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<document id="0CA6685FEBAA73D2AC9B8C3BAD38D86C" ID-DOI="10.5281/zenodo.199263" ID-GBIF-Dataset="6907b192-2120-45f7-aa79-7fb04b007561" ID-ISSN="1175-5326" ID-Zenodo-Dep="199263" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460332548641" checkinUser="plazi" docAuthor="Ayón-Parente, Manuel &amp; Hendrickx, Michel E." docDate="2010" docId="03C587ED9F2EFFF8FF47F298FC5EFA70" docLanguage="en" docName="zt02677p059.pdf" docOrigin="Zootaxa 2677" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Tetralobistes Ayón-Parente &amp; Hendrickx, 2010, gen. nov." docType="treatment" docVersion="9" lastPageNumber="52" masterDocId="FFFCFF959F2FFFFBFFD0F32FFFD4FFBC" masterDocTitle="A new genus and new species of hermit crab (Crustacea: Anomura: Paguroidea: Diogenidae) from the eastern tropical Pacific" masterLastPageNumber="59" masterPageNumber="49" pageNumber="50" updateTime="1698258954691" updateUser="plazi">
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<mods:title id="8CDEBA4ACD62AE9CF1D77C9A7BFE627B">A new genus and new species of hermit crab (Crustacea: Anomura: Paguroidea: Diogenidae) from the eastern tropical Pacific</mods:title>
</mods:titleInfo>
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<mods:namePart id="0961BE89355BFC97972DB85C169BA989">Ayón-Parente, Manuel</mods:namePart>
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<mods:namePart id="B3D8708D25F9383BD66FA57FBB31CFF6">Hendrickx, Michel E.</mods:namePart>
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<mods:date id="391F67BF6DCE3F26814B332CE3501F8D">2010</mods:date>
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<mods:identifier id="3E92F1F1F6CCD74E1AAC818156862BB9" type="DOI">10.5281/zenodo.199263</mods:identifier>
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<mods:identifier id="5780127002CFB31F0AA14AD3967C11ED" type="ISSN">1175-5326</mods:identifier>
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<treatment id="03C587ED9F2EFFF8FF47F298FC5EFA70" ID-DOI="http://doi.org/10.5281/zenodo.6208345" ID-GBIF-Taxon="119404672" ID-Zenodo-Dep="6208345" LSID="urn:lsid:plazi:treatment:03C587ED9F2EFFF8FF47F298FC5EFA70" httpUri="http://treatment.plazi.org/id/03C587ED9F2EFFF8FF47F298FC5EFA70" lastPageId="3" lastPageNumber="52" pageId="1" pageNumber="50">
<subSubSection id="C37665709F2EFFFAFF47F298FF3BFE48" pageId="1" pageNumber="50" type="nomenclature">
<paragraph id="8BD336FB9F2EFFFAFF47F298FE70FE6D" blockId="1.[151,420,439,500]" box="[151,420,439,465]" pageId="1" pageNumber="50">
<heading id="D09B81979F2EFFFAFF47F298FE70FE6D" bold="true" box="[151,420,439,465]" fontSize="11" level="1" pageId="1" pageNumber="50" reason="1">
<emphasis id="B918EAE99F2EFFFAFF47F298FE70FE6D" bold="true" box="[151,420,439,465]" pageId="1" pageNumber="50">
<taxonomicName id="4C6C4D789F2EFFFAFF47F298FEFBFE6D" ID-CoL="44ead6da-d490-48d9-9ada-29280282c9ea" box="[151,303,439,465]" class="Malacostraca" family="Diogenidae" genus="Tetralobistes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="50" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B918EAE99F2EFFFAFF47F298FEFBFE6D" bold="true" box="[151,303,439,465]" italics="true" pageId="1" pageNumber="50">Tetralobistes</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22B57929F2EFFFAFEE6F298FE70FE6D" box="[310,420,439,465]" pageId="1" pageNumber="50" rank="genus">gen. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BD336FB9F2EFFFAFF47F2F5FF3BFE48" blockId="1.[151,420,439,500]" box="[151,239,474,500]" pageId="1" pageNumber="50">
(
<figureCitation id="13572A7E9F2EFFFAFF4FF2F5FF31FE48" box="[159,229,474,500]" captionStart="FIGURE 2" captionStartId="6.[151,255,1981,2005]" captionTargetBox="[223,1377,194,1968]" captionTargetId="figure@6.[175,1419,194,1970]" captionTargetPageId="6" captionText="FIGURE 2. Tetralobistes bicentenarius gen. nov. and sp. nov., A – E, holotype female (SL, 2.10 mm) (EMU- 8730). Left mouthparts, inner view. A, maxillule; B, maxilla; C, first maxilliped; D, second maxilliped; E, third maxilliped." httpUri="https://zenodo.org/record/199265/files/figure.png" pageId="1" pageNumber="50">Fig. 2</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C37665709F2EFFFAFF47F10AFB9EFB28" pageId="1" pageNumber="50" type="diagnosis">
<paragraph id="8BD336FB9F2EFFFAFF47F10AFE80FCF8" blockId="1.[151,1437,549,2031]" pageId="1" pageNumber="50">
<emphasis id="B918EAE99F2EFFFAFF47F10AFECCFD83" bold="true" box="[151,280,549,575]" pageId="1" pageNumber="50">Diagnosis.</emphasis>
Thirteen pairs of biserial gills; shield well calcified; rostrum short, subtriangular. Ocular acicles well developed, terminating in strong multifid projection, mesial margins contiguous. Antennal peduncles with supernumerary segmentation; acicles short, spinose. Antennal flagella of medium length, longer than carapace, with long setae ventrally. Mandible with unarmed mesial margin, palp 3-segmented. Endopod of maxillule (
<figureCitation id="13572A7E9F2EFFFAFEC9F195FEB5FD68" box="[281,353,698,724]" captionStart="FIGURE 2" captionStartId="6.[151,255,1981,2005]" captionTargetBox="[223,1377,194,1968]" captionTargetId="figure@6.[175,1419,194,1970]" captionTargetPageId="6" captionText="FIGURE 2. Tetralobistes bicentenarius gen. nov. and sp. nov., A – E, holotype female (SL, 2.10 mm) (EMU- 8730). Left mouthparts, inner view. A, maxillule; B, maxilla; C, first maxilliped; D, second maxilliped; E, third maxilliped." httpUri="https://zenodo.org/record/199265/files/figure.png" pageId="1" pageNumber="50">Fig. 2</figureCitation>
A) with well developed, recurved external lobe. Maxilla (
<figureCitation id="13572A7E9F2EFFFAFBC6F195FB8AFD68" box="[1046,1118,698,724]" captionStart="FIGURE 2" captionStartId="6.[151,255,1981,2005]" captionTargetBox="[223,1377,194,1968]" captionTargetId="figure@6.[175,1419,194,1970]" captionTargetPageId="6" captionText="FIGURE 2. Tetralobistes bicentenarius gen. nov. and sp. nov., A – E, holotype female (SL, 2.10 mm) (EMU- 8730). Left mouthparts, inner view. A, maxillule; B, maxilla; C, first maxilliped; D, second maxilliped; E, third maxilliped." httpUri="https://zenodo.org/record/199265/files/figure.png" pageId="1" pageNumber="50">Fig. 2</figureCitation>
B) with elongate endopod. First maxilliped (
<figureCitation id="13572A7E9F2EFFFAFEB6F1CFFE7AFD46" box="[358,430,736,762]" captionStart="FIGURE 2" captionStartId="6.[151,255,1981,2005]" captionTargetBox="[223,1377,194,1968]" captionTargetId="figure@6.[175,1419,194,1970]" captionTargetPageId="6" captionText="FIGURE 2. Tetralobistes bicentenarius gen. nov. and sp. nov., A – E, holotype female (SL, 2.10 mm) (EMU- 8730). Left mouthparts, inner view. A, maxillule; B, maxilla; C, first maxilliped; D, second maxilliped; E, third maxilliped." httpUri="https://zenodo.org/record/199265/files/figure.png" pageId="1" pageNumber="50">Fig. 2</figureCitation>
C) with well developed epipod. Second maxilliped (
<figureCitation id="13572A7E9F2EFFFAFBF0F1CFFBBCFD46" box="[1056,1128,736,762]" captionStart="FIGURE 2" captionStartId="6.[151,255,1981,2005]" captionTargetBox="[223,1377,194,1968]" captionTargetId="figure@6.[175,1419,194,1970]" captionTargetPageId="6" captionText="FIGURE 2. Tetralobistes bicentenarius gen. nov. and sp. nov., A – E, holotype female (SL, 2.10 mm) (EMU- 8730). Left mouthparts, inner view. A, maxillule; B, maxilla; C, first maxilliped; D, second maxilliped; E, third maxilliped." httpUri="https://zenodo.org/record/199265/files/figure.png" pageId="1" pageNumber="50">Fig. 2</figureCitation>
D) without distinguishing characters. Third maxilliped (
<figureCitation id="13572A7E9F2EFFFAFE2EF02AFD92FCA3" box="[510,582,773,799]" captionStart="FIGURE 2" captionStartId="6.[151,255,1981,2005]" captionTargetBox="[223,1377,194,1968]" captionTargetId="figure@6.[175,1419,194,1970]" captionTargetPageId="6" captionText="FIGURE 2. Tetralobistes bicentenarius gen. nov. and sp. nov., A – E, holotype female (SL, 2.10 mm) (EMU- 8730). Left mouthparts, inner view. A, maxillule; B, maxilla; C, first maxilliped; D, second maxilliped; E, third maxilliped." httpUri="https://zenodo.org/record/199265/files/figure.png" pageId="1" pageNumber="50">Fig. 2</figureCitation>
E) with basal segments approximate; crista dentata well developed; no accessory tooth.
</paragraph>
<paragraph id="8BD336FB9F2EFFFAFF15F07FFE66FC08" blockId="1.[151,1437,549,2031]" pageId="1" pageNumber="50">Chelipeds subequal, right usually slightly longer than left, similar in armature; fingers opening in horizontal plane, fingertips acuminate. Fourth pereopods simple, without preungual process at base of claw. Fifth pereopods chelate.</paragraph>
<paragraph id="8BD336FB9F2EFFFAFF15F0EFFB7AFC43" blockId="1.[151,1437,549,2031]" pageId="1" pageNumber="50">Males with paired gonopores; paired pleopods modified as gonopods on first pleonal somite; no pleopods on the second somite; unpaired, well developed, uniramous left pleopods on somites 3–5.</paragraph>
<paragraph id="8BD336FB9F2EFFFAFF15F725FAACFBF6" blockId="1.[151,1437,549,2031]" pageId="1" pageNumber="50">Females with paired gonopores; first pleonal somite without modified pleopods; following 3 pleomeres with unequally biramous left pleopods; fifth pleopod uniramous; brood pouch represented by row of setae.</paragraph>
<paragraph id="8BD336FB9F2EFFFAFF15F77AFB9EFB28" blockId="1.[151,1437,549,2031]" pageId="1" pageNumber="50">Uropods asymmetrical. Telson asymmetrical, with lateral indentation; posterior margin divided into four lobes (tetralobular); external lobes unequal, larger than symmetrical, inner lobes.</paragraph>
</subSubSection>
<subSubSection id="C37665709F2EFFFAFF15F7B0FBC7FB06" box="[197,1043,1183,1210]" pageId="1" pageNumber="50" type="materials_examined">
<paragraph id="8BD336FB9F2EFFFAFF15F7B0FBC7FB06" blockId="1.[151,1437,549,2031]" box="[197,1043,1183,1210]" pageId="1" pageNumber="50">
<emphasis id="B918EAE99F2EFFFAFF15F7B0FEB3FB05" bold="true" box="[197,359,1183,1209]" pageId="1" pageNumber="50">
<typeStatus id="54D788599F2EFFFAFF15F7B0FED6FB05" box="[197,258,1183,1209]" pageId="1" pageNumber="50">Type</typeStatus>
species.
</emphasis>
<taxonomicName id="4C6C4D789F2EFFFAFEBEF78FFD7FFB05" box="[366,683,1184,1209]" class="Malacostraca" family="Diogenidae" genus="Tetralobistes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="50" phylum="Arthropoda" rank="species" species="bicentenarius" status="sp. nov.">
<emphasis id="B918EAE99F2EFFFAFEBEF78FFD7FFB05" box="[366,683,1184,1209]" italics="true" pageId="1" pageNumber="50">Tetralobistes bicentenarius</emphasis>
</taxonomicName>
<emphasis id="B918EAE99F2EFFFAFD63F7B0FD21FB05" bold="true" box="[691,757,1183,1209]" pageId="1" pageNumber="50">
<taxonomicNameLabel id="A22B57929F2EFFFAFD63F7B0FD21FB05" box="[691,757,1183,1209]" pageId="1" pageNumber="50" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, by present designation.
</paragraph>
</subSubSection>
<subSubSection id="C37665709F2EFFFAFF15F7EAFDC1FA96" pageId="1" pageNumber="50" type="etymology">
<paragraph id="8BD336FB9F2EFFFAFF15F7EAFDC1FA96" blockId="1.[151,1437,549,2031]" pageId="1" pageNumber="50">
<emphasis id="B918EAE99F2EFFFAFF15F7EAFE9FFB63" bold="true" box="[197,331,1221,1247]" pageId="1" pageNumber="50">Etymology</emphasis>
. The name of the genus refer to the posterior margin of the telson which is divided into four lobes (tetralobular), and is a combination of the Greek words
<emphasis id="B918EAE99F2EFFFAFC59F7C4FC17FAB8" box="[905,963,1259,1284]" italics="true" pageId="1" pageNumber="50">tetra</emphasis>
(four) and
<emphasis id="B918EAE99F2EFFFAFB9EF7C4FB44FAB8" box="[1102,1168,1259,1284]" italics="true" pageId="1" pageNumber="50">lobos</emphasis>
(lobe), and the Greek suffix –
<emphasis id="B918EAE99F2EFFFAFF21F63FFEF1FA95" box="[241,293,1296,1321]" italics="true" pageId="1" pageNumber="50">istes</emphasis>
. Gender, masculine.
</paragraph>
</subSubSection>
<subSubSection id="C37665709F2EFFFAFF15F61AFCABFAF3" box="[197,895,1333,1359]" pageId="1" pageNumber="50" type="distribution">
<paragraph id="8BD336FB9F2EFFFAFF15F61AFCABFAF3" blockId="1.[151,1437,549,2031]" box="[197,895,1333,1359]" pageId="1" pageNumber="50">
<emphasis id="B918EAE99F2EFFFAFF15F61AFEB1FAF3" bold="true" box="[197,357,1333,1359]" pageId="1" pageNumber="50">Distribution.</emphasis>
Matanchen Bay, Nayarit,
<collectingCountry id="F37B766B9F2EFFFAFD48F61AFD21FAF3" box="[664,757,1333,1359]" name="Mexico" pageId="1" pageNumber="50">Mexico</collectingCountry>
;
<quantity id="4C949B1E9F2EFFFAFCD1F61AFCAEFAF3" box="[769,890,1333,1359]" metricMagnitude="0" metricUnit="m" metricValue="1.0" metricValueMax="1.5" metricValueMin="0.5" pageId="1" pageNumber="50" unit="m" value="1.0" valueMax="1.5" valueMin="0.5">0.5–1.5 m</quantity>
.
</paragraph>
</subSubSection>
<subSubSection id="C37665709F2EFFF8FF15F675FC5EFA70" lastPageId="3" lastPageNumber="52" pageId="1" pageNumber="50" type="discussion">
<paragraph id="8BD336FB9F2EFFFAFF15F675FDDEF9E8" blockId="1.[151,1437,549,2031]" pageId="1" pageNumber="50">
<emphasis id="B918EAE99F2EFFFAFF15F675FEEAFAC8" bold="true" box="[197,318,1370,1396]" pageId="1" pageNumber="50">Remarks.</emphasis>
<taxonomicName id="4C6C4D789F2EFFFAFE96F675FE19FAC8" box="[326,461,1370,1396]" class="Malacostraca" family="Diogenidae" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="50" phylum="Arthropoda" rank="family">Diogenidae</taxonomicName>
have long been considered as evolutionarily more primitive than
<taxonomicName id="4C6C4D789F2EFFFAFB02F675FA9FFAC8" box="[1234,1355,1370,1396]" class="Malacostraca" family="Paguridae" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="50" phylum="Arthropoda" rank="family">Paguridae</taxonomicName>
. In her review of characters used to define affinities among hermit crabs, de Saint Laurent (1970) recognised the importance of the presence and number of pleurobranchiae on pereopods, the modification of pleopods into gonopods in males, the presence or absence of first (males and females) and second (males only) pleopods, and relative lengths of the endo- and exopodites of the pleopods. Additionally, the presence, reduction or absence of the brood pouch in females has also been considered an important phylogenetic character (
<bibRefCitation id="EFFD4B0A9F2EFFFAFF4EF515FE29F9E8" author="McLaughlin" box="[158,509,1594,1620]" pageId="1" pageNumber="50" refString="McLaughlin, P. A. &amp; Hogarth, P. J. (1998) Hermit crabs (Decapoda: Anomura: Paguridea) from the Seychelles. Zoologische Verhandelingen Leiden 318, 14. iv, 1 - 48." type="journal article" year="1998">McLaughlin &amp; Hogarth 1998</bibRefCitation>
).
</paragraph>
<paragraph id="8BD336FB9F2EFFFAFF16F54FFD1DF888" blockId="1.[151,1437,549,2031]" pageId="1" pageNumber="50">
Considering the most significant characters that separate the genera of
<taxonomicName id="4C6C4D789F2EFFFAFBFDF54FFB63F9C6" box="[1069,1207,1632,1658]" class="Malacostraca" family="Diogenidae" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="50" phylum="Arthropoda" rank="family">Diogenidae</taxonomicName>
,
<taxonomicName id="4C6C4D789F2EFFFAFB17F54FFA89F9C5" box="[1223,1373,1632,1657]" class="Malacostraca" family="Diogenidae" genus="Tetralobistes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="50" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B918EAE99F2EFFFAFB17F54FFA89F9C5" box="[1223,1373,1632,1657]" italics="true" pageId="1" pageNumber="50">Tetralobistes</emphasis>
</taxonomicName>
<emphasis id="B918EAE99F2EFFFAFAB9F570FF1FF923" bold="true" pageId="1" pageNumber="50">
<taxonomicNameLabel id="A22B57929F2EFFFAFAB9F570FF1FF923" pageId="1" pageNumber="50" rank="genus">gen. nov.</taxonomicNameLabel>
</emphasis>
is more closely related to
<taxonomicName id="4C6C4D789F2EFFFAFDDEF5AAFD11F922" box="[526,709,1669,1694]" class="Malacostraca" family="Diogenidae" genus="Areopaguristes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="50" phylum="Arthropoda" rank="genus">
<emphasis id="B918EAE99F2EFFFAFDDEF5AAFD11F922" box="[526,709,1669,1694]" italics="true" pageId="1" pageNumber="50">Areopaguristes</emphasis>
</taxonomicName>
,
<taxonomicName id="4C6C4D789F2EFFFAFD05F5AAFC82F922" box="[725,854,1669,1694]" class="Malacostraca" family="Diogenidae" genus="Paguristes" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="50" phylum="Arthropoda" rank="genus">
<emphasis id="B918EAE99F2EFFFAFD05F5AAFC82F922" box="[725,854,1669,1694]" italics="true" pageId="1" pageNumber="50">Paguristes</emphasis>
</taxonomicName>
and
<taxonomicName id="4C6C4D789F2EFFFAFC42F5AAFBBCF922" box="[914,1128,1669,1694]" class="Malacostraca" family="Diogenidae" genus="Pseudopaguristes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="50" phylum="Arthropoda" rank="genus">
<emphasis id="B918EAE99F2EFFFAFC42F5AAFBBCF922" box="[914,1128,1669,1694]" italics="true" pageId="1" pageNumber="50">Pseudopaguristes</emphasis>
</taxonomicName>
than to other genera. For comparative purposes, 12 characters were selected and analysed (see
<tableCitation id="C6EE03409F2EFFFAFC13F585FBDFF978" box="[963,1035,1706,1732]" captionStart="TABLE 1" captionStartId="2.[151,239,151,175]" captionText="TABLE 1. Selected comparative characters observed in genera of Diogenidae closely related to Tetralobistes gen. nov." pageId="1" pageNumber="50">Tab. 1</tableCitation>
). In
<taxonomicName id="4C6C4D789F2EFFFAFB94F584FB0EF978" box="[1092,1242,1707,1732]" class="Malacostraca" family="Diogenidae" genus="Tetralobistes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="50" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B918EAE99F2EFFFAFB94F584FB0EF978" box="[1092,1242,1707,1732]" italics="true" pageId="1" pageNumber="50">Tetralobistes</emphasis>
</taxonomicName>
<emphasis id="B918EAE99F2EFFFAFB32F585FA9BF978" bold="true" box="[1250,1359,1706,1732]" pageId="1" pageNumber="50">
<taxonomicNameLabel id="A22B57929F2EFFFAFB32F585FA9BF978" box="[1250,1359,1706,1732]" pageId="1" pageNumber="50" rank="genus">gen. nov.</taxonomicNameLabel>
</emphasis>
, some of these characters are similar to one, two or all these three other genera, but other characters are clearly distinct or even unique. In
<taxonomicName id="4C6C4D789F2EFFFAFE06F5DAFDB9F8B2" box="[470,621,1781,1806]" class="Malacostraca" family="Diogenidae" genus="Tetralobistes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="50" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B918EAE99F2EFFFAFE06F5DAFDB9F8B2" box="[470,621,1781,1806]" italics="true" pageId="1" pageNumber="50">Tetralobistes</emphasis>
</taxonomicName>
<emphasis id="B918EAE99F2EFFFAFDA6F5DAFD31F8B3" bold="true" box="[630,741,1781,1807]" pageId="1" pageNumber="50">
<taxonomicNameLabel id="A22B57929F2EFFFAFDA6F5DAFD31F8B3" box="[630,741,1781,1807]" pageId="1" pageNumber="50" rank="genus">gen. nov.</taxonomicNameLabel>
</emphasis>
, for example, the presence of 13 pairs of phyllobranchiate biserial gills is unique among these four genera.
</paragraph>
<paragraph id="8BD336FB9F2EFFF9FF16F46FFA48F86E" blockId="1.[151,1437,549,2031]" lastBlockId="2.[151,1437,1714,2002]" lastPageId="2" lastPageNumber="51" pageId="1" pageNumber="50">
In her review of
<taxonomicName id="4C6C4D789F2EFFFAFE74F46FFDF5F8E5" box="[420,545,1856,1881]" class="Malacostraca" family="Diogenidae" genus="Paguristes" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="50" phylum="Arthropoda" rank="genus" sensu="lato">
<emphasis id="B918EAE99F2EFFFAFE74F46FFDF5F8E5" box="[420,545,1856,1881]" italics="true" pageId="1" pageNumber="50">Paguristes</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22B57929F2EFFFAFDFEF46FFD7DF8E6" box="[558,681,1856,1882]" pageId="1" pageNumber="50" sensu="lato">sensu lato</taxonomicNameLabel>
<bibRefCitation id="EFFD4B0A9F2EFFFAFD67F46FFCB8F8E6" author="Rahayu" box="[695,876,1856,1882]" pageId="1" pageNumber="50" refString="Rahayu, D. L. (2005) Additions to the Indonesian fauna of the hermit crab genus Pseudopaguristes McLaughlin and a further division of the genus Paguristes Dana (Crustacea: Decapoda: Paguroidea: Diogenidae). Zootaxa, 831, 1 - 42." type="journal article" year="2005">Rahayu (2005)</bibRefCitation>
considered reduction of the number of gills (branchiae) as an evolutionary trend. On this basis, she suggested splitting
<taxonomicName id="4C6C4D789F2EFFFAFBF0F44AFB74F8C2" box="[1056,1184,1893,1918]" class="Malacostraca" family="Diogenidae" genus="Paguristes" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="50" phylum="Arthropoda" rank="genus" sensu="lato">
<emphasis id="B918EAE99F2EFFFAFBF0F44AFB74F8C2" box="[1056,1184,1893,1918]" italics="true" pageId="1" pageNumber="50">Paguristes</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22B57929F2EFFFAFB79F44AFAF5F8C3" box="[1193,1313,1893,1919]" pageId="1" pageNumber="50" sensu="lato">sensu lato</taxonomicNameLabel>
into three genera:
<taxonomicName id="4C6C4D789F2EFFFAFF26F4A4FEA1F818" box="[246,373,1931,1956]" class="Malacostraca" family="Diogenidae" genus="Paguristes" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="50" phylum="Arthropoda" rank="genus" sensu="stricto">
<emphasis id="B918EAE99F2EFFFAFF26F4A4FEA1F818" box="[246,373,1931,1956]" italics="true" pageId="1" pageNumber="50">Paguristes</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22B57929F2EFFFAFEAEF4A5FDC2F818" box="[382,534,1930,1956]" pageId="1" pageNumber="50" sensu="stricto">sensu stricto</taxonomicNameLabel>
, with 13 pairs of gills,
<taxonomicName id="4C6C4D789F2EFFFAFCFCF4A4FC34F818" box="[812,992,1931,1956]" class="Malacostraca" family="Diogenidae" genus="Areopaguristes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="50" phylum="Arthropoda" rank="genus">
<emphasis id="B918EAE99F2EFFFAFCFCF4A4FC34F818" box="[812,992,1931,1956]" italics="true" pageId="1" pageNumber="50">Areopaguristes</emphasis>
</taxonomicName>
with 12 pairs, and
<taxonomicName id="4C6C4D789F2EFFFAFB19F4A4FA48F818" box="[1225,1436,1931,1956]" class="Malacostraca" family="Diogenidae" genus="Pseudopaguristes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="50" phylum="Arthropoda" rank="genus">
<emphasis id="B918EAE99F2EFFFAFB19F4A4FA48F818" box="[1225,1436,1931,1956]" italics="true" pageId="1" pageNumber="50">Pseudopaguristes</emphasis>
</taxonomicName>
with only 8 pairs. This is also supported by
<bibRefCitation id="EFFD4B0A9F2EFFFAFD78F49FFC67F876" box="[680,947,1968,1994]" pageId="1" pageNumber="50" refString="Lemaitre, R., McLaughlin, P. A. &amp; Sorhannus, U. (2009) Phylogenetic relationships within the Pylochelidae (Decapoda: Anomura: Paguroidea): A cladistic analysis based on morphological characters. Zootaxa, 2022, 1 - 14." type="journal article">
Lemaitre
<emphasis id="B918EAE99F2EFFFAFCCCF49FFC85F875" box="[796,849,1968,1993]" italics="true" pageId="1" pageNumber="50">et al</emphasis>
. (2009)
</bibRefCitation>
, who considered
<taxonomicName id="4C6C4D789F2EFFFAFB55F49FFF00F853" authority="Henderson, 1888" authorityName="Henderson" authorityYear="1888" class="Malacostraca" family="Diogenidae" genus="Paguropsis" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="50" phylum="Arthropoda" rank="genus">
<emphasis id="B918EAE99F2EFFFAFB55F49FFADFF875" box="[1157,1291,1968,1993]" italics="true" pageId="1" pageNumber="50">Paguropsis</emphasis>
<bibRefCitation id="EFFD4B0A9F2EFFFAFAC3F49FFF00F853" author="Henderson" pageId="1" pageNumber="50" refString="Henderson, J. R. (1888) Report on the Anomura collected by H. M. S. Challenger during the years 1873 - 76. Scientific Results of the Exploratory Voyage of HMS Challenger, (Zoology), 27, i - xi, 1 - 221. Her Majesty´s Stationary Office, Edinburgh etc." type="journal article" year="1888">Henderson, 1888</bibRefCitation>
</taxonomicName>
, and
<taxonomicName id="4C6C4D789F2EFFFAFEC1F4FAFE5AF852" box="[273,398,2005,2030]" class="Malacostraca" family="Diogenidae" genus="Paguristes" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="50" phylum="Arthropoda" rank="genus">
<emphasis id="B918EAE99F2EFFFAFEC1F4FAFE5AF852" box="[273,398,2005,2030]" italics="true" pageId="1" pageNumber="50">Paguristes</emphasis>
</taxonomicName>
(both with 13 pairs of gills) among the most primitive of diogenids. Species of these two genera have first and second pairs of males pleopods modified as gonopods, a character considered by
<bibRefCitation id="EFFD4B0A9F2DFFF9FF47F5F7FDFAF94E" author="McLaughlin" box="[151,558,1752,1778]" pageId="2" pageNumber="51" refString="McLaughlin, P. A. &amp; Lemaitre, R. (1997) Carcinization in the Anomura - fact or fiction? I. Evidence from adult morphology. Contributions to Zoology, 67 (29), 79 - 123." type="journal article" year="1997">McLaughlin &amp; Lemaitre (1997)</bibRefCitation>
to be a plesiomorphic condition that evolved into a more advanced, apomorphic condition in which reduction and ultimate loss are eventually observed. The first and second pairs of pleopods (modified as gonopods) are also conserved in
<taxonomicName id="4C6C4D789F2DFFF9FC9FF40CFBD6F880" box="[847,1026,1827,1852]" class="Malacostraca" family="Diogenidae" genus="Areopaguristes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="51" phylum="Arthropoda" rank="genus">
<emphasis id="B918EAE99F2DFFF9FC9FF40CFBD6F880" box="[847,1026,1827,1852]" italics="true" pageId="2" pageNumber="51">Areopaguristes</emphasis>
</taxonomicName>
. However, this is not true in some species of
<taxonomicName id="4C6C4D789F2DFFF9FEC6F467FE3EF8DD" box="[278,490,1864,1889]" class="Malacostraca" family="Diogenidae" genus="Pseudopaguristes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="51" phylum="Arthropoda" rank="genus">
<emphasis id="B918EAE99F2DFFF9FEC6F467FE3EF8DD" box="[278,490,1864,1889]" italics="true" pageId="2" pageNumber="51">Pseudopaguristes</emphasis>
</taxonomicName>
[e.g.,
<emphasis id="B918EAE99F2DFFF9FDE8F467FD47F8DD" box="[568,659,1864,1889]" italics="true" pageId="2" pageNumber="51">P. h i a n s</emphasis>
(
<bibRefCitation id="EFFD4B0A9F2DFFF9FD77F467FCADF8DE" author="Henderson" box="[679,889,1864,1890]" pageId="2" pageNumber="51" refString="Henderson, J. R. (1888) Report on the Anomura collected by H. M. S. Challenger during the years 1873 - 76. Scientific Results of the Exploratory Voyage of HMS Challenger, (Zoology), 27, i - xi, 1 - 221. Her Majesty´s Stationary Office, Edinburgh etc." type="journal article" year="1888">Henderson, 1888</bibRefCitation>
),
<taxonomicName id="4C6C4D789F2DFFF9FC5FF467FB74F8DE" authority="Rahayu, 2005" authorityName="Rahayu" authorityYear="2005" box="[911,1184,1864,1890]" class="Malacostraca" family="Diogenidae" genus="Pseudopaguristes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="51" phylum="Arthropoda" rank="species" species="asper">
<emphasis id="B918EAE99F2DFFF9FC5FF467FC3BF8DD" box="[911,1007,1864,1889]" italics="true" pageId="2" pageNumber="51">P. asper</emphasis>
<bibRefCitation id="EFFD4B0A9F2DFFF9FC28F467FB74F8DE" author="Rahayu" box="[1016,1184,1864,1890]" pageId="2" pageNumber="51" refString="Rahayu, D. L. (2005) Additions to the Indonesian fauna of the hermit crab genus Pseudopaguristes McLaughlin and a further division of the genus Paguristes Dana (Crustacea: Decapoda: Paguroidea: Diogenidae). Zootaxa, 831, 1 - 42." type="journal article" year="2005">Rahayu, 2005</bibRefCitation>
</taxonomicName>
] and in
<taxonomicName id="4C6C4D789F2DFFF9FAD3F467FA48F8DD" box="[1283,1436,1864,1889]" class="Malacostraca" family="Diogenidae" genus="Tetralobistes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="51" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B918EAE99F2DFFF9FAD3F467FA48F8DD" box="[1283,1436,1864,1889]" italics="true" pageId="2" pageNumber="51">Tetralobistes</emphasis>
</taxonomicName>
<emphasis id="B918EAE99F2DFFF9FF47F442FED9F83B" bold="true" box="[151,269,1901,1927]" pageId="2" pageNumber="51">
<taxonomicNameLabel id="A22B57929F2DFFF9FF47F442FED9F83B" box="[151,269,1901,1927]" pageId="2" pageNumber="51" rank="genus">gen. nov.</taxonomicNameLabel>
</emphasis>
<bibRefCitation id="EFFD4B0A9F2DFFF9FEC5F442FE1BF83B" author="Rahayu" box="[277,463,1901,1927]" pageId="2" pageNumber="51" refString="Rahayu, D. L. (2005) Additions to the Indonesian fauna of the hermit crab genus Pseudopaguristes McLaughlin and a further division of the genus Paguristes Dana (Crustacea: Decapoda: Paguroidea: Diogenidae). Zootaxa, 831, 1 - 42." type="journal article" year="2005">Rahayu (2005)</bibRefCitation>
concluded that
<taxonomicName id="4C6C4D789F2DFFF9FD47F442FC87F83A" box="[663,851,1901,1926]" class="Malacostraca" family="Diogenidae" genus="Areopaguristes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="51" phylum="Arthropoda" rank="genus">
<emphasis id="B918EAE99F2DFFF9FD47F442FC87F83A" box="[663,851,1901,1926]" italics="true" pageId="2" pageNumber="51">Areopaguristes</emphasis>
</taxonomicName>
and
<taxonomicName id="4C6C4D789F2DFFF9FC41F442FBB9F83A" box="[913,1133,1901,1926]" class="Malacostraca" family="Diogenidae" genus="Pseudopaguristes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="51" phylum="Arthropoda" rank="genus">
<emphasis id="B918EAE99F2DFFF9FC41F442FBB9F83A" box="[913,1133,1901,1926]" italics="true" pageId="2" pageNumber="51">Pseudopaguristes</emphasis>
</taxonomicName>
(12 and 8 pairs of gills, respectivelly; first and second pair of gonopods usually present in males) are more derived than
<taxonomicName id="4C6C4D789F2DFFF9FACCF4BCFA4FF810" box="[1308,1435,1939,1964]" class="Malacostraca" family="Diogenidae" genus="Paguristes" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="51" phylum="Arthropoda" rank="genus">
<emphasis id="B918EAE99F2DFFF9FACCF4BCFA4FF810" box="[1308,1435,1939,1964]" italics="true" pageId="2" pageNumber="51">Paguristes</emphasis>
</taxonomicName>
and also suggested that a well developed brood pouch, as is found in
<taxonomicName id="4C6C4D789F2DFFF9FC0EF497FB8AF86D" box="[990,1118,1976,2001]" class="Malacostraca" family="Diogenidae" genus="Paguristes" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="51" phylum="Arthropoda" rank="genus">
<emphasis id="B918EAE99F2DFFF9FC0EF497FB8AF86D" box="[990,1118,1976,2001]" italics="true" pageId="2" pageNumber="51">Paguristes</emphasis>
</taxonomicName>
, denotes a primitive state.
</paragraph>
<caption id="DF1366739F2DFFF9FF47F3B8FA59FF13" box="[151,1421,151,175]" pageId="2" pageNumber="51">
<paragraph id="8BD336FB9F2DFFF9FF47F3B8FA59FF13" blockId="2.[151,1421,151,175]" box="[151,1421,151,175]" pageId="2" pageNumber="51">
<emphasis id="B918EAE99F2DFFF9FF47F3B8FEDCFF13" bold="true" box="[151,264,151,175]" pageId="2" pageNumber="51">TABLE 1.</emphasis>
Selected comparative characters observed in genera of
<taxonomicName id="4C6C4D789F2DFFF9FC8BF3B7FC02FF13" box="[859,982,152,175]" class="Malacostraca" family="Diogenidae" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="51" phylum="Arthropoda" rank="family">Diogenidae</taxonomicName>
closely related to
<taxonomicName id="4C6C4D789F2DFFF9FB4AF3B7FAF7FF13" box="[1178,1315,152,175]" class="Malacostraca" family="Diogenidae" genus="Tetralobistes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="51" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B918EAE99F2DFFF9FB4AF3B7FAF7FF13" box="[1178,1315,152,175]" italics="true" pageId="2" pageNumber="51">Tetralobistes</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22B57929F2DFFF9FAF9F3B8FA59FF13" box="[1321,1421,151,175]" pageId="2" pageNumber="51" rank="genus">gen. nov.</taxonomicNameLabel>
</paragraph>
</caption>
<paragraph id="8BD336FB9F2DFFF9FE55F3E5FAD4F9D0" pageId="2" pageNumber="51">
<table id="F96CC45B9F2D0004FF47F3E5FA43F9D0" box="[151,1431,202,1644]" gridcols="5" gridrows="14" pageId="2" pageNumber="51">
<tr id="355C34B99F2D0004FF47F3E5FA43FEBE" box="[151,1431,202,258]" gridrow="0" pageId="2" pageNumber="51" rowspan-0="1">
<th id="768D5DC59F2D0004FE55F3E5FD89FEBE" box="[389,605,202,258]" gridcol="1" gridrow="0" pageId="2" pageNumber="51">
<taxonomicName id="4C6C4D789F2DFFF9FE55F3E5FE6EFEBD" authority="Dana, 1851" authorityName="Dana" authorityYear="1851" class="Malacostraca" family="Diogenidae" genus="Paguristes" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="51" phylum="Arthropoda" rank="genus">
<emphasis id="B918EAE99F2DFFF9FE55F3E5FE23FF5D" box="[389,503,202,225]" italics="true" pageId="2" pageNumber="51">Paguristes</emphasis>
Dana, 1851
</taxonomicName>
</th>
<th id="768D5DC59F2D0004FDA2F3E5FC46FEBE" box="[626,914,202,258]" gridcol="2" gridrow="0" pageId="2" pageNumber="51">
<taxonomicName id="4C6C4D789F2DFFF9FDA2F3E5FCECFEBD" authority="Rahayu &amp; McLaughlin, 2010" authorityName="Rahayu &amp; McLaughlin" authorityYear="2010" class="Malacostraca" family="Diogenidae" genus="Areopaguristes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="51" phylum="Arthropoda" rank="genus">
<emphasis id="B918EAE99F2DFFF9FDA2F3E5FCC0FF5D" box="[626,788,202,225]" italics="true" pageId="2" pageNumber="51">Areopaguristes</emphasis>
Rahayu &amp; McLaughlin, 2010
</taxonomicName>
</th>
<th id="768D5DC59F2D0004FC73F3E5FB4BFEBE" box="[931,1183,202,258]" gridcol="3" gridrow="0" pageId="2" pageNumber="51">
<taxonomicName id="4C6C4D789F2DFFF9FC73F3E5FBBDFEBD" authority="McLaughlin, 2002" authorityName="McLaughlin" authorityYear="2002" box="[931,1129,202,257]" class="Malacostraca" family="Diogenidae" genus="Pseudopaguristes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="51" phylum="Arthropoda" rank="genus">
<emphasis id="B918EAE99F2DFFF9FC73F3E5FBB5FF5D" box="[931,1121,202,225]" italics="true" pageId="2" pageNumber="51">Pseudopaguristes</emphasis>
McLaughlin, 2002
</taxonomicName>
</th>
<th id="768D5DC59F2D0004FB63F3E5FA43FEBE" box="[1203,1431,202,258]" gridcol="4" gridrow="0" pageId="2" pageNumber="51">
<taxonomicName id="4C6C4D789F2DFFF9FB63F3E5FAE8FF5D" box="[1203,1340,202,225]" class="Malacostraca" family="Diogenidae" genus="Tetralobistes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="51" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B918EAE99F2DFFF9FB63F3E5FAE8FF5D" box="[1203,1340,202,225]" italics="true" pageId="2" pageNumber="51">Tetralobistes</emphasis>
</taxonomicName>
<emphasis id="B918EAE99F2DFFF9FA92F3E5FB36FEBE" bold="true" pageId="2" pageNumber="51">
<taxonomicNameLabel id="A22B57929F2DFFF9FA92F3E5FB36FEBE" pageId="2" pageNumber="51" rank="genus">gen. nov.</taxonomicNameLabel>
</emphasis>
</th>
</tr>
<tr id="355C34B99F2D0004FF47F23AFA43FE90" box="[151,1431,277,300]" gridrow="1" pageId="2" pageNumber="51">
<th id="768D5DC59F2D0004FF47F23AFEBAFE90" box="[151,366,277,300]" gridcol="0" gridrow="1" pageId="2" pageNumber="51">Number of gills</th>
<td id="768D5DC59F2D0004FE55F23AFD89FE90" box="[389,605,277,300]" gridcol="1" gridrow="1" pageId="2" pageNumber="51">13</td>
<td id="768D5DC59F2D0004FDA2F23AFC46FE90" box="[626,914,277,300]" gridcol="2" gridrow="1" pageId="2" pageNumber="51">12</td>
<td id="768D5DC59F2D0004FC73F23AFB4BFE90" box="[931,1183,277,300]" gridcol="3" gridrow="1" pageId="2" pageNumber="51">8</td>
<td id="768D5DC59F2D0004FB63F23AFA43FE90" box="[1203,1431,277,300]" gridcol="4" gridrow="1" pageId="2" pageNumber="51">13</td>
</tr>
<tr id="355C34B99F2D0004FF47F26FFA43FECB" box="[151,1431,320,375]" gridrow="2" pageId="2" pageNumber="51">
<th id="768D5DC59F2D0004FF47F26FFEBAFECB" box="[151,366,320,375]" gridcol="0" gridrow="2" pageId="2" pageNumber="51">Type of gills</th>
<td id="768D5DC59F2D0004FE55F26FFD89FECB" box="[389,605,320,375]" gridcol="1" gridrow="2" pageId="2" pageNumber="51">Phyllobranchiate; bi or quadriserial</td>
<td id="768D5DC59F2D0004FDA2F26FFC46FECB" box="[626,914,320,375]" gridcol="2" gridrow="2" pageId="2" pageNumber="51">Phyllobranchiate; bi or quadriserial</td>
<td id="768D5DC59F2D0004FC73F26FFB4BFECB" box="[931,1183,320,375]" gridcol="3" gridrow="2" pageId="2" pageNumber="51">Phyllobranchiate; bi or quadriserial</td>
<td id="768D5DC59F2D0004FB63F26FFA43FECB" box="[1203,1431,320,375]" gridcol="4" gridrow="2" pageId="2" pageNumber="51">Phyllobranchiate; biserial</td>
</tr>
<tr id="355C34B99F2D0004FF47F2A5FA43FDBD" box="[151,1431,394,513]" gridrow="3" pageId="2" pageNumber="51">
<th id="768D5DC59F2D0004FF47F2A5FEBAFDBD" box="[151,366,394,513]" gridcol="0" gridrow="3" pageId="2" pageNumber="51">Ocular acicles</th>
<td id="768D5DC59F2D0004FE55F2A5FD89FDBD" box="[389,605,394,513]" gridcol="1" gridrow="3" pageId="2" pageNumber="51">Well developed, often multispinous, more or less separated</td>
<td id="768D5DC59F2D0004FDA2F2A5FC46FDBD" box="[626,914,394,513]" gridcol="2" gridrow="3" pageId="2" pageNumber="51">Well developed or reduced, simple, bi or multidenticulate, conti- guous or widely separated</td>
<td id="768D5DC59F2D0004FC73F2A5FB4BFDBD" box="[931,1183,394,513]" gridcol="3" gridrow="3" pageId="2" pageNumber="51">Well developed, simple, bi or multidenticulate, widely separated</td>
<td id="768D5DC59F2D0004FB63F2A5FA43FDBD" box="[1203,1431,394,513]" gridcol="4" gridrow="3" pageId="2" pageNumber="51">Well developed, multidenticulate, contiguous</td>
</tr>
<tr id="355C34B99F2D0004FF47F13AFA43FDD0" box="[151,1431,533,620]" gridrow="4" pageId="2" pageNumber="51">
<th id="768D5DC59F2D0004FF47F13AFEBAFDD0" box="[151,366,533,620]" gridcol="0" gridrow="4" pageId="2" pageNumber="51">Maxillules</th>
<td id="768D5DC59F2D0004FE55F13AFD89FDD0" box="[389,605,533,620]" gridcol="1" gridrow="4" pageId="2" pageNumber="51">External lobe of endopod well developed, recurved</td>
<td id="768D5DC59F2D0004FDA2F13AFC46FDD0" box="[626,914,533,620]" gridcol="2" gridrow="4" pageId="2" pageNumber="51">External lobe of endopod prominently recurved</td>
<td id="768D5DC59F2D0004FC73F13AFB4BFDD0" box="[931,1183,533,620]" gridcol="3" gridrow="4" pageId="2" pageNumber="51">External lobe of endopod well developed, recurved</td>
<td id="768D5DC59F2D0004FB63F13AFA43FDD0" box="[1203,1431,533,620]" gridcol="4" gridrow="4" pageId="2" pageNumber="51">External lobe of endopod well developed, recurved</td>
</tr>
<tr id="355C34B99F2D0004FF47F1AFFA43FD0B" box="[151,1431,640,695]" gridrow="5" pageId="2" pageNumber="51">
<th id="768D5DC59F2D0004FF47F1AFFEBAFD0B" box="[151,366,640,695]" gridcol="0" gridrow="5" pageId="2" pageNumber="51">First maxillipeds</th>
<td id="768D5DC59F2D0004FE55F1AFFD89FD0B" box="[389,605,640,695]" gridcol="1" gridrow="5" pageId="2" pageNumber="51">Epipod well developed</td>
<td id="768D5DC59F2D0004FDA2F1AFFC46FD0B" box="[626,914,640,695]" gridcol="2" gridrow="5" pageId="2" pageNumber="51">Epipod well developed or absent</td>
<td id="768D5DC59F2D0004FC73F1AFFB4BFD0B" box="[931,1183,640,695]" gridcol="3" gridrow="5" pageId="2" pageNumber="51">Epipod well developed</td>
<td id="768D5DC59F2D0004FB63F1AFFA43FD0B" box="[1203,1431,640,695]" gridcol="4" gridrow="5" pageId="2" pageNumber="51">Epipod well developed</td>
</tr>
<tr id="355C34B99F2D0004FF47F1E5FA43FCFD" box="[151,1431,714,833]" gridrow="6" pageId="2" pageNumber="51">
<th id="768D5DC59F2D0004FF47F1E5FEBAFCFD" box="[151,366,714,833]" gridcol="0" gridrow="6" pageId="2" pageNumber="51">Third maxillipeds</th>
<td id="768D5DC59F2D0004FE55F1E5FD89FCFD" box="[389,605,714,833]" gridcol="1" gridrow="6" pageId="2" pageNumber="51">Crista dentata well developed, no accesory tooth</td>
<td id="768D5DC59F2D0004FDA2F1E5FC46FCFD" box="[626,914,714,833]" gridcol="2" gridrow="6" pageId="2" pageNumber="51">Crista dentata well developed, no accesory tooth</td>
<td id="768D5DC59F2D0004FC73F1E5FB4BFCFD" box="[931,1183,714,833]" gridcol="3" gridrow="6" pageId="2" pageNumber="51">Crista dentata consisting of row of quite small teeth, no accesory tooth</td>
<td id="768D5DC59F2D0004FB63F1E5FA43FCFD" box="[1203,1431,714,833]" gridcol="4" gridrow="6" pageId="2" pageNumber="51">Crista dentata well developed, no accesory tooth</td>
</tr>
<tr id="355C34B99F2D0004FF47F07AFA43FC70" box="[151,1431,853,972]" gridrow="7" pageId="2" pageNumber="51">
<th id="768D5DC59F2D0004FF47F07AFEBAFC70" box="[151,366,853,972]" gridcol="0" gridrow="7" pageId="2" pageNumber="51">Chelipeds</th>
<td id="768D5DC59F2D0004FE55F07AFD89FC70" box="[389,605,853,972]" gridcol="1" gridrow="7" pageId="2" pageNumber="51">Usually subequal; claws with fingers opening in horizontal plane</td>
<td id="768D5DC59F2D0004FDA2F07AFC46FC70" box="[626,914,853,972]" gridcol="2" gridrow="7" pageId="2" pageNumber="51">Equal, subequal or unequal; claws with fingers opening in horizontal or oblique plane</td>
<td id="768D5DC59F2D0004FC73F07AFB4BFC70" box="[931,1183,853,972]" gridcol="3" gridrow="7" pageId="2" pageNumber="51">Subequal or unequal; claws with fingers opening in horizontal or oblique plane</td>
<td id="768D5DC59F2D0004FB63F07AFA43FC70" box="[1203,1431,853,972]" gridcol="4" gridrow="7" pageId="2" pageNumber="51">Subequal; claws with fingers opening in horizontal plane</td>
</tr>
<tr id="355C34B99F2D0004FF47F0CFFA43FBEB" box="[151,1431,992,1111]" gridrow="8" pageId="2" pageNumber="51">
<th id="768D5DC59F2D0004FF47F0CFFEBAFBEB" box="[151,366,992,1111]" gridcol="0" gridrow="8" pageId="2" pageNumber="51">Fourth pereopods</th>
<td id="768D5DC59F2D0004FE55F0CFFD89FBEB" box="[389,605,992,1111]" gridcol="1" gridrow="8" pageId="2" pageNumber="51">Simple, with or without preungual process at base of claw</td>
<td id="768D5DC59F2D0004FDA2F0CFFC46FBEB" box="[626,914,992,1111]" gridcol="2" gridrow="8" pageId="2" pageNumber="51">Subchelate or weakly semichelate, with or without preungual process at base of claw</td>
<td id="768D5DC59F2D0004FC73F0CFFB4BFBEB" box="[931,1183,992,1111]" gridcol="3" gridrow="8" pageId="2" pageNumber="51">Simple, with or without preungual process at base of claw</td>
<td id="768D5DC59F2D0004FB63F0CFFA43FBEB" box="[1203,1431,992,1111]" gridcol="4" gridrow="8" pageId="2" pageNumber="51">Simple, without preungual process at base of claw</td>
</tr>
<tr id="355C34B99F2D0004FF47F745FA43FB1D" box="[151,1431,1130,1185]" gridrow="9" pageId="2" pageNumber="51">
<th id="768D5DC59F2D0004FF47F745FEBAFB1D" box="[151,366,1130,1185]" gridcol="0" gridrow="9" pageId="2" pageNumber="51">Male first pleopods</th>
<td id="768D5DC59F2D0004FE55F745FD89FB1D" box="[389,605,1130,1185]" gridcol="1" gridrow="9" pageId="2" pageNumber="51">Modified as gonopod</td>
<td id="768D5DC59F2D0004FDA2F745FC46FB1D" box="[626,914,1130,1185]" gridcol="2" gridrow="9" pageId="2" pageNumber="51">Modified as gonopod</td>
<td id="768D5DC59F2D0004FC73F745FB4BFB1D" box="[931,1183,1130,1185]" gridcol="3" gridrow="9" pageId="2" pageNumber="51">Modified as gonopod</td>
<td id="768D5DC59F2D0004FB63F745FA43FB1D" box="[1203,1431,1130,1185]" gridcol="4" gridrow="9" pageId="2" pageNumber="51">Modified as gonopod</td>
</tr>
<tr id="355C34B99F2D0004FF47F79AFA43FB50" box="[151,1431,1205,1260]" gridrow="10" pageId="2" pageNumber="51">
<th id="768D5DC59F2D0004FF47F79AFEBAFB50" box="[151,366,1205,1260]" gridcol="0" gridrow="10" pageId="2" pageNumber="51">Male second pleopods</th>
<td id="768D5DC59F2D0004FE55F79AFD89FB50" box="[389,605,1205,1260]" gridcol="1" gridrow="10" pageId="2" pageNumber="51">Modified as gonopod</td>
<td id="768D5DC59F2D0004FDA2F79AFC46FB50" box="[626,914,1205,1260]" gridcol="2" gridrow="10" pageId="2" pageNumber="51">Modified as gonopod</td>
<td id="768D5DC59F2D0004FC73F79AFB4BFB50" box="[931,1183,1205,1260]" gridcol="3" gridrow="10" pageId="2" pageNumber="51">Modified as gonopod or absent</td>
<td id="768D5DC59F2D0004FB63F79AFA43FB50" box="[1203,1431,1205,1260]" gridcol="4" gridrow="10" pageId="2" pageNumber="51">Absent</td>
</tr>
<tr id="355C34B99F2D0004FF47F62FFA43FA8B" box="[151,1431,1280,1335]" gridrow="11" pageId="2" pageNumber="51">
<th id="768D5DC59F2D0004FF47F62FFEBAFA8B" box="[151,366,1280,1335]" gridcol="0" gridrow="11" pageId="2" pageNumber="51">Female first pleopods</th>
<td id="768D5DC59F2D0004FE55F62FFD89FA8B" box="[389,605,1280,1335]" gridcol="1" gridrow="11" pageId="2" pageNumber="51">Modified as gonopod</td>
<td id="768D5DC59F2D0004FDA2F62FFC46FA8B" box="[626,914,1280,1335]" gridcol="2" gridrow="11" pageId="2" pageNumber="51">Modified as gonopod or absent</td>
<td id="768D5DC59F2D0004FC73F62FFB4BFA8B" box="[931,1183,1280,1335]" gridcol="3" gridrow="11" pageId="2" pageNumber="51">Modified as gonopod</td>
<td id="768D5DC59F2D0004FB63F62FFA43FA8B" box="[1203,1431,1280,1335]" gridcol="4" gridrow="11" pageId="2" pageNumber="51">Absent</td>
</tr>
<tr id="355C34B99F2D0004FF47F665FA43FA1D" box="[151,1431,1354,1441]" gridrow="12" pageId="2" pageNumber="51">
<th id="768D5DC59F2D0004FF47F665FEBAFA1D" box="[151,366,1354,1441]" gridcol="0" gridrow="12" pageId="2" pageNumber="51">Brood pouch</th>
<td id="768D5DC59F2D0004FE55F665FD89FA1D" box="[389,605,1354,1441]" gridcol="1" gridrow="12" pageId="2" pageNumber="51">Usually well developed or absent</td>
<td id="768D5DC59F2D0004FDA2F665FC46FA1D" box="[626,914,1354,1441]" gridcol="2" gridrow="12" pageId="2" pageNumber="51">Usually well developed, occasionally entirely absent</td>
<td id="768D5DC59F2D0004FC73F665FB4BFA1D" box="[931,1183,1354,1441]" gridcol="3" gridrow="12" pageId="2" pageNumber="51">Present or absent</td>
<td id="768D5DC59F2D0004FB63F665FA43FA1D" box="[1203,1431,1354,1441]" gridcol="4" gridrow="12" pageId="2" pageNumber="51">Absent</td>
</tr>
<tr id="355C34B99F2D0004FF47F69AFA43F9D0" box="[151,1431,1461,1644]" gridrow="13" pageId="2" pageNumber="51">
<th id="768D5DC59F2D0004FF47F69AFEBAF9D0" box="[151,366,1461,1644]" gridcol="0" gridrow="13" pageId="2" pageNumber="51">Telson</th>
<td id="768D5DC59F2D0004FE55F69AFD89F9D0" box="[389,605,1461,1644]" gridcol="1" gridrow="13" pageId="2" pageNumber="51">Posterior margin divided into two subequal to markedly unequal lobes</td>
<td id="768D5DC59F2D0004FDA2F69AFC46F9D0" box="[626,914,1461,1644]" gridcol="2" gridrow="13" pageId="2" pageNumber="51">Posterior margin divided into two subequal to markedly unequal lobes</td>
<td id="768D5DC59F2D0004FC73F69AFB4BF9D0" box="[931,1183,1461,1644]" gridcol="3" gridrow="13" pageId="2" pageNumber="51">Posterior margin divided into two slightly to markedly asymmetrical lobes</td>
<td id="768D5DC59F2D0004FB63F69AFA43F9D0" box="[1203,1431,1461,1644]" gridcol="4" gridrow="13" pageId="2" pageNumber="51">Posterior margin tetralobular; external lobules unequal, larger than symmetrical, inner lobules</td>
</tr>
</table>
</paragraph>
<paragraph id="8BD336FB9F2CFFF8FF47F3B7FC7CFDF0" blockId="3.[151,1437,151,1484]" pageId="3" pageNumber="52">
Although in
<taxonomicName id="4C6C4D789F2CFFF8FEFFF3B7FE1CFF0D" box="[303,456,152,177]" class="Malacostraca" family="Diogenidae" genus="Tetralobistes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="52" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B918EAE99F2CFFF8FEFFF3B7FE1CFF0D" box="[303,456,152,177]" italics="true" pageId="3" pageNumber="52">Tetralobistes</emphasis>
</taxonomicName>
<emphasis id="B918EAE99F2CFFF8FE00F3B8FD94FF0D" bold="true" box="[464,576,151,177]" pageId="3" pageNumber="52">
<taxonomicNameLabel id="A22B57929F2CFFF8FE00F3B8FD94FF0D" box="[464,576,151,177]" pageId="3" pageNumber="52" rank="genus">gen. nov.</taxonomicNameLabel>
</emphasis>
the absence of the female brood pouch (reduced to a tufts of setae) is a distinctive character, the latter can be occasionally absent or also reduced to tufts of setae in some species of
<taxonomicName id="4C6C4D789F2CFFF8FF47F3CDFE86FF47" box="[151,338,226,251]" class="Malacostraca" family="Diogenidae" genus="Areopaguristes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="52" phylum="Arthropoda" rank="genus">
<emphasis id="B918EAE99F2CFFF8FF47F3CDFE86FF47" box="[151,338,226,251]" italics="true" pageId="3" pageNumber="52">Areopaguristes</emphasis>
</taxonomicName>
[e.g.,
<taxonomicName id="4C6C4D789F2CFFF8FE72F3CDFCF3FF40" authority="Rahayu, 2005" authorityName="Rahayu" authorityYear="2005" box="[418,807,226,252]" class="Malacostraca" family="Diogenidae" genus="Areopaguristes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="52" phylum="Arthropoda" rank="species" species="tuberculatus">
<emphasis id="B918EAE99F2CFFF8FE72F3CDFD8BFF47" box="[418,607,226,251]" italics="true" pageId="3" pageNumber="52">A. tuberculatus</emphasis>
(
<bibRefCitation id="EFFD4B0A9F2CFFF8FDA0F3CDFCCBFF40" author="Rahayu" box="[624,799,226,252]" pageId="3" pageNumber="52" refString="Rahayu, D. L. (2005) Additions to the Indonesian fauna of the hermit crab genus Pseudopaguristes McLaughlin and a further division of the genus Paguristes Dana (Crustacea: Decapoda: Paguroidea: Diogenidae). Zootaxa, 831, 1 - 42." type="journal article" year="2005">Rahayu, 2005</bibRefCitation>
)
</taxonomicName>
and
<taxonomicName id="4C6C4D789F2CFFF8FCB5F3CDFF37FE9E" authority="Ayon-Parente &amp; Hendrickx, 2006" authorityName="Ayon-Parente &amp; Hendrickx" authorityYear="2006" class="Malacostraca" family="Diogenidae" genus="Areopaguristes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="52" phylum="Arthropoda" rank="species" species="mclaughlinae">
<emphasis id="B918EAE99F2CFFF8FCB5F3CDFBFBFF47" box="[869,1071,226,251]" italics="true" pageId="3" pageNumber="52">A. mclaughlinae</emphasis>
(
<bibRefCitation id="EFFD4B0A9F2CFFF8FB90F3CDFF0FFE9E" author="Ayon-Parente" pageId="3" pageNumber="52" refString="Ayon-Parente, M. &amp; Hendrickx, M. E. (2006) A new species of Stratiotes Thomson, 1899 (Anomura, Paguroidea, Diogenidae) from the eastern tropical Pacific. Zoosystema, 28 (2), 487 - 494." type="journal article" year="2006">Ayón-Parente &amp; Hendrickx, 2006</bibRefCitation>
)
</taxonomicName>
],
<taxonomicName id="4C6C4D789F2CFFF8FF28F227FEAEFE9D" box="[248,378,264,289]" class="Malacostraca" family="Diogenidae" genus="Paguristes" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="52" phylum="Arthropoda" rank="genus">
<emphasis id="B918EAE99F2CFFF8FF28F227FEAEFE9D" box="[248,378,264,289]" italics="true" pageId="3" pageNumber="52">Paguristes</emphasis>
</taxonomicName>
(e.g.,
<taxonomicName id="4C6C4D789F2CFFF8FE19F227FCC7FE9E" authority="Bouvier, 1918" authorityName="Bouvier" authorityYear="1918" box="[457,787,264,290]" class="Malacostraca" family="Diogenidae" genus="Paguristes" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="52" phylum="Arthropoda" rank="species" species="anomalus">
<emphasis id="B918EAE99F2CFFF8FE19F227FD89FE9D" box="[457,605,264,289]" italics="true" pageId="3" pageNumber="52">P. anomalus</emphasis>
<bibRefCitation id="EFFD4B0A9F2CFFF8FDB6F227FCC7FE9E" author="Bouvier" box="[614,787,264,290]" pageId="3" pageNumber="52" refString="Bouvier, E. L. (1918) Sur une petite collection de Crustaces de Cuba offerte au Museum par M. de Boury. Bulletin du Museum national d'Histoire naturelle, 24, 6 - 15." type="journal article" year="1918">Bouvier, 1918</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="4C6C4D789F2CFFF8FC80F227FB85FE9E" authority="Forest, 1954" authorityName="Forest" authorityYear="1954" box="[848,1105,264,290]" class="Malacostraca" family="Diogenidae" genus="Paguristes" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="52" phylum="Arthropoda" rank="species" species="fagei">
<emphasis id="B918EAE99F2CFFF8FC80F227FC7FFE9D" box="[848,939,264,289]" italics="true" pageId="3" pageNumber="52">P. fagei</emphasis>
<bibRefCitation id="EFFD4B0A9F2CFFF8FC64F227FB85FE9E" author="Forest" box="[948,1105,264,290]" pageId="3" pageNumber="52" refString="Forest, J. (1954) Les Paguristes des cotes occidentales et meridionales d'Afrique. Annals of the South African Museum, 41, 159 - 213." type="journal article" year="1954">Forest, 1954</bibRefCitation>
</taxonomicName>
), and
<taxonomicName id="4C6C4D789F2CFFF8FB4CF227FAA0FE9D" box="[1180,1396,264,289]" class="Malacostraca" family="Diogenidae" genus="Pseudopaguristes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="52" phylum="Arthropoda" rank="genus">
<emphasis id="B918EAE99F2CFFF8FB4CF227FAA0FE9D" box="[1180,1396,264,289]" italics="true" pageId="3" pageNumber="52">Pseudopaguristes</emphasis>
</taxonomicName>
[
<taxonomicName id="4C6C4D789F2CFFF8FA56F227FE19FEFB" authority="Rahayu, 2005" authorityName="Rahayu" authorityYear="2005" class="Malacostraca" family="Diogenidae" genus="Pseudopaguristes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="52" phylum="Arthropoda" rank="species" species="janetkae">
<emphasis id="B918EAE99F2CFFF8FA56F227FED4FEFA" italics="true" pageId="3" pageNumber="52">P. janetkae</emphasis>
(
<bibRefCitation id="EFFD4B0A9F2CFFF8FEC1F202FE10FEFB" author="Rahayu" box="[273,452,301,327]" pageId="3" pageNumber="52" refString="Rahayu, D. L. (2005) Additions to the Indonesian fauna of the hermit crab genus Pseudopaguristes McLaughlin and a further division of the genus Paguristes Dana (Crustacea: Decapoda: Paguroidea: Diogenidae). Zootaxa, 831, 1 - 42." type="journal article" year="2005">Rahayu, 2005</bibRefCitation>
)
</taxonomicName>
] (see also
<bibRefCitation id="EFFD4B0A9F2CFFF8FD82F202FCE0FEFB" author="McLaughlin" box="[594,820,301,327]" pageId="3" pageNumber="52" refString="McLaughlin, P. A. (2002) Pseudopaguristes, a new and aberrant genus of hermit crabs (Anomura: Paguridea: Diogenidae). Micronesica, 34 (2), 185 - 199." type="journal article" year="2002">McLaughlin 2002</bibRefCitation>
,
<bibRefCitation id="EFFD4B0A9F2CFFF8FC92F202FC3FFEFB" author="Rahayu" box="[834,1003,301,327]" pageId="3" pageNumber="52" refString="Rahayu, D. L. (2005) Additions to the Indonesian fauna of the hermit crab genus Pseudopaguristes McLaughlin and a further division of the genus Paguristes Dana (Crustacea: Decapoda: Paguroidea: Diogenidae). Zootaxa, 831, 1 - 42." type="journal article" year="2005">Rahayu 2005</bibRefCitation>
). Female first pleopods are always present and modified as gonopods in
<taxonomicName id="4C6C4D789F2CFFF8FD9EF27DFD1FFED7" box="[590,715,338,363]" class="Malacostraca" family="Diogenidae" genus="Paguristes" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="52" phylum="Arthropoda" rank="genus">
<emphasis id="B918EAE99F2CFFF8FD9EF27DFD1FFED7" box="[590,715,338,363]" italics="true" pageId="3" pageNumber="52">Paguristes</emphasis>
</taxonomicName>
and
<taxonomicName id="4C6C4D789F2CFFF8FCD5F27DFC01FED7" box="[773,981,338,363]" class="Malacostraca" family="Diogenidae" genus="Pseudopaguristes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="52" phylum="Arthropoda" rank="genus">
<emphasis id="B918EAE99F2CFFF8FCD5F27DFC01FED7" box="[773,981,338,363]" italics="true" pageId="3" pageNumber="52">Pseudopaguristes</emphasis>
</taxonomicName>
, but in
<taxonomicName id="4C6C4D789F2CFFF8FBE0F27DFB36FED7" box="[1072,1250,338,363]" class="Malacostraca" family="Diogenidae" genus="Areopaguristes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="52" phylum="Arthropoda" rank="genus">
<emphasis id="B918EAE99F2CFFF8FBE0F27DFB36FED7" box="[1072,1250,338,363]" italics="true" pageId="3" pageNumber="52">Areopaguristes</emphasis>
</taxonomicName>
these are either present or absent [e.g.,
<taxonomicName id="4C6C4D789F2CFFF8FE7AF257FC82FE2E" authority="Wass, 1955" authorityName="Wass" authorityYear="1955" box="[426,854,376,402]" class="Malacostraca" family="Diogenidae" genus="Areopaguristes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="52" phylum="Arthropoda" rank="species" species="hummi">
<emphasis id="B918EAE99F2CFFF8FE7AF257FD60FE2D" box="[426,692,376,401]" italics="true" pageId="3" pageNumber="52">Areopaguristes hummi</emphasis>
(
<bibRefCitation id="EFFD4B0A9F2CFFF8FD14F257FC9AFE2E" author="Wass" box="[708,846,376,402]" pageId="3" pageNumber="52" refString="Wass, M. L. (1955) The decapod crustaceans of Alligator Harbor and adjacent inshore areas of northwestern Florida. The Quarterly Journal of the Florida Academy of Sciences, 18 (3), 129 - 176." type="journal article" year="1955">Wass, 1955</bibRefCitation>
)
</taxonomicName>
and
<taxonomicName id="4C6C4D789F2CFFF8FC41F257FE56FE0B" authority="Ayon-Parente &amp; Hendrickx 2006" authorityName="Ayon-Parente &amp; Hendrickx" authorityYear="2006" class="Malacostraca" family="Diogenidae" genus="Areopaguristes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="52" phylum="Arthropoda" rank="species" species="mclaughlinae">
<emphasis id="B918EAE99F2CFFF8FC41F257FB3EFE2D" box="[913,1258,376,401]" italics="true" pageId="3" pageNumber="52">Areopaguristes mclaughlinae</emphasis>
(
<bibRefCitation id="EFFD4B0A9F2CFFF8FB29F257FEADFE0B" author="Ayon-Parente" pageId="3" pageNumber="52" refString="Ayon-Parente, M. &amp; Hendrickx, M. E. (2006) A new species of Stratiotes Thomson, 1899 (Anomura, Paguroidea, Diogenidae) from the eastern tropical Pacific. Zoosystema, 28 (2), 487 - 494." type="journal article" year="2006">Ayón-Parente &amp; Hendrickx 2006</bibRefCitation>
)
</taxonomicName>
], as in
<taxonomicName id="4C6C4D789F2CFFF8FE02F2B2FDBDFE0A" box="[466,617,413,438]" class="Malacostraca" family="Diogenidae" genus="Tetralobistes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="52" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B918EAE99F2CFFF8FE02F2B2FDBDFE0A" box="[466,617,413,438]" italics="true" pageId="3" pageNumber="52">Tetralobistes</emphasis>
</taxonomicName>
<emphasis id="B918EAE99F2CFFF8FDA0F2B2FD09FE0B" bold="true" box="[624,733,413,439]" pageId="3" pageNumber="52">
<taxonomicNameLabel id="A22B57929F2CFFF8FDA0F2B2FD09FE0B" box="[624,733,413,439]" pageId="3" pageNumber="52" rank="genus">gen. nov.</taxonomicNameLabel>
</emphasis>
With 13 pairs of gills,
<taxonomicName id="4C6C4D789F2CFFF8FC3BF2B2FB55FE0A" box="[1003,1153,413,438]" class="Malacostraca" family="Diogenidae" genus="Tetralobistes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="52" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B918EAE99F2CFFF8FC3BF2B2FB55FE0A" box="[1003,1153,413,438]" italics="true" pageId="3" pageNumber="52">Tetralobistes</emphasis>
</taxonomicName>
<emphasis id="B918EAE99F2CFFF8FB58F2B2FB22FE0B" bold="true" box="[1160,1270,413,439]" pageId="3" pageNumber="52">
<taxonomicNameLabel id="A22B57929F2CFFF8FB58F2B2FB22FE0B" box="[1160,1270,413,439]" pageId="3" pageNumber="52" rank="genus">gen. nov.</taxonomicNameLabel>
</emphasis>
would appear to be more primitive than
<taxonomicName id="4C6C4D789F2CFFF8FE0FF2EDFD4FFE67" box="[479,667,450,475]" class="Malacostraca" family="Diogenidae" genus="Areopaguristes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="52" phylum="Arthropoda" rank="genus">
<emphasis id="B918EAE99F2CFFF8FE0FF2EDFD4FFE67" box="[479,667,450,475]" italics="true" pageId="3" pageNumber="52">Areopaguristes</emphasis>
</taxonomicName>
and
<taxonomicName id="4C6C4D789F2CFFF8FD09F2EDFC61FE67" box="[729,949,450,475]" class="Malacostraca" family="Diogenidae" genus="Pseudopaguristes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="52" phylum="Arthropoda" rank="genus">
<emphasis id="B918EAE99F2CFFF8FD09F2EDFC61FE67" box="[729,949,450,475]" italics="true" pageId="3" pageNumber="52">Pseudopaguristes</emphasis>
</taxonomicName>
, but more derived than
<taxonomicName id="4C6C4D789F2CFFF8FB33F2EDFABCFE67" box="[1251,1384,450,475]" class="Malacostraca" family="Diogenidae" genus="Paguristes" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="52" phylum="Arthropoda" rank="genus">
<emphasis id="B918EAE99F2CFFF8FB33F2EDFABCFE67" box="[1251,1384,450,475]" italics="true" pageId="3" pageNumber="52">Paguristes</emphasis>
</taxonomicName>
and
<taxonomicName id="4C6C4D789F2CFFF8FF47F2C7FEF4FDBD" box="[151,288,488,513]" class="Malacostraca" family="Diogenidae" genus="Paguropsis" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="52" phylum="Arthropoda" rank="genus">
<emphasis id="B918EAE99F2CFFF8FF47F2C7FEF4FDBD" box="[151,288,488,513]" italics="true" pageId="3" pageNumber="52">Paguropsis</emphasis>
</taxonomicName>
. This would also be supported by the loss of the males’ second pair and females’ first pair of pleopods, and of the brood pouch in
<taxonomicName id="4C6C4D789F2CFFF8FD84F122FD3AFD9A" box="[596,750,525,550]" class="Malacostraca" family="Diogenidae" genus="Tetralobistes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="52" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B918EAE99F2CFFF8FD84F122FD3AFD9A" box="[596,750,525,550]" italics="true" pageId="3" pageNumber="52">Tetralobistes</emphasis>
</taxonomicName>
<emphasis id="B918EAE99F2CFFF8FD26F122FCB3FD9B" bold="true" box="[758,871,525,551]" pageId="3" pageNumber="52">
<taxonomicNameLabel id="A22B57929F2CFFF8FD26F122FCB3FD9B" box="[758,871,525,551]" pageId="3" pageNumber="52" rank="genus">gen. nov.</taxonomicNameLabel>
</emphasis>
. These characters would set
<taxonomicName id="4C6C4D789F2CFFF8FB15F122FA8BFD9A" box="[1221,1375,525,550]" class="Malacostraca" family="Diogenidae" genus="Tetralobistes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="52" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B918EAE99F2CFFF8FB15F122FA8BFD9A" box="[1221,1375,525,550]" italics="true" pageId="3" pageNumber="52">Tetralobistes</emphasis>
</taxonomicName>
<emphasis id="B918EAE99F2CFFF8FAB8F122FF1EFDF0" bold="true" pageId="3" pageNumber="52">
<taxonomicNameLabel id="A22B57929F2CFFF8FAB8F122FF1EFDF0" pageId="3" pageNumber="52" rank="genus">gen. nov.</taxonomicNameLabel>
</emphasis>
in an intermediate position between these two pairs of genera.
</paragraph>
<paragraph id="8BD336FB9F2CFFF8FF16F177FB1CFD00" blockId="3.[151,1437,151,1484]" pageId="3" pageNumber="52">
The size, armature and distance separating the ocular acicles in the hermit crabs had been considered a generic diagnostic character. In
<taxonomicName id="4C6C4D789F2CFFF8FDC4F152FD78FD2A" box="[532,684,637,662]" class="Malacostraca" family="Diogenidae" genus="Tetralobistes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="52" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B918EAE99F2CFFF8FDC4F152FD78FD2A" box="[532,684,637,662]" italics="true" pageId="3" pageNumber="52">Tetralobistes</emphasis>
</taxonomicName>
<emphasis id="B918EAE99F2CFFF8FD65F152FCF1FD2B" bold="true" box="[693,805,637,663]" pageId="3" pageNumber="52">
<taxonomicNameLabel id="A22B57929F2CFFF8FD65F152FCF1FD2B" box="[693,805,637,663]" pageId="3" pageNumber="52" rank="genus">gen. nov.</taxonomicNameLabel>
</emphasis>
, ocular acicles are contiguous, as in some species of
<taxonomicName id="4C6C4D789F2CFFF8FF47F18DFE9DFD07" box="[151,329,674,699]" class="Malacostraca" family="Diogenidae" genus="Areopaguristes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="52" phylum="Arthropoda" rank="genus">
<emphasis id="B918EAE99F2CFFF8FF47F18DFE9DFD07" box="[151,329,674,699]" italics="true" pageId="3" pageNumber="52">Areopaguristes</emphasis>
</taxonomicName>
, but are well, or more or less separated in
<taxonomicName id="4C6C4D789F2CFFF8FCEBF18DFC6CFD07" box="[827,952,674,699]" class="Malacostraca" family="Diogenidae" genus="Paguristes" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="52" phylum="Arthropoda" rank="genus">
<emphasis id="B918EAE99F2CFFF8FCEBF18DFC6CFD07" box="[827,952,674,699]" italics="true" pageId="3" pageNumber="52">Paguristes</emphasis>
</taxonomicName>
and
<taxonomicName id="4C6C4D789F2CFFF8FC21F18DFB15FD07" box="[1009,1217,674,699]" class="Malacostraca" family="Diogenidae" genus="Pseudopaguristes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="52" phylum="Arthropoda" rank="genus">
<emphasis id="B918EAE99F2CFFF8FC21F18DFB15FD07" box="[1009,1217,674,699]" italics="true" pageId="3" pageNumber="52">Pseudopaguristes</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8BD336FB9F2CFFF8FF15F1E7FBBFFCEE" blockId="3.[151,1437,151,1484]" pageId="3" pageNumber="52">
De Saint Laurent (1970) and
<bibRefCitation id="EFFD4B0A9F2CFFF8FDF7F1E7FCC7FD5E" author="McLaughlin" box="[551,787,712,738]" pageId="3" pageNumber="52" refString="McLaughlin, P. A. (1974) The hermit crabs (Crustacea Decapoda, Paguridea) of northwestern North America. Zoologische Verhandelingen, 130, 1 - 396." type="journal article" year="1974">McLaughlin (1974)</bibRefCitation>
mentioned that the preungual process is an important diagnostic character at genus level. The presence or absence of this preungal process in the fourth pereopod appears to be a highly variable character in
<taxonomicName id="4C6C4D789F2CFFF8FD43F03DFCC4FC97" box="[659,784,786,811]" class="Malacostraca" family="Diogenidae" genus="Paguristes" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="52" phylum="Arthropoda" rank="genus">
<emphasis id="B918EAE99F2CFFF8FD43F03DFCC4FC97" box="[659,784,786,811]" italics="true" pageId="3" pageNumber="52">Paguristes</emphasis>
</taxonomicName>
,
<taxonomicName id="4C6C4D789F2CFFF8FCCEF03DFC04FC97" box="[798,976,786,811]" class="Malacostraca" family="Diogenidae" genus="Areopaguristes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="52" phylum="Arthropoda" rank="genus">
<emphasis id="B918EAE99F2CFFF8FCCEF03DFC04FC97" box="[798,976,786,811]" italics="true" pageId="3" pageNumber="52">Areopaguristes</emphasis>
</taxonomicName>
and
<taxonomicName id="4C6C4D789F2CFFF8FBD8F03DFB0CFC97" box="[1032,1240,786,811]" class="Malacostraca" family="Diogenidae" genus="Pseudopaguristes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="52" phylum="Arthropoda" rank="genus">
<emphasis id="B918EAE99F2CFFF8FBD8F03DFB0CFC97" box="[1032,1240,786,811]" italics="true" pageId="3" pageNumber="52">Pseudopaguristes</emphasis>
</taxonomicName>
; in
<taxonomicName id="4C6C4D789F2CFFF8FAD5F03DFA4FFC97" box="[1285,1435,786,811]" class="Malacostraca" family="Diogenidae" genus="Tetralobistes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="52" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B918EAE99F2CFFF8FAD5F03DFA4FFC97" box="[1285,1435,786,811]" italics="true" pageId="3" pageNumber="52">Tetralobistes</emphasis>
</taxonomicName>
<emphasis id="B918EAE99F2CFFF8FF47F018FED0FCED" bold="true" box="[151,260,823,849]" pageId="3" pageNumber="52">
<taxonomicNameLabel id="A22B57929F2CFFF8FF47F018FED0FCED" box="[151,260,823,849]" pageId="3" pageNumber="52" rank="genus">gen. nov.</taxonomicNameLabel>
</emphasis>
, so far a monotypic genus, the value of this character is difficult to assess.
</paragraph>
<paragraph id="8BD336FB9F2CFFF8FF15F072FDB8FB7B" blockId="3.[151,1437,151,1484]" pageId="3" pageNumber="52">
The shape of the telson, including the depth of lateral constriction and posteromedian cleft, the
<typeStatus id="54D788599F2CFFF8FA97F072FAAEFCCB" box="[1351,1402,861,887]" pageId="3" pageNumber="52">type</typeStatus>
of armature, and the pilosity of posterior lobes, are important characters used to define relationships among hermit crab genera (de Saint Laurent 1970). Perhaps the most conspicuous character separating
<taxonomicName id="4C6C4D789F2CFFF8FAD5F087FA48FC7D" box="[1285,1436,936,961]" class="Malacostraca" family="Diogenidae" genus="Tetralobistes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="52" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B918EAE99F2CFFF8FAD5F087FA48FC7D" box="[1285,1436,936,961]" italics="true" pageId="3" pageNumber="52">Tetralobistes</emphasis>
</taxonomicName>
<emphasis id="B918EAE99F2CFFF8FF47F0E2FED2FC5B" bold="true" box="[151,262,973,999]" pageId="3" pageNumber="52">
<taxonomicNameLabel id="A22B57929F2CFFF8FF47F0E2FED2FC5B" box="[151,262,973,999]" pageId="3" pageNumber="52" rank="genus">gen. nov.</taxonomicNameLabel>
</emphasis>
from the other three genera of
<tableCitation id="C6EE03409F2CFFF8FDAAF0E2FD06FC5B" box="[634,722,973,999]" captionStart="TABLE 1" captionStartId="2.[151,239,151,175]" captionText="TABLE 1. Selected comparative characters observed in genera of Diogenidae closely related to Tetralobistes gen. nov." pageId="3" pageNumber="52">Table 1</tableCitation>
is the shape of its telson: posterior margin typically divided into four lobes, the two inner lobes much smaller than the two outer lobes. Actually, no genus of
<taxonomicName id="4C6C4D789F2CFFF8FAC5F0DDFA48FBB0" box="[1301,1436,1010,1036]" class="Malacostraca" family="Diogenidae" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="52" phylum="Arthropoda" rank="family">Diogenidae</taxonomicName>
features a four-lobed posterior margin. In the case of
<taxonomicName id="4C6C4D789F2CFFF8FCF4F737FA9DFB8E" authority="H. Milne Edwards, 1836" authorityName="H. Milne Edwards" authorityYear="1836" box="[804,1353,1048,1074]" class="Malacostraca" family="Diogenidae" genus="Paguristes" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="52" phylum="Arthropoda" rank="species" species="frontalis">
<emphasis id="B918EAE99F2CFFF8FCF4F737FBC0FB8D" box="[804,1044,1048,1073]" italics="true" pageId="3" pageNumber="52">Paguristes frontalis</emphasis>
H.
<bibRefCitation id="EFFD4B0A9F2CFFF8FB93F737FA9DFB8E" author="Milne" box="[1091,1353,1048,1074]" pageId="3" pageNumber="52" refString="Milne Edwards, A. (1836) Observations zoologiques sur les Pagures et description d'un nouveau genre de la tribu des Paguriens. Annales des Sciences Naturelles Zoologie, Paris 2 (6), 257 - 288." type="journal article" year="1836">Milne Edwards, 1836</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="4C6C4D789F2CFFF8FA56F737FDE3FBEB" authority="Morgan, 1987" authorityName="Morgan" authorityYear="1987" class="Malacostraca" family="Diogenidae" genus="Paguristes" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="52" phylum="Arthropoda" rank="species" species="purpureantennatus">
<emphasis id="B918EAE99F2CFFF8FA56F737FEABFBEA" italics="true" pageId="3" pageNumber="52">P. purpureantennatus</emphasis>
<bibRefCitation id="EFFD4B0A9F2CFFF8FE57F712FDE3FBEB" author="Morgan" box="[391,567,1085,1111]" pageId="3" pageNumber="52" refString="Morgan, G. J. (1987) Two new species of Paguristes (Decapoda: Anomura: Diogenidae) from southwestern Australia. Proceedings of the Biological Society of Washington, 100 (4), 726 - 734." type="journal article" year="1987">Morgan, 1987</bibRefCitation>
</taxonomicName>
, the telson is four-lobed, but these correspond to anterior (two weakly marked lobes) and posterior (two lobes) sections of the telson, not to the posterior margin. As seen here, the combination of several characters set
<taxonomicName id="4C6C4D789F2CFFF8FD87F7A7FD3BFB1D" box="[599,751,1160,1185]" class="Malacostraca" family="Diogenidae" genus="Tetralobistes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="52" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B918EAE99F2CFFF8FD87F7A7FD3BFB1D" box="[599,751,1160,1185]" italics="true" pageId="3" pageNumber="52">Tetralobistes</emphasis>
</taxonomicName>
<emphasis id="B918EAE99F2CFFF8FD27F7A8FCB2FB1D" bold="true" box="[759,870,1159,1185]" pageId="3" pageNumber="52">
<taxonomicNameLabel id="A22B57929F2CFFF8FD27F7A8FCB2FB1D" box="[759,870,1159,1185]" pageId="3" pageNumber="52" rank="genus">gen. nov.</taxonomicNameLabel>
</emphasis>
apart from the three other genera, and supports the recognition of a distinct, new genus.
</paragraph>
<paragraph id="8BD336FB9F2CFFF8FF15F7FDFC5EFA70" blockId="3.[151,1437,151,1484]" pageId="3" pageNumber="52">
A comprehensive revision of the “
<taxonomicName id="4C6C4D789F2CFFF8FDBCF7FCFD3BFB50" box="[620,751,1235,1260]" class="Malacostraca" family="Diogenidae" genus="Paguristes" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="52" phylum="Arthropoda" rank="genus">
<emphasis id="B918EAE99F2CFFF8FDBCF7FCFD3BFB50" box="[620,751,1235,1260]" italics="true" pageId="3" pageNumber="52">Paguristes</emphasis>
</taxonomicName>
” (sensu lato) species occurring in the western Atlantlic might reveal an amphi-American distribution of
<taxonomicName id="4C6C4D789F2CFFF8FD30F7D7FCADFAAD" box="[736,889,1272,1297]" class="Malacostraca" family="Diogenidae" genus="Tetralobistes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="52" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B918EAE99F2CFFF8FD30F7D7FCADFAAD" box="[736,889,1272,1297]" italics="true" pageId="3" pageNumber="52">Tetralobistes</emphasis>
</taxonomicName>
<emphasis id="B918EAE99F2CFFF8FC52F7D8FC27FAAD" bold="true" box="[898,1011,1271,1297]" pageId="3" pageNumber="52">
<taxonomicNameLabel id="A22B57929F2CFFF8FC52F7D8FC27FAAD" box="[898,1011,1271,1297]" pageId="3" pageNumber="52" rank="genus">gen. nov.</taxonomicNameLabel>
</emphasis>
, as it is known to occur with many decapod crustacean genera (see
<bibRefCitation id="EFFD4B0A9F2CFFF8FDC2F632FD7FFA8B" author="Boschi" box="[530,683,1309,1335]" pageId="3" pageNumber="52" refString="Boschi, E. E. (2000) Species of decapod crustaceans and their distribution in the American marine zoogeographic provinces. Revista de Investigacion y Desarrollo Pesquero, 13, 7 - 136." type="journal article" year="2000">Boschi 2000</bibRefCitation>
). A careful review of species of
<taxonomicName id="4C6C4D789F2CFFF8FBFBF632FB7CFA8A" box="[1067,1192,1309,1334]" class="Malacostraca" family="Diogenidae" genus="Paguristes" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="52" phylum="Arthropoda" rank="genus">
<emphasis id="B918EAE99F2CFFF8FBFBF632FB7CFA8A" box="[1067,1192,1309,1334]" italics="true" pageId="3" pageNumber="52">Paguristes</emphasis>
</taxonomicName>
,
<taxonomicName id="4C6C4D789F2CFFF8FB67F632FABDFA8A" box="[1207,1385,1309,1334]" class="Malacostraca" family="Diogenidae" genus="Areopaguristes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="52" phylum="Arthropoda" rank="genus">
<emphasis id="B918EAE99F2CFFF8FB67F632FABDFA8A" box="[1207,1385,1309,1334]" italics="true" pageId="3" pageNumber="52">Areopaguristes</emphasis>
</taxonomicName>
and
<taxonomicName id="4C6C4D789F2CFFF8FF47F66CFEBEFAE0" box="[151,362,1347,1372]" class="Malacostraca" family="Diogenidae" genus="Pseudopaguristes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="52" phylum="Arthropoda" rank="genus">
<emphasis id="B918EAE99F2CFFF8FF47F66CFEBEFAE0" box="[151,362,1347,1372]" italics="true" pageId="3" pageNumber="52">Pseudopaguristes</emphasis>
</taxonomicName>
from other regions (including both coasts of Central and South
<collectingCountry id="F37B766B9F2CFFF8FBBAF66DFB06FAE0" box="[1130,1234,1346,1372]" name="United States of America" pageId="3" pageNumber="52">America</collectingCountry>
) that share some apomorphic characters with
<taxonomicName id="4C6C4D789F2CFFF8FE23F647FD44FA3D" box="[499,656,1384,1409]" class="Malacostraca" family="Diogenidae" genus="Tetralobistes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="52" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B918EAE99F2CFFF8FE23F647FD44FA3D" box="[499,656,1384,1409]" italics="true" pageId="3" pageNumber="52">Tetralobistes</emphasis>
</taxonomicName>
<emphasis id="B918EAE99F2CFFF8FD49F648FCDFFA3D" bold="true" box="[665,779,1383,1409]" pageId="3" pageNumber="52">
<taxonomicNameLabel id="A22B57929F2CFFF8FD49F648FCDFFA3D" box="[665,779,1383,1409]" pageId="3" pageNumber="52" rank="genus">gen. nov.</taxonomicNameLabel>
</emphasis>
(e.g., absence of paired second male and female first pleopods; absence of brood pouch; presence or absence of epipod on first maxilliped) is also needed to define more clearly the affinities of the later genus with the other three.
</paragraph>
</subSubSection>
</treatment>
</document>
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