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<document id="D9001A74F84E5A7CDD8B4AA775C23C01" ID-DOI="10.5281/zenodo.13244881" ID-ISSN="2345-7600" ID-Zenodo-Dep="13244881" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1722970250919" checkinUser="felipe" docAuthor="Hendrich, Lars &amp; Balke, Michael" docDate="2004" docId="03D687B7A73CFFB13AAEF8F2BD394ADF" docLanguage="en" docName="RafflesBZool.52.1.97-145.pdf" docOrigin="Raffles Bulletin of Zoology 52 (1)" docStyle="DocumentStyle:19380AF0FBA5B2F218DD253A34ABE57F.5:RafflesBZool.2003-2005.journal_article" docStyleId="19380AF0FBA5B2F218DD253A34ABE57F" docStyleName="RafflesBZool.2003-2005.journal_article" docStyleVersion="5" docTitle="Laccophilus medialis Sharp 1882" docType="treatment" docVersion="2" lastPageNumber="117" masterDocId="FFEFFFCFA72FFFA53903FF87BF1E4871" masterDocTitle="Aquatic Coleoptera Of Singapore: Species Richness, Ecology And Conservation #" masterLastPageNumber="145" masterPageNumber="97" pageNumber="116" updateTime="1722975545229" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title id="0FF209D6D6916600C82D31287D1F2BC5">Aquatic Coleoptera Of Singapore: Species Richness, Ecology And Conservation #</mods:title>
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<mods:date id="34505897728E62095E0FDEB3B7470CEB">2004</mods:date>
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<treatment id="03D687B7A73CFFB13AAEF8F2BD394ADF" LSID="urn:lsid:plazi:treatment:03D687B7A73CFFB13AAEF8F2BD394ADF" httpUri="http://treatment.plazi.org/id/03D687B7A73CFFB13AAEF8F2BD394ADF" lastPageId="20" lastPageNumber="117" pageId="19" pageNumber="116">
<subSubSection id="C365652AA73CFFB63AAEF8F2BA354FFC" box="[941,1323,1909,1933]" pageId="19" pageNumber="116" type="nomenclature">
<paragraph id="8BC036A1A73CFFB63AAEF8F2BA354FFC" blockId="19.[941,1323,1909,1962]" box="[941,1323,1909,1933]" pageId="19" pageNumber="116">
<heading id="D08881CDA73CFFB63AAEF8F2BA354FFC" bold="true" box="[941,1323,1909,1933]" centered="true" fontSize="10" level="2" pageId="19" pageNumber="116" reason="2">
<taxonomicName id="4C7F4D22A73CFFB63AAEF8F2BA354FFC" ID-CoL="3RM8W" authority="Sharp, 1882" authorityName="Sharp" authorityYear="1882" box="[941,1323,1909,1933]" class="Insecta" family="Dytiscidae" genus="Laccophilus" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="116" phylum="Arthropoda" rank="species" species="medialis">
<emphasis id="B90BEAB3A73CFFB63AAEF8F2BA354FFC" bold="true" box="[941,1323,1909,1933]" pageId="19" pageNumber="116">
<emphasis id="B90BEAB3A73CFFB63AAEF8F2BB844FFC" bold="true" box="[941,1178,1909,1933]" italics="true" pageId="19" pageNumber="116">Laccophilus medialis</emphasis>
<bibRefCitation id="EFEE4B50A73CFFB63DA2F8F2BA354FFC" author="Sharp, D" box="[1185,1323,1909,1933]" pageId="19" pageNumber="116" pagination="6 - 18" refId="ref41075" refString="Sharp, D., 1882. On aquatic carnivorous Coleoptera or Dytiscidae. Transactions of the Royal Dublin Society, (2) 2: 179 - 1003, pl. 6 - 18." type="journal article" year="1882">Sharp, 1882</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C365652AA73CFFB63D3FF814BB824FDB" box="[1084,1180,1939,1962]" pageId="19" pageNumber="116" type="description">
<paragraph id="8BC036A1A73CFFB63D3FF814BB824FDB" blockId="19.[941,1323,1909,1962]" box="[1084,1180,1939,1962]" pageId="19" pageNumber="116">
(
<figureCitation id="13442A24A73CFFB63D47F814BB8A4FDB" box="[1092,1172,1939,1962]" captionStart="Figs" captionStartId="15.[113,158,2006,2027]" captionTargetBox="[253,1312,191,1946]" captionTargetPageId="15" captionText="Figs. 18-23. 18.Hydaticus fabricii (MacLeay, 1825) (Fam. Dytiscidae, 10.5 mm); 19. Leiodytes nicobaricus (Redtenbacher, 1867) (Fam. Dytiscidae, 2.1 mm); 20. Hydrovatus rufoniger rufoniger (Clark, 1863) (Fam. Dytiscidae, 3.6 mm); 21. Laccophilus medialis Sharp, 1882 (Fam. Dytiscidae, 3.3 mm); 22. Copelatus minutissimus Balfour-Browne, 1939 (Fam. Dytiscidae, 4.1 mm); 23. Eretes griseus (Fabricius, 1781) (Fam. Dytiscidae, 13.5 mm)." figureDoi="http://doi.org/10.5281/zenodo.13244912" httpUri="https://zenodo.org/record/13244912/files/figure.png" pageId="19" pageNumber="116">Fig. 21</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C365652AA73CFFB63A2EF855BA6D4030" pageId="19" pageNumber="116" type="reference_group">
<paragraph id="8BC036A1A73CFFB63A2EF855BBAC4F97" blockId="19.[813,1452,2001,2113]" box="[813,1202,2001,2023]" pageId="19" pageNumber="116">
<treatmentCitationGroup id="AB6F118FA73CFFB63A2EF855BBAC4F97" box="[813,1202,2001,2023]" pageId="19" pageNumber="116">
<treatmentCitation id="0ADE10B0A73CFFB63A2EF855BBB04F97" author="Sharp, D" box="[813,1198,2001,2023]" page="309" pageId="19" pageNumber="116" year="1882">
<taxonomicName id="4C7F4D22A73CFFB63A2EF855BBB04F97" ID-CoL="3RM8W" authority="Sharp, 1882: 309" authorityName="Sharp" authorityPageNumber="309" authorityYear="1882" box="[813,1198,2001,2023]" class="Insecta" family="Dytiscidae" genus="Laccophilus" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="116" phylum="Arthropoda" rank="species" species="medialis">
<emphasis id="B90BEAB3A73CFFB63A2EF855BCE34F96" box="[813,1021,2002,2023]" italics="true" pageId="19" pageNumber="116">Laccophilus medialis</emphasis>
<bibRefCitation id="EFEE4B50A73CFFB63D00F856BBB04F97" author="Sharp, D" box="[1027,1198,2001,2022]" pageId="19" pageNumber="116" pagination="6 - 18" refId="ref41075" refString="Sharp, D., 1882. On aquatic carnivorous Coleoptera or Dytiscidae. Transactions of the Royal Dublin Society, (2) 2: 179 - 1003, pl. 6 - 18." type="journal article" year="1882">Sharp, 1882: 309</bibRefCitation>
</taxonomicName>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BC036A1A73CFFB63A2EF877BB2F4052" blockId="19.[813,1452,2001,2113]" pageId="19" pageNumber="116">
<treatmentCitationGroup id="AB6F118FA73CFFB63A2EF877BB2F4052" pageId="19" pageNumber="116">
<treatmentCitation id="0ADE10B0A73CFFB63A2EF877BBD24074" author="Guignot, F" box="[813,1228,2032,2053]" page="60" pageId="19" pageNumber="116" year="1956">
<taxonomicName id="4C7F4D22A73CFFB63A2EF877BBD24074" authority="Guignot, 1956 a: 60" authorityName="Guignot" authorityPageNumber="60" authorityYear="1956" box="[813,1228,2032,2053]" class="Insecta" family="Dytiscidae" genus="Laccophilus" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="116" phylum="Arthropoda" rank="species" species="cingulifer">
<emphasis id="B90BEAB3A73CFFB63A2EF877BB164074" box="[813,1032,2032,2053]" italics="true" pageId="19" pageNumber="116">Laccophilus cingulifer</emphasis>
<bibRefCitation id="EFEE4B50A73CFFB63D0DF877BBD24074" author="Guignot, F" box="[1038,1228,2032,2053]" pageId="19" pageNumber="116" pagination="51 - 60" refId="ref37377" refString="Guignot, F., 1956 a. Dytiscides recoltes par le Dr. L. Biro en Nouvelle Guinee et dans l'ile de Java (Coleoptera). Annales historico- Naturales Musei Nationales Hungarici (N. S), 7: 51 - 60." type="journal article" year="1956">Guignot, 1956a: 60</bibRefCitation>
</taxonomicName>
</treatmentCitation>
;
<treatmentCitation id="0ADE10B0A73CFFB63DD5F877BAB64074" author="Brancucci, M" box="[1238,1448,2032,2053]" page="326" pageId="19" pageNumber="116" year="1983">
<bibRefCitation id="EFEE4B50A73CFFB63DD5F877BAB64074" author="Brancucci, M" box="[1238,1448,2032,2053]" pageId="19" pageNumber="116" pagination="241 - 426" refId="ref36064" refString="Brancucci, M., 1983. Revision des especes est-palearctiques, orientales et australiennes du genre Laccophilus (Col. Dytiscidae). Entomologische Arbeiten aus dem Museum Georg Frey, 31 / 32: 241 - 426." type="journal article" year="1983">Brancucci, 1983: 326</bibRefCitation>
</treatmentCitation>
, and references therein.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BC036A1A73CFFB63A2EF7ABBA6D4030" blockId="19.[813,1452,2001,2113]" box="[813,1395,2092,2113]" pageId="19" pageNumber="116">
<treatmentCitationGroup id="AB6F118FA73CFFB63A2EF7ABBA6D4030" box="[813,1395,2092,2113]" pageId="19" pageNumber="116">
<treatmentCitation id="0ADE10B0A73CFFB63A2EF7ABBA6E4030" author="Nilsson, A" box="[813,1392,2092,2113]" page="309" pageId="19" pageNumber="116" year="1882">
<taxonomicName id="4C7F4D22A73CFFB63A2EF7ABBA6E4030" ID-CoL="3RM8W" authority="Sharp, 1882: 309 Nilsson, 2001: 246" authorityName="Sharp, 1882: 309 Nilsson" authorityYear="2001" box="[813,1392,2092,2113]" class="Insecta" family="Dytiscidae" genus="Laccophilus" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="116" phylum="Arthropoda" rank="species" species="medialis">
<emphasis id="B90BEAB3A73CFFB63A2EF7ABBCE04030" box="[813,1022,2092,2113]" italics="true" pageId="19" pageNumber="116">Laccophilus medialis</emphasis>
<bibRefCitation id="EFEE4B50A73CFFB63D07F7ABBBB54030" author="Sharp, D" box="[1028,1195,2092,2113]" pageId="19" pageNumber="116" pagination="6 - 18" refId="ref41075" refString="Sharp, D., 1882. On aquatic carnivorous Coleoptera or Dytiscidae. Transactions of the Royal Dublin Society, (2) 2: 179 - 1003, pl. 6 - 18." type="journal article" year="1882">Sharp, 1882: 309</bibRefCitation>
<bibRefCitation id="EFEE4B50A73CFFB63DB1F7ABBA6E4030" author="Nilsson, A" box="[1202,1392,2092,2113]" pageId="19" pageNumber="116" refId="ref39046" refString="Nilsson, A. N., 2001. Dytiscidae (Coleoptera). In: World Catalogue of Insects Volume 3. Apollo Books, Stenstrup, Denmark, 395 pp." type="book" year="2001">Nilsson, 2001: 246</bibRefCitation>
</taxonomicName>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C365652AA73BFFB1398BFF46BD564883" pageId="20" pageNumber="117" type="materials_examined">
<paragraph id="8BC036A1A73BFFB1398BFF46BD564883" blockId="20.[136,776,192,242]" pageId="20" pageNumber="117">
<emphasis id="B90BEAB3A73BFFB1398BFF46BE5548A4" bold="true" box="[136,331,192,214]" pageId="20" pageNumber="117">
<emphasis id="B90BEAB3A73BFFB1398BFF46BE5B48A7" bold="true" box="[136,325,193,214]" italics="true" pageId="20" pageNumber="117">Material examined</emphasis>
.
</emphasis>
-
<specimenCount id="9D79FD28A73BFFB1385EFF46BEAD48A7" box="[349,435,193,214]" count="1" pageId="20" pageNumber="117" type="female">1 female</specimenCount>
, “
<collectingCountry id="F3687631A73BFFB138C7FF46BD3548A7" box="[452,555,193,214]" name="Singapore" pageId="20" pageNumber="117">Singapore</collectingCountry>
, 1897, Biro” (
<collectionCode id="ED6EAE64A73BFFB13BB2FF46BDE248A7" box="[689,764,193,214]" pageId="20" pageNumber="117">TDMB</collectionCode>
). (
<bibRefCitation id="EFEE4B50A73BFFB1398CFF5ABE354883" author="Guignot, F" box="[143,299,221,242]" pageId="20" pageNumber="117" pagination="51 - 60" refId="ref37377" refString="Guignot, F., 1956 a. Dytiscides recoltes par le Dr. L. Biro en Nouvelle Guinee et dans l'ile de Java (Coleoptera). Annales historico- Naturales Musei Nationales Hungarici (N. S), 7: 51 - 60." type="journal article" year="1956">Guignot, 1956a</bibRefCitation>
;
<bibRefCitation id="EFEE4B50A73BFFB13835FF5ABEC74883" author="Brancucci, M" box="[310,473,221,242]" pageId="20" pageNumber="117" pagination="241 - 426" refId="ref36064" refString="Brancucci, M., 1983. Revision des especes est-palearctiques, orientales et australiennes du genre Laccophilus (Col. Dytiscidae). Entomologische Arbeiten aus dem Museum Georg Frey, 31 / 32: 241 - 426." type="journal article" year="1983">Brancucci, 1983</bibRefCitation>
). Not seen.
</paragraph>
</subSubSection>
<subSubSection id="C365652AA73BFFB1398BFE90BD00491F" pageId="20" pageNumber="117" type="distribution">
<paragraph id="8BC036A1A73BFFB1398BFE90BD00491F" blockId="20.[136,775,279,366]" pageId="20" pageNumber="117">
<emphasis id="B90BEAB3A73BFFB1398BFE90BE30495E" bold="true" box="[136,302,279,303]" italics="true" pageId="20" pageNumber="117">Distribution. </emphasis>
<collectingCountry id="F3687631A73BFFB1383BFE90BE6C495F" box="[312,370,279,302]" name="India" pageId="20" pageNumber="117">India</collectingCountry>
,
<collectingCountry id="F3687631A73BFFB13882FE90BECC495F" box="[385,466,279,302]" name="Bhutan" pageId="20" pageNumber="117">Bhutan</collectingCountry>
,
<collectingCountry id="F3687631A73BFFB138E3FE90BD52495F" box="[480,588,279,302]" name="Myanmar" pageId="20" pageNumber="117">Myanmar</collectingCountry>
,
<collectingCountry id="F3687631A73BFFB13B59FE90BDA3495F" box="[602,701,279,302]" name="Thailand" pageId="20" pageNumber="117">Thailand</collectingCountry>
,
<collectingCountry id="F3687631A73BFFB13BCFFE90BC1D495F" box="[716,771,279,302]" name="Laos" pageId="20" pageNumber="117">Laos</collectingCountry>
,
<collectingCountry id="F3687631A73BFFB1398BFEB0BFF3493F" box="[136,237,311,334]" name="Vietnam" pageId="20" pageNumber="117">Vietnam</collectingCountry>
, West
<collectingCountry id="F3687631A73BFFB13840FEB0BEB1493F" box="[323,431,311,334]" name="Malaysia" pageId="20" pageNumber="117">Malaysia</collectingCountry>
,
<collectingCountry id="F3687631A73BFFB138C3FEB0BD26493F" box="[448,568,311,334]" name="Singapore" pageId="20" pageNumber="117">Singapore</collectingCountry>
,
<collectingCountry id="F3687631A73BFFB13B44FEB0BDA9493F" box="[583,695,311,334]" name="Indonesia" pageId="20" pageNumber="117">Indonesia</collectingCountry>
[
<collectingRegion id="49BBF843A73BFFB13BC9FEB0BC1C493F" box="[714,770,311,334]" country="Indonesia" name="Jawa Timur" pageId="20" pageNumber="117">Java</collectingRegion>
, Sumbawa, Timor] (
<bibRefCitation id="EFEE4B50A73BFFB13858FED0BD0C491F" author="Brancucci, M" box="[347,530,343,366]" pageId="20" pageNumber="117" pagination="241 - 426" refId="ref36064" refString="Brancucci, M., 1983. Revision des especes est-palearctiques, orientales et australiennes du genre Laccophilus (Col. Dytiscidae). Entomologische Arbeiten aus dem Museum Georg Frey, 31 / 32: 241 - 426." type="journal article" year="1983">Brancucci, 1983</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C365652AA73BFFB1398BFE10BC1F4A3F" pageId="20" pageNumber="117" type="biology_ecology">
<paragraph id="8BC036A1A73BFFB1398BFE10BC1F4A3F" blockId="20.[136,775,407,590]" pageId="20" pageNumber="117">
<emphasis id="B90BEAB3A73BFFB1398BFE10BFE049DE" bold="true" box="[136,254,407,431]" italics="true" pageId="20" pageNumber="117">Ecology. </emphasis>
A lentic species restricted to forested areas. In West
<collectingCountry id="F3687631A73BFFB139C4FE30BE3449BF" box="[199,298,439,462]" name="Malaysia" pageId="20" pageNumber="117">Malaysia</collectingCountry>
the species was collected from shallow and shaded, more permanent forest pools in primary and second growth (older rubber plantations) lowland forest. The bottom consists of mud and a thick layer of rotten leaves. Probably the species still occurs in the Swamp Forest of Nee Soon.
</paragraph>
</subSubSection>
<subSubSection id="C365652AA73BFFB1398BFDF0BD394ADF" pageId="20" pageNumber="117" type="discussion">
<paragraph id="8BC036A1A73BFFB1398BFDF0BD394ADF" blockId="20.[136,775,631,686]" pageId="20" pageNumber="117">
<emphasis id="B90BEAB3A73BFFB1398BFDF0BE194AFE" bold="true" box="[136,263,631,655]" italics="true" pageId="20" pageNumber="117">Remarks. </emphasis>
We have not found this species in our survey and it is perhaps extinct in
<collectingCountry id="F3687631A73BFFB138B1FD10BD3D4ADF" box="[434,547,663,686]" name="Singapore" pageId="20" pageNumber="117">Singapore</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>