357 lines
36 KiB
XML
357 lines
36 KiB
XML
<document id="E7F7FA3B2DF0687C62D98C8FC125B66E" ID-DOI="10.1206/3953.1" ID-ISSN="0003-0082" ID-Zenodo-Dep="5370347" approvalRequired="6" approvalRequired_for_textStreams="6" checkinTime="1630268857110" checkinUser="carolina" docAuthor="Long, Khuat Dang, Van Achterberg, Cornelis, Carpenter, James M. & Oanh, Nguyen Thi" docDate="2020" docId="03D3F52DF62AFFC1BDC9FF4FFDC7FCFD" docLanguage="en" docName="N3953.pdf" docOrigin="American Museum Novitates 2020 (3953)" docSource="https://bioone.org/journals/american-museum-novitates/volume-2020/issue-3953/3953.1/Review-of-the-Genus-Wroughtonia-Cameron-1899-Hymenoptera-Braconidae-Helconinae/10.1206/3953.1.full" docStyle="DocumentStyle:26C3580A17D4BBE58BD8D8AC6AE09AC4.4:AmMusNovit.2010-.journal_article" docStyleId="26C3580A17D4BBE58BD8D8AC6AE09AC4" docStyleName="AmMusNovit.2010-.journal_article" docStyleVersion="4" docTitle="Wroughtonia aspera Long & Van Achterberg & Carpenter & Oanh 2020, sp. nov." docType="treatment" docVersion="4" lastPageNumber="11" masterDocId="FFEA8D55F622FFCBBC2CFFACFFC7FE19" masterDocTitle="Review of the genus Wroughtonia Cameron, 1899 (Hymenoptera, Braconidae, Helconinae), with the description of 12 new species from Vietnam" masterLastPageNumber="56" masterPageNumber="1" pageNumber="9" updateTime="1698468785609" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title id="D79B47E11AD04E644D54ED980D6F6E9F">Review of the genus Wroughtonia Cameron, 1899 (Hymenoptera, Braconidae, Helconinae), with the description of 12 new species from Vietnam</mods:title>
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<mods:namePart id="E56A19EA03F28FF95A90EB914109E7EB">Long, Khuat Dang</mods:namePart>
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<mods:namePart id="D6DFF94418E62E0800AE3154930ABFF4">Van Achterberg, Cornelis</mods:namePart>
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<mods:namePart id="A94CF9ABA9F4A9252395FA8A784E1407">Carpenter, James M.</mods:namePart>
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<mods:namePart id="3F1E5DC4956E6896DBC191793E91A925">Oanh, Nguyen Thi</mods:namePart>
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<mods:title id="8D11C168C533F6AC049593F0A00B5403">American Museum Novitates</mods:title>
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<mods:part id="D78256ACD1331EBB54AE1D22534BCD1E">
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<mods:date id="FABC9435BE530BC2020313EE92EB1D64">2020</mods:date>
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<mods:number id="486030EF9532721E5BAAF52ED01AE5FF">2020-06-16</mods:number>
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<mods:number id="6A1660879ACC3230EF23EFE4437C824F">2020</mods:number>
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<mods:detail id="CABE6F5D6DE378C37CC53D4494EEF20A" type="issue">
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<mods:number id="95F6AD79A2ECB461B1243CCE80DC1F1D">3953</mods:number>
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<mods:url id="907B12EC69EEA766D6DCA6A2B86FB5E7">https://bioone.org/journals/american-museum-novitates/volume-2020/issue-3953/3953.1/Review-of-the-Genus-Wroughtonia-Cameron-1899-Hymenoptera-Braconidae-Helconinae/10.1206/3953.1.full</mods:url>
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<mods:classification id="D063C70035BD9726FE218D58FF118F1C">journal article</mods:classification>
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<mods:identifier id="C5D94156129FF29BD804864B206C0E47" type="DOI">10.1206/3953.1</mods:identifier>
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<mods:identifier id="CC50AEC1F17472128D77A7EA9795FEDE" type="ISSN">0003-0082</mods:identifier>
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<treatment id="03D3F52DF62AFFC1BDC9FF4FFDC7FCFD" ID-DOI="http://doi.org/10.5281/zenodo.5461923" ID-Zenodo-Dep="5461923" LSID="urn:lsid:plazi:treatment:03D3F52DF62AFFC1BDC9FF4FFDC7FCFD" httpUri="http://treatment.plazi.org/id/03D3F52DF62AFFC1BDC9FF4FFDC7FCFD" lastPageId="10" lastPageNumber="11" pageId="8" pageNumber="9">
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<subSubSection id="C36017B0F62AFFC3BDC9FF4FFC84FEE6" box="[485,835,227,255]" pageId="8" pageNumber="9" type="nomenclature">
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<paragraph id="8BC5443BF62AFFC3BDC9FF4FFC84FEE6" blockId="8.[485,835,227,255]" box="[485,835,227,255]" pageId="8" pageNumber="9">
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<heading id="D08DF357F62AFFC3BDC9FF4FFC84FEE6" box="[485,835,227,255]" centered="true" fontSize="11" level="2" pageId="8" pageNumber="9" reason="2">
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<emphasis id="B90E9829F62AFFC3BDC9FF4FFD25FEE7" bold="true" box="[485,738,227,254]" pageId="8" pageNumber="9">
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<taxonomicName id="4C7A3FB8F62AFFC3BDC9FF4FFD1CFEE7" authority="Long & Van Achterberg & Carpenter & Oanh, 2020" authorityName="Long & Van Achterberg & Carpenter & Oanh" authorityYear="2020" box="[485,731,227,254]" class="Insecta" family="Braconidae" genus="Wroughtonia" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="aspera" status="sp. nov.">
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<emphasis id="B90E9829F62AFFC3BDC9FF4FFD1CFEE7" bold="true" box="[485,731,227,254]" italics="true" pageId="8" pageNumber="9">Wroughtonia aspera</emphasis>
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</taxonomicName>
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,
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</emphasis>
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<taxonomicNameLabel id="A23D2552F62AFFC3BEC6FF49FC84FEE6" box="[746,835,229,255]" pageId="8" pageNumber="9" rank="species">sp. nov.</taxonomicNameLabel>
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</heading>
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</paragraph>
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</subSubSection>
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||
<subSubSection id="C36017B0F62AFFC3BE6DFEB0FD20FF2F" box="[577,743,284,310]" pageId="8" pageNumber="9" type="description">
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<paragraph id="8BC5443BF62AFFC3BE6DFEB0FD20FF2F" blockId="8.[577,743,284,310]" box="[577,743,284,310]" pageId="8" pageNumber="9">
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||
<heading id="D08DF357F62AFFC3BE6DFEB0FD20FF2F" box="[577,743,284,310]" centered="true" fontSize="36" level="3" pageId="8" pageNumber="9" reason="7">
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<figureCitation id="134158BEF62AFFC3BE6DFEB0FD20FF2F" box="[577,743,284,310]" captionStart-0="FIGURE 14" captionStart-1="FIGURES 15–26" captionStartId-0="9.[108,194,792,813]" captionStartId-1="11.[108,206,1526,1547]" captionTargetBox-0="[207,1114,227,765]" captionTargetBox-1="[184,1138,225,1499]" captionTargetId-0="figure-338@9.[207,1114,227,765]" captionTargetId-1="figure-50@11.[181,1141,223,1503]" captionTargetPageId-0="9" captionTargetPageId-1="11" captionText-0="FIGURE 14. Wroughtonia aspera, sp. nov., habitus (holotype, female, lateral)." captionText-1="FIGURES 15–26. Wroughtonia aspera (holotype, female, lateral). 15. Head, dorsal. 16. Head, frontal 17. Head, lateral. 18. Mesoscutum. 19. Mesopleuron. 20. Forewing. 21. Propodeum. 22. Hind wing. 23. Metasoma. 24. Hind coxa, lateral. 25. Hind femur, lateral. 26. Hind tibia, lateral." figureDoi-0="http://doi.org/10.5281/zenodo.5370353" figureDoi-1="http://doi.org/10.5281/zenodo.5370355" httpUri-0="https://zenodo.org/record/5370353/files/figure.png" httpUri-1="https://zenodo.org/record/5370355/files/figure.png" pageId="8" pageNumber="9">Figures 14–26</figureCitation>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C36017B0F62AFFC3BCB0FEF9FBC8FF8D" pageId="8" pageNumber="9" type="materials_examined">
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<paragraph id="8BC5443BF62AFFC3BCB0FEF9FBC8FF8D" blockId="8.[108,1213,339,1748]" pageId="8" pageNumber="9">
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<materialsCitation id="3B124E66F62AFFC3BCB0FEF9FBC8FF8D" collectingDate="1998-04-20" collectingDateMax="1998-04-28" collectingDateMin="1998-04-20" collectionCode="IEBR" collectorName="Malaise & K. Long." country="Vietnam" elevation="900" latitude="18.366667" location="Huong Son" longLatPrecision="1275" longitude="106.21667" pageId="8" pageNumber="9" specimenCount="1" specimenCount-female="1" stateProvince="Ha Tinh" typeStatus="holotype">
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TYPE MATERIAL:
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<typeStatus id="54C1FA99F62AFFC3BD5FFEF8FE23FF77" box="[371,484,340,366]" pageId="8" pageNumber="9">Holotype</typeStatus>
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,
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<specimenCount id="9D7C8FB2F62AFFC3BDD8FEFFFDCBFF77" box="[500,524,339,366]" pageId="8" pageNumber="9" type="female">♀</specimenCount>
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, “Hel.049” (
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<collectionCode id="ED6BDCFEF62AFFC3BE88FEF8FD2DFF77" box="[676,746,340,366]" pageId="8" pageNumber="9">IEBR</collectionCode>
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), NC
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<collectingCountry id="F36D04ABF62AFFC3BF1BFEF8FC66FF77" box="[823,929,340,366]" name="Vietnam" pageId="8" pageNumber="9">Vietnam</collectingCountry>
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:
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<collectingRegion id="49BE8AD9F62AFFC3BF9CFEF9FBDDFF77" box="[944,1050,340,367]" country="Vietnam" name="Ha Tinh" pageId="8" pageNumber="9">Ha Tinh</collectingRegion>
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,
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<location id="8EA512E0F62AFFC3B806FEF9FB7FFF77" LSID="urn:lsid:plazi:treatment:03D3F52DF62AFFC1BDC9FF4FFDC7FCFD:8EA512E0F62AFFC3B806FEF9FB7FFF77" box="[1066,1208,340,367]" country="Vietnam" latitude="18.366667" longLatPrecision="1275" longitude="106.21667" name="Huong Son" pageId="8" pageNumber="9" stateProvince="Ha Tinh">Huong Son</location>
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,
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<geoCoordinate id="EE4E22FCF62AFFC3BC40FED5FF08FF8D" box="[108,207,377,404]" degrees="18" direction="north" minutes="22" orientation="latitude" pageId="8" pageNumber="9" precision="925" value="18.366667">
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18
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<emphasis id="B90E9829F62AFFC3BCA4FED5FF68FF8A" bold="true" box="[136,175,377,403]" pageId="8" pageNumber="9">°22</emphasis>
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′N
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</geoCoordinate>
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<geoCoordinate id="EE4E22FCF62AFFC3BCFBFED5FE83FF8D" box="[215,324,377,404]" degrees="106" direction="east" minutes="13" orientation="longitude" pageId="8" pageNumber="9" precision="925" value="106.21667">106°13′E</geoCoordinate>
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,
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<quantity id="4C82E9DEF62AFFC3BD7CFED5FE5AFF8D" box="[336,413,377,404]" metricMagnitude="2" metricUnit="m" metricValue="9.0" pageId="8" pageNumber="9" unit="m" value="900.0">
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<elevation id="0057A308F62AFFC3BD7CFED5FE5AFF8D" box="[336,413,377,404]" metricMagnitude="2" metricUnit="m" metricValue="9.0" pageId="8" pageNumber="9" unit="m" value="900.0">900 m</elevation>
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</quantity>
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,
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<date id="FFC462FBF62AFFC3BD85FED5FDB8FF8A" box="[425,639,377,403]" pageId="8" pageNumber="9" value="1998-04-20" valueMax="1998-04-28" valueMin="1998-04-20">
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<collectingDate id="EF809B13F62AFFC3BD85FED5FDB8FF8A" box="[425,639,377,403]" pageId="8" pageNumber="9" value="1998-04-20" valueMax="1998-04-28" valueMin="1998-04-20">April 20–28, 1998</collectingDate>
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</date>
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,
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<collectorName id="268F21EDF62AFFC3BEA7FED5FD22FF8A" box="[651,741,377,403]" pageId="8" pageNumber="9">Malaise</collectorName>
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[trap], AMNH,
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<collectorName id="268F21EDF62AFFC3BF85FED6FBC8FF8D" box="[937,1039,378,404]" pageId="8" pageNumber="9">K. Long.</collectorName>
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</materialsCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="C36017B0F62AFFC3BCB0FE33FCCFFFA0" box="[156,776,415,441]" pageId="8" pageNumber="9" type="distribution">
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<paragraph id="8BC5443BF62AFFC3BCB0FE33FCCFFFA0" blockId="8.[108,1213,339,1748]" box="[156,776,415,441]" pageId="8" pageNumber="9">
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DISTRIBUTION: NC
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<collectingCountry id="F36D04ABF62AFFC3BDABFE33FE37FFA0" box="[391,496,415,441]" name="Vietnam" pageId="8" pageNumber="9">Vietnam</collectingCountry>
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:
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<collectingRegion id="49BE8AD9F62AFFC3BDD7FE33FD98FFA0" box="[507,607,415,441]" country="Vietnam" name="Ha Tinh" pageId="8" pageNumber="9">Ha Tinh</collectingRegion>
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(Huong Son).
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</paragraph>
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</subSubSection>
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<subSubSection id="C36017B0F62AFFC3BCB0FE68FE55FFC7" box="[156,402,452,479]" pageId="8" pageNumber="9" type="biology_ecology">
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<paragraph id="8BC5443BF62AFFC3BCB0FE68FE55FFC7" blockId="8.[108,1213,339,1748]" box="[156,402,452,479]" pageId="8" pageNumber="9">BIOLOGY: Unknown.</paragraph>
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</subSubSection>
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<subSubSection id="C36017B0F62AFFC3BCB0FE45FE85FBD6" pageId="8" pageNumber="9" type="diagnosis">
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<paragraph id="8BC5443BF62AFFC3BCB0FE45FC2EFA17" blockId="8.[108,1213,339,1748]" pageId="8" pageNumber="9">
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DIAGNOSIS: Antenna with 38 segments, with antennal segments 11–20 cream white; maxillary palp 1.6
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<emphasis id="B90E9829F62AFFC3BCD3FDA3FED7FC30" bold="true" box="[255,272,527,553]" pageId="8" pageNumber="9">×</emphasis>
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as long as head; frontal protuberance long, acute and raised higher than lateral carinae of frons; in dorsal view width of head 1.6
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<emphasis id="B90E9829F62AFFC3BEE2FD98FD18FC57" bold="true" box="[718,735,564,590]" pageId="8" pageNumber="9">×</emphasis>
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its median length; height of eye 1.55
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<emphasis id="B90E9829F62AFFC3B887FD98FB7BFC57" bold="true" box="[1195,1212,564,590]" pageId="8" pageNumber="9">×</emphasis>
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temple. Length of mesosoma 1.9
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<emphasis id="B90E9829F62AFFC3BDDFFDF5FDC3FC6A" bold="true" box="[499,516,601,627]" pageId="8" pageNumber="9">×</emphasis>
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as long as high; notauli wide and shallow, largely rugose, fused posteriorly with longitudinal rugosities; scutellum rather short, rugose; propodeum coarsely rugose. Foretarsus 1.2
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<emphasis id="B90E9829F62AFFC3BDFEFD08FE24FCA7" bold="true" box="[466,483,676,702]" pageId="8" pageNumber="9">×</emphasis>
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longer than foretibia; forewing 3.1
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<emphasis id="B90E9829F62AFFC3BF56FD08FC4CFCA7" bold="true" box="[890,907,676,702]" pageId="8" pageNumber="9">×</emphasis>
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longer than its maximum width; vein 3-SR short, 0.6
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<emphasis id="B90E9829F62AFFC3BD89FD66FE70FCFD" bold="true" box="[421,439,714,740]" italics="true" pageId="8" pageNumber="9">×</emphasis>
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vein r; second submarginal cell trapezium-shaped, vein 2-M 3.7
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<emphasis id="B90E9829F62AFFC3B887FD65FB7BFCFA" bold="true" box="[1195,1212,713,739]" pageId="8" pageNumber="9">×</emphasis>
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3-SR; vein 1-M of hind wing 0.55
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<emphasis id="B90E9829F62AFFC3BE22FD43FDE7FD10" bold="true" box="[526,544,751,777]" italics="true" pageId="8" pageNumber="9">×</emphasis>
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vein 1r-m; vein 2-SC+R transverse; hind wing with 4 hamuli. Hind femur robust, with a rather wide obtuse protuberance, without distinct ventral serrations; length of hind femur (without protuberance) 3.1
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<emphasis id="B90E9829F62AFFC3BF1EFC96FC83FD4D" bold="true" box="[818,836,826,852]" italics="true" pageId="8" pageNumber="9">×</emphasis>
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its maximum width; outer side of hind coxa sparsely finely punctate; hind femur densely punctate laterally. First tergite 1.1
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<emphasis id="B90E9829F62AFFC3B887FCF3FB7BFD60" bold="true" box="[1195,1212,863,889]" pageId="8" pageNumber="9">×</emphasis>
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longer than its apical width; dorsal carinae in basal 0.7 of tergite; median length of second tergite 0.6
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<emphasis id="B90E9829F62AFFC3BCF3FC05FF37FDDA" bold="true" box="[223,240,937,963]" pageId="8" pageNumber="9">×</emphasis>
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as long as its basal width, and 1.15
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<emphasis id="B90E9829F62AFFC3BEAEFC05FD54FDDA" bold="true" box="[642,659,937,963]" pageId="8" pageNumber="9">×</emphasis>
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third tergite; first tergite largely smooth basally, transversely rugose mediolaterally and nearly areolate-rugulose medioapically; second tergite punctate-reticulate, but smooth medially and remainder shiny and smooth.
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</paragraph>
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<paragraph id="8BC5443BF62AFFC3BCB1FBB5FE9DFAF6" blockId="8.[108,1213,339,1748]" pageId="8" pageNumber="9">
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NOTES:
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<emphasis id="B90E9829F62AFFC3BCD5FBB6FEB7FA2A" box="[249,368,1050,1075]" italics="true" pageId="8" pageNumber="9">
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<taxonomicName id="4C7A3FB8F62AFFC3BCD5FBB6FEABFA2A" authority="Long & Van Achterberg & Carpenter & Oanh, 2020" authorityName="Long & Van Achterberg & Carpenter & Oanh" authorityYear="2020" box="[249,364,1050,1075]" class="Insecta" family="Braconidae" genus="Wroughtonia" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="aspera" status="sp. nov.">W. aspera</taxonomicName>
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,
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</emphasis>
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<taxonomicNameLabel id="A23D2552F62AFFC3BD5BFBB6FE16FA2D" box="[375,465,1050,1076]" pageId="8" pageNumber="9" rank="species">sp. nov.</taxonomicNameLabel>
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, runs in the key by
|
||
<bibRefCitation id="EFEB39CAF62AFFC3BE9AFBB6FCB3FA2A" author="Yan, C. - J. & C. van Achterberg & J. - H. He & X. - X. Chen" box="[694,884,1049,1076]" pageId="8" pageNumber="9" pagination="401 - 457" refId="ref27775" refString="Yan, C. - J., C. van Achterberg, J. - H. He, and X. - X. Chen. 2017. Review of the tribe Helconini Foerster s. s. from China, with the description of 18 new species. Zootaxa 4291 (3): 401 - 457." type="journal article" year="2017">Yan et al. (2017)</bibRefCitation>
|
||
to
|
||
<taxonomicName id="4C7A3FB8F62AFFC3BFB6FBB6FEA9FA40" authority="Yan and van Achterberg, 2017" authorityName="Yan and van Achterberg" authorityYear="2017" class="Insecta" family="Braconidae" genus="Wroughtonia" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="zhejiangensis">
|
||
<emphasis id="B90E9829F62AFFC3BFB6FBB6FB91FA2A" box="[922,1110,1049,1075]" italics="true" pageId="8" pageNumber="9">W. zhejiangensis</emphasis>
|
||
<bibRefCitation id="EFEB39CAF62AFFC3B871FBB6FEA9FA40" author="Yan, C. - J. & C. van Achterberg & J. - H. He & X. - X. Chen" pageId="8" pageNumber="9" pagination="401 - 457" refId="ref27775" refString="Yan, C. - J., C. van Achterberg, J. - H. He, and X. - X. Chen. 2017. Review of the tribe Helconini Foerster s. s. from China, with the description of 18 new species. Zootaxa 4291 (3): 401 - 457." type="journal article" year="2017">Yan and van Achterberg, 2017</bibRefCitation>
|
||
</taxonomicName>
|
||
, from
|
||
<collectingCountry id="F36D04ABF62AFFC3BD93FB93FDC0FA40" box="[447,519,1087,1113]" name="China" pageId="8" pageNumber="9">China</collectingCountry>
|
||
and can be separated as follows: (1) frontal protuberance near antennal sockets evenly narrowed from base to apex, roundly pointed apically (frontal protuberance with posterior protruding narrow lamella in
|
||
<taxonomicName id="4C7A3FB8F62AFFC3BF31FB26FC1DFABA" authorityName="Yan and van Achterberg" authorityYear="2017" box="[797,986,1161,1187]" class="Insecta" family="Braconidae" genus="Wroughtonia" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="zhejiangensis">
|
||
<emphasis id="B90E9829F62AFFC3BF31FB26FC1DFABA" box="[797,986,1161,1187]" italics="true" pageId="8" pageNumber="9">W. zhejiangensis</emphasis>
|
||
</taxonomicName>
|
||
); (2) forewing vein r 1.7
|
||
<emphasis id="B90E9829F62AFFC3BC8DFB03FF75FAD0" bold="true" box="[161,178,1199,1225]" pageId="8" pageNumber="9">×</emphasis>
|
||
vein 3-SR (1.1
|
||
<emphasis id="B90E9829F62AFFC3BD73FB03FEB7FAD0" bold="true" box="[351,368,1199,1225]" pageId="8" pageNumber="9">×</emphasis>
|
||
in
|
||
<taxonomicName id="4C7A3FB8F62AFFC3BDBBFB03FD94FAD0" authorityName="Yan and van Achterberg" authorityYear="2017" box="[407,595,1199,1225]" class="Insecta" family="Braconidae" genus="Wroughtonia" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="zhejiangensis">
|
||
<emphasis id="B90E9829F62AFFC3BDBBFB03FD94FAD0" box="[407,595,1199,1225]" italics="true" pageId="8" pageNumber="9">W. zhejiangensis</emphasis>
|
||
</taxonomicName>
|
||
); and (3) hind wing vein 1-M 0.55
|
||
<emphasis id="B90E9829F62AFFC3BFC7FB03FC3BFAD0" bold="true" box="[1003,1020,1199,1225]" pageId="8" pageNumber="9">×</emphasis>
|
||
vein 1r-m (0.4
|
||
<emphasis id="B90E9829F62AFFC3B887FB03FB7BFAD0" bold="true" box="[1195,1212,1199,1225]" pageId="8" pageNumber="9">×</emphasis>
|
||
in
|
||
<taxonomicName id="4C7A3FB8F62AFFC3BCA0FB79FE8EFAF7" authorityName="Yan and van Achterberg" authorityYear="2017" box="[140,329,1236,1262]" class="Insecta" family="Braconidae" genus="Wroughtonia" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="zhejiangensis">
|
||
<emphasis id="B90E9829F62AFFC3BCA0FB79FE8EFAF7" box="[140,329,1236,1262]" italics="true" pageId="8" pageNumber="9">W. zhejiangensis</emphasis>
|
||
</taxonomicName>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BC5443BF62AFFC3BCB0FB55FE85FBD6" blockId="8.[108,1213,339,1748]" pageId="8" pageNumber="9">
|
||
The new species is also similar to
|
||
<taxonomicName id="4C7A3FB8F62AFFC3BE11FB56FBB4FB0A" authority="van Achterberg and Chen, 2017" authorityName="van Achterberg and Chen" authorityYear="2017" box="[573,1139,1273,1300]" class="Insecta" family="Braconidae" genus="Wroughtonia" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="albobasalis">
|
||
<emphasis id="B90E9829F62AFFC3BE11FB56FD21FB0A" box="[573,742,1273,1299]" italics="true" pageId="8" pageNumber="9">W. albobasalis</emphasis>
|
||
van Achterberg and Chen, 2017
|
||
</taxonomicName>
|
||
, from
|
||
<collectingCountry id="F36D04ABF62AFFC3BC40FAB3FF71FB20" box="[108,182,1311,1337]" name="China" pageId="8" pageNumber="9">China</collectingCountry>
|
||
, but differs from the latter by having: (1) hind femur slender, length 3.1
|
||
<emphasis id="B90E9829F62AFFC3B82AFAB3FBD0FB20" bold="true" box="[1030,1047,1311,1337]" pageId="8" pageNumber="9">×</emphasis>
|
||
its maximum width (2.8
|
||
<emphasis id="B90E9829F62AFFC3BCC9FAE8FF31FB47" bold="true" box="[229,246,1348,1374]" pageId="8" pageNumber="9">×</emphasis>
|
||
in
|
||
<taxonomicName id="4C7A3FB8F62AFFC3BD0DFAE9FE02FB47" authorityName="van Achterberg and Chen" authorityYear="2017" box="[289,453,1348,1374]" class="Insecta" family="Braconidae" genus="Wroughtonia" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="albobasalis">
|
||
<emphasis id="B90E9829F62AFFC3BD0DFAE9FE02FB47" box="[289,453,1348,1374]" italics="true" pageId="8" pageNumber="9">W. albobasalis</emphasis>
|
||
</taxonomicName>
|
||
); (2) vein r of forewing 1.7
|
||
<emphasis id="B90E9829F62AFFC3BF26FAE8FCDCFB47" bold="true" box="[778,795,1348,1374]" pageId="8" pageNumber="9">×</emphasis>
|
||
vein 3-SR (1.1
|
||
<emphasis id="B90E9829F62AFFC3BFE7FAE8FC1BFB47" bold="true" box="[971,988,1348,1374]" pageId="8" pageNumber="9">×</emphasis>
|
||
in
|
||
<taxonomicName id="4C7A3FB8F62AFFC3B82BFAE9FB6CFB47" authorityName="van Achterberg and Chen" authorityYear="2017" box="[1031,1195,1348,1374]" class="Insecta" family="Braconidae" genus="Wroughtonia" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="albobasalis">
|
||
<emphasis id="B90E9829F62AFFC3B82BFAE9FB6CFB47" box="[1031,1195,1348,1374]" italics="true" pageId="8" pageNumber="9">W. albobasalis</emphasis>
|
||
</taxonomicName>
|
||
); (3) vein 1-M of hind wing 0.55
|
||
<emphasis id="B90E9829F62AFFC3BDCEFAC5FE34FB9A" bold="true" box="[482,499,1385,1411]" pageId="8" pageNumber="9">×</emphasis>
|
||
vein 1r-m (0.4
|
||
<emphasis id="B90E9829F62AFFC3BE8BFAC5FD7FFB9A" bold="true" box="[679,696,1385,1411]" pageId="8" pageNumber="9">×</emphasis>
|
||
in
|
||
<taxonomicName id="4C7A3FB8F62AFFC3BEC9FAC6FC4EFB9A" authorityName="van Achterberg and Chen" authorityYear="2017" box="[741,905,1385,1411]" class="Insecta" family="Braconidae" genus="Wroughtonia" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="albobasalis">
|
||
<emphasis id="B90E9829F62AFFC3BEC9FAC6FC4EFB9A" box="[741,905,1385,1411]" italics="true" pageId="8" pageNumber="9">W. albobasalis</emphasis>
|
||
</taxonomicName>
|
||
); and (4) occiput smooth (sparsely punctate in
|
||
<taxonomicName id="4C7A3FB8F62AFFC3BD40FA23FDD7FBB0" authorityName="van Achterberg and Chen" authorityYear="2017" box="[364,528,1423,1449]" class="Insecta" family="Braconidae" genus="Wroughtonia" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="albobasalis">
|
||
<emphasis id="B90E9829F62AFFC3BD40FA23FDD7FBB0" box="[364,528,1423,1449]" italics="true" pageId="8" pageNumber="9">W. albobasalis</emphasis>
|
||
</taxonomicName>
|
||
), 11th–20th antennal segments 11–20 cream white (
|
||
<quantity id="4C82E9DEF62AFFC3B8ADFA23FF41FBD7" metricMagnitude="-1" metricUnit="m" metricValue="1.778" metricValueMax="2.794" metricValueMin="0.762" pageId="8" pageNumber="9" unit="in" value="7.0" valueMax="11.0" valueMin="3.0">3–11 in</quantity>
|
||
<taxonomicName id="4C7A3FB8F62AFFC3BCA0FA19FEF6FBD7" authorityName="van Achterberg and Chen" authorityYear="2017" box="[140,305,1460,1486]" class="Insecta" family="Braconidae" genus="Wroughtonia" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="albobasalis">
|
||
<emphasis id="B90E9829F62AFFC3BCA0FA19FEF6FBD7" box="[140,305,1460,1486]" italics="true" pageId="8" pageNumber="9">W. albobasalis</emphasis>
|
||
</taxonomicName>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C36017B0F62AFFC1BCB0FA75FEA5FC80" lastPageId="10" lastPageNumber="11" pageId="8" pageNumber="9" type="description">
|
||
<paragraph id="8BC5443BF62AFFC3BCB0FA75FE41F800" blockId="8.[108,1213,339,1748]" pageId="8" pageNumber="9">
|
||
DESCRIPTION:
|
||
<typeStatus id="54C1FA99F62AFFC3BD62FA75FE79FBEA" box="[334,446,1497,1523]" pageId="8" pageNumber="9">Holotype</typeStatus>
|
||
,
|
||
<specimenCount id="9D7C8FB2F62AFFC3BDE1FA75FE22FBED" box="[461,485,1497,1524]" pageId="8" pageNumber="9" type="female">♀</specimenCount>
|
||
, body length
|
||
<quantity id="4C82E9DEF62AFFC3BEBDFA75FD38FBED" box="[657,767,1497,1524]" metricMagnitude="-2" metricUnit="m" metricValue="1.14" pageId="8" pageNumber="9" unit="mm" value="11.4">11.4 mm</quantity>
|
||
, forewing length
|
||
<quantity id="4C82E9DEF62AFFC3BFFAFA75FBF2FBED" box="[982,1077,1497,1524]" metricMagnitude="-3" metricUnit="m" metricValue="7.5" pageId="8" pageNumber="9" unit="mm" value="7.5">7.5 mm</quantity>
|
||
, ovipositor sheath
|
||
<quantity id="4C82E9DEF62AFFC3BCECFA53FEDCF800" box="[192,283,1535,1561]" metricMagnitude="-3" metricUnit="m" metricValue="7.7" pageId="8" pageNumber="9" unit="mm" value="7.7">7.7 mm</quantity>
|
||
(fig. 14).
|
||
</paragraph>
|
||
<paragraph id="8BC5443BF62AFFC2BCB0F988FBFAFDE7" blockId="8.[108,1213,339,1748]" lastBlockId="9.[108,1215,846,1732]" lastPageId="9" lastPageNumber="10" pageId="8" pageNumber="9">
|
||
<emphasis id="B90E9829F62AFFC3BCB0F988FF27F827" box="[156,224,1572,1598]" italics="true" pageId="8" pageNumber="9">Head:</emphasis>
|
||
Antenna with 38 segments; third antennal segment 1.2
|
||
<emphasis id="B90E9829F62AFFC3BF56F988FC4CF827" bold="true" box="[890,907,1572,1598]" pageId="8" pageNumber="9">×</emphasis>
|
||
fourth (19:17); in frontal view height of eye 2.1
|
||
<emphasis id="B90E9829F62AFFC3BD5AF9E5FE40F87A" bold="true" box="[374,391,1609,1635]" pageId="8" pageNumber="9">×</emphasis>
|
||
its transverse width (27:13); width of face 1.85
|
||
<emphasis id="B90E9829F62AFFC3BFEAF9E5FC10F87A" bold="true" box="[966,983,1609,1635]" pageId="8" pageNumber="9">×</emphasis>
|
||
length of face and clypeus combined (39:21); malar space 1.2
|
||
<emphasis id="B90E9829F62AFFC3BE77F9C3FDABF890" bold="true" box="[603,620,1647,1673]" pageId="8" pageNumber="9">×</emphasis>
|
||
as long as mandible width (18:15) and 0.5
|
||
<emphasis id="B90E9829F62AFFC3B876F9C3FBACF890" bold="true" box="[1114,1131,1647,1673]" pageId="8" pageNumber="9">×</emphasis>
|
||
height of eye (18:39) (fig. 16); maxillary palp 1.6
|
||
<emphasis id="B90E9829F62AFFC3BE4CF938FDB6F8B7" bold="true" box="[608,625,1684,1710]" pageId="8" pageNumber="9">×</emphasis>
|
||
as long as head (50:32); in dorsal view, width of head 1.6
|
||
<emphasis id="B90E9829F62AFFC3BCF8F915FF22F8CA" bold="true" box="[212,229,1721,1747]" pageId="8" pageNumber="9">×</emphasis>
|
||
as long as median length (75:47); height of eye 1.5
|
||
<emphasis id="B90E9829F62AFFC3BF48F915FCB2F8CA" bold="true" box="[868,885,1721,1747]" pageId="8" pageNumber="9">×</emphasis>
|
||
as long as temple (30:20); POL:OD:OOL = 10:7:19 (fig. 15); distance between front and hind ocelli 0.3
|
||
<emphasis id="B90E9829F62BFFC2BFC1FCE3FC39FD70" bold="true" box="[1005,1022,847,873]" pageId="9" pageNumber="10">×</emphasis>
|
||
as long as OOL (6:19); in lateral view, length of eye 1.3
|
||
<emphasis id="B90E9829F62BFFC2BE6DFCD8FD95FD97" bold="true" box="[577,594,884,910]" pageId="9" pageNumber="10">×</emphasis>
|
||
as long as transverse width (38:30) (fig. 17); frontal protuberance long, acute and raised above lateral carinae of frons; face coarsely rugose; frons deeply depressed, smooth; vertex rugose-punctate anteriorly and smooth medioposteriorly; occiput largely smooth; temple nearly rugose-punctate, but coarsely rugose ventrally.
|
||
</paragraph>
|
||
<caption id="DF0514B3F62BFFC2BC40FCB4FCBDFD37" ID-DOI="http://doi.org/10.5281/zenodo.5370353" ID-Zenodo-Dep="5370353" box="[108,890,790,814]" httpUri="https://zenodo.org/record/5370353/files/figure.png" pageId="9" pageNumber="10" startId="9.[108,194,792,813]" targetBox="[207,1114,227,765]" targetPageId="9">
|
||
<paragraph id="8BC5443BF62BFFC2BC40FCB4FCBDFD37" blockId="9.[108,890,790,814]" box="[108,890,790,814]" pageId="9" pageNumber="10">
|
||
FIGURE 14.
|
||
<taxonomicName id="4C7A3FB8F62BFFC2BCC3FCBAFE7FFD34" authority="Long & Van Achterberg & Carpenter & Oanh, 2020" authorityName="Long & Van Achterberg & Carpenter & Oanh" authorityYear="2020" box="[239,440,790,814]" class="Insecta" family="Braconidae" genus="Wroughtonia" kingdom="Animalia" order="Hymenoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="aspera" status="sp. nov.">
|
||
<emphasis id="B90E9829F62BFFC2BCC3FCBAFE7FFD34" box="[239,440,790,814]" italics="true" pageId="9" pageNumber="10">Wroughtonia aspera</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="A23D2552F62BFFC2BDE9FCBAFDD3FD34" box="[453,532,790,813]" pageId="9" pageNumber="10" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, habitus (holotype, female, lateral).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BC5443BF62BFFC2BCB0FBA6FE41FAC7" blockId="9.[108,1215,846,1732]" pageId="9" pageNumber="10">
|
||
<taxonomicName id="4C7A3FB8F62BFFC2BCB0FBA6FE5AFA3D" authority=": Mesosoma" authorityName="Mesosoma" box="[156,413,1034,1060]" class="Arachnida" family="Sclerosomatidae" genus="Mesosoma" higherTaxonomySource="CoL" kingdom="Animalia" order="Opiliones" pageId="9" pageNumber="10" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B90E9829F62BFFC2BCB0FBA6FEDEFA3A" box="[156,281,1034,1059]" italics="true" pageId="9" pageNumber="10">Mesosoma:</emphasis>
|
||
Mesosoma
|
||
</taxonomicName>
|
||
1.9
|
||
<emphasis id="B90E9829F62BFFC2BDEBFBA5FE1FFA3A" bold="true" box="[455,472,1033,1059]" pageId="9" pageNumber="10">×</emphasis>
|
||
longer than high (100:52); pronotal side coarsely rugose; mesopleuron largely smooth medially, rugose-punctate ventrally (fig. 19); metapleuron coarsely rugose; median lobe of mesoscutum nearly finely punctate, but rugose lateroapically; notauli wide and shallow, largely rugose, fused posteriorly with longitudinal rugosities (fig. 18); scutellar sulcus with 4 carinae, 0.5
|
||
<emphasis id="B90E9829F62BFFC2BD67FB33FE9BFAA0" bold="true" box="[331,348,1183,1209]" pageId="9" pageNumber="10">×</emphasis>
|
||
as long as scutellum (7:13); scutellum rather short, rugose; propodeum coarsely rugose (fig. 21).
|
||
</paragraph>
|
||
<paragraph id="8BC5443BF62BFFC2BCB0FB46FF02FBA7" blockId="9.[108,1215,846,1732]" pageId="9" pageNumber="10">
|
||
<emphasis id="B90E9829F62BFFC2BCB0FB46FF2DFB1A" box="[156,234,1258,1283]" italics="true" pageId="9" pageNumber="10">Wings.</emphasis>
|
||
Forewing: length of forewing 3.1
|
||
<emphasis id="B90E9829F62BFFC2BE5BFB45FD4FFB1A" bold="true" box="[631,648,1257,1283]" pageId="9" pageNumber="10">×</emphasis>
|
||
its maximum width (140:45); pterostigma 2.9
|
||
<emphasis id="B90E9829F62BFFC2B887FB45FB7BFB1A" bold="true" box="[1195,1212,1257,1283]" pageId="9" pageNumber="10">×</emphasis>
|
||
longer than wide (38:13); vein 3-SR short, 0.6
|
||
<emphasis id="B90E9829F62BFFC2BEADFAA3FD55FB30" bold="true" box="[641,658,1295,1321]" pageId="9" pageNumber="10">×</emphasis>
|
||
vein r (7:12); r:2-SR:3-SR:SR1:r-m = 12:17:7:65:16; vein cu-a oblique, 1-CU1:cu-a:2-CU1 = 6:17:28; second submarginal cell trapezium-shaped, vein 2-M 3.7
|
||
<emphasis id="B90E9829F62BFFC2BD2FFAF5FED3FB6A" bold="true" box="[259,276,1369,1395]" pageId="9" pageNumber="10">×</emphasis>
|
||
3-SR (26:7) (fig. 20). Hind wing with vein m-cu as an antefurcal spectral trace; vein 1-M 0.55
|
||
<emphasis id="B90E9829F62BFFC2BD21FAD3FED9FB80" bold="true" box="[269,286,1407,1433]" pageId="9" pageNumber="10">×</emphasis>
|
||
as long as vein 1r-m (10:18); vein 2-SC+R transverse (fig. 22); hind wing with 4 hamuli.
|
||
</paragraph>
|
||
<paragraph id="8BC5443BF62BFFC2BCB0FA66FD92F8DA" blockId="9.[108,1215,846,1732]" pageId="9" pageNumber="10">
|
||
<emphasis id="B90E9829F62BFFC2BCB0FA66FF14FBFA" box="[156,211,1482,1507]" italics="true" pageId="9" pageNumber="10">Legs:</emphasis>
|
||
Foretarsus 1.2
|
||
<emphasis id="B90E9829F62BFFC2BD5AFA65FE40FBFA" bold="true" box="[374,391,1481,1507]" pageId="9" pageNumber="10">×</emphasis>
|
||
as long as foretibia (43:35); hind femur robust, with rather a wide obtuse protuberance, without distinct ventral serrations (fig. 25); length of hind femur (without protuberance), tibia, and basitarsus 3.1 (74:24), 9.2 (110:12), and 5.5
|
||
<emphasis id="B90E9829F62BFFC2BF1FF9B8FC83F837" bold="true" box="[819,836,1556,1582]" pageId="9" pageNumber="10">×</emphasis>
|
||
(33:6) as long as their maximum width, respectively; hind tibia evenly curved basally (fig. 26); hind basitarsus 0.3
|
||
<emphasis id="B90E9829F62BFFC2BFD7F995FBCBF84A" bold="true" box="[1019,1036,1593,1619]" pageId="9" pageNumber="10">×</emphasis>
|
||
as long as hind tibia (33:110); hind basitarsus 0.7
|
||
<emphasis id="B90E9829F62BFFC2BDCAF9F3FE30F860" bold="true" box="[486,503,1631,1657]" pageId="9" pageNumber="10">×</emphasis>
|
||
as long as hind tarsal segments 2–5 (33:49); fourth hind tarsal segment 0.4
|
||
<emphasis id="B90E9829F62BFFC2BCDBF928FECFF887" bold="true" box="[247,264,1668,1694]" pageId="9" pageNumber="10">×</emphasis>
|
||
as long as telotarsus (6:16). Outer side of hind coxa sparsely finely punctate (fig. 24); hind femur densely punctate laterally.
|
||
</paragraph>
|
||
<paragraph id="8BC5443BF628FFC1BCB0FF49FB7BFFA3" blockId="10.[108,1214,228,740]" pageId="10" pageNumber="11">
|
||
<emphasis id="B90E9829F628FFC1BCB0FF49FEDEFEE7" box="[156,281,229,254]" italics="true" pageId="10" pageNumber="11">Metasoma:</emphasis>
|
||
Metasoma about as long as head and mesosoma combined (65:66); first tergite 1.1
|
||
<emphasis id="B90E9829F628FFC1BCA2FEA6FF58FF3D" bold="true" box="[142,159,266,292]" pageId="10" pageNumber="11">×</emphasis>
|
||
longer than its apical width (36:33) (fig. 23); dorsal carinae in basal 0.7 of tergite (31:43); median length of second tergite 0.6
|
||
<emphasis id="B90E9829F628FFC1BE27FE83FDDBFF50" bold="true" box="[523,540,303,329]" pageId="10" pageNumber="11">×</emphasis>
|
||
as long as its basal width (31:52), and 1.15
|
||
<emphasis id="B90E9829F628FFC1B83BFE83FBEFFF50" bold="true" box="[1047,1064,303,329]" pageId="10" pageNumber="11">×</emphasis>
|
||
third tergite (31:27); first tergite almost smooth basally, transversely rugose mediolaterally, nearly areolaterugulose medioapically; second tergite as long as third (19:19); second tergite punctate-reticulate (fig. 23); remainder of metasoma shiny, smooth; ovipositor sheath slightly longer forewing (77:75).
|
||
</paragraph>
|
||
<paragraph id="8BC5443BF628FFC1BCB0FE68FBA4FC6D" blockId="10.[108,1214,228,740]" pageId="10" pageNumber="11">
|
||
<emphasis id="B90E9829F628FFC1BCB0FE68FF23FFC7" box="[156,228,452,478]" italics="true" pageId="10" pageNumber="11">Color:</emphasis>
|
||
Body black, scapus brown; antenna brown with antennal segments 11–20 cream white; palpi whitish yellow, but first and second segments yellowish brown; fore- and middle legs yellow, but tarsus whitish yellow; hind coxa yellow; hind femur yellow basally and brown apically (fig. 25); hind tibia whitish yellow, but infuscate apically (fig. 26); pterostigma brown, but yellow basally; wing veins brownish yellow; wing membrane hyaline (figs. 20, 22).
|
||
</paragraph>
|
||
<paragraph id="8BC5443BF628FFC1BCB0FDD3FEA5FC80" blockId="10.[108,1214,228,740]" box="[156,354,639,665]" pageId="10" pageNumber="11">
|
||
<emphasis id="B90E9829F628FFC1BCB0FDD3FF1AFC80" box="[156,221,639,665]" italics="true" pageId="10" pageNumber="11">Male:</emphasis>
|
||
Unknown.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C36017B0F628FFC1BCB0FD09FDC7FCFD" pageId="10" pageNumber="11" type="etymology">
|
||
<paragraph id="8BC5443BF628FFC1BCB0FD09FDC7FCFD" blockId="10.[108,1214,228,740]" pageId="10" pageNumber="11">
|
||
ETYMOLOGY: From
|
||
<emphasis id="B90E9829F628FFC1BDAFFD09FE07FCA7" box="[387,448,677,702]" italics="true" pageId="10" pageNumber="11">asper</emphasis>
|
||
(Latin for “rough,” “harsh”), because the flat medioposterior area of mesoscutum is coarsely rugose.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |