497 lines
78 KiB
XML
497 lines
78 KiB
XML
<document id="EED4F1195A1A5DC6F97727A0B966DFA7" ID-CLB-Dataset="2931" ID-DOI="10.11646/zootaxa.5079.1.1" ID-GBIF-Dataset="010695ef-a142-42ad-81d7-2e7ed1f7e89e" ID-ISSN="1175-5326" ID-Zenodo-Dep="5766402" ID-ZooBank="2B118B81-7353-4A2C-A892-5466DFC83230" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1638952199435" checkinUser="plazi" docAuthor="Takano, Kohei Takenaka, Gao, Jian-Jun, Hu, Yao-Guang, Li, Nan-Nan, Yafuso, Masako, Suwito, Awit, Repin, Rimi, Pungga, Runi Anak Sylvester, Meleng, Paulus Ak, Kaliang, Clement Het, Chong, Lucy & Toda, Masanori J." docDate="2021" docId="03D75C2D457F2E28FF5260DDB70DB79D" docLanguage="en" docName="zootaxa.5079.1.1.pdf" docOrigin="Zootaxa 5079 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Colocasiomyia ecornuta Toda & Takano 2021, sp. nov." docType="treatment" docVersion="9" lastPageNumber="24" masterDocId="FFEE2455456A2E3FFFC56005B34CB33C" masterDocTitle="Phylogeny, taxonomy and flower-breeding ecology of the Colocasiomyia cristata species group (Diptera: Drosophilidae), with descriptions of ten new species" masterLastPageNumber="70" masterPageNumber="1" pageNumber="22" updateTime="1699472124804" updateUser="ExternalLinkService">
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<mods:title id="12FB06E0AE81E5D0D0253861F1EAF9BC">Phylogeny, taxonomy and flower-breeding ecology of the Colocasiomyia cristata species group (Diptera: Drosophilidae), with descriptions of ten new species</mods:title>
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<mods:namePart id="BC798E946605544044C388C6B71A2552">Takano, Kohei Takenaka</mods:namePart>
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<mods:namePart id="97F9A0C1058AA4307E67EC9322E8189F">Hu, Yao-Guang</mods:namePart>
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<mods:namePart id="C682F4F105B4324548008400D3BAC0F1">Yafuso, Masako</mods:namePart>
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<mods:namePart id="2D59FA2B105635946DE1CF442CD1736B">Toda, Masanori J.</mods:namePart>
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<mods:title id="D5B80B7B2C7EF90A642F5C7F0AF2FE1D">Zootaxa</mods:title>
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<mods:date id="E07662DCEB7CE9E9D31DC6689FDB6687">2021</mods:date>
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<mods:number id="9F6C04B25B0860C66302517A869ACE75">2021-12-08</mods:number>
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<mods:number id="A73EA4968A4EFF16790539517FA43391">5079</mods:number>
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<treatment id="03D75C2D457F2E28FF5260DDB70DB79D" ID-DOI="http://doi.org/10.5281/zenodo.5786114" ID-GBIF-Taxon="191651663" ID-Zenodo-Dep="5786114" LSID="urn:lsid:plazi:treatment:03D75C2D457F2E28FF5260DDB70DB79D" httpUri="http://treatment.plazi.org/id/03D75C2D457F2E28FF5260DDB70DB79D" lastPageId="23" lastPageNumber="24" pageId="21" pageNumber="22">
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<emphasis id="B90A3129457F2E2AFF5260DDB05DB3CF" bold="true" box="[151,785,216,243]" pageId="21" pageNumber="22">
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1)
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<taxonomicName id="4C7E96B8457F2E2AFF7260DDB1EAB3CF" ID-CoL="9HXQ3" authority="Toda & Takano" authorityName="Toda & Takano" authorityYear="2021" box="[183,678,216,243]" class="Insecta" family="Drosophilidae" genus="Colocasiomyia" kingdom="Animalia" order="Diptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="ecornuta" status="sp. nov.">
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<emphasis id="B90A3129457F2E2AFF7260DDB290B3CF" bold="true" box="[183,476,216,243]" italics="true" pageId="21" pageNumber="22">Colocasiomyia ecornuta</emphasis>
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Toda & Takano
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</taxonomicName>
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,
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<taxonomicNameLabel id="A2398C52457F2E2AFD7160DCB05DB3CF" box="[692,785,217,243]" pageId="21" pageNumber="22" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</paragraph>
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<subSubSection id="C364BEB0457F2E2AFF5260F9B2D0B22B" box="[151,412,252,279]" pageId="21" pageNumber="22" type="description">
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<paragraph id="8BC1ED3B457F2E2AFF5260F9B2D0B22B" blockId="21.[151,785,216,279]" box="[151,412,252,279]" pageId="21" pageNumber="22">
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(
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<figureCitation id="1345F1BE457F2E2AFF5A60F9B3B0B22B" box="[159,252,252,279]" captionStart="FIGURE 5" captionStartId="10.[151,250,1874,1898]" captionTargetBox="[151,1435,920,1850]" captionTargetId="figure-75@10.[151,1435,919,1851]" captionTargetPageId="10" captionText="FIGURE 5. Characters 7‒11 on the thorax (A‒D, dorsal view; E‒G, lateral view) of Colocasiomyia colocasiae (Duda) (A, E), C. grandis sp. nov. (B), C. cristata de Meijere (C, F), C. diconica (Toda & Okada) (D) and C. ecornuta sp. nov. (G). Abbreviations: a dc s, anterior dorsocentral setae; ap sctl s, apical scutellar seta; b sctl s, basal scutellar seta; p kepst s, posterior katepisternal seta; pprn s, postpronotal seta; trn sut, transverse suture. Scale bars: 0.5 mm in A‒D, 0.1 mm in E‒G." figureDoi="http://doi.org/10.5281/zenodo.5766412" httpUri="https://zenodo.org/record/5766412/files/figure.png" pageId="21" pageNumber="22">Figs 5G</figureCitation>
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,
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<figureCitation id="1345F1BE457F2E2AFECD60F8B261B22B" box="[264,301,253,279]" captionStart="FIGURE 9" captionStartId="13.[151,250,1780,1804]" captionTargetBox="[151,1435,181,1757]" captionTargetId="figure-18@13.[151,1435,181,1757]" captionTargetPageId="13" captionText="FIGURE 9. Male abdominal sternite VIs of Colocasiomyia colocasiae (Duda) (B in anteroventral view, C in lateral view), C. stamenicola (Carson & Okada) (D in anterior view) and all species of the C. cristata subgroup (A in ventral view, E‒R in anterior view): A, C. ecornuta sp. nov.; E, C. grandis sp. nov.; F, C. vieti sp. nov.; G, C. cristata de Meijere; H, C. matthewsi sp. nov.; I, C. sulawesiana Okada & Yafuso; J, C. kinabaluana sp. nov.; K, C. kotana sp. nov.; L, C. sabahana sp. nov.; M, C. sarawakana sp. nov.; N, C. pistilicola (Carson & Okada); O, C. xenalocasiae (Okada); P, C. diconica (Toda & Okada); Q, C. sumatrana sp. nov.; and R, C. leucocasiae sp. nov. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5766420" httpUri="https://zenodo.org/record/5766420/files/figure.png" pageId="21" pageNumber="22">9A</figureCitation>
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,
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<figureCitation id="1345F1BE457F2E2AFEFD60F9B227B22B" box="[312,363,252,279]" captionStart="FIGURE 15" captionStartId="21.[151,250,1936,1960]" captionTargetBox="[151,1435,860,1912]" captionTargetId="figure-242@21.[151,1435,860,1912]" captionTargetPageId="21" captionText="FIGURE 15. Habitus (upper, lateral view; lower, dorsal view) of species of the Colocasiomyia cristata subgroup.A, C. ecornuta sp. nov. ♂; B, C. grandis sp. nov. ♂; C, C. vieti sp. nov. ♂; D, C. cristata de Meijere ♂; E, C. sulawesiana Okada & Yafuso ♂; F, C. kinabaluana sp. nov. ♂; G, C. kotana sp. nov. ♂; and H, C. matthewsi sp. nov. ♂. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5766434" httpUri="https://zenodo.org/record/5766434/files/figure.png" pageId="21" pageNumber="22">15A</figureCitation>
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,
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<figureCitation id="1345F1BE457F2E2AFEB360F9B2DFB22B" box="[374,403,252,279]" captionStart="FIGURE 16" captionStartId="24.[151,250,1785,1809]" captionTargetBox="[242,1345,181,1761]" captionTargetId="figure-22@24.[242,1345,181,1761]" captionTargetPageId="24" captionText="FIGURE 16. Colocasiomyia ecornuta Toda & Takano, sp. nov. (♂ and ♀ paratypes and puparium from Soppeng, South Sulawesi, Indonesia). A, Head (frontal view); B, scutum and scutellum (dorsal view); C, D, abdominal sternites in ventral view (C, male; D, female); E, periphallic organs (caudolateral view; inset: cercal ventral lobe); F, phallal sheath (dorsal view); G, H, phallic organs (G, lateral view; H, ventral view); I peri-oviscapt (caudal view); J, oviscapt (ventral view); K, puparium (ventral habitus; inset: caudal-most portion at lower right and ventrolateral portion of segments at the upper right); L, mouth hook; M, cephalopharyngeal skeleton. Abbreviations: i vt s, inner vertical setae; sctl, scutellum. Scale bars: 0.1 mm in A–J and M, 1.0 mm in K." figureDoi="http://doi.org/10.5281/zenodo.5766438" httpUri="https://zenodo.org/record/5766438/files/figure.png" pageId="21" pageNumber="22">16</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="C364BEB0457F2E2AFF52613FB05BB26E" box="[151,791,314,338]" pageId="21" pageNumber="22" type="reference_group">
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<paragraph id="8BC1ED3B457F2E2AFF52613FB05BB26E" blockId="21.[151,791,314,338]" box="[151,791,314,338]" pageId="21" pageNumber="22">
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<treatmentCitationGroup id="AB6ECA15457F2E2AFF52613FB05BB26E" box="[151,791,314,338]" pageId="21" pageNumber="22">
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<taxonomicName id="4C7E96B8457F2E2AFF52613FB05FB26E" authority=": Sultana et al., 2006: 694" authorityName="Sultana" authorityPageNumber="694" authorityYear="2006" box="[151,787,314,338]" class="Insecta" family="Drosophilidae" genus="Colocasiomyia" kingdom="Animalia" order="Diptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="sp.3">
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<emphasis id="B90A3129457F2E2AFF52613FB263B26E" box="[151,303,314,338]" italics="true" pageId="21" pageNumber="22">Colocasiomyia</emphasis>
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sp.3
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<treatmentCitation id="0ADFCB2A457F2E2AFEA0613FB05FB26E" author="Sultana, F. & Hu, Y. G. & Toda, M. J. & Takenaka, K. & Yafuso, M." box="[357,787,314,338]" page="694" pageId="21" pageNumber="22" year="2006">
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aff.
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<emphasis id="B90A3129457F2E2AFE4B613FB147B26E" box="[398,523,314,338]" italics="true" pageId="21" pageNumber="22">sulawesiana</emphasis>
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:
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<bibRefCitation id="EFEF90CA457F2E2AFDDD613FB05FB26E" author="Sultana, F. & Hu, Y. G. & Toda, M. J. & Takenaka, K. & Yafuso, M." box="[536,787,314,338]" pageId="21" pageNumber="22" pagination="684 - 702" refId="ref33499" refString="Sultana, F., Hu, Y. G., Toda, M. J., Takenaka, K. & Yafuso, M. (2006) Phylogeny and classification of Colocasiomyia (Diptera, Drosophilidae), and its evolution of pollination mutualism with aroid plants. Systematic Entomology, 31, 684 - 702. https: // doi. org / 10.1111 / j. 1365 - 3113.2006.00344. x" type="journal article" year="2006">
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Sultana
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<emphasis id="B90A3129457F2E2AFDAE613EB1DBB26E" box="[619,663,314,338]" italics="true" pageId="21" pageNumber="22">et al</emphasis>
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., 2006: 694
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</bibRefCitation>
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</treatmentCitation>
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</taxonomicName>
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.
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</treatmentCitationGroup>
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</paragraph>
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</subSubSection>
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<subSubSection id="C364BEB0457F2E2AFF52617CB668B151" pageId="21" pageNumber="22" type="diagnosis">
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<paragraph id="8BC1ED3B457F2E2AFF52617CB668B151" blockId="21.[151,1437,377,836]" pageId="21" pageNumber="22">
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<emphasis id="B90A3129457F2E2AFF52617CB25EB2AF" bold="true" box="[151,274,377,403]" pageId="21" pageNumber="22">Diagnosis.</emphasis>
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Outer vertical seta absent (
|
||
<figureCitation id="1345F1BE457F2E2AFD92617FB18FB2A9" box="[599,707,378,405]" captionStart="FIGURE 16" captionStartId="24.[151,250,1785,1809]" captionTargetBox="[242,1345,181,1761]" captionTargetId="figure-22@24.[242,1345,181,1761]" captionTargetPageId="24" captionText="FIGURE 16. Colocasiomyia ecornuta Toda & Takano, sp. nov. (♂ and ♀ paratypes and puparium from Soppeng, South Sulawesi, Indonesia). A, Head (frontal view); B, scutum and scutellum (dorsal view); C, D, abdominal sternites in ventral view (C, male; D, female); E, periphallic organs (caudolateral view; inset: cercal ventral lobe); F, phallal sheath (dorsal view); G, H, phallic organs (G, lateral view; H, ventral view); I peri-oviscapt (caudal view); J, oviscapt (ventral view); K, puparium (ventral habitus; inset: caudal-most portion at lower right and ventrolateral portion of segments at the upper right); L, mouth hook; M, cephalopharyngeal skeleton. Abbreviations: i vt s, inner vertical setae; sctl, scutellum. Scale bars: 0.1 mm in A–J and M, 1.0 mm in K." figureDoi="http://doi.org/10.5281/zenodo.5766438" httpUri="https://zenodo.org/record/5766438/files/figure.png" pageId="21" pageNumber="22">Fig. 16A</figureCitation>
|
||
). Katepisternal setae minute (
|
||
<figureCitation id="1345F1BE457F2E2AFBDB617FB731B2A9" box="[1054,1149,378,405]" captionStart="FIGURE 5" captionStartId="10.[151,250,1874,1898]" captionTargetBox="[151,1435,920,1850]" captionTargetId="figure-75@10.[151,1435,919,1851]" captionTargetPageId="10" captionText="FIGURE 5. Characters 7‒11 on the thorax (A‒D, dorsal view; E‒G, lateral view) of Colocasiomyia colocasiae (Duda) (A, E), C. grandis sp. nov. (B), C. cristata de Meijere (C, F), C. diconica (Toda & Okada) (D) and C. ecornuta sp. nov. (G). Abbreviations: a dc s, anterior dorsocentral setae; ap sctl s, apical scutellar seta; b sctl s, basal scutellar seta; p kepst s, posterior katepisternal seta; pprn s, postpronotal seta; trn sut, transverse suture. Scale bars: 0.5 mm in A‒D, 0.1 mm in E‒G." figureDoi="http://doi.org/10.5281/zenodo.5766412" httpUri="https://zenodo.org/record/5766412/files/figure.png" pageId="21" pageNumber="22">Fig. 5G</figureCitation>
|
||
). Scutellum with several setulae scattered on dorsal surface (
|
||
<figureCitation id="1345F1BE457F2E2AFDDD619BB130B285" box="[536,636,414,441]" captionStart="FIGURE 16" captionStartId="24.[151,250,1785,1809]" captionTargetBox="[242,1345,181,1761]" captionTargetId="figure-22@24.[242,1345,181,1761]" captionTargetPageId="24" captionText="FIGURE 16. Colocasiomyia ecornuta Toda & Takano, sp. nov. (♂ and ♀ paratypes and puparium from Soppeng, South Sulawesi, Indonesia). A, Head (frontal view); B, scutum and scutellum (dorsal view); C, D, abdominal sternites in ventral view (C, male; D, female); E, periphallic organs (caudolateral view; inset: cercal ventral lobe); F, phallal sheath (dorsal view); G, H, phallic organs (G, lateral view; H, ventral view); I peri-oviscapt (caudal view); J, oviscapt (ventral view); K, puparium (ventral habitus; inset: caudal-most portion at lower right and ventrolateral portion of segments at the upper right); L, mouth hook; M, cephalopharyngeal skeleton. Abbreviations: i vt s, inner vertical setae; sctl, scutellum. Scale bars: 0.1 mm in A–J and M, 1.0 mm in K." figureDoi="http://doi.org/10.5281/zenodo.5766438" httpUri="https://zenodo.org/record/5766438/files/figure.png" pageId="21" pageNumber="22">Fig. 16B</figureCitation>
|
||
). Male abdominal sternite VI without any process (
|
||
<figureCitation id="1345F1BE457F2E2AFB62619BB64DB284" box="[1191,1281,414,440]" captionStart="FIGURE 9" captionStartId="13.[151,250,1780,1804]" captionTargetBox="[151,1435,181,1757]" captionTargetId="figure-18@13.[151,1435,181,1757]" captionTargetPageId="13" captionText="FIGURE 9. Male abdominal sternite VIs of Colocasiomyia colocasiae (Duda) (B in anteroventral view, C in lateral view), C. stamenicola (Carson & Okada) (D in anterior view) and all species of the C. cristata subgroup (A in ventral view, E‒R in anterior view): A, C. ecornuta sp. nov.; E, C. grandis sp. nov.; F, C. vieti sp. nov.; G, C. cristata de Meijere; H, C. matthewsi sp. nov.; I, C. sulawesiana Okada & Yafuso; J, C. kinabaluana sp. nov.; K, C. kotana sp. nov.; L, C. sabahana sp. nov.; M, C. sarawakana sp. nov.; N, C. pistilicola (Carson & Okada); O, C. xenalocasiae (Okada); P, C. diconica (Toda & Okada); Q, C. sumatrana sp. nov.; and R, C. leucocasiae sp. nov. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5766420" httpUri="https://zenodo.org/record/5766420/files/figure.png" pageId="21" pageNumber="22">Figs 9A</figureCitation>
|
||
,
|
||
<figureCitation id="1345F1BE457F2E2AFAC9619BB671B285" box="[1292,1341,414,441]" captionStart="FIGURE 16" captionStartId="24.[151,250,1785,1809]" captionTargetBox="[242,1345,181,1761]" captionTargetId="figure-22@24.[242,1345,181,1761]" captionTargetPageId="24" captionText="FIGURE 16. Colocasiomyia ecornuta Toda & Takano, sp. nov. (♂ and ♀ paratypes and puparium from Soppeng, South Sulawesi, Indonesia). A, Head (frontal view); B, scutum and scutellum (dorsal view); C, D, abdominal sternites in ventral view (C, male; D, female); E, periphallic organs (caudolateral view; inset: cercal ventral lobe); F, phallal sheath (dorsal view); G, H, phallic organs (G, lateral view; H, ventral view); I peri-oviscapt (caudal view); J, oviscapt (ventral view); K, puparium (ventral habitus; inset: caudal-most portion at lower right and ventrolateral portion of segments at the upper right); L, mouth hook; M, cephalopharyngeal skeleton. Abbreviations: i vt s, inner vertical setae; sctl, scutellum. Scale bars: 0.1 mm in A–J and M, 1.0 mm in K." figureDoi="http://doi.org/10.5281/zenodo.5766438" httpUri="https://zenodo.org/record/5766438/files/figure.png" pageId="21" pageNumber="22">16C</figureCitation>
|
||
). Cercus ventrally narrowed, strongly curved posteriad, apically forming somewhat triangular lobe bearing 1 long and many short setae (
|
||
<figureCitation id="1345F1BE457F2E2AFEDA61E3B2CFB13D" box="[287,387,486,513]" captionStart="FIGURE 16" captionStartId="24.[151,250,1785,1809]" captionTargetBox="[242,1345,181,1761]" captionTargetId="figure-22@24.[242,1345,181,1761]" captionTargetPageId="24" captionText="FIGURE 16. Colocasiomyia ecornuta Toda & Takano, sp. nov. (♂ and ♀ paratypes and puparium from Soppeng, South Sulawesi, Indonesia). A, Head (frontal view); B, scutum and scutellum (dorsal view); C, D, abdominal sternites in ventral view (C, male; D, female); E, periphallic organs (caudolateral view; inset: cercal ventral lobe); F, phallal sheath (dorsal view); G, H, phallic organs (G, lateral view; H, ventral view); I peri-oviscapt (caudal view); J, oviscapt (ventral view); K, puparium (ventral habitus; inset: caudal-most portion at lower right and ventrolateral portion of segments at the upper right); L, mouth hook; M, cephalopharyngeal skeleton. Abbreviations: i vt s, inner vertical setae; sctl, scutellum. Scale bars: 0.1 mm in A–J and M, 1.0 mm in K." figureDoi="http://doi.org/10.5281/zenodo.5766438" httpUri="https://zenodo.org/record/5766438/files/figure.png" pageId="21" pageNumber="22">Fig. 16E</figureCitation>
|
||
). Phallal sheath pubescent on dorsal surface, gently curved ventrad, wider than thick, distally narrowing, basally roundly dilated, apically knobbed, subapically with small, acute projection on right margin (
|
||
<figureCitation id="1345F1BE457F2E2AFAB5620FB3ABB175" captionStart="FIGURE 16" captionStartId="24.[151,250,1785,1809]" captionTargetBox="[242,1345,181,1761]" captionTargetId="figure-22@24.[242,1345,181,1761]" captionTargetPageId="24" captionText="FIGURE 16. Colocasiomyia ecornuta Toda & Takano, sp. nov. (♂ and ♀ paratypes and puparium from Soppeng, South Sulawesi, Indonesia). A, Head (frontal view); B, scutum and scutellum (dorsal view); C, D, abdominal sternites in ventral view (C, male; D, female); E, periphallic organs (caudolateral view; inset: cercal ventral lobe); F, phallal sheath (dorsal view); G, H, phallic organs (G, lateral view; H, ventral view); I peri-oviscapt (caudal view); J, oviscapt (ventral view); K, puparium (ventral habitus; inset: caudal-most portion at lower right and ventrolateral portion of segments at the upper right); L, mouth hook; M, cephalopharyngeal skeleton. Abbreviations: i vt s, inner vertical setae; sctl, scutellum. Scale bars: 0.1 mm in A–J and M, 1.0 mm in K." figureDoi="http://doi.org/10.5281/zenodo.5766438" httpUri="https://zenodo.org/record/5766438/files/figure.png" pageId="21" pageNumber="22">Fig. 16F–H</figureCitation>
|
||
). Gonocoxites broad, somewhat triangular plates (
|
||
<figureCitation id="1345F1BE457F2E2AFCD3622BB031B175" box="[790,893,558,585]" captionStart="FIGURE 16" captionStartId="24.[151,250,1785,1809]" captionTargetBox="[242,1345,181,1761]" captionTargetId="figure-22@24.[242,1345,181,1761]" captionTargetPageId="24" captionText="FIGURE 16. Colocasiomyia ecornuta Toda & Takano, sp. nov. (♂ and ♀ paratypes and puparium from Soppeng, South Sulawesi, Indonesia). A, Head (frontal view); B, scutum and scutellum (dorsal view); C, D, abdominal sternites in ventral view (C, male; D, female); E, periphallic organs (caudolateral view; inset: cercal ventral lobe); F, phallal sheath (dorsal view); G, H, phallic organs (G, lateral view; H, ventral view); I peri-oviscapt (caudal view); J, oviscapt (ventral view); K, puparium (ventral habitus; inset: caudal-most portion at lower right and ventrolateral portion of segments at the upper right); L, mouth hook; M, cephalopharyngeal skeleton. Abbreviations: i vt s, inner vertical setae; sctl, scutellum. Scale bars: 0.1 mm in A–J and M, 1.0 mm in K." figureDoi="http://doi.org/10.5281/zenodo.5766438" httpUri="https://zenodo.org/record/5766438/files/figure.png" pageId="21" pageNumber="22">Fig. 16G</figureCitation>
|
||
). Neither epiproct nor hypoproct pubescent (
|
||
<figureCitation id="1345F1BE457F2E2AFAB5622BB3F3B151" captionStart="FIGURE 16" captionStartId="24.[151,250,1785,1809]" captionTargetBox="[242,1345,181,1761]" captionTargetId="figure-22@24.[242,1345,181,1761]" captionTargetPageId="24" captionText="FIGURE 16. Colocasiomyia ecornuta Toda & Takano, sp. nov. (♂ and ♀ paratypes and puparium from Soppeng, South Sulawesi, Indonesia). A, Head (frontal view); B, scutum and scutellum (dorsal view); C, D, abdominal sternites in ventral view (C, male; D, female); E, periphallic organs (caudolateral view; inset: cercal ventral lobe); F, phallal sheath (dorsal view); G, H, phallic organs (G, lateral view; H, ventral view); I peri-oviscapt (caudal view); J, oviscapt (ventral view); K, puparium (ventral habitus; inset: caudal-most portion at lower right and ventrolateral portion of segments at the upper right); L, mouth hook; M, cephalopharyngeal skeleton. Abbreviations: i vt s, inner vertical setae; sctl, scutellum. Scale bars: 0.1 mm in A–J and M, 1.0 mm in K." figureDoi="http://doi.org/10.5281/zenodo.5766438" httpUri="https://zenodo.org/record/5766438/files/figure.png" pageId="21" pageNumber="22">Fig. 16I</figureCitation>
|
||
). Oviscapt distally narrowing, apically obliquely truncated and with 6–7 trichoid ovisensilla (
|
||
<figureCitation id="1345F1BE457F2E2AFB7E6257B65BB151" box="[1211,1303,594,621]" captionStart="FIGURE 16" captionStartId="24.[151,250,1785,1809]" captionTargetBox="[242,1345,181,1761]" captionTargetId="figure-22@24.[242,1345,181,1761]" captionTargetPageId="24" captionText="FIGURE 16. Colocasiomyia ecornuta Toda & Takano, sp. nov. (♂ and ♀ paratypes and puparium from Soppeng, South Sulawesi, Indonesia). A, Head (frontal view); B, scutum and scutellum (dorsal view); C, D, abdominal sternites in ventral view (C, male; D, female); E, periphallic organs (caudolateral view; inset: cercal ventral lobe); F, phallal sheath (dorsal view); G, H, phallic organs (G, lateral view; H, ventral view); I peri-oviscapt (caudal view); J, oviscapt (ventral view); K, puparium (ventral habitus; inset: caudal-most portion at lower right and ventrolateral portion of segments at the upper right); L, mouth hook; M, cephalopharyngeal skeleton. Abbreviations: i vt s, inner vertical setae; sctl, scutellum. Scale bars: 0.1 mm in A–J and M, 1.0 mm in K." figureDoi="http://doi.org/10.5281/zenodo.5766438" httpUri="https://zenodo.org/record/5766438/files/figure.png" pageId="21" pageNumber="22">Fig. 16J</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C364BEB0457F2E28FF026270B67AB09A" lastPageId="23" lastPageNumber="24" pageId="21" pageNumber="22" type="description">
|
||
<paragraph id="8BC1ED3B457F2E2AFF026270B2E2B078" blockId="21.[151,1437,377,836]" pageId="21" pageNumber="22">
|
||
<emphasis id="B90A3129457F2E2AFF026270B21DB1B3" bold="true" box="[199,337,629,655]" pageId="21" pageNumber="22">Description</emphasis>
|
||
(
|
||
<specimenCount id="9D7826B2457F2E2AFEA56273B239B1AC" box="[352,373,630,656]" pageId="21" pageNumber="22" type="male">♂</specimenCount>
|
||
and
|
||
<specimenCount id="9D7826B2457F2E2AFE6F6273B2F2B1AC" box="[426,446,630,656]" pageId="21" pageNumber="22" type="female">♀</specimenCount>
|
||
). Head. Supracervical setae 7–11 per side, nearly straight, as long as inner occipital setae. Anterior reclinate orbital seta long, situated slightly anteriorly to proclinate orbital seta; additional, interfrontal setulae on fronto-orbital plate present (
|
||
<figureCitation id="1345F1BE457F2E2AFD9362BBB18DB1E5" box="[598,705,702,729]" captionStart="FIGURE 16" captionStartId="24.[151,250,1785,1809]" captionTargetBox="[242,1345,181,1761]" captionTargetId="figure-22@24.[242,1345,181,1761]" captionTargetPageId="24" captionText="FIGURE 16. Colocasiomyia ecornuta Toda & Takano, sp. nov. (♂ and ♀ paratypes and puparium from Soppeng, South Sulawesi, Indonesia). A, Head (frontal view); B, scutum and scutellum (dorsal view); C, D, abdominal sternites in ventral view (C, male; D, female); E, periphallic organs (caudolateral view; inset: cercal ventral lobe); F, phallal sheath (dorsal view); G, H, phallic organs (G, lateral view; H, ventral view); I peri-oviscapt (caudal view); J, oviscapt (ventral view); K, puparium (ventral habitus; inset: caudal-most portion at lower right and ventrolateral portion of segments at the upper right); L, mouth hook; M, cephalopharyngeal skeleton. Abbreviations: i vt s, inner vertical setae; sctl, scutellum. Scale bars: 0.1 mm in A–J and M, 1.0 mm in K." figureDoi="http://doi.org/10.5281/zenodo.5766438" httpUri="https://zenodo.org/record/5766438/files/figure.png" pageId="21" pageNumber="22">Fig. 16A</figureCitation>
|
||
). Eye with fine, sparse interfacetal setulae (
|
||
<figureCitation id="1345F1BE457F2E2AFB7C62BBB668B1E5" box="[1209,1316,702,729]" captionStart="FIGURE 16" captionStartId="24.[151,250,1785,1809]" captionTargetBox="[242,1345,181,1761]" captionTargetId="figure-22@24.[242,1345,181,1761]" captionTargetPageId="24" captionText="FIGURE 16. Colocasiomyia ecornuta Toda & Takano, sp. nov. (♂ and ♀ paratypes and puparium from Soppeng, South Sulawesi, Indonesia). A, Head (frontal view); B, scutum and scutellum (dorsal view); C, D, abdominal sternites in ventral view (C, male; D, female); E, periphallic organs (caudolateral view; inset: cercal ventral lobe); F, phallal sheath (dorsal view); G, H, phallic organs (G, lateral view; H, ventral view); I peri-oviscapt (caudal view); J, oviscapt (ventral view); K, puparium (ventral habitus; inset: caudal-most portion at lower right and ventrolateral portion of segments at the upper right); L, mouth hook; M, cephalopharyngeal skeleton. Abbreviations: i vt s, inner vertical setae; sctl, scutellum. Scale bars: 0.1 mm in A–J and M, 1.0 mm in K." figureDoi="http://doi.org/10.5281/zenodo.5766438" httpUri="https://zenodo.org/record/5766438/files/figure.png" pageId="21" pageNumber="22">Fig. 16A</figureCitation>
|
||
). Distance between antennal sockets narrower than half of socket width (
|
||
<figureCitation id="1345F1BE457F2E2AFCA562E7B086B1C1" box="[864,970,738,765]" captionStart="FIGURE 16" captionStartId="24.[151,250,1785,1809]" captionTargetBox="[242,1345,181,1761]" captionTargetId="figure-22@24.[242,1345,181,1761]" captionTargetPageId="24" captionText="FIGURE 16. Colocasiomyia ecornuta Toda & Takano, sp. nov. (♂ and ♀ paratypes and puparium from Soppeng, South Sulawesi, Indonesia). A, Head (frontal view); B, scutum and scutellum (dorsal view); C, D, abdominal sternites in ventral view (C, male; D, female); E, periphallic organs (caudolateral view; inset: cercal ventral lobe); F, phallal sheath (dorsal view); G, H, phallic organs (G, lateral view; H, ventral view); I peri-oviscapt (caudal view); J, oviscapt (ventral view); K, puparium (ventral habitus; inset: caudal-most portion at lower right and ventrolateral portion of segments at the upper right); L, mouth hook; M, cephalopharyngeal skeleton. Abbreviations: i vt s, inner vertical setae; sctl, scutellum. Scale bars: 0.1 mm in A–J and M, 1.0 mm in K." figureDoi="http://doi.org/10.5281/zenodo.5766438" httpUri="https://zenodo.org/record/5766438/files/figure.png" pageId="21" pageNumber="22">Fig. 16A</figureCitation>
|
||
). Cibarial, medial sensilla (1–3 per side) in parallel rows narrower than sensilla campaniformia; posterior sensillum 1 per side. Supralateral setae outside prementum 3–4 per side.
|
||
</paragraph>
|
||
<caption id="DF01BDB3457F2E2AFF526795B780B4CC" ID-DOI="http://doi.org/10.5281/zenodo.5766434" ID-Zenodo-Dep="5766434" httpUri="https://zenodo.org/record/5766434/files/figure.png" pageId="21" pageNumber="22" startId="21.[151,250,1936,1960]" targetBox="[151,1435,860,1912]" targetPageId="21">
|
||
<paragraph id="8BC1ED3B457F2E2AFF526795B780B4CC" blockId="21.[151,1436,1936,2032]" pageId="21" pageNumber="22">
|
||
<emphasis id="B90A3129457F2E2AFF526795B252B494" bold="true" box="[151,286,1936,1960]" pageId="21" pageNumber="22">FIGURE 15.</emphasis>
|
||
Habitus (upper, lateral view; lower, dorsal view) of species of the
|
||
<taxonomicName id="4C7E96B8457F2E2AFC746795B7D7B494" authorityName="de Meijere" authorityYear="1914" box="[945,1179,1936,1960]" class="Insecta" family="Drosophilidae" genus="Colocasiomyia" kingdom="Animalia" order="Diptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="cristata">
|
||
<emphasis id="B90A3129457F2E2AFC746795B7D7B494" box="[945,1179,1936,1960]" italics="true" pageId="21" pageNumber="22">Colocasiomyia cristata</emphasis>
|
||
</taxonomicName>
|
||
subgroup.A,
|
||
<taxonomicName id="4C7E96B8457F2E2AFAE36794B6D0B494" authority="Takano & Gao & Hu & Li & Yafuso & Suwito & Repin & Pungga & Meleng & Kaliang & Chong & Toda, 2021" authorityName="Takano & Gao & Hu & Li & Yafuso & Suwito & Repin & Pungga & Meleng & Kaliang & Chong & Toda" authorityYear="2021" box="[1318,1436,1937,1960]" class="Insecta" family="Drosophilidae" genus="Colocasiomyia" kingdom="Animalia" order="Diptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="ecornuta" status="sp. nov.">
|
||
<emphasis id="B90A3129457F2E2AFAE36794B67BB494" box="[1318,1335,1937,1960]" italics="true" pageId="21" pageNumber="22">C</emphasis>
|
||
.
|
||
<emphasis id="B90A3129457F2E2AFA876794B6D0B494" box="[1346,1436,1937,1960]" italics="true" pageId="21" pageNumber="22">ecornuta</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B90A3129457F2E2AFF5267B1B3ABB4F0" bold="true" box="[151,231,1972,1996]" pageId="21" pageNumber="22">
|
||
<taxonomicNameLabel id="A2398C52457F2E2AFF5267B1B3ABB4F0" box="[151,231,1972,1996]" pageId="21" pageNumber="22" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
♂; B,
|
||
<taxonomicName id="4C7E96B8457F2E2AFEEE67B0B2DAB4F0" authority="Takano & Gao & Hu & Li & Yafuso & Suwito & Repin & Pungga & Meleng & Kaliang & Chong & Toda, 2021" authorityName="Takano & Gao & Hu & Li & Yafuso & Suwito & Repin & Pungga & Meleng & Kaliang & Chong & Toda" authorityYear="2021" box="[299,406,1972,1996]" class="Insecta" family="Drosophilidae" genus="Colocasiomyia" kingdom="Animalia" order="Diptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="grandis" status="sp. nov.">
|
||
<emphasis id="B90A3129457F2E2AFEEE67B0B270B4F0" box="[299,316,1973,1996]" italics="true" pageId="21" pageNumber="22">C</emphasis>
|
||
.
|
||
<emphasis id="B90A3129457F2E2AFE8C67B1B2DAB4F0" box="[329,406,1972,1996]" italics="true" pageId="21" pageNumber="22">grandis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B90A3129457F2E2AFE5967B1B2A0B4F0" bold="true" box="[412,492,1972,1996]" pageId="21" pageNumber="22">
|
||
<taxonomicNameLabel id="A2398C52457F2E2AFE5967B1B2A0B4F0" box="[412,492,1972,1996]" pageId="21" pageNumber="22" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
♂; C,
|
||
<taxonomicName id="4C7E96B8457F2E2AFDF567B0B135B4F0" authority="Takano & Gao & Hu & Li & Yafuso & Suwito & Repin & Pungga & Meleng & Kaliang & Chong & Toda, 2021" authorityName="Takano & Gao & Hu & Li & Yafuso & Suwito & Repin & Pungga & Meleng & Kaliang & Chong & Toda" authorityYear="2021" box="[560,633,1973,1996]" class="Insecta" family="Drosophilidae" genus="Colocasiomyia" kingdom="Animalia" order="Diptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="vieti" status="sp. nov.">
|
||
<emphasis id="B90A3129457F2E2AFDF567B0B10DB4F0" box="[560,577,1973,1996]" italics="true" pageId="21" pageNumber="22">C</emphasis>
|
||
.
|
||
<emphasis id="B90A3129457F2E2AFD8867B0B135B4F0" box="[589,633,1973,1996]" italics="true" pageId="21" pageNumber="22">vieti</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B90A3129457F2E2AFDBA67B1B183B4F0" bold="true" box="[639,719,1972,1996]" pageId="21" pageNumber="22">
|
||
<taxonomicNameLabel id="A2398C52457F2E2AFDBA67B1B183B4F0" box="[639,719,1972,1996]" pageId="21" pageNumber="22" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
♂; D,
|
||
<taxonomicName id="4C7E96B8457F2E2AFCD067B0B0BEB4F0" authority="de Meijere" authorityName="de Meijere" authorityYear="1914" box="[789,1010,1972,1996]" class="Insecta" family="Drosophilidae" genus="Colocasiomyia" kingdom="Animalia" order="Diptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="cristata">
|
||
<emphasis id="B90A3129457F2E2AFCD067B0B06AB4F0" box="[789,806,1973,1996]" italics="true" pageId="21" pageNumber="22">C</emphasis>
|
||
.
|
||
<emphasis id="B90A3129457F2E2AFCF767B0B033B4F0" box="[818,895,1973,1996]" italics="true" pageId="21" pageNumber="22">cristata</emphasis>
|
||
de Meijere
|
||
</taxonomicName>
|
||
♂; E,
|
||
<taxonomicName id="4C7E96B8457F2E2AFBF167B0B630B4F0" authority="Okada & Yafuso" authorityName="Okada & Yafuso" authorityYear="1989" box="[1076,1404,1972,1996]" class="Insecta" family="Drosophilidae" genus="Colocasiomyia" kingdom="Animalia" order="Diptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="sulawesiana">
|
||
<emphasis id="B90A3129457F2E2AFBF167B0B709B4F0" box="[1076,1093,1973,1996]" italics="true" pageId="21" pageNumber="22">C</emphasis>
|
||
.
|
||
<emphasis id="B90A3129457F2E2AFB9467B1B782B4F0" box="[1105,1230,1972,1996]" italics="true" pageId="21" pageNumber="22">sulawesiana</emphasis>
|
||
Okada & Yafuso
|
||
</taxonomicName>
|
||
♂; F,
|
||
<taxonomicName id="4C7E96B8457F2E2AFF7567DCB200B4CC" authority="Takano & Gao & Hu & Li & Yafuso & Suwito & Repin & Pungga & Meleng & Kaliang & Chong & Toda, 2021" authorityName="Takano & Gao & Hu & Li & Yafuso & Suwito & Repin & Pungga & Meleng & Kaliang & Chong & Toda" authorityYear="2021" box="[176,332,2008,2032]" class="Insecta" family="Drosophilidae" genus="Colocasiomyia" kingdom="Animalia" order="Diptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="kinabaluana" status="sp. nov.">
|
||
<emphasis id="B90A3129457F2E2AFF7567DCB38DB4CC" box="[176,193,2009,2032]" italics="true" pageId="21" pageNumber="22">C</emphasis>
|
||
.
|
||
<emphasis id="B90A3129457F2E2AFF0867DDB200B4CC" box="[205,332,2008,2032]" italics="true" pageId="21" pageNumber="22">kinabaluana</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B90A3129457F2E2AFE9767DDB2EFB4CC" bold="true" box="[338,419,2008,2032]" pageId="21" pageNumber="22">
|
||
<taxonomicNameLabel id="A2398C52457F2E2AFE9767DDB2EFB4CC" box="[338,419,2008,2032]" pageId="21" pageNumber="22" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
♂; G,
|
||
<taxonomicName id="4C7E96B8457F2E2AFE2C67DCB100B4CC" authority="Takano & Gao & Hu & Li & Yafuso & Suwito & Repin & Pungga & Meleng & Kaliang & Chong & Toda, 2021" authorityName="Takano & Gao & Hu & Li & Yafuso & Suwito & Repin & Pungga & Meleng & Kaliang & Chong & Toda" authorityYear="2021" box="[489,588,2008,2032]" class="Insecta" family="Drosophilidae" genus="Colocasiomyia" kingdom="Animalia" order="Diptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="kotana" status="sp. nov.">
|
||
<emphasis id="B90A3129457F2E2AFE2C67DCB2B6B4CC" box="[489,506,2009,2032]" italics="true" pageId="21" pageNumber="22">C</emphasis>
|
||
.
|
||
<emphasis id="B90A3129457F2E2AFDC267DDB100B4CC" box="[519,588,2008,2032]" italics="true" pageId="21" pageNumber="22">kotana</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B90A3129457F2E2AFD9767DDB1EEB4CC" bold="true" box="[594,674,2008,2032]" pageId="21" pageNumber="22">
|
||
<taxonomicNameLabel id="A2398C52457F2E2AFD9767DDB1EEB4CC" box="[594,674,2008,2032]" pageId="21" pageNumber="22" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
♂; and H,
|
||
<taxonomicName id="4C7E96B8457F2E2AFCD667DCB0D4B4CC" authority="Takano & Gao & Hu & Li & Yafuso & Suwito & Repin & Pungga & Meleng & Kaliang & Chong & Toda, 2021" authorityName="Takano & Gao & Hu & Li & Yafuso & Suwito & Repin & Pungga & Meleng & Kaliang & Chong & Toda" authorityYear="2021" box="[787,920,2008,2032]" class="Insecta" family="Drosophilidae" genus="Colocasiomyia" kingdom="Animalia" order="Diptera" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="matthewsi" status="sp. nov.">
|
||
<emphasis id="B90A3129457F2E2AFCD667DCB068B4CC" box="[787,804,2009,2032]" italics="true" pageId="21" pageNumber="22">C</emphasis>
|
||
.
|
||
<emphasis id="B90A3129457F2E2AFCF467DDB0D4B4CC" box="[817,920,2008,2032]" italics="true" pageId="21" pageNumber="22">matthewsi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B90A3129457F2E2AFC5B67DDB0A3B4CC" bold="true" box="[926,1007,2008,2032]" pageId="21" pageNumber="22">
|
||
<taxonomicNameLabel id="A2398C52457F2E2AFC5B67DDB0A3B4CC" box="[926,1007,2008,2032]" pageId="21" pageNumber="22" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
♂. Scale bars: 1 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF01BDB3457C2E29FF526580B61FB6FD" ID-DOI="http://doi.org/10.5281/zenodo.5766436" ID-Zenodo-Dep="5766436" httpUri="https://zenodo.org/record/5766436/files/figure.png" pageId="22" pageNumber="23" startId="22.[151,250,1413,1437]" targetBox="[151,1435,181,1389]" targetPageId="22">
|
||
<paragraph id="8BC1ED3B457C2E29FF526580B61FB6FD" blockId="22.[151,1436,1413,1473]" pageId="22" pageNumber="23">
|
||
<emphasis id="B90A3129457C2E29FF526580B26EB6A1" bold="true" box="[151,290,1413,1437]" pageId="22" pageNumber="23">FIGURE 15.</emphasis>
|
||
(Continued). I,
|
||
<taxonomicName id="4C7E96B8457C2E29FE086583B11DB6A1" authority="Takano & Gao & Hu & Li & Yafuso & Suwito & Repin & Pungga & Meleng & Kaliang & Chong & Toda, 2021" authorityName="Takano & Gao & Hu & Li & Yafuso & Suwito & Repin & Pungga & Meleng & Kaliang & Chong & Toda" authorityYear="2021" box="[461,593,1413,1437]" class="Insecta" family="Drosophilidae" genus="Colocasiomyia" kingdom="Animalia" order="Diptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="sabahana" status="sp. nov.">
|
||
<emphasis id="B90A3129457C2E29FE086583B292B6A1" box="[461,478,1414,1437]" italics="true" pageId="22" pageNumber="23">C</emphasis>
|
||
.
|
||
<emphasis id="B90A3129457C2E29FE2B6580B11DB6A1" box="[494,593,1413,1437]" italics="true" pageId="22" pageNumber="23">sabahana</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B90A3129457C2E29FD9C6580B1E0B6A1" bold="true" box="[601,684,1413,1437]" pageId="22" pageNumber="23">
|
||
<taxonomicNameLabel id="A2398C52457C2E29FD9C6580B1E0B6A1" box="[601,684,1413,1437]" pageId="22" pageNumber="23" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
♂; J,
|
||
<taxonomicName id="4C7E96B8457C2E29FD376583B0C3B6A1" authority="Takano & Gao & Hu & Li & Yafuso & Suwito & Repin & Pungga & Meleng & Kaliang & Chong & Toda, 2021" authorityName="Takano & Gao & Hu & Li & Yafuso & Suwito & Repin & Pungga & Meleng & Kaliang & Chong & Toda" authorityYear="2021" box="[754,911,1413,1437]" class="Insecta" family="Drosophilidae" genus="Colocasiomyia" kingdom="Animalia" order="Diptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="sarawakana" status="sp. nov.">
|
||
<emphasis id="B90A3129457C2E29FD376583B04FB6A1" box="[754,771,1414,1437]" italics="true" pageId="22" pageNumber="23">C</emphasis>
|
||
.
|
||
<emphasis id="B90A3129457C2E29FCD66580B0C3B6A1" box="[787,911,1413,1437]" italics="true" pageId="22" pageNumber="23">sarawakana</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B90A3129457C2E29FC5D6580B0A7B6A1" bold="true" box="[920,1003,1413,1437]" pageId="22" pageNumber="23">
|
||
<taxonomicNameLabel id="A2398C52457C2E29FC5D6580B0A7B6A1" box="[920,1003,1413,1437]" pageId="22" pageNumber="23" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
♂; K,
|
||
<taxonomicName id="4C7E96B8457C2E29FBFC6583B676B6A1" authority="(Okada)" baseAuthorityName="Okada" baseAuthorityYear="1980" box="[1081,1338,1413,1437]" class="Insecta" family="Drosophilidae" genus="Colocasiomyia" kingdom="Animalia" order="Diptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="xenalocasiae">
|
||
<emphasis id="B90A3129457C2E29FBFC6583B706B6A1" box="[1081,1098,1414,1437]" italics="true" pageId="22" pageNumber="23">C</emphasis>
|
||
.
|
||
<emphasis id="B90A3129457C2E29FB9F6580B792B6A1" box="[1114,1246,1413,1437]" italics="true" pageId="22" pageNumber="23">xenalocasiae</emphasis>
|
||
(Okada)
|
||
</taxonomicName>
|
||
♂; L,
|
||
<taxonomicName id="4C7E96B8457C2E29FA406583B2D7B6FD" authority="(Toda & Okada)" baseAuthorityName="Toda & Okada" baseAuthorityYear="1983" class="Insecta" family="Drosophilidae" genus="Colocasiomyia" kingdom="Animalia" order="Diptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="diconica">
|
||
<emphasis id="B90A3129457C2E29FA406583B6DAB6A1" box="[1413,1430,1414,1437]" italics="true" pageId="22" pageNumber="23">C</emphasis>
|
||
.
|
||
<emphasis id="B90A3129457C2E29FF5265ACB3A2B6FD" box="[151,238,1449,1473]" italics="true" pageId="22" pageNumber="23">diconica</emphasis>
|
||
(Toda & Okada)
|
||
</taxonomicName>
|
||
♂; M,
|
||
<taxonomicName id="4C7E96B8457C2E29FE2165AFB122B6FD" authority="Takano & Gao & Hu & Li & Yafuso & Suwito & Repin & Pungga & Meleng & Kaliang & Chong & Toda, 2021" authorityName="Takano & Gao & Hu & Li & Yafuso & Suwito & Repin & Pungga & Meleng & Kaliang & Chong & Toda" authorityYear="2021" box="[484,622,1450,1473]" class="Insecta" family="Sphaeroceridae" genus="Colocasiomia" kingdom="Animalia" order="Diptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="sumatrana" status="sp. nov.">
|
||
<emphasis id="B90A3129457C2E29FE2165AFB2B9B6FD" box="[484,501,1450,1473]" italics="true" pageId="22" pageNumber="23">C</emphasis>
|
||
.
|
||
<emphasis id="B90A3129457C2E29FDC765AFB122B6FD" box="[514,622,1450,1473]" italics="true" pageId="22" pageNumber="23">sumatrana</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B90A3129457C2E29FDB065ACB189B6FD" bold="true" box="[629,709,1449,1473]" pageId="22" pageNumber="23">
|
||
<taxonomicNameLabel id="A2398C52457C2E29FDB065ACB189B6FD" box="[629,709,1449,1473]" pageId="22" pageNumber="23" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
♂; N, ditto ♀; and O,
|
||
<taxonomicName id="4C7E96B8457C2E29FC6F65AFB70CB6FD" authority="Takano & Gao & Hu & Li & Yafuso & Suwito & Repin & Pungga & Meleng & Kaliang & Chong & Toda, 2021" authorityName="Takano & Gao & Hu & Li & Yafuso & Suwito & Repin & Pungga & Meleng & Kaliang & Chong & Toda" authorityYear="2021" box="[938,1088,1449,1473]" class="Insecta" family="Drosophilidae" genus="Colocasiomyia" kingdom="Animalia" order="Diptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="leucocasiae" status="sp. nov.">
|
||
<emphasis id="B90A3129457C2E29FC6F65AFB0F7B6FD" box="[938,955,1450,1473]" italics="true" pageId="22" pageNumber="23">C</emphasis>
|
||
.
|
||
<emphasis id="B90A3129457C2E29FC0D65ACB70CB6FD" box="[968,1088,1449,1473]" italics="true" pageId="22" pageNumber="23">leucocasiae</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B90A3129457C2E29FB8365ACB7DAB6FD" bold="true" box="[1094,1174,1449,1473]" pageId="22" pageNumber="23">
|
||
<taxonomicNameLabel id="A2398C52457C2E29FB8365ACB7DAB6FD" box="[1094,1174,1449,1473]" pageId="22" pageNumber="23" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
Scale bars: 1 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BC1ED3B457C2E29FF0265EAB1F7B54A" blockId="22.[151,1437,1519,2014]" pageId="22" pageNumber="23">
|
||
Thorax.Anterior dorsocentral setae just beside or slightly behind transverse suture (
|
||
<figureCitation id="1345F1BE457C2E29FB9865EAB78DB536" box="[1117,1217,1519,1546]" captionStart="FIGURE 16" captionStartId="24.[151,250,1785,1809]" captionTargetBox="[242,1345,181,1761]" captionTargetId="figure-22@24.[242,1345,181,1761]" captionTargetPageId="24" captionText="FIGURE 16. Colocasiomyia ecornuta Toda & Takano, sp. nov. (♂ and ♀ paratypes and puparium from Soppeng, South Sulawesi, Indonesia). A, Head (frontal view); B, scutum and scutellum (dorsal view); C, D, abdominal sternites in ventral view (C, male; D, female); E, periphallic organs (caudolateral view; inset: cercal ventral lobe); F, phallal sheath (dorsal view); G, H, phallic organs (G, lateral view; H, ventral view); I peri-oviscapt (caudal view); J, oviscapt (ventral view); K, puparium (ventral habitus; inset: caudal-most portion at lower right and ventrolateral portion of segments at the upper right); L, mouth hook; M, cephalopharyngeal skeleton. Abbreviations: i vt s, inner vertical setae; sctl, scutellum. Scale bars: 0.1 mm in A–J and M, 1.0 mm in K." figureDoi="http://doi.org/10.5281/zenodo.5766438" httpUri="https://zenodo.org/record/5766438/files/figure.png" pageId="22" pageNumber="23">Fig. 16B</figureCitation>
|
||
). Neither additional dorsocentral setae on presutural area nor prescutellar acrostichal setae present; acrostichal setulae in 4 rows (
|
||
<figureCitation id="1345F1BE457C2E29FAAA6616B385B56E" captionStart="FIGURE 16" captionStartId="24.[151,250,1785,1809]" captionTargetBox="[242,1345,181,1761]" captionTargetId="figure-22@24.[242,1345,181,1761]" captionTargetPageId="24" captionText="FIGURE 16. Colocasiomyia ecornuta Toda & Takano, sp. nov. (♂ and ♀ paratypes and puparium from Soppeng, South Sulawesi, Indonesia). A, Head (frontal view); B, scutum and scutellum (dorsal view); C, D, abdominal sternites in ventral view (C, male; D, female); E, periphallic organs (caudolateral view; inset: cercal ventral lobe); F, phallal sheath (dorsal view); G, H, phallic organs (G, lateral view; H, ventral view); I peri-oviscapt (caudal view); J, oviscapt (ventral view); K, puparium (ventral habitus; inset: caudal-most portion at lower right and ventrolateral portion of segments at the upper right); L, mouth hook; M, cephalopharyngeal skeleton. Abbreviations: i vt s, inner vertical setae; sctl, scutellum. Scale bars: 0.1 mm in A–J and M, 1.0 mm in K." figureDoi="http://doi.org/10.5281/zenodo.5766438" httpUri="https://zenodo.org/record/5766438/files/figure.png" pageId="22" pageNumber="23">Fig. 16B</figureCitation>
|
||
). Basal scutellar setae short, not reaching to half of apical scutellar setae; apical scutellar setae much nearer to each other than to basal scutellar seta (
|
||
<figureCitation id="1345F1BE457C2E29FD82665EB1E2B54A" box="[583,686,1627,1654]" captionStart="FIGURE 16" captionStartId="24.[151,250,1785,1809]" captionTargetBox="[242,1345,181,1761]" captionTargetId="figure-22@24.[242,1345,181,1761]" captionTargetPageId="24" captionText="FIGURE 16. Colocasiomyia ecornuta Toda & Takano, sp. nov. (♂ and ♀ paratypes and puparium from Soppeng, South Sulawesi, Indonesia). A, Head (frontal view); B, scutum and scutellum (dorsal view); C, D, abdominal sternites in ventral view (C, male; D, female); E, periphallic organs (caudolateral view; inset: cercal ventral lobe); F, phallal sheath (dorsal view); G, H, phallic organs (G, lateral view; H, ventral view); I peri-oviscapt (caudal view); J, oviscapt (ventral view); K, puparium (ventral habitus; inset: caudal-most portion at lower right and ventrolateral portion of segments at the upper right); L, mouth hook; M, cephalopharyngeal skeleton. Abbreviations: i vt s, inner vertical setae; sctl, scutellum. Scale bars: 0.1 mm in A–J and M, 1.0 mm in K." figureDoi="http://doi.org/10.5281/zenodo.5766438" httpUri="https://zenodo.org/record/5766438/files/figure.png" pageId="22" pageNumber="23">Fig. 16B</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BC1ED3B457C2E29FF02667AB043B5A5" blockId="22.[151,1437,1519,2014]" box="[199,783,1663,1690]" pageId="22" pageNumber="23">Wing. Costal setae in middle row all weak, trichoid.</paragraph>
|
||
<paragraph id="8BC1ED3B457C2E29FF0266A1B2FFB5DE" blockId="22.[151,1437,1519,2014]" pageId="22" pageNumber="23">Legs. Patch covered with only minute pubescence absent on anterodorsal portion of fore tarsomere I. Mid tibia with 2 apical, stout setae.</paragraph>
|
||
<paragraph id="8BC1ED3B457C2E29FF0266EEB7ACB416" blockId="22.[151,1437,1519,2014]" pageId="22" pageNumber="23">
|
||
Abdomen. Male: sternite III longer than wide; IV as wide as long (
|
||
<figureCitation id="1345F1BE457C2E29FC0266EEB77CB43A" box="[967,1072,1771,1798]" captionStart="FIGURE 16" captionStartId="24.[151,250,1785,1809]" captionTargetBox="[242,1345,181,1761]" captionTargetId="figure-22@24.[242,1345,181,1761]" captionTargetPageId="24" captionText="FIGURE 16. Colocasiomyia ecornuta Toda & Takano, sp. nov. (♂ and ♀ paratypes and puparium from Soppeng, South Sulawesi, Indonesia). A, Head (frontal view); B, scutum and scutellum (dorsal view); C, D, abdominal sternites in ventral view (C, male; D, female); E, periphallic organs (caudolateral view; inset: cercal ventral lobe); F, phallal sheath (dorsal view); G, H, phallic organs (G, lateral view; H, ventral view); I peri-oviscapt (caudal view); J, oviscapt (ventral view); K, puparium (ventral habitus; inset: caudal-most portion at lower right and ventrolateral portion of segments at the upper right); L, mouth hook; M, cephalopharyngeal skeleton. Abbreviations: i vt s, inner vertical setae; sctl, scutellum. Scale bars: 0.1 mm in A–J and M, 1.0 mm in K." figureDoi="http://doi.org/10.5281/zenodo.5766438" httpUri="https://zenodo.org/record/5766438/files/figure.png" pageId="22" pageNumber="23">Fig. 16C</figureCitation>
|
||
). Female: sternites III–V longer than wide; VI wider than long (
|
||
<figureCitation id="1345F1BE457C2E29FE32670AB113B416" box="[503,607,1807,1834]" captionStart="FIGURE 16" captionStartId="24.[151,250,1785,1809]" captionTargetBox="[242,1345,181,1761]" captionTargetId="figure-22@24.[242,1345,181,1761]" captionTargetPageId="24" captionText="FIGURE 16. Colocasiomyia ecornuta Toda & Takano, sp. nov. (♂ and ♀ paratypes and puparium from Soppeng, South Sulawesi, Indonesia). A, Head (frontal view); B, scutum and scutellum (dorsal view); C, D, abdominal sternites in ventral view (C, male; D, female); E, periphallic organs (caudolateral view; inset: cercal ventral lobe); F, phallal sheath (dorsal view); G, H, phallic organs (G, lateral view; H, ventral view); I peri-oviscapt (caudal view); J, oviscapt (ventral view); K, puparium (ventral habitus; inset: caudal-most portion at lower right and ventrolateral portion of segments at the upper right); L, mouth hook; M, cephalopharyngeal skeleton. Abbreviations: i vt s, inner vertical setae; sctl, scutellum. Scale bars: 0.1 mm in A–J and M, 1.0 mm in K." figureDoi="http://doi.org/10.5281/zenodo.5766438" httpUri="https://zenodo.org/record/5766438/files/figure.png" pageId="22" pageNumber="23">Fig. 16D</figureCitation>
|
||
). Male sternite VI not pubescent at least partly (
|
||
<figureCitation id="1345F1BE457C2E29FBBC670AB79EB416" box="[1145,1234,1807,1834]" captionStart="FIGURE 9" captionStartId="13.[151,250,1780,1804]" captionTargetBox="[151,1435,181,1757]" captionTargetId="figure-18@13.[151,1435,181,1757]" captionTargetPageId="13" captionText="FIGURE 9. Male abdominal sternite VIs of Colocasiomyia colocasiae (Duda) (B in anteroventral view, C in lateral view), C. stamenicola (Carson & Okada) (D in anterior view) and all species of the C. cristata subgroup (A in ventral view, E‒R in anterior view): A, C. ecornuta sp. nov.; E, C. grandis sp. nov.; F, C. vieti sp. nov.; G, C. cristata de Meijere; H, C. matthewsi sp. nov.; I, C. sulawesiana Okada & Yafuso; J, C. kinabaluana sp. nov.; K, C. kotana sp. nov.; L, C. sabahana sp. nov.; M, C. sarawakana sp. nov.; N, C. pistilicola (Carson & Okada); O, C. xenalocasiae (Okada); P, C. diconica (Toda & Okada); Q, C. sumatrana sp. nov.; and R, C. leucocasiae sp. nov. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5766420" httpUri="https://zenodo.org/record/5766420/files/figure.png" pageId="22" pageNumber="23">Fig. 9A</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BC1ED3B457C2E28FF026736B394B3C6" blockId="22.[151,1437,1519,2014]" lastBlockId="23.[151,1437,151,1185]" lastPageId="23" lastPageNumber="24" pageId="22" pageNumber="23">
|
||
Male terminalia. Epandrium pubescent on dorsal and posterolateral portions, roundish on posteroventral margin, with 2–3 setae near posterodorsal margin and 11–12 setae thicker than cercal setae on posteroventral portion of each side, but lacking phragma (
|
||
<figureCitation id="1345F1BE457C2E29FE0B677EB179B4AA" box="[462,565,1915,1942]" captionStart="FIGURE 16" captionStartId="24.[151,250,1785,1809]" captionTargetBox="[242,1345,181,1761]" captionTargetId="figure-22@24.[242,1345,181,1761]" captionTargetPageId="24" captionText="FIGURE 16. Colocasiomyia ecornuta Toda & Takano, sp. nov. (♂ and ♀ paratypes and puparium from Soppeng, South Sulawesi, Indonesia). A, Head (frontal view); B, scutum and scutellum (dorsal view); C, D, abdominal sternites in ventral view (C, male; D, female); E, periphallic organs (caudolateral view; inset: cercal ventral lobe); F, phallal sheath (dorsal view); G, H, phallic organs (G, lateral view; H, ventral view); I peri-oviscapt (caudal view); J, oviscapt (ventral view); K, puparium (ventral habitus; inset: caudal-most portion at lower right and ventrolateral portion of segments at the upper right); L, mouth hook; M, cephalopharyngeal skeleton. Abbreviations: i vt s, inner vertical setae; sctl, scutellum. Scale bars: 0.1 mm in A–J and M, 1.0 mm in K." figureDoi="http://doi.org/10.5281/zenodo.5766438" httpUri="https://zenodo.org/record/5766438/files/figure.png" pageId="22" pageNumber="23">Fig. 16E</figureCitation>
|
||
). Surstylus long, somewhat triangular plate with 2 stout teeth dorsally and 3 long, recurved, trichoid setae ventrally on distal margin (
|
||
<figureCitation id="1345F1BE457C2E29FCE4679AB0C4B486" box="[801,904,1951,1978]" captionStart="FIGURE 16" captionStartId="24.[151,250,1785,1809]" captionTargetBox="[242,1345,181,1761]" captionTargetId="figure-22@24.[242,1345,181,1761]" captionTargetPageId="24" captionText="FIGURE 16. Colocasiomyia ecornuta Toda & Takano, sp. nov. (♂ and ♀ paratypes and puparium from Soppeng, South Sulawesi, Indonesia). A, Head (frontal view); B, scutum and scutellum (dorsal view); C, D, abdominal sternites in ventral view (C, male; D, female); E, periphallic organs (caudolateral view; inset: cercal ventral lobe); F, phallal sheath (dorsal view); G, H, phallic organs (G, lateral view; H, ventral view); I peri-oviscapt (caudal view); J, oviscapt (ventral view); K, puparium (ventral habitus; inset: caudal-most portion at lower right and ventrolateral portion of segments at the upper right); L, mouth hook; M, cephalopharyngeal skeleton. Abbreviations: i vt s, inner vertical setae; sctl, scutellum. Scale bars: 0.1 mm in A–J and M, 1.0 mm in K." figureDoi="http://doi.org/10.5281/zenodo.5766438" httpUri="https://zenodo.org/record/5766438/files/figure.png" pageId="22" pageNumber="23">Fig. 16E</figureCitation>
|
||
). Median piece of subepandrial sclerite mostly bilobed into long, highly wrinkled lobes connected only at posterior end; lateral pieces less sclerotized broad plates. Cercus nearly entirely pubescent, with approximately 22 setae, including especially long, most ventral one, on dorsal, oval portion (
|
||
<figureCitation id="1345F1BE457D2E28FE4160BEB2A6B3EA" box="[388,490,187,214]" captionStart="FIGURE 16" captionStartId="24.[151,250,1785,1809]" captionTargetBox="[242,1345,181,1761]" captionTargetId="figure-22@24.[242,1345,181,1761]" captionTargetPageId="24" captionText="FIGURE 16. Colocasiomyia ecornuta Toda & Takano, sp. nov. (♂ and ♀ paratypes and puparium from Soppeng, South Sulawesi, Indonesia). A, Head (frontal view); B, scutum and scutellum (dorsal view); C, D, abdominal sternites in ventral view (C, male; D, female); E, periphallic organs (caudolateral view; inset: cercal ventral lobe); F, phallal sheath (dorsal view); G, H, phallic organs (G, lateral view; H, ventral view); I peri-oviscapt (caudal view); J, oviscapt (ventral view); K, puparium (ventral habitus; inset: caudal-most portion at lower right and ventrolateral portion of segments at the upper right); L, mouth hook; M, cephalopharyngeal skeleton. Abbreviations: i vt s, inner vertical setae; sctl, scutellum. Scale bars: 0.1 mm in A–J and M, 1.0 mm in K." figureDoi="http://doi.org/10.5281/zenodo.5766438" httpUri="https://zenodo.org/record/5766438/files/figure.png" pageId="23" pageNumber="24">Fig. 16E</figureCitation>
|
||
). Phallapodeme less sclerotized thin plate, nearly perpendicular to phallal axis (
|
||
<figureCitation id="1345F1BE457D2E28FAAA60BEB387B3C6" captionStart="FIGURE 16" captionStartId="24.[151,250,1785,1809]" captionTargetBox="[242,1345,181,1761]" captionTargetId="figure-22@24.[242,1345,181,1761]" captionTargetPageId="24" captionText="FIGURE 16. Colocasiomyia ecornuta Toda & Takano, sp. nov. (♂ and ♀ paratypes and puparium from Soppeng, South Sulawesi, Indonesia). A, Head (frontal view); B, scutum and scutellum (dorsal view); C, D, abdominal sternites in ventral view (C, male; D, female); E, periphallic organs (caudolateral view; inset: cercal ventral lobe); F, phallal sheath (dorsal view); G, H, phallic organs (G, lateral view; H, ventral view); I peri-oviscapt (caudal view); J, oviscapt (ventral view); K, puparium (ventral habitus; inset: caudal-most portion at lower right and ventrolateral portion of segments at the upper right); L, mouth hook; M, cephalopharyngeal skeleton. Abbreviations: i vt s, inner vertical setae; sctl, scutellum. Scale bars: 0.1 mm in A–J and M, 1.0 mm in K." figureDoi="http://doi.org/10.5281/zenodo.5766438" httpUri="https://zenodo.org/record/5766438/files/figure.png" pageId="23" pageNumber="24">Fig. 16G</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BC1ED3B457D2E28FF026106B731B222" blockId="23.[151,1437,151,1185]" box="[199,1149,259,286]" pageId="23" pageNumber="24">
|
||
Female terminalia. Oviscapt shorter than phallus (apodeme + sheath) (
|
||
<figureCitation id="1345F1BE457D2E28FC1E6106B73CB222" box="[987,1136,259,286]" captionStart="FIGURE 16" captionStartId="24.[151,250,1785,1809]" captionTargetBox="[242,1345,181,1761]" captionTargetId="figure-22@24.[242,1345,181,1761]" captionTargetPageId="24" captionText="FIGURE 16. Colocasiomyia ecornuta Toda & Takano, sp. nov. (♂ and ♀ paratypes and puparium from Soppeng, South Sulawesi, Indonesia). A, Head (frontal view); B, scutum and scutellum (dorsal view); C, D, abdominal sternites in ventral view (C, male; D, female); E, periphallic organs (caudolateral view; inset: cercal ventral lobe); F, phallal sheath (dorsal view); G, H, phallic organs (G, lateral view; H, ventral view); I peri-oviscapt (caudal view); J, oviscapt (ventral view); K, puparium (ventral habitus; inset: caudal-most portion at lower right and ventrolateral portion of segments at the upper right); L, mouth hook; M, cephalopharyngeal skeleton. Abbreviations: i vt s, inner vertical setae; sctl, scutellum. Scale bars: 0.1 mm in A–J and M, 1.0 mm in K." figureDoi="http://doi.org/10.5281/zenodo.5766438" httpUri="https://zenodo.org/record/5766438/files/figure.png" pageId="23" pageNumber="24">Fig. 16G,H,J</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BC1ED3B457D2E28FF026122B77AB029" blockId="23.[151,1437,151,1185]" pageId="23" pageNumber="24">
|
||
Indices (range of
|
||
<specimenCount id="9D7826B2457D2E28FE426122B2E6B27D" box="[391,426,295,321]" pageId="23" pageNumber="24" type="male">4♂</specimenCount>
|
||
and
|
||
<specimenCount id="9D7826B2457D2E28FE186122B14EB27D" box="[477,514,295,321]" pageId="23" pageNumber="24" type="female">4♀</specimenCount>
|
||
): FW/HW (frontal width / head width) = 0.59–0.66, ch/o (maximum width of gena / maximum diameter of eye) = 0.49–0.57, prorb (proclinate orbital seta length / posterior reclinate orbital seta length) = 0.86–0.95, rcorb (anterior reclinate orbital seta length / posterior reclinate orbital seta length) = 0.54–0.70, vb (subvibrissal seta length / vibrissa length) = 0.30–0.51, dcl (anterior dorsocentral seta length / posterior dorsocentral seta length) = 0.69–1.00, sctl (basal scutellar seta length / apical scutellar seta length) = 0.69–0.78, sterno (anterior katepisternal seta length / posterior katepisternal seta length) = 0.60–0.79, orbito (distance between proclinate and posterior reclinate orbital setae / distance between inner vertical and posterior reclinate orbital setae) = 0.42–0.58, dcp (distance between ipsilateral dorsocentral setae / distance between anterior dorsocentral setae) = 0.98–1.19, sctlp (distance between ipsilateral scutellar setae / distance between apical scutellar setae) = 2.00, C (2nd costal section between subcostal break and R
|
||
<subScript id="17FAEF7E457D2E28FD906278B122B1B7" attach="left" box="[597,622,637,651]" fontSize="6" pageId="23" pageNumber="24">2+3</subScript>
|
||
/ 3rd costal section between R
|
||
<subScript id="17FAEF7E457D2E28FC146278B0A6B1B7" attach="left" box="[977,1002,637,651]" fontSize="6" pageId="23" pageNumber="24">2+3</subScript>
|
||
and R
|
||
<subScript id="17FAEF7E457D2E28FBFC6278B71FB1B7" attach="left" box="[1081,1107,637,651]" fontSize="6" pageId="23" pageNumber="24">4+5</subScript>
|
||
) = 1.08–1.56, 4c (3rd costal section between R
|
||
<subScript id="17FAEF7E457D2E28FEA062A4B232B193" attach="left" box="[357,382,673,687]" fontSize="6" pageId="23" pageNumber="24">2+3</subScript>
|
||
and R
|
||
<subScript id="17FAEF7E457D2E28FE0C62A4B2AFB193" attach="left" box="[457,483,673,687]" fontSize="6" pageId="23" pageNumber="24">4+5</subScript>
|
||
/ M
|
||
<subScript id="17FAEF7E457D2E28FDCA62A4B15BB193" attach="left" box="[527,535,673,687]" fontSize="6" pageId="23" pageNumber="24">1</subScript>
|
||
between r-m and dm-m) = 1.16–1.51, 4v (M
|
||
<subScript id="17FAEF7E457D2E28FBD462A4B755B193" attach="left" box="[1041,1049,673,687]" fontSize="6" pageId="23" pageNumber="24">1</subScript>
|
||
between dm-m and wing margin / M
|
||
<subScript id="17FAEF7E457D2E28FF7562C0B3F4B1EF" attach="left" box="[176,184,709,723]" fontSize="6" pageId="23" pageNumber="24">1</subScript>
|
||
between r-m and dm-m) = 1.70–1.97, 5x (M
|
||
<subScript id="17FAEF7E457D2E28FD7D62C0B18CB1EF" attach="left" box="[696,704,709,723]" fontSize="6" pageId="23" pageNumber="24">4</subScript>
|
||
between dm-m and wing margin / dm-m between M
|
||
<subScript id="17FAEF7E457D2E28FADC62C0B66DB1EF" attach="left" box="[1305,1313,709,723]" fontSize="6" pageId="23" pageNumber="24">1</subScript>
|
||
and M
|
||
<subScript id="17FAEF7E457D2E28FAB662C0B637B1EF" attach="left" box="[1395,1403,709,723]" fontSize="6" pageId="23" pageNumber="24">4</subScript>
|
||
) = 1.76–2.00, ac (3rd costal section between R
|
||
<subScript id="17FAEF7E457D2E28FD4662ECB1D0B1CB" attach="left" box="[643,668,745,759]" fontSize="6" pageId="23" pageNumber="24">2+3</subScript>
|
||
and R
|
||
<subScript id="17FAEF7E457D2E28FD2362ECB04CB1CB" attach="left" box="[742,768,745,759]" fontSize="6" pageId="23" pageNumber="24">4+5</subScript>
|
||
/ distance between distal ends of R
|
||
<subScript id="17FAEF7E457D2E28FB4E62ECB7E9B1CB" attach="left" box="[1163,1189,745,759]" fontSize="6" pageId="23" pageNumber="24">4+5</subScript>
|
||
and M
|
||
<subScript id="17FAEF7E457D2E28FB3062ECB7B1B1CB" attach="left" box="[1269,1277,745,759]" fontSize="6" pageId="23" pageNumber="24">1</subScript>
|
||
) = 2.67–3.38, M (M
|
||
<subScript id="17FAEF7E457D2E28FF1C6308B3ADB027" attach="left" box="[217,225,781,795]" fontSize="6" pageId="23" pageNumber="24">4</subScript>
|
||
between dm-m and wing margin / M
|
||
<subScript id="17FAEF7E457D2E28FD466308B1C7B027" attach="left" box="[643,651,781,795]" fontSize="6" pageId="23" pageNumber="24">1</subScript>
|
||
between r-m and dm-m) = 0.49–0.59.
|
||
</paragraph>
|
||
<paragraph id="8BC1ED3B457D2E28FF02631AB67AB09A" blockId="23.[151,1437,151,1185]" pageId="23" pageNumber="24">
|
||
Puparium (3rd instar larva). Segments with stout spicules on ventral surface; some lateral ones large, clawlike; anterior spiracle absent; caudal segments elongate, very narrow, without spicules, ending in a pair of minute posterior spiracles (
|
||
<figureCitation id="1345F1BE457D2E28FEB16362B291B0BE" box="[372,477,871,898]" captionStart="FIGURE 16" captionStartId="24.[151,250,1785,1809]" captionTargetBox="[242,1345,181,1761]" captionTargetId="figure-22@24.[242,1345,181,1761]" captionTargetPageId="24" captionText="FIGURE 16. Colocasiomyia ecornuta Toda & Takano, sp. nov. (♂ and ♀ paratypes and puparium from Soppeng, South Sulawesi, Indonesia). A, Head (frontal view); B, scutum and scutellum (dorsal view); C, D, abdominal sternites in ventral view (C, male; D, female); E, periphallic organs (caudolateral view; inset: cercal ventral lobe); F, phallal sheath (dorsal view); G, H, phallic organs (G, lateral view; H, ventral view); I peri-oviscapt (caudal view); J, oviscapt (ventral view); K, puparium (ventral habitus; inset: caudal-most portion at lower right and ventrolateral portion of segments at the upper right); L, mouth hook; M, cephalopharyngeal skeleton. Abbreviations: i vt s, inner vertical setae; sctl, scutellum. Scale bars: 0.1 mm in A–J and M, 1.0 mm in K." figureDoi="http://doi.org/10.5281/zenodo.5766438" httpUri="https://zenodo.org/record/5766438/files/figure.png" pageId="23" pageNumber="24">Fig. 16K</figureCitation>
|
||
). Mouth hook: distal blade shorter than basal portion, apically more or less roundish, gently curved downward, with two lines of blunt teeth on submedial portion of ventral margin (
|
||
<figureCitation id="1345F1BE457D2E28FB01638EB665B09A" box="[1220,1321,907,934]" captionStart="FIGURE 16" captionStartId="24.[151,250,1785,1809]" captionTargetBox="[242,1345,181,1761]" captionTargetId="figure-22@24.[242,1345,181,1761]" captionTargetPageId="24" captionText="FIGURE 16. Colocasiomyia ecornuta Toda & Takano, sp. nov. (♂ and ♀ paratypes and puparium from Soppeng, South Sulawesi, Indonesia). A, Head (frontal view); B, scutum and scutellum (dorsal view); C, D, abdominal sternites in ventral view (C, male; D, female); E, periphallic organs (caudolateral view; inset: cercal ventral lobe); F, phallal sheath (dorsal view); G, H, phallic organs (G, lateral view; H, ventral view); I peri-oviscapt (caudal view); J, oviscapt (ventral view); K, puparium (ventral habitus; inset: caudal-most portion at lower right and ventrolateral portion of segments at the upper right); L, mouth hook; M, cephalopharyngeal skeleton. Abbreviations: i vt s, inner vertical setae; sctl, scutellum. Scale bars: 0.1 mm in A–J and M, 1.0 mm in K." figureDoi="http://doi.org/10.5281/zenodo.5766438" httpUri="https://zenodo.org/record/5766438/files/figure.png" pageId="23" pageNumber="24">Fig. 16L</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C364BEB0457D2E28FF0263AAB0B3B72D" pageId="23" pageNumber="24" type="materials_examined">
|
||
<paragraph id="8BC1ED3B457D2E28FF0263AAB1B7B0D1" blockId="23.[151,1437,151,1185]" pageId="23" pageNumber="24">
|
||
<materialsCitation id="3B16E766457D2E28FF0263AAB1B7B0D1" ID-GBIF-Occurrence="3418896845" collectingDate="2005-01-09" collectionCode="MZB, T" country="Indonesia" county="Emerged" location="Soppeng" municipality="Takano" pageId="23" pageNumber="24" specimenCount="1" specimenCount-male="1" stateProvince="Sulawesi Selatan" typeStatus="holotype">
|
||
<emphasis id="B90A3129457D2E28FF0263AAB277B0F5" bold="true" box="[199,315,943,969]" pageId="23" pageNumber="24">
|
||
<typeStatus id="54C55399457D2E28FF0263AAB27AB0F5" box="[199,310,943,969]" pageId="23" pageNumber="24" type="holotype">Holotype</typeStatus>
|
||
.
|
||
</emphasis>
|
||
<specimenCount id="9D7826B2457D2E28FE8663AAB214B0F5" box="[323,344,943,969]" pageId="23" pageNumber="24" type="male">♂</specimenCount>
|
||
(
|
||
<collectionCode id="ED6F75FE457D2E28FEAC63AAB2E7B0F5" box="[361,427,943,969]" country="Indonesia" httpUri="http://grbio.org/cool/b9gz-izvg" name="Museum Zoologicum Bogoriense" pageId="23" pageNumber="24" type="Museum">MZB</collectionCode>
|
||
), “
|
||
<location id="8EA1BBE0457D2E28FE0C63AAB17CB0F5" LSID="urn:lsid:plazi:treatment:03D75C2D457F2E28FF5260DDB70DB79D:8EA1BBE0457D2E28FE0C63AAB17CB0F5" box="[457,560,943,969]" country="Indonesia" county="Emerged" municipality="Takano" name="Soppeng" pageId="23" pageNumber="24" stateProvince="Sulawesi Selatan">Soppeng</location>
|
||
,
|
||
<collectingRegion id="49BA23D9457D2E28FDF863AAB1A2B0F5" box="[573,750,943,969]" country="Indonesia" name="Sulawesi Selatan" pageId="23" pageNumber="24">South Sulawesi</collectingRegion>
|
||
,
|
||
<collectingCountry id="F369ADAB457D2E28FD3E63AAB027B0F5" box="[763,875,943,969]" name="Indonesia" pageId="23" pageNumber="24">Indonesia</collectingCountry>
|
||
,
|
||
<date id="FFC0CBFB457D2E28FCB263ABB099B0F5" box="[887,981,942,969]" pageId="23" pageNumber="24" value="2005-01-09">
|
||
<collectingDate id="EF843213457D2E28FCB263ABB099B0F5" box="[887,981,942,969]" pageId="23" pageNumber="24" value="2005-01-09">9.i.2005</collectingDate>
|
||
</date>
|
||
, K.
|
||
<collectionCode id="ED6F75FE457D2E28FC3863AAB75CB0F5" box="[1021,1040,943,969]" name="Tavera, Department of Geology and Geophysics" pageId="23" pageNumber="24">T</collectionCode>
|
||
.
|
||
<collectingMunicipality id="6BA57741457D2E28FBDE63AAB721B0F5" box="[1051,1133,943,969]" pageId="23" pageNumber="24">Takano</collectingMunicipality>
|
||
/
|
||
<collectingCounty id="62A095B7457D2E28FB4063AAB7A6B0F5" box="[1157,1258,943,969]" pageId="23" pageNumber="24">Emerged</collectingCounty>
|
||
from puparium collected from an inflorescence of
|
||
<taxonomicName id="4C7E96B8457D2E28FDD863D6B1A4B0D1" box="[541,744,979,1005]" class="Liliopsida" family="Araceae" genus="Alocasia" kingdom="Plantae" order="Alismatales" pageId="23" pageNumber="24" phylum="Tracheophyta" rank="species" species="balgooyi">
|
||
<emphasis id="B90A3129457D2E28FDD863D6B1A4B0D1" box="[541,744,979,1005]" italics="true" pageId="23" pageNumber="24">Alocasia balgooyi</emphasis>
|
||
</taxonomicName>
|
||
”.
|
||
</materialsCitation>
|
||
</paragraph>
|
||
<paragraph id="8BC1ED3B457D2E28FF0263F2B0B3B72D" blockId="23.[151,1437,151,1185]" box="[199,1023,1015,1041]" pageId="23" pageNumber="24">
|
||
<materialsCitation id="3B16E766457D2E28FF0263F2B0B7B72D" ID-GBIF-Occurrence="3418896371" box="[199,1019,1015,1041]" collectingDate="2005-01-09" collectionCode="MZB, SEHU" country="Indonesia" county="Emerged" location="Soppeng" municipality="Takano" pageId="23" pageNumber="24" specimenCount="7" specimenCount-female="4" specimenCount-male="3" stateProvince="Sulawesi Selatan" typeStatus="paratype">
|
||
<emphasis id="B90A3129457D2E28FF0263F2B20AB72D" bold="true" box="[199,326,1015,1041]" pageId="23" pageNumber="24">
|
||
<typeStatus id="54C55399457D2E28FF0263F2B20DB72D" box="[199,321,1015,1041]" pageId="23" pageNumber="24" type="paratype">Paratypes</typeStatus>
|
||
.
|
||
</emphasis>
|
||
<collectingCountry id="F369ADAB457D2E28FE8863F2B2F2B72D" box="[333,446,1015,1041]" name="Indonesia" pageId="23" pageNumber="24">Indonesia</collectingCountry>
|
||
:
|
||
<specimenCount id="9D7826B2457D2E28FE0C63F2B2A2B72D" box="[457,494,1015,1041]" pageId="23" pageNumber="24" type="male">3♂</specimenCount>
|
||
,
|
||
<specimenCount id="9D7826B2457D2E28FE3F63F2B152B72D" box="[506,542,1015,1041]" pageId="23" pageNumber="24" type="female">4♀</specimenCount>
|
||
, same data as the holotype (
|
||
<collectionCode id="ED6F75FE457D2E28FC9C63F2B0D6B72D" box="[857,922,1015,1041]" country="Indonesia" httpUri="http://grbio.org/cool/b9gz-izvg" name="Museum Zoologicum Bogoriense" pageId="23" pageNumber="24" type="Museum">MZB</collectionCode>
|
||
,
|
||
<collectionCode id="ED6F75FE457D2E28FC6063F2B0BFB72D" box="[933,1011,1015,1041]" pageId="23" pageNumber="24">SEHU</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C364BEB0457D2E28FF02641FB1D7B70A" box="[199,667,1050,1078]" pageId="23" pageNumber="24" type="distribution">
|
||
<paragraph id="8BC1ED3B457D2E28FF02641FB1D7B70A" blockId="23.[151,1437,151,1185]" box="[199,667,1050,1078]" pageId="23" pageNumber="24">
|
||
<emphasis id="B90A3129457D2E28FF02641FB22DB708" bold="true" box="[199,353,1050,1076]" pageId="23" pageNumber="24">Distribution.</emphasis>
|
||
<collectingRegion id="49BA23D9457D2E28FEA2641EB280B709" box="[359,460,1051,1077]" country="Indonesia" name="Sulawesi Utara" pageId="23" pageNumber="24">Sulawesi</collectingRegion>
|
||
(
|
||
<collectingRegion id="49BA23D9457D2E28FE1E641EB1C3B70A" box="[475,655,1051,1078]" country="Indonesia" name="Sulawesi Selatan" pageId="23" pageNumber="24">South Sulawesi</collectingRegion>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C364BEB0457D2E28FF02643AB61FB741" pageId="23" pageNumber="24" type="discussion">
|
||
<paragraph id="8BC1ED3B457D2E28FF02643AB61FB741" blockId="23.[151,1437,151,1185]" pageId="23" pageNumber="24">
|
||
<emphasis id="B90A3129457D2E28FF02643AB277B765" bold="true" box="[199,315,1087,1113]" pageId="23" pageNumber="24">Remarks.</emphasis>
|
||
This species is peculiar in having many unusual characters as described in the diagnosis. Especially, the absence of any process on the male abdominal sternite VI is exceptional in the
|
||
<taxonomicName id="4C7E96B8457D2E28FBF06466B7E7B740" authorityName="de Meijere" authorityYear="1914" box="[1077,1195,1123,1149]" class="Insecta" family="Drosophilidae" genus="Colocasiomyia" kingdom="Animalia" order="Diptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="cristata">
|
||
<emphasis id="B90A3129457D2E28FBF06466B704B741" box="[1077,1096,1123,1149]" italics="true" pageId="23" pageNumber="24">C</emphasis>
|
||
.
|
||
<emphasis id="B90A3129457D2E28FB906466B7E7B740" box="[1109,1195,1123,1148]" italics="true" pageId="23" pageNumber="24">cristata</emphasis>
|
||
</taxonomicName>
|
||
species group.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C364BEB0457D2E28FF026482B70DB79D" box="[199,1089,1159,1185]" pageId="23" pageNumber="24" type="etymology">
|
||
<paragraph id="8BC1ED3B457D2E28FF026482B70DB79D" blockId="23.[151,1437,151,1185]" box="[199,1089,1159,1185]" pageId="23" pageNumber="24">
|
||
<emphasis id="B90A3129457D2E28FF026482B202B79D" bold="true" box="[199,334,1159,1185]" pageId="23" pageNumber="24">Etymology.</emphasis>
|
||
Referring to the lack of process on the male abdominal sternite VI.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |