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<document id="C02BD9217FCA7D7E1E4A78D8AB66E50E" ID-CLB-Dataset="80831" ID-DOI="10.5281/zenodo.6727957" ID-GBIF-Dataset="a9b14435-eb35-414e-8820-b9eda51268f3" ID-ISBN="978-84-96553-89-7" ID-Zenodo-Dep="6727957" IM.metadata_requiresApprovalFor="plazi" IM.taxonomicNames_requiresApprovalFor="plazi" checkinTime="1656102006799" checkinUser="jonas" docAuthor="Russell A. Mittermeier, Anthony B. Rylands &amp; Don E. Wilson" docDate="2013" docId="03D787BA0E3AFFC3FFE5F732F651C728" docLanguage="en" docName="hbmw_3_Hylobatidae_0778.pdf.imf" docOrigin="Handbook of the Mammals of the World Volume 3 Primates, Barcelona: Lynx Edicions" docTitle="Hylobates klossii" docType="treatment" docVersion="8" lastPageNumber="781" masterDocId="FFEEFFC20E39FFC0FF86FFAFFFB9CB7E" masterDocTitle="Hylobatidae" masterLastPageNumber="791" masterPageNumber="778" pageNumber="781" updateTime="1699350032655" updateUser="ExternalLinkService">
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<mods:title id="9767A38C9BED4F65FEABDB0ACE651731">Hylobatidae</mods:title>
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<mods:namePart id="84627B05796B6CB406D23638BB98D4EC">Russell A. Mittermeier</mods:namePart>
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<mods:namePart id="B242BECC77A0B5E0AF181E83A53E6180">Anthony B. Rylands</mods:namePart>
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<mods:namePart id="95DA885FD302AAB7263F8568852516CD">Don E. Wilson</mods:namePart>
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<subSubSection id="C36465270E3AFFC3FFE5F732FF3AC3B5" box="[99,131,2205,2251]" pageId="3" pageNumber="781" type="multiple">
<paragraph id="8BC136AC0E3AFFC3FFE5F732FF3AC3B5" blockId="3.[96,870,2205,2329]" box="[99,131,2205,2251]" pageId="3" pageNumber="781">
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<figureCitation id="13452A290E3AFFC3FFE5F732FF3AC3B5" box="[99,131,2205,2251]" captionStart="Plate 53: Hylobatidae" captionStartId="1.[100,130,3414,3439]" captionTargetBox="[11,2743,13,3642]" captionTargetPageId="0" captionText="1. Western Hoolock Gibbon (Hoolock hoolock), 2. Eastern Hoolock Gibbon (Hoolock leuconedys), 3. Agile Gibbon (Hylobates agilis), 4. Bornean White-bearded Gibbon (Hylobates albibarbs), 5. Klosss Gibbon (Hylobates klossu), 6. Lar Gibbon (Hylobates lar), 7. Moloch Gibbon (Hylobates moloch), 8. Mullers Gibbon (Hylobates muelleri), 9. Abbott's Gray Gibbon (Hylobates abbotti), 10. East Bornean Gray Gibbon (Hylobates funereus), 11. Pileated Gibbon (Hylobates pileatus)" figureDoi="http://doi.org/10.5281/zenodo.6727999" httpUri="https://zenodo.org/record/6727999/files/figure.png" pageId="3" pageNumber="781">5.</figureCitation>
</heading>
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<paragraph id="8BC136AC0E3AFFC3FF12F732FE7DC3B5" blockId="3.[96,870,2205,2329]" box="[148,452,2205,2251]" pageId="3" pageNumber="781">
<heading id="D08981C00E3AFFC3FF12F732FE7DC3B5" box="[148,452,2205,2251]" pageId="3" pageNumber="781">
<vernacularName id="057D46820E3AFFC3FF12F732FE7DC3B5" box="[148,452,2205,2251]" pageId="3" pageNumber="781">Klosss Gibbon</vernacularName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C36465270E3AFFC3FD8CF732FC91C3B5" box="[522,808,2205,2251]" pageId="3" pageNumber="781" type="nomenclature">
<paragraph id="8BC136AC0E3AFFC3FD8CF732FC91C3B5" blockId="3.[96,870,2205,2329]" box="[522,808,2205,2251]" pageId="3" pageNumber="781">
<heading id="D08981C00E3AFFC3FD8CF732FC91C3B5" box="[522,808,2205,2251]" pageId="3" pageNumber="781">
<taxonomicName id="4C7E4D2F0E3AFFC3FD8CF732FC91C3B5" ID-CoL="3NFRR" baseAuthorityName="G. S. Miller" baseAuthorityYear="1903" box="[522,808,2205,2251]" class="Mammalia" family="Hylobatidae" genus="Hylobates" kingdom="Animalia" order="Primates" pageId="3" pageNumber="781" phylum="Chordata" rank="species" species="klossii">
<emphasis id="B90AEABE0E3AFFC3FD8CF732FC91C3B5" box="[522,808,2205,2251]" italics="true" pageId="3" pageNumber="781">Hylobates klossii</emphasis>
</taxonomicName>
</heading>
</paragraph>
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<subSubSection id="C36465270E3AFFC3FFE4F772FD12C267" pageId="3" pageNumber="781" type="vernacular_names">
<paragraph id="8BC136AC0E3AFFC3FFE4F772FCDCC38C" blockId="3.[96,870,2205,2329]" box="[98,869,2269,2290]" pageId="3" pageNumber="781">
<heading id="D08981C00E3AFFC3FFE4F772FCDCC38C" box="[98,869,2269,2290]" pageId="3" pageNumber="781">
<emphasis id="B90AEABE0E3AFFC3FFE4F772FF16C38C" bold="true" box="[98,175,2269,2290]" pageId="3" pageNumber="781">French:</emphasis>
<vernacularName id="057D46820E3AFFC3FF3EF772FEE3C38C" box="[184,346,2269,2290]" pageId="3" pageNumber="781">Gibbon de Kloss</vernacularName>
/
<emphasis id="B90AEABE0E3AFFC3FEE9F772FE73C38C" bold="true" box="[367,458,2269,2290]" pageId="3" pageNumber="781">German:</emphasis>
<vernacularName id="057D46820E3AFFC3FE52F772FDECC38C" box="[468,597,2269,2290]" pageId="3" pageNumber="781">Kloss-Gibbon</vernacularName>
/
<emphasis id="B90AEABE0E3AFFC3FDEDF772FD7FC38C" bold="true" box="[619,710,2269,2290]" pageId="3" pageNumber="781">Spanish:</emphasis>
<vernacularName id="057D46820E3AFFC3FD56F772FCDCC38C" box="[720,869,2269,2290]" pageId="3" pageNumber="781">Gibon de Kloss</vernacularName>
</heading>
</paragraph>
<paragraph id="8BC136AC0E3AFFC3FFE4F6ABFD12C267" blockId="3.[96,870,2205,2329]" box="[98,683,2308,2329]" pageId="3" pageNumber="781">
<heading id="D08981C00E3AFFC3FFE4F6ABFD12C267" box="[98,683,2308,2329]" pageId="3" pageNumber="781">
<emphasis id="B90AEABE0E3AFFC3FFE4F6ABFEE1C267" bold="true" box="[98,344,2308,2329]" pageId="3" pageNumber="781">Other common names:</emphasis>
<vernacularName id="057D46820E3AFFC3FEE4F6ABFE57C267" box="[354,494,2308,2329]" pageId="3" pageNumber="781">Dwarf Gibbon</vernacularName>
,
<vernacularName id="057D46820E3AFFC3FE7AF6ABFD12C267" box="[508,683,2308,2329]" pageId="3" pageNumber="781">Mentawai Gibbon</vernacularName>
</heading>
</paragraph>
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<subSubSection id="C36465270E3AFFC3FD4CF6E4FCF9C2EE" pageId="3" pageNumber="781" type="reference_group">
<paragraph id="8BC136AC0E3AFFC3FD4CF6E4FCF9C2EE" blockId="3.[713,1300,2379,2802]" pageId="3" pageNumber="781">
<emphasis id="B90AEABE0E3AFFC3FD4CF6E4FCDCC216" bold="true" box="[714,869,2379,2408]" pageId="3" pageNumber="781">Taxonomy.</emphasis>
<taxonomicName id="4C7E4D2F0E3AFFC3FCF5F6E4FC85C2EE" authorityName="G.S.Miller" authorityYear="1903" class="Mammalia" family="Hylobatidae" genus="Symphalangus" kingdom="Animalia" order="Primates" pageId="3" pageNumber="781" phylum="Chordata" rank="species" species="klossii">Symphalangus klossii G. S. Miller, 1903</taxonomicName>
,
</paragraph>
</subSubSection>
<subSubSection id="C36465270E3AFFC3FCC8F6DCFCC3C2C8" pageId="3" pageNumber="781" type="materials_examined">
<paragraph id="8BC136AC0E3AFFC3FCC8F6DCFCC3C2C8" blockId="3.[713,1300,2379,2802]" pageId="3" pageNumber="781">
<materialsCitation id="3B163CF10E3AFFC3FCC8F6DCFCC3C2C8" ID-GBIF-Occurrence="3818819303" pageId="3" pageNumber="781">Indonesia, West Sumatra, South Pagai Island.</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C36465270E3AFFC3FD4CF612FC92C067" pageId="3" pageNumber="781" type="discussion">
<paragraph id="8BC136AC0E3AFFC3FD4CF612FC92C067" blockId="3.[713,1300,2379,2802]" lastBlockId="3.[96,1301,2812,3470]" pageId="3" pageNumber="781">Little to no geographic variation has been observed among populations on differ ent Mentawai islands. In a very small sample of specimens, some variation in the direction of the hair on the forearm was observed. Genetic and vocal data suggest that no differentiation has occurred between the islands, likely due to historic gene flow as recently as 7000 years ago. Monotypic.</paragraph>
</subSubSection>
<subSubSection id="C36465270E3AFFC3FFE6F48CFBC3C016" pageId="3" pageNumber="781" type="distribution">
<caption id="DF0166240E3AFFC3FFE6F48CFBC3C016" ID-DOI="http://doi.org/10.5281/zenodo.6727967" ID-Zenodo-Dep="6727967" httpUri="https://zenodo.org/record/6727967/files/figure.png" inLine="true" pageId="3" pageNumber="781" targetBox="[100,681,2384,2791]" targetPageId="3">
<paragraph id="8BC136AC0E3AFFC3FFE6F48CFBC3C016" blockId="3.[96,1301,2812,3470]" pageId="3" pageNumber="781">
<emphasis id="B90AEABE0E3AFFC3FFE6F48CFEA9C03E" bold="true" box="[96,272,2851,2880]" pageId="3" pageNumber="781">Distribution.</emphasis>
Endemic to the MentawaiIs off the W coast of Sumatra (Siberut, Sipura, North Pagai, South Pagai, and Sinakakislet off the E coast of South Pagai).
</paragraph>
</caption>
</subSubSection>
<subSubSection id="C36465270E3AFFC3FFE6F4DDFB42C78F" pageId="3" pageNumber="781" type="description">
<paragraph id="8BC136AC0E3AFFC3FFE6F4DDFB42C78F" blockId="3.[96,1301,2812,3470]" pageId="3" pageNumber="781">
<emphasis id="B90AEABE0E3AFFC3FFE6F4DDFEE0C0F1" bold="true" box="[96,345,2930,2959]" pageId="3" pageNumber="781">Descriptive notes.</emphasis>
Head-body 43.5-58.5 cm (males) and 42-58 cm (females); weight 5.7-8 kg (males) and 4.4-6.8 kg (females). Female Klosss Gibbonsare slightly smaller than males, but they have canine teeth that are similar in size. Klosss
<taxonomicName id="4C7E4D2F0E3AFFC3FBA4F46AFB32C0A0" box="[1058,1163,3013,3038]" class="Mammalia" family="Hylobatidae" genus="Gibbon" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Primates" pageId="3" pageNumber="781" phylum="Chordata" rank="genus">Gibbon</taxonomicName>
is considered one of the smallest gibbons, although its body size is within the range observed in other species of
<taxonomicName id="4C7E4D2F0E3AFFC3FEECF3A3FE5AC753" authorityName="Illiger" authorityYear="1811" box="[362,483,3084,3117]" class="Mammalia" family="Hylobatidae" genus="Hylobates" kingdom="Animalia" order="Primates" pageId="3" pageNumber="781" phylum="Chordata" rank="genus">Hylobates</taxonomicName>
. Both sexes and all ages have completely black fur with no markings. Klosss
<taxonomicName id="4C7E4D2F0E3AFFC3FED1F39CFE06C72A" box="[343,447,3123,3156]" class="Mammalia" family="Hylobatidae" genus="Gibbon" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Primates" pageId="3" pageNumber="781" phylum="Chordata" rank="genus">Gibbon</taxonomicName>
was originally described as a “dwarf siamang” due to its black fur, but morphologically it more closely resembles other species of
<taxonomicName id="4C7E4D2F0E3AFFC3FB9CF3F5FB2BC705" authorityName="Illiger" authorityYear="1811" box="[1050,1170,3162,3195]" class="Mammalia" family="Hylobatidae" genus="Hylobates" kingdom="Animalia" order="Primates" pageId="3" pageNumber="781" phylum="Chordata" rank="genus">Hylobates</taxonomicName>
than the Siamang (
<taxonomicName id="4C7E4D2F0E3AFFC3FF76F32DFDF7C7DD" baseAuthorityName="Raffles" baseAuthorityYear="1821" box="[240,590,3202,3235]" class="Mammalia" family="Hylobatidae" genus="Symphalangus" kingdom="Animalia" order="Primates" pageId="3" pageNumber="781" phylum="Chordata" rank="species" species="syndactylus">Symphalangus syndactylus</taxonomicName>
). One notable difference is that the fur is sparser than is typical of other species of
<taxonomicName id="4C7E4D2F0E3AFFC3FDAEF306FD18C7B4" authorityName="Illiger" authorityYear="1811" box="[552,673,3241,3274]" class="Mammalia" family="Hylobatidae" genus="Hylobates" kingdom="Animalia" order="Primates" pageId="3" pageNumber="781" phylum="Chordata" rank="genus">Hylobates</taxonomicName>
, and there is a nearly bare area on the throat. In some respects, the skull of the Klosss
<taxonomicName id="4C7E4D2F0E3AFFC3FD1DF37FFCBAC78F" box="[667,771,3280,3313]" class="Mammalia" family="Hylobatidae" genus="Gibbon" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Primates" pageId="3" pageNumber="781" phylum="Chordata" rank="genus">Gibbon</taxonomicName>
also resembles that of the Siamang.
</paragraph>
</subSubSection>
<subSubSection id="C36465270E3AFFC3FFE6F358F6A2CAF1" pageId="3" pageNumber="781" type="biology_ecology">
<paragraph id="8BC136AC0E3AFFC3FFE6F358F6A2CAF1" blockId="3.[96,1301,2812,3470]" lastBlockId="3.[1370,2581,291,3160]" pageId="3" pageNumber="781">
<emphasis id="B90AEABE0E3AFFC3FFE6F358FF76C666" bold="true" box="[96,207,3319,3352]" pageId="3" pageNumber="781">Habitat.</emphasis>
Primary semi-deciduous monsoon and tropical evergreen forest at all elevations. Klosss Gibbons generally prefer to remain in the upper canopy. They have also been observed in swamp forest. Mentawai forests are ever-wet, with annual rainfall of up to 4000 mm. Throughoutits distribution, Klosss
<taxonomicName id="4C7E4D2F0E3AFFC3FCC2F2C2FC15C6F0" box="[836,940,3437,3470]" class="Mammalia" family="Hylobatidae" genus="Gibbon" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Primates" pageId="3" pageNumber="781" phylum="Chordata" rank="genus">Gibbon</taxonomicName>
is sympatric with the Pigtailed Langur (Simias
<taxonomicName id="4C7E4D2F0E3AFFC3F909FE8CF943CA3E" baseAuthorityName="Harlan" baseAuthorityYear="1826" box="[1679,1786,291,320]" class="Mammalia" family="Hylobatidae" genus="Hylobates" kingdom="Animalia" order="Primates" pageId="3" pageNumber="781" phylum="Chordata" rank="species" species="concolor">concolor</taxonomicName>
), and the Siberut Langur (Presbytis siberu) and the Siberut Macaque (Macaca siberu) on Siberut and the Mentawai Langur (P. potenziani) and the Pagai Macaque (Macaca pagensis) on the southern Mentawai Islands.
</paragraph>
</subSubSection>
<subSubSection id="C36465270E3AFFC3FAD9FE39F960C92D" pageId="3" pageNumber="781" type="food_feeding">
<paragraph id="8BC136AC0E3AFFC3FAD9FE39F960C92D" blockId="3.[1370,2581,291,3160]" pageId="3" pageNumber="781">
<emphasis id="B90AEABE0E3AFFC3FAD9FE39F9D4CAC9" bold="true" box="[1375,1645,406,439]" pageId="3" pageNumber="781">Food and Feeding.</emphasis>
Klosss Gibbons are predominantly frugivorous. Fruits make up c.73% of their feeding time, arthropods and other small animals ¢.25% (their primary source of protein), and leaves c.2% (less than is typical of other gibbons). Klosss
<taxonomicName id="4C7E4D2F0E3AFFC3F65FFE47FA2CC953" class="Mammalia" family="Hylobatidae" genus="Gibbon" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Primates" pageId="3" pageNumber="781" phylum="Chordata" rank="genus">Gibbon</taxonomicName>
is dominant to the Siberut and Mentawai surelis, always displacing them from food trees when they meet.
</paragraph>
</subSubSection>
<subSubSection id="C36465270E3AFFC3FAD8FDF4F9A6C819" pageId="3" pageNumber="781" type="breeding">
<paragraph id="8BC136AC0E3AFFC3FAD8FDF4F9A6C819" blockId="3.[1370,2581,291,3160]" pageId="3" pageNumber="781">
<emphasis id="B90AEABE0E3AFFC3FAD8FDF4FA5FC902" bold="true" box="[1374,1510,603,636]" pageId="3" pageNumber="781">Breeding.</emphasis>
Klosss
<taxonomicName id="4C7E4D2F0E3AFFC3F9DEFDF4F978C902" box="[1624,1729,603,636]" class="Mammalia" family="Hylobatidae" genus="Gibbon" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Primates" pageId="3" pageNumber="781" phylum="Chordata" rank="genus">Gibbon</taxonomicName>
appears to breed throughout the year. Single births are the norm. Age classification is the same as for other gibbons: infants up to two years old are dependent on the mother;juveniles, age 2—4 years, are independent; adolescents, age 4-6 years, display increased aggression toward same-sex parents; the subadult stage begins when about six years old; and adult independence is typically achieved at about eight years old. Adult males do not carry offspring. Individuals may live more than 30 years.
</paragraph>
</subSubSection>
<subSubSection id="C36465270E3AFFC3FADAFCC1F642CD7B" pageId="3" pageNumber="781" type="activity">
<paragraph id="8BC136AC0E3AFFC3FADAFCC1F642CD7B" blockId="3.[1370,2581,291,3160]" pageId="3" pageNumber="781">
<emphasis id="B90AEABE0E3AFFC3FADAFCC1F9FBC8F1" bold="true" box="[1372,1602,878,911]" pageId="3" pageNumber="781">Activity patterns.</emphasis>
Klosss Gibbons are diurnal and arboreal. Groups spend much of their day at rest. They do not use the same sleeping tree two nights in a row and choose sleeping sites that do not have biting ants. They also prefer sleeping trees that lack hanging lianas, which may be an adaptation to avoid human predation because Mentawai hunters often climb lianas to shoot primates. Unlike most other gibbons, Klosss Gibbons do not sing duets. Instead, adult males sing from sleeping trees in the hours before dawn, sometimesas early as 01:00 h. Adult females typically chorus after the first feeding bout of the day (07:00-09:00 h). Singing appears to be contagious, and male bouts typically occur in “countersinging” interactions, in which adjacent individuals take turns singing and avoid overlapping. Neighboring females may produce songs in synchrony. The greatcall of the female Klosss
<taxonomicName id="4C7E4D2F0E3AFFC3F9FDFB57F95BCE67" box="[1659,1762,1272,1305]" class="Mammalia" family="Hylobatidae" genus="Gibbon" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Primates" pageId="3" pageNumber="781" phylum="Chordata" rank="genus">Gibbon</taxonomicName>
has been described as “the finest music uttered by any land mammal,” and itis often accompanied by an acrobatic display. Females exhibit structural individuality in their songs, such that each female can be distinguished by their song alone. The males song is simpler than the females. As in other gibbons, the primary function of the female's song is to defend the territory and possibly to advertise mated status. Solitary, “floating” male Klosss Gibbons that do not occupy a territory have been observed to sing, suggesting that mate attraction may be a function of the males song.
</paragraph>
</subSubSection>
<subSubSection id="C36465270E3AFFC3FADBF9A3F6F1C303" pageId="3" pageNumber="781" type="biology_ecology">
<paragraph id="8BC136AC0E3AFFC3FADBF9A3F6F1C303" blockId="3.[1370,2581,291,3160]" pageId="3" pageNumber="781">
<emphasis id="B90AEABE0E3AFFC3FADBF9A3F7ABCD53" bold="true" box="[1373,2066,1548,1581]" pageId="3" pageNumber="781">Movements, Home range and Social organization.</emphasis>
Home ranges of Klosss Gibbons are 7-32 ha, probably varying in relation to habitat quality and floristic composition. Average daily movement is 1514 m. Group size is typically 4-6 individuals. Groups as large as 10-15 individuals have been observed on North Pagai and Siberut, butit is unclearif these are stable units. In general, males are aggressive toward other males and defend their territories from predators (humans), while females tend to lead groups movements during the day, and also exclude other adult females from the group. At about eight years old, Klosss Gibbons find a mate and establish their own territory. Young adult Klosss Gibbons may establish territories adjacent to their parents territories, with their parents assistance. They may also replace a deceased same-sex adult in their parental territory to live with the remaining opposite-sex adult; in the observed cases,it was not known whether that surviving adult was the genetic parent of the young adult. Disturbance levels in the habitats of Klosss Gibbons vary on the different islands, but recent surveys have detected similar densities in unlogged forests, forests logged ten years ago, and those logged 20 years ago. The highest densities have been observed in northern Siberut. Density in the Mentawai islands averages 12 ind/km?,
</paragraph>
</subSubSection>
<subSubSection id="C36465270E3AFFC3FADAF72BF648C0C6" pageId="3" pageNumber="781" type="conservation">
<paragraph id="8BC136AC0E3AFFC3FADAF72BF648C0C6" blockId="3.[1370,2581,291,3160]" pageId="3" pageNumber="781">
<emphasis id="B90AEABE0E3AFFC3FADAF72BF907C3DB" bold="true" box="[1372,1726,2180,2213]" pageId="3" pageNumber="781">Status and Conservation.</emphasis>
CITES Appendix I. Classified as Endangered on The IUCN Red List. Klosss
<taxonomicName id="4C7E4D2F0E3AFFC3F9C6F704F910C3B2" box="[1600,1705,2219,2252]" class="Mammalia" family="Hylobatidae" genus="Gibbon" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Primates" pageId="3" pageNumber="781" phylum="Chordata" rank="genus">Gibbon</taxonomicName>
is protected under Indonesian law, but it occurs in only one protected area, Siberut National Park. The total population, based on limited survey effort using auditory sampling, is 20,000-25,000 individuals, with island totals of 18,210-21,345 for Siberut, 753-880 for Sipora, and 1680-1974 on North and South Pagai together (another estimate for these last two islands, using a transect-based approach, was 2029 individuals). The vast majority of the global population is believed to be in Siberut National Park, with 13,190-15,413 individuals, but this estimate is based on a survey of less than 0-1% of the park and therefore needs to be viewed as preliminary. Total population decline of Klosss
<taxonomicName id="4C7E4D2F0E3AFFC3F841F649F789C179" box="[1991,2096,2534,2567]" class="Mammalia" family="Hylobatidae" genus="Gibbon" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Primates" pageId="3" pageNumber="781" phylum="Chordata" rank="genus">Gibbon</taxonomicName>
may be ¢.50% since 1980. Threats include hunting, commercial logging, forest conversion for agroindustry, especially oil palm, and forest clearance and disturbance at the local scale. Selective logging is widespread across the distribution of Klosss
<taxonomicName id="4C7E4D2F0E3AFFC3F851F5F3F7FBC103" box="[2007,2114,2652,2685]" class="Mammalia" family="Hylobatidae" genus="Gibbon" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Primates" pageId="3" pageNumber="781" phylum="Chordata" rank="genus">Gibbon</taxonomicName>
, but one study suggests that this may have limited impact on carrying capacity. Hunting pressure has been exacerbated by increased access to forested areas through logging infrastructure and a move away from traditional hunting methods in the use of guns. Infant Klosss Gibbons are often taken as pets by hunters after they kill a parent. Traded Klosss Gibbons mostly remain in the Mentawai islands but sometimes go to neighboring western Sumatra. Increased legal protection of the remaining populations, including those in the Peleonan Forest, increased commitment and planning for conservation from logging companies, and improved enforcement in Siberut National Park are key conservation interventions.
</paragraph>
</subSubSection>
<subSubSection id="C36465270E3AFFC3FADDF468F651C728" pageId="3" pageNumber="781" type="bibRefCitation_list">
<paragraph id="8BC136AC0E3AFFC3FADDF468F651C728" blockId="3.[1370,2581,291,3160]" pageId="3" pageNumber="781">
<emphasis id="B90AEABE0E3AFFC3FADDF468FA49C09E" bold="true" box="[1371,1520,3015,3040]" pageId="3" pageNumber="781">Bibliography.</emphasis>
Chasen &amp; Kloss (1927), Chivers (2001), Groves (2001), Haimoff &amp; Tilson (1985), Keith et al. (2009), Marshall &amp; Marshall (1976), Miller (1903), Nijman (2005b), Paciulli (2004), Schultz (1932), Tenaza (1974, 1975, 1976, 1987), Tenaza &amp; Hamilton (1971), Tenaza &amp; Tilson (1977, 1985), Tilson (1980, 1981), Tilson &amp; Tenaza (1982), Whittaker (2005, 2006, 2009), Whittaker &amp; Geissmann (2008), Whitten (1982a, 1982b, 1982c, 1982d, 1982¢, 1984).
</paragraph>
</subSubSection>
</treatment>
</document>