218 lines
29 KiB
XML
218 lines
29 KiB
XML
<document id="6B78523EA7516F52BAA6E3037F757980" ID-CLB-Dataset="8562" ID-DOI="10.5252/geodiversitas2020v42a20" ID-GBIF-Dataset="ad4dbfb6-14ce-48e2-8fe9-99fa26a9bcf4" ID-ISSN="1638-9395" ID-Zenodo-Dep="4447563" ID-ZooBank="urn:lsid:zoobank.org:pub:8FF2A078-CE45-4BF1-A681-00136F57375E" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1610989940068" checkinUser="felipe" docAuthor="Ivanov, Martin, Čerňanský, Andrej, Bonilla-Salomón, Isaac & Luján, Àngel Hernández" docDate="2020" docId="03C587C74312FFFCFE2CF9AF497BFEE0" docLanguage="en" docName="Geodiversitas.42.20.343-376.pdf" docOrigin="Geodiversitas 42 (20)" docStyle="DocumentStyle:F830B10FF475E64C1F1601E3B32DDC00.4:Geodiversitas.2018-.journal_article" docStyleId="F830B10FF475E64C1F1601E3B32DDC00" docStyleName="Geodiversitas.2018-.journal_article" docStyleVersion="4" docTitle="Natrix sp. Laurenti 1768" docType="treatment" docVersion="10" lastPageNumber="363" masterDocId="FFFCFFBF4304FFE4FFBCFF9C484CFFD2" masterDocTitle="Early Miocene squamate assemblage from the Mokrá-Western Quarry (Czech Republic) and its palaeobiogeographical and palaeoenvironmental implications" masterLastPageNumber="376" masterPageNumber="343" pageNumber="363" updateTime="1698925529958" updateUser="ExternalLinkService" zenodo-license-document="CC0-1.0" zenodo-license-figures="CC0-1.0">
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<mods:title id="75E180B39FEFCEC29B0E6AB4172A996A">Early Miocene squamate assemblage from the Mokrá-Western Quarry (Czech Republic) and its palaeobiogeographical and palaeoenvironmental implications</mods:title>
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<mods:namePart id="652F619C83108FEC0E95C1A7FF7580FE">Ivanov, Martin</mods:namePart>
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<mods:namePart id="9C70E1888B47515A908AC40FCF0FA2EA">Čerňanský, Andrej</mods:namePart>
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<mods:namePart id="FDCF58BDE8A254FDB654F7CD86483B88">Bonilla-Salomón, Isaac</mods:namePart>
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<mods:namePart id="2458C8EEF3631B5825E8BB9BF3645ED5">Luján, Àngel Hernández</mods:namePart>
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<mods:title id="6004297BF01C3E910872D5695E5532A8">Geodiversitas</mods:title>
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<mods:date id="06B3F8CDCD70A46740648D14FAF86733">2020</mods:date>
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<mods:number id="7EC3FCB8E8C93E8519FED9C5997029E0">2020-08-28</mods:number>
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<treatment id="03C587C74312FFFCFE2CF9AF497BFEE0" ID-DOI="http://doi.org/10.5281/zenodo.4447730" ID-GBIF-Taxon="177200735" ID-Zenodo-Dep="4447730" LSID="urn:lsid:plazi:treatment:03C587C74312FFFCFE2CF9AF497BFEE0" httpUri="http://treatment.plazi.org/id/03C587C74312FFFCFE2CF9AF497BFEE0" lastPageId="24" lastPageNumber="363" pageId="22" pageNumber="363">
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<subSubSection id="C376655A4312FFF2FE2CF9AF49B0F99F" box="[400,508,1587,1613]" pageId="22" pageNumber="363" type="nomenclature">
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<paragraph id="8BD336D14312FFF2FE2CF9AF49B0F99F" blockId="22.[400,508,1587,1613]" box="[400,508,1587,1613]" pageId="22" pageNumber="363">
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<heading id="D09B81BD4312FFF2FE2CF9AF49B0F99F" box="[400,508,1587,1613]" centered="true" fontSize="11" level="2" pageId="22" pageNumber="363" reason="2">
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<taxonomicName id="4C6C4D524312FFF2FE2CF9AF499AF99E" authorityName="Laurenti" authorityYear="1768" box="[400,470,1587,1612]" class="Reptilia" family="Colubridae" genus="Natrix" kingdom="Animalia" order="Squamata" pageId="22" pageNumber="363" phylum="Chordata" rank="species" species="sp.">
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<emphasis id="B918EAC34312FFF2FE2CF9AF499AF99E" box="[400,470,1587,1612]" italics="true" pageId="22" pageNumber="363">Natrix</emphasis>
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</taxonomicName>
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sp.
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C376655A4312FFF2FE3DF9CE4A46F9BF" box="[385,522,1618,1645]" pageId="22" pageNumber="363" type="description">
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<paragraph id="8BD336D14312FFF2FE3DF9CE4A46F9BF" blockId="22.[385,522,1618,1645]" box="[385,522,1618,1645]" pageId="22" pageNumber="363">
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(
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<figureCitation id="13572A544312FFF2FE36F9CE499BF9BE" box="[394,471,1618,1644]" captionStart="FIG" captionStartId="23.[132,143,1530,1548]" captionTargetBox="[132,1455,215,1487]" captionTargetId="figure-224@23.[698,1267,758,1306]" captionTargetPageId="23" captionText="FIG. 12. — Natrix sp. and “Natricinae” indet. from the early Miocene (MN 4) of MWQ. Natrix sp.: trunk vertebra Pal. 1968 (2/2003 Reptile Joint) in left lateral (A), dorsal (B), ventral (C), cranial (D) and caudal (E) views.“Natricinae” indet.:trunk vertebra Pal. 1483 (1/2001 Turtle Joint) with preserved hypapophysis in left lateral (F), dorsal (G), ventral (H) and caudal (I) views. Abbreviations: ak, anterior keel; cd, condyle; ct, cotyle; es, epizypapophyseal spine; hy, hypapophysis; lf, lateral foramen;nc, neural canal;ns, neural spine;pctf, paracotylar foramen;po, postzygapophysis;pof, postzygapophyseal articular facet;prf, prezygapophyseal articular facet; prp, prezygapophyseal process;scf, subcentral foramen;scg, subcentral groove;scr, subcentral ridge;zy, zygosphene;zyg, zygantrum.Scale bars:2 mm." figureDoi="http://doi.org/10.5281/zenodo.4447589" httpUri="https://zenodo.org/record/4447589/files/figure.png" pageId="22" pageNumber="363">Fig. 12</figureCitation>
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A-E)
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</paragraph>
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</subSubSection>
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<subSubSection id="C376655A4312FFF2FF38F9104A9EF901" pageId="22" pageNumber="363" type="reference_group">
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<paragraph id="8BD336D14312FFF2FF38F9104A47F971" blockId="22.[132,523,1675,1699]" box="[132,523,1675,1699]" pageId="22" pageNumber="363">
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<treatmentCitationGroup id="AB7C11FF4312FFF2FF38F9104A47F971" box="[132,523,1675,1699]" pageId="22" pageNumber="363">
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<taxonomicName id="4C6C4D524312FFF2FF38F910488FF971" authorityName="Laurenti" authorityYear="1768" box="[132,195,1676,1699]" class="Reptilia" family="Colubridae" genus="Natrix" kingdom="Animalia" order="Squamata" pageId="22" pageNumber="363" phylum="Chordata" rank="species" species="sp.">
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<emphasis id="B918EAC34312FFF2FF38F910488FF971" box="[132,195,1676,1699]" italics="true" pageId="22" pageNumber="363">Natrix</emphasis>
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</taxonomicName>
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sp. –
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<treatmentCitation id="0ACD10C04312FFF2FF42F9104A4BF971" author="IVANOV M. & MUSIL R." box="[254,519,1675,1699]" page="229" pageId="22" pageNumber="363" year="2004">
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<bibRefCitation id="EFFD4B204312FFF2FF42F9104A4BF971" author="IVANOV M. & MUSIL R." box="[254,519,1675,1699]" pageId="22" pageNumber="363" pagination="223 - 236" refId="ref24802" refString="IVANOV M. & MUSIL R. 2004. - Preliminary results of investigation of Neogene vertebrates from the Mokra-Quarry site. Acta Musei Moraviae, Scientiae geologicae 89: 223 - 236 (in Czech)." type="journal article" year="2004">Ivanov & Musil 2004: 229</bibRefCitation>
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</treatmentCitation>
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.
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</treatmentCitationGroup>
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</paragraph>
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<paragraph id="8BD336D14312FFF2FF38F9204A9EF901" blockId="22.[132,722,1723,1747]" box="[132,722,1723,1747]" pageId="22" pageNumber="363">
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<treatmentCitationGroup id="AB7C11FF4312FFF2FF38F9204A9EF901" box="[132,722,1723,1747]" pageId="22" pageNumber="363">
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<taxonomicName id="4C6C4D524312FFF2FF38F920488FF901" authorityName="Laurenti" authorityYear="1768" box="[132,195,1724,1747]" class="Reptilia" family="Colubridae" genus="Natrix" kingdom="Animalia" order="Squamata" pageId="22" pageNumber="363" phylum="Chordata" rank="species" species="sp.">
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<emphasis id="B918EAC34312FFF2FF38F920488FF901" box="[132,195,1724,1747]" italics="true" pageId="22" pageNumber="363">Natrix</emphasis>
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</taxonomicName>
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sp.,
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<typeStatus id="54D788734312FFF2FF4DF9274957F901" box="[241,283,1723,1747]" pageId="22" pageNumber="363">type</typeStatus>
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I –
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<treatmentCitation id="0ACD10C04312FFF2FEFFF9274A6AF901" author="IVANOV M. & MUSIL R. & BRZOBOHATY R." box="[323,550,1723,1747]" page="229" pageId="22" pageNumber="363" year="2006">
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||
<bibRefCitation id="EFFD4B204312FFF2FEFFF9274A6AF901" author="IVANOV M. & MUSIL R. & BRZOBOHATY R." box="[323,550,1723,1747]" pageId="22" pageNumber="363" pagination="223 - 239" refId="ref24955" refString="IVANOV M., MUSIL R. & BRZOBOHATY R. 2006. - Terrestrial and Marine Faunas from the Miocene Deposits of the Mokra Plateau (Drahany Upland, Czech Republic) - Impact on Palaeogeography. Beitrage zur Palaontologie 30: 223 - 239." type="journal article" year="2006">
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Ivanov
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<emphasis id="B918EAC34312FFF2FE36F92049F5F901" box="[394,441,1723,1747]" italics="true" pageId="22" pageNumber="363">et al.</emphasis>
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2006: 229
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</bibRefCitation>
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</treatmentCitation>
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||
, table 2 (in part).
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</treatmentCitationGroup>
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C376655A4312FFF2FF38F9774ACAF8E5" pageId="22" pageNumber="363" type="materials_examined">
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<paragraph id="8BD336D14312FFF2FF38F9774ACAF8E5" blockId="22.[130,776,1771,1847]" pageId="22" pageNumber="363">
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MATERIAL. —
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<materialsCitation id="3B043C8C4312FFF2FEB3F9774ACAF8E5" ID-GBIF-Occurrence="3027945312" pageId="22" pageNumber="363">
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MWQ, early Miocene, Burdigalian, Orleanian,
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<collectionCode id="ED7DAE144312FFF2FD7AF9774ABCF8D0" box="[710,752,1771,1794]" country="Brazil" name="Museu Nacional, Universidade Federal do Rio de Janeiro" pageId="22" pageNumber="363">MN</collectionCode>
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4: 1/2001 Turtle Joint:Two trunk vertebrae (Pal. 1481-1482); 2/2003 Reptile Joint: Two trunk vertebrae (Pal. 1968-1969).
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</materialsCitation>
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</paragraph>
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||
</subSubSection>
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<subSubSection id="C376655A4312FFF2FF38F8EE4DC4F99F" pageId="22" pageNumber="363" type="description">
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<paragraph id="8BD336D14312FFF2FF38F8EE4959F858" blockId="22.[132,775,1906,2028]" box="[132,277,1906,1932]" pageId="22" pageNumber="363">DESCRIPTION</paragraph>
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<paragraph id="8BD336D14312FFF2FF38F80D4965F879" blockId="22.[132,775,1906,2028]" box="[132,297,1937,1963]" pageId="22" pageNumber="363">
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<emphasis id="B918EAC34312FFF2FF38F80D4965F879" box="[132,297,1937,1963]" italics="true" pageId="22" pageNumber="363">Trunk vertebrae</emphasis>
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</paragraph>
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<paragraph id="8BD336D14312FFF2FF38F82D4DC4F99F" blockId="22.[132,775,1906,2028]" lastBlockId="22.[810,1457,821,1613]" pageId="22" pageNumber="363">
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The most complete specimen, Pal. 1968, is fragmentary with the hypapophysis and right prezygapophyseal process broken off, and strongly damaged paradiapophyses (
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<figureCitation id="13572A544312FFF2FB42FCA94D0BFC9D" box="[1278,1351,821,847]" captionStart="FIG" captionStartId="23.[132,143,1530,1548]" captionTargetBox="[132,1455,215,1487]" captionTargetId="figure-224@23.[698,1267,758,1306]" captionTargetPageId="23" captionText="FIG. 12. — Natrix sp. and “Natricinae” indet. from the early Miocene (MN 4) of MWQ. Natrix sp.: trunk vertebra Pal. 1968 (2/2003 Reptile Joint) in left lateral (A), dorsal (B), ventral (C), cranial (D) and caudal (E) views.“Natricinae” indet.:trunk vertebra Pal. 1483 (1/2001 Turtle Joint) with preserved hypapophysis in left lateral (F), dorsal (G), ventral (H) and caudal (I) views. Abbreviations: ak, anterior keel; cd, condyle; ct, cotyle; es, epizypapophyseal spine; hy, hypapophysis; lf, lateral foramen;nc, neural canal;ns, neural spine;pctf, paracotylar foramen;po, postzygapophysis;pof, postzygapophyseal articular facet;prf, prezygapophyseal articular facet; prp, prezygapophyseal process;scf, subcentral foramen;scg, subcentral groove;scr, subcentral ridge;zy, zygosphene;zyg, zygantrum.Scale bars:2 mm." figureDoi="http://doi.org/10.5281/zenodo.4447589" httpUri="https://zenodo.org/record/4447589/files/figure.png" pageId="22" pageNumber="363">Fig. 12</figureCitation>
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A-E). The vertebral centrum is cylindrical. In lateral view, the interzygapophyseal ridges are moderately developed. The completely preserved neural spine is about twice as long as high. Its cranial margin is inclined anteriorly whereas the caudal margin is inclined posteriorly. The small lateral foramina are situated in shallow depressions. The well-developed subcentral ridges are arched dorsally. In dorsal view, the prezygapophyseal articular facets are almost oval.The prezygapophyseal processes are about two thirds of the prezygapophyseal facets length. The cranial margin of the mostly damaged zygosphene has developed a wide medial lobe and small lateral lobes. The epizygapophyseal spines are absent. In ventral view, the hypapophysis expands laterally in a cranial direction to form a triangular anterior keel. The very small subcentral foramina are situated on both sides of the hypapophysis.In cranial view, the neural arch is slightly vaulted, and the neural canal is rounded with short lateral sinuses. The parapophyseal processes are separated from the rounded cotyle by deep furrows.The paracotylar foramina are situated on both sides of the cotyle. The left paracotylar foramen is doubled in Pal. 1968. The small paracotylar tubercles occur at the ventral margin of the cotyle. The vertebral dimensions of the figured specimen (Pal. 1968) are as follows: cl =
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<quantity id="4C949B344312FFF2FB71FA6F4D7DF9DF" box="[1229,1329,1523,1549]" metricMagnitude="-3" metricUnit="m" metricValue="4.6" pageId="22" pageNumber="363" unit="mm" value="4.6">4.60 mm</quantity>
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; naw =
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<quantity id="4C949B344312FFF2FAC3FA6F4B15F9FF" metricMagnitude="-3" metricUnit="m" metricValue="2.72" pageId="22" pageNumber="363" unit="mm" value="2.72">2.72 mm</quantity>
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; cl/naw = 1.69. The largest specimen (Pal.1481) measures as follows: cl =
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<quantity id="4C949B344312FFF2FC76F9AF4C62F99F" box="[970,1070,1587,1613]" metricMagnitude="-3" metricUnit="m" metricValue="4.93" pageId="22" pageNumber="363" unit="mm" value="4.93">4.93 mm</quantity>
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||
; naw =
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||
<quantity id="4C949B344312FFF2FB3CF9AE4CA9F99F" box="[1152,1253,1586,1613]" metricMagnitude="-3" metricUnit="m" metricValue="2.85" pageId="22" pageNumber="363" unit="mm" value="2.85">2.85 mm</quantity>
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||
; cl/naw = 1.73.
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</paragraph>
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</subSubSection>
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<subSubSection id="C376655A4312FFFCFC90F9EF497BFEE0" lastPageId="24" lastPageNumber="365" pageId="22" pageNumber="363" type="discussion">
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<paragraph id="8BD336D14312FFF2FC90F9EF4BDDF959" blockId="22.[811,1457,1651,2028]" box="[812,913,1651,1677]" pageId="22" pageNumber="363">REMARKS</paragraph>
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<paragraph id="8BD336D14312FFFCFC91F90E497BFEE0" blockId="22.[811,1457,1651,2028]" lastBlockId="24.[132,777,215,306]" lastPageId="24" lastPageNumber="365" pageId="22" pageNumber="363">
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Assignment to the “natricine” snakes is based on the presence of hypapophyses in precloacal vertebrae, the vaulted neural arch, the high neural spine, and the presence of paracotylar foramina on either side of the cotyle. The most complete vertebra with laterally directed prezygapophyses, an elongated centrum with a triangular anterior keel on the hypapophysis, and a neural spine inclined both anteriorly and posteriorly permit identification of the preserved vertebrae to the genus
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<taxonomicName id="4C6C4D524312FFF2FC91F80E4B3DF879" authorityName="Laurenti" authorityYear="1768" box="[813,881,1938,1963]" class="Reptilia" family="Colubridae" genus="Natrix" kingdom="Animalia" order="Squamata" pageId="22" pageNumber="363" phylum="Chordata" rank="genus">
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<emphasis id="B918EAC34312FFF2FC91F80E4B3DF879" box="[813,881,1938,1963]" italics="true" pageId="22" pageNumber="363">Natrix</emphasis>
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</taxonomicName>
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whose fossil remains are abundant in Central Europe as early as the early Miocene (
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<bibRefCitation id="EFFD4B204312FFF2FBDBF82E4CBBF819" author="IVANOV M." box="[1127,1271,1969,1996]" pageId="22" pageNumber="363" pagination="513 - 534" refId="ref24652" refString="IVANOV M. 2002 a. - The oldest known Miocene snake fauna from Central Europe: Merkur-North locality, Czech Republic. Acta Palaeontologica Polonica 47 (3): 513 - 534." type="journal article" year="2002">Ivanov 2002a</bibRefCitation>
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||
;
|
||
<bibRefCitation id="EFFD4B204312FFF2FAB9F82F4B28F83E" author="CERNANSKY A. & RAGE J. C. & KLEMBARA J." pageId="22" pageNumber="363" pagination="97 - 128" refId="ref23201" refString="CERNANSKY A., RAGE J. C. & KLEMBARA J. 2015. - The Early Miocene squamates of Amoneburg (Germany): the first stages of modern squamates in Europe. The Journal of Systematic Palaeontology 13 (2): 97 - 128. https: // doi. org / 10.1080 / 14772019.20 14.897266" type="journal article" year="2015">
|
||
Čerňanský
|
||
<emphasis id="B918EAC34312FFF2FAC1F82E4DFCF819" box="[1405,1456,1969,1995]" italics="true" pageId="22" pageNumber="363">et al.</emphasis>
|
||
2015
|
||
</bibRefCitation>
|
||
).
|
||
<taxonomicName id="4C6C4D524312FFF2FCC0F84E4B8CF839" authorityName="Laurenti" authorityYear="1768" box="[892,960,2002,2027]" class="Reptilia" family="Colubridae" genus="Natrix" kingdom="Animalia" order="Squamata" pageId="22" pageNumber="363" phylum="Chordata" rank="genus">
|
||
<emphasis id="B918EAC34312FFF2FCC0F84E4B8CF839" box="[892,960,2002,2027]" italics="true" pageId="22" pageNumber="363">Natrix</emphasis>
|
||
</taxonomicName>
|
||
sp. differs from early Miocene
|
||
<taxonomicName id="4C6C4D524312FFF3FAA9F84E4942F91E" authority="Ivanov, 2002" authorityName="Ivanov" authorityYear="2002" class="Reptilia" family="Colubridae" genus="Natrix" kingdom="Animalia" lastPageId="23" lastPageNumber="364" order="Squamata" pageId="22" pageNumber="363" phylum="Chordata" rank="species" species="merkurensis">
|
||
<emphasis id="B918EAC34312FFF2FAA9F84E4DE3F839" box="[1301,1455,2001,2027]" italics="true" pageId="22" pageNumber="363">N. merkurensis</emphasis>
|
||
Ivanov, 2002
|
||
</taxonomicName>
|
||
(MN 3a-?MN 4;
|
||
<bibRefCitation id="EFFD4B204313FFF3FE70F92F4A10F91E" author="IVANOV M." box="[460,604,1714,1741]" pageId="23" pageNumber="364" pagination="513 - 534" refId="ref24652" refString="IVANOV M. 2002 a. - The oldest known Miocene snake fauna from Central Europe: Merkur-North locality, Czech Republic. Acta Palaeontologica Polonica 47 (3): 513 - 534." type="journal article" year="2002">Ivanov 2002a</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFFD4B204313FFF3FDD6F92F48F7F93F" author="RAGE J. - C. & BAILON S." pageId="23" pageNumber="364" pagination="413 - 441" refId="ref26650" refString="RAGE J. - C. & BAILON S. 2005. - Amphibians and squamate reptiles from the late early Miocene (MN 4) of Beon 1 (Montrealdu-Gers, southwestern France). Geodiversitas 27 (3): 413 - 441." type="journal article" year="2005">Rage & Bailon 2005</bibRefCitation>
|
||
) in its smaller dimensions, elongated prezygapophyseal facets, shorter prezygapophyseal processes, and more distinct lateral lobes of the zygosphene. It differs from
|
||
<taxonomicName id="4C6C4D524313FFF3FDD0F88E4951F89E" authority="(Lartet, 1851)" baseAuthorityName="Lartet" baseAuthorityYear="1851" class="Reptilia" family="Colubridae" genus="Natrix" kingdom="Animalia" order="Squamata" pageId="23" pageNumber="364" phylum="Chordata" rank="species" species="sansaniensis">
|
||
<emphasis id="B918EAC34313FFF3FDD0F88E4B4AF8F9" box="[620,774,1810,1835]" italics="true" pageId="23" pageNumber="364">N. sansaniensis</emphasis>
|
||
(Lartet, 1851)
|
||
</taxonomicName>
|
||
, reported from the early and middle Miocene (MN 3-?MN 7+8;
|
||
<bibRefCitation id="EFFD4B204313FFF3FEF3F8CE4A50F8BE" author="AUGE M. & RAGE J. - C." box="[335,540,1874,1900]" pageId="23" pageNumber="364" refId="ref22449" refString="AUGE M. & RAGE J. - C. 2000. - Les Squamates (Reptilia) du Miocene moyen de Sansan (Gers, France), in GINSBURG L. (ed.), La faune miocene de Sansan et son environnement. Museum national d'Histoire naturelle, Paris: 263 - 313 (Memoires du Museum national d'Histoire naturelle; 183)." type="journal volume" year="2000">Augé & Rage 2000</bibRefCitation>
|
||
; Ivanov 2000, 2002a, b), in the less vaulted neural arch and flattened prezygapophyseal processes (
|
||
<bibRefCitation id="EFFD4B204313FFF3FEFFF80D4994F87E" author="SZYNDLAR Z." box="[323,472,1937,1964]" pageId="23" pageNumber="364" pagination="31 - 52" refId="ref27643" refString="SZYNDLAR Z. 2005. - Snake fauna from the Late Miocene of Rudabanya. Palaeontographia Italica 90 (2003): 31 - 52." type="journal article" year="2005">Szyndlar 2005</bibRefCitation>
|
||
).
|
||
<taxonomicName id="4C6C4D524313FFF3FE51F80E48F7F819" authority="Szyndlar, 1984" authorityName="Szyndlar" authorityYear="1984" class="Reptilia" family="Colubridae" genus="Natrix" kingdom="Animalia" order="Squamata" pageId="23" pageNumber="364" phylum="Chordata" rank="species" species="longivertebrata">
|
||
<emphasis id="B918EAC34313FFF3FE51F80E4AEFF879" box="[493,675,1937,1963]" italics="true" pageId="23" pageNumber="364">N. longivertebrata</emphasis>
|
||
<bibRefCitation id="EFFD4B204313FFF3FD1BF80D48F7F819" author="SZYNDLAR Z." pageId="23" pageNumber="364" pagination="3 - 156" refId="ref27272" refString="SZYNDLAR Z. 1984. - Fossil snakes from Poland. Acta Zoologica Cracoviensia 28 (1): 3 - 156." type="journal article" year="1984">Szyndlar, 1984</bibRefCitation>
|
||
</taxonomicName>
|
||
reported from the late Miocene (MN 9, MN 10/11;
|
||
<bibRefCitation id="EFFD4B204313FFF3FF3AF84E4945F839" author="IVANOV M." box="[134,265,2001,2028]" pageId="23" pageNumber="364" refId="ref24525" refString="IVANOV M. 1997. - Hadi evropskeho kenozoika. Unpublished PhD thesis, Masaryk University, Department of Geology and Palaeontology), Brno, 217 p. (in Czech)." type="book" year="1997">Ivanov 1997</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFFD4B204313FFF3FEAAF84D49E1F83E" author="TEMPFER P. M." box="[278,429,2001,2028]" pageId="23" pageNumber="364" pagination="145 - 253" refId="ref27985" refString="TEMPFER P. M. 2005. - The Herpetofauna (Amphibia: Caudata, Anura; Reptilia: Scleroglossa) of the Upper Miocene Locality Kohfidisch (Burgerland, Austria). Beitrage zur Palaontologie 29: 145 - 253." type="journal article" year="2005">Tempfer 2005</bibRefCitation>
|
||
) and Pliocene (MN 14-MN 16;
|
||
<bibRefCitation id="EFFD4B204313FFF3FC90F92E4B88F91E" author="SZYNDLAR Z." box="[812,964,1714,1740]" pageId="23" pageNumber="364" pagination="3 - 156" refId="ref27272" refString="SZYNDLAR Z. 1984. - Fossil snakes from Poland. Acta Zoologica Cracoviensia 28 (1): 3 - 156." type="journal article" year="1984">Szyndlar 1984</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="EFFD4B204313FFF3FC6CF92E4C59F91E" author="SZYNDLAR Z." box="[976,1045,1714,1740]" pageId="23" pageNumber="364" pagination="237 - 266" refId="ref27471" refString="SZYNDLAR Z. 1991 b. - A rewiew of Neogene and Quaternary snakes of Central and Eastern Europe. Part II: Natricinae, Elapidae, Viperidae. Estudios geologicos 47 (3 - 4): 237 - 266. https: // doi. org / 10.3989 / egeol. 91473 - 4422" type="journal article" year="1991">1991b</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFFD4B204313FFF3FB9DF92E4CE1F91E" author="VENCZEL M." box="[1057,1197,1714,1740]" pageId="23" pageNumber="364" pagination="79 - 90" refId="ref28151" refString="VENCZEL M. 2001. - Anurans and squamates from the Lower Pliocene (MN 14) Osztramos 1 locality (Northern Hungary). Fragmenta Palaeontologica Hungarica 19: 79 - 90." type="journal article" year="2001">Venczel 2001</bibRefCitation>
|
||
), and doubtfully as early as the middle Miocene (?MN 6-?MN 7+8;
|
||
<bibRefCitation id="EFFD4B204313FFF3FB47F94E4B28F8DE" author="RAGE J. - C. & SZYNDLAR Z." pageId="23" pageNumber="364" pagination="56 - 64" refId="ref26588" refString="RAGE J. - C. & SZYNDLAR Z. 1986. - Natrix longivertebrata from the European Neogene, a snake with one of the longest known stratigraphic ranges. Neues Jahrbuch fur Geologie und Palaontologie, Mh. 1: 56 - 64. https: // doi. org / 10.1127 / njgpm / 1986 / 1986 / 56" type="journal article" year="1986">Rage & Szyndlar 1986</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFFD4B204313FFF3FCC9F96E4C6CF8DE" author="SZYNDLAR Z." box="[885,1056,1778,1804]" pageId="23" pageNumber="364" pagination="412 - 418" refId="ref27527" refString="SZYNDLAR Z. 1991 c. - Ancestry of the Grass Snake (Natrix natrix): Paleontological evidence. Journal of Herpetology 25 (4): 412 - 418. https: // doi. org / 10.2307 / 1564762" type="journal article" year="1991">Szyndlar 1991c</bibRefCitation>
|
||
), has more elongated vertebrae and a much lower neural spine.
|
||
<taxonomicName id="4C6C4D524313FFF3FBE2F88E4CEEF8F9" authorityName="Laurenti" authorityYear="1768" box="[1118,1186,1810,1835]" class="Reptilia" family="Colubridae" genus="Natrix" kingdom="Animalia" order="Squamata" pageId="23" pageNumber="364" phylum="Chordata" rank="genus">
|
||
<emphasis id="B918EAC34313FFF3FBE2F88E4CEEF8F9" box="[1118,1186,1810,1835]" italics="true" pageId="23" pageNumber="364">Natrix</emphasis>
|
||
</taxonomicName>
|
||
sp. closely resembles the extinct
|
||
<taxonomicName id="4C6C4D524313FFF3FCC2F8AE4CAAF89E" authority="Szyndlar, 2005" authorityName="Szyndlar" authorityYear="2005" box="[894,1254,1841,1868]" class="Reptilia" family="Colubridae" genus="Natrix" kingdom="Animalia" order="Squamata" pageId="23" pageNumber="364" phylum="Chordata" rank="species" species="rudabanyaensis">
|
||
<emphasis id="B918EAC34313FFF3FCC2F8AE4C71F899" box="[894,1085,1841,1867]" italics="true" pageId="23" pageNumber="364">N. rudabanyaensis</emphasis>
|
||
<bibRefCitation id="EFFD4B204313FFF3FBF9F8AD4CAAF89E" author="SZYNDLAR Z." box="[1093,1254,1841,1868]" pageId="23" pageNumber="364" pagination="31 - 52" refId="ref27643" refString="SZYNDLAR Z. 2005. - Snake fauna from the Late Miocene of Rudabanya. Palaeontographia Italica 90 (2003): 31 - 52." type="journal article" year="2005">Szyndlar, 2005</bibRefCitation>
|
||
</taxonomicName>
|
||
reported from the early late Miocene of Rudabánya,
|
||
<collectingCountry id="F37B76414313FFF3FB27F8CE4CB6F8BE" box="[1179,1274,1874,1900]" name="Hungary" pageId="23" pageNumber="364">Hungary</collectingCountry>
|
||
(MN 9a;
|
||
<bibRefCitation id="EFFD4B204313FFF3FADAF8CD4BC9F85E" author="SZYNDLAR Z." pageId="23" pageNumber="364" pagination="31 - 52" refId="ref27643" refString="SZYNDLAR Z. 2005. - Snake fauna from the Late Miocene of Rudabanya. Palaeontographia Italica 90 (2003): 31 - 52." type="journal article" year="2005">Szyndlar 2005</bibRefCitation>
|
||
) and perhaps the middle Miocene of Tauţ,
|
||
<collectingCountry id="F37B76414313FFF3FAF3F8ED4DE3F859" box="[1359,1455,1905,1931]" name="Romania" pageId="23" pageNumber="364">Romania</collectingCountry>
|
||
(MN 7+8;
|
||
<bibRefCitation id="EFFD4B204313FFF3FC22F80D4CC2F879" author="VENCZEL M. & STIUCA E." box="[926,1166,1937,1963]" pageId="23" pageNumber="364" pagination="731 - 763" refId="ref28185" refString="VENCZEL M. & STIUCA E. 2008. - Late middle Miocene amphibians and squamate reptiles from Taut, Romania. Geodiversitas 30 (4): 731 - 763." type="journal article" year="2008">Venczel & Ştiucă 2008</bibRefCitation>
|
||
) in: 1) the same height and shape of the neural spine; 2) moderately developed subcentral ridges; 3) elongated prezygapophyseal facets; and 4) flattened prezygapophyseal processes (
|
||
<bibRefCitation id="EFFD4B20431CFFFCFE02FF4B4A10FF20" author="SZYNDLAR Z." box="[446,604,215,242]" pageId="24" pageNumber="365" pagination="31 - 52" refId="ref27643" refString="SZYNDLAR Z. 2005. - Snake fauna from the Late Miocene of Rudabanya. Palaeontographia Italica 90 (2003): 31 - 52." type="journal article" year="2005">Szyndlar 2005</bibRefCitation>
|
||
). However, the absence of the hypapophysis and paradiapophyses prevents alpha taxonomy.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |