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<mods:title id="1BE1CC2F88A12D674A2E3B4B8647EC1D">Key to species of the leafhopper genus Fistulatus Zhang, Zhang &amp; Chen (Hemiptera: Cicadellidae: Deltocephalinae: Drabescini) with description of a new species</mods:title>
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Key to species (males) of
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<emphasis id="B90AEA94FFF2CF741F9FF9BC0F670159" bold="true" box="[465,587,1557,1584]" italics="true" pageId="1" pageNumber="364">Fistulatus</emphasis>
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(modified from
<bibRefCitation id="EFEF4B77FFF2CF741D59F9BF0ED20146" author="Lu, L. &amp; Zhang, Y. L." box="[791,1022,1557,1584]" pageId="1" pageNumber="364" pagination="247 - 250" refId="ref5506" refString="Lu, L. &amp; Zhang, Y. L. (2014) Taxonomy of the Oriental leafhopper genus Fistulatus (Hemiptera: Cicadellidae: Deltocephalinae), with description of a new species from China. Zootaxa, 3838 (2), 247 - 250. https: // doi. org / 10.11646 / zootaxa. 3838.2.10" type="journal article" year="2014">Lu &amp; Zhang, 2014</bibRefCitation>
)
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<paragraph id="8BC13686FFF2CF741ED9F9F208B00119" blockId="1.[151,1437,1627,2030]" box="[151,1436,1627,1649]" pageId="1" pageNumber="364">
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1 Pygofer with two pairs of processes (
<figureCitation id="13452A03FFF2CF741C08F9F20FA00118" box="[582,652,1627,1649]" captionStart="FIGURES 1439" captionStartId="5.[151,264,1631,1655]" captionTargetBox="[157,1429,187,1602]" captionTargetId="figure-17@5.[151,1435,181,1608]" captionTargetPageId="5" captionText="FIGURES 1439. Fistulatus species. 1419: F. athena (after Shang &amp; Zhang, 2003), 14: male pygofer, lateral view; 15: style, dorsal view; 16: connective, dorsal view; 17: aedeagus, ventral view; 18: aedeagus, dorsal view; 19: aedeagus, lateral view; 20 24: F. bidentatus (after Cen &amp; Cai, 2002), 20: male pygofer, dorsolateral view; 21: style, ventral view; 22: connective, ventral view; 23: aedeagus, ventral view; 24: aedeagus, lateral view; 2529: F. biconjugara (after Yang et al. 2024), 25: male pygofer, lateral view; 26: style, dorsal view; 27: connective, dorsal view; 28: aedeagus, ventral view; 29: aedeagus, lateral view; 3034: F. brevimarginalis (after Yang et al. 2024), 30: male pygofer, lateral view; 31. style, dorsal view; 32: connective, dorsal view; 33: aedeagus, ventral view; 34: aedeagus, lateral view; 3539: F. luteolus (after Cen &amp; Cai, 2002), 35: male pygofer, lateral view; 36: style, ventral view; 37: connective, ventral view; 38: aedeagus, ventral view; 39: aedeagus, lateral view." figureDoi="http://doi.org/10.5281/zenodo.12758869" httpUri="https://zenodo.org/record/12758869/files/figure.png" pageId="1" pageNumber="364">Fig. 35</figureCitation>
)............................................................... 2
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<paragraph id="8BC13686FFF2CF741ED9F9D008B001E7" blockId="1.[151,1437,1627,2030]" box="[151,1436,1656,1678]" pageId="1" pageNumber="364">
<keyLead id="308F95B3FFF2CF741ED9F9D008B001E7" box="[151,1436,1656,1678]" pageId="1" pageNumber="364">
- Pygofer with one pair of processes (
<figureCitation id="13452A03FFF2CF741C72F9D10F5901E7" box="[572,629,1656,1678]" captionStart="FIGURES 513" captionStartId="3.[151,264,1846,1870]" captionTargetBox="[158,1429,188,1816]" captionTargetId="figure-17@3.[151,1435,181,1822]" captionTargetPageId="3" captionText="FIGURES 513. Fistulatus motuoensis Hassan &amp; Xing sp. nov. Male. 5. Genitalia (after maceration), lateral view; 6. Male pygofer, lateral view; 7. Valve and subgenital plate, ventral view; 8. Style, lateral view; 9. Connective, dorsal view; 10.Aedeagus, dorsal view; 11. Aedeagus, ventral view; 12. Aedeagus, lateral view; 13. Aedeagus, anteroventral view focusing on the apex." figureDoi="http://doi.org/10.5281/zenodo.12758861" httpUri="https://zenodo.org/record/12758861/files/figure.png" pageId="1" pageNumber="364">Fig. 6</figureCitation>
)................................................................. 5
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<paragraph id="8BC13686FFF2CF741ED9F93F08B001C2" blockId="1.[151,1437,1627,2030]" box="[151,1436,1685,1707]" pageId="1" pageNumber="364">
<keyLead id="308F95B3FFF2CF741ED9F93F08B001C2" box="[151,1436,1685,1707]" pageId="1" pageNumber="364">2 Aedeagal shaft with one or more processes in distal half; pygofer processes extended mesad......................... 3</keyLead>
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<paragraph id="8BC13686FFF2CF741ED9F91A08B101A1" blockId="1.[151,1437,1627,2030]" box="[151,1437,1714,1737]" pageId="1" pageNumber="364">
<keyLead id="308F95B3FFF2CF741ED9F91A08B101A1" box="[151,1437,1714,1737]" pageId="1" pageNumber="364">
- Aedeagal shaft without processes in distal half (
<figureCitation id="13452A03FFF2CF741CE3F91A0E3101A1" box="[685,797,1715,1737]" captionStart="FIGURES 1439" captionStartId="5.[151,264,1631,1655]" captionTargetBox="[157,1429,187,1602]" captionTargetId="figure-17@5.[151,1435,181,1608]" captionTargetPageId="5" captionText="FIGURES 1439. Fistulatus species. 1419: F. athena (after Shang &amp; Zhang, 2003), 14: male pygofer, lateral view; 15: style, dorsal view; 16: connective, dorsal view; 17: aedeagus, ventral view; 18: aedeagus, dorsal view; 19: aedeagus, lateral view; 20 24: F. bidentatus (after Cen &amp; Cai, 2002), 20: male pygofer, dorsolateral view; 21: style, ventral view; 22: connective, ventral view; 23: aedeagus, ventral view; 24: aedeagus, lateral view; 2529: F. biconjugara (after Yang et al. 2024), 25: male pygofer, lateral view; 26: style, dorsal view; 27: connective, dorsal view; 28: aedeagus, ventral view; 29: aedeagus, lateral view; 3034: F. brevimarginalis (after Yang et al. 2024), 30: male pygofer, lateral view; 31. style, dorsal view; 32: connective, dorsal view; 33: aedeagus, ventral view; 34: aedeagus, lateral view; 3539: F. luteolus (after Cen &amp; Cai, 2002), 35: male pygofer, lateral view; 36: style, ventral view; 37: connective, ventral view; 38: aedeagus, ventral view; 39: aedeagus, lateral view." figureDoi="http://doi.org/10.5281/zenodo.12758869" httpUri="https://zenodo.org/record/12758869/files/figure.png" pageId="1" pageNumber="364">Figs 3839</figureCitation>
); pygofer processes extended distad (
<figureCitation id="13452A03FFF2CF741ACAF91A09E601A0" box="[1156,1226,1715,1737]" captionStart="FIGURES 1439" captionStartId="5.[151,264,1631,1655]" captionTargetBox="[157,1429,187,1602]" captionTargetId="figure-17@5.[151,1435,181,1608]" captionTargetPageId="5" captionText="FIGURES 1439. Fistulatus species. 1419: F. athena (after Shang &amp; Zhang, 2003), 14: male pygofer, lateral view; 15: style, dorsal view; 16: connective, dorsal view; 17: aedeagus, ventral view; 18: aedeagus, dorsal view; 19: aedeagus, lateral view; 20 24: F. bidentatus (after Cen &amp; Cai, 2002), 20: male pygofer, dorsolateral view; 21: style, ventral view; 22: connective, ventral view; 23: aedeagus, ventral view; 24: aedeagus, lateral view; 2529: F. biconjugara (after Yang et al. 2024), 25: male pygofer, lateral view; 26: style, dorsal view; 27: connective, dorsal view; 28: aedeagus, ventral view; 29: aedeagus, lateral view; 3034: F. brevimarginalis (after Yang et al. 2024), 30: male pygofer, lateral view; 31. style, dorsal view; 32: connective, dorsal view; 33: aedeagus, ventral view; 34: aedeagus, lateral view; 3539: F. luteolus (after Cen &amp; Cai, 2002), 35: male pygofer, lateral view; 36: style, ventral view; 37: connective, ventral view; 38: aedeagus, ventral view; 39: aedeagus, lateral view." figureDoi="http://doi.org/10.5281/zenodo.12758869" httpUri="https://zenodo.org/record/12758869/files/figure.png" pageId="1" pageNumber="364">Fig. 35</figureCitation>
).......
<taxonomicName id="4C7E4D05FFF2CF741B7BF91A08B101A1" ID-CoL="6J6X4" authorityName="Cen &amp; Cai" authorityYear="2002" box="[1333,1437,1714,1736]" class="Insecta" family="Cicadellidae" genus="Fistulatus" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="364" phylum="Arthropoda" rank="species" species="luteolus">
<emphasis id="B90AEA94FFF2CF741B7BF91A08B101A1" bold="true" box="[1333,1437,1714,1736]" italics="true" pageId="1" pageNumber="364">F. luteolus</emphasis>
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3 Aedeagal shaft with pair of short, slender preapical processes (
<figureCitation id="13452A03FFF2CF741D75F9790E85018F" box="[827,937,1744,1766]" captionStart="FIGURES 4059" captionStartId="7.[151,264,1421,1445]" captionTargetBox="[157,1430,187,1391]" captionTargetId="figure-17@7.[151,1435,181,1397]" captionTargetPageId="7" captionText="FIGURES 4059. Fistulatus species. 4044: F. monkoboensis (after Shang &amp; Zhang, 2003), 40: male pygofer, lateral view; 41: subgenital plate, valve, style and connective, ventral view; 42: aedeagus, dorsal view; 43: aedeagus, ventral view; 44: aedeagus, lateral view; 4549: F. rectilineus (after Shang et al., 2006), 45: male pygofer, lateral view; 46: style, dorsal view; 47: connective, dorsal view; 48: aedeagus, ventral view; 49: aedeagus, lateral view; 5054: F. sinensis (after Zhang et al., 1997), 50: male pygofer, lateral view; 51: style, dorsal view; 52: connective, dorsal view; 53: aedeagus, posteroventral view; 54: aedeagus, lateral view; 5559: F. quadrispinosus (after Lu &amp; Zhang, 2014), 55: male pygofer, lateral view; 56: subgenital plate, valve, style and connective, ventral view; 57: aedeagus, lateral view; 58: aedeagus, ventral view; 59: aedeagus, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.12758871" httpUri="https://zenodo.org/record/12758871/files/figure.png" pageId="1" pageNumber="364">Figs 5759</figureCitation>
).........................
<taxonomicName id="4C7E4D05FFF2CF741AA3F97908B0018F" ID-CoL="8RKR6" authorityName="Lu &amp; Zhang" authorityYear="2014" box="[1261,1436,1744,1766]" class="Insecta" family="Cicadellidae" genus="Fistulatus" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="364" phylum="Arthropoda" rank="species" species="quadrispinosus">
<emphasis id="B90AEA94FFF2CF741AA3F97908B0018F" bold="true" box="[1261,1436,1744,1766]" italics="true" pageId="1" pageNumber="364">F. quadrispinosus</emphasis>
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<keyLead id="308F95B3FFF2CF741ED9F94708B0006A" box="[151,1436,1773,1795]" pageId="1" pageNumber="364">- Aedeagal shaft with process or process flangelike............................................................ 4</keyLead>
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4 Aedeagal shaft with a pair of short apical lateral processes (
<figureCitation id="13452A03FFF2CF741D53F8A20EA20049" box="[797,910,1803,1825]" captionStart="FIGURES 1439" captionStartId="5.[151,264,1631,1655]" captionTargetBox="[157,1429,187,1602]" captionTargetId="figure-17@5.[151,1435,181,1608]" captionTargetPageId="5" captionText="FIGURES 1439. Fistulatus species. 1419: F. athena (after Shang &amp; Zhang, 2003), 14: male pygofer, lateral view; 15: style, dorsal view; 16: connective, dorsal view; 17: aedeagus, ventral view; 18: aedeagus, dorsal view; 19: aedeagus, lateral view; 20 24: F. bidentatus (after Cen &amp; Cai, 2002), 20: male pygofer, dorsolateral view; 21: style, ventral view; 22: connective, ventral view; 23: aedeagus, ventral view; 24: aedeagus, lateral view; 2529: F. biconjugara (after Yang et al. 2024), 25: male pygofer, lateral view; 26: style, dorsal view; 27: connective, dorsal view; 28: aedeagus, ventral view; 29: aedeagus, lateral view; 3034: F. brevimarginalis (after Yang et al. 2024), 30: male pygofer, lateral view; 31. style, dorsal view; 32: connective, dorsal view; 33: aedeagus, ventral view; 34: aedeagus, lateral view; 3539: F. luteolus (after Cen &amp; Cai, 2002), 35: male pygofer, lateral view; 36: style, ventral view; 37: connective, ventral view; 38: aedeagus, ventral view; 39: aedeagus, lateral view." figureDoi="http://doi.org/10.5281/zenodo.12758869" httpUri="https://zenodo.org/record/12758869/files/figure.png" pageId="1" pageNumber="364">Figs 2829</figureCitation>
); ventral process of pygofer side extended toward base (
<figureCitation id="13452A03FFF2CF741EA7F8810C030057" box="[233,303,1832,1854]" captionStart="FIGURES 1439" captionStartId="5.[151,264,1631,1655]" captionTargetBox="[157,1429,187,1602]" captionTargetId="figure-17@5.[151,1435,181,1608]" captionTargetPageId="5" captionText="FIGURES 1439. Fistulatus species. 1419: F. athena (after Shang &amp; Zhang, 2003), 14: male pygofer, lateral view; 15: style, dorsal view; 16: connective, dorsal view; 17: aedeagus, ventral view; 18: aedeagus, dorsal view; 19: aedeagus, lateral view; 20 24: F. bidentatus (after Cen &amp; Cai, 2002), 20: male pygofer, dorsolateral view; 21: style, ventral view; 22: connective, ventral view; 23: aedeagus, ventral view; 24: aedeagus, lateral view; 2529: F. biconjugara (after Yang et al. 2024), 25: male pygofer, lateral view; 26: style, dorsal view; 27: connective, dorsal view; 28: aedeagus, ventral view; 29: aedeagus, lateral view; 3034: F. brevimarginalis (after Yang et al. 2024), 30: male pygofer, lateral view; 31. style, dorsal view; 32: connective, dorsal view; 33: aedeagus, ventral view; 34: aedeagus, lateral view; 3539: F. luteolus (after Cen &amp; Cai, 2002), 35: male pygofer, lateral view; 36: style, ventral view; 37: connective, ventral view; 38: aedeagus, ventral view; 39: aedeagus, lateral view." figureDoi="http://doi.org/10.5281/zenodo.12758869" httpUri="https://zenodo.org/record/12758869/files/figure.png" pageId="1" pageNumber="364">Fig. 25</figureCitation>
).................................................................................
<taxonomicName id="4C7E4D05FFF2CF741B45F88108B00057" ID-CoL="CCMX6" authorityName="Yang &amp; Dai" authorityYear="2024" box="[1291,1436,1832,1854]" class="Insecta" family="Cicadellidae" genus="Fistulatus" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="364" phylum="Arthropoda" rank="species" species="biconjugara">
<emphasis id="B90AEA94FFF2CF741B45F88108B00057" bold="true" box="[1291,1436,1832,1854]" italics="true" pageId="1" pageNumber="364">F. biconjugara</emphasis>
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<paragraph id="8BC13686FFF2CF741ED9F8EF08B00011" blockId="1.[151,1437,1627,2030]" pageId="1" pageNumber="364">
<keyLead id="308F95B3FFF2CF741ED9F8EF08B00011" pageId="1" pageNumber="364">
- Aedeagal shaft with one dorsal preapical process (
<figureCitation id="13452A03FFF2CF741C8AF8EC0E190032" box="[708,821,1861,1883]" captionStart="FIGURES 1439" captionStartId="5.[151,264,1631,1655]" captionTargetBox="[157,1429,187,1602]" captionTargetId="figure-17@5.[151,1435,181,1608]" captionTargetPageId="5" captionText="FIGURES 1439. Fistulatus species. 1419: F. athena (after Shang &amp; Zhang, 2003), 14: male pygofer, lateral view; 15: style, dorsal view; 16: connective, dorsal view; 17: aedeagus, ventral view; 18: aedeagus, dorsal view; 19: aedeagus, lateral view; 20 24: F. bidentatus (after Cen &amp; Cai, 2002), 20: male pygofer, dorsolateral view; 21: style, ventral view; 22: connective, ventral view; 23: aedeagus, ventral view; 24: aedeagus, lateral view; 2529: F. biconjugara (after Yang et al. 2024), 25: male pygofer, lateral view; 26: style, dorsal view; 27: connective, dorsal view; 28: aedeagus, ventral view; 29: aedeagus, lateral view; 3034: F. brevimarginalis (after Yang et al. 2024), 30: male pygofer, lateral view; 31. style, dorsal view; 32: connective, dorsal view; 33: aedeagus, ventral view; 34: aedeagus, lateral view; 3539: F. luteolus (after Cen &amp; Cai, 2002), 35: male pygofer, lateral view; 36: style, ventral view; 37: connective, ventral view; 38: aedeagus, ventral view; 39: aedeagus, lateral view." figureDoi="http://doi.org/10.5281/zenodo.12758869" httpUri="https://zenodo.org/record/12758869/files/figure.png" pageId="1" pageNumber="364">Figs 3334</figureCitation>
); ventral process of pygofer extended toward apex (
<figureCitation id="13452A03FFF2CF741B68F8EC08400032" box="[1318,1388,1861,1883]" captionStart="FIGURES 1439" captionStartId="5.[151,264,1631,1655]" captionTargetBox="[157,1429,187,1602]" captionTargetId="figure-17@5.[151,1435,181,1608]" captionTargetPageId="5" captionText="FIGURES 1439. Fistulatus species. 1419: F. athena (after Shang &amp; Zhang, 2003), 14: male pygofer, lateral view; 15: style, dorsal view; 16: connective, dorsal view; 17: aedeagus, ventral view; 18: aedeagus, dorsal view; 19: aedeagus, lateral view; 20 24: F. bidentatus (after Cen &amp; Cai, 2002), 20: male pygofer, dorsolateral view; 21: style, ventral view; 22: connective, ventral view; 23: aedeagus, ventral view; 24: aedeagus, lateral view; 2529: F. biconjugara (after Yang et al. 2024), 25: male pygofer, lateral view; 26: style, dorsal view; 27: connective, dorsal view; 28: aedeagus, ventral view; 29: aedeagus, lateral view; 3034: F. brevimarginalis (after Yang et al. 2024), 30: male pygofer, lateral view; 31. style, dorsal view; 32: connective, dorsal view; 33: aedeagus, ventral view; 34: aedeagus, lateral view; 3539: F. luteolus (after Cen &amp; Cai, 2002), 35: male pygofer, lateral view; 36: style, ventral view; 37: connective, ventral view; 38: aedeagus, ventral view; 39: aedeagus, lateral view." figureDoi="http://doi.org/10.5281/zenodo.12758869" httpUri="https://zenodo.org/record/12758869/files/figure.png" pageId="1" pageNumber="364">Fig. 30</figureCitation>
)........................................................................................
<taxonomicName id="4C7E4D05FFF2CF741AA9F8CA08B00011" ID-CoL="CCMWX" authorityName="Yang &amp; Dai" authorityYear="2024" box="[1255,1436,1890,1912]" class="Insecta" family="Cicadellidae" genus="Fistulatus" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="364" phylum="Arthropoda" rank="species" species="brevimarginalis">
<emphasis id="B90AEA94FFF2CF741AA9F8CA08B00011" bold="true" box="[1255,1436,1890,1912]" italics="true" pageId="1" pageNumber="364">F. brevimarginalis</emphasis>
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<paragraph id="8BC13686FFF2CF741ED9F82908B000FF" blockId="1.[151,1437,1627,2030]" box="[151,1436,1920,1942]" pageId="1" pageNumber="364">
<keyLead id="308F95B3FFF2CF741ED9F82908B000FF" box="[151,1436,1920,1942]" pageId="1" pageNumber="364">5 Aedeagal shaft with one pair of processes or flanges.......................................................... 6</keyLead>
</paragraph>
<paragraph id="8BC13686FFF2CF741ED9F83708B100DA" blockId="1.[151,1437,1627,2030]" box="[151,1437,1949,1971]" pageId="1" pageNumber="364">
<keyLead id="308F95B3FFF2CF741ED9F83708B100DA" box="[151,1437,1949,1971]" pageId="1" pageNumber="364">
- Aedeagal shaft with two pairs of processes (
<figureCitation id="13452A03FFF2CF741CC4F8340FE300DA" box="[650,719,1949,1971]" captionStart="FIGURES 1439" captionStartId="5.[151,264,1631,1655]" captionTargetBox="[157,1429,187,1602]" captionTargetId="figure-17@5.[151,1435,181,1608]" captionTargetPageId="5" captionText="FIGURES 1439. Fistulatus species. 1419: F. athena (after Shang &amp; Zhang, 2003), 14: male pygofer, lateral view; 15: style, dorsal view; 16: connective, dorsal view; 17: aedeagus, ventral view; 18: aedeagus, dorsal view; 19: aedeagus, lateral view; 20 24: F. bidentatus (after Cen &amp; Cai, 2002), 20: male pygofer, dorsolateral view; 21: style, ventral view; 22: connective, ventral view; 23: aedeagus, ventral view; 24: aedeagus, lateral view; 2529: F. biconjugara (after Yang et al. 2024), 25: male pygofer, lateral view; 26: style, dorsal view; 27: connective, dorsal view; 28: aedeagus, ventral view; 29: aedeagus, lateral view; 3034: F. brevimarginalis (after Yang et al. 2024), 30: male pygofer, lateral view; 31. style, dorsal view; 32: connective, dorsal view; 33: aedeagus, ventral view; 34: aedeagus, lateral view; 3539: F. luteolus (after Cen &amp; Cai, 2002), 35: male pygofer, lateral view; 36: style, ventral view; 37: connective, ventral view; 38: aedeagus, ventral view; 39: aedeagus, lateral view." figureDoi="http://doi.org/10.5281/zenodo.12758869" httpUri="https://zenodo.org/record/12758869/files/figure.png" pageId="1" pageNumber="364">Fig. 24</figureCitation>
)................................................
<taxonomicName id="4C7E4D05FFF2CF741B53F83708B100DA" ID-CoL="6J6WS" authorityName="Cen &amp; Cai" authorityYear="2002" box="[1309,1437,1949,1971]" class="Insecta" family="Cicadellidae" genus="Fistulatus" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="364" phylum="Arthropoda" rank="species" species="bidentatus">
<emphasis id="B90AEA94FFF2CF741B53F83708B100DA" bold="true" box="[1309,1437,1949,1971]" italics="true" pageId="1" pageNumber="364">F. bidentatus</emphasis>
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<keyStep id="308A2E23FFF2CF741ED9F81208B00084" pageId="1" pageNumber="364">
<paragraph id="8BC13686FFF2CF741ED9F81208B000B9" blockId="1.[151,1437,1627,2030]" box="[151,1436,1979,2001]" pageId="1" pageNumber="364">
<keyLead id="308F95B3FFF2CF741ED9F81208B000B9" box="[151,1436,1979,2001]" pageId="1" pageNumber="364">
6 Aedeagal shaft with spikelike lateral preapical processes (
<figureCitation id="13452A03FFF2CF741D43F8120E5700B9" box="[781,891,1979,2001]" captionStart="FIGURES 513" captionStartId="3.[151,264,1846,1870]" captionTargetBox="[158,1429,188,1816]" captionTargetId="figure-17@3.[151,1435,181,1822]" captionTargetPageId="3" captionText="FIGURES 513. Fistulatus motuoensis Hassan &amp; Xing sp. nov. Male. 5. Genitalia (after maceration), lateral view; 6. Male pygofer, lateral view; 7. Valve and subgenital plate, ventral view; 8. Style, lateral view; 9. Connective, dorsal view; 10.Aedeagus, dorsal view; 11. Aedeagus, ventral view; 12. Aedeagus, lateral view; 13. Aedeagus, anteroventral view focusing on the apex." figureDoi="http://doi.org/10.5281/zenodo.12758861" httpUri="https://zenodo.org/record/12758861/files/figure.png" pageId="1" pageNumber="364">Figs 1013</figureCitation>
)........................................... 7
</keyLead>
</paragraph>
<paragraph id="8BC13686FFF2CF741ED9F87008B00084" blockId="1.[151,1437,1627,2030]" box="[151,1436,2008,2030]" pageId="1" pageNumber="364">
<keyLead id="308F95B3FFF2CF741ED9F87008B00084" box="[151,1436,2008,2030]" pageId="1" pageNumber="364">
- Aedeagal shaft not as above (
<figureCitation id="13452A03FFF2CF741FB2F8710F470084" box="[508,619,2008,2030]" captionStart="FIGURES 1439" captionStartId="5.[151,264,1631,1655]" captionTargetBox="[157,1429,187,1602]" captionTargetId="figure-17@5.[151,1435,181,1608]" captionTargetPageId="5" captionText="FIGURES 1439. Fistulatus species. 1419: F. athena (after Shang &amp; Zhang, 2003), 14: male pygofer, lateral view; 15: style, dorsal view; 16: connective, dorsal view; 17: aedeagus, ventral view; 18: aedeagus, dorsal view; 19: aedeagus, lateral view; 20 24: F. bidentatus (after Cen &amp; Cai, 2002), 20: male pygofer, dorsolateral view; 21: style, ventral view; 22: connective, ventral view; 23: aedeagus, ventral view; 24: aedeagus, lateral view; 2529: F. biconjugara (after Yang et al. 2024), 25: male pygofer, lateral view; 26: style, dorsal view; 27: connective, dorsal view; 28: aedeagus, ventral view; 29: aedeagus, lateral view; 3034: F. brevimarginalis (after Yang et al. 2024), 30: male pygofer, lateral view; 31. style, dorsal view; 32: connective, dorsal view; 33: aedeagus, ventral view; 34: aedeagus, lateral view; 3539: F. luteolus (after Cen &amp; Cai, 2002), 35: male pygofer, lateral view; 36: style, ventral view; 37: connective, ventral view; 38: aedeagus, ventral view; 39: aedeagus, lateral view." figureDoi="http://doi.org/10.5281/zenodo.12758869" httpUri="https://zenodo.org/record/12758869/files/figure.png" pageId="1" pageNumber="364">Figs 1718</figureCitation>
,
<figureCitation id="13452A03FFF2CF741C3AF8710FF90084" box="[628,725,2008,2029]" captionStart="FIGURES 4059" captionStartId="7.[151,264,1421,1445]" captionTargetBox="[157,1430,187,1391]" captionTargetId="figure-17@7.[151,1435,181,1397]" captionTargetPageId="7" captionText="FIGURES 4059. Fistulatus species. 4044: F. monkoboensis (after Shang &amp; Zhang, 2003), 40: male pygofer, lateral view; 41: subgenital plate, valve, style and connective, ventral view; 42: aedeagus, dorsal view; 43: aedeagus, ventral view; 44: aedeagus, lateral view; 4549: F. rectilineus (after Shang et al., 2006), 45: male pygofer, lateral view; 46: style, dorsal view; 47: connective, dorsal view; 48: aedeagus, ventral view; 49: aedeagus, lateral view; 5054: F. sinensis (after Zhang et al., 1997), 50: male pygofer, lateral view; 51: style, dorsal view; 52: connective, dorsal view; 53: aedeagus, posteroventral view; 54: aedeagus, lateral view; 5559: F. quadrispinosus (after Lu &amp; Zhang, 2014), 55: male pygofer, lateral view; 56: subgenital plate, valve, style and connective, ventral view; 57: aedeagus, lateral view; 58: aedeagus, ventral view; 59: aedeagus, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.12758871" httpUri="https://zenodo.org/record/12758871/files/figure.png" pageId="1" pageNumber="364">4243, 48</figureCitation>
)......................................................... 8
</keyLead>
</paragraph>
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<keyStep id="308A2E23FFF1CF771ED9FF3108B107A3" pageId="2" pageNumber="365">
<paragraph id="8BC13686FFF1CF771ED9FF3108B007C4" blockId="2.[151,1437,151,378]" box="[151,1436,151,173]" pageId="2" pageNumber="365">
<keyLead id="308F95B3FFF1CF771ED9FF3108B007C4" box="[151,1436,151,173]" pageId="2" pageNumber="365">
7 Aedeagal shaft with hammer-shaped apex in lateral view (
<figureCitation id="13452A03FFF1CF771D5EFF3E0E7A07C4" box="[784,854,151,173]" captionStart="FIGURES 513" captionStartId="3.[151,264,1846,1870]" captionTargetBox="[158,1429,188,1816]" captionTargetId="figure-17@3.[151,1435,181,1822]" captionTargetPageId="3" captionText="FIGURES 513. Fistulatus motuoensis Hassan &amp; Xing sp. nov. Male. 5. Genitalia (after maceration), lateral view; 6. Male pygofer, lateral view; 7. Valve and subgenital plate, ventral view; 8. Style, lateral view; 9. Connective, dorsal view; 10.Aedeagus, dorsal view; 11. Aedeagus, ventral view; 12. Aedeagus, lateral view; 13. Aedeagus, anteroventral view focusing on the apex." figureDoi="http://doi.org/10.5281/zenodo.12758861" httpUri="https://zenodo.org/record/12758861/files/figure.png" pageId="2" pageNumber="365">Fig. 12</figureCitation>
).............................
<emphasis id="B90AEA94FFF1CF771A8FFF3108B007C4" bold="true" box="[1217,1436,151,173]" pageId="2" pageNumber="365">
<taxonomicName id="4C7E4D05FFF1CF771A8FFF31086507C5" authorityName="Hassan &amp; Shah &amp; Xing" authorityYear="2024" box="[1217,1353,151,173]" class="Insecta" family="Cicadellidae" genus="Fistulatus" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="365" phylum="Arthropoda" rank="species" species="motuoensis" status="sp. nov.">
<emphasis id="B90AEA94FFF1CF771A8FFF31086507C5" bold="true" box="[1217,1353,151,173]" italics="true" pageId="2" pageNumber="365">F. motuoensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23957EFFFF1CF771B1EFF3108B007C4" box="[1360,1436,152,173]" pageId="2" pageNumber="365" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
<paragraph id="8BC13686FFF1CF771ED9FF1C08B107A3" blockId="2.[151,1437,151,378]" box="[151,1437,181,203]" pageId="2" pageNumber="365">
<keyLead id="308F95B3FFF1CF771ED9FF1C08B107A3" box="[151,1437,181,203]" pageId="2" pageNumber="365">
- Aedeagal shaft with somewhat digitate apex in lateral view (
<figureCitation id="13452A03FFF1CF771D69FF1C0E4107A2" box="[807,877,181,203]" captionStart="FIGURES 4059" captionStartId="7.[151,264,1421,1445]" captionTargetBox="[157,1430,187,1391]" captionTargetId="figure-17@7.[151,1435,181,1397]" captionTargetPageId="7" captionText="FIGURES 4059. Fistulatus species. 4044: F. monkoboensis (after Shang &amp; Zhang, 2003), 40: male pygofer, lateral view; 41: subgenital plate, valve, style and connective, ventral view; 42: aedeagus, dorsal view; 43: aedeagus, ventral view; 44: aedeagus, lateral view; 4549: F. rectilineus (after Shang et al., 2006), 45: male pygofer, lateral view; 46: style, dorsal view; 47: connective, dorsal view; 48: aedeagus, ventral view; 49: aedeagus, lateral view; 5054: F. sinensis (after Zhang et al., 1997), 50: male pygofer, lateral view; 51: style, dorsal view; 52: connective, dorsal view; 53: aedeagus, posteroventral view; 54: aedeagus, lateral view; 5559: F. quadrispinosus (after Lu &amp; Zhang, 2014), 55: male pygofer, lateral view; 56: subgenital plate, valve, style and connective, ventral view; 57: aedeagus, lateral view; 58: aedeagus, ventral view; 59: aedeagus, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.12758871" httpUri="https://zenodo.org/record/12758871/files/figure.png" pageId="2" pageNumber="365">Fig. 54</figureCitation>
)....................................
<taxonomicName id="4C7E4D05FFF1CF771B7BFF1C08B107A3" ID-CoL="6J6WR" authorityName="Zhang, Zhang &amp; Chen" authorityYear="1997" box="[1333,1437,181,202]" class="Insecta" family="Cicadellidae" genus="Fistulatus" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="365" phylum="Arthropoda" rank="species" species="sinensis">
<emphasis id="B90AEA94FFF1CF771B7BFF1C08B107A3" bold="true" box="[1333,1437,181,202]" italics="true" pageId="2" pageNumber="365">F. sinensis</emphasis>
</taxonomicName>
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</paragraph>
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<keyStep id="308A2E23FFF1CF771ED9FF7B08B0064B" pageId="2" pageNumber="365">
<paragraph id="8BC13686FFF1CF771ED9FF7B08B0066C" blockId="2.[151,1437,151,378]" pageId="2" pageNumber="365">
<keyLead id="308F95B3FFF1CF771ED9FF7B08B0066C" pageId="2" pageNumber="365">
8 Pygofer with processes directed medially (
<figureCitation id="13452A03FFF1CF771C35FF7B0FEF0781" box="[635,707,210,232]" captionStart="FIGURES 4059" captionStartId="7.[151,264,1421,1445]" captionTargetBox="[157,1430,187,1391]" captionTargetId="figure-17@7.[151,1435,181,1397]" captionTargetPageId="7" captionText="FIGURES 4059. Fistulatus species. 4044: F. monkoboensis (after Shang &amp; Zhang, 2003), 40: male pygofer, lateral view; 41: subgenital plate, valve, style and connective, ventral view; 42: aedeagus, dorsal view; 43: aedeagus, ventral view; 44: aedeagus, lateral view; 4549: F. rectilineus (after Shang et al., 2006), 45: male pygofer, lateral view; 46: style, dorsal view; 47: connective, dorsal view; 48: aedeagus, ventral view; 49: aedeagus, lateral view; 5054: F. sinensis (after Zhang et al., 1997), 50: male pygofer, lateral view; 51: style, dorsal view; 52: connective, dorsal view; 53: aedeagus, posteroventral view; 54: aedeagus, lateral view; 5559: F. quadrispinosus (after Lu &amp; Zhang, 2014), 55: male pygofer, lateral view; 56: subgenital plate, valve, style and connective, ventral view; 57: aedeagus, lateral view; 58: aedeagus, ventral view; 59: aedeagus, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.12758871" httpUri="https://zenodo.org/record/12758871/files/figure.png" pageId="2" pageNumber="365">Fig. 45</figureCitation>
); aedeagus with shaft relatively straight in lateral view (
<figureCitation id="13452A03FFF1CF771A93FF7B080F078E" box="[1245,1315,210,232]" captionStart="FIGURES 4059" captionStartId="7.[151,264,1421,1445]" captionTargetBox="[157,1430,187,1391]" captionTargetId="figure-17@7.[151,1435,181,1397]" captionTargetPageId="7" captionText="FIGURES 4059. Fistulatus species. 4044: F. monkoboensis (after Shang &amp; Zhang, 2003), 40: male pygofer, lateral view; 41: subgenital plate, valve, style and connective, ventral view; 42: aedeagus, dorsal view; 43: aedeagus, ventral view; 44: aedeagus, lateral view; 4549: F. rectilineus (after Shang et al., 2006), 45: male pygofer, lateral view; 46: style, dorsal view; 47: connective, dorsal view; 48: aedeagus, ventral view; 49: aedeagus, lateral view; 5054: F. sinensis (after Zhang et al., 1997), 50: male pygofer, lateral view; 51: style, dorsal view; 52: connective, dorsal view; 53: aedeagus, posteroventral view; 54: aedeagus, lateral view; 5559: F. quadrispinosus (after Lu &amp; Zhang, 2014), 55: male pygofer, lateral view; 56: subgenital plate, valve, style and connective, ventral view; 57: aedeagus, lateral view; 58: aedeagus, ventral view; 59: aedeagus, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.12758871" httpUri="https://zenodo.org/record/12758871/files/figure.png" pageId="2" pageNumber="365">Fig. 49</figureCitation>
)..................................................................................................
<taxonomicName id="4C7E4D05FFF1CF771B55FF5908B0066C" ID-CoL="6J794" authorityName="Shang &amp; Zhang" authorityYear="2006" box="[1307,1436,239,261]" class="Insecta" family="Cicadellidae" genus="Fistulatus" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="365" phylum="Arthropoda" rank="species" species="rectilineus">
<emphasis id="B90AEA94FFF1CF771B55FF5908B0066C" bold="true" box="[1307,1436,239,261]" italics="true" pageId="2" pageNumber="365">F. rectilineus</emphasis>
</taxonomicName>
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</paragraph>
<paragraph id="8BC13686FFF1CF771ED9FEA408B0064B" blockId="2.[151,1437,151,378]" box="[151,1436,269,291]" pageId="2" pageNumber="365">
<keyLead id="308F95B3FFF1CF771ED9FEA408B0064B" box="[151,1436,269,291]" pageId="2" pageNumber="365">
- Pygofer with processes directed ventrally (
<figureCitation id="13452A03FFF1CF771C32FEA40FE8064B" box="[636,708,269,291]" captionStart="FIGURES 4059" captionStartId="7.[151,264,1421,1445]" captionTargetBox="[157,1430,187,1391]" captionTargetId="figure-17@7.[151,1435,181,1397]" captionTargetPageId="7" captionText="FIGURES 4059. Fistulatus species. 4044: F. monkoboensis (after Shang &amp; Zhang, 2003), 40: male pygofer, lateral view; 41: subgenital plate, valve, style and connective, ventral view; 42: aedeagus, dorsal view; 43: aedeagus, ventral view; 44: aedeagus, lateral view; 4549: F. rectilineus (after Shang et al., 2006), 45: male pygofer, lateral view; 46: style, dorsal view; 47: connective, dorsal view; 48: aedeagus, ventral view; 49: aedeagus, lateral view; 5054: F. sinensis (after Zhang et al., 1997), 50: male pygofer, lateral view; 51: style, dorsal view; 52: connective, dorsal view; 53: aedeagus, posteroventral view; 54: aedeagus, lateral view; 5559: F. quadrispinosus (after Lu &amp; Zhang, 2014), 55: male pygofer, lateral view; 56: subgenital plate, valve, style and connective, ventral view; 57: aedeagus, lateral view; 58: aedeagus, ventral view; 59: aedeagus, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.12758871" httpUri="https://zenodo.org/record/12758871/files/figure.png" pageId="2" pageNumber="365">Fig. 40</figureCitation>
); aedeagus with shaft curved dorsally in lateral view (
<figureCitation id="13452A03FFF1CF771A84FEA4083C064B" box="[1226,1296,269,291]" captionStart="FIGURES 4059" captionStartId="7.[151,264,1421,1445]" captionTargetBox="[157,1430,187,1391]" captionTargetId="figure-17@7.[151,1435,181,1397]" captionTargetPageId="7" captionText="FIGURES 4059. Fistulatus species. 4044: F. monkoboensis (after Shang &amp; Zhang, 2003), 40: male pygofer, lateral view; 41: subgenital plate, valve, style and connective, ventral view; 42: aedeagus, dorsal view; 43: aedeagus, ventral view; 44: aedeagus, lateral view; 4549: F. rectilineus (after Shang et al., 2006), 45: male pygofer, lateral view; 46: style, dorsal view; 47: connective, dorsal view; 48: aedeagus, ventral view; 49: aedeagus, lateral view; 5054: F. sinensis (after Zhang et al., 1997), 50: male pygofer, lateral view; 51: style, dorsal view; 52: connective, dorsal view; 53: aedeagus, posteroventral view; 54: aedeagus, lateral view; 5559: F. quadrispinosus (after Lu &amp; Zhang, 2014), 55: male pygofer, lateral view; 56: subgenital plate, valve, style and connective, ventral view; 57: aedeagus, lateral view; 58: aedeagus, ventral view; 59: aedeagus, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.12758871" httpUri="https://zenodo.org/record/12758871/files/figure.png" pageId="2" pageNumber="365">Fig. 44</figureCitation>
).......... 9
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<keyStep id="308A2E23FFF1CF771ED9FE8308B00613" pageId="2" pageNumber="365">
<paragraph id="8BC13686FFF1CF771ED9FE8308B00629" blockId="2.[151,1437,151,378]" box="[151,1436,298,320]" pageId="2" pageNumber="365">
<keyLead id="308F95B3FFF1CF771ED9FE8308B00629" box="[151,1436,298,320]" pageId="2" pageNumber="365">
9 Connective with lateral processes (
<figureCitation id="13452A03FFF1CF771C7FFE830F540629" box="[561,632,298,320]" captionStart="FIGURES 1439" captionStartId="5.[151,264,1631,1655]" captionTargetBox="[157,1429,187,1602]" captionTargetId="figure-17@5.[151,1435,181,1608]" captionTargetPageId="5" captionText="FIGURES 1439. Fistulatus species. 1419: F. athena (after Shang &amp; Zhang, 2003), 14: male pygofer, lateral view; 15: style, dorsal view; 16: connective, dorsal view; 17: aedeagus, ventral view; 18: aedeagus, dorsal view; 19: aedeagus, lateral view; 20 24: F. bidentatus (after Cen &amp; Cai, 2002), 20: male pygofer, dorsolateral view; 21: style, ventral view; 22: connective, ventral view; 23: aedeagus, ventral view; 24: aedeagus, lateral view; 2529: F. biconjugara (after Yang et al. 2024), 25: male pygofer, lateral view; 26: style, dorsal view; 27: connective, dorsal view; 28: aedeagus, ventral view; 29: aedeagus, lateral view; 3034: F. brevimarginalis (after Yang et al. 2024), 30: male pygofer, lateral view; 31. style, dorsal view; 32: connective, dorsal view; 33: aedeagus, ventral view; 34: aedeagus, lateral view; 3539: F. luteolus (after Cen &amp; Cai, 2002), 35: male pygofer, lateral view; 36: style, ventral view; 37: connective, ventral view; 38: aedeagus, ventral view; 39: aedeagus, lateral view." figureDoi="http://doi.org/10.5281/zenodo.12758869" httpUri="https://zenodo.org/record/12758869/files/figure.png" pageId="2" pageNumber="365">Fig. 16</figureCitation>
); style with apical process slightly curved laterally, digitate (
<figureCitation id="13452A03FFF1CF771AF1FE8308280629" box="[1215,1284,298,320]" captionStart="FIGURES 1439" captionStartId="5.[151,264,1631,1655]" captionTargetBox="[157,1429,187,1602]" captionTargetId="figure-17@5.[151,1435,181,1608]" captionTargetPageId="5" captionText="FIGURES 1439. Fistulatus species. 1419: F. athena (after Shang &amp; Zhang, 2003), 14: male pygofer, lateral view; 15: style, dorsal view; 16: connective, dorsal view; 17: aedeagus, ventral view; 18: aedeagus, dorsal view; 19: aedeagus, lateral view; 20 24: F. bidentatus (after Cen &amp; Cai, 2002), 20: male pygofer, dorsolateral view; 21: style, ventral view; 22: connective, ventral view; 23: aedeagus, ventral view; 24: aedeagus, lateral view; 2529: F. biconjugara (after Yang et al. 2024), 25: male pygofer, lateral view; 26: style, dorsal view; 27: connective, dorsal view; 28: aedeagus, ventral view; 29: aedeagus, lateral view; 3034: F. brevimarginalis (after Yang et al. 2024), 30: male pygofer, lateral view; 31. style, dorsal view; 32: connective, dorsal view; 33: aedeagus, ventral view; 34: aedeagus, lateral view; 3539: F. luteolus (after Cen &amp; Cai, 2002), 35: male pygofer, lateral view; 36: style, ventral view; 37: connective, ventral view; 38: aedeagus, ventral view; 39: aedeagus, lateral view." figureDoi="http://doi.org/10.5281/zenodo.12758869" httpUri="https://zenodo.org/record/12758869/files/figure.png" pageId="2" pageNumber="365">Fig. 15</figureCitation>
)...
<taxonomicName id="4C7E4D05FFF1CF771B71FE8308B00629" ID-CoL="6J795" authorityName="Shang &amp; Zhang" authorityYear="2003" box="[1343,1436,298,320]" class="Insecta" family="Cicadellidae" genus="Fistulatus" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="365" phylum="Arthropoda" rank="species" species="athena">
<emphasis id="B90AEA94FFF1CF771B71FE8308B00629" bold="true" box="[1343,1436,298,320]" italics="true" pageId="2" pageNumber="365">F. athena</emphasis>
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</paragraph>
<paragraph id="8BC13686FFF1CF771ED9FEE108B00613" blockId="2.[151,1437,151,378]" pageId="2" pageNumber="365">
<keyLead id="308F95B3FFF1CF771ED9FEE108B00613" pageId="2" pageNumber="365">
- Connective without lateral processes; style with apical process strongly curved laterally, foot-like (
<figureCitation id="13452A03FFF1CF771ADBFEEE09F70634" box="[1173,1243,327,349]" captionStart="FIGURES 4059" captionStartId="7.[151,264,1421,1445]" captionTargetBox="[157,1430,187,1391]" captionTargetId="figure-17@7.[151,1435,181,1397]" captionTargetPageId="7" captionText="FIGURES 4059. Fistulatus species. 4044: F. monkoboensis (after Shang &amp; Zhang, 2003), 40: male pygofer, lateral view; 41: subgenital plate, valve, style and connective, ventral view; 42: aedeagus, dorsal view; 43: aedeagus, ventral view; 44: aedeagus, lateral view; 4549: F. rectilineus (after Shang et al., 2006), 45: male pygofer, lateral view; 46: style, dorsal view; 47: connective, dorsal view; 48: aedeagus, ventral view; 49: aedeagus, lateral view; 5054: F. sinensis (after Zhang et al., 1997), 50: male pygofer, lateral view; 51: style, dorsal view; 52: connective, dorsal view; 53: aedeagus, posteroventral view; 54: aedeagus, lateral view; 5559: F. quadrispinosus (after Lu &amp; Zhang, 2014), 55: male pygofer, lateral view; 56: subgenital plate, valve, style and connective, ventral view; 57: aedeagus, lateral view; 58: aedeagus, ventral view; 59: aedeagus, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.12758871" httpUri="https://zenodo.org/record/12758871/files/figure.png" pageId="2" pageNumber="365">Fig. 41</figureCitation>
).....................................................................................................
<taxonomicName id="4C7E4D05FFF1CF771AB8FECC08B00613" ID-CoL="6J78S" authorityName="Shang &amp; Zhang" authorityYear="2003" box="[1270,1436,356,378]" class="Insecta" family="Cicadellidae" genus="Fistulatus" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="365" phylum="Arthropoda" rank="species" species="monkoboensis">
<emphasis id="B90AEA94FFF1CF771AB8FECC08B00613" bold="true" box="[1270,1436,356,378]" italics="true" pageId="2" pageNumber="365">F. monkoboensis</emphasis>
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