photographs (AC, EG: fresh specimens; D: ethanol-preserved specimen): (A) Singhaplax ockelmanni (Serène, 1971), male (IO/SS/BRC/00360, 4.8 × 2.5 mm); (B) Urnalana angulata (Rathbun, 1911), undetermined sex (IO/SS/BRC/00184, 6.9 × 7.0 mm); (C) Pseudolambrus confragosus (Calman, 1900), male (IO/SS/BRC/00323, 22.2 × 18.0 mm; (D) Cavoportunus dubius (Laurie, 1906), male (IO/SS/BRC/00191, 26.9 × 20.4 mm); (E) Cyrtocarcinus truncatus (Rathbun, 1906), male (IO/SS/BRC/00193, 8.6 × 6.1 mm); (F) Visayax osteodictyon Mendoza &amp; Ng, 2008, male (IO/SS/ BRC/00194, 5.2 × 3.6 mm); (G) Xanthias joanneae Mendoza, 2013, male (IO/SS/BRC/00197, 6.4 × 4.2 mm). Scale bars: AB, E, G, 2 mm, C, 10 mm, D, F, 5 mm." figureDoi="http://doi.org/10.5281/zenodo.13210274" httpUri="https://zenodo.org/record/13210274/files/figure.png" pageId="7" pageNumber="483">Figs. 2C</figureCitation>
,
<figureCitation id="13782A027F2CFF8FFEE3FC26B41DD4C7" box="[269,340,967,994]" captionStart="FIGURE 5" captionStartId="8.[152,255,1812,1836]" captionTargetBox="[151,1436,181,1788]" captionTargetId="figure-18@8.[151,1436,181,1788]" captionTargetPageId="8" captionText="FIGURE 5. Pseudolambrus confragosus (Calman, 1900), male (IO/SS/BRC/00323, 22.2 × 18.0 mm),Andaman Sea, India: (A) carapace, dorsal view; (B) carapace, lateral view; (C) carapace, frontal view; (D) right mxp3, ventral view; (E) right chela and carpus, subdorsal view; (F) right P2, dorsal view; (G) male pleon and telson, ventral view. Scale bars: AG, 2 mm." figureDoi="http://doi.org/10.5281/zenodo.13210284" httpUri="https://zenodo.org/record/13210284/files/figure.png" pageId="7" pageNumber="483">5AG</figureCitation>
,
<figureCitation id="13782A027F2CFF8FFE8EFC29B4FBD4C7" box="[352,434,968,994]" captionStart="FIGURE 10" captionStartId="17.[151,250,1667,1691]" captionTargetBox="[304,1294,191,1633]" captionTargetId="figure-37@17.[292,1295,181,1645]" captionTargetPageId="17" captionText="FIGURE 10. First and second gonopods: Singhaplax ockelmanni (Serène, 1971), male (IO/SS/BRC/00360, 4.8 × 2.5 mm): (A) entire G1, pleonal view; (B) entire G1, sternal view; (C) entire G2, pleonal view. Pseudolambrus confragosus (Calman, 1900), male (IO/SS/BRC/00323, 22.2 × 18.0 mm): (D) entire G1, pleonal view; (E) G1 distal portion, pleonal view; (F) G1 distal portion, sternal view; (G) entire G1, sternal view; (H) entire G2, pleonal view. Cavoportunus dubius (Laurie, 1906), male (IO/SS/BRC/00191, 26.9 × 20.4 mm): (I) entire G1, pleonal view; (J) entire G1, lateral view; (K) entire G1, mesial view; (L) entire G2, pleonal view. Cyrtocarcinus truncatus (Rathbun, 1906), male (IO/SS/BRC/00193, 8.6 × 6.1 mm): (M) entire G1, pleonal view; (N) entire G1, sternal view; (O) entire G2, sternal view. Visayax osteodictyon Mendoza &amp; Ng, 2008, male (IO/ SS/BRC/00194, 5.2 × 3.6 mm): (P) entire G1, pleonal view; (Q) G1 distal portion, pleonal view; (R) entire G1, sternal view. Xanthias joanneae Mendoza, 2013, male (IO/SS/BRC/00197, 6.4 × 4.2 mm). (S) entire G1, pleonal view; (T) G1 distal portion, pleonal view; (U) G1 distal portion, sternal view; (V) entire G1, sternal view. Scale bars: AC, PV, 0.2 mm, D, GN, 1 mm, E, F, O, 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13210300" httpUri="https://zenodo.org/record/13210300/files/figure.png" pageId="7" pageNumber="483">10DH</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C359650C7F2CFF8FFF79FBEDB069D398" pageId="7" pageNumber="483" type="reference_group">
<paragraph id="8BFC36877F2CFF8FFF79FBEDB72FD366" blockId="7.[151,1436,1036,1213]" pageId="7" pageNumber="483">
<treatmentCitationGroup id="AB5311A97F2CFF8FFF79FBEDB72FD366" pageId="7" pageNumber="483">
<treatmentCitation id="0AE210967F2CFF8FFF79FBEDB7B7D301" author="Calman, W. T." box="[151,766,1036,1060]" page="42" pageId="7" pageNumber="483" year="1900">
<taxonomicName id="4C434D047F2CFF8FFF79FBEDB7B7D301" ID-CoL="3RYTC" authority="Calman, 1900: 42" authorityName="Calman" authorityPageNumber="42" authorityYear="1900" box="[151,766,1036,1060]" class="Malacostraca" family="Parthenopidae" genus="Lambrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="483" phylum="Arthropoda" rank="species" species="confragosus" subGenus="Parthenolambrus">
<emphasis id="B937EA957F2CFF8FFF79FBEDB5B8D301" box="[151,241,1036,1060]" italics="true" pageId="7" pageNumber="483">Lambrus</emphasis>
(
<emphasis id="B937EA957F2CFF8FFEEFFBEDB4FBD301" box="[257,434,1036,1060]" italics="true" pageId="7" pageNumber="483">Parthenolambrus</emphasis>
)
<emphasis id="B937EA957F2CFF8FFE2CFBEDB777D301" box="[450,574,1036,1060]" italics="true" pageId="7" pageNumber="483">confragosus</emphasis>
<bibRefCitation id="EFD24B767F2CFF8FFDABFBEDB7B7D301" author="Calman, W. T." box="[581,766,1036,1060]" pageId="7" pageNumber="483" pagination="1 - 50" refId="ref10828" refString="Calman, W. T. (1900) On a collection of Brachyura from Torres Straits. Transactions of the Linnean Society of London, Series 2, Zoology, 8, 1 - 50, pls. 1 - 3. https: // doi. org / 10.1111 / j. 1096 - 3642.1900. tb 00307. x" type="journal article" year="1900">Calman, 1900: 42</bibRefCitation>
</taxonomicName>
</treatmentCitation>
, pl. 3 figs. 27, 28 (
<typeStatus id="54F888257F2CFF8FFC2FFBEDB6A7D301" box="[961,1006,1036,1060]" pageId="7" pageNumber="483">type</typeStatus>
locality: Torres Strait,
<collectingCountry id="F35476177F2CFF8FFB37FBEDB071D301" box="[1241,1336,1036,1060]" name="Australia" pageId="7" pageNumber="483">Australia</collectingCountry>
).—
<treatmentCitation id="0AE210967F2CFF8FFAB3FBEDB469D366" author="Flipse, H. J." page="48" pageId="7" pageNumber="483" year="1930">
<bibRefCitation id="EFD24B767F2CFF8FFAB3FBEDB469D366" author="Flipse, H. J." pageId="7" pageNumber="483" pagination="1 - 96" refId="ref11616" refString="Flipse, H. J. (1930) Die Decapoda Brachyura der Siboga-Expedition. VI. Oxyrhyncha: Parthenopidae. Siboga Expeditie, Monograph, 39 (c 2), 1 - 96." type="journal article" year="1930">Flipse 1930: 48</bibRefCitation>
</treatmentCitation>
(identification key), 87 (index).
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BFC36877F2CFF8FFF79FBA8B1CAD344" blockId="7.[151,1436,1036,1213]" box="[151,1155,1097,1121]" pageId="7" pageNumber="483">
<treatmentCitationGroup id="AB5311A97F2CFF8FFF79FBA8B1CAD344" box="[151,1155,1097,1121]" pageId="7" pageNumber="483">
<taxonomicName id="4C434D047F2CFF8FFF79FBA8B71FD344" ID-CoL="4DTX6" authorityName="Adams &amp; White" authorityYear="1849" box="[151,598,1097,1121]" class="Malacostraca" family="Parthenopidae" genus="Parthenope" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="483" phylum="Arthropoda" rank="species" species="calappoides" subGenus="Parthenolambrus">
<emphasis id="B937EA957F2CFF8FFF79FBA8B444D344" box="[151,269,1097,1121]" italics="true" pageId="7" pageNumber="483">Parthenope</emphasis>
(
<emphasis id="B937EA957F2CFF8FFEF2FBA8B484D344" box="[284,461,1097,1121]" italics="true" pageId="7" pageNumber="483">Parthenolambrus</emphasis>
)
<emphasis id="B937EA957F2CFF8FFE32FBA8B71FD344" box="[476,598,1097,1121]" italics="true" pageId="7" pageNumber="483">calappoides</emphasis>
</taxonomicName>
.—
<treatmentCitation id="0AE210967F2CFF8FFD98FBA8B670D344" author="Rathbun, M. J." box="[630,825,1097,1121]" page="832" pageId="7" pageNumber="483" year="1906">
<bibRefCitation id="EFD24B767F2CFF8FFD98FBA8B670D344" author="Rathbun, M. J." box="[630,825,1097,1121]" pageId="7" pageNumber="483" pagination="827 - 930" refId="ref14062" refString="Rathbun, M. J. (1906) The Brachyura and Macrura of the Hawaiian Islands. Bulletin of the United States Fish Commission, 23 (3), 827 - 930, pls. 1 - 24." type="journal article" year="1906">Rathbun 1906: 832</bibRefCitation>
</treatmentCitation>
(list), 886, pl. 15 fig. 6 (in part).
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BFC36877F2CFF8FFF79FB89B789D3A5" blockId="7.[151,1436,1036,1213]" box="[151,704,1128,1152]" pageId="7" pageNumber="483">
<treatmentCitationGroup id="AB5311A97F2CFF8FFF79FB89B789D3A5" box="[151,704,1128,1152]" pageId="7" pageNumber="483">
<taxonomicName id="4C434D047F2CFF8FFF79FB89B43DD3A5" ID-CoL="3RYTC" authorityName="Calman" authorityYear="1900" box="[151,372,1128,1152]" class="Malacostraca" family="Parthenopidae" genus="Lambrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="483" phylum="Arthropoda" rank="species" species="confragosus">
<emphasis id="B937EA957F2CFF8FFF79FB89B43DD3A5" box="[151,372,1128,1152]" italics="true" pageId="7" pageNumber="483">Lambrus confragosus</emphasis>
</taxonomicName>
<treatmentCitation id="0AE210967F2CFF8FFE63FB89B7F2D3A5" author="Grant, F. E. &amp; McCulloch A. R." box="[397,699,1128,1152]" page="32" pageId="7" pageNumber="483" year="1906">
<bibRefCitation id="EFD24B767F2CFF8FFE63FB89B7F2D3A5" author="Grant, F. E. &amp; McCulloch A. R." box="[397,699,1128,1152]" pageId="7" pageNumber="483" pagination="2 - 53" refId="ref11694" refString="Grant, F. E. &amp; McCulloch A. R. (1906) On a collection of Crustacea from the Port Curtis district, Queensland. Proceedings of the Linnean Society of New South Wales, 31 (1), 2 - 53." type="journal article" year="1906">Grant &amp; McCulloch 1906: 32</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BFC36877F2CFF8FFF79FB66B609D3BA" blockId="7.[151,1436,1036,1213]" box="[151,832,1159,1183]" pageId="7" pageNumber="483">
<treatmentCitationGroup id="AB5311A97F2CFF8FFF79FB66B609D3BA" box="[151,832,1159,1183]" pageId="7" pageNumber="483">
<taxonomicName id="4C434D047F2CFF8FFF79FB66B70CD3BA" box="[151,581,1159,1183]" class="Malacostraca" family="Parthenopidae" genus="Parthenope" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="483" phylum="Arthropoda" rank="species" species="confragosus" subGenus="Pseudolambrus">
<emphasis id="B937EA957F2CFF8FFF79FB66B444D3BA" box="[151,269,1159,1183]" italics="true" pageId="7" pageNumber="483">Parthenope</emphasis>
(
<emphasis id="B937EA957F2CFF8FFEF2FB66B70CD3BA" box="[284,581,1159,1183]" italics="true" pageId="7" pageNumber="483">Pseudolambrus) confragosus</emphasis>
</taxonomicName>
<treatmentCitation id="0AE210967F2CFF8FFDB0FB66B64BD3BA" author="Serene, R." box="[606,770,1159,1183]" page="61" pageId="7" pageNumber="483" year="1968">
<bibRefCitation id="EFD24B767F2CFF8FFDB0FB66B64BD3BA" author="Serene, R." box="[606,770,1159,1183]" pageId="7" pageNumber="483" pagination="1 - 122" refId="ref14708" refString="Serene, R. (1968) Prodromus for a check list of the non-planctonic marine fauna of South East Asia. Singapore National Academy of Science, Special Publication, 1, 1 - 122." type="journal article" year="1968">Serène 1968: 61</bibRefCitation>
</treatmentCitation>
(list).
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BFC36877F2CFF8FFF79FB44B069D398" blockId="7.[151,1436,1036,1213]" box="[151,1312,1189,1213]" pageId="7" pageNumber="483">
<treatmentCitationGroup id="AB5311A97F2CFF8FFF79FB44B069D398" box="[151,1312,1189,1213]" pageId="7" pageNumber="483">
<taxonomicName id="4C434D047F2CFF8FFF79FB44B4FED398" ID-CoL="6WFQ8" baseAuthorityName="Calman" baseAuthorityYear="1900" box="[151,439,1189,1213]" class="Malacostraca" family="Parthenopidae" genus="Pseudolambrus" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="483" phylum="Arthropoda" rank="species" species="confragosus">
<emphasis id="B937EA957F2CFF8FFF79FB44B4FED398" box="[151,439,1189,1213]" italics="true" pageId="7" pageNumber="483">Pseudolambrus confragosus</emphasis>
</taxonomicName>
—Davie 2002: 389 (list).—
<treatmentCitation id="0AE210967F2CFF8FFD21FB47B62AD398" author="Tan, S. H." box="[719,867,1189,1213]" page="367" pageId="7" pageNumber="483" year="2004">
<bibRefCitation id="EFD24B767F2CFF8FFD21FB47B62AD398" author="Tan, S. H." box="[719,867,1189,1213]" pageId="7" pageNumber="483" refId="ref15550" refString="Tan, S. H. (2004) A Systematic Revision of the Parthenopidae (Crustacea: Decapoda: Brachyura). Unpublished PhD Thesis, National University of Singapore, Singapore, 729 pp." type="book" year="2004">Tan 2004: 367</bibRefCitation>
</treatmentCitation>
, figs. 96G, 101.—
<treatmentCitation id="0AE210967F2CFF8FFBF2FB44B1AAD398" author="Ng, P. K. L. &amp; Guinot, D. &amp; Davie, P. J. F." box="[1052,1251,1189,1213]" page="131" pageId="7" pageNumber="483" year="2008">
<bibRefCitation id="EFD24B767F2CFF8FFBF2FB44B1AAD398" author="Ng, P. K. L. &amp; Guinot, D. &amp; Davie, P. J. F." box="[1052,1251,1189,1213]" pageId="7" pageNumber="483" pagination="1 - 286" refId="ref13438" refString="Ng, P. K. L., Guinot, D. &amp; Davie, P. J. F. (2008) Systema Brachyurorum: Part I. An annotated checklist of extant brachyuran crabs of the world. Raffles Bulletin of Zoology, 17, 1 - 286." type="journal article" year="2008">
Ng
<emphasis id="B937EA957F2CFF8FFBAAFB47B13ED398" box="[1092,1143,1189,1213]" italics="true" pageId="7" pageNumber="483">et al.</emphasis>
2008: 131
</bibRefCitation>
</treatmentCitation>
(list).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C359650C7F2CFF8FFF79FB0AB009D20F" pageId="7" pageNumber="483" type="materials_examined">
<paragraph id="8BFC36877F2CFF8FFF79FB0AB009D20F" blockId="7.[151,1437,1259,2042]" pageId="7" pageNumber="483">
<emphasis id="B937EA957F2CFF8FFF79FB0AB4CBD220" bold="true" box="[151,386,1259,1285]" pageId="7" pageNumber="483">Material examined.</emphasis>
<materialsCitation id="3B2B3CDA7F2CFF8FFE64FB0DB074D20F" collectingDate="2019-08-16" collectionCode="FORVSS" collectorName="V. P. Padate" country="India" elevation="56" latitude="6.65" location="Great Nicobar Island" longLatPrecision="782" longitude="93.83" municipality="Nicobars" pageId="7" pageNumber="483" specimenCount="1" specimenCount-male="1">
<specimenCount id="9D45FD0E7F2CFF8FFE64FB0DB49FD220" box="[394,470,1259,1286]" count="1" pageId="7" pageNumber="483" type="male">1 male</specimenCount>
(22.2 × 18.0 mm) (IO/SS/BRC/00323),
<collectionCode id="ED52AE427F2CFF8FFC4DFB0DB142D223" box="[931,1035,1260,1286]" pageId="7" pageNumber="483">FORVSS</collectionCode>
stn. 38818, south of
<location id="8E9C605C7F2CFF8FFB12FB0DB597D20C" LSID="urn:lsid:plazi:treatment:03EA87917F2CFF81FF79FC42B5A1D4EC:8E9C605C7F2CFF8FFB12FB0DB597D20C" country="India" latitude="6.65" longLatPrecision="782" longitude="93.83" municipality="Nicobars" name="Great Nicobar Island" pageId="7" pageNumber="483">Great Nicobar Island</location>
,
<collectingMunicipality id="6B98ACFD7F2CFF8FFF04FAEEB41BD20C" box="[234,338,1295,1321]" pageId="7" pageNumber="483">Nicobars</collectingMunicipality>
,
<collectingCountry id="F35476177F2CFF8FFEB3FAEEB4D0D20C" box="[349,409,1295,1321]" name="India" pageId="7" pageNumber="483">India</collectingCountry>
,
<geoCoordinate id="EE7750407F2CFF8FFE4AFAEEB4BED20F" box="[420,503,1295,1322]" degrees="6.65" direction="north" orientation="latitude" pageId="7" pageNumber="483" precision="555" value="6.65">6.65°N</geoCoordinate>
,
<geoCoordinate id="EE7750407F2CFF8FFDEDFAF1B717D20F" box="[515,606,1296,1322]" degrees="93.83" direction="east" orientation="longitude" pageId="7" pageNumber="483" precision="555" value="93.83">93.83°E</geoCoordinate>
,
<elevation id="006ED1B47F2CFF8FFD82FAEEB7A4D20C" box="[620,749,1295,1322]" metricMagnitude="1" metricUnit="m" metricValue="5.6" pageId="7" pageNumber="483" unit="m" value="56.0">
<quantity id="4CBB9B627F2CFF8FFD82FAEEB7ECD20F" box="[620,677,1295,1322]" metricMagnitude="1" metricUnit="m" metricValue="5.6" pageId="7" pageNumber="483" unit="m" value="56.0">56 m</quantity>
depth
</elevation>
, chain bag dredge, coll.
<collectorName id="26B653517F2CFF8FFC13FAF1B134D20C" box="[1021,1149,1295,1322]" pageId="7" pageNumber="483">V.P. Padate</collectorName>
,
<date id="FFFD10477F2CFF8FFB69FAEEB074D20F" box="[1159,1341,1295,1322]" pageId="7" pageNumber="483" value="2019-08-16">
<collectingDate id="EFB9E9AF7F2CFF8FFB69FAEEB074D20F" box="[1159,1341,1295,1322]" pageId="7" pageNumber="483" value="2019-08-16">16 August 2019</collectingDate>
</date>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C359650C7F2CFF81FF29FAD2B581D7DD" lastPageId="9" lastPageNumber="485" pageId="7" pageNumber="483" type="description">
<paragraph id="8BFC36877F2CFF8FFF29FAD2B5BED100" blockId="7.[151,1437,1259,2042]" pageId="7" pageNumber="483">
<emphasis id="B937EA957F2CFF8FFF29FAD2B410D268" bold="true" box="[199,345,1331,1357]" pageId="7" pageNumber="483">Description.</emphasis>
Carapace subtriangular, 1.2 times as wide as long, widest at posterolateral angle, dorsal surface irregularly tuberculate (
<figureCitation id="13782A027F2CFF8FFE48FAB6B742D257" box="[422,523,1367,1394]" captionStart="FIGURE 2" captionStartId="3.[152,255,1683,1707]" captionTargetBox="[177,1410,181,1659]" captionTargetId="figure-37@3.[177,1410,181,1659]" captionTargetPageId="3" captionText="FIGURE 2. Dorsal habitus photographs (AC, EG: fresh specimens; D: ethanol-preserved specimen): (A) Singhaplax ockelmanni (Serène, 1971), male (IO/SS/BRC/00360, 4.8 × 2.5 mm); (B) Urnalana angulata (Rathbun, 1911), undetermined sex (IO/SS/BRC/00184, 6.9 × 7.0 mm); (C) Pseudolambrus confragosus (Calman, 1900), male (IO/SS/BRC/00323, 22.2 × 18.0 mm; (D) Cavoportunus dubius (Laurie, 1906), male (IO/SS/BRC/00191, 26.9 × 20.4 mm); (E) Cyrtocarcinus truncatus (Rathbun, 1906), male (IO/SS/BRC/00193, 8.6 × 6.1 mm); (F) Visayax osteodictyon Mendoza &amp; Ng, 2008, male (IO/SS/ BRC/00194, 5.2 × 3.6 mm); (G) Xanthias joanneae Mendoza, 2013, male (IO/SS/BRC/00197, 6.4 × 4.2 mm). Scale bars: AB, E, G, 2 mm, C, 10 mm, D, F, 5 mm." figureDoi="http://doi.org/10.5281/zenodo.13210274" httpUri="https://zenodo.org/record/13210274/files/figure.png" pageId="7" pageNumber="483">Figs. 2C</figureCitation>
,
<figureCitation id="13782A027F2CFF8FFDF4FAB6B709D257" box="[538,576,1367,1394]" captionStart="FIGURE 5" captionStartId="8.[152,255,1812,1836]" captionTargetBox="[151,1436,181,1788]" captionTargetId="figure-18@8.[151,1436,181,1788]" captionTargetPageId="8" captionText="FIGURE 5. Pseudolambrus confragosus (Calman, 1900), male (IO/SS/BRC/00323, 22.2 × 18.0 mm),Andaman Sea, India: (A) carapace, dorsal view; (B) carapace, lateral view; (C) carapace, frontal view; (D) right mxp3, ventral view; (E) right chela and carpus, subdorsal view; (F) right P2, dorsal view; (G) male pleon and telson, ventral view. Scale bars: AG, 2 mm." figureDoi="http://doi.org/10.5281/zenodo.13210284" httpUri="https://zenodo.org/record/13210284/files/figure.png" pageId="7" pageNumber="483">5A</figureCitation>
); hepatic region inflated, with strongly projected, medially angled margin; gastric region with a tall median, distally tapering tubercle (
<figureCitation id="13782A027F2CFF8FFCD4FA9AB6DBD2B3" box="[826,914,1403,1430]" captionStart="FIGURE 5" captionStartId="8.[152,255,1812,1836]" captionTargetBox="[151,1436,181,1788]" captionTargetId="figure-18@8.[151,1436,181,1788]" captionTargetPageId="8" captionText="FIGURE 5. Pseudolambrus confragosus (Calman, 1900), male (IO/SS/BRC/00323, 22.2 × 18.0 mm),Andaman Sea, India: (A) carapace, dorsal view; (B) carapace, lateral view; (C) carapace, frontal view; (D) right mxp3, ventral view; (E) right chela and carpus, subdorsal view; (F) right P2, dorsal view; (G) male pleon and telson, ventral view. Scale bars: AG, 2 mm." figureDoi="http://doi.org/10.5281/zenodo.13210284" httpUri="https://zenodo.org/record/13210284/files/figure.png" pageId="7" pageNumber="483">Fig. 5B</figureCitation>
); hepatobranchial and gastrobranchial notches deep; cardiac region with anterior median tubercle 2.0 times as high as posterior median tubercle. Frontal margin relatively long, deflexed, ventral surface with 2 depressions (
<figureCitation id="13782A027F2CFF8FFCD1FA22B6DED2FB" box="[831,919,1475,1502]" captionStart="FIGURE 5" captionStartId="8.[152,255,1812,1836]" captionTargetBox="[151,1436,181,1788]" captionTargetId="figure-18@8.[151,1436,181,1788]" captionTargetPageId="8" captionText="FIGURE 5. Pseudolambrus confragosus (Calman, 1900), male (IO/SS/BRC/00323, 22.2 × 18.0 mm),Andaman Sea, India: (A) carapace, dorsal view; (B) carapace, lateral view; (C) carapace, frontal view; (D) right mxp3, ventral view; (E) right chela and carpus, subdorsal view; (F) right P2, dorsal view; (G) male pleon and telson, ventral view. Scale bars: AG, 2 mm." figureDoi="http://doi.org/10.5281/zenodo.13210284" httpUri="https://zenodo.org/record/13210284/files/figure.png" pageId="7" pageNumber="483">Fig. 5C</figureCitation>
); lateral margins distinctly granular, diverging anteriorly, subparallel posteriorly; posterolateral corners with blunt spines; posterior margin with granulate tubercles dorsally.
</paragraph>
<paragraph id="8BFC36877F2CFF8FFF29F9CEB44ED193" blockId="7.[151,1437,1259,2042]" pageId="7" pageNumber="483">
Antennules folding obliquely; basal antennal article short, article 4 margin with 2 mesial and 1 lateral teeth (
<figureCitation id="13782A027F2CFF8FFA81F9CEB5F2D14B" captionStart="FIGURE 5" captionStartId="8.[152,255,1812,1836]" captionTargetBox="[151,1436,181,1788]" captionTargetId="figure-18@8.[151,1436,181,1788]" captionTargetPageId="8" captionText="FIGURE 5. Pseudolambrus confragosus (Calman, 1900), male (IO/SS/BRC/00323, 22.2 × 18.0 mm),Andaman Sea, India: (A) carapace, dorsal view; (B) carapace, lateral view; (C) carapace, frontal view; (D) right mxp3, ventral view; (E) right chela and carpus, subdorsal view; (F) right P2, dorsal view; (G) male pleon and telson, ventral view. Scale bars: AG, 2 mm." figureDoi="http://doi.org/10.5281/zenodo.13210284" httpUri="https://zenodo.org/record/13210284/files/figure.png" pageId="7" pageNumber="483">Fig. 5C</figureCitation>
); epistome well-developed, posterior margin elevated, median portion with two rounded protuberances (
<figureCitation id="13782A027F2CFF8FFA81F9B2B5F2D1B7" captionStart="FIGURE 5" captionStartId="8.[152,255,1812,1836]" captionTargetBox="[151,1436,181,1788]" captionTargetId="figure-18@8.[151,1436,181,1788]" captionTargetPageId="8" captionText="FIGURE 5. Pseudolambrus confragosus (Calman, 1900), male (IO/SS/BRC/00323, 22.2 × 18.0 mm),Andaman Sea, India: (A) carapace, dorsal view; (B) carapace, lateral view; (C) carapace, frontal view; (D) right mxp3, ventral view; (E) right chela and carpus, subdorsal view; (F) right P2, dorsal view; (G) male pleon and telson, ventral view. Scale bars: AG, 2 mm." figureDoi="http://doi.org/10.5281/zenodo.13210284" httpUri="https://zenodo.org/record/13210284/files/figure.png" pageId="7" pageNumber="483">Fig. 5C</figureCitation>
); mxp3 merus subquadrate, half as long as ischium, with median depression and irregularly arranged granules (
<figureCitation id="13782A027F2CFF8FFF71F97AB5B0D193" box="[159,249,1691,1718]" captionStart="FIGURE 5" captionStartId="8.[152,255,1812,1836]" captionTargetBox="[151,1436,181,1788]" captionTargetId="figure-18@8.[151,1436,181,1788]" captionTargetPageId="8" captionText="FIGURE 5. Pseudolambrus confragosus (Calman, 1900), male (IO/SS/BRC/00323, 22.2 × 18.0 mm),Andaman Sea, India: (A) carapace, dorsal view; (B) carapace, lateral view; (C) carapace, frontal view; (D) right mxp3, ventral view; (E) right chela and carpus, subdorsal view; (F) right P2, dorsal view; (G) male pleon and telson, ventral view. Scale bars: AG, 2 mm." figureDoi="http://doi.org/10.5281/zenodo.13210284" httpUri="https://zenodo.org/record/13210284/files/figure.png" pageId="7" pageNumber="483">Fig. 5D</figureCitation>
).
</paragraph>
<paragraph id="8BFC36877F2CFF8FFF29F95EB4EAD04F" blockId="7.[151,1437,1259,2042]" pageId="7" pageNumber="483">
Chelipeds subequal, 1.7 times CL, merus outer margin with large teeth, inner margin with 3 large triangular teeth (
<figureCitation id="13782A027F2CFF8FFF32F902B47DD1DB" box="[220,308,1763,1790]" captionStart="FIGURE 2" captionStartId="3.[152,255,1683,1707]" captionTargetBox="[177,1410,181,1659]" captionTargetId="figure-37@3.[177,1410,181,1659]" captionTargetPageId="3" captionText="FIGURE 2. Dorsal habitus photographs (AC, EG: fresh specimens; D: ethanol-preserved specimen): (A) Singhaplax ockelmanni (Serène, 1971), male (IO/SS/BRC/00360, 4.8 × 2.5 mm); (B) Urnalana angulata (Rathbun, 1911), undetermined sex (IO/SS/BRC/00184, 6.9 × 7.0 mm); (C) Pseudolambrus confragosus (Calman, 1900), male (IO/SS/BRC/00323, 22.2 × 18.0 mm; (D) Cavoportunus dubius (Laurie, 1906), male (IO/SS/BRC/00191, 26.9 × 20.4 mm); (E) Cyrtocarcinus truncatus (Rathbun, 1906), male (IO/SS/BRC/00193, 8.6 × 6.1 mm); (F) Visayax osteodictyon Mendoza &amp; Ng, 2008, male (IO/SS/ BRC/00194, 5.2 × 3.6 mm); (G) Xanthias joanneae Mendoza, 2013, male (IO/SS/BRC/00197, 6.4 × 4.2 mm). Scale bars: AB, E, G, 2 mm, C, 10 mm, D, F, 5 mm." figureDoi="http://doi.org/10.5281/zenodo.13210274" httpUri="https://zenodo.org/record/13210274/files/figure.png" pageId="7" pageNumber="483">Fig. 2C</figureCitation>
); carpus with dentate carinae on inner and outer margins; palm with large crested lobe at midlength of outer margin, dactylus half as long as palm, with toothed crest along proximal half (
<figureCitation id="13782A027F2CFF8FFBD4F8E6B1C5D007" box="[1082,1164,1799,1826]" captionStart="FIGURE 5" captionStartId="8.[152,255,1812,1836]" captionTargetBox="[151,1436,181,1788]" captionTargetId="figure-18@8.[151,1436,181,1788]" captionTargetPageId="8" captionText="FIGURE 5. Pseudolambrus confragosus (Calman, 1900), male (IO/SS/BRC/00323, 22.2 × 18.0 mm),Andaman Sea, India: (A) carapace, dorsal view; (B) carapace, lateral view; (C) carapace, frontal view; (D) right mxp3, ventral view; (E) right chela and carpus, subdorsal view; (F) right P2, dorsal view; (G) male pleon and telson, ventral view. Scale bars: AG, 2 mm." figureDoi="http://doi.org/10.5281/zenodo.13210284" httpUri="https://zenodo.org/record/13210284/files/figure.png" pageId="7" pageNumber="483">Fig. 5E</figureCitation>
). Pereopods 25 shorter than chelipeds, densely tomentose, outer margins of meri, carpi and propodi armed with larger teeth compared with inner margins (
<figureCitation id="13782A027F2CFF8FFEAFF8AEB4DFD04F" box="[321,406,1871,1898]" captionStart="FIGURE 5" captionStartId="8.[152,255,1812,1836]" captionTargetBox="[151,1436,181,1788]" captionTargetId="figure-18@8.[151,1436,181,1788]" captionTargetPageId="8" captionText="FIGURE 5. Pseudolambrus confragosus (Calman, 1900), male (IO/SS/BRC/00323, 22.2 × 18.0 mm),Andaman Sea, India: (A) carapace, dorsal view; (B) carapace, lateral view; (C) carapace, frontal view; (D) right mxp3, ventral view; (E) right chela and carpus, subdorsal view; (F) right P2, dorsal view; (G) male pleon and telson, ventral view. Scale bars: AG, 2 mm." figureDoi="http://doi.org/10.5281/zenodo.13210284" httpUri="https://zenodo.org/record/13210284/files/figure.png" pageId="7" pageNumber="483">Fig. 5F</figureCitation>
).
</paragraph>
<paragraph id="8BFC36877F2CFF8FFF29F892B037D0DF" blockId="7.[151,1437,1259,2042]" pageId="7" pageNumber="483">
Male pleon elongate sub-triangular, somites 35 fused, somite 6 longest, ventral surfaces of somites 35 with a transverse row of 4 granules each, that of somite 6 with median granule (
<figureCitation id="13782A027F2CFF8FFC31F876B173D097" box="[991,1082,1943,1970]" captionStart="FIGURE 5" captionStartId="8.[152,255,1812,1836]" captionTargetBox="[151,1436,181,1788]" captionTargetId="figure-18@8.[151,1436,181,1788]" captionTargetPageId="8" captionText="FIGURE 5. Pseudolambrus confragosus (Calman, 1900), male (IO/SS/BRC/00323, 22.2 × 18.0 mm),Andaman Sea, India: (A) carapace, dorsal view; (B) carapace, lateral view; (C) carapace, frontal view; (D) right mxp3, ventral view; (E) right chela and carpus, subdorsal view; (F) right P2, dorsal view; (G) male pleon and telson, ventral view. Scale bars: AG, 2 mm." figureDoi="http://doi.org/10.5281/zenodo.13210284" httpUri="https://zenodo.org/record/13210284/files/figure.png" pageId="7" pageNumber="483">Fig. 5G</figureCitation>
); telson bluntly triangular, twothirds as long as wide (
<figureCitation id="13782A027F2CFF8FFE72F85AB4BED0F3" box="[412,503,1979,2006]" captionStart="FIGURE 5" captionStartId="8.[152,255,1812,1836]" captionTargetBox="[151,1436,181,1788]" captionTargetId="figure-18@8.[151,1436,181,1788]" captionTargetPageId="8" captionText="FIGURE 5. Pseudolambrus confragosus (Calman, 1900), male (IO/SS/BRC/00323, 22.2 × 18.0 mm),Andaman Sea, India: (A) carapace, dorsal view; (B) carapace, lateral view; (C) carapace, frontal view; (D) right mxp3, ventral view; (E) right chela and carpus, subdorsal view; (F) right P2, dorsal view; (G) male pleon and telson, ventral view. Scale bars: AG, 2 mm." figureDoi="http://doi.org/10.5281/zenodo.13210284" httpUri="https://zenodo.org/record/13210284/files/figure.png" pageId="7" pageNumber="483">Fig. 5G</figureCitation>
). G1 stout, nearly straight (
<figureCitation id="13782A027F2CFF8FFCC2F85AB6FED0F3" box="[812,951,1979,2006]" captionStart="FIGURE 10" captionStartId="17.[151,250,1667,1691]" captionTargetBox="[304,1294,191,1633]" captionTargetId="figure-37@17.[292,1295,181,1645]" captionTargetPageId="17" captionText="FIGURE 10. First and second gonopods: Singhaplax ockelmanni (Serène, 1971), male (IO/SS/BRC/00360, 4.8 × 2.5 mm): (A) entire G1, pleonal view; (B) entire G1, sternal view; (C) entire G2, pleonal view. Pseudolambrus confragosus (Calman, 1900), male (IO/SS/BRC/00323, 22.2 × 18.0 mm): (D) entire G1, pleonal view; (E) G1 distal portion, pleonal view; (F) G1 distal portion, sternal view; (G) entire G1, sternal view; (H) entire G2, pleonal view. Cavoportunus dubius (Laurie, 1906), male (IO/SS/BRC/00191, 26.9 × 20.4 mm): (I) entire G1, pleonal view; (J) entire G1, lateral view; (K) entire G1, mesial view; (L) entire G2, pleonal view. Cyrtocarcinus truncatus (Rathbun, 1906), male (IO/SS/BRC/00193, 8.6 × 6.1 mm): (M) entire G1, pleonal view; (N) entire G1, sternal view; (O) entire G2, sternal view. Visayax osteodictyon Mendoza &amp; Ng, 2008, male (IO/ SS/BRC/00194, 5.2 × 3.6 mm): (P) entire G1, pleonal view; (Q) G1 distal portion, pleonal view; (R) entire G1, sternal view. Xanthias joanneae Mendoza, 2013, male (IO/SS/BRC/00197, 6.4 × 4.2 mm). (S) entire G1, pleonal view; (T) G1 distal portion, pleonal view; (U) G1 distal portion, sternal view; (V) entire G1, sternal view. Scale bars: AC, PV, 0.2 mm, D, GN, 1 mm, E, F, O, 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13210300" httpUri="https://zenodo.org/record/13210300/files/figure.png" pageId="7" pageNumber="483">Fig. 10D, G</figureCitation>
), distal portion blunt with subdistal ventral spinose patch, spines on mesial margin extending to proximal half (
<figureCitation id="13782A027F2CFF8FFC62F83EB146D0DF" box="[908,1039,2015,2042]" captionStart="FIGURE 10" captionStartId="17.[151,250,1667,1691]" captionTargetBox="[304,1294,191,1633]" captionTargetId="figure-37@17.[292,1295,181,1645]" captionTargetPageId="17" captionText="FIGURE 10. First and second gonopods: Singhaplax ockelmanni (Serène, 1971), male (IO/SS/BRC/00360, 4.8 × 2.5 mm): (A) entire G1, pleonal view; (B) entire G1, sternal view; (C) entire G2, pleonal view. Pseudolambrus confragosus (Calman, 1900), male (IO/SS/BRC/00323, 22.2 × 18.0 mm): (D) entire G1, pleonal view; (E) G1 distal portion, pleonal view; (F) G1 distal portion, sternal view; (G) entire G1, sternal view; (H) entire G2, pleonal view. Cavoportunus dubius (Laurie, 1906), male (IO/SS/BRC/00191, 26.9 × 20.4 mm): (I) entire G1, pleonal view; (J) entire G1, lateral view; (K) entire G1, mesial view; (L) entire G2, pleonal view. Cyrtocarcinus truncatus (Rathbun, 1906), male (IO/SS/BRC/00193, 8.6 × 6.1 mm): (M) entire G1, pleonal view; (N) entire G1, sternal view; (O) entire G2, sternal view. Visayax osteodictyon Mendoza &amp; Ng, 2008, male (IO/ SS/BRC/00194, 5.2 × 3.6 mm): (P) entire G1, pleonal view; (Q) G1 distal portion, pleonal view; (R) entire G1, sternal view. Xanthias joanneae Mendoza, 2013, male (IO/SS/BRC/00197, 6.4 × 4.2 mm). (S) entire G1, pleonal view; (T) G1 distal portion, pleonal view; (U) G1 distal portion, sternal view; (V) entire G1, sternal view. Scale bars: AC, PV, 0.2 mm, D, GN, 1 mm, E, F, O, 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13210300" httpUri="https://zenodo.org/record/13210300/files/figure.png" pageId="7" pageNumber="483">Fig. 10EF</figureCitation>
). G2 shorter than G1 (
<figureCitation id="13782A027F2CFF8FFAE7F83EB024D0DF" box="[1289,1389,2015,2042]" captionStart="FIGURE 10" captionStartId="17.[151,250,1667,1691]" captionTargetBox="[304,1294,191,1633]" captionTargetId="figure-37@17.[292,1295,181,1645]" captionTargetPageId="17" captionText="FIGURE 10. First and second gonopods: Singhaplax ockelmanni (Serène, 1971), male (IO/SS/BRC/00360, 4.8 × 2.5 mm): (A) entire G1, pleonal view; (B) entire G1, sternal view; (C) entire G2, pleonal view. Pseudolambrus confragosus (Calman, 1900), male (IO/SS/BRC/00323, 22.2 × 18.0 mm): (D) entire G1, pleonal view; (E) G1 distal portion, pleonal view; (F) G1 distal portion, sternal view; (G) entire G1, sternal view; (H) entire G2, pleonal view. Cavoportunus dubius (Laurie, 1906), male (IO/SS/BRC/00191, 26.9 × 20.4 mm): (I) entire G1, pleonal view; (J) entire G1, lateral view; (K) entire G1, mesial view; (L) entire G2, pleonal view. Cyrtocarcinus truncatus (Rathbun, 1906), male (IO/SS/BRC/00193, 8.6 × 6.1 mm): (M) entire G1, pleonal view; (N) entire G1, sternal view; (O) entire G2, sternal view. Visayax osteodictyon Mendoza &amp; Ng, 2008, male (IO/ SS/BRC/00194, 5.2 × 3.6 mm): (P) entire G1, pleonal view; (Q) G1 distal portion, pleonal view; (R) entire G1, sternal view. Xanthias joanneae Mendoza, 2013, male (IO/SS/BRC/00197, 6.4 × 4.2 mm). (S) entire G1, pleonal view; (T) G1 distal portion, pleonal view; (U) G1 distal portion, sternal view; (V) entire G1, sternal view. Scale bars: AC, PV, 0.2 mm, D, GN, 1 mm, E, F, O, 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13210300" httpUri="https://zenodo.org/record/13210300/files/figure.png" pageId="7" pageNumber="483">Fig. 10H</figureCitation>
).
</paragraph>
<caption id="DF3C660F7F23FF80FF76F8F5B068D051" ID-DOI="http://doi.org/10.5281/zenodo.13210284" ID-Zenodo-Dep="13210284" httpUri="https://zenodo.org/record/13210284/files/figure.png" pageId="8" pageNumber="484" startId="8.[152,255,1812,1836]" targetBox="[151,1436,181,1788]" targetPageId="8" targetType="figure">
<paragraph id="8BFC36877F23FF80FF76F8F5B068D051" blockId="8.[151,1437,1812,1908]" pageId="8" pageNumber="484">
<emphasis id="B937EA957F23FF80FF76F8F5B458D009" bold="true" box="[152,273,1812,1836]" pageId="8" pageNumber="484">FIGURE 5</emphasis>
.
<taxonomicName id="4C434D047F23FF80FEF2F8F5B7A9D009" authority="(Calman, 1900)" baseAuthorityName="Calman" baseAuthorityYear="1900" box="[284,736,1812,1836]" class="Malacostraca" family="Parthenopidae" genus="Pseudolambrus" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="484" phylum="Arthropoda" rank="species" species="confragosus">
<emphasis id="B937EA957F23FF80FEF2F8F5B772D009" box="[284,571,1812,1836]" italics="true" pageId="8" pageNumber="484">Pseudolambrus confragosus</emphasis>
(
<bibRefCitation id="EFD24B767F23FF80FDA9F8F5B790D009" author="Calman, W. T." box="[583,729,1812,1836]" pageId="8" pageNumber="484" pagination="1 - 50" refId="ref10828" refString="Calman, W. T. (1900) On a collection of Brachyura from Torres Straits. Transactions of the Linnean Society of London, Series 2, Zoology, 8, 1 - 50, pls. 1 - 3. https: // doi. org / 10.1111 / j. 1096 - 3642.1900. tb 00307. x" type="journal article" year="1900">Calman, 1900</bibRefCitation>
)
</taxonomicName>
, male (IO/SS/BRC/00323, 22.2 × 18.0 mm), Andaman Sea, India: (A) carapace, dorsal view; (B) carapace, lateral view; (C) carapace, frontal view; (D) right mxp3, ventral view; (E) right chela and carpus, subdorsal view; (F) right P2, dorsal view; (G) male pleon and telson, ventral view. Scale bars: AG, 2 mm.
</paragraph>
</caption>
<paragraph id="8BFC36877F22FF81FF29FF77B581D7DD" blockId="9.[151,1437,150,969]" pageId="9" pageNumber="485">
<emphasis id="B937EA957F22FF81FF29FF77B416D795" bold="true" box="[199,351,150,176]" pageId="9" pageNumber="485">Colouration.</emphasis>
Fresh specimen: Carapace brick red, chelipeds brick red with whitish bands on proximal half of merus and midlength of palm, fingers whitish with brown tips, pereopods whitish with brick red patch on meri (
<figureCitation id="13782A027F22FF81FA81FF5BB5F5D7DD" captionStart="FIGURE 2" captionStartId="3.[152,255,1683,1707]" captionTargetBox="[177,1410,181,1659]" captionTargetId="figure-37@3.[177,1410,181,1659]" captionTargetPageId="3" captionText="FIGURE 2. Dorsal habitus photographs (AC, EG: fresh specimens; D: ethanol-preserved specimen): (A) Singhaplax ockelmanni (Serène, 1971), male (IO/SS/BRC/00360, 4.8 × 2.5 mm); (B) Urnalana angulata (Rathbun, 1911), undetermined sex (IO/SS/BRC/00184, 6.9 × 7.0 mm); (C) Pseudolambrus confragosus (Calman, 1900), male (IO/SS/BRC/00323, 22.2 × 18.0 mm; (D) Cavoportunus dubius (Laurie, 1906), male (IO/SS/BRC/00191, 26.9 × 20.4 mm); (E) Cyrtocarcinus truncatus (Rathbun, 1906), male (IO/SS/BRC/00193, 8.6 × 6.1 mm); (F) Visayax osteodictyon Mendoza &amp; Ng, 2008, male (IO/SS/ BRC/00194, 5.2 × 3.6 mm); (G) Xanthias joanneae Mendoza, 2013, male (IO/SS/BRC/00197, 6.4 × 4.2 mm). Scale bars: AB, E, G, 2 mm, C, 10 mm, D, F, 5 mm." figureDoi="http://doi.org/10.5281/zenodo.13210274" httpUri="https://zenodo.org/record/13210274/files/figure.png" pageId="9" pageNumber="485">Fig. 2C</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C359650C7F22FF81FF29FEE3B486D641" pageId="9" pageNumber="485" type="biology_ecology">
<paragraph id="8BFC36877F22FF81FF29FEE3B486D641" blockId="9.[151,1437,150,969]" pageId="9" pageNumber="485">
<emphasis id="B937EA957F22FF81FF29FEE3B46ED639" bold="true" box="[199,295,258,284]" pageId="9" pageNumber="485">Biology.</emphasis>
The present specimen was collected from rocky substrate in the vicinity of crinoids, at
<quantity id="4CBB9B627F22FF81FAFAFEE3B006D638" box="[1300,1359,258,285]" metricMagnitude="1" metricUnit="m" metricValue="5.6" pageId="9" pageNumber="485" unit="m" value="56.0">56 m</quantity>
depth. Known to inhabit coral and sponge colonies, and coarse calcareous sand in reefs (
<bibRefCitation id="EFD24B767F22FF81FBC7FEC7B1FDD665" author="Laurie, R. D." box="[1065,1204,294,320]" pageId="9" pageNumber="485" pagination="349 - 432" refId="ref12375" refString="Laurie, R. D. (1906) Report on the Brachyura collected by Professor Herdman, at Ceylon, in 1902. In: Herdman, W. A. (Ed.), Report to the Government of Ceylon on the Pearl Oyster Fisheries of the Gulf of Manaar with Supplementary Reports Upon the Marine Biology of Ceylon by Other Naturalists. Part 5. The Royal Society, London, pp. 349 - 432." type="book chapter" year="1906">Laurie 1906</bibRefCitation>
;
<bibRefCitation id="EFD24B767F22FF81FB51FEC6B063D665" author="Tan, S. H." box="[1215,1322,294,321]" pageId="9" pageNumber="485" refId="ref15550" refString="Tan, S. H. (2004) A Systematic Revision of the Parthenopidae (Crustacea: Decapoda: Brachyura). Unpublished PhD Thesis, National University of Singapore, Singapore, 729 pp." type="book" year="2004">Tan 2004</bibRefCitation>
) at depths from
<quantity id="4CBB9B627F22FF81FF3BFEABB40FD640" box="[213,326,330,357]" metricMagnitude="2" metricUnit="m" metricValue="2.59" metricValueMax="5.0" metricValueMin="0.18" pageId="9" pageNumber="485" unit="m" value="259.0" valueMax="500.0" valueMin="18.0">18500 m</quantity>
(
<bibRefCitation id="EFD24B767F22FF81FEB8FEAAB48AD641" author="Tan, S. H." box="[342,451,330,357]" pageId="9" pageNumber="485" refId="ref15550" refString="Tan, S. H. (2004) A Systematic Revision of the Parthenopidae (Crustacea: Decapoda: Brachyura). Unpublished PhD Thesis, National University of Singapore, Singapore, 729 pp." type="book" year="2004">Tan 2004</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C359650C7F22FF81FF29FE8FB5A1D4EC" pageId="9" pageNumber="485" type="discussion">
<paragraph id="8BFC36877F22FF81FF29FE8FB01ED41D" blockId="9.[151,1437,150,969]" pageId="9" pageNumber="485">
<emphasis id="B937EA957F22FF81FF29FE8FB472D6AD" bold="true" box="[199,315,366,392]" pageId="9" pageNumber="485">Remarks.</emphasis>
<bibRefCitation id="EFD24B767F22FF81FEAAFE8FB4A6D6AD" author="Calman, W. T." box="[324,495,366,392]" pageId="9" pageNumber="485" pagination="1 - 50" refId="ref10828" refString="Calman, W. T. (1900) On a collection of Brachyura from Torres Straits. Transactions of the Linnean Society of London, Series 2, Zoology, 8, 1 - 50, pls. 1 - 3. https: // doi. org / 10.1111 / j. 1096 - 3642.1900. tb 00307. x" type="journal article" year="1900">Calman (1900)</bibRefCitation>
described
<taxonomicName id="4C434D047F22FF81FD83FE8FB10AD6AD" authorityName="Calman" authorityYear="1900" box="[621,1091,366,392]" class="Malacostraca" family="Parthenopidae" genus="Lambrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="485" phylum="Arthropoda" rank="species" species="confragosus" subGenus="Parthenolambrus">
<emphasis id="B937EA957F22FF81FD83FE8FB798D6AD" box="[621,721,366,392]" italics="true" pageId="9" pageNumber="485">Lambrus</emphasis>
(
<emphasis id="B937EA957F22FF81FD0AFE8FB6E1D6AD" box="[740,936,366,392]" italics="true" pageId="9" pageNumber="485">Parthenolambrus</emphasis>
)
<emphasis id="B937EA957F22FF81FC54FE8FB10AD6AD" box="[954,1091,366,392]" italics="true" pageId="9" pageNumber="485">confragosus</emphasis>
</taxonomicName>
from a female collected from Torres Strait,
<collectingCountry id="F35476177F22FF81FEDEFE73B4D3D689" box="[304,410,402,428]" name="Australia" pageId="9" pageNumber="485">Australia</collectingCountry>
, on the basis of a highly rugged carapace, compressed dentiform hepatic prominences, and the flattened lamelliform lobes on the outer margin of the cheliped palm. According to
<bibRefCitation id="EFD24B767F22FF81FB8BFE56B1AAD6F5" author="Tan, S. H." box="[1125,1251,438,465]" pageId="9" pageNumber="485" refId="ref15550" refString="Tan, S. H. (2004) A Systematic Revision of the Parthenopidae (Crustacea: Decapoda: Brachyura). Unpublished PhD Thesis, National University of Singapore, Singapore, 729 pp." type="book" year="2004">Tan (2004)</bibRefCitation>
,
<taxonomicName id="4C434D047F22FF81FB00FE57B469D6D1" baseAuthorityName="Calman" baseAuthorityYear="1900" class="Malacostraca" family="Parthenopidae" genus="Pseudolambrus" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="485" phylum="Arthropoda" rank="species" species="confragosus">
<emphasis id="B937EA957F22FF81FB00FE57B469D6D1" italics="true" pageId="9" pageNumber="485">Pseudolambrus confragosus</emphasis>
</taxonomicName>
resembles
<taxonomicName id="4C434D047F22FF81FE73FE3AB661D6D1" authority="(Adams &amp; White, 1849)" baseAuthorityName="Adams &amp; White" baseAuthorityYear="1849" box="[413,808,474,500]" class="Malacostraca" family="Parthenopidae" genus="Pseudolambrus" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="485" phylum="Arthropoda" rank="species" species="tarpeius">
<emphasis id="B937EA957F22FF81FE73FE3AB75DD6D1" box="[413,532,475,500]" italics="true" pageId="9" pageNumber="485">P. tarpeius</emphasis>
(Adams &amp; White, 1849)
</taxonomicName>
in the carapace armed with irregular, granular tubercles, and the presence of granular tubercles on the dorsal surface of carapace posterior margin. However, it differs from
<taxonomicName id="4C434D047F22FF81FF79FDC2B459D519" baseAuthorityName="Adams &amp; White" baseAuthorityYear="1849" box="[151,272,547,572]" class="Malacostraca" family="Parthenopidae" genus="Pseudolambrus" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="485" phylum="Arthropoda" rank="species" species="tarpeius">
<emphasis id="B937EA957F22FF81FF79FDC2B459D519" box="[151,272,547,572]" italics="true" pageId="9" pageNumber="485">P. tarpeius</emphasis>
</taxonomicName>
in having a narrower hepatic region, relatively more inflated epibranchial region, and the presence of a lamelliform lobe on the outer surface of cheliped palm (
<figureCitation id="13782A027F22FF81FCD6FDA7B6FAD544" box="[824,947,582,609]" captionStart="FIGURE 5" captionStartId="8.[152,255,1812,1836]" captionTargetBox="[151,1436,181,1788]" captionTargetId="figure-18@8.[151,1436,181,1788]" captionTargetPageId="8" captionText="FIGURE 5. Pseudolambrus confragosus (Calman, 1900), male (IO/SS/BRC/00323, 22.2 × 18.0 mm),Andaman Sea, India: (A) carapace, dorsal view; (B) carapace, lateral view; (C) carapace, frontal view; (D) right mxp3, ventral view; (E) right chela and carpus, subdorsal view; (F) right P2, dorsal view; (G) male pleon and telson, ventral view. Scale bars: AG, 2 mm." figureDoi="http://doi.org/10.5281/zenodo.13210284" httpUri="https://zenodo.org/record/13210284/files/figure.png" pageId="9" pageNumber="485">Fig. 5A, E</figureCitation>
) (versus broader hepatic region, relatively less inflated epibranchial region, and the absence of a lamelliform lobe on the outer surface of cheliped palm in
<taxonomicName id="4C434D047F22FF81FF79FD6EB459D58D" baseAuthorityName="Adams &amp; White" baseAuthorityYear="1849" box="[151,272,655,680]" class="Malacostraca" family="Parthenopidae" genus="Pseudolambrus" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="485" phylum="Arthropoda" rank="species" species="tarpeius">
<emphasis id="B937EA957F22FF81FF79FD6EB459D58D" box="[151,272,655,680]" italics="true" pageId="9" pageNumber="485">P. tarpeius</emphasis>
</taxonomicName>
; see
<bibRefCitation id="EFD24B767F22FF81FEA3FD6EB4F2D58D" author="Tan, S. H." box="[333,443,654,681]" pageId="9" pageNumber="485" refId="ref15550" refString="Tan, S. H. (2004) A Systematic Revision of the Parthenopidae (Crustacea: Decapoda: Brachyura). Unpublished PhD Thesis, National University of Singapore, Singapore, 729 pp." type="book" year="2004">Tan 2004</bibRefCitation>
: fig. 104A). The Indian specimen conforms to the description of the
<typeStatus id="54F888257F22FF81FB3FFD6FB07AD58D" box="[1233,1331,654,680]" pageId="9" pageNumber="485" type="holotype">holotype</typeStatus>
(
<bibRefCitation id="EFD24B767F22FF81FAADFD6FB599D5E9" author="Calman, W. T." pageId="9" pageNumber="485" pagination="1 - 50" refId="ref10828" refString="Calman, W. T. (1900) On a collection of Brachyura from Torres Straits. Transactions of the Linnean Society of London, Series 2, Zoology, 8, 1 - 50, pls. 1 - 3. https: // doi. org / 10.1111 / j. 1096 - 3642.1900. tb 00307. x" type="journal article" year="1900">Calman 1900</bibRefCitation>
) as well as subsequent descriptions (
<bibRefCitation id="EFD24B767F22FF81FD97FD53B64FD5E9" author="Laurie, R. D." box="[633,774,690,716]" pageId="9" pageNumber="485" pagination="349 - 432" refId="ref12375" refString="Laurie, R. D. (1906) Report on the Brachyura collected by Professor Herdman, at Ceylon, in 1902. In: Herdman, W. A. (Ed.), Report to the Government of Ceylon on the Pearl Oyster Fisheries of the Gulf of Manaar with Supplementary Reports Upon the Marine Biology of Ceylon by Other Naturalists. Part 5. The Royal Society, London, pp. 349 - 432." type="book chapter" year="1906">Laurie 1906</bibRefCitation>
;
<bibRefCitation id="EFD24B767F22FF81FCFDFD53B6FFD5E9" author="Rathbun, M. J." box="[787,950,690,716]" pageId="9" pageNumber="485" pagination="827 - 930" refId="ref14062" refString="Rathbun, M. J. (1906) The Brachyura and Macrura of the Hawaiian Islands. Bulletin of the United States Fish Commission, 23 (3), 827 - 930, pls. 1 - 24." type="journal article" year="1906">Rathbun 1906</bibRefCitation>
;
<bibRefCitation id="EFD24B767F22FF81FC2DFD52B166D5E9" author="Tan, S. H." box="[963,1071,690,717]" pageId="9" pageNumber="485" refId="ref15550" refString="Tan, S. H. (2004) A Systematic Revision of the Parthenopidae (Crustacea: Decapoda: Brachyura). Unpublished PhD Thesis, National University of Singapore, Singapore, 729 pp." type="book" year="2004">Tan 2004</bibRefCitation>
) in the distinctly tubercular and rugged surface of the carapace and chelipeds. However, the ornamentation on the G1 differs from the G1 of a male specimen from the Balicasag Island,
<collectingCountry id="F35476177F22FF81FDD0FD1BB7F5D431" box="[574,700,762,788]" name="Philippines" pageId="9" pageNumber="485">Philippines</collectingCountry>
in the presence of a subdistal spiny patch on the ventral surface (
<figureCitation id="13782A027F22FF81FF71FCFFB46AD41C" box="[159,291,798,825]" captionStart="FIGURE 10" captionStartId="17.[151,250,1667,1691]" captionTargetBox="[304,1294,191,1633]" captionTargetId="figure-37@17.[292,1295,181,1645]" captionTargetPageId="17" captionText="FIGURE 10. First and second gonopods: Singhaplax ockelmanni (Serène, 1971), male (IO/SS/BRC/00360, 4.8 × 2.5 mm): (A) entire G1, pleonal view; (B) entire G1, sternal view; (C) entire G2, pleonal view. Pseudolambrus confragosus (Calman, 1900), male (IO/SS/BRC/00323, 22.2 × 18.0 mm): (D) entire G1, pleonal view; (E) G1 distal portion, pleonal view; (F) G1 distal portion, sternal view; (G) entire G1, sternal view; (H) entire G2, pleonal view. Cavoportunus dubius (Laurie, 1906), male (IO/SS/BRC/00191, 26.9 × 20.4 mm): (I) entire G1, pleonal view; (J) entire G1, lateral view; (K) entire G1, mesial view; (L) entire G2, pleonal view. Cyrtocarcinus truncatus (Rathbun, 1906), male (IO/SS/BRC/00193, 8.6 × 6.1 mm): (M) entire G1, pleonal view; (N) entire G1, sternal view; (O) entire G2, sternal view. Visayax osteodictyon Mendoza &amp; Ng, 2008, male (IO/ SS/BRC/00194, 5.2 × 3.6 mm): (P) entire G1, pleonal view; (Q) G1 distal portion, pleonal view; (R) entire G1, sternal view. Xanthias joanneae Mendoza, 2013, male (IO/SS/BRC/00197, 6.4 × 4.2 mm). (S) entire G1, pleonal view; (T) G1 distal portion, pleonal view; (U) G1 distal portion, sternal view; (V) entire G1, sternal view. Scale bars: AC, PV, 0.2 mm, D, GN, 1 mm, E, F, O, 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13210300" httpUri="https://zenodo.org/record/13210300/files/figure.png" pageId="9" pageNumber="485">Fig. 10D, E</figureCitation>
) (versus G1 with smooth ventral surface in the Philippine specimen; cf.
<bibRefCitation id="EFD24B767F22FF81FBA1FCFEB1F5D41D" author="Tan, S. H." box="[1103,1212,798,825]" pageId="9" pageNumber="485" refId="ref15550" refString="Tan, S. H. (2004) A Systematic Revision of the Parthenopidae (Crustacea: Decapoda: Brachyura). Unpublished PhD Thesis, National University of Singapore, Singapore, 729 pp." type="book" year="2004">Tan 2004</bibRefCitation>
: fig. 96G, H).
</paragraph>
<paragraph id="8BFC36877F22FF81FF29FCA3B5A1D4EC" blockId="9.[151,1437,150,969]" pageId="9" pageNumber="485">
<emphasis id="B937EA957F22FF81FF29FCA3B742D479" bold="true" box="[199,523,834,860]" pageId="9" pageNumber="485">Geographical distribution.</emphasis>
Torres Straits (
<bibRefCitation id="EFD24B767F22FF81FD24FCA3B625D479" author="Calman, W. T." box="[714,876,834,860]" pageId="9" pageNumber="485" pagination="1 - 50" refId="ref10828" refString="Calman, W. T. (1900) On a collection of Brachyura from Torres Straits. Transactions of the Linnean Society of London, Series 2, Zoology, 8, 1 - 50, pls. 1 - 3. https: // doi. org / 10.1111 / j. 1096 - 3642.1900. tb 00307. x" type="journal article" year="1900">Calman 1900</bibRefCitation>
),
<collectingCountry id="F35476177F22FF81FC68FCA3B6B3D479" box="[902,1018,834,860]" name="Sri Lanka" pageId="9" pageNumber="485">Sri Lanka</collectingCountry>
(
<bibRefCitation id="EFD24B767F22FF81FBFFFCA3B1EAD479" author="Laurie, R. D." box="[1041,1187,834,860]" pageId="9" pageNumber="485" pagination="349 - 432" refId="ref12375" refString="Laurie, R. D. (1906) Report on the Brachyura collected by Professor Herdman, at Ceylon, in 1902. In: Herdman, W. A. (Ed.), Report to the Government of Ceylon on the Pearl Oyster Fisheries of the Gulf of Manaar with Supplementary Reports Upon the Marine Biology of Ceylon by Other Naturalists. Part 5. The Royal Society, London, pp. 349 - 432." type="book chapter" year="1906">Laurie 1906</bibRefCitation>
),
<collectingCountry id="F35476177F22FF81FB52FCA3B06AD479" box="[1212,1315,834,860]" name="Australia" pageId="9" pageNumber="485">Australia</collectingCountry>
(
<bibRefCitation id="EFD24B767F22FF81FAD9FCA3B413D4A5" author="Grant, F. E. &amp; McCulloch A. R." pageId="9" pageNumber="485" pagination="2 - 53" refId="ref11694" refString="Grant, F. E. &amp; McCulloch A. R. (1906) On a collection of Crustacea from the Port Curtis district, Queensland. Proceedings of the Linnean Society of New South Wales, 31 (1), 2 - 53." type="journal article" year="1906">Grant &amp; McCulloch 1906</bibRefCitation>
) and Hawaii (
<bibRefCitation id="EFD24B767F22FF81FDE9FC87B7E4D4A5" author="Rathbun, M. J." box="[519,685,870,896]" pageId="9" pageNumber="485" pagination="827 - 930" refId="ref14062" refString="Rathbun, M. J. (1906) The Brachyura and Macrura of the Hawaiian Islands. Bulletin of the United States Fish Commission, 23 (3), 827 - 930, pls. 1 - 24." type="journal article" year="1906">Rathbun 1906</bibRefCitation>
),
<collectingCountry id="F35476177F22FF81FD2AFC87B676D4A5" box="[708,831,870,896]" name="Seychelles" pageId="9" pageNumber="485">Seychelles</collectingCountry>
,
<collectingCountry id="F35476177F22FF81FCA1FC87B6F6D4A5" box="[847,959,870,896]" name="Mauritius" pageId="9" pageNumber="485">Mauritius</collectingCountry>
, Gulf of Aden, the
<collectingCountry id="F35476177F22FF81FB44FC87B061D4A5" box="[1194,1320,870,896]" name="Philippines" pageId="9" pageNumber="485">Philippines</collectingCountry>
and
<collectingCountry id="F35476177F22FF81FA89FC86B440D481" name="New Caledonia" pageId="9" pageNumber="485">New Caledonia</collectingCountry>
(
<bibRefCitation id="EFD24B767F22FF81FEF4FC6AB4C1D481" author="Tan, S. H." box="[282,392,906,933]" pageId="9" pageNumber="485" refId="ref15550" refString="Tan, S. H. (2004) A Systematic Revision of the Parthenopidae (Crustacea: Decapoda: Brachyura). Unpublished PhD Thesis, National University of Singapore, Singapore, 729 pp." type="book" year="2004">Tan 2004</bibRefCitation>
); and Nicobars,
<collectingCountry id="F35476177F22FF81FDAEFC6BB733D481" box="[576,634,906,932]" name="India" pageId="9" pageNumber="485">India</collectingCountry>
(present study). The present observation is the first record from Indian waters.
</paragraph>
</subSubSection>
</treatment>
</document>