787 lines
128 KiB
XML
787 lines
128 KiB
XML
<document id="3327BCA2AD821FFDC44CE6EB3CAB18DE" ID-DOI="10.11646/zootaxa.4019.1.10" ID-GBIF-Dataset="52ee815e-9e7e-40c3-b900-e0e03f07c8ae" ID-ISSN="1175-5326" ID-Zenodo-Dep="245139" ID-ZooBank="28811FA6-340B-42AE-9D3A-12BA3BFC7EF6" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461260114132" checkinUser="plazi" docAuthor="Capa, María, Faroni-Perez, Larisse & Hutchings, Pat" docDate="2015" docId="03D787B1FFFC2C5AFF22AD21FE1AFC07" docLanguage="en" docName="zt04019p206.pdf" docOrigin="Zootaxa 4019 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Lygdamis nasutus Capa, Faroni-Perez & Hutchings, 2015, n. sp." docType="treatment" docVersion="11" lastPageNumber="198" masterDocId="FFEEFFC9FFF82C54FFB5AA7DFFE5FF82" masterDocTitle="Sabellariidae from Lizard Island, Great Barrier Reef, including a new species of Lygdamis and notes on external morphology of the median organ" masterLastPageNumber="206" masterPageNumber="184" pageNumber="188" updateTime="1698609282051" updateUser="plazi">
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<mods:mods id="B99FB9F716780ACC2B3C4BC9554F4678" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="FCC92A180B9BCAE9973FEFCE67FAF16B">
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<mods:title id="561F68469E35E1724418F75116A0264A">Sabellariidae from Lizard Island, Great Barrier Reef, including a new species of Lygdamis and notes on external morphology of the median organ</mods:title>
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<mods:name id="8507B7426CB90F4ADABDAAF2196E6551" type="personal">
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<mods:namePart id="6A50EB1E1AD5C87382C032046CC9C31B">Capa, María</mods:namePart>
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<mods:namePart id="A24AC76EA79CDE8FDBF5E5DE9979C441">Faroni-Perez, Larisse</mods:namePart>
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<mods:roleTerm id="1A0D402D831F87A5157A130F0D15ECE1">Author</mods:roleTerm>
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<mods:namePart id="729AABD1F5125568648B8126C0C2FA9F">Hutchings, Pat</mods:namePart>
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<mods:title id="2931A6C1236E21CA7391D92EDF346779">Zootaxa</mods:title>
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<mods:part id="4AA35C16805C76F8639DAD479F779C1F">
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<mods:date id="305AE2F99653C8C1306DDC1EF3A468DD">2015</mods:date>
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<mods:number id="CFC8C9C4C41738A701569D9428F31683">4019</mods:number>
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<mods:classification id="2C6574BF1759543779D15989934C2202">journal article</mods:classification>
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<mods:identifier id="32B2D13B8AAB8333A6F087FB8115BA2D" type="DOI">10.11646/zootaxa.4019.1.10</mods:identifier>
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<treatment id="03D787B1FFFC2C5AFF22AD21FE1AFC07" ID-DOI="http://doi.org/10.5281/zenodo.6109689" ID-GBIF-Taxon="119636838" ID-Zenodo-Dep="6109689" LSID="urn:lsid:plazi:treatment:03D787B1FFFC2C5AFF22AD21FE1AFC07" httpUri="http://treatment.plazi.org/id/03D787B1FFFC2C5AFF22AD21FE1AFC07" lastPageId="14" lastPageNumber="198" pageId="4" pageNumber="188">
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<subSubSection id="C364652CFFFC2C50FF22AD21FEEEF815" pageId="4" pageNumber="188" type="nomenclature">
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<paragraph id="8BC136A7FFFC2C50FF22AD21FE5FF8F4" blockId="4.[151,442,1884,1943]" box="[151,442,1884,1910]" pageId="4" pageNumber="188">
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<heading id="D08981CBFFFC2C50FF22AD21FE5FF8F4" bold="true" box="[151,442,1884,1910]" fontSize="11" level="1" pageId="4" pageNumber="188" reason="1">
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<emphasis id="B90AEAB5FFFC2C50FF22AD21FE5FF8F4" bold="true" box="[151,442,1884,1910]" pageId="4" pageNumber="188">
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<taxonomicName id="4C7E4D24FFFC2C50FF22AD21FE95F8F4" box="[151,368,1884,1910]" class="Polychaeta" family="Sabellariidae" genus="Lygdamis" kingdom="Animalia" order="Sabellida" pageId="4" pageNumber="188" phylum="Annelida" rank="species" species="nasutus" status="sp. nov.">
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||
<emphasis id="B90AEAB5FFFC2C50FF22AD21FE95F8F4" bold="true" box="[151,368,1884,1910]" italics="true" pageId="4" pageNumber="188">Lygdamis nasutus</emphasis>
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||
</taxonomicName>
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||
<taxonomicNameLabel id="A23957CEFFFC2C50FECDAD21FE5FF8F4" box="[376,442,1884,1910]" pageId="4" pageNumber="188" rank="species">n. sp.</taxonomicNameLabel>
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</emphasis>
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||
</heading>
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</paragraph>
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||
<paragraph id="8BC136A7FFFC2C50FF22AD03FEEEF815" blockId="4.[151,442,1884,1943]" box="[151,267,1918,1943]" pageId="4" pageNumber="188">
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(
|
||
<figureCitation id="13452A22FFFC2C50FF2AAD03FEE7F815" box="[159,258,1918,1943]" captionStart-0="FIGURE 2" captionStart-1="FIGURE 3. A – D" captionStart-2="FIGURE 4" captionStart-3="FIGURE 5" captionStart-4="FIGURE 6" captionStart-5="FIGURE 7" captionStartId-0="5.[151,250,1499,1521]" captionStartId-1="6.[151,250,1449,1471]" captionStartId-2="7.[151,250,1902,1924]" captionStartId-3="8.[151,250,1820,1842]" captionStartId-4="9.[151,250,1737,1759]" captionStartId-5="10.[151,250,1877,1899]" captionTargetBox-0="[153,1422,188,1471]" captionTargetBox-1="[154,1433,195,1423]" captionTargetBox-2="[153,1431,412,1878]" captionTargetBox-3="[191,1407,198,1798]" captionTargetBox-4="[153,1435,341,1713]" captionTargetBox-5="[183,1410,288,1860]" captionTargetId-0="figure@5.[151,1424,186,1474]" captionTargetId-1="figure@6.[151,1436,193,1428]" captionTargetId-2="figure@7.[151,1436,409,1881]" captionTargetId-3="figure@8.[187,1408,193,1805]" captionTargetId-4="figure@9.[150,1437,337,1716]" captionTargetId-5="figure@10.[178,1412,283,1861]" captionTargetPageId-0="5" captionTargetPageId-1="6" captionTargetPageId-2="7" captionTargetPageId-3="8" captionTargetPageId-4="9" captionTargetPageId-5="10" captionText-0="FIGURE 2. Lygdamis nasutus n. sp., paratype, AM W. 47750, live. A. Whole specimen, dorsal view, arrowheads point to notopodial cirri; B. Anterior end, lateral view, with median organ protruding from operculum (arrow); C. Anterior end, dorsal view; D. Operculum, frontal view; E. Posterior abdominal chaetigers and cauda, ventral view. Photos by A. Semenov." captionText-1="FIGURE 3. A – D. Lygdamis nasutus n. sp., holotype, AM W. 47372, preserved. A. Operculum, thoracic and parathoracic chaetigers, lateral view; arrowheads point to notopodial cirri; B. Anterior end, ventral view; C. Same, dorsal view; D. Same, but with exposed well-developed median organ. E – F Paratype AM W. 44467. E. Detail of median organ, dorsal view; F. Cross section of paratypes, showing half of the operculum with whole median organ. Abbreviations: br = branchia, bo = building organ, cn 1 = cirrus neuropodia segment 1, chn 1 = chaetae neuropodium segment 1, dc = dorsal cirrus, es = eyespots, ip = inner paleae, li = lips, m = mouth, mo = median organ, nh = nuchal hook, op = outer paleae, opa = opercular papillae, pa = palp, rtf = row of tentacular filaments, tf = tentacular filaments. Small arrow indicates conical dorsal cirrus. Photo: A – D—E. Wong." captionText-2="FIGURE 4. Median organ, Lygdamis nasutus n. sp., paratypes, AM W. 44467. A. Lateral view with a close up of the eyespots along its length; B. Details of the distal tip, showing the flat corona; C. Distal end, showing teardrop coronal shape frontal view. D – G. SEM images, AM W. 47750. D. Complete median organ, lateral view; E. Distal end, lateral view; F. Detail of the base of median organ with a few sparsely distributed ciliary tufts; G. Close up of ciliary tufts." captionText-3="FIGURE 5. Chaetae, Lygdamis nasutus n. sp., paratypes. A. Outer paleae; B. Detail of tips of outer paleae; C. Inner paleae, distal end; D. Notopodium, parathoracic chaetiger; E. Detail of tips of lanceolate parathoracic notopodial chaeta; F. Parathoracic notopodial simple capillaries, interspersed with lanceolate chaetae; G. Neuropodium, parathoracic chaetiger; H. Detail of tips of lanceolate parathoracic neuropodial chaetae; I. Parathoracic neuropodial simple capillaries, interspersed with lanceolate chaetae; J. Abdominal notochaetae (two types); K. Neuropodial abdominal uncini; L. Detail of abdominal uncinus, side view." captionText-4="FIGURE 6. Chaetae, Lygdamis nasutus n. sp., paratypes, SEM images, AM W. 44467. A. Operculum, lateral view; B. Outer paleae; C. Thecal sculpture, outer paleae; D. Inner paleae; E. Close up of distal end, inner paleae; F. First thoracic neuropodial chaetae; G. Parathoracic notopodia; H. Parathoracic neuropodium. Abbreviations: cn 1 = cirrus neuropodia segment 1, chn 1 = chaetae neuropodium segment 1, ip = inner paleae, mo = median organ, nh = nuchal hook, op = outer paleae, opa = opercular papillae, pa = palp." captionText-5="FIGURE 7. Chaetae, Lygdamis nasutus n. sp., paratypes, SEM images, AM W. 44467. A. Anterior abdominal uncini with number of rows of teeth provided in one uncinus, close up of distal dentition; B. Detached uncini, with number of rows of teeth provided in one uncinus; C. Posterior abdominal uncini; D. Detail of posterior abdominal uncini; E. Abdominal notopodium; F. Detail of the two types of abdominal notochaetae; G. Posterior abdominal chaetiger and cauda." httpUri-0="https://zenodo.org/record/245141/files/figure.png" httpUri-1="https://zenodo.org/record/245142/files/figure.png" httpUri-2="https://zenodo.org/record/245143/files/figure.png" httpUri-3="https://zenodo.org/record/245144/files/figure.png" httpUri-4="https://zenodo.org/record/245145/files/figure.png" httpUri-5="https://zenodo.org/record/245146/files/figure.png" pageId="4" pageNumber="188">Figs 2–7</figureCitation>
|
||
)
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C364652CFFFC2C51FF22ADBDFC7DF92C" lastPageId="5" lastPageNumber="189" pageId="4" pageNumber="188" type="materials_examined">
|
||
<paragraph id="8BC136A7FFFC2C50FF22ADBDFB1CF876" blockId="4.[151,1436,1984,2036]" pageId="4" pageNumber="188">
|
||
<emphasis id="B90AEAB5FFFC2C50FF22ADBDFED6F854" bold="true" box="[151,307,1984,2006]" pageId="4" pageNumber="188">
|
||
<typeStatus id="54C58805FFFC2C50FF22ADBDFF29F854" box="[151,204,1984,2006]" pageId="4" pageNumber="188">Type</typeStatus>
|
||
material.
|
||
</emphasis>
|
||
<typeStatus id="54C58805FFFC2C50FE8CADBDFE7EF854" box="[313,411,1984,2006]" pageId="4" pageNumber="188" type="holotype">Holotype</typeStatus>
|
||
: AM W.47372, Lizard Island, North Point, off Mermaid Cove,
|
||
<geoCoordinate id="EE4A5060FFFC2C50FBA2ADBDFB8DF854" box="[1047,1128,1984,2006]" direction="south" orientation="latitude" pageId="4" pageNumber="188" precision="925" value="-14.65">14°39'S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE4A5060FFFC2C50FBC6ADBDFB37F854" box="[1139,1234,1984,2006]" direction="east" orientation="longitude" pageId="4" pageNumber="188" precision="925" value="145.45">145°27'E</geoCoordinate>
|
||
, coral rubble,
|
||
<specimenCount id="9D78FD2EFFFC2C50FAD7ADBDFA79F854" box="[1378,1436,1984,2006]" pageId="4" pageNumber="188" type="generic">
|
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<quantity id="4C869B42FFFC2C50FAD7ADBDFA7DF854" box="[1378,1432,1984,2006]" metricMagnitude="1" metricUnit="m" metricValue="2.0" pageId="4" pageNumber="188" unit="m" value="20.0">20 m</quantity>
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||
.
|
||
</specimenCount>
|
||
<typeStatus id="54C58805FFFC2C50FF72ADA2FECBF876" box="[199,302,2015,2036]" pageId="4" pageNumber="188" type="paratype">Paratypes</typeStatus>
|
||
: AM W.44467, MI QLD 2424, partially mounted on a SEM stub; AM W.47750, MI QLD 2435.
|
||
</paragraph>
|
||
<caption id="DF01662FFFFD2C51FF22AFA6FAA5F9AC" httpUri="https://zenodo.org/record/245141/files/figure.png" pageId="5" pageNumber="189" targetBox="[153,1422,188,1471]" targetPageId="5">
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<paragraph id="8BC136A7FFFD2C51FF22AFA6FAA5F9AC" blockId="5.[151,1436,1499,1582]" pageId="5" pageNumber="189">
|
||
<emphasis id="B90AEAB5FFFD2C51FF22AFA6FEF3FA73" bold="true" box="[151,278,1499,1521]" pageId="5" pageNumber="189">FIGURE 2.</emphasis>
|
||
<taxonomicName id="4C7E4D24FFFD2C51FEAAAFA6FE3DFA73" box="[287,472,1499,1521]" class="Polychaeta" family="Sabellariidae" genus="Lygdamis" kingdom="Animalia" order="Sabellida" pageId="5" pageNumber="189" phylum="Annelida" rank="species" species="nasutus" status="sp. nov.">
|
||
<emphasis id="B90AEAB5FFFD2C51FEAAAFA6FE3DFA73" box="[287,472,1499,1521]" italics="true" pageId="5" pageNumber="189">Lygdamis nasutus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A23957CEFFFD2C51FE57AFA1FDF8FA73" box="[482,541,1500,1521]" pageId="5" pageNumber="189" rank="species">n. sp.</taxonomicNameLabel>
|
||
, paratype, AM W.47750, live. A. Whole specimen, dorsal view, arrowheads point to notopodial cirri; B. Anterior end, lateral view, with median organ protruding from operculum (arrow); C. Anterior end, dorsal view; D. Operculum, frontal view; E. Posterior abdominal chaetigers and cauda, ventral view. Photos by A. Semenov.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BC136A7FFFD2C51FF22AC0CFC7DF92C" blockId="5.[151,1437,1649,2034]" pageId="5" pageNumber="189">
|
||
<emphasis id="B90AEAB5FFFD2C51FF22AC0CFDC9F908" bold="true" box="[151,556,1649,1674]" pageId="5" pageNumber="189">Comparative material examined.</emphasis>
|
||
<typeStatus id="54C58805FFFD2C51FD8DAC0CFD45F908" box="[568,672,1649,1674]" pageId="5" pageNumber="189" type="holotype">Holotype</typeStatus>
|
||
of
|
||
<taxonomicName id="4C7E4D24FFFD2C51FD65AC0CFB87F908" authority="McIntosh, 1885" authorityName="McIntosh" authorityYear="1885" box="[720,1122,1649,1674]" class="Polychaeta" family="Sabellariidae" genus="Lygdamis" kingdom="Animalia" order="Sabellida" pageId="5" pageNumber="189" phylum="Annelida" rank="species" species="giardi">
|
||
<emphasis id="B90AEAB5FFFD2C51FD65AC0CFC69F908" box="[720,908,1649,1674]" italics="true" pageId="5" pageNumber="189">Lygdamis giardi</emphasis>
|
||
(
|
||
<bibRefCitation id="EFEF4B56FFFD2C51FC15AC0CFBBFF908" author="McIntosh" box="[928,1114,1649,1674]" pageId="5" pageNumber="189" refString="McIntosh, W. C. (1885) Report on the Annelida Polychaeta collected by H. M. S. ' Challenger' during the years 1873 - 76. Report on the Scientific Results of the Voyage of H. M. S. Challenger, Zoology, 12, 1 - 554." type="journal article" year="1885">McIntosh, 1885</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
<collectionCode id="ED6FAE62FFFD2C51FBDBAC0FFB27F90B" box="[1134,1218,1650,1673]" pageId="5" pageNumber="189">BMNH</collectionCode>
|
||
1885.12.1.6, New South
|
||
<collectingCountry id="F3697637FFFD2C51FF6AACE8FEC3F92C" box="[223,294,1685,1710]" name="United Kingdom" pageId="5" pageNumber="189">Wales</collectingCountry>
|
||
, off Port Jackson,
|
||
<geoCoordinate id="EE4A5060FFFD2C51FE41ACE8FDABF92C" box="[500,590,1685,1710]" direction="south" orientation="latitude" pageId="5" pageNumber="189" precision="925" value="-33.85">33°51'S</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE4A5060FFFD2C51FDEFACE8FD26F92C" box="[602,707,1685,1710]" direction="east" orientation="longitude" pageId="5" pageNumber="189" precision="925" value="151.36667">151°22'E</geoCoordinate>
|
||
,
|
||
<quantity id="4C869B42FFFD2C51FD7AACE8FCEFF92F" box="[719,778,1685,1710]" metricMagnitude="1" metricUnit="m" metricValue="6.4" pageId="5" pageNumber="189" unit="m" value="64.0">64 m</quantity>
|
||
,
|
||
<date id="FFC01067FFFD2C51FCA3ACEBFC70F92C" box="[790,917,1686,1710]" pageId="5" pageNumber="189" value="1874-06-03">3 Jun 1874</date>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C364652CFFFD2C5FFF72ACC4FBDEFDE2" lastPageId="11" lastPageNumber="195" pageId="5" pageNumber="189" type="description">
|
||
<paragraph id="8BC136A7FFFD2C51FF72ACC4FD6FF870" blockId="5.[151,1437,1649,2034]" pageId="5" pageNumber="189">
|
||
<emphasis id="B90AEAB5FFFD2C51FF72ACC4FEBCF950" bold="true" box="[199,345,1721,1746]" pageId="5" pageNumber="189">Description.</emphasis>
|
||
<emphasis id="B90AEAB5FFFD2C51FED7ACC4FDE9F950" box="[354,524,1721,1746]" italics="true" pageId="5" pageNumber="189">Colour pattern</emphasis>
|
||
. Live specimens with bright pigmentation pattern on epithelium of operculum, consisting of white opercular lobes with 3–4 orange transverse bands on outer and inner side of opercular lobes (
|
||
<figureCitation id="13452A22FFFD2C51FF2AAD7CFF00F898" box="[159,229,1793,1818]" captionStart="FIGURE 2" captionStartId="5.[151,250,1499,1521]" captionTargetBox="[153,1422,188,1471]" captionTargetId="figure@5.[151,1424,186,1474]" captionTargetPageId="5" captionText="FIGURE 2. Lygdamis nasutus n. sp., paratype, AM W. 47750, live. A. Whole specimen, dorsal view, arrowheads point to notopodial cirri; B. Anterior end, lateral view, with median organ protruding from operculum (arrow); C. Anterior end, dorsal view; D. Operculum, frontal view; E. Posterior abdominal chaetigers and cauda, ventral view. Photos by A. Semenov." httpUri="https://zenodo.org/record/245141/files/figure.png" pageId="5" pageNumber="189">Fig. 2</figureCitation>
|
||
A–D) and 7–9 short brown and white transversal bands on ventral side of lobes, flanking the tentacular filaments (
|
||
<figureCitation id="13452A22FFFD2C51FEBBAD58FEB6F8BC" box="[270,339,1829,1854]" captionStart="FIGURE 2" captionStartId="5.[151,250,1499,1521]" captionTargetBox="[153,1422,188,1471]" captionTargetId="figure@5.[151,1424,186,1474]" captionTargetPageId="5" captionText="FIGURE 2. Lygdamis nasutus n. sp., paratype, AM W. 47750, live. A. Whole specimen, dorsal view, arrowheads point to notopodial cirri; B. Anterior end, lateral view, with median organ protruding from operculum (arrow); C. Anterior end, dorsal view; D. Operculum, frontal view; E. Posterior abdominal chaetigers and cauda, ventral view. Photos by A. Semenov." httpUri="https://zenodo.org/record/245141/files/figure.png" pageId="5" pageNumber="189">Fig. 2</figureCitation>
|
||
B). Median organ deep orange at base, becoming paler towards tip, dark ring around the tip with a white pigmented flattened top. Base of opercular lobes, around insertion of median organ, strongly pigmented in black. First two chaetigers with white pigment. Rest of body unpigmented except for red-brown branchiae (
|
||
<figureCitation id="13452A22FFFD2C51FADAAD10FF40F828" captionStart="FIGURE 2" captionStartId="5.[151,250,1499,1521]" captionTargetBox="[153,1422,188,1471]" captionTargetId="figure@5.[151,1424,186,1474]" captionTargetPageId="5" captionText="FIGURE 2. Lygdamis nasutus n. sp., paratype, AM W. 47750, live. A. Whole specimen, dorsal view, arrowheads point to notopodial cirri; B. Anterior end, lateral view, with median organ protruding from operculum (arrow); C. Anterior end, dorsal view; D. Operculum, frontal view; E. Posterior abdominal chaetigers and cauda, ventral view. Photos by A. Semenov." httpUri="https://zenodo.org/record/245141/files/figure.png" pageId="5" pageNumber="189">Fig. 2</figureCitation>
|
||
A, E). Preserved specimens lack white pigmentation (including the top of the median organ, distally dark on preserved specimens,
|
||
<figureCitation id="13452A22FFFD2C51FE20ADC8FE14F84C" box="[405,497,1973,1998]" captionStart="FIGURE 3. A – D" captionStartId="6.[151,250,1449,1471]" captionTargetBox="[154,1433,195,1423]" captionTargetId="figure@6.[151,1436,193,1428]" captionTargetPageId="6" captionText="FIGURE 3. A – D. Lygdamis nasutus n. sp., holotype, AM W. 47372, preserved. A. Operculum, thoracic and parathoracic chaetigers, lateral view; arrowheads point to notopodial cirri; B. Anterior end, ventral view; C. Same, dorsal view; D. Same, but with exposed well-developed median organ. E – F Paratype AM W. 44467. E. Detail of median organ, dorsal view; F. Cross section of paratypes, showing half of the operculum with whole median organ. Abbreviations: br = branchia, bo = building organ, cn 1 = cirrus neuropodia segment 1, chn 1 = chaetae neuropodium segment 1, dc = dorsal cirrus, es = eyespots, ip = inner paleae, li = lips, m = mouth, mo = median organ, nh = nuchal hook, op = outer paleae, opa = opercular papillae, pa = palp, rtf = row of tentacular filaments, tf = tentacular filaments. Small arrow indicates conical dorsal cirrus. Photo: A – D—E. Wong." httpUri="https://zenodo.org/record/245142/files/figure.png" pageId="5" pageNumber="189">Figs 3D</figureCitation>
|
||
–F, 4A–C) but retain the colour bands on operculum described above, now dark brown instead of bright orange (
|
||
<figureCitation id="13452A22FFFD2C51FE4AADA4FDBFF870" box="[511,602,2009,2034]" captionStart="FIGURE 3. A – D" captionStartId="6.[151,250,1449,1471]" captionTargetBox="[154,1433,195,1423]" captionTargetId="figure@6.[151,1436,193,1428]" captionTargetPageId="6" captionText="FIGURE 3. A – D. Lygdamis nasutus n. sp., holotype, AM W. 47372, preserved. A. Operculum, thoracic and parathoracic chaetigers, lateral view; arrowheads point to notopodial cirri; B. Anterior end, ventral view; C. Same, dorsal view; D. Same, but with exposed well-developed median organ. E – F Paratype AM W. 44467. E. Detail of median organ, dorsal view; F. Cross section of paratypes, showing half of the operculum with whole median organ. Abbreviations: br = branchia, bo = building organ, cn 1 = cirrus neuropodia segment 1, chn 1 = chaetae neuropodium segment 1, dc = dorsal cirrus, es = eyespots, ip = inner paleae, li = lips, m = mouth, mo = median organ, nh = nuchal hook, op = outer paleae, opa = opercular papillae, pa = palp, rtf = row of tentacular filaments, tf = tentacular filaments. Small arrow indicates conical dorsal cirrus. Photo: A – D—E. Wong." httpUri="https://zenodo.org/record/245142/files/figure.png" pageId="5" pageNumber="189">Figs 3A</figureCitation>
|
||
–E).
|
||
</paragraph>
|
||
<caption id="DF01662FFFFE2C52FF22AFD4FA86F9F5" httpUri="https://zenodo.org/record/245142/files/figure.png" pageId="6" pageNumber="190" targetBox="[154,1433,195,1423]" targetPageId="6">
|
||
<paragraph id="8BC136A7FFFE2C52FF22AFD4FA86F9F5" blockId="6.[151,1436,1449,1655]" pageId="6" pageNumber="190">
|
||
<emphasis id="B90AEAB5FFFE2C52FF22AFD4FEF3FA3D" bold="true" box="[151,278,1449,1471]" pageId="6" pageNumber="190">FIGURE 3.</emphasis>
|
||
A–D.
|
||
<taxonomicName id="4C7E4D24FFFE2C52FED6AFD4FDF9FA3D" box="[355,540,1449,1471]" class="Polychaeta" family="Sabellariidae" genus="Lygdamis" kingdom="Animalia" order="Sabellida" pageId="6" pageNumber="190" phylum="Annelida" rank="species" species="nasutus" status="sp. nov.">
|
||
<emphasis id="B90AEAB5FFFE2C52FED6AFD4FDF9FA3D" box="[355,540,1449,1471]" italics="true" pageId="6" pageNumber="190">Lygdamis nasutus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A23957CEFFFE2C52FD92AFD7FD86FA3D" box="[551,611,1450,1471]" pageId="6" pageNumber="190" rank="species">n. sp.</taxonomicNameLabel>
|
||
, holotype, AM W.47372, preserved. A. Operculum, thoracic and parathoracic chaetigers, lateral view; arrowheads point to notopodial cirri; B. Anterior end, ventral view; C. Same, dorsal view; D. Same, but with exposed well-developed median organ. E–F Paratype AM W.44467. E. Detail of median organ, dorsal view; F. Cross section of paratypes, showing half of the operculum with whole median organ. Abbreviations: br = branchia, bo = building organ, cn1 = cirrus neuropodia segment 1, chn1 = chaetae neuropodium segment 1, dc = dorsal cirrus, es = eyespots, ip = inner paleae, li = lips, m = mouth, mo = median organ, nh = nuchal hook, op = outer paleae, opa = opercular papillae, pa = palp, rtf = row of tentacular filaments, tf = tentacular filaments. Small arrow indicates conical dorsal cirrus. Photo: A–D—E. Wong.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BC136A7FFFE2C52FF72ACBAFB13F886" blockId="6.[151,1437,1735,2013]" pageId="6" pageNumber="190">
|
||
<emphasis id="B90AEAB5FFFE2C52FF72ACBAFF1AF962" box="[199,255,1735,1760]" italics="true" pageId="6" pageNumber="190">Body</emphasis>
|
||
.
|
||
<typeStatus id="54C58805FFFE2C52FEBAACBAFE92F962" box="[271,375,1735,1760]" pageId="6" pageNumber="190" type="holotype">Holotype</typeStatus>
|
||
incomplete but well preserved, 39 chaetigers (32–43),
|
||
<quantity id="4C869B42FFFE2C52FC41ACB5FBA0F95D" box="[1012,1093,1736,1760]" metricMagnitude="-2" metricUnit="m" metricValue="3.2" pageId="6" pageNumber="190" unit="mm" value="32.0">32 mm</quantity>
|
||
long (
|
||
<quantity id="4C869B42FFFE2C52FB24ACB5FAF7F962" box="[1169,1298,1736,1760]" metricMagnitude="-2" metricUnit="m" metricValue="3.65" metricValueMax="4.3" metricValueMin="3.0" pageId="6" pageNumber="190" unit="mm" value="36.5" valueMax="43.0" valueMin="30.0">30–43 mm</quantity>
|
||
), and
|
||
<quantity id="4C869B42FFFE2C52FAECACB5FA78F95D" box="[1369,1437,1736,1760]" metricMagnitude="-3" metricUnit="m" metricValue="4.0" pageId="6" pageNumber="190" unit="mm" value="4.0">4 mm</quantity>
|
||
maximum width excluding parapodia (
|
||
<quantity id="4C869B42FFFE2C52FDFCAC96FD4EF886" box="[585,683,1771,1796]" metricMagnitude="-3" metricUnit="m" metricValue="7.0" metricValueMax="8.0" metricValueMin="6.0" pageId="6" pageNumber="190" unit="mm" value="7.0" valueMax="8.0" valueMin="6.0">6–8 mm</quantity>
|
||
); posterior abdominal chaetigers and cauda missing.
|
||
</paragraph>
|
||
<paragraph id="8BC136A7FFFE2C53FF72AD72FBB4FEE6" blockId="6.[151,1437,1735,2013]" lastBlockId="7.[151,1436,151,357]" lastPageId="7" lastPageNumber="191" pageId="6" pageNumber="190">
|
||
<emphasis id="B90AEAB5FFFE2C52FF72AD72FEA1F8AA" box="[199,324,1807,1832]" italics="true" pageId="6" pageNumber="190">Operculum</emphasis>
|
||
. Operculum completely divided into two free long lobes (
|
||
<figureCitation id="13452A22FFFE2C52FC6AAD72FBC6F8AA" box="[991,1059,1807,1832]" captionStart="FIGURE 2" captionStartId="5.[151,250,1499,1521]" captionTargetBox="[153,1422,188,1471]" captionTargetId="figure@5.[151,1424,186,1474]" captionTargetPageId="5" captionText="FIGURE 2. Lygdamis nasutus n. sp., paratype, AM W. 47750, live. A. Whole specimen, dorsal view, arrowheads point to notopodial cirri; B. Anterior end, lateral view, with median organ protruding from operculum (arrow); C. Anterior end, dorsal view; D. Operculum, frontal view; E. Posterior abdominal chaetigers and cauda, ventral view. Photos by A. Semenov." httpUri="https://zenodo.org/record/245141/files/figure.png" pageId="6" pageNumber="190">Fig. 2</figureCitation>
|
||
D) with distal ends oblique (
|
||
<figureCitation id="13452A22FFFE2C52FADFAD72FF43F8CE" captionStart="FIGURE 2" captionStartId="5.[151,250,1499,1521]" captionTargetBox="[153,1422,188,1471]" captionTargetId="figure@5.[151,1424,186,1474]" captionTargetPageId="5" captionText="FIGURE 2. Lygdamis nasutus n. sp., paratype, AM W. 47750, live. A. Whole specimen, dorsal view, arrowheads point to notopodial cirri; B. Anterior end, lateral view, with median organ protruding from operculum (arrow); C. Anterior end, dorsal view; D. Operculum, frontal view; E. Posterior abdominal chaetigers and cauda, ventral view. Photos by A. Semenov." httpUri="https://zenodo.org/record/245141/files/figure.png" pageId="6" pageNumber="190">Figs 2</figureCitation>
|
||
B, 3A, B). Two rows of golden paleae, outer row with 19 (18–19) pairs of paleae, cylindrical at base and flattened, with curved tips, and some fine thecal sculpture (
|
||
<figureCitation id="13452A22FFFE2C52FD68AD2AFCC2F8F2" box="[733,807,1879,1904]" captionStart="FIGURE 5" captionStartId="8.[151,250,1820,1842]" captionTargetBox="[191,1407,198,1798]" captionTargetId="figure@8.[187,1408,193,1805]" captionTargetPageId="8" captionText="FIGURE 5. Chaetae, Lygdamis nasutus n. sp., paratypes. A. Outer paleae; B. Detail of tips of outer paleae; C. Inner paleae, distal end; D. Notopodium, parathoracic chaetiger; E. Detail of tips of lanceolate parathoracic notopodial chaeta; F. Parathoracic notopodial simple capillaries, interspersed with lanceolate chaetae; G. Neuropodium, parathoracic chaetiger; H. Detail of tips of lanceolate parathoracic neuropodial chaetae; I. Parathoracic neuropodial simple capillaries, interspersed with lanceolate chaetae; J. Abdominal notochaetae (two types); K. Neuropodial abdominal uncini; L. Detail of abdominal uncinus, side view." httpUri="https://zenodo.org/record/245144/files/figure.png" pageId="6" pageNumber="190">Figs 5</figureCitation>
|
||
A, B, 6A–C); inner row with 7 (7–8) pairs of paleae, cylindrical shafts with blunt tips and smooth surface (
|
||
<figureCitation id="13452A22FFFE2C52FCA2AD01FC86F817" box="[791,867,1916,1941]" captionStart="FIGURE 5" captionStartId="8.[151,250,1820,1842]" captionTargetBox="[191,1407,198,1798]" captionTargetId="figure@8.[187,1408,193,1805]" captionTargetPageId="8" captionText="FIGURE 5. Chaetae, Lygdamis nasutus n. sp., paratypes. A. Outer paleae; B. Detail of tips of outer paleae; C. Inner paleae, distal end; D. Notopodium, parathoracic chaetiger; E. Detail of tips of lanceolate parathoracic notopodial chaeta; F. Parathoracic notopodial simple capillaries, interspersed with lanceolate chaetae; G. Neuropodium, parathoracic chaetiger; H. Detail of tips of lanceolate parathoracic neuropodial chaetae; I. Parathoracic neuropodial simple capillaries, interspersed with lanceolate chaetae; J. Abdominal notochaetae (two types); K. Neuropodial abdominal uncini; L. Detail of abdominal uncinus, side view." httpUri="https://zenodo.org/record/245144/files/figure.png" pageId="6" pageNumber="190">Figs 5</figureCitation>
|
||
C, 6D, E). Opercular papillae, 10 pairs (10–12), peripheral to outer paleae of each lobe, conical, similar in size except for the 2–3 pairs closer to nuchal spines larger than the rest (
|
||
<figureCitation id="13452A22FFFE2C52FE36ADB9FE2BF85E" box="[387,462,1988,2013]" captionStart="FIGURE 2" captionStartId="5.[151,250,1499,1521]" captionTargetBox="[153,1422,188,1471]" captionTargetId="figure@5.[151,1424,186,1474]" captionTargetPageId="5" captionText="FIGURE 2. Lygdamis nasutus n. sp., paratype, AM W. 47750, live. A. Whole specimen, dorsal view, arrowheads point to notopodial cirri; B. Anterior end, lateral view, with median organ protruding from operculum (arrow); C. Anterior end, dorsal view; D. Operculum, frontal view; E. Posterior abdominal chaetigers and cauda, ventral view. Photos by A. Semenov." httpUri="https://zenodo.org/record/245141/files/figure.png" pageId="6" pageNumber="190">Figs 2</figureCitation>
|
||
B, 3A–B, D, F, 6A). One pair of brown nuchal spines on each side (
|
||
<figureCitation id="13452A22FFFE2C52FB45ADB9FADFF85E" box="[1264,1338,1988,2013]" captionStart="FIGURE 2" captionStartId="5.[151,250,1499,1521]" captionTargetBox="[153,1422,188,1471]" captionTargetId="figure@5.[151,1424,186,1474]" captionTargetPageId="5" captionText="FIGURE 2. Lygdamis nasutus n. sp., paratype, AM W. 47750, live. A. Whole specimen, dorsal view, arrowheads point to notopodial cirri; B. Anterior end, lateral view, with median organ protruding from operculum (arrow); C. Anterior end, dorsal view; D. Operculum, frontal view; E. Posterior abdominal chaetigers and cauda, ventral view. Photos by A. Semenov." httpUri="https://zenodo.org/record/245141/files/figure.png" pageId="6" pageNumber="190">Figs 2</figureCitation>
|
||
A, C–D, 3C–D), stout, strongly recurved without sharp limbations on concave margin (
|
||
<figureCitation id="13452A22FFFF2C53FB8CAAEAFB67FF32" box="[1081,1154,151,176]" captionStart="FIGURE 6" captionStartId="9.[151,250,1737,1759]" captionTargetBox="[153,1435,341,1713]" captionTargetId="figure@9.[150,1437,337,1716]" captionTargetPageId="9" captionText="FIGURE 6. Chaetae, Lygdamis nasutus n. sp., paratypes, SEM images, AM W. 44467. A. Operculum, lateral view; B. Outer paleae; C. Thecal sculpture, outer paleae; D. Inner paleae; E. Close up of distal end, inner paleae; F. First thoracic neuropodial chaetae; G. Parathoracic notopodia; H. Parathoracic neuropodium. Abbreviations: cn 1 = cirrus neuropodia segment 1, chn 1 = chaetae neuropodium segment 1, ip = inner paleae, mo = median organ, nh = nuchal hook, op = outer paleae, opa = opercular papillae, pa = palp." httpUri="https://zenodo.org/record/245145/files/figure.png" pageId="7" pageNumber="191">Fig. 6</figureCitation>
|
||
A). Tentacular filaments compound, arranged in 11 rows (11–12) (
|
||
<figureCitation id="13452A22FFFF2C53FD31AAC1FD34FF56" box="[644,721,188,213]" captionStart="FIGURE 2" captionStartId="5.[151,250,1499,1521]" captionTargetBox="[153,1422,188,1471]" captionTargetId="figure@5.[151,1424,186,1474]" captionTargetPageId="5" captionText="FIGURE 2. Lygdamis nasutus n. sp., paratype, AM W. 47750, live. A. Whole specimen, dorsal view, arrowheads point to notopodial cirri; B. Anterior end, lateral view, with median organ protruding from operculum (arrow); C. Anterior end, dorsal view; D. Operculum, frontal view; E. Posterior abdominal chaetigers and cauda, ventral view. Photos by A. Semenov." httpUri="https://zenodo.org/record/245141/files/figure.png" pageId="7" pageNumber="191">Figs 2</figureCitation>
|
||
B, 3B). Median ridge inconspicuous. Well-developed median organ, arising abruptly from the base of opercular lobes, near edge of upper lip; slightly shorter than lobes. Median organ cylindrical, with a planar, teardrop coronal shape when viewed head on. A row of eyespots along both sides of its stalk (
|
||
<figureCitation id="13452A22FFFF2C53FEABAB5AFE81FEC2" box="[286,356,295,320]" captionStart="FIGURE 3. A – D" captionStartId="6.[151,250,1449,1471]" captionTargetBox="[154,1433,195,1423]" captionTargetId="figure@6.[151,1436,193,1428]" captionTargetPageId="6" captionText="FIGURE 3. A – D. Lygdamis nasutus n. sp., holotype, AM W. 47372, preserved. A. Operculum, thoracic and parathoracic chaetigers, lateral view; arrowheads point to notopodial cirri; B. Anterior end, ventral view; C. Same, dorsal view; D. Same, but with exposed well-developed median organ. E – F Paratype AM W. 44467. E. Detail of median organ, dorsal view; F. Cross section of paratypes, showing half of the operculum with whole median organ. Abbreviations: br = branchia, bo = building organ, cn 1 = cirrus neuropodia segment 1, chn 1 = chaetae neuropodium segment 1, dc = dorsal cirrus, es = eyespots, ip = inner paleae, li = lips, m = mouth, mo = median organ, nh = nuchal hook, op = outer paleae, opa = opercular papillae, pa = palp, rtf = row of tentacular filaments, tf = tentacular filaments. Small arrow indicates conical dorsal cirrus. Photo: A – D—E. Wong." httpUri="https://zenodo.org/record/245142/files/figure.png" pageId="7" pageNumber="191">Figs 3</figureCitation>
|
||
E–F, 4A–C), and tufts of cilia on its surface, more densely arranged on coronal distal end (
|
||
<figureCitation id="13452A22FFFF2C53FADAAB5AFF43FEE6" captionStart="FIGURE 4" captionStartId="7.[151,250,1902,1924]" captionTargetBox="[153,1431,412,1878]" captionTargetId="figure@7.[151,1436,409,1881]" captionTargetPageId="7" captionText="FIGURE 4. Median organ, Lygdamis nasutus n. sp., paratypes, AM W. 44467. A. Lateral view with a close up of the eyespots along its length; B. Details of the distal tip, showing the flat corona; C. Distal end, showing teardrop coronal shape frontal view. D – G. SEM images, AM W. 47750. D. Complete median organ, lateral view; E. Distal end, lateral view; F. Detail of the base of median organ with a few sparsely distributed ciliary tufts; G. Close up of ciliary tufts." httpUri="https://zenodo.org/record/245143/files/figure.png" pageId="7" pageNumber="191">Fig. 4</figureCitation>
|
||
D–G). Pair of grooved, ringed palps, half the length of opercular lobes (
|
||
<figureCitation id="13452A22FFFF2C53FC7CAB31FBA1FEE6" box="[969,1092,332,357]" captionStart="FIGURE 3. A – D" captionStartId="6.[151,250,1449,1471]" captionTargetBox="[154,1433,195,1423]" captionTargetId="figure@6.[151,1436,193,1428]" captionTargetPageId="6" captionText="FIGURE 3. A – D. Lygdamis nasutus n. sp., holotype, AM W. 47372, preserved. A. Operculum, thoracic and parathoracic chaetigers, lateral view; arrowheads point to notopodial cirri; B. Anterior end, ventral view; C. Same, dorsal view; D. Same, but with exposed well-developed median organ. E – F Paratype AM W. 44467. E. Detail of median organ, dorsal view; F. Cross section of paratypes, showing half of the operculum with whole median organ. Abbreviations: br = branchia, bo = building organ, cn 1 = cirrus neuropodia segment 1, chn 1 = chaetae neuropodium segment 1, dc = dorsal cirrus, es = eyespots, ip = inner paleae, li = lips, m = mouth, mo = median organ, nh = nuchal hook, op = outer paleae, opa = opercular papillae, pa = palp, rtf = row of tentacular filaments, tf = tentacular filaments. Small arrow indicates conical dorsal cirrus. Photo: A – D—E. Wong." httpUri="https://zenodo.org/record/245142/files/figure.png" pageId="7" pageNumber="191">Fig. 3A–B</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<caption id="DF01662FFFFF2C53FF22AD13FC1DF86A" httpUri="https://zenodo.org/record/245143/files/figure.png" pageId="7" pageNumber="191" targetBox="[153,1431,412,1878]" targetPageId="7">
|
||
<paragraph id="8BC136A7FFFF2C53FF22AD13FC1DF86A" blockId="7.[151,1437,1902,2024]" pageId="7" pageNumber="191">
|
||
<emphasis id="B90AEAB5FFFF2C53FF22AD13FEF6F806" bold="true" box="[151,275,1902,1924]" pageId="7" pageNumber="191">FIGURE 4.</emphasis>
|
||
Median organ,
|
||
<taxonomicName id="4C7E4D24FFFF2C53FE01AD13FD8FF806" box="[436,618,1902,1924]" class="Polychaeta" family="Sabellariidae" genus="Lygdamis" kingdom="Animalia" order="Sabellida" pageId="7" pageNumber="191" phylum="Annelida" rank="species" species="nasutus" status="sp. nov.">
|
||
<emphasis id="B90AEAB5FFFF2C53FE01AD13FD8FF806" box="[436,618,1902,1924]" italics="true" pageId="7" pageNumber="191">Lygdamis nasutus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A23957CEFFFF2C53FDC4AD12FD4FF806" box="[625,682,1903,1924]" pageId="7" pageNumber="191" rank="species">n. sp.</taxonomicNameLabel>
|
||
, paratypes, AM W.44467. A. Lateral view with a close up of the eyespots along its length; B. Details of the distal tip, showing the flat corona; C. Distal end, showing teardrop coronal shape frontal view. D–G. SEM images, AM W.47750. D. Complete median organ, lateral view; E. Distal end, lateral view; F. Detail of the base of median organ with a few sparsely distributed ciliary tufts; G. Close up of ciliary tufts.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF01662FFFF02C5CFF22AD61FF18F85B" httpUri="https://zenodo.org/record/245144/files/figure.png" pageId="8" pageNumber="192" targetBox="[191,1407,198,1798]" targetPageId="8">
|
||
<paragraph id="8BC136A7FFF02C5CFF22AD61FF18F85B" blockId="8.[151,1437,1820,2009]" pageId="8" pageNumber="192">
|
||
<emphasis id="B90AEAB5FFF02C5CFF22AD61FEF6F8B0" bold="true" box="[151,275,1820,1842]" pageId="8" pageNumber="192">FIGURE 5.</emphasis>
|
||
Chaetae,
|
||
<taxonomicName id="4C7E4D24FFF02C5CFEC9AD61FDD7F8B0" box="[380,562,1820,1842]" class="Polychaeta" family="Sabellariidae" genus="Lygdamis" kingdom="Animalia" order="Sabellida" pageId="8" pageNumber="192" phylum="Annelida" rank="species" species="nasutus" status="sp. nov.">
|
||
<emphasis id="B90AEAB5FFF02C5CFEC9AD61FDD7F8B0" box="[380,562,1820,1842]" italics="true" pageId="8" pageNumber="192">Lygdamis nasutus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A23957CEFFF02C5CFD8FAD60FD91F8B0" box="[570,628,1821,1842]" pageId="8" pageNumber="192" rank="species">n. sp.</taxonomicNameLabel>
|
||
, paratypes. A. Outer paleae; B. Detail of tips of outer paleae; C. Inner paleae, distal end; D. Notopodium, parathoracic chaetiger; E. Detail of tips of lanceolate parathoracic notopodial chaeta; F. Parathoracic notopodial simple capillaries, interspersed with lanceolate chaetae; G. Neuropodium, parathoracic chaetiger; H. Detail of tips of lanceolate parathoracic neuropodial chaetae; I. Parathoracic neuropodial simple capillaries, interspersed with lanceolate chaetae; J. Abdominal notochaetae (two types); K. Neuropodial abdominal uncini; L. Detail of abdominal uncinus, side view.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BC136A7FFF12C5DFF72AAEAFD00FE9E" blockId="9.[151,1437,151,285]" pageId="9" pageNumber="193">
|
||
<emphasis id="B90AEAB5FFF12C5DFF72AAEAFEFDFF32" box="[199,280,151,176]" italics="true" pageId="9" pageNumber="193">Thorax</emphasis>
|
||
. Chaetiger 1 with lobe-shaped neuropodia and elongated cirrus (
|
||
<figureCitation id="13452A22FFF12C5DFC52AAEAFBCBFF32" box="[999,1070,151,176]" captionStart="FIGURE 2" captionStartId="5.[151,250,1499,1521]" captionTargetBox="[153,1422,188,1471]" captionTargetId="figure@5.[151,1424,186,1474]" captionTargetPageId="5" captionText="FIGURE 2. Lygdamis nasutus n. sp., paratype, AM W. 47750, live. A. Whole specimen, dorsal view, arrowheads point to notopodial cirri; B. Anterior end, lateral view, with median organ protruding from operculum (arrow); C. Anterior end, dorsal view; D. Operculum, frontal view; E. Posterior abdominal chaetigers and cauda, ventral view. Photos by A. Semenov." httpUri="https://zenodo.org/record/245141/files/figure.png" pageId="9" pageNumber="193">Figs 2</figureCitation>
|
||
B, 3A–B); capillary chaetae with fibres forming distal hood (
|
||
<figureCitation id="13452A22FFF12C5DFE6EAAC1FDC7FF57" box="[475,546,188,213]" captionStart="FIGURE 6" captionStartId="9.[151,250,1737,1759]" captionTargetBox="[153,1435,341,1713]" captionTargetId="figure@9.[150,1437,337,1716]" captionTargetPageId="9" captionText="FIGURE 6. Chaetae, Lygdamis nasutus n. sp., paratypes, SEM images, AM W. 44467. A. Operculum, lateral view; B. Outer paleae; C. Thecal sculpture, outer paleae; D. Inner paleae; E. Close up of distal end, inner paleae; F. First thoracic neuropodial chaetae; G. Parathoracic notopodia; H. Parathoracic neuropodium. Abbreviations: cn 1 = cirrus neuropodia segment 1, chn 1 = chaetae neuropodium segment 1, ip = inner paleae, mo = median organ, nh = nuchal hook, op = outer paleae, opa = opercular papillae, pa = palp." httpUri="https://zenodo.org/record/245145/files/figure.png" pageId="9" pageNumber="193">Fig. 6</figureCitation>
|
||
F). Chaetiger 2 with three pairs of elongated triangular-shaped lateral lobes, connecting branchiae to neuropodia (
|
||
<figureCitation id="13452A22FFF12C5DFD88AAA2FD60FF7A" box="[573,645,223,248]" captionStart="FIGURE 2" captionStartId="5.[151,250,1499,1521]" captionTargetBox="[153,1422,188,1471]" captionTargetId="figure@5.[151,1424,186,1474]" captionTargetPageId="5" captionText="FIGURE 2. Lygdamis nasutus n. sp., paratype, AM W. 47750, live. A. Whole specimen, dorsal view, arrowheads point to notopodial cirri; B. Anterior end, lateral view, with median organ protruding from operculum (arrow); C. Anterior end, dorsal view; D. Operculum, frontal view; E. Posterior abdominal chaetigers and cauda, ventral view. Photos by A. Semenov." httpUri="https://zenodo.org/record/245141/files/figure.png" pageId="9" pageNumber="193">Figs 2</figureCitation>
|
||
A–B, 3A); capillary chaetae similar to those in chaetiger 1. About 15 pairs of branchiae present from segment 2 (
|
||
<figureCitation id="13452A22FFF12C5DFDC8AB79FD24FE9E" box="[637,705,260,285]" captionStart="FIGURE 2" captionStartId="5.[151,250,1499,1521]" captionTargetBox="[153,1422,188,1471]" captionTargetId="figure@5.[151,1424,186,1474]" captionTargetPageId="5" captionText="FIGURE 2. Lygdamis nasutus n. sp., paratype, AM W. 47750, live. A. Whole specimen, dorsal view, arrowheads point to notopodial cirri; B. Anterior end, lateral view, with median organ protruding from operculum (arrow); C. Anterior end, dorsal view; D. Operculum, frontal view; E. Posterior abdominal chaetigers and cauda, ventral view. Photos by A. Semenov." httpUri="https://zenodo.org/record/245141/files/figure.png" pageId="9" pageNumber="193">Fig. 2</figureCitation>
|
||
A).
|
||
</paragraph>
|
||
<caption id="DF01662FFFF12C5DFF22ACB4FEBCF8D8" httpUri="https://zenodo.org/record/245145/files/figure.png" pageId="9" pageNumber="193" targetBox="[153,1435,341,1713]" targetPageId="9">
|
||
<paragraph id="8BC136A7FFF12C5DFF22ACB4FEBCF8D8" blockId="9.[151,1436,1737,1882]" pageId="9" pageNumber="193">
|
||
<emphasis id="B90AEAB5FFF12C5DFF22ACB4FEF1F95D" bold="true" box="[151,276,1737,1759]" pageId="9" pageNumber="193">FIGURE 6.</emphasis>
|
||
Chaetae,
|
||
<taxonomicName id="4C7E4D24FFF12C5DFEC9ACB4FDD7F95D" box="[380,562,1737,1759]" class="Polychaeta" family="Sabellariidae" genus="Lygdamis" kingdom="Animalia" order="Sabellida" pageId="9" pageNumber="193" phylum="Annelida" rank="species" species="nasutus" status="sp. nov.">
|
||
<emphasis id="B90AEAB5FFF12C5DFEC9ACB4FDD7F95D" box="[380,562,1737,1759]" italics="true" pageId="9" pageNumber="193">Lygdamis nasutus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A23957CEFFF12C5DFD8EACB7FD91F95D" box="[571,628,1738,1759]" pageId="9" pageNumber="193" rank="species">n. sp.</taxonomicNameLabel>
|
||
, paratypes, SEM images, AM W.44467. A. Operculum, lateral view; B. Outer paleae; C. Thecal sculpture, outer paleae; D. Inner paleae; E. Close up of distal end, inner paleae; F. First thoracic neuropodial chaetae; G. Parathoracic notopodia; H. Parathoracic neuropodium. Abbreviations: cn1 = cirrus neuropodia segment 1, chn1 = chaetae neuropodium segment 1, ip = inner paleae, mo = median organ, nh = nuchal hook, op = outer paleae, opa = opercular papillae, pa = palp.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BC136A7FFF12C5EFF72ADD7FD3DFF7A" blockId="9.[151,1436,1962,2023]" lastBlockId="10.[151,1436,151,248]" lastPageId="10" lastPageNumber="194" pageId="9" pageNumber="193">
|
||
<emphasis id="B90AEAB5FFF12C5DFF72ADD7FEADF841" box="[199,328,1962,1987]" italics="true" pageId="9" pageNumber="193">Parathorax</emphasis>
|
||
. Chaetiger 3–6, with notopodia bearing small, conical dorsal cirrus (
|
||
<figureCitation id="13452A22FFF12C5DFBF3ADD7FB69F841" box="[1094,1164,1962,1987]" captionStart="FIGURE 2" captionStartId="5.[151,250,1499,1521]" captionTargetBox="[153,1422,188,1471]" captionTargetId="figure@5.[151,1424,186,1474]" captionTargetPageId="5" captionText="FIGURE 2. Lygdamis nasutus n. sp., paratype, AM W. 47750, live. A. Whole specimen, dorsal view, arrowheads point to notopodial cirri; B. Anterior end, lateral view, with median organ protruding from operculum (arrow); C. Anterior end, dorsal view; D. Operculum, frontal view; E. Posterior abdominal chaetigers and cauda, ventral view. Photos by A. Semenov." httpUri="https://zenodo.org/record/245141/files/figure.png" pageId="9" pageNumber="193">Figs 2</figureCitation>
|
||
A–B, 3A, C). Two
|
||
<typeStatus id="54C58805FFF12C5DFAD4ADD6FA79F841" box="[1377,1436,1963,1987]" pageId="9" pageNumber="193">types</typeStatus>
|
||
of notochaetae arranged transversely, six lanceolate (6–7), distally concave, with frayed twisted tips (
|
||
<figureCitation id="13452A22FFF12C5DFA8DADB3FA66F865" box="[1336,1411,1998,2023]" captionStart="FIGURE 2" captionStartId="5.[151,250,1499,1521]" captionTargetBox="[153,1422,188,1471]" captionTargetId="figure@5.[151,1424,186,1474]" captionTargetPageId="5" captionText="FIGURE 2. Lygdamis nasutus n. sp., paratype, AM W. 47750, live. A. Whole specimen, dorsal view, arrowheads point to notopodial cirri; B. Anterior end, lateral view, with median organ protruding from operculum (arrow); C. Anterior end, dorsal view; D. Operculum, frontal view; E. Posterior abdominal chaetigers and cauda, ventral view. Photos by A. Semenov." httpUri="https://zenodo.org/record/245141/files/figure.png" pageId="9" pageNumber="193">Figs 2</figureCitation>
|
||
B, 5D–E, 6G), and interspersed with minute capillaries (6–7) (
|
||
<figureCitation id="13452A22FFF22C5EFCF3AAEAFC6AFF32" box="[838,911,151,176]" captionStart="FIGURE 5" captionStartId="8.[151,250,1820,1842]" captionTargetBox="[191,1407,198,1798]" captionTargetId="figure@8.[187,1408,193,1805]" captionTargetPageId="8" captionText="FIGURE 5. Chaetae, Lygdamis nasutus n. sp., paratypes. A. Outer paleae; B. Detail of tips of outer paleae; C. Inner paleae, distal end; D. Notopodium, parathoracic chaetiger; E. Detail of tips of lanceolate parathoracic notopodial chaeta; F. Parathoracic notopodial simple capillaries, interspersed with lanceolate chaetae; G. Neuropodium, parathoracic chaetiger; H. Detail of tips of lanceolate parathoracic neuropodial chaetae; I. Parathoracic neuropodial simple capillaries, interspersed with lanceolate chaetae; J. Abdominal notochaetae (two types); K. Neuropodial abdominal uncini; L. Detail of abdominal uncinus, side view." httpUri="https://zenodo.org/record/245144/files/figure.png" pageId="10" pageNumber="194">Figs 5</figureCitation>
|
||
F, 6G). Neuropodia with large, rounded lobes directed anteriorly (
|
||
<figureCitation id="13452A22FFF22C5EFECDAAC1FE24FF56" box="[376,449,188,213]" captionStart="FIGURE 2" captionStartId="5.[151,250,1499,1521]" captionTargetBox="[153,1422,188,1471]" captionTargetId="figure@5.[151,1424,186,1474]" captionTargetPageId="5" captionText="FIGURE 2. Lygdamis nasutus n. sp., paratype, AM W. 47750, live. A. Whole specimen, dorsal view, arrowheads point to notopodial cirri; B. Anterior end, lateral view, with median organ protruding from operculum (arrow); C. Anterior end, dorsal view; D. Operculum, frontal view; E. Posterior abdominal chaetigers and cauda, ventral view. Photos by A. Semenov." httpUri="https://zenodo.org/record/245141/files/figure.png" pageId="10" pageNumber="194">Figs 2</figureCitation>
|
||
B, 3A); two alternating transverse rows of 6 (5–6) chaetae, lanceolate with frayed tips (
|
||
<figureCitation id="13452A22FFF22C5EFF2AAAA2FF01FF7A" box="[159,228,223,248]" captionStart="FIGURE 5" captionStartId="8.[151,250,1820,1842]" captionTargetBox="[191,1407,198,1798]" captionTargetId="figure@8.[187,1408,193,1805]" captionTargetPageId="8" captionText="FIGURE 5. Chaetae, Lygdamis nasutus n. sp., paratypes. A. Outer paleae; B. Detail of tips of outer paleae; C. Inner paleae, distal end; D. Notopodium, parathoracic chaetiger; E. Detail of tips of lanceolate parathoracic notopodial chaeta; F. Parathoracic notopodial simple capillaries, interspersed with lanceolate chaetae; G. Neuropodium, parathoracic chaetiger; H. Detail of tips of lanceolate parathoracic neuropodial chaetae; I. Parathoracic neuropodial simple capillaries, interspersed with lanceolate chaetae; J. Abdominal notochaetae (two types); K. Neuropodial abdominal uncini; L. Detail of abdominal uncinus, side view." httpUri="https://zenodo.org/record/245144/files/figure.png" pageId="10" pageNumber="194">Figs 5</figureCitation>
|
||
G–H, 6H), and thin capillaries (
|
||
<figureCitation id="13452A22FFF22C5EFDF2AAA2FD6AFF7A" box="[583,655,223,248]" captionStart="FIGURE 5" captionStartId="8.[151,250,1820,1842]" captionTargetBox="[191,1407,198,1798]" captionTargetId="figure@8.[187,1408,193,1805]" captionTargetPageId="8" captionText="FIGURE 5. Chaetae, Lygdamis nasutus n. sp., paratypes. A. Outer paleae; B. Detail of tips of outer paleae; C. Inner paleae, distal end; D. Notopodium, parathoracic chaetiger; E. Detail of tips of lanceolate parathoracic notopodial chaeta; F. Parathoracic notopodial simple capillaries, interspersed with lanceolate chaetae; G. Neuropodium, parathoracic chaetiger; H. Detail of tips of lanceolate parathoracic neuropodial chaetae; I. Parathoracic neuropodial simple capillaries, interspersed with lanceolate chaetae; J. Abdominal notochaetae (two types); K. Neuropodial abdominal uncini; L. Detail of abdominal uncinus, side view." httpUri="https://zenodo.org/record/245144/files/figure.png" pageId="10" pageNumber="194">
|
||
Figs
|
||
<date id="FFC01067FFF22C5EFD35AAA2FD6AFF7A" box="[640,655,223,248]" pageId="10" pageNumber="194">5</date>
|
||
</figureCitation>
|
||
I, 6H).
|
||
</paragraph>
|
||
<caption id="DF01662FFFF22C5EFF22AD28FBB3F845" httpUri="https://zenodo.org/record/245146/files/figure.png" pageId="10" pageNumber="194" targetBox="[183,1410,288,1860]" targetPageId="10">
|
||
<paragraph id="8BC136A7FFF22C5EFF22AD28FBB3F845" blockId="10.[151,1437,1877,1991]" pageId="10" pageNumber="194">
|
||
<emphasis id="B90AEAB5FFF22C5EFF22AD28FEF0F8E9" bold="true" box="[151,277,1877,1899]" pageId="10" pageNumber="194">FIGURE 7.</emphasis>
|
||
Chaetae,
|
||
<taxonomicName id="4C7E4D24FFF22C5EFE34AD28FDDCF8E9" box="[385,569,1877,1899]" class="Polychaeta" family="Sabellariidae" genus="Lygdamis" kingdom="Animalia" order="Sabellida" pageId="10" pageNumber="194" phylum="Annelida" rank="species" species="nasutus" status="sp. nov.">
|
||
<emphasis id="B90AEAB5FFF22C5EFE34AD28FDDCF8E9" box="[385,569,1877,1899]" italics="true" pageId="10" pageNumber="194">Lygdamis nasutus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A23957CEFFF22C5EFDF6AD2BFD9BF8E9" box="[579,638,1878,1899]" pageId="10" pageNumber="194" rank="species">n. sp.</taxonomicNameLabel>
|
||
, paratypes, SEM images, AM W.44467. A. Anterior abdominal uncini with number of rows of teeth provided in one uncinus, close up of distal dentition; B. Detached uncini, with number of rows of teeth provided in one uncinus; C. Posterior abdominal uncini; D. Detail of posterior abdominal uncini; E. Abdominal notopodium; F. Detail of the two types of abdominal notochaetae; G. Posterior abdominal chaetiger and cauda.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BC136A7FFF32C5FFF72AAEAFBDBFE52" blockId="11.[151,1437,151,1185]" pageId="11" pageNumber="195">
|
||
<emphasis id="B90AEAB5FFF32C5FFF72AAEAFED4FF32" box="[199,305,151,176]" italics="true" pageId="11" pageNumber="195">Abdomen</emphasis>
|
||
. Abdominal notopodia lobe-like, each with a transverse row of uncini decreasing in number posteriorly. Uncini of anterior abdominal chaetigers with 8–9 transverse rows of teeth with characteristic arrangement: single basal tooth, median 5–6 rows with two teeth each (not aligned but alternating), and distal 2–3 rows with 3–4 teeth each (
|
||
<figureCitation id="13452A22FFF32C5FFE7DAB79FDEAFE9F" box="[456,527,260,285]" captionStart="FIGURE 5" captionStartId="8.[151,250,1820,1842]" captionTargetBox="[191,1407,198,1798]" captionTargetId="figure@8.[187,1408,193,1805]" captionTargetPageId="8" captionText="FIGURE 5. Chaetae, Lygdamis nasutus n. sp., paratypes. A. Outer paleae; B. Detail of tips of outer paleae; C. Inner paleae, distal end; D. Notopodium, parathoracic chaetiger; E. Detail of tips of lanceolate parathoracic notopodial chaeta; F. Parathoracic notopodial simple capillaries, interspersed with lanceolate chaetae; G. Neuropodium, parathoracic chaetiger; H. Detail of tips of lanceolate parathoracic neuropodial chaetae; I. Parathoracic neuropodial simple capillaries, interspersed with lanceolate chaetae; J. Abdominal notochaetae (two types); K. Neuropodial abdominal uncini; L. Detail of abdominal uncinus, side view." httpUri="https://zenodo.org/record/245144/files/figure.png" pageId="11" pageNumber="195">Figs 5</figureCitation>
|
||
K–L, 7A–B). Uncini of posterior chaetigers with two rows of seven teeth each, thicker than those of anterior uncini (
|
||
<figureCitation id="13452A22FFF32C5FFDD4AB5AFD4EFEC2" box="[609,683,295,320]" captionStart="FIGURE 7" captionStartId="10.[151,250,1877,1899]" captionTargetBox="[183,1410,288,1860]" captionTargetId="figure@10.[178,1412,283,1861]" captionTargetPageId="10" captionText="FIGURE 7. Chaetae, Lygdamis nasutus n. sp., paratypes, SEM images, AM W. 44467. A. Anterior abdominal uncini with number of rows of teeth provided in one uncinus, close up of distal dentition; B. Detached uncini, with number of rows of teeth provided in one uncinus; C. Posterior abdominal uncini; D. Detail of posterior abdominal uncini; E. Abdominal notopodium; F. Detail of the two types of abdominal notochaetae; G. Posterior abdominal chaetiger and cauda." httpUri="https://zenodo.org/record/245146/files/figure.png" pageId="11" pageNumber="195">Fig. 7</figureCitation>
|
||
C–D). Abdominal neuropodia with capillaries, arranged in two longitudinal rows; dorsal row with slightly larger chaetae, ornamented with irregular thecal laminar extensions and ventral row with thinner and with fibres forming distal hood (
|
||
<figureCitation id="13452A22FFF32C5FFCC3AB12FC24FE0A" box="[886,961,367,392]" captionStart="FIGURE 5" captionStartId="8.[151,250,1820,1842]" captionTargetBox="[191,1407,198,1798]" captionTargetId="figure@8.[187,1408,193,1805]" captionTargetPageId="8" captionText="FIGURE 5. Chaetae, Lygdamis nasutus n. sp., paratypes. A. Outer paleae; B. Detail of tips of outer paleae; C. Inner paleae, distal end; D. Notopodium, parathoracic chaetiger; E. Detail of tips of lanceolate parathoracic notopodial chaeta; F. Parathoracic notopodial simple capillaries, interspersed with lanceolate chaetae; G. Neuropodium, parathoracic chaetiger; H. Detail of tips of lanceolate parathoracic neuropodial chaetae; I. Parathoracic neuropodial simple capillaries, interspersed with lanceolate chaetae; J. Abdominal notochaetae (two types); K. Neuropodial abdominal uncini; L. Detail of abdominal uncinus, side view." httpUri="https://zenodo.org/record/245144/files/figure.png" pageId="11" pageNumber="195">Figs 5</figureCitation>
|
||
J, 7E–F). Last 5–7 abdominal chaetigers thinner and compacted (
|
||
<figureCitation id="13452A22FFF32C5FFE1AABE9FE1FFE2E" box="[431,506,404,429]" captionStart="FIGURE 2" captionStartId="5.[151,250,1499,1521]" captionTargetBox="[153,1422,188,1471]" captionTargetId="figure@5.[151,1424,186,1474]" captionTargetPageId="5" captionText="FIGURE 2. Lygdamis nasutus n. sp., paratype, AM W. 47750, live. A. Whole specimen, dorsal view, arrowheads point to notopodial cirri; B. Anterior end, lateral view, with median organ protruding from operculum (arrow); C. Anterior end, dorsal view; D. Operculum, frontal view; E. Posterior abdominal chaetigers and cauda, ventral view. Photos by A. Semenov." httpUri="https://zenodo.org/record/245141/files/figure.png" pageId="11" pageNumber="195">Figs 2</figureCitation>
|
||
E, 7G), not bearing chaetae or uncini. Branchiae diminishing in size posteriorly, present to chaetiger 14 (12–18), and absent in posterior-most abdominal chaetigers.
|
||
</paragraph>
|
||
<paragraph id="8BC136A7FFF32C5FFF72ABA1FDB6FD9A" blockId="11.[151,1437,151,1185]" pageId="11" pageNumber="195">
|
||
<emphasis id="B90AEAB5FFF32C5FFF72ABA1FEF7FE77" box="[199,274,476,501]" italics="true" pageId="11" pageNumber="195">Cauda</emphasis>
|
||
. Cauda smooth, about the length of ten posterior abdomen segments, with no signs of segmentation but some longitudinal ridges (
|
||
<figureCitation id="13452A22FFF32C5FFE0FAB82FDE7FD9A" box="[442,514,511,536]" captionStart="FIGURE 2" captionStartId="5.[151,250,1499,1521]" captionTargetBox="[153,1422,188,1471]" captionTargetId="figure@5.[151,1424,186,1474]" captionTargetPageId="5" captionText="FIGURE 2. Lygdamis nasutus n. sp., paratype, AM W. 47750, live. A. Whole specimen, dorsal view, arrowheads point to notopodial cirri; B. Anterior end, lateral view, with median organ protruding from operculum (arrow); C. Anterior end, dorsal view; D. Operculum, frontal view; E. Posterior abdominal chaetigers and cauda, ventral view. Photos by A. Semenov." httpUri="https://zenodo.org/record/245141/files/figure.png" pageId="11" pageNumber="195">Figs 2</figureCitation>
|
||
E, 7G).
|
||
</paragraph>
|
||
<paragraph id="8BC136A7FFF32C5FFF72A858FBDEFDE2" blockId="11.[151,1437,151,1185]" pageId="11" pageNumber="195">
|
||
<emphasis id="B90AEAB5FFF32C5FFF72A858FF14FDBE" box="[199,241,549,572]" italics="true" pageId="11" pageNumber="195">Gut</emphasis>
|
||
. Straight tube running from mouth to anus, with a broadened structure or ‘proventriculus’ between the last parathoracic segment and the first abdominal. Not obvious in preserved specimens.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C364652CFFF32C5AFF72A811FAD3FDA2" lastPageId="14" lastPageNumber="198" pageId="11" pageNumber="195" type="discussion">
|
||
<paragraph id="8BC136A7FFF32C5FFF72A811FC7DFC97" blockId="11.[151,1437,151,1185]" pageId="11" pageNumber="195">
|
||
<emphasis id="B90AEAB5FFF32C5FFF72A811FEDEFD07" bold="true" box="[199,315,620,645]" pageId="11" pageNumber="195">Remarks.</emphasis>
|
||
<taxonomicName id="4C7E4D24FFF32C5FFEF1A811FDEBFD06" box="[324,526,620,645]" class="Polychaeta" family="Sabellariidae" genus="Lygdamis" kingdom="Animalia" order="Sabellida" pageId="11" pageNumber="195" phylum="Annelida" rank="species" species="nasutus" status="sp. nov.">
|
||
<emphasis id="B90AEAB5FFF32C5FFEF1A811FDEBFD06" box="[324,526,620,645]" italics="true" pageId="11" pageNumber="195">Lygdamis nasutus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A23957CEFFF32C5FFDA2A810FDB3FD06" box="[535,598,621,644]" pageId="11" pageNumber="195" rank="species">n. sp.</taxonomicNameLabel>
|
||
is distinguished from its congeners by a unique combination of features: well-developed cylindrical median organ distally pigmented, with a flattened, teardrop corona, outer paleae (18–19 pairs) with asymmetrical pointed tips and subtle thecal sculpture, inner paleae (7–8 pairs) with blunt tips and smooth surface, three elongated and pointed lateral lobes on chaetiger 2, and abdominal chaetigers with two
|
||
<typeStatus id="54C58805FFF32C5FFAD4A8A5FA79FD72" box="[1377,1436,728,752]" pageId="11" pageNumber="195">types</typeStatus>
|
||
of neurochaetae and notopodial uncini with three or four distal teeth.
|
||
</paragraph>
|
||
<paragraph id="8BC136A7FFF32C5FFF72A962FA89FBB6" blockId="11.[151,1437,151,1185]" pageId="11" pageNumber="195">
|
||
The three species of
|
||
<taxonomicName id="4C7E4D24FFF32C5FFE03A962FDC7FCBA" box="[438,546,799,824]" class="Polychaeta" family="Sabellariidae" genus="Lygdamis" kingdom="Animalia" order="Sabellida" pageId="11" pageNumber="195" phylum="Annelida" rank="genus">
|
||
<emphasis id="B90AEAB5FFF32C5FFE03A962FDC7FCBA" box="[438,546,799,824]" italics="true" pageId="11" pageNumber="195">Lygdamis</emphasis>
|
||
</taxonomicName>
|
||
previously reported in
|
||
<collectingCountry id="F3697637FFF32C5FFC9BA962FC70FCBA" box="[814,917,799,824]" name="Australia" pageId="11" pageNumber="195">Australia</collectingCountry>
|
||
are characterised by a small or medium size, conical and unpigmented median organ. In contrast, the new species bears a conspicuous, well-developed, cylindrical and strongly pigmented median organ.
|
||
<taxonomicName id="4C7E4D24FFF32C5FFD7EA91AFC76FC02" box="[715,915,871,896]" class="Polychaeta" family="Sabellariidae" genus="Lygdamis" kingdom="Animalia" order="Sabellida" pageId="11" pageNumber="195" phylum="Annelida" rank="species" species="nasutus" status="sp. nov.">
|
||
<emphasis id="B90AEAB5FFF32C5FFD7EA91AFC76FC02" box="[715,915,871,896]" italics="true" pageId="11" pageNumber="195">Lygdamis nasutus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A23957CEFFF32C5FFC2EA915FC32FCFD" box="[923,983,872,895]" pageId="11" pageNumber="195" rank="species">n. sp.</taxonomicNameLabel>
|
||
is provided with rows of eyespots along both sides of the median organ, whereas
|
||
<taxonomicName id="4C7E4D24FFF32C5FFDCDA9F0FC7BFC26" authority="Kirtley, 1994" authorityName="Kirtley" authorityYear="1994" box="[632,926,908,933]" class="Polychaeta" family="Sabellariidae" genus="Lygdamis" kingdom="Animalia" order="Sabellida" pageId="11" pageNumber="195" phylum="Annelida" rank="species" species="augeneri">
|
||
<emphasis id="B90AEAB5FFF32C5FFDCDA9F0FD18FC26" box="[632,765,909,932]" italics="true" pageId="11" pageNumber="195">L. augeneri</emphasis>
|
||
<bibRefCitation id="EFEF4B56FFF32C5FFCB2A9F1FC7BFC26" author="Kirtley" box="[775,926,908,933]" pageId="11" pageNumber="195" refString="Kirtley, D. W. (1994) A review and taxonomic revision of the family Sabellariidae Johnston, 1865 (Annelida; Polychaeta). Sabecon Press Science Series, 1, 1 - 223." type="journal article" year="1994">Kirtley, 1994</bibRefCitation>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C7E4D24FFF32C5FFC69A9F0FAF1FC27" authority="McIntosh, 1885" authorityName="McIntosh" authorityYear="1885" box="[988,1300,908,933]" class="Polychaeta" family="Sabellariidae" genus="Lygdamis" kingdom="Animalia" order="Sabellida" pageId="11" pageNumber="195" phylum="Annelida" rank="species" species="giardi">
|
||
<emphasis id="B90AEAB5FFF32C5FFC69A9F0FBDAFC27" box="[988,1087,908,933]" italics="true" pageId="11" pageNumber="195">L. giardi</emphasis>
|
||
(
|
||
<bibRefCitation id="EFEF4B56FFF32C5FFBE7A9F1FAE9FC27" author="McIntosh" box="[1106,1292,908,933]" pageId="11" pageNumber="195" refString="McIntosh, W. C. (1885) Report on the Annelida Polychaeta collected by H. M. S. ' Challenger' during the years 1873 - 76. Report on the Scientific Results of the Voyage of H. M. S. Challenger, Zoology, 12, 1 - 554." type="journal article" year="1885">McIntosh, 1885</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
, two of the species previously reported from
|
||
<collectingCountry id="F3697637FFF32C5FFDA6A9D2FD99FC4A" box="[531,636,943,968]" name="Australia" pageId="11" pageNumber="195">Australia</collectingCountry>
|
||
, bear eyes on the median ridge, not the median organ (free distal end) (Hutchings
|
||
<emphasis id="B90AEAB5FFF32C5FFEAEA9A8FEB6FC6F" box="[283,339,980,1005]" italics="true" pageId="11" pageNumber="195">et al.</emphasis>
|
||
2012:
|
||
<figureCitation id="13452A22FFF32C5FFE17A9A9FE02FC6E" box="[418,487,980,1005]" captionStart="FIGURE 8" captionStartId="15.[151,250,1815,1837]" captionTargetBox="[190,1395,626,1787]" captionTargetId="figure@15.[188,1399,622,1794]" captionTargetPageId="15" captionText="FIGURE 8. Median organ and ridge diversity in Australian Idanthyrsus. A. Idanthyrsus australiensis, AM W. 201770, with conical median organ (MO), eyespots (white arrow) along median ridge (MR); B. Idanthyrsus sp. cf. I. australiensis AM W. 29153, with flattened, triangular median organ; eyespots absent along median ridge; C. Idanthyrsus sp. cf. I. australiensis, AM W. 29125 with flattened, triangular median organ, eyespots present along median ridge; D. Idanthyrsus willora, scanning electron micrograph, with inconspicuous median organ." httpUri="https://zenodo.org/record/245147/files/figure.png" pageId="11" pageNumber="195">Figs 8</figureCitation>
|
||
A, B), and in the third one,
|
||
<taxonomicName id="4C7E4D24FFF32C5FFC95A9A8FB45FC6E" authority="Hutchings et al., 2012" authorityName="Hutchings et al." authorityYear="2012" box="[800,1184,980,1005]" class="Polychaeta" family="Sabellariidae" genus="Lygdamis" kingdom="Animalia" order="Sabellida" pageId="11" pageNumber="195" phylum="Annelida" rank="species" species="wambiri">
|
||
<emphasis id="B90AEAB5FFF32C5FFC95A9A8FC7EFC6F" box="[800,923,980,1005]" italics="true" pageId="11" pageNumber="195">L. wambiri</emphasis>
|
||
Hutchings
|
||
<emphasis id="B90AEAB5FFF32C5FFBA8A9A8FBB3FC6F" box="[1053,1110,980,1005]" italics="true" pageId="11" pageNumber="195">et al.</emphasis>
|
||
, 2012
|
||
</taxonomicName>
|
||
, eyespots seem absent both on the median organ and median ridge. Moreover,
|
||
<taxonomicName id="4C7E4D24FFF32C5FFCA2A984FC7FFB92" box="[791,922,1017,1040]" class="Polychaeta" family="Sabellariidae" genus="Lygdamis" kingdom="Animalia" order="Sabellida" pageId="11" pageNumber="195" phylum="Annelida" rank="species" species="augeneri">
|
||
<emphasis id="B90AEAB5FFF32C5FFCA2A984FC7FFB92" box="[791,922,1017,1040]" italics="true" pageId="11" pageNumber="195">L. augeneri</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C7E4D24FFF32C5FFC61A984FBD3FB92" box="[980,1078,1015,1040]" class="Polychaeta" family="Sabellariidae" genus="Lygdamis" kingdom="Animalia" order="Sabellida" pageId="11" pageNumber="195" phylum="Annelida" rank="species" species="giardi">
|
||
<emphasis id="B90AEAB5FFF32C5FFC61A984FBD3FB92" box="[980,1078,1015,1040]" italics="true" pageId="11" pageNumber="195">L. giardi</emphasis>
|
||
</taxonomicName>
|
||
bear symmetrical outer paleae, unlike the new species. Other differences between the Australian species of
|
||
<taxonomicName id="4C7E4D24FFF32C5FFC5FAE61FBB3FBB7" box="[1002,1110,1052,1077]" class="Polychaeta" family="Sabellariidae" genus="Lygdamis" kingdom="Animalia" order="Sabellida" pageId="11" pageNumber="195" phylum="Annelida" rank="genus">
|
||
<emphasis id="B90AEAB5FFF32C5FFC5FAE61FBB3FBB7" box="[1002,1110,1052,1077]" italics="true" pageId="11" pageNumber="195">Lygdamis</emphasis>
|
||
</taxonomicName>
|
||
can be found in
|
||
<tableCitation id="C6FC031CFFF32C5FFAA7AE61FA83FBB6" box="[1298,1382,1052,1077]" captionStart="TABLE 1" captionStartId="12.[151,235,1088,1110]" captionText="TABLE 1. Comparison of morphological traits in Lygdamis species reported from Australia" pageId="11" pageNumber="195">Table 1</tableCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BC136A7FFF32C5FFF72AE42FAF2FB22" blockId="11.[151,1437,151,1185]" pageId="11" pageNumber="195">
|
||
There are several attributes that are outstanding in
|
||
<taxonomicName id="4C7E4D24FFF32C5FFCA5AE3CFC63FBDA" box="[784,902,1089,1112]" class="Polychaeta" family="Sabellariidae" genus="Lygdamis" kingdom="Animalia" order="Sabellida" pageId="11" pageNumber="195" phylum="Annelida" rank="species" species="nasutus" status="sp. nov.">
|
||
<emphasis id="B90AEAB5FFF32C5FFCA5AE3CFC63FBDA" box="[784,902,1089,1112]" italics="true" pageId="11" pageNumber="195">L. nasutus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A23957CEFFF32C5FFC3AAE3DFC2AFBD5" box="[911,975,1088,1111]" pageId="11" pageNumber="195" rank="species">n. sp.</taxonomicNameLabel>
|
||
, but seem not to have been noticed and described in other congeners. A comprehensive revision of these features in
|
||
<taxonomicName id="4C7E4D24FFF32C5FFC53AE19FBB4FBFF" box="[998,1105,1124,1149]" class="Polychaeta" family="Sabellariidae" genus="Lygdamis" kingdom="Animalia" order="Sabellida" pageId="11" pageNumber="195" phylum="Annelida" rank="genus">
|
||
<emphasis id="B90AEAB5FFF32C5FFC53AE19FBB4FBFF" box="[998,1105,1124,1149]" italics="true" pageId="11" pageNumber="195">Lygdamis</emphasis>
|
||
</taxonomicName>
|
||
species and other sabellariids may provide informative characters for taxonomic purposes and phylogenetic reconstruction. These are:
|
||
</paragraph>
|
||
<paragraph id="8BC136A7FFF32C5FFF22AEADFD3EFACF" blockId="11.[151,1437,1231,2023]" pageId="11" pageNumber="195">
|
||
1) The new species has small and conical cirri on the distal dorsal margin of parathoracic notopodia (
|
||
<figureCitation id="13452A22FFF32C5FFA8EAEB2FA66FB6A" box="[1339,1411,1231,1256]" captionStart="FIGURE 2" captionStartId="5.[151,250,1499,1521]" captionTargetBox="[153,1422,188,1471]" captionTargetId="figure@5.[151,1424,186,1474]" captionTargetPageId="5" captionText="FIGURE 2. Lygdamis nasutus n. sp., paratype, AM W. 47750, live. A. Whole specimen, dorsal view, arrowheads point to notopodial cirri; B. Anterior end, lateral view, with median organ protruding from operculum (arrow); C. Anterior end, dorsal view; D. Operculum, frontal view; E. Posterior abdominal chaetigers and cauda, ventral view. Photos by A. Semenov." httpUri="https://zenodo.org/record/245141/files/figure.png" pageId="11" pageNumber="195">Figs 2</figureCitation>
|
||
B, 3A). These structures have also been described in other
|
||
<taxonomicName id="4C7E4D24FFF32C5FFCDAAE8CFC3EFA88" box="[879,987,1265,1290]" class="Polychaeta" family="Sabellariidae" genus="Lygdamis" kingdom="Animalia" order="Sabellida" pageId="11" pageNumber="195" phylum="Annelida" rank="genus">
|
||
<emphasis id="B90AEAB5FFF32C5FFCDAAE8CFC3EFA88" box="[879,987,1265,1290]" italics="true" pageId="11" pageNumber="195">Lygdamis</emphasis>
|
||
</taxonomicName>
|
||
species (i.e.,
|
||
<taxonomicName id="4C7E4D24FFF32C5FFBC9AE8FFE27FAA9" authority="Kinberg, 1867 sensu Kirtley, 1994" authorityName="Kinberg, 1867 sensu Kirtley" authorityYear="1994" class="Polychaeta" family="Sabellariidae" genus="Lygdamis" kingdom="Animalia" order="Sabellida" pageId="11" pageNumber="195" phylum="Annelida" rank="species" species="indicus">
|
||
<emphasis id="B90AEAB5FFF32C5FFBC9AE8FFB0BFA88" box="[1148,1262,1265,1290]" italics="true" pageId="11" pageNumber="195">L. indicus</emphasis>
|
||
<bibRefCitation id="EFEF4B56FFF32C5FFB42AE8CFA79FA88" author="Kinberg" box="[1271,1436,1265,1290]" pageId="11" pageNumber="195" refString="Kinberg, J. G. H. (1867) Annulata nova. Ofversigt af Koniglich Vetenskapsakademiens forhandlingar, Stockholm, 23, 337 - 357." type="journal article" year="1867">Kinberg, 1867</bibRefCitation>
|
||
<emphasis id="B90AEAB5FFF32C5FFF54AF69FEC5FAA9" box="[225,288,1300,1323]" italics="true" pageId="11" pageNumber="195">sensu</emphasis>
|
||
<bibRefCitation id="EFEF4B56FFF32C5FFE9CAF6FFE27FAA9" author="Kirtley" box="[297,450,1298,1323]" pageId="11" pageNumber="195" refString="Kirtley, D. W. (1994) A review and taxonomic revision of the family Sabellariidae Johnston, 1865 (Annelida; Polychaeta). Sabecon Press Science Series, 1, 1 - 223." type="journal article" year="1994">Kirtley, 1994</bibRefCitation>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C7E4D24FFF32C5FFE65AF69FC75FAA9" authority="Lechapt & Kirtley, 1998" authorityName="Lechapt & Kirtley" authorityYear="1998" box="[464,912,1298,1323]" class="Polychaeta" family="Sabellariidae" genus="Lygdamis" kingdom="Animalia" order="Sabellida" pageId="11" pageNumber="195" phylum="Annelida" rank="species" species="splendidus">
|
||
<emphasis id="B90AEAB5FFF32C5FFE65AF69FD8CFAA9" box="[464,617,1298,1323]" italics="true" pageId="11" pageNumber="195">L. splendidus</emphasis>
|
||
<bibRefCitation id="EFEF4B56FFF32C5FFDC6AF6FFC75FAA9" author="Lechapt" box="[627,912,1298,1323]" pageId="11" pageNumber="195" refString="Lechapt, J-P. & Kirtley, D. W. (1998) New species of bathyal and abyssal Sabellariidae (Annelida: Polychaeta) from near New Caledonia (southwest Pacific Ocean). Proceedings of the Biological Society of Washington, 111, 807 - 822." type="journal article" year="1998">Lechapt & Kirtley, 1998</bibRefCitation>
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="4C7E4D24FFF32C5FFC65AF69FA83FAA9" authority="Nishi & Kirtley, 1999" authorityName="Nishi & Kirtley" authorityYear="1999" box="[976,1382,1298,1323]" class="Polychaeta" family="Sabellariidae" genus="Lygdamis" kingdom="Animalia" order="Sabellida" pageId="11" pageNumber="195" phylum="Annelida" rank="species" species="japonicus">
|
||
<emphasis id="B90AEAB5FFF32C5FFC65AF69FBBBFAA9" box="[976,1118,1300,1323]" italics="true" pageId="11" pageNumber="195">L. japonicus</emphasis>
|
||
<bibRefCitation id="EFEF4B56FFF32C5FFBDDAF6FFA83FAA9" author="Nishi" box="[1128,1382,1298,1323]" pageId="11" pageNumber="195" refString="Nishi, E. & Kirtley, D. W. (1999) Three new species of Sabellariidae (Polychaeta) from Japan. Natural History Research, 5, 93 - 105." type="journal article" year="1999">Nishi & Kirtley, 1999</bibRefCitation>
|
||
</taxonomicName>
|
||
) but have often not been reported in the literature.
|
||
</paragraph>
|
||
<paragraph id="8BC136A7FFF32C5FFF22AF2BFED0FA71" blockId="11.[151,1437,1231,2023]" pageId="11" pageNumber="195">
|
||
2)
|
||
<taxonomicName id="4C7E4D24FFF32C5FFF54AF28FE4FFAEF" box="[225,426,1365,1390]" class="Polychaeta" family="Sabellariidae" genus="Lygdamis" kingdom="Animalia" order="Sabellida" pageId="11" pageNumber="195" phylum="Annelida" rank="species" species="nasutus" status="sp. nov.">
|
||
<emphasis id="B90AEAB5FFF32C5FFF54AF28FE4FFAEF" box="[225,426,1365,1390]" italics="true" pageId="11" pageNumber="195">Lygdamis nasutus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A23957CEFFF32C5FFE04AF2BFE08FAEF" box="[433,493,1366,1389]" pageId="11" pageNumber="195" rank="species">n. sp.</taxonomicNameLabel>
|
||
bears chaetae of two different morphologies on abdominal neuropodia (
|
||
<figureCitation id="13452A22FFF32C5FFAA3AF28FAB8FAEC" box="[1302,1373,1365,1390]" captionStart="FIGURE 5" captionStartId="8.[151,250,1820,1842]" captionTargetBox="[191,1407,198,1798]" captionTargetId="figure@8.[187,1408,193,1805]" captionTargetPageId="8" captionText="FIGURE 5. Chaetae, Lygdamis nasutus n. sp., paratypes. A. Outer paleae; B. Detail of tips of outer paleae; C. Inner paleae, distal end; D. Notopodium, parathoracic chaetiger; E. Detail of tips of lanceolate parathoracic notopodial chaeta; F. Parathoracic notopodial simple capillaries, interspersed with lanceolate chaetae; G. Neuropodium, parathoracic chaetiger; H. Detail of tips of lanceolate parathoracic neuropodial chaetae; I. Parathoracic neuropodial simple capillaries, interspersed with lanceolate chaetae; J. Abdominal notochaetae (two types); K. Neuropodial abdominal uncini; L. Detail of abdominal uncinus, side view." httpUri="https://zenodo.org/record/245144/files/figure.png" pageId="11" pageNumber="195">Figs 5</figureCitation>
|
||
J, 7E, F), similar to those described in
|
||
<taxonomicName id="4C7E4D24FFF32C5FFDFDAF0AFC7DFA0D" authority="Johansson, 1922" authorityName="Johansson" authorityYear="1922" box="[584,920,1398,1423]" class="Polychaeta" family="Sabellariidae" genus="Lygdamis" kingdom="Animalia" order="Sabellida" pageId="11" pageNumber="195" phylum="Annelida" rank="species" species="curvatus">
|
||
<emphasis id="B90AEAB5FFF32C5FFDFDAF0AFD23FA0C" box="[584,710,1399,1422]" italics="true" pageId="11" pageNumber="195">L. curvatus</emphasis>
|
||
(
|
||
<bibRefCitation id="EFEF4B56FFF32C5FFD61AF0BFC75FA0D" author="Johansson" box="[724,912,1398,1423]" pageId="11" pageNumber="195" refString="Johansson, K. E. (1922) On some new tubicolous annelids from Japan, the Bonin Islands and the Antarctic. Arkiv for Zoologi, 15, 1 - 11." type="journal article" year="1922">Johansson, 1922</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
and also
|
||
<taxonomicName id="4C7E4D24FFF32C5FFBB7AF0BFE9DFA32" authority="Lechapt & Kirtley, 1996" authorityName="Lechapt & Kirtley" authorityYear="1996" class="Polychaeta" family="Sabellariidae" genus="Bathysabellaria" kingdom="Animalia" order="Sabellida" pageId="11" pageNumber="195" phylum="Annelida" rank="species" species="spinifera">
|
||
<emphasis id="B90AEAB5FFF32C5FFBB7AF0BFAFAFA0D" box="[1026,1311,1398,1423]" italics="true" pageId="11" pageNumber="195">Bathysabellaria spinifera</emphasis>
|
||
<bibRefCitation id="EFEF4B56FFF32C5FFA90AF0BFE9DFA32" author="Lechapt" pageId="11" pageNumber="195" refString="Lechapt, J-P. & Kirtley, D. W. (1996) Bathysabellaria spinifera (Polychaeta: Sabellariidae), a new species from deep water off New Caledonia, Southwest Pacific Ocean. Proceedings of the Biological Society of Washington, 109, 560 - 574." type="journal article" year="1996">Lechapt & Kirtley, 1996</bibRefCitation>
|
||
</taxonomicName>
|
||
. Remaining
|
||
<taxonomicName id="4C7E4D24FFF32C5FFDB1AFEAFD95FA32" box="[516,624,1431,1456]" class="Polychaeta" family="Sabellariidae" genus="Lygdamis" kingdom="Animalia" order="Sabellida" pageId="11" pageNumber="195" phylum="Annelida" rank="genus">
|
||
<emphasis id="B90AEAB5FFF32C5FFDB1AFEAFD95FA32" box="[516,624,1431,1456]" italics="true" pageId="11" pageNumber="195">Lygdamis</emphasis>
|
||
</taxonomicName>
|
||
species have been described with a single
|
||
<typeStatus id="54C58805FFF32C5FFBFBAFE4FB9BFA33" box="[1102,1150,1433,1457]" pageId="11" pageNumber="195">type</typeStatus>
|
||
of abdominal neurochaetae (i.e., the other Australian species), but it is probable that in other species this feature has been overlooked.
|
||
</paragraph>
|
||
<paragraph id="8BC136A7FFF32C5FFF22AF81FE22F9FA" blockId="11.[151,1437,1231,2023]" pageId="11" pageNumber="195">
|
||
3) Posterior abdominal chaetigers in
|
||
<taxonomicName id="4C7E4D24FFF32C5FFDD7AF80FD33F996" box="[610,726,1533,1556]" class="Polychaeta" family="Sabellariidae" genus="Lygdamis" kingdom="Animalia" order="Sabellida" pageId="11" pageNumber="195" phylum="Annelida" rank="species" species="nasutus" status="sp. nov.">
|
||
<emphasis id="B90AEAB5FFF32C5FFDD7AF80FD33F996" box="[610,726,1533,1556]" italics="true" pageId="11" pageNumber="195">L. nasutus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A23957CEFFF32C5FFD68AF80FCFFF996" box="[733,794,1533,1556]" pageId="11" pageNumber="195" rank="species">n. sp.</taxonomicNameLabel>
|
||
specimens examined lack uncini in their notopodial tori. Since new abdominal segments are added between the abdomen and the cauda during development (
|
||
<bibRefCitation id="EFEF4B56FFF32C5FFAE7AC60FE8FF9D5" author="Eckelbarger" pageId="11" pageNumber="195" refString="Eckelbarger, K. J. (1975) Developmental studies of the postsettling stages of Sabellaria vulgaris (Polychaeta: Sabellariidae). Marine Biology, 30, 137 - 149. http: // dx. doi. org / 10.1007 / BF 00391588" type="journal article" year="1975">Eckelbarger 1975</bibRefCitation>
|
||
), it is possible that uncini are formed after the addition of chaetigers, but it could also be a species specific feature.
|
||
</paragraph>
|
||
<paragraph id="8BC136A7FFF32C58FF22ACFFFE41FED5" blockId="11.[151,1437,1231,2023]" lastBlockId="12.[151,1437,151,1043]" lastPageId="12" lastPageNumber="196" pageId="11" pageNumber="195">
|
||
4) Abdominal uncini are generally described as having two longitudinal rows of teeth (
|
||
<bibRefCitation id="EFEF4B56FFF32C5FFB05ACFFFA72F918" author="Lana" box="[1200,1431,1666,1690]" pageId="11" pageNumber="195" refString="Lana, P. C. & Gruet, Y. (1989) Sabellaria wilsoni sp. n. (Polychaeta, Sabellariidae) from the southeast coast of Brazil. Zoologica Scripta, 18, 239 - 244. http: // dx. doi. org / 10.1111 / j. 1463 - 6409.1989. tb 00449. x" type="journal article" year="1989">Lana & Gruet 1989</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFEF4B56FFF32C5FFF54ACDFFE09F939" author="Lechapt" box="[225,492,1698,1723]" pageId="11" pageNumber="195" refString="Lechapt, J-P. & Gruet, Y. (1993) Bathysabellaria neocaledoniensis, a new genus and species of Sabellariidae (Annelida, Polychaeta) from the bathyal zones off New Caledonia (Southwest Pacific Ocean). Zoologica Scripta, 22, 243 - 247. http: // dx. doi. org / 10.1111 / j. 1463 - 6409.1993. tb 00355. x" type="journal article" year="1993">Lechapt & Gruet 1993</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFEF4B56FFF32C5FFE49ACDFFD77F939" author="Kirtley" box="[508,658,1698,1723]" pageId="11" pageNumber="195" refString="Kirtley, D. W. (1994) A review and taxonomic revision of the family Sabellariidae Johnston, 1865 (Annelida; Polychaeta). Sabecon Press Science Series, 1, 1 - 223." type="journal article" year="1994">Kirtley 1994</bibRefCitation>
|
||
). However, variations to this typical pattern have been found after reviewing the literature, suggesting it has not always been studied in detail. For instance,
|
||
<taxonomicName id="4C7E4D24FFF32C5FFB62ACB9FE61F97C" authority="Kinberg, 1867" authorityName="Kinberg" authorityYear="1867" class="Polychaeta" family="Sabellariidae" genus="Lygdamis" kingdom="Animalia" order="Sabellida" pageId="11" pageNumber="195" phylum="Annelida" rank="species" species="indicus">
|
||
<emphasis id="B90AEAB5FFF32C5FFB62ACB9FA79F95F" box="[1239,1436,1732,1757]" italics="true" pageId="11" pageNumber="195">Lygdamis indicus</emphasis>
|
||
<bibRefCitation id="EFEF4B56FFF32C5FFF54AC98FE61F97C" author="Kinberg" box="[225,388,1765,1790]" pageId="11" pageNumber="195" refString="Kinberg, J. G. H. (1867) Annulata nova. Ofversigt af Koniglich Vetenskapsakademiens forhandlingar, Stockholm, 23, 337 - 357." type="journal article" year="1867">Kinberg, 1867</bibRefCitation>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C7E4D24FFF32C5FFE25AC9BFE14F97C" box="[400,497,1765,1790]" class="Polychaeta" family="Sabellariidae" genus="Lygdamis" kingdom="Animalia" order="Sabellida" pageId="11" pageNumber="195" phylum="Annelida" rank="species" species="giardi">
|
||
<emphasis id="B90AEAB5FFF32C5FFE25AC9BFE14F97C" box="[400,497,1765,1790]" italics="true" pageId="11" pageNumber="195">L. giardi</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C7E4D24FFF32C5FFE4AAC9BFDB8F97F" box="[511,605,1766,1789]" class="Polychaeta" family="Sabellariidae" genus="Lygdamis" kingdom="Animalia" order="Sabellida" pageId="11" pageNumber="195" phylum="Annelida" rank="species" species="wirtzi">
|
||
<emphasis id="B90AEAB5FFF32C5FFE4AAC9BFDB8F97F" box="[511,605,1766,1789]" italics="true" pageId="11" pageNumber="195">L. wirtzi</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C7E4D24FFF32C5FFD21AC9BFCE2F97F" box="[660,775,1766,1789]" class="Polychaeta" family="Sabellariidae" genus="Lygdamis" kingdom="Animalia" order="Sabellida" pageId="11" pageNumber="195" phylum="Annelida" rank="species" species="nasutus" status="sp. nov.">
|
||
<emphasis id="B90AEAB5FFF32C5FFD21AC9BFCE2F97F" box="[660,775,1766,1789]" italics="true" pageId="11" pageNumber="195">L. nasutus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A23957CEFFF32C5FFCBBAC9BFCAFF97F" box="[782,842,1766,1789]" pageId="11" pageNumber="195" rank="species">n. sp.</taxonomicNameLabel>
|
||
have up to three or four longitudinal rows of teeth at the distal end of the uncini. Moreover,
|
||
<taxonomicName id="4C7E4D24FFF32C5FFD2FAD7BFB98F89D" authority="Nishi & Kirtley, 1999" authorityName="Nishi & Kirtley" authorityYear="1999" box="[666,1149,1798,1823]" class="Polychaeta" family="Sabellariidae" genus="Lygdamis" kingdom="Animalia" order="Sabellida" pageId="11" pageNumber="195" phylum="Annelida" rank="species" species="japonicus">
|
||
<emphasis id="B90AEAB5FFF32C5FFD2FAD7BFC9EF89C" box="[666,891,1798,1823]" italics="true" pageId="11" pageNumber="195">Lygdamis japonicus</emphasis>
|
||
<bibRefCitation id="EFEF4B56FFF32C5FFC36AD7BFB98F89D" author="Nishi" box="[899,1149,1798,1823]" pageId="11" pageNumber="195" refString="Nishi, E. & Kirtley, D. W. (1999) Three new species of Sabellariidae (Polychaeta) from Japan. Natural History Research, 5, 93 - 105." type="journal article" year="1999">Nishi & Kirtley, 1999</bibRefCitation>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C7E4D24FFF32C5FFB3CAD7AFAEFF89C" box="[1161,1290,1799,1822]" class="Polychaeta" family="Sabellariidae" genus="Lygdamis" kingdom="Animalia" order="Sabellida" pageId="11" pageNumber="195" phylum="Annelida" rank="species" species="augeneri">
|
||
<emphasis id="B90AEAB5FFF32C5FFB3CAD7AFAEFF89C" box="[1161,1290,1799,1822]" italics="true" pageId="11" pageNumber="195">L. augeneri</emphasis>
|
||
</taxonomicName>
|
||
and the new species present a single tooth proximally. This structural variation in uncini has not always been formally described but is clearly visible in species illustrations (e.g.,
|
||
<bibRefCitation id="EFEF4B56FFF32C5FFCCFAD34FBDEF8E0" author="Uebelacker" box="[890,1083,1865,1890]" pageId="11" pageNumber="195" refString="Uebelacker, J. M. (1984) Chapter 49. Family Sabellariidae. In: Uebelacker, J. M. & Johnson, P. G. (Eds.), Taxonomic guide to the polychaetes of the northern Gulf of Mexico. Vol. 9. Barry A. Vittor & Associates, Inc., Mobile, AL, pp. 1 - 49." type="book" year="1984">Uebelacker 1984</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFEF4B56FFF32C5FFBF2AD34FAD2F8E0" author="Nishi" box="[1095,1335,1865,1890]" pageId="11" pageNumber="195" refString="Nishi, E. & Kirtley, D. W. (1999) Three new species of Sabellariidae (Polychaeta) from Japan. Natural History Research, 5, 93 - 105." type="journal article" year="1999">Nishi & Kirtley 1999</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFEF4B56FFF32C5FFAF6AD34FE95F801" author="Nishi" pageId="11" pageNumber="195" refString="Nishi, E. & Nunez, J. (1999) A new species of shallow water Sabellariidae (Annelida: Polychaeta) from Madeira Island, Portugal, and Canary Islands, Spain. Arquipelago Boletim da Universidade dos Acores Ciencias Biologicas e Marinhas, 17 a, 37 - 42." type="book chapter" year="1999">Nishi & Núñez 1999</bibRefCitation>
|
||
; Hutchings
|
||
<emphasis id="B90AEAB5FFF32C5FFE4FAD11FDD0F801" box="[506,565,1898,1923]" italics="true" pageId="11" pageNumber="195">et al.</emphasis>
|
||
2012). Other sabellariids, for example
|
||
<emphasis id="B90AEAB5FFF32C5FFC48AD17FAFAF801" box="[1021,1311,1898,1923]" italics="true" pageId="11" pageNumber="195">
|
||
<taxonomicName id="4C7E4D24FFF32C5FFC48AD17FAFEF801" box="[1021,1307,1898,1923]" class="Polychaeta" family="Sabellariidae" genus="Phalacrostemma" kingdom="Animalia" order="Sabellida" pageId="11" pageNumber="195" phylum="Annelida" rank="species" species="maloga">Phalacrostemma maloga</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
have been described as having uncini with three rows of teeth (Hutchings
|
||
<emphasis id="B90AEAB5FFF32C5FFC1EADF0FC01F827" box="[939,996,1932,1957]" italics="true" pageId="11" pageNumber="195">et al.</emphasis>
|
||
2012). Moreover,
|
||
<taxonomicName id="4C7E4D24FFF32C5FFB03ADF1FA9AF826" box="[1206,1407,1932,1957]" class="Polychaeta" family="Sabellariidae" genus="Lygdamis" kingdom="Animalia" order="Sabellida" pageId="11" pageNumber="195" phylum="Annelida" rank="species" species="nasutus" status="sp. nov.">
|
||
<emphasis id="B90AEAB5FFF32C5FFB03ADF1FA9AF826" box="[1206,1407,1932,1957]" italics="true" pageId="11" pageNumber="195">Lygdamis nasutus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A23957CEFFF32C5FFA32ADF0FEE4F847" pageId="11" pageNumber="195" rank="species">n. sp.</taxonomicNameLabel>
|
||
shows a large variation range in the morphology of uncini along the abdominal chaetigers, a character not often commented in the literature. The structural complexity in morphology of uncini is due to the development and linked to differences in chaetoblasts and chaetogenesis (
|
||
<bibRefCitation id="EFEF4B56FFF42C58FC69AAEAFB17FF32" author="O'Clair" box="[988,1266,151,176]" pageId="12" pageNumber="196" refString="O'Clair, R. M. & Cloney, R. A. (1974) Patterns of morphogenesis mediated by dynamic microvilli: chaetogenesis in Nereis vexillosa. Cell and Tissue Research, 151 (2), 141 - 157." type="journal article" year="1974">O’Clair & Cloney 1974</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFEF4B56FFF42C58FAB4AAEAFEF9FF50" author="Bartolomaeus" pageId="12" pageNumber="196" refString="Bartolomaeus, T. (1995) Structure and formation of the uncini in Pectinaria koreni, Pectinaria auricoma (Terebellida) and Spirorbis spirorbis (Sabellida): implications for annelid phylogeny and the position of the Pogonophora. Zoomorphology, 115 (3), 161 - 177." type="journal article" year="1995">Bartolomaeus 1995</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="EFEF4B56FFF42C58FE93AAC4FE84FF53" author="Bartolomaeus" box="[294,353,185,209]" pageId="12" pageNumber="196" refString="Bartolomaeus, T. (2002) Structure and formation of thoracic and abdominal uncini in Fabricia stellaris (Muller, 1774) - Implication for the evolution of Sabellida (Annelida). Zoologischer Anzeiger, 241, 1 - 17." type="journal article" year="2002">2002</bibRefCitation>
|
||
; Hausen 2005) and is considered a diagnostic feature among other polychaete families (e.g.,
|
||
<bibRefCitation id="EFEF4B56FFF42C58FADEAAC7FE72FF71" author="Bartolomaeus" pageId="12" pageNumber="196" refString="Bartolomaeus, T. (1995) Structure and formation of the uncini in Pectinaria koreni, Pectinaria auricoma (Terebellida) and Spirorbis spirorbis (Sabellida): implications for annelid phylogeny and the position of the Pogonophora. Zoomorphology, 115 (3), 161 - 177." type="journal article" year="1995">Bartolomaeus 1995</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="EFEF4B56FFF42C58FE16AAA6FE3AFF71" author="Bartolomaeus" box="[419,479,219,243]" pageId="12" pageNumber="196" refString="Bartolomaeus, T. (2002) Structure and formation of thoracic and abdominal uncini in Fabricia stellaris (Muller, 1774) - Implication for the evolution of Sabellida (Annelida). Zoologischer Anzeiger, 241, 1 - 17." type="journal article" year="2002">2002</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFEF4B56FFF42C58FE5EAAA7FCE6FF71" author="Garraffoni" box="[491,771,218,243]" pageId="12" pageNumber="196" refString="Garraffoni, A. R. S. & Lana, P. D. C. (2004) Cladistic analysis of the subfamily Trichobranchinae (Polychaeta: Terebellidae). Journal of the Marine Biological Association of the United Kingdom, 84 (5), 973 - 982. http: // dx. doi. org / 10.1017 / S 0025315404010264 h" type="journal article" year="2004">Garraffoni & Lana 2004</bibRefCitation>
|
||
; Hausen 2005; ten
|
||
<bibRefCitation id="EFEF4B56FFF42C58FC68AAA6FAF0FF71" author="Hove" box="[989,1301,218,243]" pageId="12" pageNumber="196" refString="ten Hove, H. A. & Kupriyanova, E. K. (2009) Taxonomy of Serpulidae (Annelida, Polychaeta): the state of affairs. Zootaxa, 2036, 1 - 126." type="journal article" year="2009">Hove & Kupriyanova 2009</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFEF4B56FFF42C58FA97AAA6FEF8FE96" author="Capa" pageId="12" pageNumber="196" refString="Capa, M., Hutchings, P., Aguado, M. T. & Bott, N. (2011) Phylogeny of Sabellidae (Annelida) and relationships with related taxa inferred from morphology and multiple genes. Cladistics, 27, 449 - 469. http: // dx. doi. org / doi: 10.1111 / j. 1096 - 0031.2010.00341. x" type="journal article" year="2011">
|
||
Capa
|
||
<emphasis id="B90AEAB5FFF42C58FAD6AAA1FA79FF71" box="[1379,1436,218,243]" italics="true" pageId="12" pageNumber="196">et al.</emphasis>
|
||
2011
|
||
</bibRefCitation>
|
||
). Since the variability of uncinial morphology seems to be greater than previously documented among sabellariids, further detailed revisions are required to reveal the morphological traits among species not previously analysed.
|
||
</paragraph>
|
||
<paragraph id="8BC136A7FFF42C58FF22AB22FEBCFE7C" blockId="12.[151,1437,151,1043]" pageId="12" pageNumber="196">
|
||
5) The presence of an abdominal expansion of the gut often referred to the ‘proventriculus’ (e.g.,
|
||
<bibRefCitation id="EFEF4B56FFF42C58FAB1AB22FA71FEFA" author="Kirtley" box="[1284,1428,351,376]" pageId="12" pageNumber="196" refString="Kirtley, D. W. (1994) A review and taxonomic revision of the family Sabellariidae Johnston, 1865 (Annelida; Polychaeta). Sabecon Press Science Series, 1, 1 - 223." type="journal article" year="1994">Kirtley 1994</bibRefCitation>
|
||
) has been observed in
|
||
<taxonomicName id="4C7E4D24FFF42C58FE7AABFCFD72FE1B" box="[463,663,385,410]" class="Polychaeta" family="Sabellariidae" genus="Lygdamis" kingdom="Animalia" order="Sabellida" pageId="12" pageNumber="196" phylum="Annelida" rank="species" species="nasutus" status="sp. nov.">
|
||
<emphasis id="B90AEAB5FFF42C58FE7AABFCFD72FE1B" box="[463,663,385,410]" italics="true" pageId="12" pageNumber="196">Lygdamis nasutus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A23957CEFFF42C58FD28ABFFFD3CFE1B" box="[669,729,386,409]" pageId="12" pageNumber="196" rank="species">n. sp.</taxonomicNameLabel>
|
||
and other
|
||
<taxonomicName id="4C7E4D24FFF42C58FCF8ABFCFC5CFE18" box="[845,953,385,410]" class="Polychaeta" family="Sabellariidae" genus="Lygdamis" kingdom="Animalia" order="Sabellida" pageId="12" pageNumber="196" phylum="Annelida" rank="genus">
|
||
<emphasis id="B90AEAB5FFF42C58FCF8ABFCFC5CFE18" box="[845,953,385,410]" italics="true" pageId="12" pageNumber="196">Lygdamis</emphasis>
|
||
</taxonomicName>
|
||
species, but not as conspicuous as found in other
|
||
<taxonomicName id="4C7E4D24FFF42C58FE96ABDFFE50FE39" box="[291,437,418,443]" class="Polychaeta" family="Sabellariidae" kingdom="Animalia" order="Sabellida" pageId="12" pageNumber="196" phylum="Annelida" rank="family">Sabellariidae</taxonomicName>
|
||
(e.g., Hutchings
|
||
<emphasis id="B90AEAB5FFF42C58FDCCABD9FD56FE39" box="[633,691,418,443]" italics="true" pageId="12" pageNumber="196">et al.</emphasis>
|
||
2012: Fig 14C). Since this organ is involved in the food processing, its morphology and development may be related to some functional traits (i.e., feeding preferences or digestion).
|
||
</paragraph>
|
||
<paragraph id="8BC136A7FFF42C58FF22A87BFAC7FB90" blockId="12.[151,1437,151,1043]" pageId="12" pageNumber="196">
|
||
6) The median organ described for this species resemble that described for
|
||
<taxonomicName id="4C7E4D24FFF42C58FBA0A87AFAA7FD9D" authority="Caullery, 1913" authorityName="Caullery" authorityYear="1913" box="[1045,1346,518,543]" class="Polychaeta" family="Sabellariidae" genus="Lygdamis" kingdom="Animalia" order="Sabellida" pageId="12" pageNumber="196" phylum="Annelida" rank="species" species="ehlersi">
|
||
<emphasis id="B90AEAB5FFF42C58FBA0A87AFB65FD9D" box="[1045,1152,518,543]" italics="true" pageId="12" pageNumber="196">L. ehlersi</emphasis>
|
||
(
|
||
<bibRefCitation id="EFEF4B56FFF42C58FB3AA87BFADFFD9D" author="Caullery" box="[1167,1338,518,543]" pageId="12" pageNumber="196" refString="Caullery, M. (1913) Sur le genre Pallasia Qfg. et la region prostomiale des Sabellariens. Bulletin de la Societe Zoologique de France, 38, 198 - 203." type="journal article" year="1913">Caullery, 1913</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C7E4D24FFF42C58FAFAA87AFE30FDC2" authority="Kirtley, 1994" authorityName="Kirtley" authorityYear="1994" class="Polychaeta" family="Sabellariidae" genus="Lygdamis" kingdom="Animalia" order="Sabellida" pageId="12" pageNumber="196" phylum="Annelida" rank="species" species="rayrobertsi">
|
||
<emphasis id="B90AEAB5FFF42C58FAFAA87AFEDCFDC2" italics="true" pageId="12" pageNumber="196">L. rayrobertsi</emphasis>
|
||
<bibRefCitation id="EFEF4B56FFF42C58FE8AA85AFE30FDC2" author="Kirtley" box="[319,469,551,576]" pageId="12" pageNumber="196" refString="Kirtley, D. W. (1994) A review and taxonomic revision of the family Sabellariidae Johnston, 1865 (Annelida; Polychaeta). Sabecon Press Science Series, 1, 1 - 223." type="journal article" year="1994">Kirtley, 1994</bibRefCitation>
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="4C7E4D24FFF42C58FDBAA85AFC5FFDC2" authority="Nishi & Nunez, 1999" authorityName="Nishi & Nunez" authorityYear="1999" box="[527,954,551,576]" class="Polychaeta" family="Sabellariidae" genus="Lygdamis" kingdom="Animalia" order="Sabellida" pageId="12" pageNumber="196" phylum="Annelida" rank="species" species="wirtzi">
|
||
<emphasis id="B90AEAB5FFF42C58FDBAA85AFD27FDC2" box="[527,706,551,576]" italics="true" pageId="12" pageNumber="196">Lygdamis wirtzi</emphasis>
|
||
<bibRefCitation id="EFEF4B56FFF42C58FD7DA85AFC5FFDC2" author="Nishi" box="[712,954,551,576]" pageId="12" pageNumber="196" refString="Nishi, E. & Nunez, J. (1999) A new species of shallow water Sabellariidae (Annelida: Polychaeta) from Madeira Island, Portugal, and Canary Islands, Spain. Arquipelago Boletim da Universidade dos Acores Ciencias Biologicas e Marinhas, 17 a, 37 - 42." type="book chapter" year="1999">Nishi & Núñez, 1999</bibRefCitation>
|
||
</taxonomicName>
|
||
. In these species, the median organ is welldeveloped, arising from the dorsal sagittal suture between opercular lobes and is cylindrical with a planar coronal shape. In at least
|
||
<taxonomicName id="4C7E4D24FFF42C58FDBCA811FD8CFD01" box="[521,617,620,643]" class="Polychaeta" family="Sabellariidae" genus="Lygdamis" kingdom="Animalia" order="Sabellida" pageId="12" pageNumber="196" phylum="Annelida" rank="species" species="wirtzi">
|
||
<emphasis id="B90AEAB5FFF42C58FDBCA811FD8CFD01" box="[521,617,620,643]" italics="true" pageId="12" pageNumber="196">L. wirtzi</emphasis>
|
||
</taxonomicName>
|
||
, this organ is provided with eyespots on both sides, but these have not been described in the other two species (
|
||
<bibRefCitation id="EFEF4B56FFF42C58FD07A8F1FCADFD26" author="Kirtley" box="[690,840,652,677]" pageId="12" pageNumber="196" refString="Kirtley, D. W. (1994) A review and taxonomic revision of the family Sabellariidae Johnston, 1865 (Annelida; Polychaeta). Sabecon Press Science Series, 1, 1 - 223." type="journal article" year="1994">Kirtley 1994</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFEF4B56FFF42C58FCE0A8F1FBA3FD26" author="Nishi" box="[853,1094,652,677]" pageId="12" pageNumber="196" refString="Nishi, E. & Nunez, J. (1999) A new species of shallow water Sabellariidae (Annelida: Polychaeta) from Madeira Island, Portugal, and Canary Islands, Spain. Arquipelago Boletim da Universidade dos Acores Ciencias Biologicas e Marinhas, 17 a, 37 - 42." type="book chapter" year="1999">Nishi & Núñez 1999</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFEF4B56FFF42C58FBE6A8F0FAB9FD26" author="Dos" box="[1107,1372,652,677]" pageId="12" pageNumber="196" refString="Dos Santos, A. S., Dos Santos-Brasil, A. C. & Christoffersen, M. L. (2014) Sabellaria and Lygdamis (Polychaeta: Sabellariidae) from reefs off northeastern Brazil including a new species of Sabellaria. Zootaxa, 3881 (2), 125 - 144. http: // dx. doi. org / 10.11646 / zootaxa. 3881.2.2" type="journal article" year="2014">
|
||
Dos Santos
|
||
<emphasis id="B90AEAB5FFF42C58FB69A8F0FAF3FD27" box="[1244,1302,652,677]" italics="true" pageId="12" pageNumber="196">et al.</emphasis>
|
||
2014
|
||
</bibRefCitation>
|
||
). The new species differs from
|
||
<taxonomicName id="4C7E4D24FFF42C58FDB7A8D3FD88FD44" box="[514,621,685,710]" class="Polychaeta" family="Sabellariidae" genus="Lygdamis" kingdom="Animalia" order="Sabellida" pageId="12" pageNumber="196" phylum="Annelida" rank="species" species="ehlersi">
|
||
<emphasis id="B90AEAB5FFF42C58FDB7A8D3FD88FD44" box="[514,621,685,710]" italics="true" pageId="12" pageNumber="196">L. ehlersi</emphasis>
|
||
</taxonomicName>
|
||
in the relative size of the median organ, which is thinner in the former species (
|
||
<bibRefCitation id="EFEF4B56FFF42C58FEF5A8B3FE36FD65" author="Kirtley" box="[320,467,718,743]" pageId="12" pageNumber="196" refString="Kirtley, D. W. (1994) A review and taxonomic revision of the family Sabellariidae Johnston, 1865 (Annelida; Polychaeta). Sabecon Press Science Series, 1, 1 - 223." type="journal article" year="1994">Kirtley 1994</bibRefCitation>
|
||
).
|
||
<taxonomicName id="4C7E4D24FFF42C58FE53A8B3FD30FD65" box="[486,725,718,743]" class="Polychaeta" family="Sabellariidae" genus="Lygdamis" kingdom="Animalia" order="Sabellida" pageId="12" pageNumber="196" phylum="Annelida" rank="species" species="rayrobertsi">
|
||
<emphasis id="B90AEAB5FFF42C58FE53A8B3FD30FD65" box="[486,725,718,743]" italics="true" pageId="12" pageNumber="196">Lygdamis rayrobertsi</emphasis>
|
||
</taxonomicName>
|
||
is distinguished from
|
||
<taxonomicName id="4C7E4D24FFF42C58FC78A8B2FBA5FD64" box="[973,1088,719,742]" class="Polychaeta" family="Sabellariidae" genus="Lygdamis" kingdom="Animalia" order="Sabellida" pageId="12" pageNumber="196" phylum="Annelida" rank="species" species="nasutus" status="sp. nov.">
|
||
<emphasis id="B90AEAB5FFF42C58FC78A8B2FBA5FD64" box="[973,1088,719,742]" italics="true" pageId="12" pageNumber="196">L. nasutus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A23957CEFFF42C58FBF3A8B2FB67FD64" box="[1094,1154,719,742]" pageId="12" pageNumber="196" rank="species">n. sp.</taxonomicNameLabel>
|
||
in the morphology of the outer paleae, with more pointed tips in the former. The most similar species is
|
||
<taxonomicName id="4C7E4D24FFF42C58FBF9A88CFB4CFC8A" box="[1100,1193,753,776]" class="Polychaeta" family="Sabellariidae" genus="Lygdamis" kingdom="Animalia" order="Sabellida" pageId="12" pageNumber="196" phylum="Annelida" rank="species" species="wirtzi">
|
||
<emphasis id="B90AEAB5FFF42C58FBF9A88CFB4CFC8A" box="[1100,1193,753,776]" italics="true" pageId="12" pageNumber="196">L. wirtzi</emphasis>
|
||
</taxonomicName>
|
||
and both species also share the opercular colour pattern, with a white stalk and dark distal end, and a similar morphology of the parathoracic lanceolate notochaetae with concave blades. Differences between
|
||
<taxonomicName id="4C7E4D24FFF42C58FBE1A949FB22FCC9" box="[1108,1223,820,843]" class="Polychaeta" family="Sabellariidae" genus="Lygdamis" kingdom="Animalia" order="Sabellida" pageId="12" pageNumber="196" phylum="Annelida" rank="species" species="nasutus" status="sp. nov.">
|
||
<emphasis id="B90AEAB5FFF42C58FBE1A949FB22FCC9" box="[1108,1223,820,843]" italics="true" pageId="12" pageNumber="196">L. nasutus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A23957CEFFF42C58FB78A94EFAECFCC8" box="[1229,1289,819,842]" pageId="12" pageNumber="196" rank="species">n. sp.</taxonomicNameLabel>
|
||
and
|
||
<taxonomicName id="4C7E4D24FFF42C58FA8AA949FA79FCC9" box="[1343,1436,820,843]" class="Polychaeta" family="Sabellariidae" genus="Lygdamis" kingdom="Animalia" order="Sabellida" pageId="12" pageNumber="196" phylum="Annelida" rank="species" species="wirtzi">
|
||
<emphasis id="B90AEAB5FFF42C58FA8AA949FA79FCC9" box="[1343,1436,820,843]" italics="true" pageId="12" pageNumber="196">L. wirtzi</emphasis>
|
||
</taxonomicName>
|
||
include the shape of the outer paleae with a thinner and acute tip in the new species and the number of eyespots on the median organ (
|
||
<taxonomicName id="4C7E4D24FFF42C58FDBEA90BFD8DFC0F" box="[523,616,886,909]" class="Polychaeta" family="Sabellariidae" genus="Lygdamis" kingdom="Animalia" order="Sabellida" pageId="12" pageNumber="196" phylum="Annelida" rank="species" species="wirtzi">
|
||
<emphasis id="B90AEAB5FFF42C58FDBEA90BFD8DFC0F" box="[523,616,886,909]" italics="true" pageId="12" pageNumber="196">L. wirtzi</emphasis>
|
||
</taxonomicName>
|
||
bears only two at the base and
|
||
<taxonomicName id="4C7E4D24FFF42C58FC73A90BFBDCFC0F" box="[966,1081,886,909]" class="Polychaeta" family="Sabellariidae" genus="Lygdamis" kingdom="Animalia" order="Sabellida" pageId="12" pageNumber="196" phylum="Annelida" rank="species" species="nasutus" status="sp. nov.">
|
||
<emphasis id="B90AEAB5FFF42C58FC73A90BFBDCFC0F" box="[966,1081,886,909]" italics="true" pageId="12" pageNumber="196">L. nasutus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A23957CEFFF42C58FB8AA90BFB9EFC0F" box="[1087,1147,886,909]" pageId="12" pageNumber="196" rank="species">n. sp.</taxonomicNameLabel>
|
||
has numerous ones along both sides of the organ) (
|
||
<bibRefCitation id="EFEF4B56FFF42C58FE42A9EBFD00FC2D" author="Nishi" box="[503,741,918,943]" pageId="12" pageNumber="196" refString="Nishi, E. & Nunez, J. (1999) A new species of shallow water Sabellariidae (Annelida: Polychaeta) from Madeira Island, Portugal, and Canary Islands, Spain. Arquipelago Boletim da Universidade dos Acores Ciencias Biologicas e Marinhas, 17 a, 37 - 42." type="book chapter" year="1999">Nishi & Núñez 1999</bibRefCitation>
|
||
).
|
||
<taxonomicName id="4C7E4D24FFF42C58FD4DA9EBFC4EFC2C" box="[760,939,918,943]" class="Polychaeta" family="Sabellariidae" genus="Lygdamis" kingdom="Animalia" order="Sabellida" pageId="12" pageNumber="196" phylum="Annelida" rank="species" species="wirtzi">
|
||
<emphasis id="B90AEAB5FFF42C58FD4DA9EBFC4EFC2C" box="[760,939,918,943]" italics="true" pageId="12" pageNumber="196">Lygdamis wirtzi</emphasis>
|
||
</taxonomicName>
|
||
seems to be a larger species (three times the size of
|
||
<taxonomicName id="4C7E4D24FFF42C58FE86A9C4FE42FC52" box="[307,423,953,976]" class="Polychaeta" family="Sabellariidae" genus="Lygdamis" kingdom="Animalia" order="Sabellida" pageId="12" pageNumber="196" phylum="Annelida" rank="species" species="nasutus" status="sp. nov.">
|
||
<emphasis id="B90AEAB5FFF42C58FE86A9C4FE42FC52" box="[307,423,953,976]" italics="true" pageId="12" pageNumber="196">L. nasutus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A23957CEFFF42C58FE18A9C5FE0FFC52" box="[429,490,952,976]" pageId="12" pageNumber="196" rank="species">n. sp.</taxonomicNameLabel>
|
||
) also bearing a larger number of paleae than the new species. Even though
|
||
<taxonomicName id="4C7E4D24FFF42C58FA8BA9C4FA79FC52" box="[1342,1436,953,976]" class="Polychaeta" family="Sabellariidae" genus="Lygdamis" kingdom="Animalia" order="Sabellida" pageId="12" pageNumber="196" phylum="Annelida" rank="species" species="wirtzi">
|
||
<emphasis id="B90AEAB5FFF42C58FA8BA9C4FA79FC52" box="[1342,1436,953,976]" italics="true" pageId="12" pageNumber="196">L. wirtzi</emphasis>
|
||
</taxonomicName>
|
||
was described as presenting over 100 pairs of tentacular filaments, these are clustered in around 13 oblique compound rows (
|
||
<bibRefCitation id="EFEF4B56FFF42C58FE16A987FD3AFB91" author="Nishi" box="[419,735,1018,1043]" pageId="12" pageNumber="196" refString="Nishi, E. & Nunez, J. (1999) A new species of shallow water Sabellariidae (Annelida: Polychaeta) from Madeira Island, Portugal, and Canary Islands, Spain. Arquipelago Boletim da Universidade dos Acores Ciencias Biologicas e Marinhas, 17 a, 37 - 42." type="book chapter" year="1999">
|
||
Nishi & Núñez 1999:
|
||
<figureCitation id="13452A22FFF42C58FD29A987FD3AFB91" box="[668,735,1018,1043]" captionStart="FIGURE 3. A – D" captionStartId="6.[151,250,1449,1471]" captionTargetBox="[154,1433,195,1423]" captionTargetId="figure@6.[151,1436,193,1428]" captionTargetPageId="6" captionText="FIGURE 3. A – D. Lygdamis nasutus n. sp., holotype, AM W. 47372, preserved. A. Operculum, thoracic and parathoracic chaetigers, lateral view; arrowheads point to notopodial cirri; B. Anterior end, ventral view; C. Same, dorsal view; D. Same, but with exposed well-developed median organ. E – F Paratype AM W. 44467. E. Detail of median organ, dorsal view; F. Cross section of paratypes, showing half of the operculum with whole median organ. Abbreviations: br = branchia, bo = building organ, cn 1 = cirrus neuropodia segment 1, chn 1 = chaetae neuropodium segment 1, dc = dorsal cirrus, es = eyespots, ip = inner paleae, li = lips, m = mouth, mo = median organ, nh = nuchal hook, op = outer paleae, opa = opercular papillae, pa = palp, rtf = row of tentacular filaments, tf = tentacular filaments. Small arrow indicates conical dorsal cirrus. Photo: A – D—E. Wong." httpUri="https://zenodo.org/record/245142/files/figure.png" pageId="12" pageNumber="196">Fig. 3</figureCitation>
|
||
</bibRefCitation>
|
||
), a similar number to that found in
|
||
<taxonomicName id="4C7E4D24FFF42C58FBD9A981FB3AFB91" box="[1132,1247,1020,1043]" class="Polychaeta" family="Sabellariidae" genus="Lygdamis" kingdom="Animalia" order="Sabellida" pageId="12" pageNumber="196" phylum="Annelida" rank="species" species="nasutus" status="sp. nov.">
|
||
<emphasis id="B90AEAB5FFF42C58FBD9A981FB3AFB91" box="[1132,1247,1020,1043]" italics="true" pageId="12" pageNumber="196">L. nasutus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A23957CEFFF42C58FB53A986FAC7FB90" box="[1254,1314,1019,1042]" pageId="12" pageNumber="196" rank="species">n. sp.</taxonomicNameLabel>
|
||
</paragraph>
|
||
<caption id="DF01662FFFF42C58FF22AE3DFBDEFBD4" box="[151,1083,1088,1110]" pageId="12" pageNumber="196">
|
||
<paragraph id="8BC136A7FFF42C58FF22AE3DFBDEFBD4" blockId="12.[151,1083,1088,1110]" box="[151,1083,1088,1110]" pageId="12" pageNumber="196">
|
||
<emphasis id="B90AEAB5FFF42C58FF22AE3DFEE1FBD4" bold="true" box="[151,260,1088,1110]" pageId="12" pageNumber="196">TABLE 1.</emphasis>
|
||
Comparison of morphological traits in
|
||
<taxonomicName id="4C7E4D24FFF42C58FD20AE3DFD13FBD4" box="[661,758,1088,1110]" class="Polychaeta" family="Sabellariidae" genus="Lygdamis" kingdom="Animalia" order="Sabellida" pageId="12" pageNumber="196" phylum="Annelida" rank="genus">
|
||
<emphasis id="B90AEAB5FFF42C58FD20AE3DFD13FBD4" box="[661,758,1088,1110]" italics="true" pageId="12" pageNumber="196">Lygdamis</emphasis>
|
||
</taxonomicName>
|
||
species reported from Australia
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BC136A7FFF42C58FE32AE0AFA8DFB0F" blockId="12.[160,1426,1143,2007]" box="[391,1384,1143,1165]" pageId="12" pageNumber="196">
|
||
<emphasis id="B90AEAB5FFF42C58FE32AE0AFACEFB0F" box="[391,1323,1143,1165]" italics="true" pageId="12" pageNumber="196">
|
||
<taxonomicName id="4C7E4D24FFF42C58FE32AE0AFDAFFB0F" box="[391,586,1143,1165]" class="Polychaeta" family="Sabellariidae" genus="Lygdamis" kingdom="Animalia" order="Sabellida" pageId="12" pageNumber="196" phylum="Annelida" rank="species" species="augeneri">Lygdamis augeneri</taxonomicName>
|
||
<taxonomicName id="4C7E4D24FFF42C58FD34AE0AFCCDFB0F" box="[641,808,1143,1165]" class="Polychaeta" family="Sabellariidae" genus="Lygdamis" kingdom="Animalia" order="Sabellida" pageId="12" pageNumber="196" phylum="Annelida" rank="species" species="giardi">Lygdamis giardi</taxonomicName>
|
||
<taxonomicName id="4C7E4D24FFF42C58FCCEAE0AFBDDFB0F" box="[891,1080,1143,1165]" class="Polychaeta" family="Sabellariidae" genus="Lygdamis" kingdom="Animalia" order="Sabellida" pageId="12" pageNumber="196" phylum="Annelida" rank="species" species="wambiri">Lygdamis wambiri</taxonomicName>
|
||
<taxonomicName id="4C7E4D24FFF42C58FBC0AE0AFACEFB0F" box="[1141,1323,1143,1165]" class="Polychaeta" family="Sabellariidae" genus="Lygdamis" kingdom="Animalia" order="Sabellida" pageId="12" pageNumber="196" phylum="Annelida" rank="species" species="nasutus" status="sp. nov.">Lygdamis nasutus</taxonomicName>
|
||
</emphasis>
|
||
<taxonomicNameLabel id="A23957CEFFF42C58FA87AE05FA8DFB0F" box="[1330,1384,1144,1165]" pageId="12" pageNumber="196" rank="species">n. sp.</taxonomicNameLabel>
|
||
</paragraph>
|
||
<paragraph id="8BC136A7FFF42C58FF15AED8FCA6FB6A" blockId="12.[160,1426,1143,2007]" pageId="12" pageNumber="196">Colour pattern colourless* some pigmentation on colourless* as described above operculum in some</paragraph>
|
||
<paragraph id="8BC136A7FFF42C58FD34AE80FD0EFA91" blockId="12.[160,1426,1143,2007]" box="[641,747,1277,1299]" pageId="12" pageNumber="196">specimens</paragraph>
|
||
<paragraph id="8BC136A7FFF42C58FF15AF57FB50FAC2" blockId="12.[160,1426,1143,2007]" box="[160,1205,1322,1344]" pageId="12" pageNumber="196">
|
||
Length (mm; excl. 15–20
|
||
<date id="FFC01067FFF42C58FD34AF57FC71FAC2" box="[641,916,1322,1344]" pageId="12" pageNumber="196" value="1943-12-14">12–43 14</date>
|
||
30–43
|
||
</paragraph>
|
||
<paragraph id="8BC136A7FFF42C58FF15AF2BFF01FAEE" blockId="12.[160,1426,1143,2007]" box="[160,228,1366,1388]" pageId="12" pageNumber="196">cauda)</paragraph>
|
||
<paragraph id="8BC136A7FFF42C58FF15AFFEFB7EFA1A" blockId="12.[160,1426,1143,2007]" box="[160,1179,1410,1432]" pageId="12" pageNumber="196">Max width (mm) 2–3 2–5 2 6–8</paragraph>
|
||
<paragraph id="8BC136A7FFF42C58FF15AFD3FB51FA46" blockId="12.[160,1426,1143,2007]" box="[160,1204,1454,1476]" pageId="12" pageNumber="196">Number of outer 42–44 16–20 27–29 18–19</paragraph>
|
||
<paragraph id="8BC136A7FFF42C58FF15AFA7FECCFA72" blockId="12.[160,1426,1143,2007]" box="[160,297,1498,1520]" pageId="12" pageNumber="196">paleae (pairs)</paragraph>
|
||
<paragraph id="8BC136A7FFF42C58FF15AC7BFA8BF99E" blockId="12.[160,1426,1143,2007]" box="[160,1390,1542,1564]" pageId="12" pageNumber="196">Morphology of outer flattened shafts, flattened shafts, flattened shafts, flattened shafts, margins</paragraph>
|
||
<paragraph id="8BC136A7FFF42C58FF15AC4FFBF7F94E" blockId="12.[160,1426,1143,2007]" pageId="12" pageNumber="196">paleae distally tapering, with distally tapering͵ with margins symmetrical, asymmetrical, progressively nearly symmetrical nearly symmetrical progressively tapering tapering to acute tip with abruptly pointed tips abruptly pointed tips to acute tip with small fine chaetal sculpture and smooth surface and smooth surface distal denticles</paragraph>
|
||
<paragraph id="8BC136A7FFF42C58FF15AC98FB7EF979" blockId="12.[160,1426,1143,2007]" box="[160,1179,1765,1787]" pageId="12" pageNumber="196">
|
||
Number of inner 30–
|
||
<date id="FFC01067FFF42C58FE1BAC98FE22F979" box="[430,455,1765,1787]" pageId="12" pageNumber="196" value="1936-12-10" valueMax="1936-12-14" valueMin="1936-12-10">36</date>
|
||
10–14 12–16 7–8
|
||
</paragraph>
|
||
<paragraph id="8BC136A7FFF42C58FF15AD6CFECCF8A5" blockId="12.[160,1426,1143,2007]" box="[160,297,1809,1831]" pageId="12" pageNumber="196">paleae (pairs)</paragraph>
|
||
<paragraph id="8BC136A7FFF42C58FF15AD40FADCF8D1" blockId="12.[160,1426,1143,2007]" box="[160,1337,1853,1875]" pageId="12" pageNumber="196">Morphology of inner cylindrical, distally cylindrical, distally cylindrical, distally cylindrical, distally</paragraph>
|
||
<paragraph id="8BC136A7FFF42C58FF15AD14FC21F829" blockId="12.[160,1426,1143,2007]" pageId="12" pageNumber="196">paleae tapering, smooth tapering, smooth tapering, smooth tapering, smooth surface surface surface</paragraph>
|
||
<paragraph id="8BC136A7FFF42C59FB3BADBFFA8DFF76" blockId="12.[160,1426,1143,2007]" lastBlockId="13.[160,1421,222,1259]" lastPageId="13" lastPageNumber="197" pageId="12" pageNumber="196">
|
||
<emphasis id="B90AEAB5FFF42C59FB3BADBFFACEFF76" italics="true" lastPageId="13" lastPageNumber="197" pageId="12" pageNumber="196">
|
||
......continue on next page
|
||
<taxonomicName id="4C7E4D24FFF52C59FE32AAA3FDAFFF76" box="[391,586,222,244]" class="Polychaeta" family="Sabellariidae" genus="Lygdamis" kingdom="Animalia" order="Sabellida" pageId="13" pageNumber="197" phylum="Annelida" rank="species" species="augeneri">Lygdamis augeneri</taxonomicName>
|
||
<taxonomicName id="4C7E4D24FFF52C59FD34AAA3FCCDFF76" box="[641,808,222,244]" class="Polychaeta" family="Sabellariidae" genus="Lygdamis" kingdom="Animalia" order="Sabellida" pageId="13" pageNumber="197" phylum="Annelida" rank="species" species="giardi">Lygdamis giardi</taxonomicName>
|
||
<taxonomicName id="4C7E4D24FFF52C59FCCEAAA3FBDDFF76" box="[891,1080,222,244]" class="Polychaeta" family="Sabellariidae" genus="Lygdamis" kingdom="Animalia" order="Sabellida" pageId="13" pageNumber="197" phylum="Annelida" rank="species" species="wambiri">Lygdamis wambiri</taxonomicName>
|
||
<taxonomicName id="4C7E4D24FFF52C59FBC0AAA3FACEFF76" box="[1141,1323,222,244]" class="Polychaeta" family="Sabellariidae" genus="Lygdamis" kingdom="Animalia" order="Sabellida" pageId="13" pageNumber="197" phylum="Annelida" rank="species" species="nasutus" status="sp. nov.">Lygdamis nasutus</taxonomicName>
|
||
</emphasis>
|
||
<taxonomicNameLabel id="A23957CEFFF52C59FA87AAA2FA8DFF76" box="[1330,1384,223,244]" pageId="13" pageNumber="197" rank="species">n. sp.</taxonomicNameLabel>
|
||
</paragraph>
|
||
<caption id="DF01662FFFF52C59FF15AAD8FE75FF39" box="[160,400,165,187]" pageId="13" pageNumber="197">
|
||
<paragraph id="8BC136A7FFF52C59FF15AAD8FE75FF39" blockId="13.[160,400,165,187]" box="[160,400,165,187]" pageId="13" pageNumber="197">
|
||
<emphasis id="B90AEAB5FFF52C59FF15AAD8FEEAFF39" bold="true" box="[160,271,165,187]" pageId="13" pageNumber="197">TABLE 1.</emphasis>
|
||
(Continued)
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BC136A7FFF52C59FF15AB70FB50FEA1" blockId="13.[160,1421,222,1259]" box="[160,1205,269,291]" pageId="13" pageNumber="197">
|
||
Opercular papillae
|
||
<date id="FFC01067FFF52C59FE32AB70FC71FEA1" box="[391,916,269,291]" pageId="13" pageNumber="197" value="1918-08-12" valueMax="1918-12-12" valueMin="1918-08-12">12 8–12 18</date>
|
||
10–12
|
||
</paragraph>
|
||
<paragraph id="8BC136A7FFF52C59FF15AB44FF07FECD" blockId="13.[160,1421,222,1259]" box="[160,226,313,335]" pageId="13" pageNumber="197">(pairs)</paragraph>
|
||
<paragraph id="8BC136A7FFF52C59FF15AB18FB67FEF9" blockId="13.[160,1421,222,1259]" box="[160,1154,357,379]" pageId="13" pageNumber="197">Nuchal spines 1 1 1 1</paragraph>
|
||
<paragraph id="8BC136A7FFF52C59FF15ABECFF07FE25" blockId="13.[160,1421,222,1259]" box="[160,226,401,423]" pageId="13" pageNumber="197">(pairs)</paragraph>
|
||
<paragraph id="8BC136A7FFF52C59FF15ABC0FA60FE51" blockId="13.[160,1421,222,1259]" box="[160,1413,445,467]" pageId="13" pageNumber="197">Morphology of strongly recurved, strongly recurved, strongly recurved, strongly recurved, margins</paragraph>
|
||
<paragraph id="8BC136A7FFF52C59FF15AB94FA64FE7D" blockId="13.[160,1421,222,1259]" box="[160,1409,489,511]" pageId="13" pageNumber="197">nuchal spines margins smooth, margins smooth, margins smooth, smooth, without limbation</paragraph>
|
||
<paragraph id="8BC136A7FFF52C59FE32A868FBCAFDA9" blockId="13.[160,1421,222,1259]" box="[391,1071,533,555]" pageId="13" pageNumber="197">without limbation without limbation without limbation</paragraph>
|
||
<paragraph id="8BC136A7FFF52C59FF15A83CFB51FDD5" blockId="13.[160,1421,222,1259]" box="[160,1204,577,599]" pageId="13" pageNumber="197">
|
||
Tentacular filaments 15–19 15–
|
||
<date id="FFC01067FFF52C59FD1DA83CFD24FDD5" box="[680,705,577,599]" pageId="13" pageNumber="197" value="1919-12-15" valueMax="1919-12-30" valueMin="1919-12-15">19</date>
|
||
15–30 12–19
|
||
</paragraph>
|
||
<paragraph id="8BC136A7FFF52C59FF15A810FF07FD01" blockId="13.[160,1421,222,1259]" box="[160,226,621,643]" pageId="13" pageNumber="197">(rows)</paragraph>
|
||
<paragraph id="8BC136A7FFF52C59FF15A8E4FB18FCB1" blockId="13.[160,1421,222,1259]" pageId="13" pageNumber="197">Median ridge present present, short, not present, as a basal present, as a basal reaching the distal continuation of MO continuation of MO, margin of operculum reaching the distal margin of operculum</paragraph>
|
||
<paragraph id="8BC136A7FFF52C59FF15A934FBE5FB8D" blockId="13.[160,1421,222,1259]" pageId="13" pageNumber="197">Median organ present? small, conical; at small, conical; at large, cylindrical, with junction of lobes of junction of lobes of planar teardrop-shape opercular stalk, as a opercular stalk, as a corona; at junction of lobes continuation of continuation of of opercular stalk. median ridge median ridge</paragraph>
|
||
<paragraph id="8BC136A7FFF52C59FF15AE5BFA68FBB9" blockId="13.[160,1421,222,1259]" box="[160,1421,1061,1083]" pageId="13" pageNumber="197">Eyespots present along sides of present on either side absent present along either sides of</paragraph>
|
||
<paragraph id="8BC136A7FFF52C59FE32AE2CFB1AFBE5" blockId="13.[160,1421,222,1259]" box="[391,1279,1105,1127]" pageId="13" pageNumber="197">median ridge of median ridge median organ</paragraph>
|
||
<paragraph id="8BC136A7FFF52C59FF15AE00FA69FB11" blockId="13.[160,1421,222,1259]" box="[160,1420,1149,1171]" pageId="13" pageNumber="197">Segment 2 (lateral 3 elongate triangular 3 elongate triangular 3 elongate triangular 3 elongate triangular lateral</paragraph>
|
||
<paragraph id="8BC136A7FFF52C59FF15AED4FB0AFB3D" blockId="13.[160,1421,222,1259]" box="[160,1263,1193,1215]" pageId="13" pageNumber="197">lobes)** lateral lobes (short) lateral lobes (short) lateral lobes (long) lobes (long)</paragraph>
|
||
<paragraph id="8BC136A7FFF52C59FF15AEA8FB6AFB69" blockId="13.[160,1421,222,1259]" box="[160,1167,1237,1259]" pageId="13" pageNumber="197">
|
||
Branchiae (pairs)
|
||
<date id="FFC01067FFF52C59FE32AEA8FC71FB69" box="[391,916,1237,1259]" pageId="13" pageNumber="197" value="2012-09-08" valueMax="2012-10-08" valueMin="2012-09-08">8 9–10 12</date>
|
||
15
|
||
</paragraph>
|
||
<paragraph id="8BC136A7FFF52C59FF15AF77FA9BFACE" blockId="13.[160,1413,1290,1400]" pageId="13" pageNumber="197">Segments 3–6 6 lanceolate chaetae, 7–8 lanceolate chaetae 8 lanceolate chaetae 6–7 lanceolate chaetae and (notochaetae) and 6 capillaries in and 7–8 capillaries in and fine capillaries in 6–7 capillaries in between</paragraph>
|
||
<paragraph id="8BC136A7FFF52C59FE32AF1FFC2AFAFA" blockId="13.[160,1413,1290,1400]" box="[391,975,1378,1400]" pageId="13" pageNumber="197">between between between</paragraph>
|
||
<paragraph id="8BC136A7FFF52C59FF15AFE9FA9AFA54" blockId="13.[160,1414,1428,1929]" pageId="13" pageNumber="197">Segments 3–6 4 lanceolate chaetae 5–6 lanceolate chaetae 5 lanceolate chaetae 5 lanceolate chaetae and (neurochaetae) and 4 fine capillaries and 5 fine capillaries and fine capillaries in fine capillaries in between</paragraph>
|
||
<paragraph id="8BC136A7FFF52C59FE32AF91FC2AF980" blockId="13.[160,1414,1428,1929]" box="[391,975,1516,1538]" pageId="13" pageNumber="197">with thin flattened with compact thecae between</paragraph>
|
||
<paragraph id="8BC136A7FFF52C59FE32AC65FD0AF9AC" blockId="13.[160,1414,1428,1929]" box="[391,751,1560,1582]" pageId="13" pageNumber="197">blades with short in between</paragraph>
|
||
<paragraph id="8BC136A7FFF52C59FE32AC38FDD2F9D8" blockId="13.[160,1414,1428,1929]" box="[391,567,1604,1626]" pageId="13" pageNumber="197">narrow thecae, in</paragraph>
|
||
<paragraph id="8BC136A7FFF52C59FE32AC0DFE3EF904" blockId="13.[160,1414,1428,1929]" box="[391,475,1648,1670]" pageId="13" pageNumber="197">between</paragraph>
|
||
<paragraph id="8BC136A7FFF52C59FF15ACE1FAA9F95C" blockId="13.[160,1414,1428,1929]" pageId="13" pageNumber="197">Uncinal rows of 1–3 rows from 2 rows of teeth after a?2 2–3 rows from proximal to teeth proximal to distal end first proximal single distal end on anterior</paragraph>
|
||
<paragraph id="8BC136A7FFF52C59FD34AC89FA8AF888" blockId="13.[160,1414,1428,1929]" box="[641,1391,1780,1802]" pageId="13" pageNumber="197">
|
||
tooth chaetigers,
|
||
<quantity id="4C869B42FFF52C59FB5DAC89FAEAF888" box="[1256,1295,1780,1802]" metricMagnitude="-2" metricUnit="m" metricValue="5.08" pageId="13" pageNumber="197" unit="in" value="2.0">2 in</quantity>
|
||
posterior
|
||
</paragraph>
|
||
<paragraph id="8BC136A7FFF52C59FBC0AD5DFB67F8E0" blockId="13.[160,1414,1428,1929]" pageId="13" pageNumber="197">uncini Number of teeth/ 6–8 7–9 8 7</paragraph>
|
||
<paragraph id="8BC136A7FFF52C59FF15AD04FA77F838" blockId="13.[160,1414,1428,1929]" box="[160,1426,1913,1978]" lastBlockId="13.[1166,1426,1961,1978]" pageId="13" pageNumber="197">
|
||
row
|
||
<emphasis id="B90AEAB5FFF52C59FB3BADD8FA77F838" box="[1166,1426,1957,1978]" italics="true" pageId="13" pageNumber="197">......continue on next page</emphasis>
|
||
</paragraph>
|
||
<caption id="DF01662FFFF62C5AFF15AAD8FE75FF39" box="[160,400,165,187]" pageId="14" pageNumber="198">
|
||
<paragraph id="8BC136A7FFF62C5AFF15AAD8FE75FF39" blockId="14.[160,400,165,187]" box="[160,400,165,187]" pageId="14" pageNumber="198">
|
||
<emphasis id="B90AEAB5FFF62C5AFF15AAD8FEEAFF39" bold="true" box="[160,271,165,187]" pageId="14" pageNumber="198">TABLE 1.</emphasis>
|
||
(Continued)
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BC136A7FFF62C5AFE32AAA3FA8DFF76" blockId="14.[151,1422,222,544]" box="[391,1384,222,244]" pageId="14" pageNumber="198">
|
||
<emphasis id="B90AEAB5FFF62C5AFE32AAA3FACEFF76" box="[391,1323,222,244]" italics="true" pageId="14" pageNumber="198">
|
||
<taxonomicName id="4C7E4D24FFF62C5AFE32AAA3FDAFFF76" box="[391,586,222,244]" class="Polychaeta" family="Sabellariidae" genus="Lygdamis" kingdom="Animalia" order="Sabellida" pageId="14" pageNumber="198" phylum="Annelida" rank="species" species="augeneri">Lygdamis augeneri</taxonomicName>
|
||
<taxonomicName id="4C7E4D24FFF62C5AFD34AAA3FCCDFF76" box="[641,808,222,244]" class="Polychaeta" family="Sabellariidae" genus="Lygdamis" kingdom="Animalia" order="Sabellida" pageId="14" pageNumber="198" phylum="Annelida" rank="species" species="giardi">Lygdamis giardi</taxonomicName>
|
||
<taxonomicName id="4C7E4D24FFF62C5AFCCEAAA3FBDDFF76" box="[891,1080,222,244]" class="Polychaeta" family="Sabellariidae" genus="Lygdamis" kingdom="Animalia" order="Sabellida" pageId="14" pageNumber="198" phylum="Annelida" rank="species" species="wambiri">Lygdamis wambiri</taxonomicName>
|
||
<taxonomicName id="4C7E4D24FFF62C5AFBC0AAA3FACEFF76" box="[1141,1323,222,244]" class="Polychaeta" family="Sabellariidae" genus="Lygdamis" kingdom="Animalia" order="Sabellida" pageId="14" pageNumber="198" phylum="Annelida" rank="species" species="nasutus" status="sp. nov.">Lygdamis nasutus</taxonomicName>
|
||
</emphasis>
|
||
<taxonomicNameLabel id="A23957CEFFF62C5AFA87AAA2FA8DFF76" box="[1330,1384,223,244]" pageId="14" pageNumber="198" rank="species">n. sp.</taxonomicNameLabel>
|
||
</paragraph>
|
||
<paragraph id="8BC136A7FFF62C5AFF15AB70FA6BFEA1" blockId="14.[151,1422,222,544]" box="[160,1422,269,291]" pageId="14" pageNumber="198">
|
||
Abdominal one
|
||
<typeStatus id="54C58805FFF62C5AFE04AB73FE38FEA1" box="[433,477,270,291]" pageId="14" pageNumber="198">type</typeStatus>
|
||
of capillaries, one
|
||
<typeStatus id="54C58805FFF62C5AFD1EAB73FD32FEA1" box="[683,727,270,291]" pageId="14" pageNumber="198">type</typeStatus>
|
||
of capillaries, one
|
||
<typeStatus id="54C58805FFF62C5AFC11AB73FC35FEA1" box="[932,976,270,291]" pageId="14" pageNumber="198">type</typeStatus>
|
||
of capillaries, dorsal row ornamented with
|
||
</paragraph>
|
||
<paragraph id="8BC136A7FFF62C5AFF15AB44FB54FE51" blockId="14.[151,1422,222,544]" pageId="14" pageNumber="198">neuropodia ornamented with ornamented with ornamented with irregular thecal laminar irregular fibres irregular fibres irregular fibres extensions; ventral row (‘pilose type’) (‘pilose type’) (‘pilose type’) with irregular fibres (‘pilose type’)</paragraph>
|
||
<paragraph id="8BC136A7FFF62C5AFF22AB96FC3CFD83" blockId="14.[151,1422,222,544]" box="[151,985,491,513]" pageId="14" pageNumber="198">* Pigmentation may be related to methods of preservation and subsequent storage.</paragraph>
|
||
<paragraph id="8BC136A7FFF62C5AFF22A877FAD3FDA2" blockId="14.[151,1422,222,544]" box="[151,1334,522,544]" pageId="14" pageNumber="198">
|
||
** Short or long is just a relative comparison between the species dealt with herein (based on Hutchings
|
||
<emphasis id="B90AEAB5FFF62C5AFB0EA876FB0BFDA2" box="[1211,1262,522,544]" italics="true" pageId="14" pageNumber="198">et al.</emphasis>
|
||
2012).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C364652CFFF62C5AFF72A80DFBE0FD53" pageId="14" pageNumber="198" type="etymology">
|
||
<paragraph id="8BC136A7FFF62C5AFF72A80DFD0EFD2F" blockId="14.[151,1436,624,902]" pageId="14" pageNumber="198">
|
||
<emphasis id="B90AEAB5FFF62C5AFF72A80DFEABFD0B" bold="true" box="[199,334,624,649]" pageId="14" pageNumber="198">Etymology.</emphasis>
|
||
This species is named
|
||
<taxonomicName id="4C7E4D24FFF62C5AFDFBA80FFD46FD0B" box="[590,675,626,649]" class="Polychaeta" family="Sabellariidae" genus="Lygdamis" kingdom="Animalia" order="Sabellida" pageId="14" pageNumber="198" phylum="Annelida" rank="species" species="nasutus">
|
||
<emphasis id="B90AEAB5FFF62C5AFDFBA80FFD46FD0B" box="[590,675,626,649]" italics="true" pageId="14" pageNumber="198">nasutus</emphasis>
|
||
</taxonomicName>
|
||
(big-nosed and sensory, in Latin) in reference to the well-developed median organ, considered to have a sensory function.
|
||
</paragraph>
|
||
<paragraph id="8BC136A7FFF62C5AFF72A8C5FBE0FD53" blockId="14.[151,1436,624,902]" box="[199,1029,696,721]" pageId="14" pageNumber="198">
|
||
<emphasis id="B90AEAB5FFF62C5AFF72A8C5FECFFD53" bold="true" box="[199,298,696,721]" pageId="14" pageNumber="198">Habitat.</emphasis>
|
||
This solitary species was collected at
|
||
<quantity id="4C869B42FFF62C5AFD63A8C5FCC8FD52" box="[726,813,696,721]" metricMagnitude="0" metricUnit="m" metricValue="8.5" metricValueMax="12.0" metricValueMin="5.0" pageId="14" pageNumber="198" unit="m" value="8.5" valueMax="12.0" valueMin="5.0">5–12 m</quantity>
|
||
, from coral rubble.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C364652CFFF62C5AFF72A8A1FC78FD77" box="[199,925,732,757]" pageId="14" pageNumber="198" type="materials_examined">
|
||
<paragraph id="8BC136A7FFF62C5AFF72A8A1FC78FD77" blockId="14.[151,1436,624,902]" box="[199,925,732,757]" pageId="14" pageNumber="198">
|
||
<emphasis id="B90AEAB5FFF62C5AFF72A8A1FE80FD77" bold="true" box="[199,357,732,757]" pageId="14" pageNumber="198">
|
||
<typeStatus id="54C58805FFF62C5AFF72A8A1FEE7FD77" box="[199,258,732,757]" pageId="14" pageNumber="198">Type</typeStatus>
|
||
locality.
|
||
</emphasis>
|
||
Mermaid Cove, Lizard Island, Great Barrier Reef.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C364652CFFF62C5AFF72A97DFE1AFC07" pageId="14" pageNumber="198" type="distribution">
|
||
<paragraph id="8BC136A7FFF62C5AFF72A97DFE1AFC07" blockId="14.[151,1436,624,902]" pageId="14" pageNumber="198">
|
||
<emphasis id="B90AEAB5FFF62C5AFF72A97DFE85FC9B" bold="true" box="[199,352,768,793]" pageId="14" pageNumber="198">Distribution.</emphasis>
|
||
Only known from Lizard Island. Specimens were collected twice (in
|
||
<date id="FFC01067FFF62C5AFBDBA97DFAFDFC9B" box="[1134,1304,768,793]" pageId="14" pageNumber="198" value="1995" valueMax="2013">1995 and 2013</date>
|
||
), in spite of extensive collecting for polychaetes over the last 40 years at Lizard, indicating this species is rare. The
|
||
<typeStatus id="54C58805FFF62C5AFAA7A95BFAA7FCBC" box="[1298,1346,806,830]" pageId="14" pageNumber="198">type</typeStatus>
|
||
locality is an easily accessible site which has been dived at many times by Pat Hutchings since Lizard Island Research Station was established in 1973.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |