230 lines
42 KiB
XML
230 lines
42 KiB
XML
<document id="4DB959D666FF06DE0785215DB0608906" ID-DOI="10.1080/00222930500218664" ID-ISSN="1464-5262" ID-Zenodo-Dep="5220042" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1629351132246" checkinUser="felipe" docAuthor="Schmelz, R. M. & Römbke, J." docDate="2005" docId="03E3594E584D026B6DC1042FA9B7FD39" docLanguage="en" docName="JNatHist.39.32.2967-2986.pdf" docOrigin="Journal of Natural History 39 (32)" docSource="http://dx.doi.org/10.1080/00222930500218664" docStyle="DocumentStyle:5A9EB2ED5589A523F0236C66DE2A6BA8.4:JNatHist.2005-2007.journal_article.1cover" docStyleId="5A9EB2ED5589A523F0236C66DE2A6BA8" docStyleName="JNatHist.2005-2007.journal_article.1cover" docStyleVersion="4" docTitle="Hemienchytraeus siljae Schmelz & Römbke 2005, sp. nov." docType="treatment" docVersion="3" lastPageNumber="2973" masterDocId="FFDA21365848026C6C0B0632AA02FF9E" masterDocTitle="Three new species of Hemienchytraeus (Enchytraeidae, Oligochaeta) from Amazonian forest soil" masterLastPageNumber="2986" masterPageNumber="2967" pageNumber="2971" updateTime="1699069100524" updateUser="plazi" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0">
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<mods:title id="4F1B7A763DA49DFCC0D8D378B5B14DA6">Three new species of Hemienchytraeus (Enchytraeidae, Oligochaeta) from Amazonian forest soil</mods:title>
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<mods:name id="1BD5A855DEF19276791E95F7F7186AE5" type="personal">
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<mods:roleTerm id="ED1498869A43C8C887BAC158632B5D84">Author</mods:roleTerm>
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<mods:namePart id="98D0A694C507052E2B8CAB6348026C7B">Schmelz, R. M.</mods:namePart>
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<mods:roleTerm id="D446B079D26BABACB58BED47E0948418">Author</mods:roleTerm>
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<mods:namePart id="E8B1C0E42D724D5611BCD525FFB9AE90">Römbke, J.</mods:namePart>
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<mods:typeOfResource id="9E18B201834082C646BC3AD27FF3D7C6">text</mods:typeOfResource>
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<mods:title id="DC8A39DDEF55373CB2AAE4BA2147AAFA">Journal of Natural History</mods:title>
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<mods:part id="E46446DF15514E508D4C564A41ED8DCA">
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<mods:date id="3317BC7962C9A441075F2DA29E32047D">2005</mods:date>
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<mods:detail id="96FA47204FD845C6C345C20D77EBC7F4" type="pubDate">
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<mods:number id="59EA341A3E77807E05E0F3A0F2C97F6E">2005-09-27</mods:number>
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</mods:detail>
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<mods:detail id="4F22B2B98688F8D8C9FDE2092960021B" type="volume">
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<mods:number id="C14525355E14C1BD9430FAFE7943D995">39</mods:number>
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<mods:detail id="A7E0F7BBDEFD255459017CF8C35A17AA" type="issue">
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<mods:number id="6AF502607CCEA6605A9EBD024285AE33">32</mods:number>
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<mods:extent id="024F9108367F622A9E7CB8E5E822DBE1" unit="page">
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<mods:url id="ACBDC399521CEA2A3A3695B06050EFAF">http://dx.doi.org/10.1080/00222930500218664</mods:url>
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<mods:classification id="58C7D045D1E5D5EA0C99B97C285A4FEB">journal article</mods:classification>
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<mods:identifier id="E69D8B5677BC635E5D43C274B9D27F5D" type="DOI">10.1080/00222930500218664</mods:identifier>
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<mods:identifier id="0F2AE9D69A0B034923B4CBF4484CF55F" type="ISSN">1464-5262</mods:identifier>
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<mods:identifier id="8E1F00C8B1A65DE2770103990A07ABB4" type="Zenodo-Dep">5220042</mods:identifier>
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<treatment id="03E3594E584D026B6DC1042FA9B7FD39" LSID="urn:lsid:plazi:treatment:03E3594E584D026B6DC1042FA9B7FD39" httpUri="http://treatment.plazi.org/id/03E3594E584D026B6DC1042FA9B7FD39" lastPageId="7" lastPageNumber="2973" pageId="5" pageNumber="2971">
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<subSubSection id="C350BBD3584D02696DC1042FA963FDAB" box="[458,865,541,565]" pageId="5" pageNumber="2971" type="nomenclature">
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<paragraph id="8BF5E858584D02696DC1042FA963FDAB" blockId="5.[458,865,541,598]" box="[458,865,541,565]" pageId="5" pageNumber="2971">
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<heading id="D0BD5F34584D02696DC1042FA963FDAB" box="[458,865,541,565]" centered="true" fontSize="10" level="2" pageId="5" pageNumber="2971" reason="2">
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<taxonomicName id="4C4A93DB584D02696DC1042FA8FEFDAB" ID-CoL="852QS" authority="Schmelz & Römbke, 2005" authorityName="Schmelz & Römbke" authorityYear="2005" box="[458,764,541,565]" class="Clitellata" family="Enchytraeidae" genus="Hemienchytraeus" kingdom="Animalia" order="Enchytraeida" pageId="5" pageNumber="2971" phylum="Annelida" rank="species" species="siljae" status="sp. nov.">
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<emphasis id="B93E344A584D02696DC1042FA8FEFDAB" bold="true" box="[458,764,541,565]" italics="true" pageId="5" pageNumber="2971">Hemienchytraeus siljae</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A20D8931584D02696F0F042FA963FDAB" box="[772,865,541,565]" pageId="5" pageNumber="2971" rank="species">sp. nov.</taxonomicNameLabel>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C350BBD3584D026B6E45040CA9B7FD39" lastPageId="7" lastPageNumber="2973" pageId="5" pageNumber="2971" type="description">
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<paragraph id="8BF5E858584D02696E45040CA8C3FDC8" blockId="5.[458,865,541,598]" box="[590,705,574,598]" pageId="5" pageNumber="2971">
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(
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<figureCitation id="1371F4DD584D02696E5D040CA8B7FDC8" box="[598,693,574,598]" captionStart="Figure 1" captionStartId="4.[125,185,1504,1523]" captionTargetBox="[163,1133,121,1466]" captionTargetId="figure-0@4.[163,1134,120,1468]" captionTargetPageId="4" captionText="Figure 1. Hemienchytraeus siljae sp. nov. (A) Anterior body region of a mature specimen, dorsal view; lateral chaetae to the right omitted; (B) lateral chaeta in IV (left), ventral chaeta, 42nd segment (right); (C) clitellum and bursal slits, ventro-lateral view; (D) male copulatory organ with bursal slit, glandular body and copulatory muscles; (E) sperm funnels with sperm nuclei attached to flared collar, sperm flagella not drawn; (F) sperm funnel, submature specimen; (G) nephridium in IX; (H) nephridium in XV; (I) nephridia in hindmost segments, ventral chaetae and chaetal muscles, lateral view. All figures from whole-mounted specimens: (C–E) drawn from same specimen. Scale bars: 250 mm (A, C–F); 50 mm (B); 100 mm (G–I)." figureDoi="http://doi.org/10.5281/zenodo.5220044" httpUri="https://zenodo.org/record/5220044/files/figure.png" pageId="5" pageNumber="2971">Figure 1</figureCitation>
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)
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</paragraph>
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<paragraph id="8BF5E858584D02696C8104A2AB20FD39" blockId="5.[138,290,655,679]" box="[138,290,655,679]" pageId="5" pageNumber="2971">
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<emphasis id="B93E344A584D02696C8104A2AB20FD39" box="[138,290,655,679]" italics="true" pageId="5" pageNumber="2971">Type material</emphasis>
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</paragraph>
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<paragraph id="8BF5E858584D02696C8104F2AE15FC87" blockId="5.[138,1185,704,793]" pageId="5" pageNumber="2971">
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<materialsCitation id="3B22E205584D02696C8104F2AE1AFD46" box="[138,1048,704,728]" collectionCode="INPA" pageId="5" pageNumber="2971" specimenCode="INPA 121" specimenCount="1" typeStatus="holotype">
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<typeStatus id="54F156FA584D02696C8104F2AAF8FD46" box="[138,250,704,728]" pageId="5" pageNumber="2971" type="holotype">Holotype</typeStatus>
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:
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<specimenCode id="DBEC4023584D02696D0704F2AB8FFD46" box="[268,397,704,728]" collectionCode="INPA" country="Brazil" httpUri="http://grbio.org/cool/rm88-499z" name="Instituto Nacional de Pesquisas da Amazonia" pageId="5" pageNumber="2971">INPA 121</specimenCode>
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, fully mature specimen, stained and whole-mounted
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</materialsCitation>
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.
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<materialsCitation id="3B22E205584D0269682204F2A995FD66" collectionCode="INPA" pageId="5" pageNumber="2971" specimenCode="INPA 122" specimenCount="1" typeStatus="paratype">
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<typeStatus id="54F156FA584D0269682204F2AE9FFD46" box="[1065,1181,704,728]" pageId="5" pageNumber="2971" type="paratype">Paratypes</typeStatus>
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:
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<specimenCode id="DBEC4023584D02696C8104D2AB0AFD66" box="[138,264,736,760]" collectionCode="INPA" country="Brazil" httpUri="http://grbio.org/cool/rm88-499z" name="Instituto Nacional de Pesquisas da Amazonia" pageId="5" pageNumber="2971">INPA 122</specimenCode>
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,
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<specimenCount id="9D4C23D1584D02696D1304D2ABBFFD66" box="[280,445,736,760]" pageId="5" pageNumber="2971" type="generic">one specimen</specimenCount>
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, submature, with severed posterior end
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</materialsCitation>
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;
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<materialsCitation id="3B22E205584D02696FAC04D2AE16FC87" collectingDate="1997-12" collectionCode="ZIM" pageId="5" pageNumber="2971" specimenCode="OL 14281" specimenCount="1" typeStatus="paratype">
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<collectionCode id="ED5B709D584D02696FAC04D2A9DCFD66" box="[935,990,736,760]" country="Slovenia" httpUri="http://grbio.org/cool/3263-rkr7" name="ZIM Culture Collection of Industrial Microorganisms" pageId="5" pageNumber="2971">ZIM</collectionCode>
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<specimenCode id="DBEC4023584D02696FE304D2AE65FD66" box="[1000,1127,736,760]" collectionCode="ZIM" country="Slovenia" httpUri="http://grbio.org/cool/3263-rkr7" name="ZIM Culture Collection of Industrial Microorganisms" pageId="5" pageNumber="2971">OL 14281</specimenCode>
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,
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<specimenCount id="9D4C23D1584D0269687C04D2AAF8FC87" pageId="5" pageNumber="2971" type="generic">one specimen</specimenCount>
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, fully mature. All types from a sample collected in
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<date id="FFF4CE98584D02696F5F0533AE16FC87" box="[852,1044,769,793]" pageId="5" pageNumber="2971" value="1997-12">
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<collectingDate id="EFB03770584D02696F5F0533AE16FC87" box="[852,1044,769,793]" pageId="5" pageNumber="2971" value="1997-12">December 1997</collectingDate>
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</date>
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</materialsCitation>
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.
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</paragraph>
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<paragraph id="8BF5E858584D02696C810568ABC6FCEC" blockId="5.[138,452,858,882]" box="[138,452,858,882]" pageId="5" pageNumber="2971">
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<emphasis id="B93E344A584D02696C810568ABC6FCEC" box="[138,452,858,882]" italics="true" pageId="5" pageNumber="2971">Further material investigated</emphasis>
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</paragraph>
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<paragraph id="8BF5E858584D02696C8105B8A927FC3C" blockId="5.[138,805,906,930]" box="[138,805,906,930]" pageId="5" pageNumber="2971">Three mature specimens, examined alive, not conserved.</paragraph>
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<paragraph id="8BF5E858584D02696C8105D3AB01FC67" blockId="5.[138,259,993,1017]" box="[138,259,993,1017]" pageId="5" pageNumber="2971">
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<emphasis id="B93E344A584D02696C8105D3AB01FC67" box="[138,259,993,1017]" italics="true" pageId="5" pageNumber="2971">Description</emphasis>
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</paragraph>
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<paragraph id="8BF5E858584D02696C810223A85DFA51" blockId="5.[138,1185,1041,1682]" pageId="5" pageNumber="2971">
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Length
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<quantity id="4CB245BD584D02696CEF0223AB5DFBB7" box="[228,351,1041,1065]" metricMagnitude="-2" metricUnit="m" metricValue="1.1" metricValueMax="1.2" metricValueMin="1.0" pageId="5" pageNumber="2971" unit="mm" value="11.0" valueMax="12.0" valueMin="10.0">10–12 mm</quantity>
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(viv) or
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<quantity id="4CB245BD584D02696DC90223A82CFBB7" box="[450,558,1041,1065]" metricMagnitude="-3" metricUnit="m" metricValue="9.5" metricValueMax="11.0" metricValueMin="8.0" pageId="5" pageNumber="2971" unit="mm" value="9.5" valueMax="11.0" valueMin="8.0">8–11 mm</quantity>
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(fix), diameter (fix) ca
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<quantity id="4CB245BD584D02696F300223A994FBB7" box="[827,918,1041,1065]" metricMagnitude="-4" metricUnit="m" metricValue="3.0" pageId="5" pageNumber="2971" unit="mm" value="0.3">0.3 mm</quantity>
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, up to
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<quantity id="4CB245BD584D02696FEE0223AE4EFBB7" box="[997,1100,1041,1065]" metricMagnitude="-4" metricUnit="m" metricValue="3.5" pageId="5" pageNumber="2971" unit="mm" value="0.35">0.35 mm</quantity>
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in XII. Segment number 43–45. Two chaetae per bundle throughout, present laterally in XII also in mature specimens. Chaetae (
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<figureCitation id="1371F4DD584D02696E0F0260A894FBF4" box="[516,662,1106,1130]" captionStart="Figure 1" captionStartId="4.[125,185,1504,1523]" captionTargetBox="[163,1133,121,1466]" captionTargetId="figure-0@4.[163,1134,120,1468]" captionTargetPageId="4" captionText="Figure 1. Hemienchytraeus siljae sp. nov. (A) Anterior body region of a mature specimen, dorsal view; lateral chaetae to the right omitted; (B) lateral chaeta in IV (left), ventral chaeta, 42nd segment (right); (C) clitellum and bursal slits, ventro-lateral view; (D) male copulatory organ with bursal slit, glandular body and copulatory muscles; (E) sperm funnels with sperm nuclei attached to flared collar, sperm flagella not drawn; (F) sperm funnel, submature specimen; (G) nephridium in IX; (H) nephridium in XV; (I) nephridia in hindmost segments, ventral chaetae and chaetal muscles, lateral view. All figures from whole-mounted specimens: (C–E) drawn from same specimen. Scale bars: 250 mm (A, C–F); 50 mm (B); 100 mm (G–I)." figureDoi="http://doi.org/10.5281/zenodo.5220044" httpUri="https://zenodo.org/record/5220044/files/figure.png" pageId="5" pageNumber="2971">Figure 1B, I</figureCitation>
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) slightly sigmoid, without nodulus, pointed distally, blunt proximally; ventral chaetae distinctly larger (longer and thicker) than lateral chaetae; chaetae in posterior segments much larger than chaetae in anterior segments; lateral preclitellar chaetae
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<quantity id="4CB245BD584D02696DB40281A830FB55" box="[447,562,1202,1228]" metricMagnitude="-2" metricUnit="m" metricValue="4.2" metricValueMax="4.7" metricValueMin="3.7" pageId="5" pageNumber="2971" unit="mm" value="42.0" valueMax="47.0" valueMin="37.0">37–47 mm</quantity>
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long,
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<quantity id="4CB245BD584D02696E740281A8D6FB55" box="[639,724,1202,1228]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" metricValueMax="4.0" metricValueMin="3.0" pageId="5" pageNumber="2971" unit="mm" value="3.5" valueMax="4.0" valueMin="3.0">3–4 mm</quantity>
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thick, increasing in posterior segments from 40×
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<quantity id="4CB245BD584D02696D0F02E6AB3CFB72" box="[260,318,1234,1260]" metricMagnitude="-3" metricUnit="m" metricValue="3.0" pageId="5" pageNumber="2971" unit="mm" value="3.0">3 mm</quantity>
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(XIII) up to 85×
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<quantity id="4CB245BD584D02696E1602E6A855FB72" box="[541,599,1234,1260]" metricMagnitude="-3" metricUnit="m" metricValue="7.0" pageId="5" pageNumber="2971" unit="mm" value="7.0">7 mm</quantity>
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in terminal segments; ventral preclitellar chaetae
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<quantity id="4CB245BD584D02696C8102C6AAFFFA92" box="[138,253,1266,1292]" metricMagnitude="-2" metricUnit="m" metricValue="5.25" metricValueMax="6.5" metricValueMin="4.0" pageId="5" pageNumber="2971" unit="mm" value="52.5" valueMax="65.0" valueMin="40.0">40–65 mm</quantity>
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long and
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<quantity id="4CB245BD584D02696D8E02C6ABF3FA92" box="[389,497,1266,1292]" metricMagnitude="-3" metricUnit="m" metricValue="4.35" metricValueMax="4.7" metricValueMin="4.0" pageId="5" pageNumber="2971" unit="mm" value="4.35" valueMax="4.7" valueMin="4.0">4–4.7 mm</quantity>
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thick; gradually increasing in posterior segments up to 90×
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<quantity id="4CB245BD584D02696CC90327AAFEFAB3" box="[194,252,1299,1325]" metricMagnitude="-3" metricUnit="m" metricValue="9.0" pageId="5" pageNumber="2971" unit="mm" value="9.0">9 mm</quantity>
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in terminal segments. Head pore (
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<figureCitation id="1371F4DD584D02696E9B0327A904FAB3" box="[656,774,1301,1325]" captionStart="Figure 1" captionStartId="4.[125,185,1504,1523]" captionTargetBox="[163,1133,121,1466]" captionTargetId="figure-0@4.[163,1134,120,1468]" captionTargetPageId="4" captionText="Figure 1. Hemienchytraeus siljae sp. nov. (A) Anterior body region of a mature specimen, dorsal view; lateral chaetae to the right omitted; (B) lateral chaeta in IV (left), ventral chaeta, 42nd segment (right); (C) clitellum and bursal slits, ventro-lateral view; (D) male copulatory organ with bursal slit, glandular body and copulatory muscles; (E) sperm funnels with sperm nuclei attached to flared collar, sperm flagella not drawn; (F) sperm funnel, submature specimen; (G) nephridium in IX; (H) nephridium in XV; (I) nephridia in hindmost segments, ventral chaetae and chaetal muscles, lateral view. All figures from whole-mounted specimens: (C–E) drawn from same specimen. Scale bars: 250 mm (A, C–F); 50 mm (B); 100 mm (G–I)." figureDoi="http://doi.org/10.5281/zenodo.5220044" httpUri="https://zenodo.org/record/5220044/files/figure.png" pageId="5" pageNumber="2971">Figure 1A</figureCitation>
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) on prostomium in form of a small transverse slit. Epidermal gland cells (
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<figureCitation id="1371F4DD584D02696E510307A8D0FAD3" box="[602,722,1333,1357]" captionStart="Figure 1" captionStartId="4.[125,185,1504,1523]" captionTargetBox="[163,1133,121,1466]" captionTargetId="figure-0@4.[163,1134,120,1468]" captionTargetPageId="4" captionText="Figure 1. Hemienchytraeus siljae sp. nov. (A) Anterior body region of a mature specimen, dorsal view; lateral chaetae to the right omitted; (B) lateral chaeta in IV (left), ventral chaeta, 42nd segment (right); (C) clitellum and bursal slits, ventro-lateral view; (D) male copulatory organ with bursal slit, glandular body and copulatory muscles; (E) sperm funnels with sperm nuclei attached to flared collar, sperm flagella not drawn; (F) sperm funnel, submature specimen; (G) nephridium in IX; (H) nephridium in XV; (I) nephridia in hindmost segments, ventral chaetae and chaetal muscles, lateral view. All figures from whole-mounted specimens: (C–E) drawn from same specimen. Scale bars: 250 mm (A, C–F); 50 mm (B); 100 mm (G–I)." figureDoi="http://doi.org/10.5281/zenodo.5220044" httpUri="https://zenodo.org/record/5220044/files/figure.png" pageId="5" pageNumber="2971">Figure 1A</figureCitation>
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): in preclitellar segments, four to five segmental rows of transversely elongate, quasi-rectangular cells; most cells situated laterally, few cells mid-dorsally and mid-ventrally. First segment and part of prostomium densely covered with cells; in postclitellar segments one to three rows per segment, with fewer cells than in preclitellar segments.
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</paragraph>
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<paragraph id="8BF5E858584D026A6CAE03EAABCEFF5F" blockId="5.[138,1185,1041,1682]" lastBlockId="6.[125,1173,137,1685]" lastPageId="6" lastPageNumber="2972" pageId="5" pageNumber="2971">
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Clitellum (
|
||
<figureCitation id="1371F4DD584D02696D2D03EAAB9EFA6E" box="[294,412,1496,1520]" captionStart="Figure 1" captionStartId="4.[125,185,1504,1523]" captionTargetBox="[163,1133,121,1466]" captionTargetId="figure-0@4.[163,1134,120,1468]" captionTargetPageId="4" captionText="Figure 1. Hemienchytraeus siljae sp. nov. (A) Anterior body region of a mature specimen, dorsal view; lateral chaetae to the right omitted; (B) lateral chaeta in IV (left), ventral chaeta, 42nd segment (right); (C) clitellum and bursal slits, ventro-lateral view; (D) male copulatory organ with bursal slit, glandular body and copulatory muscles; (E) sperm funnels with sperm nuclei attached to flared collar, sperm flagella not drawn; (F) sperm funnel, submature specimen; (G) nephridium in IX; (H) nephridium in XV; (I) nephridia in hindmost segments, ventral chaetae and chaetal muscles, lateral view. All figures from whole-mounted specimens: (C–E) drawn from same specimen. Scale bars: 250 mm (A, C–F); 50 mm (B); 100 mm (G–I)." figureDoi="http://doi.org/10.5281/zenodo.5220044" httpUri="https://zenodo.org/record/5220044/files/figure.png" pageId="5" pageNumber="2971">Figure 1C</figureCitation>
|
||
) in XII–1/2 XIII, posterior end of clitellum immediately before chaetae of XIII dorsally and at level of chaetae of XIII latero-ventrally; clitellum present on all sides except in posterior half of mid-ventral part; cellular arrangement in indistinct rows to reticulate; mid-ventrally (anterior half) in dense rows, irregularly warty between bursal slits. On dorsal body half, hyalocytes and granulocytes present, hyalocytes (diameter
|
||
<quantity id="4CB245BD584D0269687D006BAAD7F90C" metricMagnitude="-2" metricUnit="m" metricValue="1.45" metricValueMax="1.9" metricValueMin="1.0" pageId="5" pageNumber="2971" unit="mm" value="14.5" valueMax="19.0" valueMin="10.0">10– 19 mm</quantity>
|
||
, height
|
||
<quantity id="4CB245BD584D02696D410048AB97F90C" box="[330,405,1656,1682]" metricMagnitude="-2" metricUnit="m" metricValue="1.6" pageId="5" pageNumber="2971" unit="mm" value="16.0">16 mm</quantity>
|
||
, fix) larger than granulocytes (diameter
|
||
<quantity id="4CB245BD584D02696FBA0048AE08F90C" box="[945,1034,1656,1682]" metricMagnitude="-3" metricUnit="m" metricValue="6.5" metricValueMax="8.0" metricValueMin="5.0" pageId="5" pageNumber="2971" unit="mm" value="6.5" valueMax="8.0" valueMin="5.0">5–8 mm</quantity>
|
||
), the latter inconspicuous, occupying interspaces between hyalocytes. Ventro-laterally only granulocytes, here clitellum thinner.
|
||
</paragraph>
|
||
<paragraph id="8BF5E858584E026A6C9306F8AAF2FD39" blockId="6.[125,1173,137,1685]" pageId="6" pageNumber="2972">
|
||
Body wall
|
||
<quantity id="4CB245BD584E026A6D1206F8AB8EFF7C" box="[281,396,200,226]" metricMagnitude="-2" metricUnit="m" metricValue="2.0" metricValueMax="2.5" metricValueMin="1.5" pageId="6" pageNumber="2972" unit="mm" value="20.0" valueMax="25.0" valueMin="15.0">15–25 mm</quantity>
|
||
thick (fix), cuticle
|
||
<quantity id="4CB245BD584E026A6E7D06F8A8E0FF7C" box="[630,738,200,226]" metricMagnitude="-3" metricUnit="m" metricValue="1.25" metricValueMax="1.5" metricValueMin="1.0" pageId="6" pageNumber="2972" unit="mm" value="1.25" valueMax="1.5" valueMin="1.0">1–1.5 mm</quantity>
|
||
thick (fix), always distinct at ×250 magnification, longitudinal muscle layer well developed,
|
||
<quantity id="4CB245BD584E026A6F1A06D8A981FE9C" box="[785,899,232,258]" metricMagnitude="-2" metricUnit="m" metricValue="1.5" metricValueMax="2.0" metricValueMin="1.0" pageId="6" pageNumber="2972" unit="mm" value="15.0" valueMax="20.0" valueMin="10.0">10–20 mm</quantity>
|
||
thick. Preclitellar septa not thickened. Brain (
|
||
<figureCitation id="1371F4DD584E026A6D770739ABF0FEBD" box="[380,498,267,291]" captionStart="Figure 1" captionStartId="4.[125,185,1504,1523]" captionTargetBox="[163,1133,121,1466]" captionTargetId="figure-0@4.[163,1134,120,1468]" captionTargetPageId="4" captionText="Figure 1. Hemienchytraeus siljae sp. nov. (A) Anterior body region of a mature specimen, dorsal view; lateral chaetae to the right omitted; (B) lateral chaeta in IV (left), ventral chaeta, 42nd segment (right); (C) clitellum and bursal slits, ventro-lateral view; (D) male copulatory organ with bursal slit, glandular body and copulatory muscles; (E) sperm funnels with sperm nuclei attached to flared collar, sperm flagella not drawn; (F) sperm funnel, submature specimen; (G) nephridium in IX; (H) nephridium in XV; (I) nephridia in hindmost segments, ventral chaetae and chaetal muscles, lateral view. All figures from whole-mounted specimens: (C–E) drawn from same specimen. Scale bars: 250 mm (A, C–F); 50 mm (B); 100 mm (G–I)." figureDoi="http://doi.org/10.5281/zenodo.5220044" httpUri="https://zenodo.org/record/5220044/files/figure.png" pageId="6" pageNumber="2972">Figure 1A</figureCitation>
|
||
) about as long as wide, or shorter (measured dimensions: 66×
|
||
<quantity id="4CB245BD584E026A6CBE0719AAFCFEDD" box="[181,254,297,323]" metricMagnitude="-2" metricUnit="m" metricValue="6.6" pageId="6" pageNumber="2972" unit="mm" value="66.0">66 mm</quantity>
|
||
and 70×
|
||
<quantity id="4CB245BD584E026A6D730719ABC1FEDD" box="[376,451,297,323]" metricMagnitude="-2" metricUnit="m" metricValue="8.2" pageId="6" pageNumber="2972" unit="mm" value="82.0">82 mm</quantity>
|
||
, fix), deeply incised anteriorly, slightly indented posteriorly, sides parallel. Post-pharyngeal bulbs (
|
||
<figureCitation id="1371F4DD584E026A6E400779A8C0FEFD" box="[587,706,331,355]" captionStart="Figure 1" captionStartId="4.[125,185,1504,1523]" captionTargetBox="[163,1133,121,1466]" captionTargetId="figure-0@4.[163,1134,120,1468]" captionTargetPageId="4" captionText="Figure 1. Hemienchytraeus siljae sp. nov. (A) Anterior body region of a mature specimen, dorsal view; lateral chaetae to the right omitted; (B) lateral chaeta in IV (left), ventral chaeta, 42nd segment (right); (C) clitellum and bursal slits, ventro-lateral view; (D) male copulatory organ with bursal slit, glandular body and copulatory muscles; (E) sperm funnels with sperm nuclei attached to flared collar, sperm flagella not drawn; (F) sperm funnel, submature specimen; (G) nephridium in IX; (H) nephridium in XV; (I) nephridia in hindmost segments, ventral chaetae and chaetal muscles, lateral view. All figures from whole-mounted specimens: (C–E) drawn from same specimen. Scale bars: 250 mm (A, C–F); 50 mm (B); 100 mm (G–I)." figureDoi="http://doi.org/10.5281/zenodo.5220044" httpUri="https://zenodo.org/record/5220044/files/figure.png" pageId="6" pageNumber="2972">Figure 1A</figureCitation>
|
||
, pb) conspicuous. Ventral nerve cord perikarya concentrated in segmental ganglia from V on, i.e. no perikarya in the region of the septa. Oesophageal appendage (
|
||
<figureCitation id="1371F4DD584E026A6DFD07BEA86EFE3A" box="[502,620,396,420]" captionStart="Figure 1" captionStartId="4.[125,185,1504,1523]" captionTargetBox="[163,1133,121,1466]" captionTargetId="figure-0@4.[163,1134,120,1468]" captionTargetPageId="4" captionText="Figure 1. Hemienchytraeus siljae sp. nov. (A) Anterior body region of a mature specimen, dorsal view; lateral chaetae to the right omitted; (B) lateral chaeta in IV (left), ventral chaeta, 42nd segment (right); (C) clitellum and bursal slits, ventro-lateral view; (D) male copulatory organ with bursal slit, glandular body and copulatory muscles; (E) sperm funnels with sperm nuclei attached to flared collar, sperm flagella not drawn; (F) sperm funnel, submature specimen; (G) nephridium in IX; (H) nephridium in XV; (I) nephridia in hindmost segments, ventral chaetae and chaetal muscles, lateral view. All figures from whole-mounted specimens: (C–E) drawn from same specimen. Scale bars: 250 mm (A, C–F); 50 mm (B); 100 mm (G–I)." figureDoi="http://doi.org/10.5281/zenodo.5220044" httpUri="https://zenodo.org/record/5220044/files/figure.png" pageId="6" pageNumber="2972">Figure 1A</figureCitation>
|
||
) with long unpaired root, reaching pharyngeal glands of IV, short primary branches and about four to five elongate secondary branches on each side. Large proximal chamber present. Primary branches thinner than unpaired trunk and thicker than secondary branches. Pharyngeal glands (
|
||
<figureCitation id="1371F4DD584E026A6F3007DCA9B2FD98" box="[827,944,494,518]" captionStart="Figure 1" captionStartId="4.[125,185,1504,1523]" captionTargetBox="[163,1133,121,1466]" captionTargetId="figure-0@4.[163,1134,120,1468]" captionTargetPageId="4" captionText="Figure 1. Hemienchytraeus siljae sp. nov. (A) Anterior body region of a mature specimen, dorsal view; lateral chaetae to the right omitted; (B) lateral chaeta in IV (left), ventral chaeta, 42nd segment (right); (C) clitellum and bursal slits, ventro-lateral view; (D) male copulatory organ with bursal slit, glandular body and copulatory muscles; (E) sperm funnels with sperm nuclei attached to flared collar, sperm flagella not drawn; (F) sperm funnel, submature specimen; (G) nephridium in IX; (H) nephridium in XV; (I) nephridia in hindmost segments, ventral chaetae and chaetal muscles, lateral view. All figures from whole-mounted specimens: (C–E) drawn from same specimen. Scale bars: 250 mm (A, C–F); 50 mm (B); 100 mm (G–I)." figureDoi="http://doi.org/10.5281/zenodo.5220044" httpUri="https://zenodo.org/record/5220044/files/figure.png" pageId="6" pageNumber="2972">Figure 1A</figureCitation>
|
||
) widely connected dorsally in IV, with narrow dorsal connection or separate in V, glands dorsally separate in VI. Three pairs of secondary ventral lobes, in V, VI, and VII, smallest in VII. Afferent fascicles (
|
||
<figureCitation id="1371F4DD584E026A6CE4047DAB64FDF9" box="[239,358,591,615]" captionStart="Figure 1" captionStartId="4.[125,185,1504,1523]" captionTargetBox="[163,1133,121,1466]" captionTargetId="figure-0@4.[163,1134,120,1468]" captionTargetPageId="4" captionText="Figure 1. Hemienchytraeus siljae sp. nov. (A) Anterior body region of a mature specimen, dorsal view; lateral chaetae to the right omitted; (B) lateral chaeta in IV (left), ventral chaeta, 42nd segment (right); (C) clitellum and bursal slits, ventro-lateral view; (D) male copulatory organ with bursal slit, glandular body and copulatory muscles; (E) sperm funnels with sperm nuclei attached to flared collar, sperm flagella not drawn; (F) sperm funnel, submature specimen; (G) nephridium in IX; (H) nephridium in XV; (I) nephridia in hindmost segments, ventral chaetae and chaetal muscles, lateral view. All figures from whole-mounted specimens: (C–E) drawn from same specimen. Scale bars: 250 mm (A, C–F); 50 mm (B); 100 mm (G–I)." figureDoi="http://doi.org/10.5281/zenodo.5220044" httpUri="https://zenodo.org/record/5220044/files/figure.png" pageId="6" pageNumber="2972">Figure 1A</figureCitation>
|
||
, af) comparatively thick. Chloragocytes from V. Dorsal vessel from XIV to XV. Inflated ventral gut epithelium from XXIII to XXX, extending over six or seven segments.
|
||
</paragraph>
|
||
<paragraph id="8BF5E858584E026A6C930482AAD4FB92" blockId="6.[125,1173,137,1685]" pageId="6" pageNumber="2972">
|
||
Nephridia (
|
||
<figureCitation id="1371F4DD584E026A6D2B0482ABDBFD56" box="[288,473,688,712]" captionStart="Figure 1" captionStartId="4.[125,185,1504,1523]" captionTargetBox="[163,1133,121,1466]" captionTargetId="figure-0@4.[163,1134,120,1468]" captionTargetPageId="4" captionText="Figure 1. Hemienchytraeus siljae sp. nov. (A) Anterior body region of a mature specimen, dorsal view; lateral chaetae to the right omitted; (B) lateral chaeta in IV (left), ventral chaeta, 42nd segment (right); (C) clitellum and bursal slits, ventro-lateral view; (D) male copulatory organ with bursal slit, glandular body and copulatory muscles; (E) sperm funnels with sperm nuclei attached to flared collar, sperm flagella not drawn; (F) sperm funnel, submature specimen; (G) nephridium in IX; (H) nephridium in XV; (I) nephridia in hindmost segments, ventral chaetae and chaetal muscles, lateral view. All figures from whole-mounted specimens: (C–E) drawn from same specimen. Scale bars: 250 mm (A, C–F); 50 mm (B); 100 mm (G–I)." figureDoi="http://doi.org/10.5281/zenodo.5220044" httpUri="https://zenodo.org/record/5220044/files/figure.png" pageId="6" pageNumber="2972">Figure 1A, G–I</figureCitation>
|
||
). Preclitellar segments: four pairs, in 6/7–9/10, anteseptale globular, with minute and numerous brownish granules at periphery; funnel orientated obliquely ventrad, with small and narrow anterior projection; postseptale twice as long as anteseptale (total lengths measured
|
||
<quantity id="4CB245BD584E026A6E2F0520A8ABFCB4" box="[548,681,784,810]" metricMagnitude="-1" metricUnit="m" metricValue="1.005" metricValueMax="1.16" metricValueMin="0.85" pageId="6" pageNumber="2972" unit="mm" value="100.5" valueMax="116.0" valueMin="85.0">85–116 mm</quantity>
|
||
, fix), adseptal to medial origin of efferent duct. Postclitellar segments: first pair at 14/15; number of nephridia reduced in mid-body region, slightly larger than preclitellar nephridia (length ca
|
||
<quantity id="4CB245BD584E026A6F520560A9B0FCF4" box="[857,946,849,875]" metricMagnitude="-1" metricUnit="m" metricValue="1.4" pageId="6" pageNumber="2972" unit="mm" value="140.0">140 mm</quantity>
|
||
, fix); nephridia in terminal segments more numerous, small, about half as long (ca
|
||
<quantity id="4CB245BD584E026A6FB70541AE30FC15" box="[956,1074,881,907]" metricMagnitude="-2" metricUnit="m" metricValue="5.25" metricValueMax="5.5" metricValueMin="5.0" pageId="6" pageNumber="2972" unit="mm" value="52.5" valueMax="55.0" valueMin="50.0">50–55 mm</quantity>
|
||
, fix) as preclitellar nephridia; postseptale compressed, bulged dorsad, rise of efferent duct terminal. Coelomocytes flat and broadly oval, with central nucleus and finely granular matrix (viv),
|
||
<quantity id="4CB245BD584E026A6D1E05E6AB8AFC72" box="[277,392,978,1004]" metricMagnitude="-2" metricUnit="m" metricValue="2.25" metricValueMax="3.0" metricValueMin="1.5" pageId="6" pageNumber="2972" unit="mm" value="22.5" valueMax="30.0" valueMin="15.0">15–30 mm</quantity>
|
||
long (fix), large cells 1.5× as long as wide, small cells with circular outline.
|
||
</paragraph>
|
||
<paragraph id="8BF5E858584E026A6C930227A90DF90B" blockId="6.[125,1173,137,1685]" pageId="6" pageNumber="2972">
|
||
Seminal vesicle very small or absent; a small, dense, conspicuous aggregation of developing sperm present around the sperm funnel collars. Sperm funnel (
|
||
<figureCitation id="1371F4DD584E026A6FF80204AE89FBD0" box="[1011,1163,1078,1102]" captionStart="Figure 1" captionStartId="4.[125,185,1504,1523]" captionTargetBox="[163,1133,121,1466]" captionTargetId="figure-0@4.[163,1134,120,1468]" captionTargetPageId="4" captionText="Figure 1. Hemienchytraeus siljae sp. nov. (A) Anterior body region of a mature specimen, dorsal view; lateral chaetae to the right omitted; (B) lateral chaeta in IV (left), ventral chaeta, 42nd segment (right); (C) clitellum and bursal slits, ventro-lateral view; (D) male copulatory organ with bursal slit, glandular body and copulatory muscles; (E) sperm funnels with sperm nuclei attached to flared collar, sperm flagella not drawn; (F) sperm funnel, submature specimen; (G) nephridium in IX; (H) nephridium in XV; (I) nephridia in hindmost segments, ventral chaetae and chaetal muscles, lateral view. All figures from whole-mounted specimens: (C–E) drawn from same specimen. Scale bars: 250 mm (A, C–F); 50 mm (B); 100 mm (G–I)." figureDoi="http://doi.org/10.5281/zenodo.5220044" httpUri="https://zenodo.org/record/5220044/files/figure.png" pageId="6" pageNumber="2972">Figure 1E, F</figureCitation>
|
||
) about as long as body diameter, 4–6× as long as wide (viv); funnel body cone-shaped, gradually and regularly tapering distad (e.g. from 65 to
|
||
<quantity id="4CB245BD584E026A6F010245A957FB11" box="[778,853,1141,1167]" metricMagnitude="-2" metricUnit="m" metricValue="2.0" pageId="6" pageNumber="2972" unit="mm" value="20.0">20 mm</quantity>
|
||
, fix), broadly oval in crosssection, not circular; collar flared, wider than funnel body in mature specimens (up to 1.5× as wide), outline wavy, irregular, like a brim, conspicuous by dense masses of attached spermatozoa. Spermatozoa at least
|
||
<quantity id="4CB245BD584E026A6E2802EAA878FB6E" box="[547,634,1238,1264]" metricMagnitude="-1" metricUnit="m" metricValue="1.25" pageId="6" pageNumber="2972" unit="mm" value="125.0">125 mm</quantity>
|
||
long (viv), head lengths not measured. Vasa deferentia in XII, in loose or tight irregular coils, diameter tapering from 13 proximally to
|
||
<quantity id="4CB245BD584E026A6C76032BAAB5FAAF" box="[125,183,1303,1329]" metricMagnitude="-3" metricUnit="m" metricValue="8.0" pageId="6" pageNumber="2972" unit="mm" value="8.0">8 mm</quantity>
|
||
distally, entering male copulatory organ dorsally. Male copulatory organs with strongly developed musculature. Male glandular body globular, diameter ca
|
||
<quantity id="4CB245BD584E026A6802030BAE53FACF" box="[1033,1105,1335,1361]" metricMagnitude="-2" metricUnit="m" metricValue="5.5" pageId="6" pageNumber="2972" unit="mm" value="55.0">55 mm</quantity>
|
||
(fix). Bursa laterally flattened, short, extending halfway dorsad into glandular body; bursal slit (
|
||
<figureCitation id="1371F4DD584E026A6C8E0348AB2BFA0C" box="[133,297,1402,1426]" captionStart="Figure 1" captionStartId="4.[125,185,1504,1523]" captionTargetBox="[163,1133,121,1466]" captionTargetId="figure-0@4.[163,1134,120,1468]" captionTargetPageId="4" captionText="Figure 1. Hemienchytraeus siljae sp. nov. (A) Anterior body region of a mature specimen, dorsal view; lateral chaetae to the right omitted; (B) lateral chaeta in IV (left), ventral chaeta, 42nd segment (right); (C) clitellum and bursal slits, ventro-lateral view; (D) male copulatory organ with bursal slit, glandular body and copulatory muscles; (E) sperm funnels with sperm nuclei attached to flared collar, sperm flagella not drawn; (F) sperm funnel, submature specimen; (G) nephridium in IX; (H) nephridium in XV; (I) nephridia in hindmost segments, ventral chaetae and chaetal muscles, lateral view. All figures from whole-mounted specimens: (C–E) drawn from same specimen. Scale bars: 250 mm (A, C–F); 50 mm (B); 100 mm (G–I)." figureDoi="http://doi.org/10.5281/zenodo.5220044" httpUri="https://zenodo.org/record/5220044/files/figure.png" pageId="6" pageNumber="2972">Figure 1C, D</figureCitation>
|
||
) longitudinal, slightly bent, tips curved outward. Musculature strongly developed (
|
||
<figureCitation id="1371F4DD584E026A6D0C03A8AB82FA2C" box="[263,384,1434,1458]" captionStart="Figure 1" captionStartId="4.[125,185,1504,1523]" captionTargetBox="[163,1133,121,1466]" captionTargetId="figure-0@4.[163,1134,120,1468]" captionTargetPageId="4" captionText="Figure 1. Hemienchytraeus siljae sp. nov. (A) Anterior body region of a mature specimen, dorsal view; lateral chaetae to the right omitted; (B) lateral chaeta in IV (left), ventral chaeta, 42nd segment (right); (C) clitellum and bursal slits, ventro-lateral view; (D) male copulatory organ with bursal slit, glandular body and copulatory muscles; (E) sperm funnels with sperm nuclei attached to flared collar, sperm flagella not drawn; (F) sperm funnel, submature specimen; (G) nephridium in IX; (H) nephridium in XV; (I) nephridia in hindmost segments, ventral chaetae and chaetal muscles, lateral view. All figures from whole-mounted specimens: (C–E) drawn from same specimen. Scale bars: 250 mm (A, C–F); 50 mm (B); 100 mm (G–I)." figureDoi="http://doi.org/10.5281/zenodo.5220044" httpUri="https://zenodo.org/record/5220044/files/figure.png" pageId="6" pageNumber="2972">Figure 1D</figureCitation>
|
||
), surrounding glandular body and obscuring its outline in living specimens; muscle system complicated, extending over entire segment of XII; dense aggregations of strands anteriorly, posteriorly, laterally and dorso-laterally of glandular bulb; orientation of strands mainly from dorso-lateral to mid-ventral; some strands connecting glandular body and body wall, some strands connecting dorso-lateral and midventral side of body wall in a regular pattern. In
|
||
<specimenCount id="9D4C23D1584E026A6EC3000EA975F9CA" box="[712,887,1596,1620]" pageId="6" pageNumber="2972" type="generic">two specimens</specimenCount>
|
||
, aggregations of hyaline bodies of varying size and with smooth outline interspersed between the muscular strand concentrations. No accessory copulatory glands present.
|
||
</paragraph>
|
||
<paragraph id="8BF5E858584F026B6CAE06BBAE23FDBB" blockId="7.[138,1185,137,549]" pageId="7" pageNumber="2973">
|
||
Spermathecae (
|
||
<figureCitation id="1371F4DD584F026B6D5206BBABCDFF3F" box="[345,463,137,161]" captionStart="Figure 1" captionStartId="4.[125,185,1504,1523]" captionTargetBox="[163,1133,121,1466]" captionTargetId="figure-0@4.[163,1134,120,1468]" captionTargetPageId="4" captionText="Figure 1. Hemienchytraeus siljae sp. nov. (A) Anterior body region of a mature specimen, dorsal view; lateral chaetae to the right omitted; (B) lateral chaeta in IV (left), ventral chaeta, 42nd segment (right); (C) clitellum and bursal slits, ventro-lateral view; (D) male copulatory organ with bursal slit, glandular body and copulatory muscles; (E) sperm funnels with sperm nuclei attached to flared collar, sperm flagella not drawn; (F) sperm funnel, submature specimen; (G) nephridium in IX; (H) nephridium in XV; (I) nephridia in hindmost segments, ventral chaetae and chaetal muscles, lateral view. All figures from whole-mounted specimens: (C–E) drawn from same specimen. Scale bars: 250 mm (A, C–F); 50 mm (B); 100 mm (G–I)." figureDoi="http://doi.org/10.5281/zenodo.5220044" httpUri="https://zenodo.org/record/5220044/files/figure.png" pageId="7" pageNumber="2973">Figure 1A</figureCitation>
|
||
) not attached to oesophagus, extending into VI, VII or VIII, each consisting of ectal duct and ampulla, the latter subdivided into ectal dilatation, connecting tube and ental reservoir. No ectal gland, epidermis thickened in a small circular field (diameter twice the ectal duct diameter) around ectal pore; thickening projecting inside. Ectal pore
|
||
<quantity id="4CB245BD584F026B6D570738AB94FEBC" box="[348,406,264,290]" metricMagnitude="-3" metricUnit="m" metricValue="3.0" pageId="7" pageNumber="2973" unit="mm" value="3.0">3 mm</quantity>
|
||
wide (fix), distalmost stretch of ectal duct canal
|
||
<quantity id="4CB245BD584F026B6FC60738AE20FEBC" box="[973,1058,264,290]" metricMagnitude="-3" metricUnit="m" metricValue="4.5" metricValueMax="5.0" metricValueMin="4.0" pageId="7" pageNumber="2973" unit="mm" value="4.5" valueMax="5.0" valueMin="4.0">4–5 mm</quantity>
|
||
wide (fix), lined with cuticle for about
|
||
<quantity id="4CB245BD584F026B6DEB0719A829FEDD" box="[480,555,297,323]" metricMagnitude="-2" metricUnit="m" metricValue="2.5" pageId="7" pageNumber="2973" unit="mm" value="25.0">25 mm</quantity>
|
||
; further proximally, cuticle light-microscopically not distinguishable; canal often inconspicuous here; ectal duct about one segment length long; ectal dilatation of ampulla
|
||
<quantity id="4CB245BD584F026B6DC90759A837FE1D" box="[450,565,361,387]" metricMagnitude="-2" metricUnit="m" metricValue="2.3" metricValueMax="3.0" metricValueMin="1.6" pageId="7" pageNumber="2973" unit="mm" value="23.0" valueMax="30.0" valueMin="16.0">16–30 mm</quantity>
|
||
wide (fix), ca twice as wide as ectal duct; connecting tube about as wide as ectal duct; ental reservoir thin-walled, usually bent dorsad, of varying length (one or two segments) and width (
|
||
<quantity id="4CB245BD584F026B6E6C079EA8E3FE5A" box="[615,737,426,452]" metricMagnitude="-2" metricUnit="m" metricValue="5.75" metricValueMax="8.0" metricValueMin="3.5" pageId="7" pageNumber="2973" unit="mm" value="57.5" valueMax="80.0" valueMin="35.0">35–80 mm</quantity>
|
||
). Sperm present in ectal dilatation and ental reservoir; in ectal dilatation, sperm arranged side-by-side in a wisp with nuclei orientated distad and flagella extending straight into proximal tube; in ental reservoir, spermatozoa arranged in a dense and irregular coil. One mature egg at a time.
|
||
</paragraph>
|
||
<paragraph id="8BF5E858584F026B6C81046CAAEBFDE8" blockId="7.[138,233,606,630]" box="[138,233,606,630]" pageId="7" pageNumber="2973">
|
||
<emphasis id="B93E344A584F026B6C81046CAAEBFDE8" box="[138,233,606,630]" italics="true" pageId="7" pageNumber="2973">Remarks</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8BF5E858584F026B6C8104BDA9B7FD39" blockId="7.[138,949,655,679]" box="[138,949,655,679]" pageId="7" pageNumber="2973">
|
||
<materialsCitation id="3B22E205584F026B6C8104BDA9B3FD39" box="[138,945,655,679]" collectionCode="VI" pageId="7" pageNumber="2973" specimenCount="1" typeStatus="paratype">
|
||
One
|
||
<typeStatus id="54F156FA584F026B6CCF04BDAB25FD39" box="[196,295,655,679]" pageId="7" pageNumber="2973" type="paratype">paratype</typeStatus>
|
||
specimen has slightly reduced pharyngeal glands in
|
||
<collectionCode id="ED5B709D584F026B6F9B04BDA9B3FD39" box="[912,945,655,679]" country="Norway" name="Mykotektet, National Veterinary Institute" pageId="7" pageNumber="2973">VI</collectionCode>
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |