644 lines
80 KiB
XML
644 lines
80 KiB
XML
<document id="1760212F1F4319BC07A7F34B5817551F" ID-CLB-Dataset="3153" ID-DOI="10.5169/seals-787071" ID-GBIF-Dataset="a3876c73-615d-4cd3-88d7-779d2175638f" ID-ISSN="2624-9170" ID-Zenodo-Dep="5749772" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1637933697018" checkinUser="carolina" docAuthor="Casale, Achille" docDate="2011" docId="03D34424C84ABE3CC4935627FDE1CAA8" docLanguage="en" docName="ContribNatHist.1.1-16.pdf.imf" docOrigin="Contributions to Natural History 16" docStyle="DocumentStyle:D319E9290ED630111D2A11900DAFC11F.4:ContribNatHist.2011-.journal_article" docStyleId="D319E9290ED630111D2A11900DAFC11F" docStyleName="ContribNatHist.2011-.journal_article" docStyleVersion="4" docTitle="Trechus (Trechus) messoulii Casale 2011, sp. nov." docType="treatment" docVersion="6" lastPageNumber="14" masterDocId="FFEA3C5CC84DBE31C4235232FFCACE40" masterDocTitle="Two new subterranean, microphthalmous trechine beetles from the Mediterranean area, and a synonymic note (Coleoptera: Carabidae, Trechini)" masterLastPageNumber="16" masterPageNumber="1" pageNumber="8" updateTime="1699470421717" updateUser="ExternalLinkService">
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<mods:mods id="AC9DBF7010AFFE41123995FB5BAD6083" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="DE7C9399E7BD38EDD4253FD659650CF7">
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<mods:title id="36F9272DB239FF449AD6E4C0B386993E">Two new subterranean, microphthalmous trechine beetles from the Mediterranean area, and a synonymic note (Coleoptera: Carabidae, Trechini)</mods:title>
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</mods:titleInfo>
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<mods:name id="F9C692D282DE1ACBEF2F45F26D28608C" type="personal">
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<mods:role id="F5929F8110E9A77773734DC656F5AA56">
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<mods:roleTerm id="18B5331DD1A5458882F1F8D48260E67B">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="B299A5A7CBFF32798EDEE11DEA901A1A">Casale, Achille</mods:namePart>
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<mods:affiliation id="7147654B967EC7F27DF19D503664C085">Università di Sassari Dipartimento di Zoologia e Genetica Evoluzionistica Via Muroni 25 I – 07100 Sassari (Italy)</mods:affiliation>
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<mods:nameIdentifier id="D8F700FBE82212DBA6B8AB47B3478B3B" type="email">casale@uniss.it & a_casale@libero.it</mods:nameIdentifier>
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</mods:name>
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<mods:typeOfResource id="29894A59E50B901256B98A4E2CCA3AD6">text</mods:typeOfResource>
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<mods:titleInfo id="6A3CE35B59B2E84166FA36D5674C04FD">
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<mods:title id="FA574EFB00C5C0E22FEBAA4C76B7476E">Contributions to Natural History</mods:title>
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</mods:titleInfo>
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<mods:part id="ABA3DAC191F39FA97F0422ACA6B6D998">
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<mods:date id="DF95ED7A54E4C70AD098468F05A09940">2011</mods:date>
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<mods:detail id="0D7263F19E7AA34992A6668D1DD5A9B6" type="pubDate">
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<mods:number id="B6C6473BDBBAC6CF2E5C0C193A2225F1">2011-03-17</mods:number>
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<mods:detail id="CAE36E2A67FD0C5DA078F8EEFF69A0E8" type="volume">
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<mods:number id="3D9FE5FF418729CB26F1DAC74DDA7F94">16</mods:number>
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<mods:start id="76ECB0F0C73A52278CEA61DF3B1E809E">1</mods:start>
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<mods:end id="149F0EDB157245DE273F6C6AA583C4CB">16</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:classification id="F35DCEC12B96788E3550952D89DA559F">journal article</mods:classification>
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<mods:identifier id="53A50C02096238477A249E373B16CFC2" type="CLB-Dataset">3153</mods:identifier>
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<mods:identifier id="29FB889F669C2911EB29DC344EF2A12B" type="DOI">10.5169/seals-787071</mods:identifier>
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<mods:identifier id="90E654B05759F72672628983063A836E" type="GBIF-Dataset">a3876c73-615d-4cd3-88d7-779d2175638f</mods:identifier>
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<mods:identifier id="113C0AF25037040589DDA86B0727AEF9" type="ISSN">2624-9170</mods:identifier>
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<mods:identifier id="5D5111B9CFD53E5A53253F5E7243CBC1" type="Zenodo-Dep">5749772</mods:identifier>
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<treatment id="03D34424C84ABE3CC4935627FDE1CAA8" ID-DOI="http://doi.org/10.5281/zenodo.5749780" ID-GBIF-Taxon="190875635" ID-Zenodo-Dep="5749780" LSID="urn:lsid:plazi:treatment:03D34424C84ABE3CC4935627FDE1CAA8" httpUri="http://treatment.plazi.org/id/03D34424C84ABE3CC4935627FDE1CAA8" lastPageId="13" lastPageNumber="14" pageId="7" pageNumber="8">
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<subSubSection id="C360A6B9C84ABE36C4935627FD81CA6E" box="[176,587,1045,1070]" pageId="7" pageNumber="8" type="nomenclature">
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<paragraph id="8BC5F532C84ABE36C4935627FD81CA6E" blockId="7.[176,587,1045,1070]" box="[176,587,1045,1070]" pageId="7" pageNumber="8">
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<heading id="D08D425EC84ABE36C4935627FD81CA6E" box="[176,587,1045,1070]" centered="true" fontSize="36" level="2" pageId="7" pageNumber="8" reason="4">
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<emphasis id="B90E2920C84ABE36C4935627FD81CA6E" bold="true" box="[176,587,1045,1070]" pageId="7" pageNumber="8">
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<taxonomicName id="4C7A8EB1C84ABE36C4935627FE27CA6E" ID-CoL="57ZTX" authority="Casale, 2011" authorityName="Casale" authorityYear="2011" box="[176,493,1045,1070]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="messoulii" status="sp. nov." subGenus="Trechus">
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<emphasis id="B90E2920C84ABE36C4935627FEC2CA6E" bold="true" box="[176,264,1045,1070]" italics="true" pageId="7" pageNumber="8">Trechus</emphasis>
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(
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<emphasis id="B90E2920C84ABE36C5345627FEA5CA6E" bold="true" box="[279,367,1045,1070]" italics="true" pageId="7" pageNumber="8">Trechus</emphasis>
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)
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<emphasis id="B90E2920C84ABE36C55C5627FE27CA6E" bold="true" box="[383,493,1045,1070]" italics="true" pageId="7" pageNumber="8">messoulii</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A23D945BC84ABE36C5D05627FD81CA6E" box="[499,587,1045,1070]" pageId="7" pageNumber="8" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C360A6B9C84ABE36C4935652FD2DCA38" box="[176,743,1120,1144]" pageId="7" pageNumber="8" type="materials_examined">
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<paragraph id="8BC5F532C84ABE36C4935652FD2DCA38" blockId="7.[176,743,1120,1144]" box="[176,743,1120,1144]" pageId="7" pageNumber="8">
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Type locality:
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<collectingCountry id="F36DB5A2C84ABE36C56F5652FE78CA38" box="[332,434,1120,1144]" name="Morocco" pageId="7" pageNumber="8">Morocco</collectingCountry>
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, Rif (Taza): Toghobeit cave.
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</paragraph>
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</subSubSection>
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<subSubSection id="C360A6B9C84ABE36C4935698FE98CA82" box="[176,338,1194,1218]" pageId="7" pageNumber="8" type="multiple">
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<paragraph id="8BC5F532C84ABE36C4935698FE98CA82" blockId="7.[175,1066,1194,1293]" box="[176,338,1194,1218]" pageId="7" pageNumber="8">Type material:</paragraph>
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</subSubSection>
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<subSubSection id="C360A6B9C84ABE36C49156E2FDCBCB4D" pageId="7" pageNumber="8" type="materials_examined">
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<paragraph id="8BC5F532C84ABE36C49156E2FDCBCB4D" blockId="7.[175,1066,1194,1293]" pageId="7" pageNumber="8">
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<materialsCitation id="3B12FF6FC84ABE36C49156E2FDCBCB4D" ID-GBIF-Occurrence="3413090302" collectingDate="1999-07-25" collectingDateMax="1999-07-30" collectingDateMin="1999-07-25" collectionCode="CCa" collectorName="M. Messouli" country="Morocco" location="Toghobeit cave" pageId="7" pageNumber="8" specimenCount="1" specimenCount-male="1" stateProvince="Rif (Taza)" typeStatus="holotype">
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<typeStatus id="54C14B90C84ABE36C49156E2FED7CAA8" box="[178,285,1232,1256]" pageId="7" pageNumber="8" type="holotype">Holotype</typeStatus>
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<specimenCount id="9D7C3EBBC84ABE36C50556FFFEF0CAA8" box="[294,314,1229,1256]" pageId="7" pageNumber="8" type="male">♂</specimenCount>
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with the following data: "
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<collectingCountry id="F36DB5A2C84ABE36C65056E2FD0CCAA8" box="[627,710,1232,1256]" name="Morocco" pageId="7" pageNumber="8">Maroc</collectingCountry>
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,
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<collectingRegion id="49BE3BD0C84ABE36C6F056E2FD3CCAA8" box="[723,758,1232,1256]" pageId="7" pageNumber="8">Rif</collectingRegion>
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,
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<location id="8EA5A3E9C84ABE36C72056E2FC62CAA8" LSID="urn:lsid:plazi:treatment:03D34424C84ABE3CC4935627FDE1CAA8:8EA5A3E9C84ABE36C72056E2FC62CAA8" box="[771,936,1232,1256]" country="Morocco" name="Gr. Toghobeit" pageId="7" pageNumber="8" stateProvince="Rif">Gr. Toghobeit</location>
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,
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<date id="FFC4D3F2C84ABE36C79756E2FF23CB4D" pageId="7" pageNumber="8" value="1999-07-25" valueMax="1999-07-30" valueMin="1999-07-25">
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<collectingDate id="EF802A1AC84ABE36C79756E2FF23CB4D" pageId="7" pageNumber="8" value="1999-07-25" valueMax="1999-07-30" valueMin="1999-07-25">25–30.VII. 1999</collectingDate>
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</date>
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leg.
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<collectorName id="268F90E4C84ABE36C53D56C7FE67CB4D" box="[286,429,1269,1293]" pageId="7" pageNumber="8">M. Messouli</collectorName>
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" (
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<collectionCode id="ED6B6DF7C84ABE36C5E256C7FE3DCB4D" box="[449,503,1269,1293]" pageId="7" pageNumber="8">CCa</collectionCode>
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).
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</materialsCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="C360A6B9C84ABE36C4915757FE5BCB3E" box="[178,401,1381,1406]" pageId="7" pageNumber="8" type="multiple">
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<paragraph id="8BC5F532C84ABE36C4915757FE5BCB3E" blockId="7.[178,401,1381,1406]" box="[178,401,1381,1406]" pageId="7" pageNumber="8">
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<emphasis id="B90E2920C84ABE36C4915757FE5BCB3E" bold="true" box="[178,401,1381,1406]" pageId="7" pageNumber="8">Diagnostic features</emphasis>
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</paragraph>
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</subSubSection>
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<subSubSection id="C360A6B9C84ABE38C4925782FE1FCF02" lastPageId="9" lastPageNumber="10" pageId="7" pageNumber="8" type="diagnosis">
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<paragraph id="8BC5F532C84ABE36C4925782FD92C81D" blockId="7.[177,1066,1456,1629]" pageId="7" pageNumber="8">
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A small-medium sized (TL:
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<quantity id="4C8258D7C84ABE36C5D05782FDABCB88" box="[499,609,1456,1480]" metricMagnitude="-3" metricUnit="m" metricValue="4.25" pageId="7" pageNumber="8" unit="mm" value="4.25">4.25 mm</quantity>
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), glabrous, depigmented, microphthalmous trechine species (
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<figureCitation id="1341E9B7C84ABE36C59C57E7FDCACBAD" box="[447,512,1493,1517]" captionStart="Fig" captionStartId="8.[215,244,1610,1628]" captionTargetBox="[268,1031,150,1568]" captionTargetId="figure-11@8.[263,1044,130,1573]" captionTargetPageId="8" captionText="Fig. 4: Trechus (Trechus) messoulii sp. nov., male holotype, habitus, dorsal aspect." figureDoi="http://doi.org/10.5281/zenodo.5749784" httpUri="https://zenodo.org/record/5749784/files/figure.png" pageId="7" pageNumber="8">Fig. 4</figureCitation>
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), with the character states of the genus
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<taxonomicName id="4C7A8EB1C84ABE36C7EE57E7FEBEC852" authority="CLAIRVILLE, 1806" authorityName="CLAIRVILLE" authorityYear="1806" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="genus">
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<emphasis id="B90E2920C84ABE36C7EE57E7FBEECBAD" box="[973,1060,1493,1517]" italics="true" pageId="7" pageNumber="8">Trechus</emphasis>
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||
CLAIRVILLE, 1806
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</taxonomicName>
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of the
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<taxonomicName id="4C7A8EB1C84ABE36C5E957C8FDDAC852" baseAuthorityName="LeConte" baseAuthorityYear="1847" box="[458,528,1530,1554]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="fulvus">
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<emphasis id="B90E2920C84ABE36C5E957C8FDDAC852" box="[458,528,1530,1554]" italics="true" pageId="7" pageNumber="8">fulvus</emphasis>
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</taxonomicName>
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species group in the sense of
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<bibRefCitation id="EFEB88C3C84ABE36C75857C8FBEFC852" author="Jeannel, R." box="[891,1061,1530,1554]" pageId="7" pageNumber="8" pagination="1 - 592" refId="ref4843" refString="Jeannel, R. (1927): Monographie des Trechinae 2. Morphologie comparee et distribution geographique d'un group de Coleopteres - L'Abeille, Paris 33: 1 - 592." type="journal article" year="1927">Jeannel (1927)</bibRefCitation>
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,
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<bibRefCitation id="EFEB88C3C84ABE36C4925412FE98C878" author="Casale, A." box="[177,338,1568,1592]" pageId="7" pageNumber="8" pagination="121 - 126" refId="ref4643" refString="Casale, A. (1983): Un nuovo Trechus d'Algeria (Coleoptera, Carabidae) - Bollettino del Museo regionale di Scienze naturali, Torino 1 (1): 121 - 126." type="journal article" year="1983">Casale (1983)</bibRefCitation>
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,
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<bibRefCitation id="EFEB88C3C84ABE36C57D5412FDCEC878" author="Ortuno, V. M." box="[350,516,1568,1592]" pageId="7" pageNumber="8" pagination="501 - 507" refId="ref5154" refString="Ortuno, V. M. (2008): Taxonomy and Systematics of a Hypogean Trechine from Southern Spain: Trechus breuili Jeannel (Coleoptera: Carabidae) - The Coleopterists Bulletin 62 (4): 501 - 507." type="journal article" year="2008">Ortuño (2008)</bibRefCitation>
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,
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<bibRefCitation id="EFEB88C3C84ABE36C6335412FCC6C878" author="Reboleira, A. S. & Ortuno, V. M. & Goncalvesi, F. & Oromi, P." box="[528,780,1568,1592]" pageId="7" pageNumber="8" pagination="15 - 26" refId="ref5345" refString="Reboleira, A. S., Ortuno, V. M., Goncalvesi, F. & Oromi, P. (2010): A hypogean new species of Trechus Clairville, 1806 (Coleoptera, Carabidae) from Portugal and considerations about the T. fulvus species group - Zootaxa 2689: 15 - 26." type="journal article" year="2010">Reboleira & al. (2010)</bibRefCitation>
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, and in the widest sense that will be discussed in Conclusions.
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</paragraph>
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<caption id="DF05A5BAC845BE39C4F45478FC6DC81C" ID-DOI="http://doi.org/10.5281/zenodo.5749784" ID-Zenodo-Dep="5749784" box="[215,935,1610,1628]" httpUri="https://zenodo.org/record/5749784/files/figure.png" pageId="8" pageNumber="9" startId="8.[215,244,1610,1628]" targetBox="[268,1031,150,1568]" targetPageId="8">
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||
<paragraph id="8BC5F532C845BE39C4F45478FC6DC81C" box="[215,935,1610,1628]" pageId="8" pageNumber="9">
|
||
Fig. 4:
|
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<taxonomicName id="4C7A8EB1C845BE39C5325478FDCEC81C" authority="Casale, 2011" authorityName="Casale" authorityYear="2011" box="[273,516,1610,1628]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="messoulii" status="sp. nov." subGenus="Trechus">
|
||
<emphasis id="B90E2920C845BE39C5325478FE9FC81C" bold="true" box="[273,341,1610,1628]" italics="true" pageId="8" pageNumber="9">Trechus</emphasis>
|
||
(
|
||
<emphasis id="B90E2920C845BE39C5435478FE6EC81C" bold="true" box="[352,420,1610,1628]" italics="true" pageId="8" pageNumber="9">Trechus</emphasis>
|
||
)
|
||
<emphasis id="B90E2920C845BE39C5935478FDCEC81C" bold="true" box="[432,516,1610,1628]" italics="true" pageId="8" pageNumber="9">messoulii</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A23D945BC845BE39C62A5478FD85C81C" box="[521,591,1610,1628]" pageId="8" pageNumber="9" rank="species">sp. nov.</taxonomicNameLabel>
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||
, male holotype, habitus, dorsal aspect.
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</paragraph>
|
||
</caption>
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<paragraph id="8BC5F532C844BE38C4F452A7FE1FCF02" blockId="9.[178,1067,149,472]" pageId="9" pageNumber="10">
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Colour dark yellow reddish. Pronotum markedly transverse, widened in front, with lateral sides regularly curved, slightly constricted and sinuate towards hind angles, which are obtuse. Elytra elongate-ovate, widened in the apical third, depressed, with humeral angles rounded. Chaetotaxy as in
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||
<figureCitation id="1341E9B7C844BE38C7C65337FBE9CF5D" box="[997,1059,261,285]" captionStart="Fig" captionStartId="8.[215,244,1610,1628]" captionTargetBox="[268,1031,150,1568]" captionTargetId="figure-11@8.[263,1044,130,1573]" captionTargetPageId="8" captionText="Fig. 4: Trechus (Trechus) messoulii sp. nov., male holotype, habitus, dorsal aspect." figureDoi="http://doi.org/10.5281/zenodo.5749784" httpUri="https://zenodo.org/record/5749784/files/figure.png" pageId="9" pageNumber="10">Fig. 4</figureCitation>
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||
. Male genitalia as in
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<figureCitation id="1341E9B7C844BE38C5B15318FE04CF02" box="[402,462,298,322]" captionStart="Fig" captionStartId="10.[215,244,154,172]" captionTargetBox="[493,1046,154,914]" captionTargetId="figure-208@10.[488,1048,149,916]" captionTargetPageId="10" captionText="Fig. 5: Trechus (Trechus) messoulii sp. nov., male holotype, aedeagus in left lateral aspect (a) and in dorsal aspect (b)." figureDoi="http://doi.org/10.5281/zenodo.5749786" httpUri="https://zenodo.org/record/5749786/files/figure.png" pageId="9" pageNumber="10">Fig. 5</figureCitation>
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.
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</paragraph>
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</subSubSection>
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||
<subSubSection id="C360A6B9C844BE38C4F45362FDB7CF98" pageId="9" pageNumber="10" type="discussion">
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||
<paragraph id="8BC5F532C844BE38C4F45362FDB7CF98" blockId="9.[178,1067,149,472]" pageId="9" pageNumber="10">
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Close to some North African species attributed to the genus
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<taxonomicName id="4C7A8EB1C844BE38C78D5362FE92CFCD" authority="JEANNEL, 1937" authorityName="JEANNEL" authorityYear="1937" class="Insecta" family="Carabidae" genus="Antoinella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="genus">
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||
<emphasis id="B90E2920C844BE38C78D5362FBEECF28" box="[942,1060,336,360]" italics="true" pageId="9" pageNumber="10">Antoinella</emphasis>
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||
<bibRefCitation id="EFEB88C3C844BE38C4915347FE92CFCD" author="Jeannel, R." box="[178,344,373,397]" pageId="9" pageNumber="10" pagination="82 - 88" refId="ref4907" refString="Jeannel, R. (1937): Nouveaux Trechinae palearctiques - Bulletin de la Societe entomologique de France 42: 82 - 88." type="journal article" year="1937">JEANNEL, 1937</bibRefCitation>
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</taxonomicName>
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, in the sense of Casale (1982),
|
||
<bibRefCitation id="EFEB88C3C844BE38C6F95347FC3ACFCD" author="Mateu, J. & Comas, J." box="[730,1008,373,397]" pageId="9" pageNumber="10" pagination="15 - 18" refId="ref4976" refString="Mateu, J. & Comas, J. (2006): Una nueva especie del genero Antoinella Jeannel, 1937 de Marruecos (Coleoptera: Carabidae: Trechinae) - Heteropterus Revista de Entomologia 6: 15 - 18." type="journal article" year="2006">Mateu & Comas (2006)</bibRefCitation>
|
||
and
|
||
<bibRefCitation id="EFEB88C3C844BE38C49153A8FE75CFF2" author="Mateu, J. & Escola, O." box="[178,447,410,434]" pageId="9" pageNumber="10" pagination="117 - 121" refId="ref5018" refString="Mateu, J. & Escola, O. (2006): El genero Antoinella Jeannel, 1937 (Coleoptera, Carabidae, Trechinae) tres especies nuevas de Marruecos: A. espanyoli sp. n., A. iblanensis sp. n. y A. fadriquei sp. n. - Animal Biodiversity and Conservation 29 (2): 117 - 121." type="journal article" year="2006">Mateu & Escolà (2006)</bibRefCitation>
|
||
, from which it is recognized by several different morphological features (see: Relationships).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C360A6B9C844BE38C491501DFEFFCC08" box="[178,309,559,584]" pageId="9" pageNumber="10" type="multiple">
|
||
<paragraph id="8BC5F532C844BE38C491501DFEFFCC08" blockId="9.[178,309,559,584]" box="[178,309,559,584]" pageId="9" pageNumber="10">
|
||
<emphasis id="B90E2920C844BE38C491501DFEFFCC08" bold="true" box="[178,309,559,584]" pageId="9" pageNumber="10">Description</emphasis>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C360A6B9C844BE3BC4925048FDFFC878" lastPageId="10" lastPageNumber="11" pageId="9" pageNumber="10" type="description">
|
||
<paragraph id="8BC5F532C844BE38C4925048FDABCD42" blockId="9.[177,1067,634,1592]" pageId="9" pageNumber="10">
|
||
General features as in
|
||
<figureCitation id="1341E9B7C844BE38C59B5048FE32CCD2" box="[440,504,634,658]" captionStart="Fig" captionStartId="8.[215,244,1610,1628]" captionTargetBox="[268,1031,150,1568]" captionTargetId="figure-11@8.[263,1044,130,1573]" captionTargetPageId="8" captionText="Fig. 4: Trechus (Trechus) messoulii sp. nov., male holotype, habitus, dorsal aspect." figureDoi="http://doi.org/10.5281/zenodo.5749784" httpUri="https://zenodo.org/record/5749784/files/figure.png" pageId="9" pageNumber="10">Fig. 4</figureCitation>
|
||
. Small-medium sized: TL:
|
||
<quantity id="4C8258D7C844BE38C7095048FC5FCCD2" box="[810,917,634,658]" metricMagnitude="-3" metricUnit="m" metricValue="4.25" pageId="9" pageNumber="10" unit="mm" value="4.25">4.25 mm</quantity>
|
||
; L:
|
||
<quantity id="4C8258D7C844BE38C79D5048FBEDCCD2" box="[958,1063,634,658]" metricMagnitude="-3" metricUnit="m" metricValue="4.5" pageId="9" pageNumber="10" unit="mm" value="4.5">4.50 mm</quantity>
|
||
. Colour uniformly dark yellow reddish. Dorsal surface moderately shiny; transversal cuticular microlines evident on frons and pronotum, scarcely visible as polygonal meshes on elytral intervals.
|
||
</paragraph>
|
||
<paragraph id="8BC5F532C844BE38C4F45122FC54CA6D" blockId="9.[177,1067,634,1592]" pageId="9" pageNumber="10">Head robust, stout, with frontal furrows markedly impressed, angulate in the anterior third; frons convex; supraorbital setiferous punctures inserted on ranges slightly convergent backwards; eyes very small, as long as 1/5 of genae, flat, oblique and whitish, with slightly distinguishable ommatidia, pigmented only at margins; genae regularly curved, slightly convex; neck wide, neck constriction moderate; labrum slightly emarginate at the anterior side; mentum tooth slightly scooped-out distally; antennae relatively long, exceeding backwards the level of the fourth humeral setiferous puncture.</paragraph>
|
||
<paragraph id="8BC5F532C844BE38C4F45608FCA8CAA8" blockId="9.[177,1067,634,1592]" pageId="9" pageNumber="10">Pronotum markedly transverse (PL/PW: 0.74), widened in front; disc convex; sides regularly curved, slightly constricted and shortly sinuate towards hind angles, which are obtuse; marginal furrows narrow and shallow; anterior margin almost straight, front angles effaced; base slightly oblique at sides, not beaded; basal foveae deep; basal area densely wrinkled.</paragraph>
|
||
<paragraph id="8BC5F532C844BE38C4F456C7FDF8C878" blockId="9.[177,1067,634,1592]" pageId="9" pageNumber="10">
|
||
Elytra elongate-ovate (EL/EW 1.47), widened in the apical third, depressed, with humeral angles rounded, widened at the apical third, much wider than prothorax; disc depressed; lateral furrows wide and deep; striae all complete and visible, moderately impressed, punctate; apical striola curved and connected with apex of stria 5; intervals flat; apical carina slightly prominent. Chaetotaxy as in
|
||
<figureCitation id="1341E9B7C844BE38C5505782FE7ACB88" box="[371,432,1456,1480]" captionStart="Fig" captionStartId="8.[215,244,1610,1628]" captionTargetBox="[268,1031,150,1568]" captionTargetId="figure-11@8.[263,1044,130,1573]" captionTargetPageId="8" captionText="Fig. 4: Trechus (Trechus) messoulii sp. nov., male holotype, habitus, dorsal aspect." figureDoi="http://doi.org/10.5281/zenodo.5749784" httpUri="https://zenodo.org/record/5749784/files/figure.png" pageId="9" pageNumber="10">Fig. 4</figureCitation>
|
||
: three setiferous discal punctures (two discal on stria 3, one pre-apical on apex of interval 3); umbilicate pores 5 and 6 very close to each other; apical and angulo-apical punctures present, forming with the preapical puncture an apical triangle.
|
||
</paragraph>
|
||
<caption id="DF05A5BAC847BE3BC4F452A8FE66CF56" ID-DOI="http://doi.org/10.5281/zenodo.5749786" ID-Zenodo-Dep="5749786" httpUri="https://zenodo.org/record/5749786/files/figure.png" pageId="10" pageNumber="11" startId="10.[215,244,154,172]" targetBox="[493,1046,154,914]" targetPageId="10">
|
||
<paragraph id="8BC5F532C847BE3BC4F452A8FE66CF56" blockId="10.[215,428,154,278]" pageId="10" pageNumber="11">
|
||
Fig. 5:
|
||
<taxonomicName id="4C7A8EB1C847BE3BC53252A8FE61CEEC" authority="(Trechus)" box="[273,427,154,172]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="subGenus" subGenus="Trechus">
|
||
<emphasis id="B90E2920C847BE3BC53252A8FE9FCEEC" bold="true" box="[273,341,154,172]" italics="true" pageId="10" pageNumber="11">Trechus</emphasis>
|
||
(
|
||
<emphasis id="B90E2920C847BE3BC54352A8FE6ECEEC" bold="true" box="[352,420,154,172]" italics="true" pageId="10" pageNumber="11">Trechus</emphasis>
|
||
)
|
||
</taxonomicName>
|
||
<emphasis id="B90E2920C847BE3BC4F45286FE66CF56" bold="true" pageId="10" pageNumber="11">
|
||
<taxonomicName id="4C7A8EB1C847BE3BC4F45286FEE1CE86" authority="Casale, 2011" authorityName="Casale" authorityYear="2011" box="[215,299,180,198]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="messoulii" status="sp. nov.">
|
||
<emphasis id="B90E2920C847BE3BC4F45286FEE1CE86" bold="true" box="[215,299,180,198]" italics="true" pageId="10" pageNumber="11">messoulii</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A23D945BC847BE3BC5135286FEBCCE86" box="[304,374,180,198]" pageId="10" pageNumber="11" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, male
|
||
<typeStatus id="54C14B90C847BE3BC4F452FDFEE2CEA1" box="[215,296,207,225]" pageId="10" pageNumber="11" type="holotype">holotype</typeStatus>
|
||
, aedeagus in left lateral aspect (a) and in dorsal aspect (b).
|
||
</emphasis>
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BC5F532C847BE3BC4DE51F8FBF8CA6D" blockId="10.[215,1105,970,1592]" pageId="10" pageNumber="11">Legs long, slender; femora robust, thickened; protibiae dilated, very shallowly grooved on the external side, and sparsely pubescent at apex; protarsi in the male each with two basal tarsomeres dilated and denticulate inwards.</paragraph>
|
||
<paragraph id="8BC5F532C847BE3BC4DE5608FEADC852" blockId="10.[215,1105,970,1592]" pageId="10" pageNumber="11">
|
||
Male genitalia as in
|
||
<figureCitation id="1341E9B7C847BE3BC5CF5608FDE6CA12" box="[492,556,1082,1106]" captionStart="Fig" captionStartId="10.[215,244,154,172]" captionTargetBox="[493,1046,154,914]" captionTargetId="figure-208@10.[488,1048,149,916]" captionTargetPageId="10" captionText="Fig. 5: Trechus (Trechus) messoulii sp. nov., male holotype, aedeagus in left lateral aspect (a) and in dorsal aspect (b)." figureDoi="http://doi.org/10.5281/zenodo.5749786" httpUri="https://zenodo.org/record/5749786/files/figure.png" pageId="10" pageNumber="11">Fig. 5</figureCitation>
|
||
. Median lobe of aedeagus elongate (
|
||
<quantity id="4C8258D7C847BE3BC7C65608FB80CA12" box="[997,1098,1082,1106]" metricMagnitude="-3" metricUnit="m" metricValue="1.05" pageId="10" pageNumber="11" unit="mm" value="1.05">1.05 mm</quantity>
|
||
long, measured from apex to the posterior margin of basal bulb, basal carina excluded), slender, slightly thickened in the middle, markedly narrowed and curved on the dorsal side at apex, with apical lamina asymmetrical widened and ventrally hooked in lateral aspect; basal bulb with big, rounded sagittal carina, not extended on the ventral side of the bulb. Endophallus with complex structure: in left lateral view, it shows a dorsal group of scales that covers a sclerotized piece of subtriangular form; in the inner part, it shows a second sclerotized piece, covered with scarcely sclerotized scales, elongate, narrowed and bent distally. Parameres robust, asymmetrical; left paramere with an elongate, ventral basal lobe on the ventral side (as is typical in
|
||
<taxonomicName id="4C7A8EB1C847BE3BC7F55782FEF8CBAD" class="Insecta" family="Carabidae" genus="Antoinella" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="undetermined">
|
||
<emphasis id="B90E2920C847BE3BC7F55782FB80CB88" box="[982,1098,1456,1480]" italics="true" pageId="10" pageNumber="11">Antoinella</emphasis>
|
||
species
|
||
</taxonomicName>
|
||
: see below), with five apical setae; right paramere shorter, with three apical setae.
|
||
</paragraph>
|
||
<paragraph id="8BC5F532C847BE3BC4DE5412FDFFC878" blockId="10.[215,1105,970,1592]" box="[253,565,1568,1592]" pageId="10" pageNumber="11">Female genitalia: unknown.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C360A6B9C846BE3AC49152A7FEAFCEEE" box="[178,357,149,174]" pageId="11" pageNumber="12" type="multiple">
|
||
<paragraph id="8BC5F532C846BE3AC49152A7FEAFCEEE" blockId="11.[178,357,149,174]" box="[178,357,149,174]" pageId="11" pageNumber="12">
|
||
<emphasis id="B90E2920C846BE3AC49152A7FEAFCEEE" bold="true" box="[178,357,149,174]" pageId="11" pageNumber="12">Specific epithet</emphasis>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C360A6B9C846BE3AC49152D2FCA2CF28" pageId="11" pageNumber="12" type="etymology">
|
||
<paragraph id="8BC5F532C846BE3AC49152D2FCA2CF28" blockId="11.[178,1067,224,360]" pageId="11" pageNumber="12">
|
||
I wish to dedicate this interesting new species to
|
||
<collectorName id="268F90E4C846BE3AC72D52D2FBEDCEB8" box="[782,1063,224,248]" pageId="11" pageNumber="12">Dr. Mohamed Messouli</collectorName>
|
||
, active speleologist in Marrakesh, who sampled the
|
||
<typeStatus id="54C14B90C846BE3AC6D65337FC90CF5D" box="[757,858,261,285]" pageId="11" pageNumber="12" type="holotype">holotype</typeStatus>
|
||
specimen several years ago, during an expedition with the
|
||
<location id="8EA5A3E9C846BE3AC6B55318FC4BCF02" LSID="urn:lsid:plazi:treatment:03D34424C84ABE3CC4935627FDE1CAA8:8EA5A3E9C846BE3AC6B55318FC4BCF02" box="[662,897,298,322]" country="France" name="Speleo Club of Blois" pageId="11" pageNumber="12">Spéléo Club of Blois</location>
|
||
(
|
||
<collectingCountry id="F36DB5A2C846BE3AC7B25318FC2DCF02" box="[913,999,298,322]" name="France" pageId="11" pageNumber="12">France</collectingCountry>
|
||
), and through our friend
|
||
<collectorName id="268F90E4C846BE3AC5A45362FD89CF28" box="[391,579,336,360]" pageId="11" pageNumber="12">Lucien C. Genest</collectorName>
|
||
offered it to me for study.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C360A6B9C846BE3AC491538DFDAACF98" box="[178,608,447,472]" pageId="11" pageNumber="12" type="multiple">
|
||
<paragraph id="8BC5F532C846BE3AC491538DFDAACF98" blockId="11.[178,608,447,472]" box="[178,608,447,472]" pageId="11" pageNumber="12">
|
||
<emphasis id="B90E2920C846BE3AC491538DFDAACF98" bold="true" box="[178,608,447,472]" pageId="11" pageNumber="12">Geographical distribution and habitat</emphasis>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C360A6B9C846BE3AC4935038FE11CD0D" pageId="11" pageNumber="12" type="distribution">
|
||
<paragraph id="8BC5F532C846BE3AC4935038FE11CD0D" blockId="11.[176,1070,522,845]" pageId="11" pageNumber="12">
|
||
The male
|
||
<typeStatus id="54C14B90C846BE3AC5065038FE46CC62" box="[293,396,522,546]" pageId="11" pageNumber="12" type="holotype">holotype</typeStatus>
|
||
of this new species was sampled in the
|
||
<location id="8EA5A3E9C846BE3AC7505038FBEFCC62" LSID="urn:lsid:plazi:treatment:03D34424C84ABE3CC4935627FDE1CAA8:8EA5A3E9C846BE3AC7505038FBEFCC62" box="[883,1061,522,546]" country="Morocco" county="African" municipality="This" name="cave Toghobeit" pageId="11" pageNumber="12">cave Toghobeit</location>
|
||
(
|
||
<location id="8EA5A3E9C846BE3AC4955002FEEACC08" LSID="urn:lsid:plazi:treatment:03D34424C84ABE3CC4935627FDE1CAA8:8EA5A3E9C846BE3AC4955002FEEACC08" box="[182,288,560,584]" country="Morocco" county="African" municipality="This" name="Northern" pageId="11" pageNumber="12">Northern</location>
|
||
<collectingCountry id="F36DB5A2C846BE3AC5045002FE47CC08" box="[295,397,560,584]" name="Morocco" pageId="11" pageNumber="12">Morocco</collectingCountry>
|
||
,
|
||
<location id="8EA5A3E9C846BE3AC5B55002FDC3CC08" LSID="urn:lsid:plazi:treatment:03D34424C84ABE3CC4935627FDE1CAA8:8EA5A3E9C846BE3AC5B55002FDC3CC08" box="[406,521,560,584]" country="Morocco" county="African" municipality="This" name="Rif Massif" pageId="11" pageNumber="12">Rif Massif</location>
|
||
,
|
||
<location id="8EA5A3E9C846BE3AC6305002FD81CC08" LSID="urn:lsid:plazi:treatment:03D34424C84ABE3CC4935627FDE1CAA8:8EA5A3E9C846BE3AC6305002FD81CC08" box="[531,587,560,584]" country="Morocco" county="African" municipality="This" name="Taza" pageId="11" pageNumber="12">Taza</location>
|
||
, near
|
||
<location id="8EA5A3E9C846BE3AC6AD5002FD38CC08" LSID="urn:lsid:plazi:treatment:03D34424C84ABE3CC4935627FDE1CAA8:8EA5A3E9C846BE3AC6AD5002FD38CC08" box="[654,754,560,584]" country="Morocco" county="African" municipality="This" name="Chaouen" pageId="11" pageNumber="12">Chaouen</location>
|
||
or
|
||
<location id="8EA5A3E9C846BE3AC7345002FC79CC08" LSID="urn:lsid:plazi:treatment:03D34424C84ABE3CC4935627FDE1CAA8:8EA5A3E9C846BE3AC7345002FC79CC08" box="[791,947,560,584]" country="Morocco" county="African" municipality="This" name="Chefchaouen" pageId="11" pageNumber="12">Chefchaouen</location>
|
||
).
|
||
<collectingMunicipality id="6BA16F48C846BE3AC7E05002FC38CC08" box="[963,1010,560,584]" pageId="11" pageNumber="12">This</collectingMunicipality>
|
||
subterranean system is so far one of the deepest caves of the
|
||
<collectingCounty id="62A48DBEC846BE3AC7795067FC61CC2D" box="[858,939,597,621]" pageId="11" pageNumber="12">African</collectingCounty>
|
||
continent: located at
|
||
<quantity id="4C8258D7C846BE3AC5095048FEB5CCD2" box="[298,383,634,658]" metricMagnitude="3" metricUnit="m" metricValue="1.7" pageId="11" pageNumber="12" unit="m" value="1700.0">
|
||
<elevation id="00571201C846BE3AC5095048FEB5CCD2" box="[298,383,634,658]" metricMagnitude="3" metricUnit="m" metricValue="1.7" pageId="11" pageNumber="12" unit="m" value="1700.0">1700 m</elevation>
|
||
</quantity>
|
||
of altitude, its actual depth is -
|
||
<quantity id="4C8258D7C846BE3AC6C35048FCE1CCD2" box="[736,811,634,658]" metricMagnitude="2" metricUnit="m" metricValue="7.32" pageId="11" pageNumber="12" unit="m" value="732.0">
|
||
<elevation id="00571201C846BE3AC6C35048FCE1CCD2" box="[736,811,634,658]" metricMagnitude="2" metricUnit="m" metricValue="7.32" pageId="11" pageNumber="12" unit="m" value="732.0">732 m</elevation>
|
||
</quantity>
|
||
, and the development reaches
|
||
<quantity id="4C8258D7C846BE3AC5305092FEA3CCF8" box="[275,361,672,696]" metricMagnitude="3" metricUnit="m" metricValue="3.05" pageId="11" pageNumber="12" unit="m" value="3050.0">
|
||
<elevation id="00571201C846BE3AC5305092FEA3CCF8" box="[275,361,672,696]" metricMagnitude="3" metricUnit="m" metricValue="3.05" pageId="11" pageNumber="12" unit="m" value="3050.0">3050 m</elevation>
|
||
</quantity>
|
||
(after: "Inventaire Spéléologique du Maroc", 1981: DC 5, Kef Toghobeit). The only examined individual was collected in the "salle Renner" at -
|
||
<elevation id="00571201C846BE3AC4FB50D8FEA5CD42" box="[216,367,746,770]" metricMagnitude="2" metricUnit="m" metricValue="1.2" pageId="11" pageNumber="12" unit="m" value="120.0">
|
||
<quantity id="4C8258D7C846BE3AC4FB50D8FEEACD42" box="[216,288,746,770]" metricMagnitude="2" metricUnit="m" metricValue="1.2" pageId="11" pageNumber="12" unit="m" value="120.0">120 m</quantity>
|
||
depth
|
||
</elevation>
|
||
, but a second individual (not available at the moment) was collected in the "salle du bivouac" and entrusted to
|
||
<collectorName id="268F90E4C846BE3AC7385122FC63CD68" box="[795,937,784,808]" pageId="11" pageNumber="12">L. C. Genest</collectorName>
|
||
(
|
||
<collectorName id="268F90E4C846BE3AC79B5122FBECCD68" box="[952,1062,784,808]" pageId="11" pageNumber="12">Messouli</collectorName>
|
||
, personal communication).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C360A6B9C846BE3AC4915197FE85CDFE" box="[178,335,933,958]" pageId="11" pageNumber="12" type="multiple">
|
||
<paragraph id="8BC5F532C846BE3AC4915197FE85CDFE" blockId="11.[178,335,933,958]" box="[178,335,933,958]" pageId="11" pageNumber="12">
|
||
<emphasis id="B90E2920C846BE3AC4915197FE85CDFE" bold="true" box="[178,335,933,958]" pageId="11" pageNumber="12">Relationships</emphasis>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C360A6B9C846BE3DC48C51C2FD2CC852" lastPageId="12" lastPageNumber="13" pageId="11" pageNumber="12" type="discussion">
|
||
<paragraph id="8BC5F532C846BE3AC48C51C2FBE9CAA8" blockId="11.[175,1067,1008,1629]" pageId="11" pageNumber="12">
|
||
<taxonomicName id="4C7A8EB1C846BE3AC48C51C2FE48CA48" authority="Casale, 2011" authorityName="Casale" authorityYear="2011" box="[175,386,1008,1032]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="messoulii" status="sp. nov.">
|
||
<emphasis id="B90E2920C846BE3AC48C51C2FE48CA48" box="[175,386,1008,1032]" italics="true" pageId="11" pageNumber="12">Trechus messoulii</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A23D945BC846BE3AC5A851C2FE21CA48" box="[395,491,1008,1032]" pageId="11" pageNumber="12" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, for both its external features and peculiar characters of male genitalia – in particular, the structure of copulatory pieces in endophallus (a sclerotized piece of triangular shape, a second elongate piece, and some groups of sclerotized scales) – is close to some
|
||
<taxonomicName id="4C7A8EB1C846BE3AC7715652FBCCCA38" box="[850,1030,1120,1144]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="undetermined">
|
||
<emphasis id="B90E2920C846BE3AC7715652FC63CA38" box="[850,937,1120,1144]" italics="true" pageId="11" pageNumber="12">Trechus</emphasis>
|
||
species
|
||
</taxonomicName>
|
||
of the
|
||
<taxonomicName id="4C7A8EB1C846BE3AC4FC56B7FEEFCADD" baseAuthorityName="LeConte" baseAuthorityYear="1847" box="[223,293,1157,1181]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="fulvus">
|
||
<emphasis id="B90E2920C846BE3AC4FC56B7FEEFCADD" box="[223,293,1157,1181]" italics="true" pageId="11" pageNumber="12">fulvus</emphasis>
|
||
</taxonomicName>
|
||
species group in the sense of
|
||
<bibRefCitation id="EFEB88C3C846BE3AC6A956B7FC42CADD" author="Reboleira, A. S. & Ortuno, V. M. & Goncalvesi, F. & Oromi, P." box="[650,904,1157,1181]" pageId="11" pageNumber="12" pagination="15 - 26" refId="ref5345" refString="Reboleira, A. S., Ortuno, V. M., Goncalvesi, F. & Oromi, P. (2010): A hypogean new species of Trechus Clairville, 1806 (Coleoptera, Carabidae) from Portugal and considerations about the T. fulvus species group - Zootaxa 2689: 15 - 26." type="journal article" year="2010">Reboleira & al. (2010)</bibRefCitation>
|
||
(in particular, for the widened and hooked apical lamina of the median lobe of the aedeagus, it recalls
|
||
<taxonomicName id="4C7A8EB1C846BE3AC53B56E2FDEFCAA8" authority="JEANNE, 1988" authorityName="JEANNE" authorityYear="1988" box="[280,549,1232,1256]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="arribasi">
|
||
<emphasis id="B90E2920C846BE3AC53B56E2FE43CAA8" box="[280,393,1232,1256]" italics="true" pageId="11" pageNumber="12">T. arribasi</emphasis>
|
||
JEANNE, 1988
|
||
</taxonomicName>
|
||
, endemic to the Northern Iberian peninsula).
|
||
</paragraph>
|
||
<paragraph id="8BC5F532C846BE3AC4F456C7FC0CC81D" blockId="11.[175,1067,1008,1629]" pageId="11" pageNumber="12">
|
||
But it is mostly related to Algerian species (
|
||
<bibRefCitation id="EFEB88C3C846BE3AC6F656C7FCBCCB4D" author="Jeannel, R." box="[725,886,1269,1293]" pageId="11" pageNumber="12" pagination="1 - 592" refId="ref4843" refString="Jeannel, R. (1927): Monographie des Trechinae 2. Morphologie comparee et distribution geographique d'un group de Coleopteres - L'Abeille, Paris 33: 1 - 592." type="journal article" year="1927">Jeannel, 1927</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFEB88C3C846BE3AC7A456C7FBD7CB4D" author="Casale, A." box="[903,1053,1269,1293]" pageId="11" pageNumber="12" pagination="121 - 126" refId="ref4643" refString="Casale, A. (1983): Un nuovo Trechus d'Algeria (Coleoptera, Carabidae) - Bollettino del Museo regionale di Scienze naturali, Torino 1 (1): 121 - 126." type="journal article" year="1983">Casale, 1983</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFEB88C3C846BE3AC4915728FE82CB72" author="Ortuno, V. M." box="[178,328,1306,1330]" pageId="11" pageNumber="12" pagination="501 - 507" refId="ref5154" refString="Ortuno, V. M. (2008): Taxonomy and Systematics of a Hypogean Trechine from Southern Spain: Trechus breuili Jeannel (Coleoptera: Carabidae) - The Coleopterists Bulletin 62 (4): 501 - 507." type="journal article" year="2008">Ortuño, 2008</bibRefCitation>
|
||
) and several subterranean North African species of the Rif massif (
|
||
<collectingCountry id="F36DB5A2C846BE3AC4955772FED7CB18" box="[182,285,1344,1368]" name="Morocco" pageId="11" pageNumber="12">Morocco</collectingCountry>
|
||
) currently treated as
|
||
<taxonomicName id="4C7A8EB1C846BE3AC62C5772FCE1CB18" authority="JEANNEL, 1937" authorityName="JEANNEL" authorityYear="1937" box="[527,811,1344,1368]" class="Insecta" family="Carabidae" genus="Antoinella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B90E2920C846BE3AC62C5772FD48CB18" box="[527,642,1344,1368]" italics="true" pageId="11" pageNumber="12">Antoinella</emphasis>
|
||
<bibRefCitation id="EFEB88C3C846BE3AC6AA5772FCE1CB18" author="Jeannel, R." box="[649,811,1344,1368]" pageId="11" pageNumber="12" pagination="82 - 88" refId="ref4907" refString="Jeannel, R. (1937): Nouveaux Trechinae palearctiques - Bulletin de la Societe entomologique de France 42: 82 - 88." type="journal article" year="1937">JEANNEL, 1937</bibRefCitation>
|
||
</taxonomicName>
|
||
(Casale, 1982;
|
||
<bibRefCitation id="EFEB88C3C846BE3AC7FE5772FE94CB3D" author="Mateu, J. & Comas, J." pageId="11" pageNumber="12" pagination="15 - 18" refId="ref4976" refString="Mateu, J. & Comas, J. (2006): Una nueva especie del genero Antoinella Jeannel, 1937 de Marruecos (Coleoptera: Carabidae: Trechinae) - Heteropterus Revista de Entomologia 6: 15 - 18." type="journal article" year="2006">Mateu & Comas, 2006</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFEB88C3C846BE3AC54E5757FDAFCB3D" author="Mateu, J. & Escola, O." box="[365,613,1381,1405]" pageId="11" pageNumber="12" pagination="117 - 121" refId="ref5018" refString="Mateu, J. & Escola, O. (2006): El genero Antoinella Jeannel, 1937 (Coleoptera, Carabidae, Trechinae) tres especies nuevas de Marruecos: A. espanyoli sp. n., A. iblanensis sp. n. y A. fadriquei sp. n. - Animal Biodiversity and Conservation 29 (2): 117 - 121." type="journal article" year="2006">Mateu & Escolà, 2006</bibRefCitation>
|
||
). In particular, it seems to be close to
|
||
<taxonomicName id="4C7A8EB1C846BE3AC0305757FDFFCBE2" authority="MATEU & ESCOLa, 2006" authorityName="MATEU & ESCOLa" authorityYear="2006" class="Insecta" family="Carabidae" genus="Antoinella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="iblanensis">
|
||
<emphasis id="B90E2920C846BE3AC0305757FEECCBE2" italics="true" pageId="11" pageNumber="12">A. iblanensis</emphasis>
|
||
<bibRefCitation id="EFEB88C3C846BE3AC50E57B8FDFFCBE2" author="Mateu, J. & Escola, O." box="[301,565,1418,1442]" pageId="11" pageNumber="12" pagination="117 - 121" refId="ref5018" refString="Mateu, J. & Escola, O. (2006): El genero Antoinella Jeannel, 1937 (Coleoptera, Carabidae, Trechinae) tres especies nuevas de Marruecos: A. espanyoli sp. n., A. iblanensis sp. n. y A. fadriquei sp. n. - Animal Biodiversity and Conservation 29 (2): 117 - 121." type="journal article" year="2006">MATEU & ESCOLà, 2006</bibRefCitation>
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="4C7A8EB1C846BE3AC65757B8FBC2CBE2" authority="MATEU & ESCOLa, 2006" authorityName="MATEU & ESCOLa" authorityYear="2006" box="[628,1032,1418,1442]" class="Insecta" family="Carabidae" genus="Antoinella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="fadriquei">
|
||
<emphasis id="B90E2920C846BE3AC65757B8FD33CBE2" box="[628,761,1418,1442]" italics="true" pageId="11" pageNumber="12">A. fadriquei</emphasis>
|
||
<bibRefCitation id="EFEB88C3C846BE3AC6DC57B8FBC2CBE2" author="Mateu, J. & Escola, O." box="[767,1032,1418,1442]" pageId="11" pageNumber="12" pagination="117 - 121" refId="ref5018" refString="Mateu, J. & Escola, O. (2006): El genero Antoinella Jeannel, 1937 (Coleoptera, Carabidae, Trechinae) tres especies nuevas de Marruecos: A. espanyoli sp. n., A. iblanensis sp. n. y A. fadriquei sp. n. - Animal Biodiversity and Conservation 29 (2): 117 - 121." type="journal article" year="2006">MATEU & ESCOLà, 2006</bibRefCitation>
|
||
</taxonomicName>
|
||
. It is, however, markedly distinct by the smaller size (
|
||
<quantity id="4C8258D7C846BE3AC6C85782FCBBCB88" box="[747,881,1456,1480]" metricMagnitude="-3" metricUnit="m" metricValue="5.85" metricValueMax="5.9" metricValueMin="5.8" pageId="11" pageNumber="12" unit="mm" value="5.85" valueMax="5.9" valueMin="5.8">5.8–5.9 mm</quantity>
|
||
in
|
||
<taxonomicName id="4C7A8EB1C846BE3AC7B05782FBEFCB88" authorityName="MATEU & ESCOLa" authorityYear="2006" box="[915,1061,1456,1480]" class="Insecta" family="Carabidae" genus="Antoinella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="iblanensis">
|
||
<emphasis id="B90E2920C846BE3AC7B05782FBEFCB88" box="[915,1061,1456,1480]" italics="true" pageId="11" pageNumber="12">A. iblanensis</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C7A8EB1C846BE3AC4C057E7FEA2CBAD" authorityName="MATEU & ESCOLa" authorityYear="2006" box="[227,360,1493,1517]" class="Insecta" family="Carabidae" genus="Antoinella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="fadriquei">
|
||
<emphasis id="B90E2920C846BE3AC4C057E7FEA2CBAD" box="[227,360,1493,1517]" italics="true" pageId="11" pageNumber="12">A. fadriquei</emphasis>
|
||
</taxonomicName>
|
||
, respectively, about
|
||
<quantity id="4C8258D7C846BE3AC67457E7FD66CBAD" box="[599,684,1493,1517]" metricMagnitude="-3" metricUnit="m" metricValue="4.5" pageId="11" pageNumber="12" unit="mm" value="4.5">4.5 mm</quantity>
|
||
in
|
||
<taxonomicName id="4C7A8EB1C846BE3AC6EC57E7FC9CCBAD" authority="Casale, 2011" authorityName="Casale" authorityYear="2011" box="[719,854,1493,1517]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="messoulii" status="sp. nov.">
|
||
<emphasis id="B90E2920C846BE3AC6EC57E7FC9CCBAD" box="[719,854,1493,1517]" italics="true" pageId="11" pageNumber="12">T. messoulii</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A23D945BC846BE3AC77D57E7FC76CBAD" box="[862,956,1493,1517]" pageId="11" pageNumber="12" rank="species">sp. nov.</taxonomicNameLabel>
|
||
), and the shape of the median lobe of the aedeagus, which in
|
||
<taxonomicName id="4C7A8EB1C846BE3AC73857C8FC6CC852" authority="Casale, 2011" authorityName="Casale" authorityYear="2011" box="[795,934,1530,1554]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="messoulii" status="sp. nov.">
|
||
<emphasis id="B90E2920C846BE3AC73857C8FC6CC852" box="[795,934,1530,1554]" italics="true" pageId="11" pageNumber="12">T. messoulii</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A23D945BC846BE3AC78D57C8FBC3C852" box="[942,1033,1530,1554]" pageId="11" pageNumber="12" rank="species">sp. nov.</taxonomicNameLabel>
|
||
is slender and elongate as in
|
||
<taxonomicName id="4C7A8EB1C846BE3AC5C15412FDBEC878" authorityName="MATEU & ESCOLa" authorityYear="2006" box="[482,628,1568,1592]" class="Insecta" family="Carabidae" genus="Antoinella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="iblanensis">
|
||
<emphasis id="B90E2920C846BE3AC5C15412FDBEC878" box="[482,628,1568,1592]" italics="true" pageId="11" pageNumber="12">A. iblanensis</emphasis>
|
||
</taxonomicName>
|
||
, but shows an apical lamina similar to that of
|
||
<taxonomicName id="4C7A8EB1C846BE3AC5215477FE4FC81D" authorityName="MATEU & ESCOLa" authorityYear="2006" box="[258,389,1605,1629]" class="Insecta" family="Carabidae" genus="Antoinella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="fadriquei">
|
||
<emphasis id="B90E2920C846BE3AC5215477FE4FC81D" box="[258,389,1605,1629]" italics="true" pageId="11" pageNumber="12">A. fadriquei</emphasis>
|
||
</taxonomicName>
|
||
, more curved dorsally and more obtusely truncate.
|
||
</paragraph>
|
||
<paragraph id="8BC5F532C841BE3DC4DE52A7FC8BCE92" blockId="12.[213,1105,149,1144]" pageId="12" pageNumber="13">In this case, the combination of both morphological features and biogeographical considerations allows the following remarks:</paragraph>
|
||
<paragraph id="8BC5F532C841BE3DC4F652D2FC47CC08" blockId="12.[213,1105,149,1144]" pageId="12" pageNumber="13">
|
||
1. From the taxonomical point of view, the so-called genus
|
||
<taxonomicName id="4C7A8EB1C841BE3DC7B452D2FBC6CEB8" box="[919,1036,224,248]" class="Insecta" family="Carabidae" genus="Antoinella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B90E2920C841BE3DC7B452D2FBC6CEB8" box="[919,1036,224,248]" italics="true" pageId="12" pageNumber="13">Antoinella</emphasis>
|
||
</taxonomicName>
|
||
JEANNEL, 1937, appears as a paraphyletic unit, without any relationships with the genus
|
||
<taxonomicName id="4C7A8EB1C841BE3DC5585318FE2ACF02" authorityName="Delarouzee" authorityYear="1859" box="[379,480,298,322]" class="Insecta" family="Carabidae" genus="Duvalius" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B90E2920C841BE3DC5585318FE2ACF02" box="[379,480,298,322]" italics="true" pageId="12" pageNumber="13">Duvalius</emphasis>
|
||
</taxonomicName>
|
||
. This datum is evident from both morphological features, as already supposed by
|
||
<bibRefCitation id="EFEB88C3C841BE3DC64A5362FCD9CF28" author="Jeannel, R." box="[617,787,336,360]" pageId="12" pageNumber="13" pagination="82 - 88" refId="ref4907" refString="Jeannel, R. (1937): Nouveaux Trechinae palearctiques - Bulletin de la Societe entomologique de France 42: 82 - 88." type="journal article" year="1937">Jeannel (1937)</bibRefCitation>
|
||
in the original description of the genus, and supported by Casale (1982, 1983); furthermore, it has been confirmed by a recent molecular phylogeny (
|
||
<bibRefCitation id="EFEB88C3C841BE3DC71653A8FC25CFF2" author="Faille, A. & Casale, A. & Ribera, I." box="[821,1007,410,434]" pageId="12" pageNumber="13" refId="ref4801" refString="Faille, A., Casale, A. & Ribera, I. (2010): Phylogenetic relationships of Western Mediterranean subterranean Trechini groundbeetles (Coleoptera: Carabidae) - Zoologica Scripta (printed, version on line)." type="book" year="2010">Faille & al. 2010</bibRefCitation>
|
||
). Therefore, either it will be demonstrated that
|
||
<taxonomicName id="4C7A8EB1C841BE3DC69453F2FCC7CF98" authorityName="CLAIRVILLE" authorityYear="1806" box="[695,781,448,472]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B90E2920C841BE3DC69453F2FCC7CF98" box="[695,781,448,472]" italics="true" pageId="12" pageNumber="13">Trechus</emphasis>
|
||
</taxonomicName>
|
||
is a polyphyletic unit, and in this case the
|
||
<taxonomicName id="4C7A8EB1C841BE3DC5BB53D7FE33CFBD" baseAuthorityName="LeConte" baseAuthorityYear="1847" box="[408,505,485,509]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="fulvus">
|
||
<emphasis id="B90E2920C841BE3DC5BB53D7FE33CFBD" box="[408,505,485,509]" italics="true" pageId="12" pageNumber="13">T. fulvus</emphasis>
|
||
</taxonomicName>
|
||
species group should merit the genus rank (named
|
||
<taxonomicName id="4C7A8EB1C841BE3DC4D85038FEBECC62" box="[251,372,522,546]" class="Insecta" family="Carabidae" genus="Antoinella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B90E2920C841BE3DC4D85038FEBECC62" box="[251,372,522,546]" italics="true" pageId="12" pageNumber="13">Antoinella</emphasis>
|
||
</taxonomicName>
|
||
), or the latter is a simple lineage within this group, and an objective junior synonym of
|
||
<taxonomicName id="4C7A8EB1C841BE3DC6185002FD58CC08" authorityName="CLAIRVILLE" authorityYear="1806" box="[571,658,560,584]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B90E2920C841BE3DC6185002FD58CC08" box="[571,658,560,584]" italics="true" pageId="12" pageNumber="13">Trechus</emphasis>
|
||
</taxonomicName>
|
||
(in the widest sense).
|
||
</paragraph>
|
||
<paragraph id="8BC5F532C841BE3DC4F55067FC73CDFD" blockId="12.[213,1105,149,1144]" pageId="12" pageNumber="13">
|
||
2. The only current distinctive character between
|
||
<taxonomicName id="4C7A8EB1C841BE3DC71B5067FC7BCC2D" box="[824,945,597,621]" class="Insecta" family="Carabidae" genus="Antoinella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B90E2920C841BE3DC71B5067FC7BCC2D" box="[824,945,597,621]" italics="true" pageId="12" pageNumber="13">Antoinella</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C7A8EB1C841BE3DC7D35067FB81CC2D" authorityName="CLAIRVILLE" authorityYear="1806" box="[1008,1099,597,621]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B90E2920C841BE3DC7D35067FB81CC2D" box="[1008,1099,597,621]" italics="true" pageId="12" pageNumber="13">Trechus</emphasis>
|
||
</taxonomicName>
|
||
of the
|
||
<taxonomicName id="4C7A8EB1C841BE3DC56D5048FE5DCCD2" baseAuthorityName="LeConte" baseAuthorityYear="1847" box="[334,407,634,658]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="fulvus">
|
||
<emphasis id="B90E2920C841BE3DC56D5048FE5DCCD2" box="[334,407,634,658]" italics="true" pageId="12" pageNumber="13">fulvus</emphasis>
|
||
</taxonomicName>
|
||
species group, i.e. protibiae pubescent in
|
||
<taxonomicName id="4C7A8EB1C841BE3DC7805048FBD6CCD2" box="[931,1052,634,658]" class="Insecta" family="Carabidae" genus="Antoinella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B90E2920C841BE3DC7805048FBD6CCD2" box="[931,1052,634,658]" italics="true" pageId="12" pageNumber="13">Antoinella</emphasis>
|
||
</taxonomicName>
|
||
(as in
|
||
<taxonomicName id="4C7A8EB1C841BE3DC5385092FE2FCCF8" box="[283,485,672,696]" class="Insecta" family="Carabidae" genus="Duvalius" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="undetermined">
|
||
<emphasis id="B90E2920C841BE3DC5385092FEB4CCF8" box="[283,382,672,696]" italics="true" pageId="12" pageNumber="13">Duvalius</emphasis>
|
||
species
|
||
</taxonomicName>
|
||
), and glabrous in
|
||
<taxonomicName id="4C7A8EB1C841BE3DC6945092FCC5CCF8" authorityName="CLAIRVILLE" authorityYear="1806" box="[695,783,672,696]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B90E2920C841BE3DC6945092FCC5CCF8" box="[695,783,672,696]" italics="true" pageId="12" pageNumber="13">Trechus</emphasis>
|
||
</taxonomicName>
|
||
, is not a constant diagnostic feature:
|
||
<bibRefCitation id="EFEB88C3C841BE3DC5A550F7FD43CC9D" author="Reboleira, A. S. & Ortuno, V. M. & Goncalvesi, F. & Oromi, P." box="[390,649,709,733]" pageId="12" pageNumber="13" pagination="15 - 26" refId="ref5345" refString="Reboleira, A. S., Ortuno, V. M., Goncalvesi, F. & Oromi, P. (2010): A hypogean new species of Trechus Clairville, 1806 (Coleoptera, Carabidae) from Portugal and considerations about the T. fulvus species group - Zootaxa 2689: 15 - 26." type="journal article" year="2010">Reboleira & al. (2010)</bibRefCitation>
|
||
for instance, who have just described
|
||
<taxonomicName id="4C7A8EB1C841BE3DC4D950D8FE80CD42" authorityName="Riboleira; Ortuno; Goncalves & Oromi" authorityYear="2010" box="[250,330,746,770]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="tatai">
|
||
<emphasis id="B90E2920C841BE3DC4D950D8FE80CD42" box="[250,330,746,770]" italics="true" pageId="12" pageNumber="13">T. tatai</emphasis>
|
||
</taxonomicName>
|
||
from
|
||
<collectingCountry id="F36DB5A2C841BE3DC5B150D8FE30CD42" box="[402,506,746,770]" name="Portugal" pageId="12" pageNumber="13">Portugal</collectingCountry>
|
||
, specified that it has "conspicuously hairy (including the protibiae)" legs. Furthermore, the structure of the median lobe of aedeagus, endophallus, and asymmetrical parameres (the left one with an elongate, ventral basal lobe on the ventral side) is absolutely the same both in
|
||
<taxonomicName id="4C7A8EB1C841BE3DC57551B2FDECCDD8" box="[342,550,896,920]" class="Insecta" family="Carabidae" genus="Antoinella" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="undetermined">
|
||
<emphasis id="B90E2920C841BE3DC57551B2FE00CDD8" box="[342,458,896,920]" italics="true" pageId="12" pageNumber="13">Antoinella</emphasis>
|
||
species
|
||
</taxonomicName>
|
||
and in some Algerian
|
||
<taxonomicName id="4C7A8EB1C841BE3DC70B51B2FCB5CDD8" authorityName="CLAIRVILLE" authorityYear="1806" box="[808,895,896,920]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B90E2920C841BE3DC70B51B2FCB5CDD8" box="[808,895,896,920]" italics="true" pageId="12" pageNumber="13">Trechus</emphasis>
|
||
</taxonomicName>
|
||
of the
|
||
<taxonomicName id="4C7A8EB1C841BE3DC7F151B2FBD2CDD8" baseAuthorityName="LeConte" baseAuthorityYear="1847" box="[978,1048,896,920]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="fulvus">
|
||
<emphasis id="B90E2920C841BE3DC7F151B2FBD2CDD8" box="[978,1048,896,920]" italics="true" pageId="12" pageNumber="13">fulvus</emphasis>
|
||
</taxonomicName>
|
||
species group, as illustrated by
|
||
<bibRefCitation id="EFEB88C3C841BE3DC6185197FD15CDFD" author="Jeannel, R." box="[571,735,933,957]" pageId="12" pageNumber="13" pagination="1 - 592" refId="ref4843" refString="Jeannel, R. (1927): Monographie des Trechinae 2. Morphologie comparee et distribution geographique d'un group de Coleopteres - L'Abeille, Paris 33: 1 - 592." type="journal article" year="1927">Jeannel (1927)</bibRefCitation>
|
||
and
|
||
<bibRefCitation id="EFEB88C3C841BE3DC7365197FC7ECDFD" author="Casale, A." box="[789,948,933,957]" pageId="12" pageNumber="13" pagination="121 - 126" refId="ref4643" refString="Casale, A. (1983): Un nuovo Trechus d'Algeria (Coleoptera, Carabidae) - Bollettino del Museo regionale di Scienze naturali, Torino 1 (1): 121 - 126." type="journal article" year="1983">Casale (1983)</bibRefCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BC5F532C841BE3DC4F651F8FBF3CA38" blockId="12.[213,1105,149,1144]" pageId="12" pageNumber="13">
|
||
3. Finally
|
||
<bibRefCitation id="EFEB88C3C841BE3DC56D51F8FE3ECDA2" author="Ortuno, V. M." box="[334,500,970,994]" pageId="12" pageNumber="13" pagination="501 - 507" refId="ref5154" refString="Ortuno, V. M. (2008): Taxonomy and Systematics of a Hypogean Trechine from Southern Spain: Trechus breuili Jeannel (Coleoptera: Carabidae) - The Coleopterists Bulletin 62 (4): 501 - 507." type="journal article" year="2008">Ortuño (2008)</bibRefCitation>
|
||
and
|
||
<bibRefCitation id="EFEB88C3C841BE3DC61351F8FCFBCDA2" author="Reboleira, A. S. & Ortuno, V. M. & Goncalvesi, F. & Oromi, P." box="[560,817,970,994]" pageId="12" pageNumber="13" pagination="15 - 26" refId="ref5345" refString="Reboleira, A. S., Ortuno, V. M., Goncalvesi, F. & Oromi, P. (2010): A hypogean new species of Trechus Clairville, 1806 (Coleoptera, Carabidae) from Portugal and considerations about the T. fulvus species group - Zootaxa 2689: 15 - 26." type="journal article" year="2010">Reboleira & al. (2010)</bibRefCitation>
|
||
, in their excellent contributions, exhaustively showed that the
|
||
<taxonomicName id="4C7A8EB1C841BE3DC69F51C2FCD7CA48" baseAuthorityName="LeConte" baseAuthorityYear="1847" box="[700,797,1008,1032]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="fulvus">
|
||
<emphasis id="B90E2920C841BE3DC69F51C2FCD7CA48" box="[700,797,1008,1032]" italics="true" pageId="12" pageNumber="13">T. fulvus</emphasis>
|
||
</taxonomicName>
|
||
species group, in its wide distribution along North Africa, the Iberian Peninsula and, thanks to the wide distribution of
|
||
<taxonomicName id="4C7A8EB1C841BE3DC5C05608FD89CA12" baseAuthorityName="LeConte" baseAuthorityYear="1847" box="[483,579,1082,1106]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="fulvus">
|
||
<emphasis id="B90E2920C841BE3DC5C05608FD89CA12" box="[483,579,1082,1106]" italics="true" pageId="12" pageNumber="13">T. fulvus</emphasis>
|
||
</taxonomicName>
|
||
, also along the Atlantic coast of northern Europe (
|
||
<bibRefCitation id="EFEB88C3C841BE3DC5105652FE00CA38" author="Jeannel, R." box="[307,458,1120,1144]" pageId="12" pageNumber="13" pagination="1 - 592" refId="ref4843" refString="Jeannel, R. (1927): Monographie des Trechinae 2. Morphologie comparee et distribution geographique d'un group de Coleopteres - L'Abeille, Paris 33: 1 - 592." type="journal article" year="1927">Jeannel 1927</bibRefCitation>
|
||
), is heterogeneous, and includes at least two lineages.
|
||
</paragraph>
|
||
<paragraph id="8BC5F532C841BE3DC4DE5698FD2CC852" blockId="12.[215,1105,1194,1554]" pageId="12" pageNumber="13">
|
||
In this context, it is evident that the so-called
|
||
<taxonomicName id="4C7A8EB1C841BE3DC72E5698FC28CA82" box="[781,994,1194,1218]" class="Insecta" family="Carabidae" genus="Antoinella" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="undetermined">
|
||
<emphasis id="B90E2920C841BE3DC72E5698FC4BCA82" box="[781,897,1194,1218]" italics="true" pageId="12" pageNumber="13">Antoinella</emphasis>
|
||
species
|
||
</taxonomicName>
|
||
, with the
|
||
<taxonomicName id="4C7A8EB1C841BE3DC4FB56E2FE8CCAA8" authorityName="Jeannel" authorityYear="1927" box="[216,326,1232,1256]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="martinezi">
|
||
<emphasis id="B90E2920C841BE3DC4FB56E2FE8CCAA8" box="[216,326,1232,1256]" italics="true" pageId="12" pageNumber="13">martinezi</emphasis>
|
||
</taxonomicName>
|
||
-lineage (sensu
|
||
<bibRefCitation id="EFEB88C3C841BE3DC5DD56E2FD25CAA8" author="Ortuno, V. M. & Arillo, A." box="[510,751,1232,1256]" pageId="12" pageNumber="13" pagination="3483 - 3500" refId="ref5196" refString="Ortuno, V. M. & Arillo, A. (2005): Description of a new hypogean species of the genus Trechus Clairville, 1806 from eastern Spain and comments on the Trechus martinezi - lineage (Coleoptera: Adephaga: Carabidae) - Journal of Natural History, 39 (40), 3483 - 3500." type="journal article" year="2005">Ortuño & Arillo 2005</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFEB88C3C841BE3DC6DC56E2FC58CAA8" author="Ortuno, V. M." box="[767,914,1232,1256]" pageId="12" pageNumber="13" pagination="501 - 507" refId="ref5154" refString="Ortuno, V. M. (2008): Taxonomy and Systematics of a Hypogean Trechine from Southern Spain: Trechus breuili Jeannel (Coleoptera: Carabidae) - The Coleopterists Bulletin 62 (4): 501 - 507." type="journal article" year="2008">Ortuño 2008</bibRefCitation>
|
||
), form a typical subterranean lineage within the
|
||
<taxonomicName id="4C7A8EB1C841BE3DC66E56C7FD39CB4D" baseAuthorityName="LeConte" baseAuthorityYear="1847" box="[589,755,1269,1293]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="fulvus">
|
||
<emphasis id="B90E2920C841BE3DC66E56C7FD39CB4D" box="[589,755,1269,1293]" italics="true" pageId="12" pageNumber="13">Trechus fulvus</emphasis>
|
||
</taxonomicName>
|
||
species group, which has survived in the south-western Mediterranean region. The present distribution of extant species suggests a Betic-Rifean origin (see also
|
||
<bibRefCitation id="EFEB88C3C841BE3DC77F5772FC3BCB18" author="Ortuno, V. M." box="[860,1009,1344,1368]" pageId="12" pageNumber="13" pagination="501 - 507" refId="ref5154" refString="Ortuno, V. M. (2008): Taxonomy and Systematics of a Hypogean Trechine from Southern Spain: Trechus breuili Jeannel (Coleoptera: Carabidae) - The Coleopterists Bulletin 62 (4): 501 - 507." type="journal article" year="2008">Ortuño 2008</bibRefCitation>
|
||
: fig. 4), with strong biogeographical coherence: a well known case in entomological literature. Another interesting fact is the surprising speciosity of this group in the Rif massif: in fact every cave, so far, revealed a separate species markedly distinct from its relatives, all with markedly reduced eyes, depigmented integument, and other troglomorphic features.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C360A6B9C840BE3CC4F452A7FC9BCEED" box="[215,849,149,174]" pageId="13" pageNumber="14" type="multiple">
|
||
<paragraph id="8BC5F532C840BE3CC4F452A7FC9BCEED" blockId="13.[176,1062,149,285]" box="[215,849,149,174]" pageId="13" pageNumber="14">
|
||
In conclusion, the following
|
||
<emphasis id="B90E2920C840BE3CC63152A7FD76CEEE" bold="true" box="[530,700,149,174]" pageId="13" pageNumber="14">new synonymy</emphasis>
|
||
is proposed:
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C360A6B9C840BE3CC4935288FBD5CF5D" pageId="13" pageNumber="14" type="reference_group">
|
||
<paragraph id="8BC5F532C840BE3CC4935288FBD5CF5D" blockId="13.[176,1062,149,285]" pageId="13" pageNumber="14">
|
||
<treatmentCitationGroup id="AB6AD21CC840BE3CC4935288FBD5CF5D" pageId="13" pageNumber="14">
|
||
<treatmentCitation id="0ADBD323C840BE3CC4935288FE11CE92" box="[176,475,186,210]" pageId="13" pageNumber="14">
|
||
<taxonomicName id="4C7A8EB1C840BE3CC4935288FE11CE92" ID-CoL="8LQVP" authority="JEANNEL, 1937" authorityName="JEANNEL" authorityYear="1937" box="[176,475,186,210]" class="Insecta" family="Carabidae" genus="Antoinella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B90E2920C840BE3CC4935288FEEDCE92" box="[176,295,186,210]" italics="true" pageId="13" pageNumber="14">Antoinella</emphasis>
|
||
<bibRefCitation id="EFEB88C3C840BE3CC5115288FE11CE92" author="Jeannel, R." box="[306,475,186,210]" pageId="13" pageNumber="14" pagination="82 - 88" refId="ref4907" refString="Jeannel, R. (1937): Nouveaux Trechinae palearctiques - Bulletin de la Societe entomologique de France 42: 82 - 88." type="journal article" year="1937">JEANNEL, 1937</bibRefCitation>
|
||
</taxonomicName>
|
||
</treatmentCitation>
|
||
(
|
||
<typeStatus id="54C14B90C840BE3CC5CF5288FDE9CE92" box="[492,547,186,210]" pageId="13" pageNumber="14">type</typeStatus>
|
||
species:
|
||
<treatmentCitation id="0ADBD323C840BE3CC6BA5288FBD7CE92" box="[665,1053,186,210]" pageId="13" pageNumber="14">
|
||
<taxonomicName id="4C7A8EB1C840BE3CC6BA5288FBD7CE92" ID-CoL="8LXDN" authority="ANTOINE, 1935" authorityName="ANTOINE" authorityYear="1935" box="[665,1053,186,210]" class="Insecta" family="Carabidae" genus="Duvalius" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="groubei">
|
||
<emphasis id="B90E2920C840BE3CC6BA5288FCAECE92" box="[665,868,186,210]" italics="true" pageId="13" pageNumber="14">Duvalius groubei</emphasis>
|
||
ANTOINE, 1935
|
||
</taxonomicName>
|
||
</treatmentCitation>
|
||
)
|
||
<emphasis id="B90E2920C840BE3CC4F452EDFE4CCEB8" bold="true" box="[215,390,223,248]" pageId="13" pageNumber="14">junior synonym</emphasis>
|
||
of
|
||
<treatmentCitation id="0ADBD323C840BE3CC58A52D2FD09CEB8" box="[425,707,224,248]" pageId="13" pageNumber="14">
|
||
<taxonomicName id="4C7A8EB1C840BE3CC58A52D2FD09CEB8" ID-CoL="8KZB9" authority="CLAIRVILLE, 1806" authorityName="CLAIRVILLE" authorityYear="1806" box="[425,707,224,248]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B90E2920C840BE3CC58A52D2FDCACEB8" box="[425,512,224,248]" italics="true" pageId="13" pageNumber="14">Trechus</emphasis>
|
||
CLAIRVILLE, 1806
|
||
</taxonomicName>
|
||
</treatmentCitation>
|
||
(
|
||
<typeStatus id="54C14B90C840BE3CC6F352D2FCCFCEB8" box="[720,773,224,248]" pageId="13" pageNumber="14">type</typeStatus>
|
||
species:
|
||
<treatmentCitation id="0ADBD323C840BE3CC75752D2FD86CF5D" pageId="13" pageNumber="14">
|
||
<taxonomicName id="4C7A8EB1C840BE3CC75752D2FD86CF5D" ID-CoL="QYQW" authority="FABRICIUS sensu CLAIRVILLE, 1806" authorityName="CLAIRVILLE" authorityYear="1806" class="Insecta" family="Carabidae" genus="Carabus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="rubens">
|
||
<emphasis id="B90E2920C840BE3CC75752D2FBEECEB8" box="[884,1060,224,248]" italics="true" pageId="13" pageNumber="14">Carabus rubens</emphasis>
|
||
FABRICIUS sensu CLAIRVILLE, 1806
|
||
</taxonomicName>
|
||
</treatmentCitation>
|
||
=
|
||
<treatmentCitation id="0ADBD323C840BE3CC64A5337FBD5CF5D" box="[617,1055,261,285]" pageId="13" pageNumber="14">
|
||
<taxonomicName id="4C7A8EB1C840BE3CC64A5337FBDCCF5D" ID-CoL="8L7X5" authority="SCHRANK, 1781" authorityName="SCHRANK" authorityYear="1781" box="[617,1046,261,285]" class="Insecta" family="Carabidae" genus="Carabus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="quadristriatus">
|
||
<emphasis id="B90E2920C840BE3CC64A5337FCADCF5D" box="[617,871,261,285]" italics="true" pageId="13" pageNumber="14">Carabus quadristriatus</emphasis>
|
||
SCHRANK, 1781
|
||
</taxonomicName>
|
||
)
|
||
</treatmentCitation>
|
||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C360A6B9C840BE3CC4F55362FDE1CAA8" pageId="13" pageNumber="14" type="discussion">
|
||
<paragraph id="8BC5F532C840BE3CC4F55362FD9FCC08" blockId="13.[176,1067,336,1256]" pageId="13" pageNumber="14">
|
||
A separate question is the validity and rank of
|
||
<taxonomicName id="4C7A8EB1C840BE3CC7335362FF21CFCD" authority="MATEU & COMAS, 2006" authorityName="MATEU & COMAS" authorityYear="2006" class="Insecta" family="Carabidae" genus="Irinea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B90E2920C840BE3CC7335362FC9ECF28" box="[784,852,336,360]" italics="true" pageId="13" pageNumber="14">Irinea</emphasis>
|
||
<bibRefCitation id="EFEB88C3C840BE3CC77D5362FF21CFCD" author="Mateu, J. & Comas, J." pageId="13" pageNumber="14" pagination="15 - 18" refId="ref4976" refString="Mateu, J. & Comas, J. (2006): Una nueva especie del genero Antoinella Jeannel, 1937 de Marruecos (Coleoptera: Carabidae: Trechinae) - Heteropterus Revista de Entomologia 6: 15 - 18." type="journal article" year="2006">MATEU & COMAS, 2006</bibRefCitation>
|
||
</taxonomicName>
|
||
, described as subgenus of
|
||
<taxonomicName id="4C7A8EB1C840BE3CC60F5347FD6ACFCD" box="[556,672,373,397]" class="Insecta" family="Carabidae" genus="Antoinella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B90E2920C840BE3CC60F5347FD6ACFCD" box="[556,672,373,397]" italics="true" pageId="13" pageNumber="14">Antoinella</emphasis>
|
||
</taxonomicName>
|
||
. The
|
||
<typeStatus id="54C14B90C840BE3CC6C05347FCDCCFCD" box="[739,790,373,397]" pageId="13" pageNumber="14">type</typeStatus>
|
||
species,
|
||
<taxonomicName id="4C7A8EB1C840BE3CC7A75347FE7DCFF2" authority="MATEU & COMAS, 2006" authorityName="MATEU & COMAS" authorityYear="2006" class="Insecta" family="Carabidae" genus="Antoinella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="aurouxi" subGenus="I.">
|
||
<emphasis id="B90E2920C840BE3CC7A75347FC59CFCD" box="[900,915,373,397]" italics="true" pageId="13" pageNumber="14">A</emphasis>
|
||
. (
|
||
<emphasis id="B90E2920C840BE3CC78E5347FC76CFCD" box="[941,956,373,397]" italics="true" pageId="13" pageNumber="14">I.</emphasis>
|
||
)
|
||
<emphasis id="B90E2920C840BE3CC7EE5347FBEECFCD" box="[973,1060,373,397]" italics="true" pageId="13" pageNumber="14">aurouxi</emphasis>
|
||
<bibRefCitation id="EFEB88C3C840BE3CC49253A8FE7DCFF2" author="Mateu, J. & Comas, J." box="[177,439,410,434]" pageId="13" pageNumber="14" pagination="15 - 18" refId="ref4976" refString="Mateu, J. & Comas, J. (2006): Una nueva especie del genero Antoinella Jeannel, 1937 de Marruecos (Coleoptera: Carabidae: Trechinae) - Heteropterus Revista de Entomologia 6: 15 - 18." type="journal article" year="2006">MATEU & COMAS, 2006</bibRefCitation>
|
||
</taxonomicName>
|
||
, shows marked distinctive features: in particular, only one basal tarsomere is dilated in fore tarsi of males, median lobe of aedeagus without basal sagittal carina – typically present in all "
|
||
<taxonomicName id="4C7A8EB1C840BE3CC76353D7FC7CCFBD" box="[832,950,485,509]" class="Insecta" family="Carabidae" genus="Antoinella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B90E2920C840BE3CC76353D7FC7CCFBD" box="[832,950,485,509]" italics="true" pageId="13" pageNumber="14">Antoinella</emphasis>
|
||
</taxonomicName>
|
||
" species and
|
||
<taxonomicName id="4C7A8EB1C840BE3CC4C15038FEF3CC62" authorityName="CLAIRVILLE" authorityYear="1806" box="[226,313,522,546]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B90E2920C840BE3CC4C15038FEF3CC62" box="[226,313,522,546]" italics="true" pageId="13" pageNumber="14">Trechus</emphasis>
|
||
</taxonomicName>
|
||
of the
|
||
<taxonomicName id="4C7A8EB1C840BE3CC5A45038FE07CC62" baseAuthorityName="LeConte" baseAuthorityYear="1847" box="[391,461,522,546]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="fulvus">
|
||
<emphasis id="B90E2920C840BE3CC5A45038FE07CC62" box="[391,461,522,546]" italics="true" pageId="13" pageNumber="14">fulvus</emphasis>
|
||
</taxonomicName>
|
||
species group – and endophallus of simple structure, with only one small copulatory piece.
|
||
</paragraph>
|
||
<paragraph id="8BC5F532C840BE3CC4F45067FDF2CADD" blockId="13.[176,1067,336,1256]" pageId="13" pageNumber="14">
|
||
Nevertheless, these characters are not unique and peculiar to
|
||
<taxonomicName id="4C7A8EB1C840BE3CC78E5067FC24CC2D" authorityName="MATEU & COMAS" authorityYear="2006" box="[941,1006,597,621]" class="Insecta" family="Carabidae" genus="Irinea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B90E2920C840BE3CC78E5067FC24CC2D" box="[941,1006,597,621]" italics="true" pageId="13" pageNumber="14">Irinea</emphasis>
|
||
</taxonomicName>
|
||
, but present in several species attributed to
|
||
<taxonomicName id="4C7A8EB1C840BE3CC6AA5048FD28CCD2" authorityName="CLAIRVILLE" authorityYear="1806" box="[649,738,634,658]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B90E2920C840BE3CC6AA5048FD28CCD2" box="[649,738,634,658]" italics="true" pageId="13" pageNumber="14">Trechus</emphasis>
|
||
</taxonomicName>
|
||
in the current (and widest) sense. In a contribution dedicated to the Ethiopian
|
||
<taxonomicName id="4C7A8EB1C840BE3CC6DA5092FC78CCF8" box="[761,946,672,696]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="undetermined">
|
||
<emphasis id="B90E2920C840BE3CC6DA5092FC9ACCF8" box="[761,848,672,696]" italics="true" pageId="13" pageNumber="14">Trechus</emphasis>
|
||
species
|
||
</taxonomicName>
|
||
, Ortuño & Novoa (in press) discuss the sexual dimorphism in species of this genus, and stress that the occurrence of a single dilated articulation on the protarsus in males is present in the Nearctic subgenus
|
||
<taxonomicName id="4C7A8EB1C840BE3CC6BA5122FC1FCD68" authority="JEANNEL, 1927" authorityName="JEANNEL" authorityYear="1927" box="[665,981,784,808]" class="Insecta" family="Carabidae" genus="Microtrechus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B90E2920C840BE3CC6BA5122FCE4CD68" box="[665,814,784,808]" italics="true" pageId="13" pageNumber="14">Microtrechus</emphasis>
|
||
<bibRefCitation id="EFEB88C3C840BE3CC7165122FC1FCD68" author="Jeannel, R." box="[821,981,784,808]" pageId="13" pageNumber="14" pagination="1 - 592" refId="ref4843" refString="Jeannel, R. (1927): Monographie des Trechinae 2. Morphologie comparee et distribution geographique d'un group de Coleopteres - L'Abeille, Paris 33: 1 - 592." type="journal article" year="1927">JEANNEL, 1927</bibRefCitation>
|
||
</taxonomicName>
|
||
, in
|
||
<taxonomicName id="4C7A8EB1C840BE3CC0235122FD15CD0D" authority="WOLLASTON, 1854" authorityName="WOLLASTON" authorityYear="1854" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="cautus" subGenus="Atlantotrechus">
|
||
<emphasis id="B90E2920C840BE3CC0235122FDCECD0D" italics="true" pageId="13" pageNumber="14">Trechus (Atlantotrechus) cautus</emphasis>
|
||
WOLLASTON, 1854
|
||
</taxonomicName>
|
||
, from Madeira and in some Ethiopian species (Ortuño, personal communication; Pawlowski 2003). Furthermore, it must be recalled that recently
|
||
<bibRefCitation id="EFEB88C3C840BE3CC68951B2FBECCDD8" author="Vigna Taglianti, A. & Magrini, P." box="[682,1062,896,920]" pageId="13" pageNumber="14" pagination="69 - 76" refId="ref5405" refString="Vigna Taglianti, A. & Magrini, P. (2009): A new subgenus and a new species of Trechus from Ethiopia (Coleoptera, Carabidae) - Fragmenta entomologica (Roma) 41 (1): 69 - 76." type="journal article" year="2009">Vigna Taglianti & Magrini (2009)</bibRefCitation>
|
||
described the subgenus
|
||
<taxonomicName id="4C7A8EB1C840BE3CC5F75197FD97CDFD" authorityName="Vigna Taglianti & Magrini" authorityYear="2009" box="[468,605,933,957]" class="Insecta" family="Carabidae" genus="Minitrechus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B90E2920C840BE3CC5F75197FD97CDFD" box="[468,605,933,957]" italics="true" pageId="13" pageNumber="14">Minitrechus</emphasis>
|
||
</taxonomicName>
|
||
for
|
||
<taxonomicName id="4C7A8EB1C840BE3CC6AD5197FC8DCDFD" authority="Casale, 2011" authorityName="Casale" authorityYear="2011" box="[654,839,933,957]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="gypaeti" status="sp. nov.">
|
||
<emphasis id="B90E2920C840BE3CC6AD5197FC8DCDFD" box="[654,839,933,957]" italics="true" pageId="13" pageNumber="14">Trechus gypaeti</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="A23D945BC840BE3CC7745197FC2CCDFD" box="[855,998,933,957]" pageId="13" pageNumber="14" rank="species">new species</taxonomicNameLabel>
|
||
from
|
||
<collectingCountry id="F36DB5A2C840BE3CC49151F8FEDECDA2" box="[178,276,970,994]" name="Ethiopia" pageId="13" pageNumber="14">Ethiopia</collectingCountry>
|
||
, in which only one basal tarsomere is dilated in fore tarsi of males. Therefore, given that the apomorphy of a single dilated tarsomere in males appears sporadically in
|
||
<taxonomicName id="4C7A8EB1C840BE3CC5E45627FDD5CA6D" authorityName="CLAIRVILLE" authorityYear="1806" box="[455,543,1045,1069]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B90E2920C840BE3CC5E45627FDD5CA6D" box="[455,543,1045,1069]" italics="true" pageId="13" pageNumber="14">Trechus</emphasis>
|
||
</taxonomicName>
|
||
belonging to different lineages and in different areas, this character seems not a valid characteristic able to distinguish just one group within
|
||
<taxonomicName id="4C7A8EB1C840BE3CC58E5652FDCECA38" authorityName="CLAIRVILLE" authorityYear="1806" box="[429,516,1120,1144]" class="Insecta" family="Carabidae" genus="Trechus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B90E2920C840BE3CC58E5652FDCECA38" box="[429,516,1120,1144]" italics="true" pageId="13" pageNumber="14">Trechus</emphasis>
|
||
</taxonomicName>
|
||
in the widest sense (Ortuño & Novoa, in press; Ortuño, personal communication).
|
||
</paragraph>
|
||
<paragraph id="8BC5F532C840BE3CC4F65698FDE1CAA8" blockId="13.[176,1067,336,1256]" pageId="13" pageNumber="14">
|
||
Therefore taxonomic position, validity, and phylogenetic relationships of
|
||
<taxonomicName id="4C7A8EB1C840BE3CC49256E2FF38CAA8" authorityName="MATEU & COMAS" authorityYear="2006" box="[177,242,1232,1256]" class="Insecta" family="Carabidae" genus="Irinea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B90E2920C840BE3CC49256E2FF38CAA8" box="[177,242,1232,1256]" italics="true" pageId="13" pageNumber="14">Irinea</emphasis>
|
||
</taxonomicName>
|
||
merit further investigation.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |