treatments-xml/data/03/AE/8A/03AE8A2DFFA8CF75FF47ED64FD31F829.xml
2024-06-21 12:22:17 +02:00

401 lines
88 KiB
XML
Raw Blame History

This file contains ambiguous Unicode characters

This file contains Unicode characters that might be confused with other characters. If you think that this is intentional, you can safely ignore this warning. Use the Escape button to reveal them.

<document id="D42D79ACE721D2525D3BDFDAEAD60225" ID-DOI="10.11646/zootaxa.5375.3.1" ID-ISSN="1175-5326" ID-Zenodo-Dep="10200036" ID-ZooBank="27D02F09-01B7-457A-8A99-D8644B7B6ADE" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1700728652862" checkinUser="plazi" docAuthor="Arimoto, Kôichi" docDate="2023" docId="03AE8A2DFFA8CF75FF47ED64FD31F829" docLanguage="en" docName="zootaxa.5375.3.1.pdf" docOrigin="Zootaxa 5375 (3)" docSource="https://mapress.com/zt/article/download/zootaxa.5375.3.1/52332" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Penia inopinata Arimoto 2023, sp. nov." docType="treatment" docVersion="2" lastPageNumber="318" masterDocId="FF97F255FFA6CF64FFD0EB1BFFFFFFED" masterDocTitle="Taxonomic revision of the genus Penia Laporte (Coleoptera, Elateridae, Dendrometrinae) from Taiwan" masterLastPageNumber="335" masterPageNumber="301" pageNumber="315" updateTime="1700729251563" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
<mods:mods id="806CC00A8D77B94D74BD40EA401434C2" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="DBE7EA5432726F5B4690241A673F9940">
<mods:title id="A567D6459A5C89FA9E51FF9DF609C338">Taxonomic revision of the genus Penia Laporte (Coleoptera, Elateridae, Dendrometrinae) from Taiwan</mods:title>
</mods:titleInfo>
<mods:name id="82244D2C319099996BC4EF0801386896" type="personal">
<mods:role id="C22AC7FAFADCC6685D0D5EB52FA78DB8">
<mods:roleTerm id="2D08A51FA1DE3C1BDD4A3BF587DCAF4F">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="1B38942B8B5F36F149A37B0D28DB5E7B">Arimoto, Kôichi</mods:namePart>
</mods:name>
<mods:typeOfResource id="7605C75F34F6CA6A98231459CA242DBA">text</mods:typeOfResource>
<mods:relatedItem id="F4E88FA75485487CFCFFB22AB1FB3F83" type="host">
<mods:titleInfo id="148D63ACEF558651541F2927CC7541C9">
<mods:title id="D105F6EF0FD219B636077D4FB0E523A8">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="21DCF222D84810D5A874A5E5AE995CF6">
<mods:date id="9403122A700D93F74C497AD86D15318F">2023</mods:date>
<mods:detail id="22DAEF89B0015741C3AB3685472CF483" type="pubDate">
<mods:number id="9EC374148254C0C8CC63BED24BACF17F">2023-11-23</mods:number>
</mods:detail>
<mods:detail id="8506112FA0E92D6333DC7906B9814D24" type="volume">
<mods:number id="29F21CFDBCE5CD105B58C3E0E074DF1A">5375</mods:number>
</mods:detail>
<mods:detail id="7B83593D212125E67C4353D38AFDC310" type="issue">
<mods:number id="6FF98ABD909D6ADA3DA4955D78C82001">3</mods:number>
</mods:detail>
<mods:extent id="692B3498793779D37525E9ECFCDB6F6C" unit="page">
<mods:start id="727DFEB7922482D11F2F0BC485816053">301</mods:start>
<mods:end id="44C30B3FA5111034B30C25D0AF714C28">335</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location id="687D82DE8FD4664EF914345F88CB1615">
<mods:url id="A437EC34840FF6DE7255142CD3E2372E">https://mapress.com/zt/article/download/zootaxa.5375.3.1/52332</mods:url>
</mods:location>
<mods:classification id="21F776B9294EF31732393D366CF4AC7D">journal article</mods:classification>
<mods:identifier id="5BF63DE38D57DD5AC416874C71437607" type="DOI">10.11646/zootaxa.5375.3.1</mods:identifier>
<mods:identifier id="EE2DAD70BDCC59BB57BA7E45692D5825" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="DF569C127D5F62F8F94EBC54823A9ADB" type="Zenodo-Dep">10200036</mods:identifier>
<mods:identifier id="D17665DFC61D77920A2A78F1D9409515" type="ZooBank">27D02F09-01B7-457A-8A99-D8644B7B6ADE</mods:identifier>
</mods:mods>
<treatment id="03AE8A2DFFA8CF75FF47ED64FD31F829" LSID="urn:lsid:plazi:treatment:03AE8A2DFFA8CF75FF47ED64FD31F829" httpUri="http://treatment.plazi.org/id/03AE8A2DFFA8CF75FF47ED64FD31F829" lastPageId="17" lastPageNumber="318" pageId="14" pageNumber="315">
<subSubSection id="C31D68B0FFA8CF6AFF47ED64FDCBF974" box="[151,564,1662,1689]" pageId="14" pageNumber="315" type="nomenclature">
<paragraph id="8BB83B3BFFA8CF6AFF47ED64FDCBF974" blockId="14.[151,564,1662,1725]" box="[151,564,1662,1689]" pageId="14" pageNumber="315">
<heading id="D0F08C57FFA8CF6AFF47ED64FDCBF974" bold="true" box="[151,564,1662,1689]" fontSize="11" level="1" pageId="14" pageNumber="315" reason="1">
<emphasis id="B973E729FFA8CF6AFF47ED64FDCBF974" bold="true" box="[151,564,1662,1689]" pageId="14" pageNumber="315">
<taxonomicName id="4C0740B8FFA8CF6AFF47ED64FE35F975" authority="Arimoto" authorityName="Arimoto" authorityYear="2023" box="[151,458,1662,1689]" class="Insecta" family="Elateridae" genus="Penia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="315" phylum="Arthropoda" rank="species" species="inopinata" status="sp. nov.">
<emphasis id="B973E729FFA8CF6AFF47ED64FEA6F974" bold="true" box="[151,345,1663,1689]" italics="true" pageId="14" pageNumber="315">Penia inopinata</emphasis>
Arimoto
</taxonomicName>
,
<taxonomicNameLabel id="A2405A52FFA8CF6AFE07ED64FDCBF974" box="[471,564,1663,1689]" pageId="14" pageNumber="315" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C31D68B0FFA8CF6AFF47EDB8FEC0F950" box="[151,319,1699,1725]" pageId="14" pageNumber="315" type="description">
<paragraph id="8BB83B3BFFA8CF6AFF47EDB8FEC0F950" blockId="14.[151,564,1662,1725]" box="[151,319,1699,1725]" pageId="14" pageNumber="315">
(
<figureCitation id="133C27BEFFA8CF6AFF4FEDB8FEF0F950" box="[159,271,1699,1725]" captionStart="FIGURE 10" captionStartId="15.[151,250,1654,1678]" captionTargetBox="[151,1436,186,1630]" captionTargetId="figure-16@15.[151,1436,181,1630]" captionTargetPageId="15" captionText="FIGURE 10. Penia inopinata, holotype, female (6349) (= paratype of Penia takasago). A: labels; B: habitus, dorsal side; C: habitus, ventral side; D: head, anterior view; E: head, dorsal view; F: basal antennomeres of left antenna; G: left maxillary palp, dorsal side; H: head, prothorax and mesothorax, lateral view (black arrow: the direction perpendicular to the base of ventral lobe of prosternal process, white arrow: the direction perpendicular to the mesosternal process between mesocoxae); I: left hypomeron (arrow: hypomeron with slight mesial projection); J: prosternal process from the direction perpendicular to base of ventral lobe (black arrow of figure 10H); K: scutellar shield; L: mesothorax from the direction perpendicular to mesosternal process between mesocoxae (white arrow of figure 10H); M: left mid tarsus." figureDoi="http://doi.org/10.5281/zenodo.10200057" httpUri="https://zenodo.org/record/10200057/files/figure.png" pageId="14" pageNumber="315">Figure 10</figureCitation>
,
<figureCitation id="133C27BEFFA8CF6AFECBEDB8FEC9F950" box="[283,310,1699,1725]" captionStart="FIGURE 11" captionStartId="17.[151,250,1351,1375]" captionTargetBox="[151,1436,741,1328]" captionTargetId="figure-276@17.[151,1436,741,1328]" captionTargetPageId="17" captionText="FIGURE 11. Penia inopinata, holotype, female (6349) (= paratype of Penia takasago). A: pregenital segments, ventral side; B: slide; C: pregenital segments and ovipositor, ventral side; D: coxites, ventral side." figureDoi="http://doi.org/10.5281/zenodo.10200059" httpUri="https://zenodo.org/record/10200059/files/figure.png" pageId="14" pageNumber="315">11</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C31D68B0FFA8CF6AFF47EDF3FAE8F912" box="[151,1303,1767,1792]" pageId="14" pageNumber="315" type="reference_group">
<paragraph id="8BB83B3BFFA8CF6AFF47EDF3FAE8F912" blockId="14.[151,1303,1767,1792]" box="[151,1303,1767,1792]" pageId="14" pageNumber="315">
<treatmentCitationGroup id="AB171C15FFA8CF6AFF47EDF3FAE8F912" box="[151,1303,1767,1792]" pageId="14" pageNumber="315">
<treatmentCitation id="0AA61D2AFFA8CF6AFF47EDF3FE21F912" author="Kishii, T." box="[151,478,1767,1791]" page="12" pageId="14" pageNumber="315" year="1997">
<taxonomicName id="4C0740B8FFA8CF6AFF47EDF3FE21F912" authority="Kishii, 1997: 12" authorityName="Kishii" authorityPageNumber="12" authorityYear="1997" box="[151,478,1767,1791]" class="Insecta" family="Elateridae" genus="Penia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="315" phylum="Arthropoda" rank="species" species="takasago">
<emphasis id="B973E729FFA8CF6AFF47EDF3FECBF912" box="[151,308,1767,1791]" italics="true" pageId="14" pageNumber="315">Penia takasago</emphasis>
<materialsCitation id="3B6F3166FFA8CF6AFEEAEDFCFE21F912" box="[314,478,1767,1791]" country="Taiwan" location="Sungkang" pageId="14" pageNumber="315" specimenCount="1" stateProvince="Nantou">
<bibRefCitation id="EF9646CAFFA8CF6AFEEAEDFCFE21F912" author="Kishii, T." box="[314,478,1767,1791]" pageId="14" pageNumber="315" pagination="9 - 14" refId="ref23002" refString="Kishii, T. (1997) A study on the elaterid-beetles of Shibata collection from Taiwan. V. (Coleoptera: Elateridae) On the subfamily Denticollinae: tribes Denticollini, Hemicrepidiini and Dimini. The Entomological Review of Japan, 52, 9 - 14." type="journal article" year="1997">Kishii, 1997: 12</bibRefCitation>
</materialsCitation>
</taxonomicName>
</treatmentCitation>
(original description;
<typeStatus id="54BC8599FFA8CF6AFD10EDF3FD13F8ED" box="[704,748,1768,1792]" pageId="14" pageNumber="315">type</typeStatus>
locality:
<collectingCountry id="F3107BABFFA8CF6AFC9BEDFCFC67F912" box="[843,920,1767,1791]" name="Taiwan" pageId="14" pageNumber="315">Taiwan</collectingCountry>
,
<collectingRegion id="49C3F5D9FFA8CF6AFC72EDF3FBC0F912" box="[930,1087,1767,1791]" country="Taiwan" name="Nantou" pageId="14" pageNumber="315">Nantou County</collectingRegion>
,
<location id="8ED86DE0FFA8CF6AFB98EDFCFB50F912" LSID="urn:lsid:plazi:treatment:03AE8A2DFFA8CF75FF47ED64FD31F829:8ED86DE0FFA8CF6AFB98EDFCFB50F912" box="[1096,1199,1767,1791]" country="Taiwan" name="Sungkang" pageId="14" pageNumber="315" stateProvince="Nantou">Sungkang</location>
) [partim].
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C31D68B0FFA8CF6AFF47EC36FB70F8AA" box="[151,1167,1837,1863]" pageId="14" pageNumber="315" type="etymology">
<paragraph id="8BB83B3BFFA8CF6AFF47EC36FB70F8AA" blockId="14.[151,1436,1837,2008]" box="[151,1167,1837,1863]" pageId="14" pageNumber="315">
<emphasis id="B973E729FFA8CF6AFF47EC36FEE1F8AA" bold="true" box="[151,286,1837,1863]" pageId="14" pageNumber="315">Etymology.</emphasis>
Specific epithet derived from the circumstances of the discover of the species.
</paragraph>
</subSubSection>
<subSubSection id="C31D68B0FFA8CF74FF17EC4AFAD4FCD4" lastPageId="16" lastPageNumber="317" pageId="14" pageNumber="315" type="materials_examined">
<paragraph id="8BB83B3BFFA8CF6AFF17EC4AFDEAF83A" blockId="14.[151,1436,1837,2008]" pageId="14" pageNumber="315">
<emphasis id="B973E729FFA8CF6AFF17EC4AFE8BF886" bold="true" box="[199,372,1873,1899]" pageId="14" pageNumber="315">Type material.</emphasis>
<materialsCitation id="3B6F3166FFA8CF6AFEAAEC4AFDEAF83A" collectingDate="1970-05-04" collectingDateMax="1970-05-08" collectingDateMin="1970-05-04" collectionCode="OMNH" collectorName="Y. Kiyoyama" country="Taiwan" location="Zhuqi Township" pageId="14" pageNumber="315" specimenCount="2" stateProvince="Chiayi" typeStatus="holotype">
<emphasis id="B973E729FFA8CF6AFEAAEC4AFE11F886" bold="true" box="[378,494,1873,1899]" pageId="14" pageNumber="315">
<typeStatus id="54BC8599FFA8CF6AFEAAEC4AFE16F886" box="[378,489,1873,1899]" pageId="14" pageNumber="315" type="holotype">Holotype</typeStatus>
.
</emphasis>
Female, Taiwan,
<collectingRegion id="49C3F5D9FFA8CF6AFD61EC4AFCAAF886" box="[689,853,1873,1899]" country="Taiwan" name="Chiayi" pageId="14" pageNumber="315">Chiayi County</collectingRegion>
,
<location id="8ED86DE0FFA8CF6AFC8FEC4AFBE7F886" LSID="urn:lsid:plazi:treatment:03AE8A2DFFA8CF75FF47ED64FD31F829:8ED86DE0FFA8CF6AFC8FEC4AFBE7F886" box="[863,1048,1873,1899]" country="Taiwan" name="Zhuqi Township" pageId="14" pageNumber="315" stateProvince="Chiayi">Zhuqi Township</location>
, Fenqihu,
<date id="FFB91DFBFFA8CF6AFB5AEC4AFB10F886" box="[1162,1263,1873,1900]" pageId="14" pageNumber="315" value="1970-05-04">
<collectingDate id="EFFDE413FFA8CF6AFB5AEC4AFB10F886" box="[1162,1263,1873,1900]" pageId="14" pageNumber="315" value="1970-05-04">4 V 1970</collectingDate>
</date>
(as
<date id="FFB91DFBFFA8CF6AFACCEC4AFA7FF886" box="[1308,1408,1873,1900]" pageId="14" pageNumber="315" value="1970-05-08">
<collectingDate id="EFFDE413FFA8CF6AFACCEC4AFA7FF886" box="[1308,1408,1873,1900]" pageId="14" pageNumber="315" value="1970-05-08">
8 V
<quantity id="4CFF96DEFFA8CF6AFA98EC4AFA7FF886" box="[1352,1408,1873,1899]" metricMagnitude="1" metricUnit="m" metricValue="5.0038" pageId="14" pageNumber="315" unit="in" value="1970.0">1970</quantity>
</collectingDate>
</date>
in original description of
<taxonomicName id="4C0740B8FFA8CF6AFE45EC6DFDBDF862" authorityName="Kishii" authorityYear="1997" box="[405,578,1909,1935]" class="Insecta" family="Elateridae" genus="Penia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="315" phylum="Arthropoda" rank="species" species="takasago">
<emphasis id="B973E729FFA8CF6AFE45EC6DFDBDF862" box="[405,578,1909,1935]" italics="true" pageId="14" pageNumber="315">Penia takasago</emphasis>
</taxonomicName>
),
<collectorName id="26F25EEDFFA8CF6AFD88EC6DFD14F862" box="[600,747,1909,1936]" pageId="14" pageNumber="315">Y. Kiyoyama</collectorName>
leg. [
<collectionCode id="ED16A3FEFFA8CF6AFCF7EC6EFC7CF862" box="[807,899,1909,1935]" country="Japan" httpUri="http://grbio.org/cool/rcjv-ktyc" name="Osaka Museum of Natural History" pageId="14" pageNumber="315" type="Museum">OMNH</collectionCode>
; 6349]. Verbatim label data (
<figureCitation id="133C27BEFFA8CF6AFB13EC6EFAD5F862" box="[1219,1322,1909,1935]" captionStart="FIGURE 10" captionStartId="15.[151,250,1654,1678]" captionTargetBox="[151,1436,186,1630]" captionTargetId="figure-16@15.[151,1436,181,1630]" captionTargetPageId="15" captionText="FIGURE 10. Penia inopinata, holotype, female (6349) (= paratype of Penia takasago). A: labels; B: habitus, dorsal side; C: habitus, ventral side; D: head, anterior view; E: head, dorsal view; F: basal antennomeres of left antenna; G: left maxillary palp, dorsal side; H: head, prothorax and mesothorax, lateral view (black arrow: the direction perpendicular to the base of ventral lobe of prosternal process, white arrow: the direction perpendicular to the mesosternal process between mesocoxae); I: left hypomeron (arrow: hypomeron with slight mesial projection); J: prosternal process from the direction perpendicular to base of ventral lobe (black arrow of figure 10H); K: scutellar shield; L: mesothorax from the direction perpendicular to mesosternal process between mesocoxae (white arrow of figure 10H); M: left mid tarsus." figureDoi="http://doi.org/10.5281/zenodo.10200057" httpUri="https://zenodo.org/record/10200057/files/figure.png" pageId="14" pageNumber="315">Fig. 10A</figureCitation>
). “Female symbol”; “6349”; “
<typeStatus id="54BC8599FFA8CF6AFEA7EC82FDF4F85E" box="[375,523,1945,1971]" pageId="14" pageNumber="315" type="paratype">PARATYPE</typeStatus>
”; “FUNCHIFO/
<collectingCountry id="F3107BABFFA8CF6AFD01EC82FCA7F85E" box="[721,856,1945,1971]" name="Taiwan" pageId="14" pageNumber="315">FORMOSA</collectingCountry>
/
<date id="FFB91DFBFFA8CF6AFCBBEC82FC19F85E" box="[875,998,1945,1972]" pageId="14" pageNumber="315" value="1970-05-04">
<collectingDate id="EFFDE413FFA8CF6AFCBBEC82FC19F85E" box="[875,998,1945,1972]" pageId="14" pageNumber="315" value="1970-05-04">4. V. 1970</collectingDate>
</date>
/ Y. KIYOYAMA”; “
<taxonomicName id="4C0740B8FFA8CF6AFB0EEC82FAE1F85E" box="[1246,1310,1945,1971]" class="Insecta" family="Elateridae" genus="Penia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="335" phylum="Arthropoda" rank="genus">Penia</taxonomicName>
/ takasago/ KISHII,
<taxonomicNameLabel id="A2405A52FFA8CF6AFF29ECA5FECCF83A" box="[249,307,1981,2008]" pageId="14" pageNumber="315" rank="species">sp. n.</taxonomicNameLabel>
/ Det. KISHII, 92”.
</materialsCitation>
</paragraph>
<caption id="DF786BB3FFA9CF6BFF47ED6DFC60F88B" ID-DOI="http://doi.org/10.5281/zenodo.10200057" ID-Zenodo-Dep="10200057" httpUri="https://zenodo.org/record/10200057/files/figure.png" pageId="15" pageNumber="316" startId="15.[151,250,1654,1678]" targetBox="[151,1436,186,1630]" targetPageId="15" targetType="figure">
<paragraph id="8BB83B3BFFA9CF6BFF47ED6DFC60F88B" blockId="15.[151,1437,1654,1894]" pageId="15" pageNumber="316">
<emphasis id="B973E729FFA9CF6BFF47ED6DFEDEF963" bold="true" box="[151,289,1654,1678]" pageId="15" pageNumber="316">FIGURE 10.</emphasis>
<taxonomicName id="4C0740B8FFA9CF6BFEF9ED6CFE33F963" authorityName="Arimoto" authorityYear="2023" box="[297,460,1655,1678]" class="Insecta" family="Elateridae" genus="Penia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="316" phylum="Arthropoda" rank="species" species="inopinata">
<emphasis id="B973E729FFA9CF6BFEF9ED6CFE33F963" box="[297,460,1655,1678]" italics="true" pageId="15" pageNumber="316">Penia inopinata</emphasis>
</taxonomicName>
, holotype, female (6349) (= paratype of
<taxonomicName id="4C0740B8FFA9CF6BFCA3ED6CFBEFF963" authorityName="Kishii" authorityYear="1997" box="[883,1040,1654,1678]" class="Insecta" family="Elateridae" genus="Penia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="316" phylum="Arthropoda" rank="species" species="takasago">
<emphasis id="B973E729FFA9CF6BFCA3ED6CFBEFF963" box="[883,1040,1654,1678]" italics="true" pageId="15" pageNumber="316">Penia takasago</emphasis>
</taxonomicName>
). A: labels; B: habitus, dorsal side; C: habitus, ventral side; D: head, anterior view; E: head, dorsal view; F: basal antennomeres of left antenna; G: left maxillary palp, dorsal side; H: head, prothorax and mesothorax, lateral view (black arrow: the direction perpendicular to the base of ventral lobe of prosternal process, white arrow: the direction perpendicular to the mesosternal process between mesocoxae); I: left hypomeron (arrow: hypomeron with slight mesial projection); J: prosternal process from the direction perpendicular to base of ventral lobe (black arrow of figure 10H); K: scutellar shield; L: mesothorax from the direction perpendicular to mesosternal process between mesocoxae (white arrow of figure 10H); M: left mid tarsus.
</paragraph>
</caption>
<paragraph id="8BB83B3BFFA9CF74FF17EC8EFD90FE40" blockId="15.[151,1436,1940,2039]" lastBlockId="16.[151,1437,151,2013]" lastPageId="16" lastPageNumber="317" pageId="15" pageNumber="316">
<emphasis id="B973E729FFA9CF6BFF17EC8EFE59F843" bold="true" box="[199,422,1940,1967]" pageId="15" pageNumber="316">Female. Diagnosis.</emphasis>
This species is characterized by the following features: eyes 0.25 x longer than interocular distance in dorsal view; antennae extending beyond pronotum posterior lateral apices by antennomere VII; antennomeres
<collectionCode id="ED16A3FEFFA9CF6BFEE8ECC6FEABF81A" box="[312,340,2013,2039]" pageId="15" pageNumber="316">III</collectionCode>
distinctly longer than
<collectionCode id="ED16A3FEFFA9CF6BFD98ECC6FDA1F81A" box="[584,606,2013,2039]" pageId="15" pageNumber="316">II</collectionCode>
;
<collectionCode id="ED16A3FEFFA9CF6BFDB8ECC6FD7AF81A" box="[616,645,2013,2039]" pageId="15" pageNumber="316">IV</collectionCode>
1.2 x longer than
<collectionCode id="ED16A3FEFFA9CF6BFC97ECC6FC99F81A" box="[839,870,2013,2039]" pageId="15" pageNumber="316">III</collectionCode>
, 0.8 x longer than
<collectionCode id="ED16A3FEFFA9CF6BFBFCECC6FBC1F81A" box="[1068,1086,2013,2039]" pageId="15" pageNumber="316">II</collectionCode>
<collectionCode id="ED16A3FEFFA9CF6BFB9DECC6FB96F81A" box="[1101,1129,2013,2039]" pageId="15" pageNumber="316">III</collectionCode>
combined; apical maxillary palpomere 2.1 x longer than wide, almost as long as maximum length of eye; pronotum straightly and slightly narrowed ahead of hind angles; posterior edge of pronotum with sublateral incisions; hind angles of pronotum broad, not protruding laterally beyond pronotal side; hypomeron with slight mesial projection; anterior angle of hypomeron rounded; hind angle of hypomeron broadly triangular; scutellar shield almost as long as wide; mesosternal process between mesocoxae a little higher than mesocoxae, visible in lateral view; posterior edge of mesosternal process 0.15 x wider than total width of mesosternum; elytron 3.4 x longer than wide, 2.7 x longer than pronotum length; abdominal ventrite
<collectionCode id="ED16A3FEFFB6CF74FEA3EA74FE78FE64" box="[371,391,367,393]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="16" pageNumber="317" type="Museum">V</collectionCode>
semicircular, rounded apically; spiculum ventrale 5.45 x longer than length of sternite VIII; ovipositor longer than length of abdomen.
</paragraph>
<paragraph id="8BB83B3BFFB6CF74FF17EAACFAD4FCD4" blockId="16.[151,1437,151,2013]" pageId="16" pageNumber="317">
This species is more similar to
<taxonomicName id="4C0740B8FFB6CF74FDE2EAACFD6BFE3C" authorityName="Kishii" authorityYear="1994" box="[562,660,439,465]" class="Insecta" family="Elateridae" genus="Penia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="317" phylum="Arthropoda" rank="species" species="babai">
<emphasis id="B973E729FFB6CF74FDE2EAACFDBCFE3D" box="[562,579,439,464]" italics="true" pageId="16" pageNumber="317">P</emphasis>
.
<emphasis id="B973E729FFB6CF74FD84EAACFD6BFE3C" box="[596,660,439,465]" italics="true" pageId="16" pageNumber="317">babai</emphasis>
</taxonomicName>
than to
<taxonomicName id="4C0740B8FFB6CF74FD26EAACFC82FE3C" authorityName="Kishii" authorityYear="1997" box="[758,893,439,465]" class="Insecta" family="Elateridae" genus="Penia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="317" phylum="Arthropoda" rank="species" species="takasago">
<emphasis id="B973E729FFB6CF74FD26EAACFCF8FE3D" box="[758,775,439,464]" italics="true" pageId="16" pageNumber="317">P</emphasis>
.
<emphasis id="B973E729FFB6CF74FCC8EAACFC82FE3C" box="[792,893,439,465]" italics="true" pageId="16" pageNumber="317">takasago</emphasis>
</taxonomicName>
in the eye size, the relative length of the basal antennomeres, the length-to-width ratios of the apical maxillary palpomere, scutellar shield and elytron, the shapes of the hind angle of the hypomeron and abdominal ventrite
<collectionCode id="ED16A3FEFFB6CF74FCF0EAE4FCCBFDF4" box="[800,820,511,537]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="16" pageNumber="317" type="Museum">V</collectionCode>
, and the lengths of the spiculum ventrale and ovipositor. It is distinguished from
<taxonomicName id="4C0740B8FFB6CF74FE64E938FDE8FDD0" authorityName="Kishii" authorityYear="1994" box="[436,535,547,573]" class="Insecta" family="Elateridae" genus="Penia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="317" phylum="Arthropoda" rank="species" species="babai">
<emphasis id="B973E729FFB6CF74FE64E938FE3AFDD1" box="[436,453,547,572]" italics="true" pageId="16" pageNumber="317">P</emphasis>
.
<emphasis id="B973E729FFB6CF74FE07E938FDE8FDD0" box="[471,535,547,573]" italics="true" pageId="16" pageNumber="317">babai</emphasis>
</taxonomicName>
by the following contrasting characteristics (
<taxonomicName id="4C0740B8FFB6CF74FBFBE938FB71FDD0" authorityName="Kishii" authorityYear="1994" box="[1067,1166,547,573]" class="Insecta" family="Elateridae" genus="Penia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="317" phylum="Arthropoda" rank="species" species="babai">
<emphasis id="B973E729FFB6CF74FBFBE938FBC3FDD1" box="[1067,1084,547,572]" italics="true" pageId="16" pageNumber="317">P</emphasis>
.
<emphasis id="B973E729FFB6CF74FB9EE938FB71FDD0" box="[1102,1166,547,573]" italics="true" pageId="16" pageNumber="317">babai</emphasis>
</taxonomicName>
in parentheses): apical maxillary palpomere almost as long as maximum length of eye (apical maxillary palpomere shorter than maximum length of eye); hind angles of pronotum not protruding laterally beyond pronotal side (hind angles of pronotum weakly protruding posterolaterad); anterior angle of hypomeron rounded (anterior angle of hypomeron nearly right angle); hypomeron with slight mesial projection (hypomeron with distinct mesial projection); mesosternal process between mesocoxae a little higher than mesocoxae (mesosternal process between mesocoxae distinctly higher than mesocoxae); posterior edge of mesosternum between mesocoxae 0.15 x wider than total width of mesosternum (posterior edge of mesosternum between mesocoxae 0.20.25 x wider than total width of mesosternum).
</paragraph>
</subSubSection>
<subSubSection id="C31D68B0FFB6CF75FF17E858FE77FA21" lastPageId="17" lastPageNumber="318" pageId="16" pageNumber="317" type="description">
<paragraph id="8BB83B3BFFB6CF74FF17E858FC91FC6C" blockId="16.[151,1437,151,2013]" pageId="16" pageNumber="317">
<emphasis id="B973E729FFB6CF74FF17E858FE81FCB0" bold="true" box="[199,382,835,861]" pageId="16" pageNumber="317">Measurements.</emphasis>
BL: 8.96, BW: 3.63, MAE: 1.54, MBE: 1.02, OI: 151, PL: 2.30, PML: 1.80, PW: 2.77, PAW: 1.72, PLI: 83.3, PWI: 161, EL: 6.17, EW: 1.83, EI: 336, BI: 268.
</paragraph>
<paragraph id="8BB83B3BFFB6CF74FF17E891FDC0FBFC" blockId="16.[151,1437,151,2013]" pageId="16" pageNumber="317">
<emphasis id="B973E729FFB6CF74FF17E891FEA6FC49" bold="true" box="[199,345,906,932]" pageId="16" pageNumber="317">Description.</emphasis>
Body broad, widest around elytral midlength (
<figureCitation id="133C27BEFFB6CF74FCA3E890FC24FC48" box="[883,987,907,933]" captionStart="FIGURE 10" captionStartId="15.[151,250,1654,1678]" captionTargetBox="[151,1436,186,1630]" captionTargetId="figure-16@15.[151,1436,181,1630]" captionTargetPageId="15" captionText="FIGURE 10. Penia inopinata, holotype, female (6349) (= paratype of Penia takasago). A: labels; B: habitus, dorsal side; C: habitus, ventral side; D: head, anterior view; E: head, dorsal view; F: basal antennomeres of left antenna; G: left maxillary palp, dorsal side; H: head, prothorax and mesothorax, lateral view (black arrow: the direction perpendicular to the base of ventral lobe of prosternal process, white arrow: the direction perpendicular to the mesosternal process between mesocoxae); I: left hypomeron (arrow: hypomeron with slight mesial projection); J: prosternal process from the direction perpendicular to base of ventral lobe (black arrow of figure 10H); K: scutellar shield; L: mesothorax from the direction perpendicular to mesosternal process between mesocoxae (white arrow of figure 10H); M: left mid tarsus." figureDoi="http://doi.org/10.5281/zenodo.10200057" httpUri="https://zenodo.org/record/10200057/files/figure.png" pageId="16" pageNumber="317">Fig. 10B</figureCitation>
); surface generally smooth; interspaces between punctures distinctly larger than fine puncture diameter (
<figureCitation id="133C27BEFFB6CF74FC5AE8B4FBE7FC24" box="[906,1048,943,969]" captionStart="FIGURE 10" captionStartId="15.[151,250,1654,1678]" captionTargetBox="[151,1436,186,1630]" captionTargetId="figure-16@15.[151,1436,181,1630]" captionTargetPageId="15" captionText="FIGURE 10. Penia inopinata, holotype, female (6349) (= paratype of Penia takasago). A: labels; B: habitus, dorsal side; C: habitus, ventral side; D: head, anterior view; E: head, dorsal view; F: basal antennomeres of left antenna; G: left maxillary palp, dorsal side; H: head, prothorax and mesothorax, lateral view (black arrow: the direction perpendicular to the base of ventral lobe of prosternal process, white arrow: the direction perpendicular to the mesosternal process between mesocoxae); I: left hypomeron (arrow: hypomeron with slight mesial projection); J: prosternal process from the direction perpendicular to base of ventral lobe (black arrow of figure 10H); K: scutellar shield; L: mesothorax from the direction perpendicular to mesosternal process between mesocoxae (white arrow of figure 10H); M: left mid tarsus." figureDoi="http://doi.org/10.5281/zenodo.10200057" httpUri="https://zenodo.org/record/10200057/files/figure.png" pageId="16" pageNumber="317">Fig. 10B, C</figureCitation>
). Color. Body, antennae and legs reddish brown. External edge of mandible, posterior edges of prosternum and anterior edge of scutellum black. Body covered with long yellow setae.
</paragraph>
<paragraph id="8BB83B3BFFB6CF74FF17EF00FD51FAB8" blockId="16.[151,1437,151,2013]" pageId="16" pageNumber="317">
<materialsCitation id="3B6F3166FFB6CF74FF17EF00FD55FAB8" collectionCode="II, III, IV" pageId="16" pageNumber="317" specimenCode="IV 3.2, V 3.4" specimenCount="1" typeStatus="holotype">
Head. Frons flatted medially (
<figureCitation id="133C27BEFFB6CF74FDCAEF00FD5CFBD8" box="[538,675,1051,1077]" captionStart="FIGURE 10" captionStartId="15.[151,250,1654,1678]" captionTargetBox="[151,1436,186,1630]" captionTargetId="figure-16@15.[151,1436,181,1630]" captionTargetPageId="15" captionText="FIGURE 10. Penia inopinata, holotype, female (6349) (= paratype of Penia takasago). A: labels; B: habitus, dorsal side; C: habitus, ventral side; D: head, anterior view; E: head, dorsal view; F: basal antennomeres of left antenna; G: left maxillary palp, dorsal side; H: head, prothorax and mesothorax, lateral view (black arrow: the direction perpendicular to the base of ventral lobe of prosternal process, white arrow: the direction perpendicular to the mesosternal process between mesocoxae); I: left hypomeron (arrow: hypomeron with slight mesial projection); J: prosternal process from the direction perpendicular to base of ventral lobe (black arrow of figure 10H); K: scutellar shield; L: mesothorax from the direction perpendicular to mesosternal process between mesocoxae (white arrow of figure 10H); M: left mid tarsus." figureDoi="http://doi.org/10.5281/zenodo.10200057" httpUri="https://zenodo.org/record/10200057/files/figure.png" pageId="16" pageNumber="317">Fig. 10D, E</figureCitation>
); frontal carina not complete (
<figureCitation id="133C27BEFFB6CF74FC2AEF00FB9BFBD8" box="[1018,1124,1051,1077]" captionStart="FIGURE 10" captionStartId="15.[151,250,1654,1678]" captionTargetBox="[151,1436,186,1630]" captionTargetId="figure-16@15.[151,1436,181,1630]" captionTargetPageId="15" captionText="FIGURE 10. Penia inopinata, holotype, female (6349) (= paratype of Penia takasago). A: labels; B: habitus, dorsal side; C: habitus, ventral side; D: head, anterior view; E: head, dorsal view; F: basal antennomeres of left antenna; G: left maxillary palp, dorsal side; H: head, prothorax and mesothorax, lateral view (black arrow: the direction perpendicular to the base of ventral lobe of prosternal process, white arrow: the direction perpendicular to the mesosternal process between mesocoxae); I: left hypomeron (arrow: hypomeron with slight mesial projection); J: prosternal process from the direction perpendicular to base of ventral lobe (black arrow of figure 10H); K: scutellar shield; L: mesothorax from the direction perpendicular to mesosternal process between mesocoxae (white arrow of figure 10H); M: left mid tarsus." figureDoi="http://doi.org/10.5281/zenodo.10200057" httpUri="https://zenodo.org/record/10200057/files/figure.png" pageId="16" pageNumber="317">Fig. 10D</figureCitation>
); frontal margin hexagonal, almost straight apically in dorsal view (
<figureCitation id="133C27BEFFB6CF74FDB0EF24FD37FBB4" box="[608,712,1087,1113]" captionStart="FIGURE 10" captionStartId="15.[151,250,1654,1678]" captionTargetBox="[151,1436,186,1630]" captionTargetId="figure-16@15.[151,1436,181,1630]" captionTargetPageId="15" captionText="FIGURE 10. Penia inopinata, holotype, female (6349) (= paratype of Penia takasago). A: labels; B: habitus, dorsal side; C: habitus, ventral side; D: head, anterior view; E: head, dorsal view; F: basal antennomeres of left antenna; G: left maxillary palp, dorsal side; H: head, prothorax and mesothorax, lateral view (black arrow: the direction perpendicular to the base of ventral lobe of prosternal process, white arrow: the direction perpendicular to the mesosternal process between mesocoxae); I: left hypomeron (arrow: hypomeron with slight mesial projection); J: prosternal process from the direction perpendicular to base of ventral lobe (black arrow of figure 10H); K: scutellar shield; L: mesothorax from the direction perpendicular to mesosternal process between mesocoxae (white arrow of figure 10H); M: left mid tarsus." figureDoi="http://doi.org/10.5281/zenodo.10200057" httpUri="https://zenodo.org/record/10200057/files/figure.png" pageId="16" pageNumber="317">Fig. 10E</figureCitation>
); frontoclypeal region protruding beyond base of labrum. Eyes relatively normal in convexity, 0.25 x longer than interocular distance in dorsal view (
<figureCitation id="133C27BEFFB6CF74FB98EF78FB52FB90" box="[1096,1197,1123,1149]" captionStart="FIGURE 10" captionStartId="15.[151,250,1654,1678]" captionTargetBox="[151,1436,186,1630]" captionTargetId="figure-16@15.[151,1436,181,1630]" captionTargetPageId="15" captionText="FIGURE 10. Penia inopinata, holotype, female (6349) (= paratype of Penia takasago). A: labels; B: habitus, dorsal side; C: habitus, ventral side; D: head, anterior view; E: head, dorsal view; F: basal antennomeres of left antenna; G: left maxillary palp, dorsal side; H: head, prothorax and mesothorax, lateral view (black arrow: the direction perpendicular to the base of ventral lobe of prosternal process, white arrow: the direction perpendicular to the mesosternal process between mesocoxae); I: left hypomeron (arrow: hypomeron with slight mesial projection); J: prosternal process from the direction perpendicular to base of ventral lobe (black arrow of figure 10H); K: scutellar shield; L: mesothorax from the direction perpendicular to mesosternal process between mesocoxae (white arrow of figure 10H); M: left mid tarsus." figureDoi="http://doi.org/10.5281/zenodo.10200057" httpUri="https://zenodo.org/record/10200057/files/figure.png" pageId="16" pageNumber="317">Fig. 10E</figureCitation>
). Antennae extending beyond pronotum posterior lateral apices by antennomere VII; antennomeres longer than wide;
<collectionCode id="ED16A3FEFFB6CF74FB7EEF9CFB3EFB4C" box="[1198,1217,1159,1185]" pageId="16" pageNumber="317">II</collectionCode>
obconical, shortest, 1.5 x longer than wide;
<collectionCode id="ED16A3FEFFB6CF74FE7CEFB0FE37FB28" box="[428,456,1195,1221]" pageId="16" pageNumber="317">III</collectionCode>
weakly serrated, 2.7 x longer than wide, 2.1 x longer than
<collectionCode id="ED16A3FEFFB6CF74FBA4EFB0FB75FB28" box="[1140,1162,1195,1221]" pageId="16" pageNumber="317">II</collectionCode>
;
<collectionCode id="ED16A3FEFFB6CF74FB47EFB0FB4AFB28" box="[1175,1205,1195,1221]" pageId="16" pageNumber="317">IV</collectionCode>
XI filiform;
<specimenCode id="DBA19340FFB6CF74FA83EFB0FA63FB28" box="[1363,1436,1195,1221]" collectionCode="IV" pageId="16" pageNumber="317">IV 3.2</specimenCode>
x longer than wide, 1.2 x longer than
<collectionCode id="ED16A3FEFFB6CF74FD95EFD4FD9BFB04" box="[581,612,1231,1257]" pageId="16" pageNumber="317">III</collectionCode>
, 0.8 x longer than
<collectionCode id="ED16A3FEFFB6CF74FCEBEFD4FCB2FB04" box="[827,845,1231,1257]" pageId="16" pageNumber="317">II</collectionCode>
<collectionCode id="ED16A3FEFFB6CF74FC8BEFD4FC88FB04" box="[859,887,1231,1257]" pageId="16" pageNumber="317">III</collectionCode>
combined (
<figureCitation id="133C27BEFFB6CF74FC2FEFD4FB9BFB04" box="[1023,1124,1231,1257]" captionStart="FIGURE 10" captionStartId="15.[151,250,1654,1678]" captionTargetBox="[151,1436,186,1630]" captionTargetId="figure-16@15.[151,1436,181,1630]" captionTargetPageId="15" captionText="FIGURE 10. Penia inopinata, holotype, female (6349) (= paratype of Penia takasago). A: labels; B: habitus, dorsal side; C: habitus, ventral side; D: head, anterior view; E: head, dorsal view; F: basal antennomeres of left antenna; G: left maxillary palp, dorsal side; H: head, prothorax and mesothorax, lateral view (black arrow: the direction perpendicular to the base of ventral lobe of prosternal process, white arrow: the direction perpendicular to the mesosternal process between mesocoxae); I: left hypomeron (arrow: hypomeron with slight mesial projection); J: prosternal process from the direction perpendicular to base of ventral lobe (black arrow of figure 10H); K: scutellar shield; L: mesothorax from the direction perpendicular to mesosternal process between mesocoxae (white arrow of figure 10H); M: left mid tarsus." figureDoi="http://doi.org/10.5281/zenodo.10200057" httpUri="https://zenodo.org/record/10200057/files/figure.png" pageId="16" pageNumber="317">Fig. 10F</figureCitation>
);
<specimenCode id="DBA19340FFB6CF74FBA9EFD4FB47FB04" box="[1145,1208,1231,1257]" collectionCode="V" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="16" pageNumber="317" type="Museum">V 3.4</specimenCode>
x longer than wide, almost as long as
<collectionCode id="ED16A3FEFFB6CF74FE8DEFE8FE83FAE0" box="[349,380,1267,1293]" pageId="16" pageNumber="317">IV</collectionCode>
; right antennomeres
<collectionCode id="ED16A3FEFFB6CF74FDB5EFE8FD87FAE0" box="[613,632,1267,1293]" pageId="16" pageNumber="317">II</collectionCode>
XI and left antennomeres IXXI lost in
<typeStatus id="54BC8599FFB6CF74FB96EFE8FB57FAE0" box="[1094,1192,1267,1293]" pageId="16" pageNumber="317" type="holotype">holotype</typeStatus>
(
<figureCitation id="133C27BEFFB6CF74FB67EFE8FAE2FAE0" box="[1207,1309,1267,1293]" captionStart="FIGURE 10" captionStartId="15.[151,250,1654,1678]" captionTargetBox="[151,1436,186,1630]" captionTargetId="figure-16@15.[151,1436,181,1630]" captionTargetPageId="15" captionText="FIGURE 10. Penia inopinata, holotype, female (6349) (= paratype of Penia takasago). A: labels; B: habitus, dorsal side; C: habitus, ventral side; D: head, anterior view; E: head, dorsal view; F: basal antennomeres of left antenna; G: left maxillary palp, dorsal side; H: head, prothorax and mesothorax, lateral view (black arrow: the direction perpendicular to the base of ventral lobe of prosternal process, white arrow: the direction perpendicular to the mesosternal process between mesocoxae); I: left hypomeron (arrow: hypomeron with slight mesial projection); J: prosternal process from the direction perpendicular to base of ventral lobe (black arrow of figure 10H); K: scutellar shield; L: mesothorax from the direction perpendicular to mesosternal process between mesocoxae (white arrow of figure 10H); M: left mid tarsus." figureDoi="http://doi.org/10.5281/zenodo.10200057" httpUri="https://zenodo.org/record/10200057/files/figure.png" pageId="16" pageNumber="317">Fig. 10B</figureCitation>
). Mandible bidentate (
<figureCitation id="133C27BEFFB6CF74FEDCEE0CFE8AFADC" box="[268,373,1303,1329]" captionStart="FIGURE 10" captionStartId="15.[151,250,1654,1678]" captionTargetBox="[151,1436,186,1630]" captionTargetId="figure-16@15.[151,1436,181,1630]" captionTargetPageId="15" captionText="FIGURE 10. Penia inopinata, holotype, female (6349) (= paratype of Penia takasago). A: labels; B: habitus, dorsal side; C: habitus, ventral side; D: head, anterior view; E: head, dorsal view; F: basal antennomeres of left antenna; G: left maxillary palp, dorsal side; H: head, prothorax and mesothorax, lateral view (black arrow: the direction perpendicular to the base of ventral lobe of prosternal process, white arrow: the direction perpendicular to the mesosternal process between mesocoxae); I: left hypomeron (arrow: hypomeron with slight mesial projection); J: prosternal process from the direction perpendicular to base of ventral lobe (black arrow of figure 10H); K: scutellar shield; L: mesothorax from the direction perpendicular to mesosternal process between mesocoxae (white arrow of figure 10H); M: left mid tarsus." figureDoi="http://doi.org/10.5281/zenodo.10200057" httpUri="https://zenodo.org/record/10200057/files/figure.png" pageId="16" pageNumber="317">Fig. 10D</figureCitation>
). Apical maxillary palpomere semicircular (
<figureCitation id="133C27BEFFB6CF74FC8EEE0CFC3AFADC" box="[862,965,1303,1329]" captionStart="FIGURE 10" captionStartId="15.[151,250,1654,1678]" captionTargetBox="[151,1436,186,1630]" captionTargetId="figure-16@15.[151,1436,181,1630]" captionTargetPageId="15" captionText="FIGURE 10. Penia inopinata, holotype, female (6349) (= paratype of Penia takasago). A: labels; B: habitus, dorsal side; C: habitus, ventral side; D: head, anterior view; E: head, dorsal view; F: basal antennomeres of left antenna; G: left maxillary palp, dorsal side; H: head, prothorax and mesothorax, lateral view (black arrow: the direction perpendicular to the base of ventral lobe of prosternal process, white arrow: the direction perpendicular to the mesosternal process between mesocoxae); I: left hypomeron (arrow: hypomeron with slight mesial projection); J: prosternal process from the direction perpendicular to base of ventral lobe (black arrow of figure 10H); K: scutellar shield; L: mesothorax from the direction perpendicular to mesosternal process between mesocoxae (white arrow of figure 10H); M: left mid tarsus." figureDoi="http://doi.org/10.5281/zenodo.10200057" httpUri="https://zenodo.org/record/10200057/files/figure.png" pageId="16" pageNumber="317">Fig. 10G</figureCitation>
), 2.1 x longer than wide, almost as long as maximum length of eye; anterior edge rounded
</materialsCitation>
.
</paragraph>
<paragraph id="8BB83B3BFFB6CF75FF17EE44FDB8FE3C" blockId="16.[151,1437,151,2013]" lastBlockId="17.[151,1437,151,717]" lastPageId="17" lastPageNumber="318" pageId="16" pageNumber="317">
Prothorax. Pronotum hexagonal, 0.8 x longer than wide, roundly widening anteriorly, widest around midlength, and then straightly and slightly narrowed ahead of hind angles (
<figureCitation id="133C27BEFFB6CF74FC8FEE98FC38FA70" box="[863,967,1411,1437]" captionStart="FIGURE 10" captionStartId="15.[151,250,1654,1678]" captionTargetBox="[151,1436,186,1630]" captionTargetId="figure-16@15.[151,1436,181,1630]" captionTargetPageId="15" captionText="FIGURE 10. Penia inopinata, holotype, female (6349) (= paratype of Penia takasago). A: labels; B: habitus, dorsal side; C: habitus, ventral side; D: head, anterior view; E: head, dorsal view; F: basal antennomeres of left antenna; G: left maxillary palp, dorsal side; H: head, prothorax and mesothorax, lateral view (black arrow: the direction perpendicular to the base of ventral lobe of prosternal process, white arrow: the direction perpendicular to the mesosternal process between mesocoxae); I: left hypomeron (arrow: hypomeron with slight mesial projection); J: prosternal process from the direction perpendicular to base of ventral lobe (black arrow of figure 10H); K: scutellar shield; L: mesothorax from the direction perpendicular to mesosternal process between mesocoxae (white arrow of figure 10H); M: left mid tarsus." figureDoi="http://doi.org/10.5281/zenodo.10200057" httpUri="https://zenodo.org/record/10200057/files/figure.png" pageId="16" pageNumber="317">Fig. 10A</figureCitation>
), tallest just behind midlength (
<figureCitation id="133C27BEFFB6CF74FAF7EE98FA70FA70" box="[1319,1423,1411,1437]" captionStart="FIGURE 10" captionStartId="15.[151,250,1654,1678]" captionTargetBox="[151,1436,186,1630]" captionTargetId="figure-16@15.[151,1436,181,1630]" captionTargetPageId="15" captionText="FIGURE 10. Penia inopinata, holotype, female (6349) (= paratype of Penia takasago). A: labels; B: habitus, dorsal side; C: habitus, ventral side; D: head, anterior view; E: head, dorsal view; F: basal antennomeres of left antenna; G: left maxillary palp, dorsal side; H: head, prothorax and mesothorax, lateral view (black arrow: the direction perpendicular to the base of ventral lobe of prosternal process, white arrow: the direction perpendicular to the mesosternal process between mesocoxae); I: left hypomeron (arrow: hypomeron with slight mesial projection); J: prosternal process from the direction perpendicular to base of ventral lobe (black arrow of figure 10H); K: scutellar shield; L: mesothorax from the direction perpendicular to mesosternal process between mesocoxae (white arrow of figure 10H); M: left mid tarsus." figureDoi="http://doi.org/10.5281/zenodo.10200057" httpUri="https://zenodo.org/record/10200057/files/figure.png" pageId="16" pageNumber="317">Fig. 10H</figureCitation>
), without median longitudinal depression posteriorly; anterior edge strongly concave; anterior angles simple, acute; punctate lateral ridge extending from anterior angles to hind angles (
<figureCitation id="133C27BEFFB6CF74FC6AEED0FBDBFA08" box="[954,1060,1483,1509]" captionStart="FIGURE 10" captionStartId="15.[151,250,1654,1678]" captionTargetBox="[151,1436,186,1630]" captionTargetId="figure-16@15.[151,1436,181,1630]" captionTargetPageId="15" captionText="FIGURE 10. Penia inopinata, holotype, female (6349) (= paratype of Penia takasago). A: labels; B: habitus, dorsal side; C: habitus, ventral side; D: head, anterior view; E: head, dorsal view; F: basal antennomeres of left antenna; G: left maxillary palp, dorsal side; H: head, prothorax and mesothorax, lateral view (black arrow: the direction perpendicular to the base of ventral lobe of prosternal process, white arrow: the direction perpendicular to the mesosternal process between mesocoxae); I: left hypomeron (arrow: hypomeron with slight mesial projection); J: prosternal process from the direction perpendicular to base of ventral lobe (black arrow of figure 10H); K: scutellar shield; L: mesothorax from the direction perpendicular to mesosternal process between mesocoxae (white arrow of figure 10H); M: left mid tarsus." figureDoi="http://doi.org/10.5281/zenodo.10200057" httpUri="https://zenodo.org/record/10200057/files/figure.png" pageId="16" pageNumber="317">Fig. 10B</figureCitation>
); hind angles simple, broad, not protruding laterally beyond pronotal side; posterior edge with a sublateral incision near each hind angle, without carinae next to sublateral incisions (
<figureCitation id="133C27BEFFB6CF74FDFBED08FD6DF9C0" box="[555,658,1555,1581]" captionStart="FIGURE 10" captionStartId="15.[151,250,1654,1678]" captionTargetBox="[151,1436,186,1630]" captionTargetId="figure-16@15.[151,1436,181,1630]" captionTargetPageId="15" captionText="FIGURE 10. Penia inopinata, holotype, female (6349) (= paratype of Penia takasago). A: labels; B: habitus, dorsal side; C: habitus, ventral side; D: head, anterior view; E: head, dorsal view; F: basal antennomeres of left antenna; G: left maxillary palp, dorsal side; H: head, prothorax and mesothorax, lateral view (black arrow: the direction perpendicular to the base of ventral lobe of prosternal process, white arrow: the direction perpendicular to the mesosternal process between mesocoxae); I: left hypomeron (arrow: hypomeron with slight mesial projection); J: prosternal process from the direction perpendicular to base of ventral lobe (black arrow of figure 10H); K: scutellar shield; L: mesothorax from the direction perpendicular to mesosternal process between mesocoxae (white arrow of figure 10H); M: left mid tarsus." figureDoi="http://doi.org/10.5281/zenodo.10200057" httpUri="https://zenodo.org/record/10200057/files/figure.png" pageId="16" pageNumber="317">Fig. 10B</figureCitation>
). Hypomeron with slight mesial projection (
<figureCitation id="133C27BEFFB6CF74FB53ED08FB1EF9C0" box="[1155,1249,1555,1581]" captionStart="FIGURE 10" captionStartId="15.[151,250,1654,1678]" captionTargetBox="[151,1436,186,1630]" captionTargetId="figure-16@15.[151,1436,181,1630]" captionTargetPageId="15" captionText="FIGURE 10. Penia inopinata, holotype, female (6349) (= paratype of Penia takasago). A: labels; B: habitus, dorsal side; C: habitus, ventral side; D: head, anterior view; E: head, dorsal view; F: basal antennomeres of left antenna; G: left maxillary palp, dorsal side; H: head, prothorax and mesothorax, lateral view (black arrow: the direction perpendicular to the base of ventral lobe of prosternal process, white arrow: the direction perpendicular to the mesosternal process between mesocoxae); I: left hypomeron (arrow: hypomeron with slight mesial projection); J: prosternal process from the direction perpendicular to base of ventral lobe (black arrow of figure 10H); K: scutellar shield; L: mesothorax from the direction perpendicular to mesosternal process between mesocoxae (white arrow of figure 10H); M: left mid tarsus." figureDoi="http://doi.org/10.5281/zenodo.10200057" httpUri="https://zenodo.org/record/10200057/files/figure.png" pageId="16" pageNumber="317">Fig. 10I</figureCitation>
: arrow); anterior angle rounded; mesial edge slightly rounded; mesial and posterior margins with impunctate ridge (
<figureCitation id="133C27BEFFB6CF74FB1BED2CFAD9F9BC" box="[1227,1318,1591,1617]" captionStart="FIGURE 10" captionStartId="15.[151,250,1654,1678]" captionTargetBox="[151,1436,186,1630]" captionTargetId="figure-16@15.[151,1436,181,1630]" captionTargetPageId="15" captionText="FIGURE 10. Penia inopinata, holotype, female (6349) (= paratype of Penia takasago). A: labels; B: habitus, dorsal side; C: habitus, ventral side; D: head, anterior view; E: head, dorsal view; F: basal antennomeres of left antenna; G: left maxillary palp, dorsal side; H: head, prothorax and mesothorax, lateral view (black arrow: the direction perpendicular to the base of ventral lobe of prosternal process, white arrow: the direction perpendicular to the mesosternal process between mesocoxae); I: left hypomeron (arrow: hypomeron with slight mesial projection); J: prosternal process from the direction perpendicular to base of ventral lobe (black arrow of figure 10H); K: scutellar shield; L: mesothorax from the direction perpendicular to mesosternal process between mesocoxae (white arrow of figure 10H); M: left mid tarsus." figureDoi="http://doi.org/10.5281/zenodo.10200057" httpUri="https://zenodo.org/record/10200057/files/figure.png" pageId="16" pageNumber="317">Fig. 10I</figureCitation>
); posterior margin with rectangular projection between two large emarginations; hind angle broadly triangular. Prosternum weakly incurved ventrally in lateral view; anterior lobe weakly protruding beyond prosternal ventral line in lateral view (
<figureCitation id="133C27BEFFB6CF74FF0EEDB8FEB8F950" box="[222,327,1699,1725]" captionStart="FIGURE 10" captionStartId="15.[151,250,1654,1678]" captionTargetBox="[151,1436,186,1630]" captionTargetId="figure-16@15.[151,1436,181,1630]" captionTargetPageId="15" captionText="FIGURE 10. Penia inopinata, holotype, female (6349) (= paratype of Penia takasago). A: labels; B: habitus, dorsal side; C: habitus, ventral side; D: head, anterior view; E: head, dorsal view; F: basal antennomeres of left antenna; G: left maxillary palp, dorsal side; H: head, prothorax and mesothorax, lateral view (black arrow: the direction perpendicular to the base of ventral lobe of prosternal process, white arrow: the direction perpendicular to the mesosternal process between mesocoxae); I: left hypomeron (arrow: hypomeron with slight mesial projection); J: prosternal process from the direction perpendicular to base of ventral lobe (black arrow of figure 10H); K: scutellar shield; L: mesothorax from the direction perpendicular to mesosternal process between mesocoxae (white arrow of figure 10H); M: left mid tarsus." figureDoi="http://doi.org/10.5281/zenodo.10200057" httpUri="https://zenodo.org/record/10200057/files/figure.png" pageId="16" pageNumber="317">Fig. 10H</figureCitation>
); anterior edge broadly rounded in ventral view (
<figureCitation id="133C27BEFFB6CF74FCA3EDB8FC24F950" box="[883,987,1699,1725]" captionStart="FIGURE 10" captionStartId="15.[151,250,1654,1678]" captionTargetBox="[151,1436,186,1630]" captionTargetId="figure-16@15.[151,1436,181,1630]" captionTargetPageId="15" captionText="FIGURE 10. Penia inopinata, holotype, female (6349) (= paratype of Penia takasago). A: labels; B: habitus, dorsal side; C: habitus, ventral side; D: head, anterior view; E: head, dorsal view; F: basal antennomeres of left antenna; G: left maxillary palp, dorsal side; H: head, prothorax and mesothorax, lateral view (black arrow: the direction perpendicular to the base of ventral lobe of prosternal process, white arrow: the direction perpendicular to the mesosternal process between mesocoxae); I: left hypomeron (arrow: hypomeron with slight mesial projection); J: prosternal process from the direction perpendicular to base of ventral lobe (black arrow of figure 10H); K: scutellar shield; L: mesothorax from the direction perpendicular to mesosternal process between mesocoxae (white arrow of figure 10H); M: left mid tarsus." figureDoi="http://doi.org/10.5281/zenodo.10200057" httpUri="https://zenodo.org/record/10200057/files/figure.png" pageId="16" pageNumber="317">Fig. 10C</figureCitation>
). Prosternal process broad, 2.1 x longer than procoxal cavity length, concave between procoxae, strongly curved dorsad from the middle of procoxal cavities in lateral view (
<figureCitation id="133C27BEFFB6CF74FE9DEDF0FE48F8E8" box="[333,439,1771,1797]" captionStart="FIGURE 10" captionStartId="15.[151,250,1654,1678]" captionTargetBox="[151,1436,186,1630]" captionTargetId="figure-16@15.[151,1436,181,1630]" captionTargetPageId="15" captionText="FIGURE 10. Penia inopinata, holotype, female (6349) (= paratype of Penia takasago). A: labels; B: habitus, dorsal side; C: habitus, ventral side; D: head, anterior view; E: head, dorsal view; F: basal antennomeres of left antenna; G: left maxillary palp, dorsal side; H: head, prothorax and mesothorax, lateral view (black arrow: the direction perpendicular to the base of ventral lobe of prosternal process, white arrow: the direction perpendicular to the mesosternal process between mesocoxae); I: left hypomeron (arrow: hypomeron with slight mesial projection); J: prosternal process from the direction perpendicular to base of ventral lobe (black arrow of figure 10H); K: scutellar shield; L: mesothorax from the direction perpendicular to mesosternal process between mesocoxae (white arrow of figure 10H); M: left mid tarsus." figureDoi="http://doi.org/10.5281/zenodo.10200057" httpUri="https://zenodo.org/record/10200057/files/figure.png" pageId="16" pageNumber="317">Fig. 10H</figureCitation>
), without subapical tooth; dorsal lobe abruptly narrowed posterad near apical region in ventral view (
<figureCitation id="133C27BEFFB6CF74FEE6EC14FE69F8C4" box="[310,406,1807,1833]" captionStart="FIGURE 10" captionStartId="15.[151,250,1654,1678]" captionTargetBox="[151,1436,186,1630]" captionTargetId="figure-16@15.[151,1436,181,1630]" captionTargetPageId="15" captionText="FIGURE 10. Penia inopinata, holotype, female (6349) (= paratype of Penia takasago). A: labels; B: habitus, dorsal side; C: habitus, ventral side; D: head, anterior view; E: head, dorsal view; F: basal antennomeres of left antenna; G: left maxillary palp, dorsal side; H: head, prothorax and mesothorax, lateral view (black arrow: the direction perpendicular to the base of ventral lobe of prosternal process, white arrow: the direction perpendicular to the mesosternal process between mesocoxae); I: left hypomeron (arrow: hypomeron with slight mesial projection); J: prosternal process from the direction perpendicular to base of ventral lobe (black arrow of figure 10H); K: scutellar shield; L: mesothorax from the direction perpendicular to mesosternal process between mesocoxae (white arrow of figure 10H); M: left mid tarsus." figureDoi="http://doi.org/10.5281/zenodo.10200057" httpUri="https://zenodo.org/record/10200057/files/figure.png" pageId="16" pageNumber="317">Fig. 10J</figureCitation>
); ventral lobe roundly expanded near base and then abruptly narrowed posterad in ventral view (
<figureCitation id="133C27BEFFB6CF74FF0EEC28FEC1F8A0" box="[222,318,1843,1869]" captionStart="FIGURE 10" captionStartId="15.[151,250,1654,1678]" captionTargetBox="[151,1436,186,1630]" captionTargetId="figure-16@15.[151,1436,181,1630]" captionTargetPageId="15" captionText="FIGURE 10. Penia inopinata, holotype, female (6349) (= paratype of Penia takasago). A: labels; B: habitus, dorsal side; C: habitus, ventral side; D: head, anterior view; E: head, dorsal view; F: basal antennomeres of left antenna; G: left maxillary palp, dorsal side; H: head, prothorax and mesothorax, lateral view (black arrow: the direction perpendicular to the base of ventral lobe of prosternal process, white arrow: the direction perpendicular to the mesosternal process between mesocoxae); I: left hypomeron (arrow: hypomeron with slight mesial projection); J: prosternal process from the direction perpendicular to base of ventral lobe (black arrow of figure 10H); K: scutellar shield; L: mesothorax from the direction perpendicular to mesosternal process between mesocoxae (white arrow of figure 10H); M: left mid tarsus." figureDoi="http://doi.org/10.5281/zenodo.10200057" httpUri="https://zenodo.org/record/10200057/files/figure.png" pageId="16" pageNumber="317">Fig. 10J</figureCitation>
); ventral margin roundly and medially expanded in lateral view (
<figureCitation id="133C27BEFFB6CF74FBF2EC28FB7EF8A0" box="[1058,1153,1843,1869]" captionStart="FIGURE 10" captionStartId="15.[151,250,1654,1678]" captionTargetBox="[151,1436,186,1630]" captionTargetId="figure-16@15.[151,1436,181,1630]" captionTargetPageId="15" captionText="FIGURE 10. Penia inopinata, holotype, female (6349) (= paratype of Penia takasago). A: labels; B: habitus, dorsal side; C: habitus, ventral side; D: head, anterior view; E: head, dorsal view; F: basal antennomeres of left antenna; G: left maxillary palp, dorsal side; H: head, prothorax and mesothorax, lateral view (black arrow: the direction perpendicular to the base of ventral lobe of prosternal process, white arrow: the direction perpendicular to the mesosternal process between mesocoxae); I: left hypomeron (arrow: hypomeron with slight mesial projection); J: prosternal process from the direction perpendicular to base of ventral lobe (black arrow of figure 10H); K: scutellar shield; L: mesothorax from the direction perpendicular to mesosternal process between mesocoxae (white arrow of figure 10H); M: left mid tarsus." figureDoi="http://doi.org/10.5281/zenodo.10200057" httpUri="https://zenodo.org/record/10200057/files/figure.png" pageId="16" pageNumber="317">Fig. 10J</figureCitation>
); apex rounded in lateral and ventral views (
<figureCitation id="133C27BEFFB6CF74FEA8EC4CFDFEF89C" box="[376,513,1879,1905]" captionStart="FIGURE 10" captionStartId="15.[151,250,1654,1678]" captionTargetBox="[151,1436,186,1630]" captionTargetId="figure-16@15.[151,1436,181,1630]" captionTargetPageId="15" captionText="FIGURE 10. Penia inopinata, holotype, female (6349) (= paratype of Penia takasago). A: labels; B: habitus, dorsal side; C: habitus, ventral side; D: head, anterior view; E: head, dorsal view; F: basal antennomeres of left antenna; G: left maxillary palp, dorsal side; H: head, prothorax and mesothorax, lateral view (black arrow: the direction perpendicular to the base of ventral lobe of prosternal process, white arrow: the direction perpendicular to the mesosternal process between mesocoxae); I: left hypomeron (arrow: hypomeron with slight mesial projection); J: prosternal process from the direction perpendicular to base of ventral lobe (black arrow of figure 10H); K: scutellar shield; L: mesothorax from the direction perpendicular to mesosternal process between mesocoxae (white arrow of figure 10H); M: left mid tarsus." figureDoi="http://doi.org/10.5281/zenodo.10200057" httpUri="https://zenodo.org/record/10200057/files/figure.png" pageId="16" pageNumber="317">Fig. 10J, H</figureCitation>
). Pronotosternal sutures not grooved (
<figureCitation id="133C27BEFFB6CF74FC6EEC4CFBE1F89C" box="[958,1054,1879,1905]" captionStart="FIGURE 10" captionStartId="15.[151,250,1654,1678]" captionTargetBox="[151,1436,186,1630]" captionTargetId="figure-16@15.[151,1436,181,1630]" captionTargetPageId="15" captionText="FIGURE 10. Penia inopinata, holotype, female (6349) (= paratype of Penia takasago). A: labels; B: habitus, dorsal side; C: habitus, ventral side; D: head, anterior view; E: head, dorsal view; F: basal antennomeres of left antenna; G: left maxillary palp, dorsal side; H: head, prothorax and mesothorax, lateral view (black arrow: the direction perpendicular to the base of ventral lobe of prosternal process, white arrow: the direction perpendicular to the mesosternal process between mesocoxae); I: left hypomeron (arrow: hypomeron with slight mesial projection); J: prosternal process from the direction perpendicular to base of ventral lobe (black arrow of figure 10H); K: scutellar shield; L: mesothorax from the direction perpendicular to mesosternal process between mesocoxae (white arrow of figure 10H); M: left mid tarsus." figureDoi="http://doi.org/10.5281/zenodo.10200057" httpUri="https://zenodo.org/record/10200057/files/figure.png" pageId="16" pageNumber="317">Fig. 10I</figureCitation>
), weakly sinuate in ventral view (
<figureCitation id="133C27BEFFB6CF74FF4FEC60FEF8F878" box="[159,263,1915,1941]" captionStart="FIGURE 10" captionStartId="15.[151,250,1654,1678]" captionTargetBox="[151,1436,186,1630]" captionTargetId="figure-16@15.[151,1436,181,1630]" captionTargetPageId="15" captionText="FIGURE 10. Penia inopinata, holotype, female (6349) (= paratype of Penia takasago). A: labels; B: habitus, dorsal side; C: habitus, ventral side; D: head, anterior view; E: head, dorsal view; F: basal antennomeres of left antenna; G: left maxillary palp, dorsal side; H: head, prothorax and mesothorax, lateral view (black arrow: the direction perpendicular to the base of ventral lobe of prosternal process, white arrow: the direction perpendicular to the mesosternal process between mesocoxae); I: left hypomeron (arrow: hypomeron with slight mesial projection); J: prosternal process from the direction perpendicular to base of ventral lobe (black arrow of figure 10H); K: scutellar shield; L: mesothorax from the direction perpendicular to mesosternal process between mesocoxae (white arrow of figure 10H); M: left mid tarsus." figureDoi="http://doi.org/10.5281/zenodo.10200057" httpUri="https://zenodo.org/record/10200057/files/figure.png" pageId="16" pageNumber="317">Fig. 10C</figureCitation>
), slightly opened anteriorly. Scutellar shield tongue-shaped (
<figureCitation id="133C27BEFFB6CF74FC6DEC60FBD8F878" box="[957,1063,1915,1941]" captionStart="FIGURE 10" captionStartId="15.[151,250,1654,1678]" captionTargetBox="[151,1436,186,1630]" captionTargetId="figure-16@15.[151,1436,181,1630]" captionTargetPageId="15" captionText="FIGURE 10. Penia inopinata, holotype, female (6349) (= paratype of Penia takasago). A: labels; B: habitus, dorsal side; C: habitus, ventral side; D: head, anterior view; E: head, dorsal view; F: basal antennomeres of left antenna; G: left maxillary palp, dorsal side; H: head, prothorax and mesothorax, lateral view (black arrow: the direction perpendicular to the base of ventral lobe of prosternal process, white arrow: the direction perpendicular to the mesosternal process between mesocoxae); I: left hypomeron (arrow: hypomeron with slight mesial projection); J: prosternal process from the direction perpendicular to base of ventral lobe (black arrow of figure 10H); K: scutellar shield; L: mesothorax from the direction perpendicular to mesosternal process between mesocoxae (white arrow of figure 10H); M: left mid tarsus." figureDoi="http://doi.org/10.5281/zenodo.10200057" httpUri="https://zenodo.org/record/10200057/files/figure.png" pageId="16" pageNumber="317">Fig. 10K</figureCitation>
), almost as long as wide, widest anteriorly, slightly constricted near base and then narrowed posterad, flat, inclined anterior-downwards, not visible in lateral view (
<figureCitation id="133C27BEFFB6CF74FE96ECD8FE50F830" box="[326,431,1987,2013]" captionStart="FIGURE 10" captionStartId="15.[151,250,1654,1678]" captionTargetBox="[151,1436,186,1630]" captionTargetId="figure-16@15.[151,1436,181,1630]" captionTargetPageId="15" captionText="FIGURE 10. Penia inopinata, holotype, female (6349) (= paratype of Penia takasago). A: labels; B: habitus, dorsal side; C: habitus, ventral side; D: head, anterior view; E: head, dorsal view; F: basal antennomeres of left antenna; G: left maxillary palp, dorsal side; H: head, prothorax and mesothorax, lateral view (black arrow: the direction perpendicular to the base of ventral lobe of prosternal process, white arrow: the direction perpendicular to the mesosternal process between mesocoxae); I: left hypomeron (arrow: hypomeron with slight mesial projection); J: prosternal process from the direction perpendicular to base of ventral lobe (black arrow of figure 10H); K: scutellar shield; L: mesothorax from the direction perpendicular to mesosternal process between mesocoxae (white arrow of figure 10H); M: left mid tarsus." figureDoi="http://doi.org/10.5281/zenodo.10200057" httpUri="https://zenodo.org/record/10200057/files/figure.png" pageId="16" pageNumber="317">Fig. 10H</figureCitation>
); anterior edge broadly rounded and slightly protruding medially; posterior edge rounded. Mesosternum: borders of mesosternal cavity slightly rounded and then curved obtusely ventrad in lateral view (
<figureCitation id="133C27BEFFB7CF75FABFEB8CFF34FF38" captionStart="FIGURE 10" captionStartId="15.[151,250,1654,1678]" captionTargetBox="[151,1436,186,1630]" captionTargetId="figure-16@15.[151,1436,181,1630]" captionTargetPageId="15" captionText="FIGURE 10. Penia inopinata, holotype, female (6349) (= paratype of Penia takasago). A: labels; B: habitus, dorsal side; C: habitus, ventral side; D: head, anterior view; E: head, dorsal view; F: basal antennomeres of left antenna; G: left maxillary palp, dorsal side; H: head, prothorax and mesothorax, lateral view (black arrow: the direction perpendicular to the base of ventral lobe of prosternal process, white arrow: the direction perpendicular to the mesosternal process between mesocoxae); I: left hypomeron (arrow: hypomeron with slight mesial projection); J: prosternal process from the direction perpendicular to base of ventral lobe (black arrow of figure 10H); K: scutellar shield; L: mesothorax from the direction perpendicular to mesosternal process between mesocoxae (white arrow of figure 10H); M: left mid tarsus." figureDoi="http://doi.org/10.5281/zenodo.10200057" httpUri="https://zenodo.org/record/10200057/files/figure.png" pageId="17" pageNumber="318">Fig. 10H</figureCitation>
); mesosternal process between mesocoxae higher than mesocoxae, visible in lateral view (
<figureCitation id="133C27BEFFB7CF75FB35EBA0FAAFFF38" box="[1253,1360,187,213]" captionStart="FIGURE 10" captionStartId="15.[151,250,1654,1678]" captionTargetBox="[151,1436,186,1630]" captionTargetId="figure-16@15.[151,1436,181,1630]" captionTargetPageId="15" captionText="FIGURE 10. Penia inopinata, holotype, female (6349) (= paratype of Penia takasago). A: labels; B: habitus, dorsal side; C: habitus, ventral side; D: head, anterior view; E: head, dorsal view; F: basal antennomeres of left antenna; G: left maxillary palp, dorsal side; H: head, prothorax and mesothorax, lateral view (black arrow: the direction perpendicular to the base of ventral lobe of prosternal process, white arrow: the direction perpendicular to the mesosternal process between mesocoxae); I: left hypomeron (arrow: hypomeron with slight mesial projection); J: prosternal process from the direction perpendicular to base of ventral lobe (black arrow of figure 10H); K: scutellar shield; L: mesothorax from the direction perpendicular to mesosternal process between mesocoxae (white arrow of figure 10H); M: left mid tarsus." figureDoi="http://doi.org/10.5281/zenodo.10200057" httpUri="https://zenodo.org/record/10200057/files/figure.png" pageId="17" pageNumber="318">Fig. 10H</figureCitation>
: white arrow); posterior edge 0.15 x wider than total width of mesosternum, straight (
<figureCitation id="133C27BEFFB7CF75FBDEEBC4FB8BFF14" box="[1038,1140,223,249]" captionStart="FIGURE 10" captionStartId="15.[151,250,1654,1678]" captionTargetBox="[151,1436,186,1630]" captionTargetId="figure-16@15.[151,1436,181,1630]" captionTargetPageId="15" captionText="FIGURE 10. Penia inopinata, holotype, female (6349) (= paratype of Penia takasago). A: labels; B: habitus, dorsal side; C: habitus, ventral side; D: head, anterior view; E: head, dorsal view; F: basal antennomeres of left antenna; G: left maxillary palp, dorsal side; H: head, prothorax and mesothorax, lateral view (black arrow: the direction perpendicular to the base of ventral lobe of prosternal process, white arrow: the direction perpendicular to the mesosternal process between mesocoxae); I: left hypomeron (arrow: hypomeron with slight mesial projection); J: prosternal process from the direction perpendicular to base of ventral lobe (black arrow of figure 10H); K: scutellar shield; L: mesothorax from the direction perpendicular to mesosternal process between mesocoxae (white arrow of figure 10H); M: left mid tarsus." figureDoi="http://doi.org/10.5281/zenodo.10200057" httpUri="https://zenodo.org/record/10200057/files/figure.png" pageId="17" pageNumber="318">Fig. 10L</figureCitation>
). Mesepisternum reaching mesocoxal cavity (
<figureCitation id="133C27BEFFB7CF75FEB9EA18FE31FEF0" box="[361,462,259,285]" captionStart="FIGURE 10" captionStartId="15.[151,250,1654,1678]" captionTargetBox="[151,1436,186,1630]" captionTargetId="figure-16@15.[151,1436,181,1630]" captionTargetPageId="15" captionText="FIGURE 10. Penia inopinata, holotype, female (6349) (= paratype of Penia takasago). A: labels; B: habitus, dorsal side; C: habitus, ventral side; D: head, anterior view; E: head, dorsal view; F: basal antennomeres of left antenna; G: left maxillary palp, dorsal side; H: head, prothorax and mesothorax, lateral view (black arrow: the direction perpendicular to the base of ventral lobe of prosternal process, white arrow: the direction perpendicular to the mesosternal process between mesocoxae); I: left hypomeron (arrow: hypomeron with slight mesial projection); J: prosternal process from the direction perpendicular to base of ventral lobe (black arrow of figure 10H); K: scutellar shield; L: mesothorax from the direction perpendicular to mesosternal process between mesocoxae (white arrow of figure 10H); M: left mid tarsus." figureDoi="http://doi.org/10.5281/zenodo.10200057" httpUri="https://zenodo.org/record/10200057/files/figure.png" pageId="17" pageNumber="318">Fig. 10L</figureCitation>
). Metasternum sulcate medially and ahead of metacoxal cavities (
<figureCitation id="133C27BEFFB7CF75FB7CEA18FAEDFEF0" box="[1196,1298,259,285]" captionStart="FIGURE 10" captionStartId="15.[151,250,1654,1678]" captionTargetBox="[151,1436,186,1630]" captionTargetId="figure-16@15.[151,1436,181,1630]" captionTargetPageId="15" captionText="FIGURE 10. Penia inopinata, holotype, female (6349) (= paratype of Penia takasago). A: labels; B: habitus, dorsal side; C: habitus, ventral side; D: head, anterior view; E: head, dorsal view; F: basal antennomeres of left antenna; G: left maxillary palp, dorsal side; H: head, prothorax and mesothorax, lateral view (black arrow: the direction perpendicular to the base of ventral lobe of prosternal process, white arrow: the direction perpendicular to the mesosternal process between mesocoxae); I: left hypomeron (arrow: hypomeron with slight mesial projection); J: prosternal process from the direction perpendicular to base of ventral lobe (black arrow of figure 10H); K: scutellar shield; L: mesothorax from the direction perpendicular to mesosternal process between mesocoxae (white arrow of figure 10H); M: left mid tarsus." figureDoi="http://doi.org/10.5281/zenodo.10200057" httpUri="https://zenodo.org/record/10200057/files/figure.png" pageId="17" pageNumber="318">Fig. 10B</figureCitation>
). Metacoxal plate narrowed toward outer side, becoming like a parallel-sided bar at its outer 1/
<quantity id="4CFF96DEFFB7CF75FB9DEA3CFB82FEAC" box="[1101,1149,295,321]" metricMagnitude="-1" metricUnit="m" metricValue="1.016" pageId="17" pageNumber="318" unit="in" value="4.0">4 in</quantity>
ventral view (
<figureCitation id="133C27BEFFB7CF75FAF6EA3CFA6FFEAC" box="[1318,1424,295,321]" captionStart="FIGURE 10" captionStartId="15.[151,250,1654,1678]" captionTargetBox="[151,1436,186,1630]" captionTargetId="figure-16@15.[151,1436,181,1630]" captionTargetPageId="15" captionText="FIGURE 10. Penia inopinata, holotype, female (6349) (= paratype of Penia takasago). A: labels; B: habitus, dorsal side; C: habitus, ventral side; D: head, anterior view; E: head, dorsal view; F: basal antennomeres of left antenna; G: left maxillary palp, dorsal side; H: head, prothorax and mesothorax, lateral view (black arrow: the direction perpendicular to the base of ventral lobe of prosternal process, white arrow: the direction perpendicular to the mesosternal process between mesocoxae); I: left hypomeron (arrow: hypomeron with slight mesial projection); J: prosternal process from the direction perpendicular to base of ventral lobe (black arrow of figure 10H); K: scutellar shield; L: mesothorax from the direction perpendicular to mesosternal process between mesocoxae (white arrow of figure 10H); M: left mid tarsus." figureDoi="http://doi.org/10.5281/zenodo.10200057" httpUri="https://zenodo.org/record/10200057/files/figure.png" pageId="17" pageNumber="318">Fig. 10C</figureCitation>
). Elytron broadly moderately convex, but with outer margin widely depressed, widest around midlength, 3.4 x longer than wide, 2.7 x longer than pronotum length; apex rounded; elytral striae defined by lines of elongated punctures. Hind wings fully developed. Tibiae with paired spurs; relative tarsomere lengths: IV&lt;III II&lt;V&lt;I; tarsomeres III and IV with lobe ventrally (
<figureCitation id="133C27BEFFB7CF75FE1EEAACFDC4FE3C" box="[462,571,439,465]" captionStart="FIGURE 10" captionStartId="15.[151,250,1654,1678]" captionTargetBox="[151,1436,186,1630]" captionTargetId="figure-16@15.[151,1436,181,1630]" captionTargetPageId="15" captionText="FIGURE 10. Penia inopinata, holotype, female (6349) (= paratype of Penia takasago). A: labels; B: habitus, dorsal side; C: habitus, ventral side; D: head, anterior view; E: head, dorsal view; F: basal antennomeres of left antenna; G: left maxillary palp, dorsal side; H: head, prothorax and mesothorax, lateral view (black arrow: the direction perpendicular to the base of ventral lobe of prosternal process, white arrow: the direction perpendicular to the mesosternal process between mesocoxae); I: left hypomeron (arrow: hypomeron with slight mesial projection); J: prosternal process from the direction perpendicular to base of ventral lobe (black arrow of figure 10H); K: scutellar shield; L: mesothorax from the direction perpendicular to mesosternal process between mesocoxae (white arrow of figure 10H); M: left mid tarsus." figureDoi="http://doi.org/10.5281/zenodo.10200057" httpUri="https://zenodo.org/record/10200057/files/figure.png" pageId="17" pageNumber="318">Fig. 10M</figureCitation>
).
</paragraph>
<paragraph id="8BB83B3BFFB7CF75FF17EAC0FC39FD20" blockId="17.[151,1437,151,717]" pageId="17" pageNumber="318">
Abdomen. Ventrite V wide semicircular, rounded apically (
<figureCitation id="133C27BEFFB7CF75FC81EAC0FC4AFE18" box="[849,949,475,501]" captionStart="FIGURE 10" captionStartId="15.[151,250,1654,1678]" captionTargetBox="[151,1436,186,1630]" captionTargetId="figure-16@15.[151,1436,181,1630]" captionTargetPageId="15" captionText="FIGURE 10. Penia inopinata, holotype, female (6349) (= paratype of Penia takasago). A: labels; B: habitus, dorsal side; C: habitus, ventral side; D: head, anterior view; E: head, dorsal view; F: basal antennomeres of left antenna; G: left maxillary palp, dorsal side; H: head, prothorax and mesothorax, lateral view (black arrow: the direction perpendicular to the base of ventral lobe of prosternal process, white arrow: the direction perpendicular to the mesosternal process between mesocoxae); I: left hypomeron (arrow: hypomeron with slight mesial projection); J: prosternal process from the direction perpendicular to base of ventral lobe (black arrow of figure 10H); K: scutellar shield; L: mesothorax from the direction perpendicular to mesosternal process between mesocoxae (white arrow of figure 10H); M: left mid tarsus." figureDoi="http://doi.org/10.5281/zenodo.10200057" httpUri="https://zenodo.org/record/10200057/files/figure.png" pageId="17" pageNumber="318">Fig. 10C</figureCitation>
), 0.4 x longer than wide. Tergite and sternite VIII yellow. Terigite VIII semicircular, almost as long as wide; sternite VIII (between base of spiculum ventrale and apex) semicircular, 0.8 x longer than wide (
<figureCitation id="133C27BEFFB7CF75FD69E938FCDEFDD0" box="[697,801,547,573]" captionStart="FIGURE 11" captionStartId="17.[151,250,1351,1375]" captionTargetBox="[151,1436,741,1328]" captionTargetId="figure-276@17.[151,1436,741,1328]" captionTargetPageId="17" captionText="FIGURE 11. Penia inopinata, holotype, female (6349) (= paratype of Penia takasago). A: pregenital segments, ventral side; B: slide; C: pregenital segments and ovipositor, ventral side; D: coxites, ventral side." figureDoi="http://doi.org/10.5281/zenodo.10200059" httpUri="https://zenodo.org/record/10200059/files/figure.png" pageId="17" pageNumber="318">Fig. 11A</figureCitation>
; the pregenital segments of
<typeStatus id="54BC8599FFB7CF75FBB2E938FB3BFDD0" box="[1122,1220,547,573]" pageId="17" pageNumber="318" type="holotype">holotype</typeStatus>
had been mounted in balsam on slides and were distorted by the pressure exerted by the coverslip,
<figureCitation id="133C27BEFFB7CF75FBE2E95CFB44FD8C" box="[1074,1211,583,609]" captionStart="FIGURE 11" captionStartId="17.[151,250,1351,1375]" captionTargetBox="[151,1436,741,1328]" captionTargetId="figure-276@17.[151,1436,741,1328]" captionTargetPageId="17" captionText="FIGURE 11. Penia inopinata, holotype, female (6349) (= paratype of Penia takasago). A: pregenital segments, ventral side; B: slide; C: pregenital segments and ovipositor, ventral side; D: coxites, ventral side." figureDoi="http://doi.org/10.5281/zenodo.10200059" httpUri="https://zenodo.org/record/10200059/files/figure.png" pageId="17" pageNumber="318">Fig. 11B, C</figureCitation>
); spiculum ventrale 5.45 x longer than length of sternite VIII (
<figureCitation id="133C27BEFFB7CF75FDA5E970FD24FD68" box="[629,731,619,645]" captionStart="FIGURE 11" captionStartId="17.[151,250,1351,1375]" captionTargetBox="[151,1436,741,1328]" captionTargetId="figure-276@17.[151,1436,741,1328]" captionTargetPageId="17" captionText="FIGURE 11. Penia inopinata, holotype, female (6349) (= paratype of Penia takasago). A: pregenital segments, ventral side; B: slide; C: pregenital segments and ovipositor, ventral side; D: coxites, ventral side." figureDoi="http://doi.org/10.5281/zenodo.10200059" httpUri="https://zenodo.org/record/10200059/files/figure.png" pageId="17" pageNumber="318">Fig. 11C</figureCitation>
). Ovipositor 1.2 x longer than length of abdomen; coxites two segmented at ventral side, with three setae each dorsally, ventrally, and apically; stylus with several setae around apex (
<figureCitation id="133C27BEFFB7CF75FF0BE9A8FEBDFD20" box="[219,322,691,717]" captionStart="FIGURE 11" captionStartId="17.[151,250,1351,1375]" captionTargetBox="[151,1436,741,1328]" captionTargetId="figure-276@17.[151,1436,741,1328]" captionTargetPageId="17" captionText="FIGURE 11. Penia inopinata, holotype, female (6349) (= paratype of Penia takasago). A: pregenital segments, ventral side; B: slide; C: pregenital segments and ovipositor, ventral side; D: coxites, ventral side." figureDoi="http://doi.org/10.5281/zenodo.10200059" httpUri="https://zenodo.org/record/10200059/files/figure.png" pageId="17" pageNumber="318">Fig. 11D</figureCitation>
). Vagina and bursa copulatrix lost in
<typeStatus id="54BC8599FFB7CF75FD32E9A8FCBBFD20" box="[738,836,691,717]" pageId="17" pageNumber="318" type="holotype">holotype</typeStatus>
(
<figureCitation id="133C27BEFFB7CF75FC84E9A8FC46FD20" box="[852,953,691,717]" captionStart="FIGURE 11" captionStartId="17.[151,250,1351,1375]" captionTargetBox="[151,1436,741,1328]" captionTargetId="figure-276@17.[151,1436,741,1328]" captionTargetPageId="17" captionText="FIGURE 11. Penia inopinata, holotype, female (6349) (= paratype of Penia takasago). A: pregenital segments, ventral side; B: slide; C: pregenital segments and ovipositor, ventral side; D: coxites, ventral side." figureDoi="http://doi.org/10.5281/zenodo.10200059" httpUri="https://zenodo.org/record/10200059/files/figure.png" pageId="17" pageNumber="318">Fig. 11C</figureCitation>
).
</paragraph>
<caption id="DF786BB3FFB7CF75FF47EE5CFC2BFA69" ID-DOI="http://doi.org/10.5281/zenodo.10200059" ID-Zenodo-Dep="10200059" httpUri="https://zenodo.org/record/10200059/files/figure.png" pageId="17" pageNumber="318" startId="17.[151,250,1351,1375]" targetBox="[151,1436,741,1328]" targetPageId="17" targetType="figure">
<paragraph id="8BB83B3BFFB7CF75FF47EE5CFC2BFA69" blockId="17.[151,1436,1351,1412]" pageId="17" pageNumber="318">
<emphasis id="B973E729FFB7CF75FF47EE5CFEE1FAB2" bold="true" box="[151,286,1351,1375]" pageId="17" pageNumber="318">FIGURE 11.</emphasis>
<taxonomicName id="4C0740B8FFB7CF75FEF3EE53FE3AFAB2" authorityName="Arimoto" authorityYear="2023" box="[291,453,1352,1375]" class="Insecta" family="Elateridae" genus="Penia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="318" phylum="Arthropoda" rank="species" species="inopinata">
<emphasis id="B973E729FFB7CF75FEF3EE53FE3AFAB2" box="[291,453,1352,1375]" italics="true" pageId="17" pageNumber="318">Penia inopinata</emphasis>
</taxonomicName>
, holotype, female (6349) (= paratype of
<taxonomicName id="4C0740B8FFB7CF75FC8FEE53FC05FA8D" authorityName="Kishii" authorityYear="1997" box="[863,1018,1352,1376]" class="Insecta" family="Elateridae" genus="Penia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="318" phylum="Arthropoda" rank="species" species="takasago">
<emphasis id="B973E729FFB7CF75FC8FEE53FC05FA8D" box="[863,1018,1352,1376]" italics="true" pageId="17" pageNumber="318">Penia takasago</emphasis>
</taxonomicName>
). A: pregenital segments, ventral side; B: slide; C: pregenital segments and ovipositor, ventral side; D: coxites, ventral side.
</paragraph>
</caption>
<paragraph id="8BB83B3BFFB7CF75FF17EEA9FE77FA21" blockId="17.[151,1437,1458,1988]" box="[199,392,1458,1484]" pageId="17" pageNumber="318">
<emphasis id="B973E729FFB7CF75FF17EEA9FEF4FA21" bold="true" box="[199,267,1458,1484]" pageId="17" pageNumber="318">Male.</emphasis>
Unknown.
</paragraph>
</subSubSection>
<subSubSection id="C31D68B0FFB7CF75FF17EECEFB96F84D" pageId="17" pageNumber="318" type="discussion">
<paragraph id="8BB83B3BFFB7CF75FF17EECEFB96F84D" blockId="17.[151,1437,1458,1988]" pageId="17" pageNumber="318">
<emphasis id="B973E729FFB7CF75FF17EECEFEB3FA02" bold="true" box="[199,332,1493,1519]" pageId="17" pageNumber="318">Discussion.</emphasis>
<specimenCount id="9D01F0B2FFB7CF75FE82EECDFE27FA1D" box="[338,472,1494,1520]" count="4" pageId="17" pageNumber="318" type="female">Four female</specimenCount>
types were assigned to
<taxonomicName id="4C0740B8FFB7CF75FD0DEECCFCA0FA1D" authorityName="Kishii" authorityYear="1997" box="[733,863,1494,1520]" class="Insecta" family="Elateridae" genus="Penia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="318" phylum="Arthropoda" rank="species" species="takasago">
<emphasis id="B973E729FFB7CF75FD0DEECCFD11FA1D" box="[733,750,1495,1520]" italics="true" pageId="17" pageNumber="318">P</emphasis>
.
<emphasis id="B973E729FFB7CF75FD2AEECDFCA0FA1D" box="[762,863,1494,1520]" italics="true" pageId="17" pageNumber="318">takasago</emphasis>
</taxonomicName>
in the original description (
<bibRefCitation id="EF9646CAFFB7CF75FB5FEECDFAE1FA1D" author="Kishii, T." box="[1167,1310,1494,1520]" pageId="17" pageNumber="318" pagination="9 - 14" refId="ref23002" refString="Kishii, T. (1997) A study on the elaterid-beetles of Shibata collection from Taiwan. V. (Coleoptera: Elateridae) On the subfamily Denticollinae: tribes Denticollini, Hemicrepidiini and Dimini. The Entomological Review of Japan, 52, 9 - 14." type="journal article" year="1997">Kishii, 1997</bibRefCitation>
). However, the
<typeStatus id="54BC8599FFB7CF75FF13EEE1FEDCF9F9" box="[195,291,1530,1556]" pageId="17" pageNumber="318" type="paratype">paratype</typeStatus>
(6349) is distinguished from the
<typeStatus id="54BC8599FFB7CF75FD76EEE1FCF7F9F9" box="[678,776,1530,1556]" pageId="17" pageNumber="318" type="holotype">holotype</typeStatus>
and other
<specimenCount id="9D01F0B2FFB7CF75FC56EEE1FBDBF9F9" box="[902,1060,1530,1556]" count="2" pageId="17" pageNumber="318" type="generic" typeStatus="paratypes">two paratypes</specimenCount>
of
<taxonomicName id="4C0740B8FFB7CF75FB9FEEE0FB2AF9F9" authorityName="Kishii" authorityYear="1997" box="[1103,1237,1530,1556]" class="Insecta" family="Elateridae" genus="Penia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="318" phylum="Arthropoda" rank="species" species="takasago">
<emphasis id="B973E729FFB7CF75FB9FEEE0FB9FF9F9" box="[1103,1120,1531,1556]" italics="true" pageId="17" pageNumber="318">P</emphasis>
.
<emphasis id="B973E729FFB7CF75FBA0EEE1FB2AF9F9" box="[1136,1237,1530,1556]" italics="true" pageId="17" pageNumber="318">takasago</emphasis>
</taxonomicName>
by the following contrasting characters (the
<typeStatus id="54BC8599FFB7CF75FE1CED05FDD1F9D5" box="[460,558,1566,1592]" pageId="17" pageNumber="318" type="holotype">holotype</typeStatus>
and other
<specimenCount id="9D01F0B2FFB7CF75FD7BED05FCB6F9D5" box="[683,841,1566,1592]" count="2" pageId="17" pageNumber="318" type="generic" typeStatus="paratypes">two paratypes</specimenCount>
in parentheses): eyes 0.25 x longer than interocular distance in dorsal view (eyes 0.4 x longer than interocular distance in dorsal view); apical maxillary palpomere almost as long as maximum length of eye (apical maxillary palpomere shorter than maximum length of eye); pronotum slightly and straightly narrowed ahead of hind angles (pronotum strongly and straightly narrowed ahead of hind angles); hind angles of pronotum broad, not protruding laterally beyond pronotal side (hind angles of pronotum acute, strongly protruding posterolaterad); hypomeron with slight mesial projection (hypomeron with moderate mesial projection); hind angle of hypomeron broadly triangular (hind angle of hypomeron claw-like shaped); mesosternal process between mesocoxae higher than mesocoxae, visible in lateral view (mesosternal process between mesocoxae lower than mesocoxae, not visible in lateral view); ovipositor longer than length of abdomen (ovipositor shorter than length of abdomen). Moreover, the
<typeStatus id="54BC8599FFB7CF75FC4BEC79FC04F891" box="[923,1019,1890,1916]" pageId="17" pageNumber="318" type="paratype">paratype</typeStatus>
is different from the other Taiwanese congeners (see diagnosis). Therefore, the
<typeStatus id="54BC8599FFB7CF75FDBAEC9DFD35F84D" box="[618,714,1926,1952]" pageId="17" pageNumber="318" type="paratype">paratype</typeStatus>
was determined to be a new species.
</paragraph>
</subSubSection>
<subSubSection id="C31D68B0FFB7CF75FF17ECB2FD31F829" box="[199,718,1961,1988]" pageId="17" pageNumber="318" type="distribution">
<paragraph id="8BB83B3BFFB7CF75FF17ECB2FD31F829" blockId="17.[151,1437,1458,1988]" box="[199,718,1961,1988]" pageId="17" pageNumber="318">
<emphasis id="B973E729FFB7CF75FF17ECB2FE9EF82E" bold="true" box="[199,353,1961,1987]" pageId="17" pageNumber="318">Distribution.</emphasis>
<collectingCountry id="F3107BABFFB7CF75FEB7ECB1FE42F829" box="[359,445,1962,1988]" name="Taiwan" pageId="17" pageNumber="318">Taiwan</collectingCountry>
:
<collectingRegion id="49C3F5D9FFB7CF75FE18ECB1FD94F829" box="[456,619,1962,1988]" country="Taiwan" name="Chiayi" pageId="17" pageNumber="318">Chiayi County</collectingRegion>
(
<figureCitation id="133C27BEFFB7CF75FDABECB1FD42F829" box="[635,701,1962,1988]" captionStart="FIGURE 1" captionStartId="3.[151,250,902,926]" captionTargetBox="[158,1429,188,872]" captionTargetId="figure-16@3.[151,1436,181,879]" captionTargetPageId="3" captionText="FIGURE 1. Map of Taiwan. A: location of Taiwan; B: the collection sites of the material used in this study." figureDoi="http://doi.org/10.5281/zenodo.10200038" httpUri="https://zenodo.org/record/10200038/files/figure.png" pageId="17" pageNumber="318">Fig. 1</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>