223 lines
30 KiB
XML
223 lines
30 KiB
XML
<document id="CC8C68C3B2887E1C0EB4B15A053DA956" ID-CLB-Dataset="94308" ID-DOI="10.11646/palaeoentomology.5.3.9" ID-GBIF-Dataset="fee19c7b-f010-45cb-9cd7-c9f70a915654" ID-ISSN="2624-2834" ID-Zenodo-Dep="6820888" ID-ZooBank="838B4FC4-1B1F-4674-B704-FC1231812052" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1657610634805" checkinUser="plazi" docAuthor="AZAR, DANY & MAKSOUD, SIBELLE" docDate="2022" docId="03D787CC053BAC3CFF3BF8DEEB74FA2C" docLanguage="en" docName="palaeoentomology.5.3.9.pdf" docOrigin="Palaeoentomology 5 (3)" docSource="http://dx.doi.org/10.11646/palaeoentomology.5.3.9" docStyle="DocumentStyle:4EE6FD780FF4521D541D0D0600705D0E.4:Palaeoentomology.2018-.journal_article" docStyleId="4EE6FD780FF4521D541D0D0600705D0E" docStyleName="Palaeoentomology.2018-.journal_article" docStyleVersion="4" docTitle="Paleopsychoda jarzembowskii AZAR & MAKSOUD 2022, sp. nov." docType="treatment" docVersion="7" lastPageNumber="264" masterDocId="FFEEFFB4053AAC3EFFB3FF84EF31FFF2" masterDocTitle="Paleopsychoda jarzembowskii, a new Lower Cretaceous species of moth flies from Lebanese amber (Diptera: Psychodidae: Psychodinae)" masterLastPageNumber="268" masterPageNumber="262" pageNumber="263" updateTime="1699358440506" updateUser="ExternalLinkService">
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<mods:title id="B4D9211404D1384B912AA59CA5344075">Paleopsychoda jarzembowskii, a new Lower Cretaceous species of moth flies from Lebanese amber (Diptera: Psychodidae: Psychodinae)</mods:title>
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<mods:namePart id="BCFCCF33D2D8DB01805409E0108C3BD1">AZAR, DANY</mods:namePart>
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<mods:namePart id="B620C5BA37F8DFF5EBF7EDEF3580F6EC">MAKSOUD, SIBELLE</mods:namePart>
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<mods:title id="A9CF153B8424DC68EB8F7E76C9671CA2">Palaeoentomology</mods:title>
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</mods:titleInfo>
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<mods:part id="5028AD2191A3BC7E2CD1E86C6B352923">
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<mods:date id="C31FA7EF3BE536C3B14476C056D8DC40">2022</mods:date>
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<mods:number id="49AE6DDAA67B0D4E082E26458A668BF1">2022-06-29</mods:number>
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<treatment id="03D787CC053BAC3CFF3BF8DEEB74FA2C" ID-DOI="http://doi.org/10.5281/zenodo.6820886" ID-GBIF-Taxon="197225870" ID-Zenodo-Dep="6820886" LSID="urn:lsid:plazi:treatment:03D787CC053BAC3CFF3BF8DEEB74FA2C" httpUri="http://treatment.plazi.org/id/03D787CC053BAC3CFF3BF8DEEB74FA2C" lastPageId="2" lastPageNumber="264" pageId="1" pageNumber="263">
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<subSubSection id="C3646551053BAC3FFF3BF8DEED70F887" box="[136,577,1882,1909]" pageId="1" pageNumber="263" type="nomenclature">
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<paragraph id="8BC136DA053BAC3FFF3BF8DEED70F887" blockId="1.[136,577,1882,1946]" box="[136,577,1882,1909]" pageId="1" pageNumber="263">
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<heading id="D08981B6053BAC3FFF3BF8DEED70F887" bold="true" box="[136,577,1882,1909]" fontSize="10" level="2" pageId="1" pageNumber="263" reason="3">
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<emphasis id="B90AEAC8053BAC3FFF3BF8DEED70F887" bold="true" box="[136,577,1882,1909]" pageId="1" pageNumber="263">
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<taxonomicName id="4C7E4D59053BAC3FFF3BF8DEEED0F887" ID-CoL="9J39N" authority="AZAR & MAKSOUD, 2022" authorityName="AZAR & MAKSOUD" authorityYear="2022" box="[136,481,1882,1909]" class="Insecta" family="Psychodidae" genus="Paleopsychoda" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="263" phylum="Arthropoda" rank="species" species="jarzembowskii" status="sp. nov.">
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<emphasis id="B90AEAC8053BAC3FFF3BF8DEEED0F887" bold="true" box="[136,481,1882,1909]" italics="true" pageId="1" pageNumber="263">Paleopsychoda jarzembowskii</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A23957B3053BAC3FFE5BF8DFED70F887" box="[488,577,1883,1909]" pageId="1" pageNumber="263" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C3646551053BAC3FFF3BF804EFCDF868" box="[136,252,1920,1946]" pageId="1" pageNumber="263" type="description">
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<paragraph id="8BC136DA053BAC3FFF3BF804EFCDF868" blockId="1.[136,577,1882,1946]" box="[136,252,1920,1946]" pageId="1" pageNumber="263">
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(
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<figureCitation id="13452A5F053BAC3FFF23F804EFC2F868" box="[144,243,1920,1946]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStart-3="FIGURE 4" captionStartId-0="2.[136,235,1320,1344]" captionStartId-1="3.[136,235,1859,1883]" captionStartId-2="4.[136,235,1086,1110]" captionStartId-3="4.[136,235,1749,1773]" captionTargetBox-0="[458,1130,174,1297]" captionTargetBox-1="[151,1436,189,1828]" captionTargetBox-2="[344,1244,189,1051]" captionTargetBox-3="[151,1436,1191,1725]" captionTargetId-0="figure-265@2.[458,1130,174,1297]" captionTargetId-1="figure-61@3.[151,1436,189,1835]" captionTargetId-2="figure-111@4.[344,1244,189,1062]" captionTargetId-3="figure-141@4.[151,1436,1191,1725]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionTargetPageId-2="4" captionTargetPageId-3="4" captionText-0="FIGURE 1. Paleopsychoda jarzembowskii sp. nov., male, holotype number 1615A (D. Azar collection). Microphotograph of habitus. Scale bar = 0.5 mm." captionText-1="FIGURE 2. Paleopsychoda jarzembowskii sp. nov., male, holotype number 1615A (D. Azar collection). A, Microphotograph of toothed mouthparts. B, Microphotograph of palp. C, Microphotograph of detail of antenna apex. D, Microphotograph of detail of basal part of anterior side of costal margin. E, Line drawing of antenna. F, Line drawing of palp. Scale bars = 0.1 mm." captionText-2="FIGURE 3. Paleopsychoda jarzembowskii sp. nov., male, holotype number 1615A (D.Azar collection), wing. A, Microphotograph. B, Line drawing. Scale bars = 0.5 mm." captionText-3="FIGURE 4. Paleopsychoda jarzembowskii sp. nov., male, holotype number 1615A (D. Azar collection), genitalia. A, Microphotograph. B, Line drawing. Cox, gonocoxite; Par, paramere; Sty, gonostylus; Sur, surstylus. Scale bars = 0.1 mm." figureDoi-0="http://doi.org/10.5281/zenodo.6820890" figureDoi-1="http://doi.org/10.5281/zenodo.6820892" figureDoi-2="http://doi.org/10.5281/zenodo.6820894" figureDoi-3="http://doi.org/10.5281/zenodo.6820896" httpUri-0="https://zenodo.org/record/6820890/files/figure.png" httpUri-1="https://zenodo.org/record/6820892/files/figure.png" httpUri-2="https://zenodo.org/record/6820894/files/figure.png" httpUri-3="https://zenodo.org/record/6820896/files/figure.png" pageId="1" pageNumber="263">Figs 1–4</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="C3646551053BAC3FFF3BF84DEE6CF7DE" pageId="1" pageNumber="263" type="etymology">
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<paragraph id="8BC136DA053BAC3FFF3BF84DEE6CF7DE" blockId="1.[136,778,1993,2092]" pageId="1" pageNumber="263">
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<emphasis id="B90AEAC8053BAC3FFF3BF84DEE3EF811" bold="true" box="[136,271,1993,2019]" pageId="1" pageNumber="263">Etymology.</emphasis>
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The species epithet is in honor of Prof. Edmund Jarzembowski, a friend and a great authority in palaeoentomology.
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</paragraph>
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</subSubSection>
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<subSubSection id="C3646551053BAC3FFCECFF1AEA2BFC96" pageId="1" pageNumber="263" type="diagnosis">
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<paragraph id="8BC136DA053BAC3FFCECFF1AEBCEFD0A" blockId="1.[810,1452,158,2092]" pageId="1" pageNumber="263">
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<emphasis id="B90AEAC8053BAC3FFCECFF1AECE8FF4A" bold="true" box="[863,985,158,184]" pageId="1" pageNumber="263">Diagnosis.</emphasis>
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Eye-bridge incomplete; antenna with 15 flagellomeres, the last one being reduced; mouthparts toothed, well-developed and of phlebotomine-type; maxillary palp four-segmented, with last palpomere twice as long as others and presenting a constriction in its basal third; anterior sub-basal part of costal vein inflated, from either side of end of humeral vein, allowing as such larger surface, with insertions of a basal wing tuft of setae(specific male character); R1 reaching costa beyond level of radial fork; M3 and M4 [CuA
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<subScript id="17FA349F053BAC3FFB9DFE71EB07FDF1" attach="left" box="[1070,1078,501,515]" fontSize="6" pageId="1" pageNumber="263">1</subScript>
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] meeting basally for a very small distance; CuA [CuA
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<subScript id="17FA349F053BAC3FFBADFD9DEB17FDD5" attach="left" box="[1054,1062,537,551]" fontSize="6" pageId="1" pageNumber="263">2</subScript>
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] well-developed, reaching wing margin beyond half of wing; male genitalia with stylus bearing two apical spines; a specialized long seta strongly curved in its middle part arising between gonocoxite and gonostylus; parameres paired with apically series of four strong setae with incurved apices; surstyli joined basally, bearing apically two strong setae on each.
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</paragraph>
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<paragraph id="8BC136DA053BAC3FFCECFC86EA2BFC96" blockId="1.[810,1452,158,2092]" pageId="1" pageNumber="263">
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<emphasis id="B90AEAC8053BAC3FFCECFC86EB42FCEE" bold="true" box="[863,1139,770,796]" pageId="1" pageNumber="263">Locality and horizon.</emphasis>
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<materialsCitation id="3B163C87053BAC3FFB36FC86EA9AFCB2" ID-GBIF-Occurrence="3862614301" country="Lebanon" county="Caza" location="Mdeyrij-Hammana" pageId="1" pageNumber="263" specimenCount="1" typeStatus="holotype">
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<location id="8EA16001053BAC3FFB36FC86EA6EFCEE" LSID="urn:lsid:plazi:treatment:03D787CC053BAC3CFF3BF8DEEB74FA2C:8EA16001053BAC3FFB36FC86EA6EFCEE" box="[1157,1375,770,796]" country="Lebanon" county="Caza" name="Mdeyrij-Hammana" pageId="1" pageNumber="263">Mdeyrij-Hammana</location>
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,
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<collectingCounty id="62A04E56053BAC3FFAC0FC86EA9AFCEE" box="[1395,1451,770,796]" pageId="1" pageNumber="263">Caza</collectingCounty>
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(= District) Baabda (Central
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<collectingCountry id="F369764A053BAC3FFBC1FCA2EBE8FCB2" box="[1138,1241,806,832]" name="Lebanon" pageId="1" pageNumber="263">Lebanon</collectingCountry>
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),
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<location id="8EA16001053BAC3FFB5EFCA2EA9AFCB2" LSID="urn:lsid:plazi:treatment:03D787CC053BAC3CFF3BF8DEEB74FA2C:8EA16001053BAC3FFB5EFCA2EA9AFCB2" box="[1261,1451,806,832]" country="Lebanon" county="Caza" name="lower Barremian" pageId="1" pageNumber="263">lower Barremian</location>
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</materialsCitation>
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(
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<bibRefCitation id="EFEF4B2B053BAC3FFC82FCCEEB23FC96" author="Granier, B. & Toland, C. & Geze, R. & Azar, D. & Maksoud, S." box="[817,1042,842,869]" pageId="1" pageNumber="263" pagination="247 - 269" refId="ref3174" refString="Granier, B., Toland, C., Geze, R., Azar, D. & Maksoud, S. (2016) Some steps toward a new story for the Jurassic - Cretaceous transition in Mount Lebanon. Carnets de Geologie, 16, 247 - 269. https: // doi. org / 10.4267 / 2042 / 59924" type="journal article" year="2016">
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Granier
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<emphasis id="B90AEAC8053BAC3FFC23FCCFECF0FC96" box="[912,961,842,868]" italics="true" pageId="1" pageNumber="263">et al</emphasis>
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., 2016
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</bibRefCitation>
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;
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<bibRefCitation id="EFEF4B2B053BAC3FFBAFFCCEEA3EFC96" author="Maksoud, S. & Azar, D & Granier, B. & Geze, R." box="[1052,1295,842,869]" pageId="1" pageNumber="263" pagination="331 - 338" refId="ref3441" refString="Maksoud, S., Azar, D, Granier, B. & Geze, R. (2017) New data on the age of the Lower Cretaceous amber outcrops of Lebanon. Palaeoworld, 26 (2), 331 - 338. https: // doi. org / 10.1016 / j. palwor. 2016.03.003" type="journal article" year="2017">
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Maksoud
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<emphasis id="B90AEAC8053BAC3FFB3FFCCFEB8CFC96" box="[1164,1213,842,868]" italics="true" pageId="1" pageNumber="263">et al</emphasis>
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., 2017
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</bibRefCitation>
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).
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</paragraph>
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</subSubSection>
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<subSubSection id="C3646551053BAC3CFCECFCEAEB74FA2C" lastPageId="2" lastPageNumber="264" pageId="1" pageNumber="263" type="description">
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<paragraph id="8BC136DA053BAC3FFCECFCEAEA16FA1E" blockId="1.[810,1452,158,2092]" pageId="1" pageNumber="263">
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<emphasis id="B90AEAC8053BAC3FFCECFCEAECC0FC7A" bold="true" box="[863,1009,878,904]" pageId="1" pageNumber="263">Description.</emphasis>
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Head
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<quantity id="4C869B3F053BAC3FFB8EFCEAEB92FC7B" box="[1085,1187,878,905]" metricMagnitude="-4" metricUnit="m" metricValue="2.7" pageId="1" pageNumber="263" unit="mm" value="0.27">0.27 mm</quantity>
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large; mouthparts well developed, with sharp toothed mandibles and laciniae (
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<figureCitation id="13452A5F053BAC3FFACDFC16EC7EFC22" captionStart="FIGURE 2" captionStartId="3.[136,235,1859,1883]" captionTargetBox="[151,1436,189,1828]" captionTargetId="figure-61@3.[151,1436,189,1835]" captionTargetPageId="3" captionText="FIGURE 2. Paleopsychoda jarzembowskii sp. nov., male, holotype number 1615A (D. Azar collection). A, Microphotograph of toothed mouthparts. B, Microphotograph of palp. C, Microphotograph of detail of antenna apex. D, Microphotograph of detail of basal part of anterior side of costal margin. E, Line drawing of antenna. F, Line drawing of palp. Scale bars = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.6820892" httpUri="https://zenodo.org/record/6820892/files/figure.png" pageId="1" pageNumber="263">Fig. 2A</figureCitation>
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), palp
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<quantity id="4C869B3F053BAC3FFC2EFC32EB33FC23" box="[925,1026,950,977]" metricMagnitude="-4" metricUnit="m" metricValue="2.8" pageId="1" pageNumber="263" unit="mm" value="0.28">0.28 mm</quantity>
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long, with four palpomeres (
|
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<figureCitation id="13452A5F053BAC3FFAFDFC32EC0DFC06" captionStart="FIGURE 2" captionStartId="3.[136,235,1859,1883]" captionTargetBox="[151,1436,189,1828]" captionTargetId="figure-61@3.[151,1436,189,1835]" captionTargetPageId="3" captionText="FIGURE 2. Paleopsychoda jarzembowskii sp. nov., male, holotype number 1615A (D. Azar collection). A, Microphotograph of toothed mouthparts. B, Microphotograph of palp. C, Microphotograph of detail of antenna apex. D, Microphotograph of detail of basal part of anterior side of costal margin. E, Line drawing of antenna. F, Line drawing of palp. Scale bars = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.6820892" httpUri="https://zenodo.org/record/6820892/files/figure.png" pageId="1" pageNumber="263">Fig. 2B, F</figureCitation>
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), with last palpomere almost twice as long as others and presenting a constriction in its basal third, palpomeres I–V lengths respectively 0.052-0.055-
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<quantity id="4C869B3F053BAC3FFB1BFBA6EA5BFBCF" box="[1192,1386,1058,1085]" metricMagnitude="-5" metricUnit="m" metricValue="8.599999999999998" metricValueMax="10.5" metricValueMin="6.7" pageId="1" pageNumber="263" unit="mm" value="0.086" valueMax="0.105" valueMin="0.067">0.067-0.105 mm</quantity>
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; eyes forming an incomplete eye bridge, separated by distance of
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<quantity id="4C869B3F053BAC3FFCF4FBEEEC9DFB77" box="[839,940,1130,1157]" metricMagnitude="-5" metricUnit="m" metricValue="5.0" pageId="1" pageNumber="263" unit="mm" value="0.05">0.05 mm</quantity>
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; antenna
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<quantity id="4C869B3F053BAC3FFBA0FBEEEB47FB77" box="[1043,1142,1130,1157]" metricMagnitude="-4" metricUnit="m" metricValue="8.6" pageId="1" pageNumber="263" unit="mm" value="0.86">0.86 mm</quantity>
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long, with 15 flagellomeres (
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<figureCitation id="13452A5F053BAC3FFC81FB0AECBDFB5A" box="[818,908,1166,1192]" captionStart="FIGURE 2" captionStartId="3.[136,235,1859,1883]" captionTargetBox="[151,1436,189,1828]" captionTargetId="figure-61@3.[151,1436,189,1835]" captionTargetPageId="3" captionText="FIGURE 2. Paleopsychoda jarzembowskii sp. nov., male, holotype number 1615A (D. Azar collection). A, Microphotograph of toothed mouthparts. B, Microphotograph of palp. C, Microphotograph of detail of antenna apex. D, Microphotograph of detail of basal part of anterior side of costal margin. E, Line drawing of antenna. F, Line drawing of palp. Scale bars = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.6820892" httpUri="https://zenodo.org/record/6820892/files/figure.png" pageId="1" pageNumber="263">Fig. 2E</figureCitation>
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); first flagellomere comparatively short, almost as long as following flagellomere; last flagellomere short, almost half of its precedent and drop like (
|
||
<figureCitation id="13452A5F053BAC3FFA8DFB52EAAFFB02" box="[1342,1438,1238,1264]" captionStart="FIGURE 2" captionStartId="3.[136,235,1859,1883]" captionTargetBox="[151,1436,189,1828]" captionTargetId="figure-61@3.[151,1436,189,1835]" captionTargetPageId="3" captionText="FIGURE 2. Paleopsychoda jarzembowskii sp. nov., male, holotype number 1615A (D. Azar collection). A, Microphotograph of toothed mouthparts. B, Microphotograph of palp. C, Microphotograph of detail of antenna apex. D, Microphotograph of detail of basal part of anterior side of costal margin. E, Line drawing of antenna. F, Line drawing of palp. Scale bars = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.6820892" httpUri="https://zenodo.org/record/6820892/files/figure.png" pageId="1" pageNumber="263">Fig. 2C</figureCitation>
|
||
); flagellomere I–XV lengths respectively: 0.064-0.063- 0.059-0.059-0.059-0.056-0.056-0.056-0.050-0.047-0.047- 0.044-0.041-0.041-0.023. Scape nearly cylindrical,
|
||
<quantity id="4C869B3F053BAC3FFADFFAC6EC67FA73" metricMagnitude="-5" metricUnit="m" metricValue="3.5" pageId="1" pageNumber="263" unit="mm" value="0.035">0.035 mm</quantity>
|
||
long and
|
||
<quantity id="4C869B3F053BAC3FFC61FAE2EB79FA73" box="[978,1096,1382,1409]" metricMagnitude="-5" metricUnit="m" metricValue="4.3999999999999995" pageId="1" pageNumber="263" unit="mm" value="0.044">0.044 mm</quantity>
|
||
wide. Pedicel ellipsoid,
|
||
<quantity id="4C869B3F053BAC3FFADFFAE2EC67FA57" metricMagnitude="-5" metricUnit="m" metricValue="6.1" pageId="1" pageNumber="263" unit="mm" value="0.061">0.061 mm</quantity>
|
||
long and
|
||
<quantity id="4C869B3F053BAC3FFC7FFA0EEB71FA57" box="[972,1088,1418,1445]" metricMagnitude="-5" metricUnit="m" metricValue="4.1" pageId="1" pageNumber="263" unit="mm" value="0.041">0.041 mm</quantity>
|
||
wide; all flagellomeres bearing curved setae, two of them distinctly elongate at basal quarter of the flagellomere (perhaps ascoids).
|
||
</paragraph>
|
||
<paragraph id="8BC136DA053BAC3CFCECFA72EFF7F8D8" blockId="1.[810,1452,158,2092]" lastBlockId="2.[136,778,1438,2093]" lastPageId="2" lastPageNumber="264" pageId="1" pageNumber="263">
|
||
Thorax
|
||
<quantity id="4C869B3F053BAC3FFC09FA72EB1EF9E3" box="[954,1071,1526,1553]" metricMagnitude="-4" metricUnit="m" metricValue="3.62" pageId="1" pageNumber="263" unit="mm" value="0.362">0.362 mm</quantity>
|
||
long,
|
||
<quantity id="4C869B3F053BAC3FFBC8FA72EBC1F9E3" box="[1147,1264,1526,1553]" metricMagnitude="-4" metricUnit="m" metricValue="3.94" pageId="1" pageNumber="263" unit="mm" value="0.394">0.394 mm</quantity>
|
||
high. Pronotum strongly gibbous. Legs distinctly longer than entire body. Wings incomplete, nearly
|
||
<quantity id="4C869B3F053BAC3FFB15F9BAEA2CF9AB" box="[1190,1309,1598,1625]" metricMagnitude="-3" metricUnit="m" metricValue="1.17" pageId="1" pageNumber="263" unit="mm" value="1.17">1.170 mm</quantity>
|
||
long,
|
||
<quantity id="4C869B3F053BAC3FFADFF9BAEC67F98F" metricMagnitude="-4" metricUnit="m" metricValue="4.1" pageId="1" pageNumber="263" unit="mm" value="0.41">0.410 mm</quantity>
|
||
wide, hyaline (
|
||
<figureCitation id="13452A5F053BAC3FFBBFF9E6EBBDF98E" box="[1036,1164,1634,1660]" captionStart="FIGURE 3" captionStartId="4.[136,235,1086,1110]" captionTargetBox="[344,1244,189,1051]" captionTargetId="figure-111@4.[344,1244,189,1062]" captionTargetPageId="4" captionText="FIGURE 3. Paleopsychoda jarzembowskii sp. nov., male, holotype number 1615A (D.Azar collection), wing. A, Microphotograph. B, Line drawing. Scale bars = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.6820894" httpUri="https://zenodo.org/record/6820894/files/figure.png" pageId="1" pageNumber="263">Fig. 3A, B</figureCitation>
|
||
). Humeral vein reaching costal margin at
|
||
<quantity id="4C869B3F053BAC3FFC56F902EB66F953" box="[997,1111,1670,1697]" metricMagnitude="-4" metricUnit="m" metricValue="2.28" pageId="1" pageNumber="263" unit="mm" value="0.228">0.228 mm</quantity>
|
||
from wing base; anterior subbasal part of costal vein inflated, from either side of end of humeral vein, allowing as such larger surface, with insertions of a basal wing tuft of setae; subcostal vein (Sc) distally fused with R1,
|
||
<quantity id="4C869B3F053BAC3FFBC1F892EBD6F8C3" box="[1138,1255,1814,1841]" metricMagnitude="-4" metricUnit="m" metricValue="5.31" pageId="1" pageNumber="263" unit="mm" value="0.531">0.531 mm</quantity>
|
||
from wing base, and with crossvein reaching costal margin. R1 reaching costal margin
|
||
<quantity id="4C869B3F053BAC3FFC6FF8DAEB68F88B" box="[988,1113,1886,1913]" metricMagnitude="-4" metricUnit="m" metricValue="9.65" pageId="1" pageNumber="263" unit="mm" value="0.965">0.965 mm</quantity>
|
||
from wing base; Rs fourbranched, with all its branches extending to wing margin. Rs bifurcating into R2+3 and R4+5
|
||
<quantity id="4C869B3F053BAC3FFB78F822EA0EF833" box="[1227,1343,1958,1985]" metricMagnitude="-4" metricUnit="m" metricValue="5.12" pageId="1" pageNumber="263" unit="mm" value="0.512">0.512 mm</quantity>
|
||
distad of wing base. R2 and R3 separating
|
||
<quantity id="4C869B3F053BAC3FFB1FF84EEA2EF817" box="[1196,1311,1994,2021]" metricMagnitude="-4" metricUnit="m" metricValue="8.35" pageId="1" pageNumber="263" unit="mm" value="0.835">0.835 mm</quantity>
|
||
distally; R4 and R5 almost straight; M1 and M2 bifurcating
|
||
<quantity id="4C869B3F053BAC3FFA88F86AEA9DF7FB" box="[1339,1452,2030,2057]" metricMagnitude="-4" metricUnit="m" metricValue="7.75" pageId="1" pageNumber="263" unit="mm" value="0.775">0.775 mm</quantity>
|
||
distally; M1 distally nearly straight; M2 slightly shorter than M1; M3 reaching wing margin at nearly
|
||
<quantity id="4C869B3F0538AC3CFD25FA1AEC3BFA4B" box="[662,778,1438,1465]" metricMagnitude="-4" metricUnit="m" metricValue="9.17" pageId="2" pageNumber="264" unit="mm" value="0.917">0.917 mm</quantity>
|
||
from wing base; M4 [CuA
|
||
<subScript id="17FA349F0538AC3CFE07FA52EE8DFA16" attach="left" box="[436,444,1494,1508]" fontSize="6" pageId="2" pageNumber="264">1</subScript>
|
||
] reaching wing margin
|
||
<quantity id="4C869B3F0538AC3CFD78FA47EF85F9F1" metricMagnitude="-4" metricUnit="m" metricValue="8.219999999999999" pageId="2" pageNumber="264" unit="mm" value="0.822">0.822 mm</quantity>
|
||
distad wing base. CuA [CuA
|
||
<subScript id="17FA349F0538AC3CFE4EFA7FED34F9FB" attach="left" box="[509,517,1531,1545]" fontSize="6" pageId="2" pageNumber="264">2</subScript>
|
||
] (Cu after
|
||
<bibRefCitation id="EFEF4B2B0538AC3CFD31FA6DEFF2F9D5" author="Byers, G. W." pageId="2" pageNumber="264" pagination="497 - 501" refId="ref3021" refString="Byers, G. W. (1989) Homologies in wing venation of primitive Diptera and Mecoptera. Proceedings of the Entomological Society of Washington, 91, 497 - 501." type="journal article" year="1989">
|
||
Byers
|
||
<emphasis id="B90AEAC80538AC3CFD79FA6DEDCDF9F0" box="[714,764,1512,1538]" italics="true" pageId="2" pageNumber="264">et al</emphasis>
|
||
., 1989
|
||
</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFEF4B2B0538AC3CFF7CF989ED76F9D5" author="Krzeminski, W. & Krzeminska, E." box="[207,583,1549,1575]" pageId="2" pageNumber="264" pagination="153 - 184" refId="ref3292" refString="Krzeminski, W. & Krzeminska, E. (2003) Triassic Diptera: Descriptions, revisions and phylogenetic relations. Acta Zoologica Cracoviensia, 46, 153 - 184." type="journal article" year="2003">Krzemiński & Krzemińska, 2003</bibRefCitation>
|
||
) well-developed, sigmoidal, curved distally, reaching wing margin at
|
||
<quantity id="4C869B3F0538AC3CFF3BF9D3EFCAF983" box="[136,251,1623,1649]" metricMagnitude="-4" metricUnit="m" metricValue="6.64" pageId="2" pageNumber="264" unit="mm" value="0.664">0.664 mm</quantity>
|
||
distally; CuP [A] (A1 after
|
||
<bibRefCitation id="EFEF4B2B0538AC3CFD8AF9D3EC34F983" author="Byers, G. W." box="[569,773,1623,1649]" pageId="2" pageNumber="264" pagination="497 - 501" refId="ref3021" refString="Byers, G. W. (1989) Homologies in wing venation of primitive Diptera and Mecoptera. Proceedings of the Entomological Society of Washington, 91, 497 - 501." type="journal article" year="1989">
|
||
Byers
|
||
<emphasis id="B90AEAC80538AC3CFD31F9DCED85F983" box="[642,692,1623,1649]" italics="true" pageId="2" pageNumber="264">et al</emphasis>
|
||
., 1989
|
||
</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFEF4B2B0538AC3CFF3BF9F8ED23F964" author="Krzeminski, W. & Krzeminska, E." box="[136,530,1660,1686]" pageId="2" pageNumber="264" pagination="153 - 184" refId="ref3292" refString="Krzeminski, W. & Krzeminska, E. (2003) Triassic Diptera: Descriptions, revisions and phylogenetic relations. Acta Zoologica Cracoviensia, 46, 153 - 184." type="journal article" year="2003">Krzemiński & Krzemińska, 2003</bibRefCitation>
|
||
) well-developed and reaching posterior margin of wing. All main veins and wing margin bearing long macrotrichiae. Halteres 0.200 mm long; knob fusiform
|
||
<quantity id="4C869B3F0538AC3CFE18F96FED11F8F7" box="[427,544,1771,1798]" metricMagnitude="-4" metricUnit="m" metricValue="1.05" pageId="2" pageNumber="264" unit="mm" value="0.105">0.105 mm</quantity>
|
||
long and
|
||
<quantity id="4C869B3F0538AC3CFD25F96FEC3BF8F7" box="[662,778,1771,1797]" metricMagnitude="-5" metricUnit="m" metricValue="2.6" pageId="2" pageNumber="264" unit="mm" value="0.026">0.026 mm</quantity>
|
||
wide.
|
||
</paragraph>
|
||
<caption id="DF0166520538AC3CFF3BFAACEE9AFA96" ID-DOI="http://doi.org/10.5281/zenodo.6820890" ID-Zenodo-Dep="6820890" httpUri="https://zenodo.org/record/6820890/files/figure.png" pageId="2" pageNumber="264" startId="2.[136,235,1320,1344]" targetBox="[458,1130,174,1297]" targetPageId="2">
|
||
<paragraph id="8BC136DA0538AC3CFF3BFAACEE9AFA96" blockId="2.[136,1451,1320,1381]" pageId="2" pageNumber="264">
|
||
<emphasis id="B90AEAC80538AC3CFF3BFAACEE36FAB2" bold="true" box="[136,263,1320,1344]" pageId="2" pageNumber="264">FIGURE 1.</emphasis>
|
||
<taxonomicName id="4C7E4D590538AC3CFEA2FAADED76FAB3" authority="AZAR & MAKSOUD, 2022" authorityName="AZAR & MAKSOUD" authorityYear="2022" box="[273,583,1321,1345]" class="Insecta" family="Psychodidae" genus="Paleopsychoda" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="264" phylum="Arthropoda" rank="species" species="jarzembowskii" status="sp. nov.">
|
||
<emphasis id="B90AEAC80538AC3CFEA2FAADED76FAB3" box="[273,583,1321,1345]" italics="true" pageId="2" pageNumber="264">Paleopsychoda jarzembowskii</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B90AEAC80538AC3CFDE2FAADED95FAB3" bold="true" box="[593,676,1321,1345]" pageId="2" pageNumber="264">
|
||
<taxonomicNameLabel id="A23957B30538AC3CFDE2FAADED95FAB3" box="[593,676,1321,1345]" pageId="2" pageNumber="264" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, male, holotype number 1615A (D. Azar collection). Microphotograph of habitus. Scale bar = 0.5 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BC136DA0538AC3CFF0EF8B1EB74FA2C" blockId="2.[136,778,1438,2093]" lastBlockId="2.[810,1452,1438,1502]" pageId="2" pageNumber="264">
|
||
Abdomen
|
||
<quantity id="4C869B3F0538AC3CFEF4F8B1EEFCF8BD" box="[327,461,1845,1871]" metricMagnitude="-4" metricUnit="m" metricValue="4.76" pageId="2" pageNumber="264" unit="mm" value="0.476">0.476 mm</quantity>
|
||
long excluding genital appendages, dorsal surfaces of all abdominal segments bearing few setae. Male genital appendages (
|
||
<figureCitation id="13452A5F0538AC3CFD15F8FAEFAAF84C" captionStart="FIGURE 4" captionStartId="4.[136,235,1749,1773]" captionTargetBox="[151,1436,1191,1725]" captionTargetId="figure-141@4.[151,1436,1191,1725]" captionTargetPageId="4" captionText="FIGURE 4. Paleopsychoda jarzembowskii sp. nov., male, holotype number 1615A (D. Azar collection), genitalia. A, Microphotograph. B, Line drawing. Cox, gonocoxite; Par, paramere; Sty, gonostylus; Sur, surstylus. Scale bars = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.6820896" httpUri="https://zenodo.org/record/6820896/files/figure.png" pageId="2" pageNumber="264">Fig. 4A, B</figureCitation>
|
||
) with stylus (
|
||
<quantity id="4C869B3F0538AC3CFE8EF820EE84F84C" box="[317,437,1956,1982]" metricMagnitude="-4" metricUnit="m" metricValue="2.14" pageId="2" pageNumber="264" unit="mm" value="0.214">0.214 mm</quantity>
|
||
) bearing two apical spines; a specialized long seta strongly curved in its middle part arising between coxite (
|
||
<quantity id="4C869B3F0538AC3CFE20F869ED39F7FA" box="[403,520,2029,2056]" metricMagnitude="-4" metricUnit="m" metricValue="2.05" pageId="2" pageNumber="264" unit="mm" value="0.205">0.205 mm</quantity>
|
||
) and stylus; parameres paired with apically series of four strong setae with incurved apices; surstyli joined basally, bearing apically two strong setae on each.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |