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<document id="682453D62A1D18F09E71FDDF2745B874" ID-DOI="10.5281/zenodo.5353544" ID-ISSN="1942-1354" ID-Zenodo-Dep="5353544" ID-ZooBank="772BDDF2-AD14-4950-B3B3-D6A2C9E403C2" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630210667410" checkinUser="felipe" docAuthor="Gordon, Robert D., F, Guillermo González &amp; Hanley, Guy A." docDate="2020" docId="03D687B4FF866A0BFF0AFE56FCCFFB5F" docLanguage="en" docName="0766.pdf" docOrigin="Insecta Mundi 2020 (766)" docStyle="DocumentStyle:6D85C66E4B52897F25DA165649BC06DC.3:InsectaMundi.2009-2020.journal_article" docStyleId="6D85C66E4B52897F25DA165649BC06DC" docStyleName="InsectaMundi.2009-2020.journal_article" docStyleVersion="3" docTitle="Calloeneis brooke Gordon and Hanley 2020, new species" docType="treatment" docVersion="4" lastPageNumber="16" masterDocId="FFEFFFCCFF976A1AFFCAFF83FFB6FF8F" masterDocTitle="South American Coccinellidae (Coleoptera), part XX: systematic revision of South American Calloeneis Grote (Cryptognathini)" masterLastPageNumber="25" masterPageNumber="1" pageNumber="16" updateTime="1699187120983" updateUser="plazi" zenodo-license-document="CC-BY-NC-3.0">
<mods:mods id="C15AAB80885F18E1BF8C6550D080F7BB" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="AB4B8645B6B2A96EEB63CBF40F57AE69">
<mods:title id="217DFFC3F7DB86C13D6850065061164D">South American Coccinellidae (Coleoptera), part XX: systematic revision of South American Calloeneis Grote (Cryptognathini)</mods:title>
</mods:titleInfo>
<mods:name id="781518A287A8DAB022F1107C58CB2297" type="personal">
<mods:role id="2DD3DFB64FAE284DE881DB32E38153D4">
<mods:roleTerm id="11C197F9558F47EB8EBBD539995FAC99">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="CEB2BA0DA08CBE90A2BFF527F4C77FEB">Gordon, Robert D.</mods:namePart>
<mods:affiliation id="3C25143383705A15B7148F6B845814DB">Northern Plains Entomology PO Box 65 Willow City, ND 58384, USA</mods:affiliation>
</mods:name>
<mods:name id="8935F7F60F1F217EE37D65010B9664E3" type="personal">
<mods:role id="739458EFF155573E634F9D1933CA8CFB">
<mods:roleTerm id="41D1FC5A2A54817A031ED275DA6AE376">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="25565D6DD3982A808FE7E5A28DCB0713">F, Guillermo González</mods:namePart>
<mods:affiliation id="FDEFB6A4A8D851FAE2FF955973EEA5E2">Nocedal 6455 La Reina Santiago, Chile</mods:affiliation>
</mods:name>
<mods:name id="35B1B47223D3559D7776F68AAAA72CDF" type="personal">
<mods:role id="91A6B209C2C96E57AEB7C5FF2B810B63">
<mods:roleTerm id="608C05328090BD3C0B85358F60FEB7D0">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="6685CA579CA149B9B3B24745E6342D17">Hanley, Guy A.</mods:namePart>
<mods:affiliation id="41A0D64490A740CA7A2801015CD92AFE">Northern Plains Entomology Minot, ND</mods:affiliation>
</mods:name>
<mods:typeOfResource id="C870EF44784B3B318C18C371E1381D43">text</mods:typeOfResource>
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<mods:titleInfo id="65909F47A17536EB1A0ED4D232C0C1FA">
<mods:title id="62BFF84E660AE81671D3DF42BF7C17A7">Insecta Mundi</mods:title>
</mods:titleInfo>
<mods:part id="EE5B02EE4A0F858D1047D3218A5BEC2C">
<mods:date id="714AC5F8B8D1BE7CF0086866B6813F98">2020</mods:date>
<mods:detail id="02AFD1C0CC054AB704EDFEA5F5BE8C14" type="pubDate">
<mods:number id="C880F1510A986762E83A3F31DE575DD8">2020-05-29</mods:number>
</mods:detail>
<mods:detail id="D8FC69980526EBA54BDA0222AF012413" type="volume">
<mods:number id="5583EE673E06CBC3BAA9EB4E85B5F32B">2020</mods:number>
</mods:detail>
<mods:detail id="4C89C194C245A6FC152904853957DAE3" type="issue">
<mods:number id="2DA0049978194B210B4B32A5593F6564">766</mods:number>
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<mods:start id="979553A3697040E4CD3D798A53478293">1</mods:start>
<mods:end id="7FC4C68E7BF95519FA326EF474DD86B9">25</mods:end>
</mods:extent>
</mods:part>
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<mods:classification id="246072AEC34EDD5C42FDD93B5E856E16">journal article</mods:classification>
<mods:identifier id="9594DF413A291E2216FC41850A91AE77" type="DOI">10.5281/zenodo.5353544</mods:identifier>
<mods:identifier id="E4AA32652D4A8364D081059850BC3893" type="ISSN">1942-1354</mods:identifier>
<mods:identifier id="32FC85E41037C5A723AC09BF68F46AA2" type="Zenodo-Dep">5353544</mods:identifier>
<mods:identifier id="07C24B0950BA1BB1BC235B27225DA3D1" type="ZooBank">772BDDF2-AD14-4950-B3B3-D6A2C9E403C2</mods:identifier>
</mods:mods>
<treatment id="03D687B4FF866A0BFF0AFE56FCCFFB5F" ID-DOI="http://doi.org/10.5281/zenodo.5451070" ID-Zenodo-Dep="5451070" LSID="urn:lsid:plazi:treatment:03D687B4FF866A0BFF0AFE56FCCFFB5F" httpUri="http://treatment.plazi.org/id/03D687B4FF866A0BFF0AFE56FCCFFB5F" lastPageNumber="16" pageId="17" pageNumber="16">
<subSubSection id="C3656529FF866A0BFF0AFE56FC11FE62" box="[192,935,469,493]" pageId="17" pageNumber="16" type="nomenclature">
<paragraph id="8BC036A2FF866A0BFF0AFE56FC11FE62" blockId="17.[192,935,469,493]" box="[192,935,469,493]" pageId="17" pageNumber="16">
<heading id="D08881CEFF866A0BFF0AFE56FC11FE62" bold="true" box="[192,935,469,493]" fontSize="10" level="2" pageId="17" pageNumber="16" reason="3">
<emphasis id="B90BEAB0FF866A0BFF0AFE56FC11FE62" bold="true" box="[192,935,469,493]" pageId="17" pageNumber="16">
20.
<taxonomicName id="4C7F4D21FF866A0BFF24FE56FD40FE62" authority="Gordon and Hanley" authorityName="Gordon and Hanley" authorityYear="2020" box="[238,758,469,493]" class="Insecta" family="Coccinellidae" genus="Calloeneis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="16" phylum="Arthropoda" rank="species" species="brooke" status="sp. nov.">
<emphasis id="B90BEAB0FF866A0BFF24FE56FE69FE62" bold="true" box="[238,479,469,493]" italics="true" pageId="17" pageNumber="16">Calloeneis brooke</emphasis>
Gordon and Hanley
</taxonomicName>
,
<taxonomicNameLabel id="A23857CBFF866A0BFCC9FE56FC11FE62" box="[771,935,469,493]" pageId="17" pageNumber="16" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3656529FF866A0BFF0AFD81FE64FBBA" pageId="17" pageNumber="16" type="description">
<paragraph id="8BC036A2FF866A0BFF0AFD81FB60FC54" blockId="17.[190,1442,514,987]" pageId="17" pageNumber="16">
<emphasis id="B90BEAB0FF866A0BFF0AFD81FD8DFD94" bold="true" box="[192,571,514,539]" pageId="17" pageNumber="16">
Description. Male
<typeStatus id="54C48800FF866A0BFE71FD80FD83FD94" box="[443,565,515,539]" pageId="17" pageNumber="16" type="holotype">holotype</typeStatus>
.
</emphasis>
Length
<quantity id="4C879B47FF866A0BFD6BFD81FD49FD95" box="[673,767,514,538]" metricMagnitude="-3" metricUnit="m" metricValue="1.6" pageId="17" pageNumber="16" unit="mm" value="1.6">1.6 mm</quantity>
, width
<quantity id="4C879B47FF866A0BFC92FD80FC00FD95" box="[856,950,515,539]" metricMagnitude="-3" metricUnit="m" metricValue="1.3" pageId="17" pageNumber="16" unit="mm" value="1.3">1.3 mm</quantity>
; dorsal surface shiny except head alutaceous. Color black; head, pronotum yellow (
<figureCitation id="13442A27FF866A0BFD73FDA1FCBDFDB5" box="[697,779,546,570]" captionStart="Figures 81–94" captionStartId="26.[192,287,1678,1700]" captionTargetBox="[208,1432,228,1604]" captionTargetId="figure-94@26.[192,1440,213,1646]" captionTargetPageId="26" captionText="Figures 81–94. Calloeneis spp. 81–85) Calloeneis alexandra. 81) Habitus. 82) Frons. 83) Penis guide ventral. 84) Penis guide lateral. 85) Penis. 86–90) Calloeneis brooke. 86) Habitus. 87) Frons. 88) Penis guide ventral. 89) Penis guide lateral. 90) Penis. 91–94) Calloeneis bethany. 91) Habitus. 92) Frons. 93) Penis guide ventral. 94) Penis guide lateral." figureDoi="http://doi.org/10.5281/zenodo.5353560" httpUri="https://zenodo.org/record/5353560/files/figure.png" pageId="17" pageNumber="16">Fig. 86</figureCitation>
); venter yellow except prosternum yellow, mesosternum reddish brown, metasternum dark brown, epipleuron yellowish brown. Head with punctures not visible; pronotal punctures small separated by a diameter or less; elytral punctures small, faint, nearly invisible, separated by about four times a diameter; prosternal punctures small, dense, separated by less than a diameter; mesosternal punctures apparently absent; metasternal punctures small, widely spaced, absent in lateral 1/3; basal abdominal ventrite with small punctures separated by about a diameter, ventrites 2–4 with fine punctures not clearly visible; ventrite 5 finely, densely punctured. Head with frons slightly wider than eye, clypeus widened, apex truncate (
<figureCitation id="13442A27FF866A0BFCC4FC81FCD2FC94" box="[782,868,770,795]" captionStart="Figures 81–94" captionStartId="26.[192,287,1678,1700]" captionTargetBox="[208,1432,228,1604]" captionTargetId="figure-94@26.[192,1440,213,1646]" captionTargetPageId="26" captionText="Figures 81–94. Calloeneis spp. 81–85) Calloeneis alexandra. 81) Habitus. 82) Frons. 83) Penis guide ventral. 84) Penis guide lateral. 85) Penis. 86–90) Calloeneis brooke. 86) Habitus. 87) Frons. 88) Penis guide ventral. 89) Penis guide lateral. 90) Penis. 91–94) Calloeneis bethany. 91) Habitus. 92) Frons. 93) Penis guide ventral. 94) Penis guide lateral." figureDoi="http://doi.org/10.5281/zenodo.5353560" httpUri="https://zenodo.org/record/5353560/files/figure.png" pageId="17" pageNumber="16">Fig. 87</figureCitation>
); eye canthus short, less than half width of eye. Prosternum with lateral carina on each side curved, extended from apex of intercoxal process to lateral margin of prosternum. Epipleuron not descending externally. Postcoxal line on basal abdominal ventrite curved, extended to rear margin of ventrite. Genitalia with penis guide slightly longer than paramere, wide, curved from base to widely rounded apex; paramere slender, slightly curved, apically rounded (
<figureCitation id="13442A27FF866A0BFF01FC21FEF9FC34" box="[203,335,930,955]" captionStart="Figures 81–94" captionStartId="26.[192,287,1678,1700]" captionTargetBox="[208,1432,228,1604]" captionTargetId="figure-94@26.[192,1440,213,1646]" captionTargetPageId="26" captionText="Figures 81–94. Calloeneis spp. 81–85) Calloeneis alexandra. 81) Habitus. 82) Frons. 83) Penis guide ventral. 84) Penis guide lateral. 85) Penis. 86–90) Calloeneis brooke. 86) Habitus. 87) Frons. 88) Penis guide ventral. 89) Penis guide lateral. 90) Penis. 91–94) Calloeneis bethany. 91) Habitus. 92) Frons. 93) Penis guide ventral. 94) Penis guide lateral." figureDoi="http://doi.org/10.5281/zenodo.5353560" httpUri="https://zenodo.org/record/5353560/files/figure.png" pageId="17" pageNumber="16">Fig. 88, 89</figureCitation>
); penis robust, apical 1/5 abruptly narrowed, apex acute; basal capsule slender, inner arm slightly curved with rounded apex, outer arm short, apex obliquely truncate (
<figureCitation id="13442A27FF866A0BFBBBFC41FB7FFC54" box="[1137,1225,962,987]" captionStart="Figures 81–94" captionStartId="26.[192,287,1678,1700]" captionTargetBox="[208,1432,228,1604]" captionTargetId="figure-94@26.[192,1440,213,1646]" captionTargetPageId="26" captionText="Figures 81–94. Calloeneis spp. 81–85) Calloeneis alexandra. 81) Habitus. 82) Frons. 83) Penis guide ventral. 84) Penis guide lateral. 85) Penis. 86–90) Calloeneis brooke. 86) Habitus. 87) Frons. 88) Penis guide ventral. 89) Penis guide lateral. 90) Penis. 91–94) Calloeneis bethany. 91) Habitus. 92) Frons. 93) Penis guide ventral. 94) Penis guide lateral." figureDoi="http://doi.org/10.5281/zenodo.5353560" httpUri="https://zenodo.org/record/5353560/files/figure.png" pageId="17" pageNumber="16">Fig. 90</figureCitation>
).
</paragraph>
<paragraph id="8BC036A2FF866A0BFF0AFC73FE05FB87" blockId="17.[192,435,1008,1032]" box="[192,435,1008,1032]" pageId="17" pageNumber="16">
<emphasis id="B90BEAB0FF866A0BFF0AFC73FE9BFB87" bold="true" box="[192,301,1008,1032]" pageId="17" pageNumber="16">Female.</emphasis>
Unknown.
</paragraph>
<paragraph id="8BC036A2FF866A0BFF0AFB9EFE64FBBA" blockId="17.[192,466,1053,1077]" box="[192,466,1053,1077]" pageId="17" pageNumber="16">
<emphasis id="B90BEAB0FF866A0BFF0AFB9EFEFAFBBA" bold="true" box="[192,332,1053,1077]" pageId="17" pageNumber="16">Variation.</emphasis>
Unknown.
</paragraph>
</subSubSection>
<subSubSection id="C3656529FF866A0BFF0AFBC8FC5DFB0C" pageId="17" pageNumber="16" type="materials_examined">
<paragraph id="8BC036A2FF866A0BFF0AFBC8FC5DFB0C" blockId="17.[190,1439,1098,1155]" pageId="17" pageNumber="16">
<emphasis id="B90BEAB0FF866A0BFF0AFBC8FE33FBED" bold="true" box="[192,389,1098,1123]" pageId="17" pageNumber="16">Type material.</emphasis>
<materialsCitation id="3B173CFFFF866A0BFE40FBC9FC5DFB0C" collectingDate="1926-06-20" collectingDateMax="1995-10" collectingDateMin="1926-06-20" collectionCode="USNM, BMNH" collectorName="H. E. Box &amp; Barbosa, G. V. &amp; Plot B Malaise" country="Venezuela" location="Petare" pageId="17" pageNumber="16" specimenCount="1" typeStatus="holotype">
<typeStatus id="54C48800FF866A0BFE40FBC9FE41FBED" box="[394,503,1098,1122]" pageId="17" pageNumber="16" type="holotype">Holotype</typeStatus>
male:
<collectingCountry id="F3687632FF866A0BFD8EFBC9FD73FBED" box="[580,709,1098,1122]" name="Venezuela" pageId="17" pageNumber="16">Venezuela</collectingCountry>
, nr.
<location id="8EA06079FF866A0BFD3CFBC8FCFFFBED" LSID="urn:lsid:plazi:treatment:03D687B4FF866A0BFF0AFE56FCCFFB5F:8EA06079FF866A0BFD3CFBC8FCFFFBED" box="[758,841,1099,1122]" country="Venezuela" name="Petare" pageId="17" pageNumber="16">Petare</location>
,
<date id="FFC11062FF866A0BFC99FBC9FC6FFBED" box="[851,985,1098,1122]" pageId="17" pageNumber="16" value="1926-06-20">
<collectingDate id="EF85E98AFF866A0BFC99FBC9FC6FFBED" box="[851,985,1098,1122]" pageId="17" pageNumber="16" value="1926-06-20">20.VI.1926</collectingDate>
</date>
,
<collectorName id="268A5374FF866A0BFC29FBC8FBE5FBED" box="[995,1107,1099,1122]" pageId="17" pageNumber="16">H. E. Box</collectorName>
collector (
<collectionCode id="ED6EAE67FF866A0BFB01FBC8FA92FBEC" box="[1227,1316,1099,1123]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="17" pageNumber="16" type="Museum">USNM</collectionCode>
).
<collectorName id="268A5374FF866A0BFAFFFBC9FF41FB0C" pageId="17" pageNumber="16">Barbosa, G.V.</collectorName>
,
<collectorName id="268A5374FF866A0BFEC9FBE9FE0EFB0D" box="[259,440,1130,1154]" pageId="17" pageNumber="16">Plot B Malaise</collectorName>
5,
<date id="FFC11062FF866A0BFE16FBE9FD37FB0D" box="[476,641,1130,1154]" pageId="17" pageNumber="16" value="1995-10">
<collectingDate id="EF85E98AFF866A0BFE16FBE9FD37FB0D" box="[476,641,1130,1154]" pageId="17" pageNumber="16" value="1995-10">October 1995</collectingDate>
</date>
, BMNH(E) 2003-84 (
<collectionCode id="ED6EAE67FF866A0BFC48FBE8FC69FB0C" box="[898,991,1131,1155]" pageId="17" pageNumber="16">BMNH</collectionCode>
).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C3656529FF866A0BFF0AFB1BFCCFFB5F" pageId="17" pageNumber="16" type="discussion">
<paragraph id="8BC036A2FF866A0BFF0AFB1BFCCFFB5F" blockId="17.[192,1440,1176,1232]" pageId="17" pageNumber="16">
<emphasis id="B90BEAB0FF866A0BFF0AFB1BFEF5FB3F" bold="true" box="[192,323,1176,1200]" pageId="17" pageNumber="16">Remarks.</emphasis>
This species has all black elytra but is distinguished from other such species by an entirely yellow pronotum. The male genitalia are also distinctive.
</paragraph>
</subSubSection>
</treatment>
</document>
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