704 lines
108 KiB
XML
704 lines
108 KiB
XML
<document ID-DOI="10.11646/zootaxa.5277.2.6" ID-ISSN="1175-5326" ID-Zenodo-Dep="7889968" ID-ZooBank="BD8D27A6-0246-44F7-852C-CB0CDDBBAE8F" IM.bibliography_approvedBy="diego" IM.illustrations_approvedBy="diego" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_approvedBy="diego" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="diego" checkinTime="1683100361008" checkinUser="plazi" docAuthor="Komai, Tomoyuki & Henmi, Yumi" docDate="2023" docId="03AE8784CC6A9D62EEA7F96DFED3322A" docLanguage="en" docName="zootaxa.5277.2.6.pdf" docOrigin="Zootaxa 5277 (2)" docSource="http://dx.doi.org/10.11646/zootaxa.5277.2.6" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Athanas exilis Komai & Henmi 2023, n. sp." docType="treatment" docVersion="4" lastPageNumber="370" masterDocId="FF97FFFCCC6B9D65EE30FFEAFFB83572" masterDocTitle="A new species of the alpheid shrimp genus Athanas Leach, 1814 (Decapoda: Caridea) from the Sea of Japan" masterLastPageNumber="373" masterPageNumber="363" pageNumber="364" updateTime="1683131623977" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:titleInfo>
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<mods:title>A new species of the alpheid shrimp genus Athanas Leach, 1814 (Decapoda: Caridea) from the Sea of Japan</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Komai, Tomoyuki</mods:namePart>
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<mods:affiliation>Natural History Museum and Institute, Chiba, 955 - 2 Aoba-cho, Chuo-ku, Chiba, 260 - 8682 Japan.</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Henmi, Yumi</mods:namePart>
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<mods:nameIdentifier type="ORCID">0000-0001-9141-1769</mods:nameIdentifier>
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<mods:affiliation>Maizuru Fisheries Research Station, Kyoto University, Nagahama, Maizuru, Kyoto, 625 - 0086 Japan. henmi. yumi. 7 c @ kyoto-u. ac. jp; https: // orcid. org / 0000 - 0001 - 9141 - 1769</mods:affiliation>
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<mods:nameIdentifier type="email">henmi.yumi.7c@kyoto-u.ac.jp</mods:nameIdentifier>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2023</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2023-05-02</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>5277</mods:number>
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</mods:detail>
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<mods:detail type="issue">
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<mods:number>2</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>363</mods:start>
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<mods:end>373</mods:end>
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<mods:location>
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<mods:url>http://dx.doi.org/10.11646/zootaxa.5277.2.6</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">10.11646/zootaxa.5277.2.6</mods:identifier>
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<mods:identifier type="ISSN">1175-5326</mods:identifier>
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<mods:identifier type="Zenodo-Dep">7889968</mods:identifier>
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<mods:identifier type="ZooBank">BD8D27A6-0246-44F7-852C-CB0CDDBBAE8F</mods:identifier>
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</mods:mods>
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<treatment ID-DOI="http://doi.org/10.5281/zenodo.7893015" ID-Zenodo-Dep="7893015" LSID="urn:lsid:plazi:treatment:03AE8784CC6A9D62EEA7F96DFED3322A" httpUri="http://treatment.plazi.org/id/03AE8784CC6A9D62EEA7F96DFED3322A" lastPageId="7" lastPageNumber="370" pageId="1" pageNumber="364">
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<subSubSection pageId="1" pageNumber="364" type="nomenclature">
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<paragraph blockId="1.[151,653,1671,1769]" box="[151,397,1671,1698]" pageId="1" pageNumber="364">
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<heading bold="true" box="[151,397,1671,1698]" fontSize="11" level="1" pageId="1" pageNumber="364" reason="1">
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<emphasis bold="true" box="[151,397,1671,1698]" pageId="1" pageNumber="364">
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<taxonomicName authority="Komai & Henmi, 2023" authorityName="Komai & Henmi" authorityYear="2023" box="[151,324,1671,1698]" class="Malacostraca" family="Alpheidae" genus="Athanas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="364" phylum="Arthropoda" rank="species" species="exilis" status="n. sp.">
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<emphasis bold="true" box="[151,324,1671,1698]" italics="true" pageId="1" pageNumber="364">Athanas exilis</emphasis>
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</taxonomicName>
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<taxonomicNameLabel box="[330,397,1671,1697]" pageId="1" pageNumber="364" rank="species">n. sp.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph blockId="1.[151,653,1671,1769]" box="[151,653,1707,1734]" pageId="1" pageNumber="364">[New Japanese name: Wakasa-murasaki-ebi]</paragraph>
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</subSubSection>
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<subSubSection box="[151,274,1743,1769]" pageId="1" pageNumber="364" type="description">
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<paragraph blockId="1.[151,653,1671,1769]" box="[151,274,1743,1769]" pageId="1" pageNumber="364">
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(
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<figureCitation box="[159,266,1743,1769]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStart-3="FIGURE 4" captionStartId-0="2.[151,250,1681,1705]" captionStartId-1="4.[151,250,1876,1900]" captionStartId-2="6.[151,250,1866,1890]" captionStartId-3="7.[151,250,1100,1124]" captionTargetBox-0="[151,1435,181,1657]" captionTargetBox-1="[211,1373,191,1840]" captionTargetBox-2="[210,1376,193,1833]" captionTargetBox-3="[156,1432,189,1072]" captionTargetId-0="figure-21@2.[151,1436,181,1657]" captionTargetId-1="figure-21@4.[201,1386,181,1852]" captionTargetId-2="figure-21@6.[202,1386,181,1842]" captionTargetId-3="figure-17@7.[151,1436,181,1076]" captionTargetPageId-0="2" captionTargetPageId-1="4" captionTargetPageId-2="6" captionTargetPageId-3="7" captionText-0="FIGURE 1. Athanas exilis n. sp., holotype, ovigerous female (cl 4.8 mm), CBM-ZC 17112, showing colouration in life. A, habitus in dorsal view; B, habitus in ventrolateral view. Photos taken by Y. Henmi." captionText-1="FIGURE 2. Athanas exilis n. sp., holotype, ovigerous female (cl 4.8 mm), CBM-ZC 17112.A, carapaceand cephalic appendages, left lateral view; B, anterior part of carapace and cephalic appendages, left lateral view; C, same, dorsal view; D, pleon, telson and pleonal appendages, left lateral view; E, posterior part of pleomere 5 and pleomere 6, left lateral view; F, telson, dorsal view; G, left antennal scaphocerite, dorsal view (setae omitted); H, right uropod, dorsal view." captionText-2="FIGURE 3. Athanas exilis n. sp., holotype, ovigerous female (cl 4.8 mm), CBM-ZC 17112. A, left maxilliped 3 and pereopod 1 (in process of regeneration), lateral view; B, coxae of chelipeds (pereopods 1) (basal part of right pereopod 2 also shown); C, basis of right cheliped, lateral view; D, right cheliped, lateral view; E, same, mesial view; F, left pereopod 2, lateral view; G–I, left pereopods 3–5, lateral view. Abbreviations: EXP = exopod; P1 = pereopod 1; P2 = pereopod 2" captionText-3="FIGURE 4. Athanas exilis n. sp., holotype, ovigerous female (cl 4.8 mm), CBM-ZC 17112. A, close up of ventrolateral margin of ischium of right cheliped; B, carpus of right cheliped, extensor view; C, merus and carpus of right cheliped, ventrolateral view; D, chela of right cheliped, extensor view; E, same, ventrolateral view; F–H, dactyli of pereopods 3–5, lateral view." figureDoi-0="http://doi.org/10.5281/zenodo.7889970" figureDoi-1="http://doi.org/10.5281/zenodo.7889974" figureDoi-2="http://doi.org/10.5281/zenodo.7889976" figureDoi-3="http://doi.org/10.5281/zenodo.7889978" httpUri-0="https://zenodo.org/record/7889970/files/figure.png" httpUri-1="https://zenodo.org/record/7889974/files/figure.png" httpUri-2="https://zenodo.org/record/7889976/files/figure.png" httpUri-3="https://zenodo.org/record/7889978/files/figure.png" pageId="1" pageNumber="364">Figs. 1–4</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection pageId="1" pageNumber="364" type="materials_examined">
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<paragraph blockId="1.[151,1436,1815,1914]" pageId="1" pageNumber="364">
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<emphasis bold="true" box="[151,383,1815,1841]" pageId="1" pageNumber="364">Material examined.</emphasis>
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<materialsCitation collectingDate="2022-03-14" collectionCode="RV, SM" collectorName="Y. Henmi." elevation="90" latitude="35.746387" location="Wakasa Bay" longLatPrecision="19" longitude="135.3361" pageId="1" pageNumber="364" specimenCount="1" typeStatus="holotype">
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<typeStatus box="[388,497,1815,1841]" pageId="1" pageNumber="364">Holotype</typeStatus>
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: CBM-ZC 17112, ovigerous female (cl
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<quantity box="[932,1021,1815,1841]" metricMagnitude="-3" metricUnit="m" metricValue="4.8" pageId="1" pageNumber="364" unit="mm" value="4.8">4.8 mm</quantity>
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),
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<location LSID="urn:lsid:plazi:treatment:03AE8784CC6A9D62EEA7F96DFED3322A:8ED86049CC6A9D64EA3DF8FDFB2F3240" box="[1037,1175,1815,1842]" latitude="35.746387" longLatPrecision="19" longitude="135.3361" name="Wakasa Bay" pageId="1" pageNumber="364">Wakasa Bay</location>
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,
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<locationDeviation box="[1185,1294,1815,1842]" location="Ine" pageId="1" pageNumber="364">NE of Ine</locationDeviation>
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,
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<geoCoordinate box="[1304,1436,1815,1842]" degrees="35" direction="north" minutes="44" orientation="latitude" pageId="1" pageNumber="364" precision="15" seconds="47" value="35.746387">35°44'47"N</geoCoordinate>
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<geoCoordinate box="[151,297,1851,1878]" degrees="135" direction="east" minutes="20" orientation="longitude" pageId="1" pageNumber="364" precision="15" seconds="10" value="135.3361">135°20'10”E</geoCoordinate>
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,
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<quantity box="[309,366,1851,1878]" metricMagnitude="1" metricUnit="m" metricValue="9.0" pageId="1" pageNumber="364" unit="m" value="90.0">
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<elevation box="[309,366,1851,1878]" metricMagnitude="1" metricUnit="m" metricValue="9.0" pageId="1" pageNumber="364" unit="m" value="90.0">90 m</elevation>
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</quantity>
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deep, mud,
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<date box="[503,677,1851,1877]" pageId="1" pageNumber="364" value="2022-03-14">
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<collectingDate box="[503,677,1851,1877]" pageId="1" pageNumber="364" value="2022-03-14">14 March 2022</collectingDate>
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</date>
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||
,
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<collectionCode box="[688,724,1852,1878]" country="Italy" name="Collection of Leptospira Strains" pageId="1" pageNumber="364">RV</collectionCode>
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“Ryokuyo-maru”,
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<collectionCode box="[939,979,1851,1877]" country="Malaysia" name="Sarawak Museum" pageId="1" pageNumber="364">SM</collectionCode>
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grab, coll.
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<collectorName box="[1105,1220,1851,1878]" pageId="1" pageNumber="364">Y. Henmi.</collectorName>
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</materialsCitation>
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</paragraph>
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||
</subSubSection>
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||
<subSubSection lastPageId="5" lastPageNumber="368" pageId="1" pageNumber="364" type="description">
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||
<paragraph blockId="1.[151,1436,1815,1914]" box="[199,664,1887,1914]" pageId="1" pageNumber="364">
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||
<emphasis bold="true" box="[199,345,1887,1913]" pageId="1" pageNumber="364">Description.</emphasis>
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Body (
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<figureCitation box="[428,505,1887,1913]" captionStart="FIGURE 1" captionStartId="2.[151,250,1681,1705]" captionTargetBox="[151,1435,181,1657]" captionTargetId="figure-21@2.[151,1436,181,1657]" captionTargetPageId="2" captionText="FIGURE 1. Athanas exilis n. sp., holotype, ovigerous female (cl 4.8 mm), CBM-ZC 17112, showing colouration in life. A, habitus in dorsal view; B, habitus in ventrolateral view. Photos taken by Y. Henmi." figureDoi="http://doi.org/10.5281/zenodo.7889970" httpUri="https://zenodo.org/record/7889970/files/figure.png" pageId="1" pageNumber="364">Figs. 1</figureCitation>
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,
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<figureCitation box="[519,586,1887,1913]" captionStart="FIGURE 2" captionStartId="4.[151,250,1876,1900]" captionTargetBox="[211,1373,191,1840]" captionTargetId="figure-21@4.[201,1386,181,1852]" captionTargetPageId="4" captionText="FIGURE 2. Athanas exilis n. sp., holotype, ovigerous female (cl 4.8 mm), CBM-ZC 17112.A, carapaceand cephalic appendages, left lateral view; B, anterior part of carapace and cephalic appendages, left lateral view; C, same, dorsal view; D, pleon, telson and pleonal appendages, left lateral view; E, posterior part of pleomere 5 and pleomere 6, left lateral view; F, telson, dorsal view; G, left antennal scaphocerite, dorsal view (setae omitted); H, right uropod, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.7889974" httpUri="https://zenodo.org/record/7889974/files/figure.png" pageId="1" pageNumber="364">2A, B</figureCitation>
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) stout.
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</paragraph>
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<caption ID-DOI="http://doi.org/10.5281/zenodo.7889970" ID-Zenodo-Dep="7889970" httpUri="https://zenodo.org/record/7889970/files/figure.png" pageId="2" pageNumber="365" startId="2.[151,250,1681,1705]" targetBox="[151,1435,181,1657]" targetPageId="2" targetType="figure">
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<paragraph blockId="2.[151,1436,1681,1741]" pageId="2" pageNumber="365">
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<emphasis bold="true" box="[151,277,1681,1705]" pageId="2" pageNumber="365">FIGURE 1.</emphasis>
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<taxonomicName authority="Komai & Henmi, 2023" authorityName="Komai & Henmi" authorityYear="2023" box="[285,429,1681,1705]" class="Malacostraca" family="Alpheidae" genus="Athanas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="365" phylum="Arthropoda" rank="species" species="exilis" status="n. sp.">
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<emphasis box="[285,429,1681,1705]" italics="true" pageId="2" pageNumber="365">Athanas exilis</emphasis>
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</taxonomicName>
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<emphasis bold="true" box="[438,497,1681,1705]" pageId="2" pageNumber="365">
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<taxonomicNameLabel box="[438,497,1681,1705]" pageId="2" pageNumber="365" rank="species">n. sp.</taxonomicNameLabel>
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</emphasis>
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, holotype, ovigerous female (cl 4.8 mm), CBM-ZC 17112, showing colouration in life. A, habitus in dorsal view; B, habitus in ventrolateral view. Photos taken by Y. Henmi.
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</paragraph>
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</caption>
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<paragraph blockId="2.[151,1436,1787,2030]" pageId="2" pageNumber="365">
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Carapace (
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<figureCitation box="[321,444,1787,1814]" captionStart="FIGURE 2" captionStartId="4.[151,250,1876,1900]" captionTargetBox="[211,1373,191,1840]" captionTargetId="figure-21@4.[201,1386,181,1852]" captionTargetPageId="4" captionText="FIGURE 2. Athanas exilis n. sp., holotype, ovigerous female (cl 4.8 mm), CBM-ZC 17112.A, carapaceand cephalic appendages, left lateral view; B, anterior part of carapace and cephalic appendages, left lateral view; C, same, dorsal view; D, pleon, telson and pleonal appendages, left lateral view; E, posterior part of pleomere 5 and pleomere 6, left lateral view; F, telson, dorsal view; G, left antennal scaphocerite, dorsal view (setae omitted); H, right uropod, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.7889974" httpUri="https://zenodo.org/record/7889974/files/figure.png" pageId="2" pageNumber="365">Fig. 2A–C</figureCitation>
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) glabrous, not setose. Rostrum 0.2 times as long as carapace, flattened dorsoventrally, directed forward, reaching beyond mid-length of basal article of antennular peduncle, proximally broadened, narrow triangular with acute apex in dorsal view; rostral dorsal carina absent. Frontal margin devoid of supracorneal spines; extracorneal spines acuminate, far falling short of distal corneal margin; infra-corneal spines reduced into low convexity. Pterygostomial margin broadly rounded, not protruding anteriorly. Cardiac notch deep.
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</paragraph>
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<paragraph blockId="2.[151,1436,1787,2030]" lastBlockId="3.[151,1437,151,2013]" lastPageId="3" lastPageNumber="366" pageId="2" pageNumber="365">
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Pleura of pleomeres 1–4 (
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<figureCitation box="[503,595,1967,1994]" captionStart="FIGURE 2" captionStartId="4.[151,250,1876,1900]" captionTargetBox="[211,1373,191,1840]" captionTargetId="figure-21@4.[201,1386,181,1852]" captionTargetPageId="4" captionText="FIGURE 2. Athanas exilis n. sp., holotype, ovigerous female (cl 4.8 mm), CBM-ZC 17112.A, carapaceand cephalic appendages, left lateral view; B, anterior part of carapace and cephalic appendages, left lateral view; C, same, dorsal view; D, pleon, telson and pleonal appendages, left lateral view; E, posterior part of pleomere 5 and pleomere 6, left lateral view; F, telson, dorsal view; G, left antennal scaphocerite, dorsal view (setae omitted); H, right uropod, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.7889974" httpUri="https://zenodo.org/record/7889974/files/figure.png" pageId="2" pageNumber="365">Fig. 2D</figureCitation>
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) rounded, unarmed, while that pleomere 5 (
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<figureCitation box="[1114,1241,1967,1994]" captionStart="FIGURE 2" captionStartId="4.[151,250,1876,1900]" captionTargetBox="[211,1373,191,1840]" captionTargetId="figure-21@4.[201,1386,181,1852]" captionTargetPageId="4" captionText="FIGURE 2. Athanas exilis n. sp., holotype, ovigerous female (cl 4.8 mm), CBM-ZC 17112.A, carapaceand cephalic appendages, left lateral view; B, anterior part of carapace and cephalic appendages, left lateral view; C, same, dorsal view; D, pleon, telson and pleonal appendages, left lateral view; E, posterior part of pleomere 5 and pleomere 6, left lateral view; F, telson, dorsal view; G, left antennal scaphocerite, dorsal view (setae omitted); H, right uropod, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.7889974" httpUri="https://zenodo.org/record/7889974/files/figure.png" pageId="2" pageNumber="365">Fig. 2D, E</figureCitation>
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) acutely pointed at posteroventral angle. Pleomere 6 (
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<figureCitation box="[579,704,2003,2030]" captionStart="FIGURE 2" captionStartId="4.[151,250,1876,1900]" captionTargetBox="[211,1373,191,1840]" captionTargetId="figure-21@4.[201,1386,181,1852]" captionTargetPageId="4" captionText="FIGURE 2. Athanas exilis n. sp., holotype, ovigerous female (cl 4.8 mm), CBM-ZC 17112.A, carapaceand cephalic appendages, left lateral view; B, anterior part of carapace and cephalic appendages, left lateral view; C, same, dorsal view; D, pleon, telson and pleonal appendages, left lateral view; E, posterior part of pleomere 5 and pleomere 6, left lateral view; F, telson, dorsal view; G, left antennal scaphocerite, dorsal view (setae omitted); H, right uropod, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.7889974" httpUri="https://zenodo.org/record/7889974/files/figure.png" pageId="2" pageNumber="365">Fig. 2D, E</figureCitation>
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) with articulated triangular flap posteroventrally; posterolateral process pointed but not acuminate. Telson (
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<figureCitation box="[643,728,151,177]" captionStart="FIGURE 2" captionStartId="4.[151,250,1876,1900]" captionTargetBox="[211,1373,191,1840]" captionTargetId="figure-21@4.[201,1386,181,1852]" captionTargetPageId="4" captionText="FIGURE 2. Athanas exilis n. sp., holotype, ovigerous female (cl 4.8 mm), CBM-ZC 17112.A, carapaceand cephalic appendages, left lateral view; B, anterior part of carapace and cephalic appendages, left lateral view; C, same, dorsal view; D, pleon, telson and pleonal appendages, left lateral view; E, posterior part of pleomere 5 and pleomere 6, left lateral view; F, telson, dorsal view; G, left antennal scaphocerite, dorsal view (setae omitted); H, right uropod, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.7889974" httpUri="https://zenodo.org/record/7889974/files/figure.png" pageId="3" pageNumber="366">Fig. 2F</figureCitation>
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) widest at proximal one-fifth, tapering distally, with 2 pairs of dorsolateral spiniform setae; posterior margin medially slightly convex, with 2 pairs of posterolateral spiniform setae, mesial seta almost twice as long as lateral seta.
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</paragraph>
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<paragraph blockId="3.[151,1437,151,2013]" box="[199,1157,259,285]" pageId="3" pageNumber="366">
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Eyes (
|
||
<figureCitation box="[269,388,259,285]" captionStart="FIGURE 2" captionStartId="4.[151,250,1876,1900]" captionTargetBox="[211,1373,191,1840]" captionTargetId="figure-21@4.[201,1386,181,1852]" captionTargetPageId="4" captionText="FIGURE 2. Athanas exilis n. sp., holotype, ovigerous female (cl 4.8 mm), CBM-ZC 17112.A, carapaceand cephalic appendages, left lateral view; B, anterior part of carapace and cephalic appendages, left lateral view; C, same, dorsal view; D, pleon, telson and pleonal appendages, left lateral view; E, posterior part of pleomere 5 and pleomere 6, left lateral view; F, telson, dorsal view; G, left antennal scaphocerite, dorsal view (setae omitted); H, right uropod, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.7889974" httpUri="https://zenodo.org/record/7889974/files/figure.png" pageId="3" pageNumber="366">Fig. 2B, C</figureCitation>
|
||
) largely exposed in dorsal and lateral views; cornea well pigmented.
|
||
</paragraph>
|
||
<paragraph blockId="3.[151,1437,151,2013]" pageId="3" pageNumber="366">
|
||
Antennular peduncle (
|
||
<figureCitation box="[447,565,295,321]" captionStart="FIGURE 2" captionStartId="4.[151,250,1876,1900]" captionTargetBox="[211,1373,191,1840]" captionTargetId="figure-21@4.[201,1386,181,1852]" captionTargetPageId="4" captionText="FIGURE 2. Athanas exilis n. sp., holotype, ovigerous female (cl 4.8 mm), CBM-ZC 17112.A, carapaceand cephalic appendages, left lateral view; B, anterior part of carapace and cephalic appendages, left lateral view; C, same, dorsal view; D, pleon, telson and pleonal appendages, left lateral view; E, posterior part of pleomere 5 and pleomere 6, left lateral view; F, telson, dorsal view; G, left antennal scaphocerite, dorsal view (setae omitted); H, right uropod, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.7889974" httpUri="https://zenodo.org/record/7889974/files/figure.png" pageId="3" pageNumber="366">Fig. 2B, C</figureCitation>
|
||
) basal article longer than distal two articles combined, article 2 slightly longer than article 3. Basal article with stylocerite acute, reaching its distal margin; ventromesial carina with well developed, acute spine. Lateral flagellum biramous, fused proximal portion composed of 8 segments, aesthetasc-bearing shorter ramus (left side damaged) composed of 3 segments.
|
||
</paragraph>
|
||
<paragraph blockId="3.[151,1437,151,2013]" pageId="3" pageNumber="366">
|
||
Antenna (
|
||
<figureCitation box="[311,469,439,465]" captionStart="FIGURE 2" captionStartId="4.[151,250,1876,1900]" captionTargetBox="[211,1373,191,1840]" captionTargetId="figure-21@4.[201,1386,181,1852]" captionTargetPageId="4" captionText="FIGURE 2. Athanas exilis n. sp., holotype, ovigerous female (cl 4.8 mm), CBM-ZC 17112.A, carapaceand cephalic appendages, left lateral view; B, anterior part of carapace and cephalic appendages, left lateral view; C, same, dorsal view; D, pleon, telson and pleonal appendages, left lateral view; E, posterior part of pleomere 5 and pleomere 6, left lateral view; F, telson, dorsal view; G, left antennal scaphocerite, dorsal view (setae omitted); H, right uropod, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.7889974" httpUri="https://zenodo.org/record/7889974/files/figure.png" pageId="3" pageNumber="366">Fig. 2B, C, G</figureCitation>
|
||
) with basicerite bearing small ventrolateral distal spine. Scaphocerite reaching distal end of antennular peduncle, oval, lateral margin straight, distal lamella produced, rounded, distinctly overreaching distolateral spine. Carpocerite exceeding distal margin of scaphocerite by 0.2 length, reaching far beyond distal margin of antennular peduncle.
|
||
</paragraph>
|
||
<paragraph blockId="3.[151,1437,151,2013]" box="[199,820,583,609]" pageId="3" pageNumber="366">Mouthparts without specific features, typical for genus.</paragraph>
|
||
<paragraph blockId="3.[151,1437,151,2013]" pageId="3" pageNumber="366">
|
||
Maxilliped 3 (
|
||
<figureCitation box="[367,460,619,645]" captionStart="FIGURE 3" captionStartId="6.[151,250,1866,1890]" captionTargetBox="[210,1376,193,1833]" captionTargetId="figure-21@6.[202,1386,181,1842]" captionTargetPageId="6" captionText="FIGURE 3. Athanas exilis n. sp., holotype, ovigerous female (cl 4.8 mm), CBM-ZC 17112. A, left maxilliped 3 and pereopod 1 (in process of regeneration), lateral view; B, coxae of chelipeds (pereopods 1) (basal part of right pereopod 2 also shown); C, basis of right cheliped, lateral view; D, right cheliped, lateral view; E, same, mesial view; F, left pereopod 2, lateral view; G–I, left pereopods 3–5, lateral view. Abbreviations: EXP = exopod; P1 = pereopod 1; P2 = pereopod 2" figureDoi="http://doi.org/10.5281/zenodo.7889976" httpUri="https://zenodo.org/record/7889976/files/figure.png" pageId="3" pageNumber="366">Fig. 3A</figureCitation>
|
||
) endopod slender, reaching distal margin of scaphocerite by tip of ultimate article. Ultimate article 1.2 times as long as penultimate article, tapering to blunt tip bearing only stiff setae (no spiniform setae). Antepenultimate article slightly sinuous, slightly widened proximally. Coxal plate not strongly produced, rounded. Exopod not reaching distal end of endopod antepenultimate article.
|
||
</paragraph>
|
||
<paragraph blockId="3.[151,1437,151,2013]" pageId="3" pageNumber="366">
|
||
Left pereopod 1 only bud-like, in process of regeneration (
|
||
<figureCitation box="[847,937,763,789]" captionStart="FIGURE 3" captionStartId="6.[151,250,1866,1890]" captionTargetBox="[210,1376,193,1833]" captionTargetId="figure-21@6.[202,1386,181,1842]" captionTargetPageId="6" captionText="FIGURE 3. Athanas exilis n. sp., holotype, ovigerous female (cl 4.8 mm), CBM-ZC 17112. A, left maxilliped 3 and pereopod 1 (in process of regeneration), lateral view; B, coxae of chelipeds (pereopods 1) (basal part of right pereopod 2 also shown); C, basis of right cheliped, lateral view; D, right cheliped, lateral view; E, same, mesial view; F, left pereopod 2, lateral view; G–I, left pereopods 3–5, lateral view. Abbreviations: EXP = exopod; P1 = pereopod 1; P2 = pereopod 2" figureDoi="http://doi.org/10.5281/zenodo.7889976" httpUri="https://zenodo.org/record/7889976/files/figure.png" pageId="3" pageNumber="366">Fig. 3A</figureCitation>
|
||
). Pereopods 1 coxae (
|
||
<figureCitation box="[1179,1264,763,789]" captionStart="FIGURE 3" captionStartId="6.[151,250,1866,1890]" captionTargetBox="[210,1376,193,1833]" captionTargetId="figure-21@6.[202,1386,181,1842]" captionTargetPageId="6" captionText="FIGURE 3. Athanas exilis n. sp., holotype, ovigerous female (cl 4.8 mm), CBM-ZC 17112. A, left maxilliped 3 and pereopod 1 (in process of regeneration), lateral view; B, coxae of chelipeds (pereopods 1) (basal part of right pereopod 2 also shown); C, basis of right cheliped, lateral view; D, right cheliped, lateral view; E, same, mesial view; F, left pereopod 2, lateral view; G–I, left pereopods 3–5, lateral view. Abbreviations: EXP = exopod; P1 = pereopod 1; P2 = pereopod 2" figureDoi="http://doi.org/10.5281/zenodo.7889976" httpUri="https://zenodo.org/record/7889976/files/figure.png" pageId="3" pageNumber="366">Fig. 3B</figureCitation>
|
||
) asymmetrical, right larger than left, suggesting notable asymmetry of pereopods 1. Right pereopod 1 (cheliped) (
|
||
<figureCitation box="[1304,1432,799,825]" captionStart="FIGURE 3" captionStartId="6.[151,250,1866,1890]" captionTargetBox="[210,1376,193,1833]" captionTargetId="figure-21@6.[202,1386,181,1842]" captionTargetPageId="6" captionText="FIGURE 3. Athanas exilis n. sp., holotype, ovigerous female (cl 4.8 mm), CBM-ZC 17112. A, left maxilliped 3 and pereopod 1 (in process of regeneration), lateral view; B, coxae of chelipeds (pereopods 1) (basal part of right pereopod 2 also shown); C, basis of right cheliped, lateral view; D, right cheliped, lateral view; E, same, mesial view; F, left pereopod 2, lateral view; G–I, left pereopods 3–5, lateral view. Abbreviations: EXP = exopod; P1 = pereopod 1; P2 = pereopod 2" figureDoi="http://doi.org/10.5281/zenodo.7889976" httpUri="https://zenodo.org/record/7889976/files/figure.png" pageId="3" pageNumber="366">Fig. 3E, D</figureCitation>
|
||
,
|
||
<figureCitation box="[151,216,835,861]" captionStart="FIGURE 4" captionStartId="7.[151,250,1100,1124]" captionTargetBox="[156,1432,189,1072]" captionTargetId="figure-17@7.[151,1436,181,1076]" captionTargetPageId="7" captionText="FIGURE 4. Athanas exilis n. sp., holotype, ovigerous female (cl 4.8 mm), CBM-ZC 17112. A, close up of ventrolateral margin of ischium of right cheliped; B, carpus of right cheliped, extensor view; C, merus and carpus of right cheliped, ventrolateral view; D, chela of right cheliped, extensor view; E, same, ventrolateral view; F–H, dactyli of pereopods 3–5, lateral view." figureDoi="http://doi.org/10.5281/zenodo.7889978" httpUri="https://zenodo.org/record/7889978/files/figure.png" pageId="3" pageNumber="366">4B–E</figureCitation>
|
||
) large, carried flexed, overreaching distal margin of scaphocerite by 0.2 length of merus when extended. Basis (
|
||
<figureCitation box="[229,315,871,897]" captionStart="FIGURE 3" captionStartId="6.[151,250,1866,1890]" captionTargetBox="[210,1376,193,1833]" captionTargetId="figure-21@6.[202,1386,181,1842]" captionTargetPageId="6" captionText="FIGURE 3. Athanas exilis n. sp., holotype, ovigerous female (cl 4.8 mm), CBM-ZC 17112. A, left maxilliped 3 and pereopod 1 (in process of regeneration), lateral view; B, coxae of chelipeds (pereopods 1) (basal part of right pereopod 2 also shown); C, basis of right cheliped, lateral view; D, right cheliped, lateral view; E, same, mesial view; F, left pereopod 2, lateral view; G–I, left pereopods 3–5, lateral view. Abbreviations: EXP = exopod; P1 = pereopod 1; P2 = pereopod 2" figureDoi="http://doi.org/10.5281/zenodo.7889976" httpUri="https://zenodo.org/record/7889976/files/figure.png" pageId="3" pageNumber="366">Fig. 3C</figureCitation>
|
||
) with rudimentary exopod. Ischium elongate, 0.8 times as long as merus; ventral margin with row of numerous spinules and 1 proximal spiniform seta, not particularly projecting; dorsal margin with row of 4 long, slender spiniform setae on proximal half; ventromesial facet clearly delimited, mesial margin with row of spinules; proximomesial face with 2 long spiniform setae. Merus elongate; dorsal, lateral and mesial surfaces smooth; ventral surface deeply excavate to accommodate carpus and proximal part of chela, ventrolateral (
|
||
<figureCitation box="[1287,1378,1015,1041]" captionStart="FIGURE 4" captionStartId="7.[151,250,1100,1124]" captionTargetBox="[156,1432,189,1072]" captionTargetId="figure-17@7.[151,1436,181,1076]" captionTargetPageId="7" captionText="FIGURE 4. Athanas exilis n. sp., holotype, ovigerous female (cl 4.8 mm), CBM-ZC 17112. A, close up of ventrolateral margin of ischium of right cheliped; B, carpus of right cheliped, extensor view; C, merus and carpus of right cheliped, ventrolateral view; D, chela of right cheliped, extensor view; E, same, ventrolateral view; F–H, dactyli of pereopods 3–5, lateral view." figureDoi="http://doi.org/10.5281/zenodo.7889978" httpUri="https://zenodo.org/record/7889978/files/figure.png" pageId="3" pageNumber="366">Fig. 4A</figureCitation>
|
||
) and ventromesial margins sharply edged, each with row of widely spaced spinules or granules. Carpus 0.8 length of merus, distally widening, somewhat compressed, without setose area; extensor surface with faint median sulcus; lateral surface smooth, unarmed; mesial surface with 2 irregular longitudinal rows of spinules or granules along flexor margin. Chela also somewhat compressed, distinctly wider than carpus, surface glabrous; mesial margin sinuous with concavity at pollex base; pollex deflexed, terminating in minute clow, with low, rectangular tooth on occlusal margin slightly proximal to midlength; dactylus gradually tapering, crossing fixed finger at tip, 0.7 times as long as palm, occlusal margin with small triangular tooth slightly distal to midlength.
|
||
</paragraph>
|
||
<paragraph blockId="3.[151,1437,151,2013]" pageId="3" pageNumber="366">
|
||
Pereopod 2 (
|
||
<figureCitation box="[349,435,1303,1329]" captionStart="FIGURE 3" captionStartId="6.[151,250,1866,1890]" captionTargetBox="[210,1376,193,1833]" captionTargetId="figure-21@6.[202,1386,181,1842]" captionTargetPageId="6" captionText="FIGURE 3. Athanas exilis n. sp., holotype, ovigerous female (cl 4.8 mm), CBM-ZC 17112. A, left maxilliped 3 and pereopod 1 (in process of regeneration), lateral view; B, coxae of chelipeds (pereopods 1) (basal part of right pereopod 2 also shown); C, basis of right cheliped, lateral view; D, right cheliped, lateral view; E, same, mesial view; F, left pereopod 2, lateral view; G–I, left pereopods 3–5, lateral view. Abbreviations: EXP = exopod; P1 = pereopod 1; P2 = pereopod 2" figureDoi="http://doi.org/10.5281/zenodo.7889976" httpUri="https://zenodo.org/record/7889976/files/figure.png" pageId="3" pageNumber="366">Fig. 3F</figureCitation>
|
||
) slender, overreaching distal margin of scaphocerite by half-length of carpus, without exopod. Ischium slightly shorter than merus. Carpus composed of 5 segments, first segment longer than four others combined; ratio of carpal segments (from proximal to distal) approximately: 6.2: 1: 1: 1.5: 1.6. Chela simple, longer than distal carpal segment, fingers slightly longer than palm.
|
||
</paragraph>
|
||
<paragraph blockId="3.[151,1437,151,2013]" pageId="3" pageNumber="366">
|
||
Pereopods 3–5 slender, generally similar. Pereopod 3 (
|
||
<figureCitation box="[815,903,1447,1473]" captionStart="FIGURE 3" captionStartId="6.[151,250,1866,1890]" captionTargetBox="[210,1376,193,1833]" captionTargetId="figure-21@6.[202,1386,181,1842]" captionTargetPageId="6" captionText="FIGURE 3. Athanas exilis n. sp., holotype, ovigerous female (cl 4.8 mm), CBM-ZC 17112. A, left maxilliped 3 and pereopod 1 (in process of regeneration), lateral view; B, coxae of chelipeds (pereopods 1) (basal part of right pereopod 2 also shown); C, basis of right cheliped, lateral view; D, right cheliped, lateral view; E, same, mesial view; F, left pereopod 2, lateral view; G–I, left pereopods 3–5, lateral view. Abbreviations: EXP = exopod; P1 = pereopod 1; P2 = pereopod 2" figureDoi="http://doi.org/10.5281/zenodo.7889976" httpUri="https://zenodo.org/record/7889976/files/figure.png" pageId="3" pageNumber="366">Fig. 3G</figureCitation>
|
||
) overreaching distal margin of scaphocerite by full length of propodus; ischium with 1 slender spiniform seta at dorsodistal end and 2 spiniform setae on lateral surface ventrally; merus, carpus and propodus unarmed, merus 7.3 times as long as broad; dactylus (
|
||
<figureCitation box="[1300,1385,1519,1545]" captionStart="FIGURE 4" captionStartId="7.[151,250,1100,1124]" captionTargetBox="[156,1432,189,1072]" captionTargetId="figure-17@7.[151,1436,181,1076]" captionTargetPageId="7" captionText="FIGURE 4. Athanas exilis n. sp., holotype, ovigerous female (cl 4.8 mm), CBM-ZC 17112. A, close up of ventrolateral margin of ischium of right cheliped; B, carpus of right cheliped, extensor view; C, merus and carpus of right cheliped, ventrolateral view; D, chela of right cheliped, extensor view; E, same, ventrolateral view; F–H, dactyli of pereopods 3–5, lateral view." figureDoi="http://doi.org/10.5281/zenodo.7889978" httpUri="https://zenodo.org/record/7889978/files/figure.png" pageId="3" pageNumber="366">Fig. 4F</figureCitation>
|
||
) 0.6 times as long as propodus, simple, very slender, gently curved, with paired minute setae slightly distal to midlength. Pereopod 4 (
|
||
<figureCitation box="[295,396,1591,1617]" captionStart="FIGURE 3" captionStartId="6.[151,250,1866,1890]" captionTargetBox="[210,1376,193,1833]" captionTargetId="figure-21@6.[202,1386,181,1842]" captionTargetPageId="6" captionText="FIGURE 3. Athanas exilis n. sp., holotype, ovigerous female (cl 4.8 mm), CBM-ZC 17112. A, left maxilliped 3 and pereopod 1 (in process of regeneration), lateral view; B, coxae of chelipeds (pereopods 1) (basal part of right pereopod 2 also shown); C, basis of right cheliped, lateral view; D, right cheliped, lateral view; E, same, mesial view; F, left pereopod 2, lateral view; G–I, left pereopods 3–5, lateral view. Abbreviations: EXP = exopod; P1 = pereopod 1; P2 = pereopod 2" figureDoi="http://doi.org/10.5281/zenodo.7889976" httpUri="https://zenodo.org/record/7889976/files/figure.png" pageId="3" pageNumber="366">Figs. 3H</figureCitation>
|
||
,
|
||
<figureCitation box="[409,444,1591,1617]" captionStart="FIGURE 4" captionStartId="7.[151,250,1100,1124]" captionTargetBox="[156,1432,189,1072]" captionTargetId="figure-17@7.[151,1436,181,1076]" captionTargetPageId="7" captionText="FIGURE 4. Athanas exilis n. sp., holotype, ovigerous female (cl 4.8 mm), CBM-ZC 17112. A, close up of ventrolateral margin of ischium of right cheliped; B, carpus of right cheliped, extensor view; C, merus and carpus of right cheliped, ventrolateral view; D, chela of right cheliped, extensor view; E, same, ventrolateral view; F–H, dactyli of pereopods 3–5, lateral view." figureDoi="http://doi.org/10.5281/zenodo.7889978" httpUri="https://zenodo.org/record/7889978/files/figure.png" pageId="3" pageNumber="366">4G</figureCitation>
|
||
) overreaching distal margin of scaphocerite by half-length of propodus; ischium with 1 dorsodistal and 1 ventrolateral spiniform setae. Pereopod 5 (
|
||
<figureCitation box="[816,904,1627,1653]" captionStart="FIGURE 3" captionStartId="6.[151,250,1866,1890]" captionTargetBox="[210,1376,193,1833]" captionTargetId="figure-21@6.[202,1386,181,1842]" captionTargetPageId="6" captionText="FIGURE 3. Athanas exilis n. sp., holotype, ovigerous female (cl 4.8 mm), CBM-ZC 17112. A, left maxilliped 3 and pereopod 1 (in process of regeneration), lateral view; B, coxae of chelipeds (pereopods 1) (basal part of right pereopod 2 also shown); C, basis of right cheliped, lateral view; D, right cheliped, lateral view; E, same, mesial view; F, left pereopod 2, lateral view; G–I, left pereopods 3–5, lateral view. Abbreviations: EXP = exopod; P1 = pereopod 1; P2 = pereopod 2" figureDoi="http://doi.org/10.5281/zenodo.7889976" httpUri="https://zenodo.org/record/7889976/files/figure.png" pageId="3" pageNumber="366">Figs. 3I</figureCitation>
|
||
,
|
||
<figureCitation box="[915,950,1627,1653]" captionStart="FIGURE 4" captionStartId="7.[151,250,1100,1124]" captionTargetBox="[156,1432,189,1072]" captionTargetId="figure-17@7.[151,1436,181,1076]" captionTargetPageId="7" captionText="FIGURE 4. Athanas exilis n. sp., holotype, ovigerous female (cl 4.8 mm), CBM-ZC 17112. A, close up of ventrolateral margin of ischium of right cheliped; B, carpus of right cheliped, extensor view; C, merus and carpus of right cheliped, ventrolateral view; D, chela of right cheliped, extensor view; E, same, ventrolateral view; F–H, dactyli of pereopods 3–5, lateral view." figureDoi="http://doi.org/10.5281/zenodo.7889978" httpUri="https://zenodo.org/record/7889978/files/figure.png" pageId="3" pageNumber="366">4H</figureCitation>
|
||
) overreaching distal margin of scaphocerite by length of dactylus; ischium with 1 dorsodistal spiniform seta, but no ventrolateral spiniform seta; propodus with grooming setae distally.
|
||
</paragraph>
|
||
<paragraph blockId="3.[151,1437,151,2013]" pageId="3" pageNumber="366">Gill formula typical for genus: pleurobranchs on thoracomeres 4–8 (above pereopods 1–5); podobranch and arthrobranch absent; lobe-like epipods on maxilliped 1 and 2, strap-like epipods on coxae of maxilliped 3 and pereopods 1–3; setobranchs on coxae of pereopods 1–4; maxillipeds 1 and 2 each with well-developed, flagellumlike exopod.</paragraph>
|
||
<paragraph blockId="3.[151,1437,151,2013]" box="[199,796,1878,1905]" pageId="3" pageNumber="366">Pleopod 2–5 each with appendix interna on endopod.</paragraph>
|
||
<paragraph blockId="3.[151,1437,151,2013]" pageId="3" pageNumber="366">
|
||
Uropod (
|
||
<figureCitation box="[305,398,1915,1941]" captionStart="FIGURE 2" captionStartId="4.[151,250,1876,1900]" captionTargetBox="[211,1373,191,1840]" captionTargetId="figure-21@4.[201,1386,181,1852]" captionTargetPageId="4" captionText="FIGURE 2. Athanas exilis n. sp., holotype, ovigerous female (cl 4.8 mm), CBM-ZC 17112.A, carapaceand cephalic appendages, left lateral view; B, anterior part of carapace and cephalic appendages, left lateral view; C, same, dorsal view; D, pleon, telson and pleonal appendages, left lateral view; E, posterior part of pleomere 5 and pleomere 6, left lateral view; F, telson, dorsal view; G, left antennal scaphocerite, dorsal view (setae omitted); H, right uropod, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.7889974" httpUri="https://zenodo.org/record/7889974/files/figure.png" pageId="3" pageNumber="366">Fig. 2H</figureCitation>
|
||
) with protopod bearing two distally acute lobes. Exopod lateral margin nearly straight, terminating in minute denticle; distolateral spiniform seta minute; diaeresis straight, unarmed. Endopod subovate, subequal in length to exopod.
|
||
</paragraph>
|
||
<caption ID-DOI="http://doi.org/10.5281/zenodo.7889974" ID-Zenodo-Dep="7889974" httpUri="https://zenodo.org/record/7889974/files/figure.png" pageId="4" pageNumber="367" startId="4.[151,250,1876,1900]" targetBox="[211,1373,191,1840]" targetPageId="4" targetType="figure">
|
||
<paragraph blockId="4.[151,1437,1876,2008]" pageId="4" pageNumber="367">
|
||
<emphasis bold="true" box="[151,273,1876,1900]" pageId="4" pageNumber="367">FIGURE 2.</emphasis>
|
||
<taxonomicName authority="Komai & Henmi, 2023" authorityName="Komai & Henmi" authorityYear="2023" box="[277,417,1876,1900]" class="Malacostraca" family="Alpheidae" genus="Athanas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="367" phylum="Arthropoda" rank="species" species="exilis" status="n. sp.">
|
||
<emphasis box="[277,417,1876,1900]" italics="true" pageId="4" pageNumber="367">Athanas exilis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis bold="true" box="[422,477,1876,1900]" pageId="4" pageNumber="367">
|
||
<taxonomicNameLabel box="[422,477,1876,1900]" pageId="4" pageNumber="367" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, holotype, ovigerous female (cl 4.8 mm), CBM-ZC 17112.A, carapace and cephalic appendages, left lateral view; B, anterior part of carapace and cephalic appendages, left lateral view; C, same, dorsal view; D, pleon, telson and pleonal appendages, left lateral view; E, posterior part of pleomere 5 and pleomere 6, left lateral view; F, telson, dorsal view; G, left antennal scaphocerite, dorsal view (setae omitted); H, right uropod, dorsal view.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph blockId="5.[151,1437,150,2036]" pageId="5" pageNumber="368">
|
||
<emphasis bold="true" box="[199,429,150,176]" pageId="5" pageNumber="368">Colouration in life.</emphasis>
|
||
Carapace and pleon semitransparent with scattered red dots (sparse on carapace), dorsal surface with yellowish white longitudinal band along midline; yellowish hepatopancreas visible through integument; antennule and antenna semitransparent; cheliped whitish with tinge of pink; pereopods 2–5 whitish semitransparent; uropods pinkish; eggs red (
|
||
<figureCitation box="[455,577,258,284]" captionStart="FIGURE 1" captionStartId="2.[151,250,1681,1705]" captionTargetBox="[151,1435,181,1657]" captionTargetId="figure-21@2.[151,1436,181,1657]" captionTargetPageId="2" captionText="FIGURE 1. Athanas exilis n. sp., holotype, ovigerous female (cl 4.8 mm), CBM-ZC 17112, showing colouration in life. A, habitus in dorsal view; B, habitus in ventrolateral view. Photos taken by Y. Henmi." figureDoi="http://doi.org/10.5281/zenodo.7889970" httpUri="https://zenodo.org/record/7889970/files/figure.png" pageId="5" pageNumber="368">Fig. 1A, B</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph blockId="5.[151,1437,150,2036]" pageId="5" pageNumber="368">
|
||
<emphasis bold="true" box="[199,298,293,319]" pageId="5" pageNumber="368">Habitat.</emphasis>
|
||
The
|
||
<typeStatus box="[352,450,293,319]" pageId="5" pageNumber="368">holotype</typeStatus>
|
||
of the new species was extracted from muddy sediment collected using a Smith McIntyre grab at a depth of
|
||
<specimenCount box="[351,415,329,356]" pageId="5" pageNumber="368" type="generic">
|
||
<quantity box="[351,411,329,356]" metricMagnitude="1" metricUnit="m" metricValue="9.0" pageId="5" pageNumber="368" unit="m" value="90.0">90 m</quantity>
|
||
.
|
||
</specimenCount>
|
||
The bottom salinity was 34.3‰ and the bottom water temperature was 11.8°C on
|
||
<date pageId="5" pageNumber="368" value="2022-03-14">14 March 2022</date>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection box="[199,1437,400,427]" pageId="5" pageNumber="368" type="distribution">
|
||
<paragraph blockId="5.[151,1437,150,2036]" box="[199,1437,400,427]" pageId="5" pageNumber="368">
|
||
<emphasis bold="true" box="[199,353,400,426]" pageId="5" pageNumber="368">Distribution.</emphasis>
|
||
Presently known only from Wakasa Bay, the Sea of
|
||
<collectingCountry box="[918,982,401,427]" name="Japan" pageId="5" pageNumber="368">Japan</collectingCountry>
|
||
side of Honshu,
|
||
<collectingCountry box="[1163,1230,401,427]" name="Japan" pageId="5" pageNumber="368">Japan</collectingCountry>
|
||
, at a depth of
|
||
<specimenCount box="[1375,1437,401,427]" pageId="5" pageNumber="368" type="generic">
|
||
<quantity box="[1375,1432,401,427]" metricMagnitude="1" metricUnit="m" metricValue="9.0" pageId="5" pageNumber="368" unit="m" value="90.0">90 m</quantity>
|
||
.
|
||
</specimenCount>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection lastPageId="7" lastPageNumber="370" pageId="5" pageNumber="368" type="discussion">
|
||
<paragraph blockId="5.[151,1437,150,2036]" pageId="5" pageNumber="368">
|
||
<emphasis bold="true" box="[199,315,437,463]" pageId="5" pageNumber="368">Remarks.</emphasis>
|
||
In the
|
||
<typeStatus box="[392,493,436,462]" pageId="5" pageNumber="368">holotype</typeStatus>
|
||
, the known unique representative of the new taxon, the left cheliped is in the process of regeneration (
|
||
<figureCitation box="[337,427,472,498]" captionStart="FIGURE 3" captionStartId="6.[151,250,1866,1890]" captionTargetBox="[210,1376,193,1833]" captionTargetId="figure-21@6.[202,1386,181,1842]" captionTargetPageId="6" captionText="FIGURE 3. Athanas exilis n. sp., holotype, ovigerous female (cl 4.8 mm), CBM-ZC 17112. A, left maxilliped 3 and pereopod 1 (in process of regeneration), lateral view; B, coxae of chelipeds (pereopods 1) (basal part of right pereopod 2 also shown); C, basis of right cheliped, lateral view; D, right cheliped, lateral view; E, same, mesial view; F, left pereopod 2, lateral view; G–I, left pereopods 3–5, lateral view. Abbreviations: EXP = exopod; P1 = pereopod 1; P2 = pereopod 2" figureDoi="http://doi.org/10.5281/zenodo.7889976" httpUri="https://zenodo.org/record/7889976/files/figure.png" pageId="5" pageNumber="368">Fig. 3A</figureCitation>
|
||
). Nevertheless, the coxae are asymmetrical (right coxa larger than left), indicating that the chelipeds are distinctly unequal and dissimilar. It is assumed that the right cheliped of the
|
||
<typeStatus box="[1166,1264,508,534]" pageId="5" pageNumber="368">holotype</typeStatus>
|
||
corresponds to a major cheliped because of the large size and stoutness.
|
||
</paragraph>
|
||
<paragraph blockId="5.[151,1437,150,2036]" pageId="5" pageNumber="368">
|
||
The important features of
|
||
<taxonomicName authority="Komai & Henmi, 2023" authorityName="Komai & Henmi" authorityYear="2023" box="[509,671,579,605]" class="Malacostraca" family="Alpheidae" genus="Athanas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="368" phylum="Arthropoda" rank="species" species="exilis" status="n. sp.">
|
||
<emphasis box="[509,671,579,605]" italics="true" pageId="5" pageNumber="368">Athanas exilis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis bold="true" box="[683,750,580,606]" pageId="5" pageNumber="368">
|
||
<taxonomicNameLabel box="[683,750,580,606]" pageId="5" pageNumber="368" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
for species differentiation are: the rostrum does not reach the distal margin of the basal article of the antennular peduncle, with a smooth, non-dentate dorsal margin; no infracorneal and supracorneal spines are present on the carapace frontal margin; the antennular peduncle is slender, with the article 2 being distinctly longer than wide; the antennal carpocerite overreaches the distal margin of the scaphocerite; the major cheliped is carried folded; the major cheliped ischium and merus have spinulose ventral margins; the major cheliped fingers are not twisted and devoid of setal brush; the major cheliped carpus is elongate, being longer than the palm; and the pereopods 3–5 are slender, with simple, noticeably elongate dactyli exceeding 0.5 length of propodi. This combination of characters is seen only in
|
||
<emphasis box="[924,1082,829,856]" italics="true" pageId="5" pageNumber="368">
|
||
<taxonomicName authorityName="Anker & Ahyong" authorityYear="2007" box="[924,1078,829,856]" class="Malacostraca" family="Alpheidae" genus="Athanas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="368" phylum="Arthropoda" rank="species" species="sydnyensis">A. sydnyensis</taxonomicName>
|
||
.
|
||
</emphasis>
|
||
</paragraph>
|
||
<paragraph blockId="5.[151,1437,150,2036]" pageId="5" pageNumber="368">
|
||
<taxonomicName authorityName="Anker & Ahyong" authorityYear="2007" box="[199,419,865,891]" class="Malacostraca" family="Alpheidae" genus="Athanas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="368" phylum="Arthropoda" rank="species" species="sydnyensis">
|
||
<emphasis box="[199,419,865,891]" italics="true" pageId="5" pageNumber="368">Athanas sydnyensis</emphasis>
|
||
</taxonomicName>
|
||
is known only by the type series from
|
||
<collectingRegion box="[867,1073,865,892]" country="Australia" name="New South Wales" pageId="5" pageNumber="368">New South Wales</collectingRegion>
|
||
,
|
||
<collectingCountry box="[1083,1189,865,891]" name="Australia" pageId="5" pageNumber="368">Australia</collectingCountry>
|
||
, at depths of
|
||
<quantity box="[1343,1431,865,892]" metricMagnitude="0" metricUnit="m" metricValue="7.0" metricValueMax="10.0" metricValueMin="4.0" pageId="5" pageNumber="368" unit="m" value="7.0" valueMax="10.0" valueMin="4.0">4–10 m</quantity>
|
||
, consisting of a female
|
||
<typeStatus box="[412,510,901,927]" pageId="5" pageNumber="368">holotype</typeStatus>
|
||
and
|
||
<specimenCount box="[568,699,901,927]" pageId="5" pageNumber="368" type="female">four female</specimenCount>
|
||
<typeStatus box="[708,815,902,928]" pageId="5" pageNumber="368">paratypes</typeStatus>
|
||
(
|
||
<bibRefCitation author="Anker, A. & Ahyong, S. T." box="[832,1101,901,927]" pageId="5" pageNumber="368" pagination="17 - 30" refId="ref5101" refString="Anker, A. & Ahyong, S. T. (2007) Description of two new species of the shrimp genus Athanas Leach, 1814 (Decapoda: Alpheidae) from Australia and West Africa, with remarks on A. amazone Holthuis, 1951. Zootaxa, 1563 (1), 17 - 30. https: // doi. org / 10.11646 / zootaxa. 1563.1.2" type="journal article" year="2007">Anker & Ahyong 2007</bibRefCitation>
|
||
).
|
||
<taxonomicName authority="Komai & Henmi, 2023" authorityName="Komai & Henmi" authorityYear="2023" box="[1122,1281,901,927]" class="Malacostraca" family="Alpheidae" genus="Athanas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="368" phylum="Arthropoda" rank="species" species="exilis" status="n. sp.">
|
||
<emphasis box="[1122,1281,901,927]" italics="true" pageId="5" pageNumber="368">Athanas exilis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis bold="true" box="[1290,1355,901,927]" pageId="5" pageNumber="368">
|
||
<taxonomicNameLabel box="[1290,1355,901,927]" pageId="5" pageNumber="368" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
differs from
|
||
<taxonomicName authorityName="Anker & Ahyong" authorityYear="2007" box="[213,364,937,963]" class="Malacostraca" family="Alpheidae" genus="Athanas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="368" phylum="Arthropoda" rank="species" species="sydnyensis">
|
||
<emphasis box="[213,364,937,963]" italics="true" pageId="5" pageNumber="368">A. sydnyensis</emphasis>
|
||
</taxonomicName>
|
||
in the following particulars: the dorsal surface of the rostrum is devoid of a middorsal carina in
|
||
<taxonomicName authority="Komai & Henmi, 2023" authorityName="Komai & Henmi" authorityYear="2023" box="[151,241,972,999]" class="Malacostraca" family="Alpheidae" genus="Athanas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="368" phylum="Arthropoda" rank="species" species="exilis" status="n. sp.">
|
||
<emphasis box="[151,241,972,999]" italics="true" pageId="5" pageNumber="368">A. exilis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis bold="true" box="[248,310,973,999]" pageId="5" pageNumber="368">
|
||
<taxonomicNameLabel box="[248,310,973,999]" pageId="5" pageNumber="368" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, which is discernible in
|
||
<taxonomicName authorityName="Anker & Ahyong" authorityYear="2007" box="[583,734,972,999]" class="Malacostraca" family="Alpheidae" genus="Athanas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="368" phylum="Arthropoda" rank="species" species="sydnyensis">
|
||
<emphasis box="[583,734,972,999]" italics="true" pageId="5" pageNumber="368">A. sydnyensis</emphasis>
|
||
</taxonomicName>
|
||
(cf.
|
||
<figureCitation box="[785,869,972,999]" captionStart="FIGURE 2" captionStartId="4.[151,250,1876,1900]" captionTargetBox="[211,1373,191,1840]" captionTargetId="figure-21@4.[201,1386,181,1852]" captionTargetPageId="4" captionText="FIGURE 2. Athanas exilis n. sp., holotype, ovigerous female (cl 4.8 mm), CBM-ZC 17112.A, carapaceand cephalic appendages, left lateral view; B, anterior part of carapace and cephalic appendages, left lateral view; C, same, dorsal view; D, pleon, telson and pleonal appendages, left lateral view; E, posterior part of pleomere 5 and pleomere 6, left lateral view; F, telson, dorsal view; G, left antennal scaphocerite, dorsal view (setae omitted); H, right uropod, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.7889974" httpUri="https://zenodo.org/record/7889974/files/figure.png" pageId="5" pageNumber="368">Fig. 2C</figureCitation>
|
||
versus
|
||
<bibRefCitation author="Anker, A. & Ahyong, S. T." box="[953,1213,972,999]" pageId="5" pageNumber="368" pagination="17 - 30" refId="ref5101" refString="Anker, A. & Ahyong, S. T. (2007) Description of two new species of the shrimp genus Athanas Leach, 1814 (Decapoda: Alpheidae) from Australia and West Africa, with remarks on A. amazone Holthuis, 1951. Zootaxa, 1563 (1), 17 - 30. https: // doi. org / 10.11646 / zootaxa. 1563.1.2" type="journal article" year="2007">Anker & Ahyong 2007</bibRefCitation>
|
||
: fig. 1B); the lateral margins of the telson are straight in
|
||
<taxonomicName authority="Komai & Henmi, 2023" authorityName="Komai & Henmi" authorityYear="2023" box="[562,654,1008,1035]" class="Malacostraca" family="Alpheidae" genus="Athanas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="368" phylum="Arthropoda" rank="species" species="exilis" status="n. sp.">
|
||
<emphasis box="[562,654,1008,1035]" italics="true" pageId="5" pageNumber="368">A. exilis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis bold="true" box="[662,726,1008,1034]" pageId="5" pageNumber="368">
|
||
<taxonomicNameLabel box="[662,726,1008,1034]" pageId="5" pageNumber="368" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, rather than gently convex in
|
||
<taxonomicName authorityName="Anker & Ahyong" authorityYear="2007" box="[1068,1220,1008,1035]" class="Malacostraca" family="Alpheidae" genus="Athanas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="368" phylum="Arthropoda" rank="species" species="sydnyensis">
|
||
<emphasis box="[1068,1220,1008,1035]" italics="true" pageId="5" pageNumber="368">A. sydnyensis</emphasis>
|
||
</taxonomicName>
|
||
(cf.
|
||
<figureCitation box="[1274,1356,1008,1034]" captionStart="FIGURE 2" captionStartId="4.[151,250,1876,1900]" captionTargetBox="[211,1373,191,1840]" captionTargetId="figure-21@4.[201,1386,181,1852]" captionTargetPageId="4" captionText="FIGURE 2. Athanas exilis n. sp., holotype, ovigerous female (cl 4.8 mm), CBM-ZC 17112.A, carapaceand cephalic appendages, left lateral view; B, anterior part of carapace and cephalic appendages, left lateral view; C, same, dorsal view; D, pleon, telson and pleonal appendages, left lateral view; E, posterior part of pleomere 5 and pleomere 6, left lateral view; F, telson, dorsal view; G, left antennal scaphocerite, dorsal view (setae omitted); H, right uropod, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.7889974" httpUri="https://zenodo.org/record/7889974/files/figure.png" pageId="5" pageNumber="368">Fig. 2F</figureCitation>
|
||
versus
|
||
<bibRefCitation author="Anker, A. & Ahyong, S. T." box="[151,414,1044,1070]" pageId="5" pageNumber="368" pagination="17 - 30" refId="ref5101" refString="Anker, A. & Ahyong, S. T. (2007) Description of two new species of the shrimp genus Athanas Leach, 1814 (Decapoda: Alpheidae) from Australia and West Africa, with remarks on A. amazone Holthuis, 1951. Zootaxa, 1563 (1), 17 - 30. https: // doi. org / 10.11646 / zootaxa. 1563.1.2" type="journal article" year="2007">Anker & Ahyong 2007</bibRefCitation>
|
||
: fig. 1J); the antennular stylocerite reaches the distal margin of the basal peduncular article in
|
||
<taxonomicName authority="Komai & Henmi, 2023" authorityName="Komai & Henmi" authorityYear="2023" box="[179,268,1080,1106]" class="Malacostraca" family="Alpheidae" genus="Athanas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="368" phylum="Arthropoda" rank="species" species="exilis" status="n. sp.">
|
||
<emphasis box="[179,268,1080,1106]" italics="true" pageId="5" pageNumber="368">A. exilis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis bold="true" box="[274,335,1080,1106]" pageId="5" pageNumber="368">
|
||
<taxonomicNameLabel box="[274,335,1080,1106]" pageId="5" pageNumber="368" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, while reaching the midlength of the peduncular article
|
||
<quantity box="[954,996,1080,1106]" metricMagnitude="-2" metricUnit="m" metricValue="5.08" pageId="5" pageNumber="368" unit="in" value="2.0">2 in</quantity>
|
||
<taxonomicName authorityName="Anker & Ahyong" authorityYear="2007" box="[1002,1152,1080,1106]" class="Malacostraca" family="Alpheidae" genus="Athanas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="368" phylum="Arthropoda" rank="species" species="sydnyensis">
|
||
<emphasis box="[1002,1152,1080,1106]" italics="true" pageId="5" pageNumber="368">A. sydnyensis</emphasis>
|
||
</taxonomicName>
|
||
(cf.
|
||
<figureCitation box="[1201,1285,1080,1106]" captionStart="FIGURE 2" captionStartId="4.[151,250,1876,1900]" captionTargetBox="[211,1373,191,1840]" captionTargetId="figure-21@4.[201,1386,181,1852]" captionTargetPageId="4" captionText="FIGURE 2. Athanas exilis n. sp., holotype, ovigerous female (cl 4.8 mm), CBM-ZC 17112.A, carapaceand cephalic appendages, left lateral view; B, anterior part of carapace and cephalic appendages, left lateral view; C, same, dorsal view; D, pleon, telson and pleonal appendages, left lateral view; E, posterior part of pleomere 5 and pleomere 6, left lateral view; F, telson, dorsal view; G, left antennal scaphocerite, dorsal view (setae omitted); H, right uropod, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.7889974" httpUri="https://zenodo.org/record/7889974/files/figure.png" pageId="5" pageNumber="368">Fig. 2C</figureCitation>
|
||
versus
|
||
<bibRefCitation author="Anker, A. & Ahyong, S. T." pageId="5" pageNumber="368" pagination="17 - 30" refId="ref5101" refString="Anker, A. & Ahyong, S. T. (2007) Description of two new species of the shrimp genus Athanas Leach, 1814 (Decapoda: Alpheidae) from Australia and West Africa, with remarks on A. amazone Holthuis, 1951. Zootaxa, 1563 (1), 17 - 30. https: // doi. org / 10.11646 / zootaxa. 1563.1.2" type="journal article" year="2007">Anker & Ahyong 2007</bibRefCitation>
|
||
: fig. 1A–C); the distal lamella of the antennal scaphocerite is more strongly produced in
|
||
<taxonomicName authority="Komai & Henmi, 2023" authorityName="Komai & Henmi" authorityYear="2023" box="[1318,1407,1115,1142]" class="Malacostraca" family="Alpheidae" genus="Athanas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="368" phylum="Arthropoda" rank="species" species="exilis" status="n. sp.">
|
||
<emphasis box="[1318,1407,1115,1142]" italics="true" pageId="5" pageNumber="368">A. exilis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis bold="true" pageId="5" pageNumber="368">
|
||
<taxonomicNameLabel pageId="5" pageNumber="368" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
than in
|
||
<taxonomicName authorityName="Anker & Ahyong" authorityYear="2007" box="[273,423,1151,1178]" class="Malacostraca" family="Alpheidae" genus="Athanas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="368" phylum="Arthropoda" rank="species" species="sydnyensis">
|
||
<emphasis box="[273,423,1151,1178]" italics="true" pageId="5" pageNumber="368">A. sydnyensis</emphasis>
|
||
</taxonomicName>
|
||
(cf.
|
||
<figureCitation box="[474,559,1151,1177]" captionStart="FIGURE 2" captionStartId="4.[151,250,1876,1900]" captionTargetBox="[211,1373,191,1840]" captionTargetId="figure-21@4.[201,1386,181,1852]" captionTargetPageId="4" captionText="FIGURE 2. Athanas exilis n. sp., holotype, ovigerous female (cl 4.8 mm), CBM-ZC 17112.A, carapaceand cephalic appendages, left lateral view; B, anterior part of carapace and cephalic appendages, left lateral view; C, same, dorsal view; D, pleon, telson and pleonal appendages, left lateral view; E, posterior part of pleomere 5 and pleomere 6, left lateral view; F, telson, dorsal view; G, left antennal scaphocerite, dorsal view (setae omitted); H, right uropod, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.7889974" httpUri="https://zenodo.org/record/7889974/files/figure.png" pageId="5" pageNumber="368">Fig. 2G</figureCitation>
|
||
versus
|
||
<bibRefCitation author="Anker, A. & Ahyong, S. T." box="[641,900,1151,1177]" pageId="5" pageNumber="368" pagination="17 - 30" refId="ref5101" refString="Anker, A. & Ahyong, S. T. (2007) Description of two new species of the shrimp genus Athanas Leach, 1814 (Decapoda: Alpheidae) from Australia and West Africa, with remarks on A. amazone Holthuis, 1951. Zootaxa, 1563 (1), 17 - 30. https: // doi. org / 10.11646 / zootaxa. 1563.1.2" type="journal article" year="2007">Anker & Ahyong 2007</bibRefCitation>
|
||
: fig. 1B, C); the pereopod 2 is much slenderer in
|
||
<taxonomicName authority="Komai & Henmi, 2023" authorityName="Komai & Henmi" authorityYear="2023" box="[151,241,1187,1213]" class="Malacostraca" family="Alpheidae" genus="Athanas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="368" phylum="Arthropoda" rank="species" species="exilis" status="n. sp.">
|
||
<emphasis box="[151,241,1187,1213]" italics="true" pageId="5" pageNumber="368">A. exilis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis bold="true" box="[247,310,1187,1213]" pageId="5" pageNumber="368">
|
||
<taxonomicNameLabel box="[247,310,1187,1213]" pageId="5" pageNumber="368" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
than in
|
||
<taxonomicName authorityName="Anker & Ahyong" authorityYear="2007" box="[399,550,1187,1213]" class="Malacostraca" family="Alpheidae" genus="Athanas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="368" phylum="Arthropoda" rank="species" species="sydnyensis">
|
||
<emphasis box="[399,550,1187,1213]" italics="true" pageId="5" pageNumber="368">A. sydnyensis</emphasis>
|
||
</taxonomicName>
|
||
(cf.
|
||
<figureCitation box="[601,681,1187,1213]" captionStart="FIGURE 3" captionStartId="6.[151,250,1866,1890]" captionTargetBox="[210,1376,193,1833]" captionTargetId="figure-21@6.[202,1386,181,1842]" captionTargetPageId="6" captionText="FIGURE 3. Athanas exilis n. sp., holotype, ovigerous female (cl 4.8 mm), CBM-ZC 17112. A, left maxilliped 3 and pereopod 1 (in process of regeneration), lateral view; B, coxae of chelipeds (pereopods 1) (basal part of right pereopod 2 also shown); C, basis of right cheliped, lateral view; D, right cheliped, lateral view; E, same, mesial view; F, left pereopod 2, lateral view; G–I, left pereopods 3–5, lateral view. Abbreviations: EXP = exopod; P1 = pereopod 1; P2 = pereopod 2" figureDoi="http://doi.org/10.5281/zenodo.7889976" httpUri="https://zenodo.org/record/7889976/files/figure.png" pageId="5" pageNumber="368">Fig. 3F</figureCitation>
|
||
versus
|
||
<bibRefCitation author="Anker, A. & Ahyong, S. T." box="[764,1024,1187,1213]" pageId="5" pageNumber="368" pagination="17 - 30" refId="ref5101" refString="Anker, A. & Ahyong, S. T. (2007) Description of two new species of the shrimp genus Athanas Leach, 1814 (Decapoda: Alpheidae) from Australia and West Africa, with remarks on A. amazone Holthuis, 1951. Zootaxa, 1563 (1), 17 - 30. https: // doi. org / 10.11646 / zootaxa. 1563.1.2" type="journal article" year="2007">Anker & Ahyong 2007</bibRefCitation>
|
||
: fig. 1H); the ventral armature on the pereopod 1 merus is less developed in
|
||
<taxonomicName authority="Komai & Henmi, 2023" authorityName="Komai & Henmi" authorityYear="2023" box="[585,675,1223,1249]" class="Malacostraca" family="Alpheidae" genus="Athanas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="368" phylum="Arthropoda" rank="species" species="exilis" status="n. sp.">
|
||
<emphasis box="[585,675,1223,1249]" italics="true" pageId="5" pageNumber="368">A. exilis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis bold="true" box="[683,745,1223,1249]" pageId="5" pageNumber="368">
|
||
<taxonomicNameLabel box="[683,745,1223,1249]" pageId="5" pageNumber="368" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
than in
|
||
<taxonomicName authorityName="Anker & Ahyong" authorityYear="2007" box="[837,988,1223,1249]" class="Malacostraca" family="Alpheidae" genus="Athanas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="368" phylum="Arthropoda" rank="species" species="sydnyensis">
|
||
<emphasis box="[837,988,1223,1249]" italics="true" pageId="5" pageNumber="368">A. sydnyensis</emphasis>
|
||
</taxonomicName>
|
||
(cf.
|
||
<figureCitation box="[1040,1157,1223,1249]" captionStart="FIGURE 3" captionStartId="6.[151,250,1866,1890]" captionTargetBox="[210,1376,193,1833]" captionTargetId="figure-21@6.[202,1386,181,1842]" captionTargetPageId="6" captionText="FIGURE 3. Athanas exilis n. sp., holotype, ovigerous female (cl 4.8 mm), CBM-ZC 17112. A, left maxilliped 3 and pereopod 1 (in process of regeneration), lateral view; B, coxae of chelipeds (pereopods 1) (basal part of right pereopod 2 also shown); C, basis of right cheliped, lateral view; D, right cheliped, lateral view; E, same, mesial view; F, left pereopod 2, lateral view; G–I, left pereopods 3–5, lateral view. Abbreviations: EXP = exopod; P1 = pereopod 1; P2 = pereopod 2" figureDoi="http://doi.org/10.5281/zenodo.7889976" httpUri="https://zenodo.org/record/7889976/files/figure.png" pageId="5" pageNumber="368">Fig. 3D, E</figureCitation>
|
||
versus
|
||
<bibRefCitation author="Anker, A. & Ahyong, S. T." pageId="5" pageNumber="368" pagination="17 - 30" refId="ref5101" refString="Anker, A. & Ahyong, S. T. (2007) Description of two new species of the shrimp genus Athanas Leach, 1814 (Decapoda: Alpheidae) from Australia and West Africa, with remarks on A. amazone Holthuis, 1951. Zootaxa, 1563 (1), 17 - 30. https: // doi. org / 10.11646 / zootaxa. 1563.1.2" type="journal article" year="2007">Anker & Ahyong 2007</bibRefCitation>
|
||
: figs. 1D–G, 2A, B). In addition, as mentioned above, the chelipeds are probably unequal and dissimilar in the new species, but they are subequal and similar in
|
||
<taxonomicName authorityName="Anker & Ahyong" authorityYear="2007" box="[706,857,1294,1320]" class="Malacostraca" family="Alpheidae" genus="Athanas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="368" phylum="Arthropoda" rank="species" species="sydnyensis">
|
||
<emphasis box="[706,857,1294,1320]" italics="true" pageId="5" pageNumber="368">A. sydnyensis</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<bibRefCitation author="Anker, A. & Ahyong, S. T." box="[871,1134,1294,1320]" pageId="5" pageNumber="368" pagination="17 - 30" refId="ref5101" refString="Anker, A. & Ahyong, S. T. (2007) Description of two new species of the shrimp genus Athanas Leach, 1814 (Decapoda: Alpheidae) from Australia and West Africa, with remarks on A. amazone Holthuis, 1951. Zootaxa, 1563 (1), 17 - 30. https: // doi. org / 10.11646 / zootaxa. 1563.1.2" type="journal article" year="2007">Anker & Ahyong 2007</bibRefCitation>
|
||
: figs. 1D–G, 2A, B).
|
||
</paragraph>
|
||
<paragraph blockId="5.[151,1437,150,2036]" pageId="5" pageNumber="368">
|
||
Besides the development of the infracorneal spine on the carapace,
|
||
<taxonomicName authorityName="Holthuis" authorityYear="1951" box="[953,1151,1330,1356]" class="Malacostraca" family="Alpheidae" genus="Athanas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="368" phylum="Arthropoda" rank="species" species="amazone">
|
||
<emphasis box="[953,1151,1330,1356]" italics="true" pageId="5" pageNumber="368">Athanas amazone</emphasis>
|
||
</taxonomicName>
|
||
is also similar to the new species. In
|
||
<taxonomicName authorityName="Holthuis" authorityYear="1951" box="[277,409,1366,1392]" class="Malacostraca" family="Alpheidae" genus="Athanas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="368" phylum="Arthropoda" rank="species" species="amazone">
|
||
<emphasis box="[277,409,1366,1392]" italics="true" pageId="5" pageNumber="368">A. amazone</emphasis>
|
||
</taxonomicName>
|
||
, the infracorneal spine is usually present, but occasionally absent (
|
||
<bibRefCitation author="Anker, A. & Ahyong, S. T." box="[1159,1425,1365,1392]" pageId="5" pageNumber="368" pagination="17 - 30" refId="ref5101" refString="Anker, A. & Ahyong, S. T. (2007) Description of two new species of the shrimp genus Athanas Leach, 1814 (Decapoda: Alpheidae) from Australia and West Africa, with remarks on A. amazone Holthuis, 1951. Zootaxa, 1563 (1), 17 - 30. https: // doi. org / 10.11646 / zootaxa. 1563.1.2" type="journal article" year="2007">Anker & Ahyong 2007</bibRefCitation>
|
||
).
|
||
<taxonomicName authorityName="Holthuis" authorityYear="1951" box="[151,350,1401,1427]" class="Malacostraca" family="Alpheidae" genus="Athanas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="368" phylum="Arthropoda" rank="species" species="amazone">
|
||
<emphasis box="[151,350,1401,1427]" italics="true" pageId="5" pageNumber="368">Athanas amazone</emphasis>
|
||
</taxonomicName>
|
||
has been reported from the eastern Atlantic, i.e.,
|
||
<collectingCountry box="[897,981,1401,1427]" name="Nigeria" pageId="5" pageNumber="368">Nigeria</collectingCountry>
|
||
(
|
||
<bibRefCitation author="Holthuis, L. B." box="[995,1159,1401,1428]" pageId="5" pageNumber="368" pagination="7 - 187" refId="ref6623" refString="Holthuis, L. B. (1951) The caridean Crustacea of tropical West Africa. Atlantide Report, 2, 7 - 187." type="journal article" year="1951">Holthuis 1951</bibRefCitation>
|
||
),
|
||
<collectingCountry box="[1178,1238,1402,1428]" name="Togo" pageId="5" pageNumber="368">Togo</collectingCountry>
|
||
,
|
||
<collectingCountry box="[1249,1324,1401,1427]" name="Democratic Republic of the Congo" pageId="5" pageNumber="368">Congo</collectingCountry>
|
||
(
|
||
<bibRefCitation author="Crosnier, A. & Forest, J." pageId="5" pageNumber="368" refId="ref6153" refString="Crosnier, A. & Forest, J. (1973) Les crevettes profondes de l'Atlantique Oriental Tropical. Faune Tropicale, 9. O. R. S. T. O. M, Paris, 409 pp." type="book" year="1973">Crosnier & Forest 1973</bibRefCitation>
|
||
),
|
||
<collectingCountry box="[340,470,1437,1463]" name="Cape Verde" pageId="5" pageNumber="368">Cape Verde</collectingCountry>
|
||
(Türkay 1982),
|
||
<collectingCountry box="[656,757,1438,1464]" name="Morocco" pageId="5" pageNumber="368">Morocco</collectingCountry>
|
||
(
|
||
<bibRefCitation author="Garcia Raso, J. E." box="[773,982,1437,1464]" pageId="5" pageNumber="368" pagination="730 - 752" refId="ref6403" refString="Garcia Raso, J. E. (1996) Crustacea Decapoda (excl. Sergestidae) from Ibero - Moroccan waters. Results of Balgim- 84 expedition. Bulletin of Marine Science, 58, 730 - 752." type="journal article" year="1996">García Raso 1996</bibRefCitation>
|
||
), and throughout the Mediterranean Sea (
|
||
<bibRefCitation author="Holthuis, L. B. & Gottlieb, E." box="[159,449,1473,1500]" pageId="5" pageNumber="368" pagination="1 - 126" refId="ref6649" refString="Holthuis, L. B. & Gottlieb, E. (1958) An annotated list of the decapod Crustacea of the Mediterranean coast of Israel, with an appendix listing the Decapoda of the eastern Mediterranean. Bulletin of the Research Council of Israel, 7 B, 1 - 126, pls. 1 - 3." type="journal article" year="1958">Holthuis & Gottlieb 1958</bibRefCitation>
|
||
;
|
||
<bibRefCitation author="Crosnier, A. & Forest, J." box="[460,727,1473,1499]" pageId="5" pageNumber="368" refId="ref6153" refString="Crosnier, A. & Forest, J. (1973) Les crevettes profondes de l'Atlantique Oriental Tropical. Faune Tropicale, 9. O. R. S. T. O. M, Paris, 409 pp." type="book" year="1973">Crosnier & Forest 1973</bibRefCitation>
|
||
;
|
||
<bibRefCitation author="Kocatas, A." box="[738,894,1473,1499]" pageId="5" pageNumber="368" pagination="161 - 162" refId="ref6834" refString="Kocatas, A. (1981) Liste preliminaires et repartition des crustaces decapodes des eaux turques. Rapports de la Commission Internationale de la Mer Mediterranee, 27, 161 - 162." type="journal article" year="1981">Kocataş 1981</bibRefCitation>
|
||
;
|
||
<bibRefCitation author="Koukouras, A. & Dounas, C. & Voultsiadou-Koukoura, E." box="[905,1156,1473,1499]" pageId="5" pageNumber="368" pagination="217 - 244" refId="ref6868" refString="Koukouras, A., Dounas, C., Tu ¨ rkay, M. & Voultsiadou-Koukoura, E. (1992) Decapod Crustacean fauna of the Aegean Sea: new information, checklist, affinities. Senckenbergiana Maritima, 22, 217 - 244." type="journal article" year="1992">
|
||
Koukouras
|
||
<emphasis box="[1035,1090,1473,1499]" italics="true" pageId="5" pageNumber="368">et al.</emphasis>
|
||
1992
|
||
</bibRefCitation>
|
||
;
|
||
<bibRefCitation author="Froglia, C & Argenti, L." box="[1167,1431,1473,1499]" pageId="5" pageNumber="368" pagination="201 - 205" refId="ref6296" refString="Froglia, C & Argenti, L. (1993) Athanas amazone Holthuis, 1951, first record for the Italian seas (Crustacea, Decapoda, Alpheidae). Bios (Macedonia, Greece). Scientific Annals of the School of Biology, 1, 201 - 205." type="journal article" year="1993">Froglia & Argenti 1993</bibRefCitation>
|
||
;
|
||
<bibRefCitation author="Garcia Raso, J. E." box="[151,352,1508,1535]" pageId="5" pageNumber="368" pagination="730 - 752" refId="ref6403" refString="Garcia Raso, J. E. (1996) Crustacea Decapoda (excl. Sergestidae) from Ibero - Moroccan waters. Results of Balgim- 84 expedition. Bulletin of Marine Science, 58, 730 - 752." type="journal article" year="1996">García Raso 1996</bibRefCitation>
|
||
;
|
||
<bibRefCitation author="Atkinson, R. J. A. & Froglia, C. & Arneri, E. & Antolini, B." box="[361,589,1508,1535]" pageId="5" pageNumber="368" pagination="337 - 359" refId="ref5530" refString="Atkinson, R. J. A., Froglia, C., Arneri, E. & Antolini, B. (1997) Observations on the burrows and burrowing behaviour of Squilla mantis (L.) (Crustacea: Stomatopoda). Publicazioni della Stazione Zoologica di Napoli: Marine Ecology, 18, 337 - 359. https: // doi. org / 10.1111 / j. 1439 - 0485.1997. tb 00446. x" type="journal article" year="1997">
|
||
Atkinson
|
||
<emphasis box="[470,525,1508,1534]" italics="true" pageId="5" pageNumber="368">et al.</emphasis>
|
||
1997
|
||
</bibRefCitation>
|
||
;
|
||
<bibRefCitation author="Froglia, C. & Atkinson, R. J. A." box="[600,880,1508,1535]" pageId="5" pageNumber="368" pagination="529 - 532" refId="ref6348" refString="Froglia, C. & Atkinson, R. J. A. (1998) Association between Athanas amazone (Decapoda: Alpheidae) and Squilla mantis (Stomatopoda: Squillidae). Journal of Crustacean Biology, 18, 529 - 532. https: // doi. org / 10.2307 / 1549417" type="journal article" year="1998">Froglia & Atkinson 1998</bibRefCitation>
|
||
; d’Udekem
|
||
<bibRefCitation author="Udekem d'Acoz, C." box="[1014,1161,1508,1535]" pageId="5" pageNumber="368" refId="ref7598" refString="d'. Udekem d'Acoz, C. (1999) Inventaire et distribution des crustaces decapodes de l'Atlantique nord - oriental, de la Mediterranee et des eaux continentales adjacentes au nord de 25 ° N. Institut d'Ecologie et de Gestion de la Biodiversite, S. P. N., Museum national d'Histoire naturelle, Paris, 383 pp." type="book" year="1999">d’Acoz 1999</bibRefCitation>
|
||
;
|
||
<bibRefCitation author="Anker, A. & Ahyong, S. T." box="[1170,1425,1508,1535]" pageId="5" pageNumber="368" pagination="17 - 30" refId="ref5101" refString="Anker, A. & Ahyong, S. T. (2007) Description of two new species of the shrimp genus Athanas Leach, 1814 (Decapoda: Alpheidae) from Australia and West Africa, with remarks on A. amazone Holthuis, 1951. Zootaxa, 1563 (1), 17 - 30. https: // doi. org / 10.11646 / zootaxa. 1563.1.2" type="journal article" year="2007">Anker &Ahyong 2007</bibRefCitation>
|
||
).
|
||
<bibRefCitation author="Holthuis, L. B. & Gottlieb, E." box="[151,466,1544,1571]" pageId="5" pageNumber="368" pagination="1 - 126" refId="ref6649" refString="Holthuis, L. B. & Gottlieb, E. (1958) An annotated list of the decapod Crustacea of the Mediterranean coast of Israel, with an appendix listing the Decapoda of the eastern Mediterranean. Bulletin of the Research Council of Israel, 7 B, 1 - 126, pls. 1 - 3." type="journal article" year="1958">Holthuis & Gottlieb (1958)</bibRefCitation>
|
||
,
|
||
<bibRefCitation author="Crosnier, A. & Forest, J." box="[480,771,1544,1571]" pageId="5" pageNumber="368" refId="ref6153" refString="Crosnier, A. & Forest, J. (1973) Les crevettes profondes de l'Atlantique Oriental Tropical. Faune Tropicale, 9. O. R. S. T. O. M, Paris, 409 pp." type="book" year="1973">Crosnier & Forest (1973)</bibRefCitation>
|
||
and
|
||
<bibRefCitation author="Anker, A. & Ahyong, S. T." box="[829,1113,1544,1570]" pageId="5" pageNumber="368" pagination="17 - 30" refId="ref5101" refString="Anker, A. & Ahyong, S. T. (2007) Description of two new species of the shrimp genus Athanas Leach, 1814 (Decapoda: Alpheidae) from Australia and West Africa, with remarks on A. amazone Holthuis, 1951. Zootaxa, 1563 (1), 17 - 30. https: // doi. org / 10.11646 / zootaxa. 1563.1.2" type="journal article" year="2007">Anker & Ahyong (2007)</bibRefCitation>
|
||
noted some variation in the relative length of the stylocerite (compared to the distal margin of the first segment of the antennular peduncle); the length of the carpocerite (compared to the scaphocerite); and the shape of the chelipeds.
|
||
<bibRefCitation author="Anker, A. & Ahyong, S. T." box="[1157,1436,1616,1642]" pageId="5" pageNumber="368" pagination="17 - 30" refId="ref5101" refString="Anker, A. & Ahyong, S. T. (2007) Description of two new species of the shrimp genus Athanas Leach, 1814 (Decapoda: Alpheidae) from Australia and West Africa, with remarks on A. amazone Holthuis, 1951. Zootaxa, 1563 (1), 17 - 30. https: // doi. org / 10.11646 / zootaxa. 1563.1.2" type="journal article" year="2007">Anker & Ahyong (2007)</bibRefCitation>
|
||
stated: “It shows that some features, e.g., the length of the carpocerite, the shape of the merus and the chela of the major cheliped, and the slenderness of the minor cheliped, may indeed be sexually dimorphic and/or age-dependent. However, more material with information on colour patterns and ecology is needed to conclude about the taxonomic status of
|
||
<taxonomicName authorityName="Holthuis" authorityYear="1951" box="[254,386,1759,1785]" class="Malacostraca" family="Alpheidae" genus="Athanas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="368" phylum="Arthropoda" rank="species" species="amazone">
|
||
<emphasis box="[254,386,1759,1785]" italics="true" pageId="5" pageNumber="368">A. amazone</emphasis>
|
||
</taxonomicName>
|
||
.” Considering the possible sexual dimorphism and the possibility that more than one species might be confounded under
|
||
<taxonomicName authorityName="Holthuis" authorityYear="1951" box="[473,605,1795,1821]" class="Malacostraca" family="Alpheidae" genus="Athanas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="368" phylum="Arthropoda" rank="species" species="amazone">
|
||
<emphasis box="[473,605,1795,1821]" italics="true" pageId="5" pageNumber="368">A. amazone</emphasis>
|
||
</taxonomicName>
|
||
, the following comparison is limited to the accounts mentioning females (
|
||
<bibRefCitation author="Holthuis, L. B." box="[159,322,1830,1857]" pageId="5" pageNumber="368" pagination="7 - 187" refId="ref6623" refString="Holthuis, L. B. (1951) The caridean Crustacea of tropical West Africa. Atlantide Report, 2, 7 - 187." type="journal article" year="1951">Holthuis 1951</bibRefCitation>
|
||
;
|
||
<bibRefCitation author="Anker, A. & Ahyong, S. T." box="[332,592,1830,1856]" pageId="5" pageNumber="368" pagination="17 - 30" refId="ref5101" refString="Anker, A. & Ahyong, S. T. (2007) Description of two new species of the shrimp genus Athanas Leach, 1814 (Decapoda: Alpheidae) from Australia and West Africa, with remarks on A. amazone Holthuis, 1951. Zootaxa, 1563 (1), 17 - 30. https: // doi. org / 10.11646 / zootaxa. 1563.1.2" type="journal article" year="2007">Anker & Ahyong 2007</bibRefCitation>
|
||
).
|
||
<taxonomicName authority="Komai & Henmi, 2023" authorityName="Komai & Henmi" authorityYear="2023" box="[610,767,1830,1856]" class="Malacostraca" family="Alpheidae" genus="Athanas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="368" phylum="Arthropoda" rank="species" species="exilis" status="n. sp.">
|
||
<emphasis box="[610,767,1830,1856]" italics="true" pageId="5" pageNumber="368">Athanas exilis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis bold="true" box="[774,836,1830,1856]" pageId="5" pageNumber="368">
|
||
<taxonomicNameLabel box="[774,836,1830,1856]" pageId="5" pageNumber="368" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
differs from
|
||
<taxonomicName authorityName="Holthuis" authorityYear="1951" box="[982,1113,1830,1856]" class="Malacostraca" family="Alpheidae" genus="Athanas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="368" phylum="Arthropoda" rank="species" species="amazone">
|
||
<emphasis box="[982,1113,1830,1856]" italics="true" pageId="5" pageNumber="368">A. amazone</emphasis>
|
||
</taxonomicName>
|
||
as follows: the distal lamella of the antennal scaphocerite is more strongly produced in
|
||
<taxonomicName authority="Komai & Henmi, 2023" authorityName="Komai & Henmi" authorityYear="2023" box="[835,929,1866,1892]" class="Malacostraca" family="Alpheidae" genus="Athanas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="368" phylum="Arthropoda" rank="species" species="exilis" status="n. sp.">
|
||
<emphasis box="[835,929,1866,1892]" italics="true" pageId="5" pageNumber="368">A. exilis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis bold="true" box="[940,1007,1866,1892]" pageId="5" pageNumber="368">
|
||
<taxonomicNameLabel box="[940,1007,1866,1892]" pageId="5" pageNumber="368" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
than in
|
||
<taxonomicName authorityName="Holthuis" authorityYear="1951" box="[1110,1245,1866,1892]" class="Malacostraca" family="Alpheidae" genus="Athanas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="368" phylum="Arthropoda" rank="species" species="amazone">
|
||
<emphasis box="[1110,1245,1866,1892]" italics="true" pageId="5" pageNumber="368">A. amazone</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation box="[1263,1354,1866,1892]" captionStart="FIGURE 2" captionStartId="4.[151,250,1876,1900]" captionTargetBox="[211,1373,191,1840]" captionTargetId="figure-21@4.[201,1386,181,1852]" captionTargetPageId="4" captionText="FIGURE 2. Athanas exilis n. sp., holotype, ovigerous female (cl 4.8 mm), CBM-ZC 17112.A, carapaceand cephalic appendages, left lateral view; B, anterior part of carapace and cephalic appendages, left lateral view; C, same, dorsal view; D, pleon, telson and pleonal appendages, left lateral view; E, posterior part of pleomere 5 and pleomere 6, left lateral view; F, telson, dorsal view; G, left antennal scaphocerite, dorsal view (setae omitted); H, right uropod, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.7889974" httpUri="https://zenodo.org/record/7889974/files/figure.png" pageId="5" pageNumber="368">Fig. 2G</figureCitation>
|
||
versus
|
||
<bibRefCitation author="Holthuis, L. B." box="[151,315,1901,1928]" pageId="5" pageNumber="368" pagination="7 - 187" refId="ref6623" refString="Holthuis, L. B. (1951) The caridean Crustacea of tropical West Africa. Atlantide Report, 2, 7 - 187." type="journal article" year="1951">Holthuis 1951</bibRefCitation>
|
||
: fig. 23c); the major cheliped merus is less expanded in
|
||
<taxonomicName authority="Komai & Henmi, 2023" authorityName="Komai & Henmi" authorityYear="2023" box="[945,1035,1901,1928]" class="Malacostraca" family="Alpheidae" genus="Athanas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="368" phylum="Arthropoda" rank="species" species="exilis" status="n. sp.">
|
||
<emphasis box="[945,1035,1901,1928]" italics="true" pageId="5" pageNumber="368">A. exilis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis bold="true" box="[1042,1105,1902,1928]" pageId="5" pageNumber="368">
|
||
<taxonomicNameLabel box="[1042,1105,1902,1928]" pageId="5" pageNumber="368" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
than in
|
||
<taxonomicName authorityName="Holthuis" authorityYear="1951" box="[1196,1328,1902,1928]" class="Malacostraca" family="Alpheidae" genus="Athanas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="368" phylum="Arthropoda" rank="species" species="amazone">
|
||
<emphasis box="[1196,1328,1902,1928]" italics="true" pageId="5" pageNumber="368">A. amazone</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation captionStart="FIGURE 3" captionStartId="6.[151,250,1866,1890]" captionTargetBox="[210,1376,193,1833]" captionTargetId="figure-21@6.[202,1386,181,1842]" captionTargetPageId="6" captionText="FIGURE 3. Athanas exilis n. sp., holotype, ovigerous female (cl 4.8 mm), CBM-ZC 17112. A, left maxilliped 3 and pereopod 1 (in process of regeneration), lateral view; B, coxae of chelipeds (pereopods 1) (basal part of right pereopod 2 also shown); C, basis of right cheliped, lateral view; D, right cheliped, lateral view; E, same, mesial view; F, left pereopod 2, lateral view; G–I, left pereopods 3–5, lateral view. Abbreviations: EXP = exopod; P1 = pereopod 1; P2 = pereopod 2" figureDoi="http://doi.org/10.5281/zenodo.7889976" httpUri="https://zenodo.org/record/7889976/files/figure.png" pageId="5" pageNumber="368">Fig. 3D, E</figureCitation>
|
||
versus
|
||
<bibRefCitation author="Holthuis, L. B." box="[253,416,1937,1964]" pageId="5" pageNumber="368" pagination="7 - 187" refId="ref6623" refString="Holthuis, L. B. (1951) The caridean Crustacea of tropical West Africa. Atlantide Report, 2, 7 - 187." type="journal article" year="1951">Holthuis 1951</bibRefCitation>
|
||
: fig. 23e); the pereopod 2 is much more slenderer in
|
||
<taxonomicName authority="Komai & Henmi, 2023" authorityName="Komai & Henmi" authorityYear="2023" box="[1001,1091,1937,1964]" class="Malacostraca" family="Alpheidae" genus="Athanas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="368" phylum="Arthropoda" rank="species" species="exilis" status="n. sp.">
|
||
<emphasis box="[1001,1091,1937,1964]" italics="true" pageId="5" pageNumber="368">A. exilis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[1097,1157,1938,1964]" pageId="5" pageNumber="368" rank="species">n. sp.</taxonomicNameLabel>
|
||
than in
|
||
<taxonomicName authorityName="Holthuis" authorityYear="1951" box="[1246,1377,1938,1964]" class="Malacostraca" family="Alpheidae" genus="Athanas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="368" phylum="Arthropoda" rank="species" species="amazone">
|
||
<emphasis box="[1246,1377,1938,1964]" italics="true" pageId="5" pageNumber="368">A. amazone</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation captionStart="FIGURE 3" captionStartId="6.[151,250,1866,1890]" captionTargetBox="[210,1376,193,1833]" captionTargetId="figure-21@6.[202,1386,181,1842]" captionTargetPageId="6" captionText="FIGURE 3. Athanas exilis n. sp., holotype, ovigerous female (cl 4.8 mm), CBM-ZC 17112. A, left maxilliped 3 and pereopod 1 (in process of regeneration), lateral view; B, coxae of chelipeds (pereopods 1) (basal part of right pereopod 2 also shown); C, basis of right cheliped, lateral view; D, right cheliped, lateral view; E, same, mesial view; F, left pereopod 2, lateral view; G–I, left pereopods 3–5, lateral view. Abbreviations: EXP = exopod; P1 = pereopod 1; P2 = pereopod 2" figureDoi="http://doi.org/10.5281/zenodo.7889976" httpUri="https://zenodo.org/record/7889976/files/figure.png" pageId="5" pageNumber="368">Fig. 3F</figureCitation>
|
||
versus
|
||
<bibRefCitation author="Holthuis, L. B." box="[265,428,1973,2000]" pageId="5" pageNumber="368" pagination="7 - 187" refId="ref6623" refString="Holthuis, L. B. (1951) The caridean Crustacea of tropical West Africa. Atlantide Report, 2, 7 - 187." type="journal article" year="1951">Holthuis 1951</bibRefCitation>
|
||
: fig.
|
||
<quantity box="[483,529,1973,1999]" metricMagnitude="-2" metricUnit="kg" metricValue="2.3" pageId="5" pageNumber="368" unit="g" value="23.0">23g</quantity>
|
||
); the major chela is longer than the carpus in
|
||
<taxonomicName authority="Komai & Henmi, 2023" authorityName="Komai & Henmi" authorityYear="2023" box="[1033,1123,1973,2000]" class="Malacostraca" family="Alpheidae" genus="Athanas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="368" phylum="Arthropoda" rank="species" species="exilis" status="n. sp.">
|
||
<emphasis box="[1033,1123,1973,2000]" italics="true" pageId="5" pageNumber="368">A. exilis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis bold="true" box="[1129,1192,1973,1999]" pageId="5" pageNumber="368">
|
||
<taxonomicNameLabel box="[1129,1192,1973,1999]" pageId="5" pageNumber="368" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, whereas shorter than the carpus in
|
||
<taxonomicName authorityName="Holthuis" authorityYear="1951" box="[301,432,2010,2036]" class="Malacostraca" family="Alpheidae" genus="Athanas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="368" phylum="Arthropoda" rank="species" species="amazone">
|
||
<emphasis box="[301,432,2010,2036]" italics="true" pageId="5" pageNumber="368">A. amazone</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation box="[447,565,2009,2036]" captionStart="FIGURE 3" captionStartId="6.[151,250,1866,1890]" captionTargetBox="[210,1376,193,1833]" captionTargetId="figure-21@6.[202,1386,181,1842]" captionTargetPageId="6" captionText="FIGURE 3. Athanas exilis n. sp., holotype, ovigerous female (cl 4.8 mm), CBM-ZC 17112. A, left maxilliped 3 and pereopod 1 (in process of regeneration), lateral view; B, coxae of chelipeds (pereopods 1) (basal part of right pereopod 2 also shown); C, basis of right cheliped, lateral view; D, right cheliped, lateral view; E, same, mesial view; F, left pereopod 2, lateral view; G–I, left pereopods 3–5, lateral view. Abbreviations: EXP = exopod; P1 = pereopod 1; P2 = pereopod 2" figureDoi="http://doi.org/10.5281/zenodo.7889976" httpUri="https://zenodo.org/record/7889976/files/figure.png" pageId="5" pageNumber="368">Fig. 3D, E</figureCitation>
|
||
versus
|
||
<bibRefCitation author="Holthuis, L. B." box="[650,856,2009,2036]" pageId="5" pageNumber="368" pagination="7 - 187" refId="ref6623" refString="Holthuis, L. B. (1951) The caridean Crustacea of tropical West Africa. Atlantide Report, 2, 7 - 187." type="journal article" year="1951">Holthuis 1951: 23</bibRefCitation>
|
||
e).
|
||
</paragraph>
|
||
<caption ID-DOI="http://doi.org/10.5281/zenodo.7889976" ID-Zenodo-Dep="7889976" httpUri="https://zenodo.org/record/7889976/files/figure.png" pageId="6" pageNumber="369" startId="6.[151,250,1866,1890]" targetBox="[210,1376,193,1833]" targetPageId="6" targetType="figure">
|
||
<paragraph blockId="6.[151,1436,1866,1998]" pageId="6" pageNumber="369">
|
||
<emphasis bold="true" box="[151,275,1866,1890]" pageId="6" pageNumber="369">FIGURE 3.</emphasis>
|
||
<taxonomicName authority="Komai & Henmi, 2023" authorityName="Komai & Henmi" authorityYear="2023" box="[282,424,1866,1890]" class="Malacostraca" family="Alpheidae" genus="Athanas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="369" phylum="Arthropoda" rank="species" species="exilis" status="n. sp.">
|
||
<emphasis box="[282,424,1866,1890]" italics="true" pageId="6" pageNumber="369">Athanas exilis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis bold="true" box="[431,488,1866,1890]" pageId="6" pageNumber="369">
|
||
<taxonomicNameLabel box="[431,488,1866,1890]" pageId="6" pageNumber="369" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, holotype, ovigerous female (cl 4.8 mm), CBM-ZC 17112. A, left maxilliped 3 and pereopod 1 (in process of regeneration), lateral view; B, coxae of chelipeds (pereopods 1) (basal part of right pereopod 2 also shown); C, basis of right cheliped, lateral view; D, right cheliped, lateral view; E, same, mesial view; F, left pereopod 2, lateral view; G–I, left pereopods 3–5, lateral view. Abbreviations: EXP = exopod; P1 = pereopod 1; P2 = pereopod 2
|
||
</paragraph>
|
||
</caption>
|
||
<caption ID-DOI="http://doi.org/10.5281/zenodo.7889978" ID-Zenodo-Dep="7889978" httpUri="https://zenodo.org/record/7889978/files/figure.png" pageId="7" pageNumber="370" startId="7.[151,250,1100,1124]" targetBox="[156,1432,189,1072]" targetPageId="7" targetType="figure">
|
||
<paragraph blockId="7.[151,1436,1100,1196]" pageId="7" pageNumber="370">
|
||
<emphasis bold="true" box="[151,274,1100,1124]" pageId="7" pageNumber="370">FIGURE 4.</emphasis>
|
||
<taxonomicName authority="Komai & Henmi, 2023" authorityName="Komai & Henmi" authorityYear="2023" box="[280,422,1100,1124]" class="Malacostraca" family="Alpheidae" genus="Athanas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="370" phylum="Arthropoda" rank="species" species="exilis" status="n. sp.">
|
||
<emphasis box="[280,422,1100,1124]" italics="true" pageId="7" pageNumber="370">Athanas exilis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis bold="true" box="[429,485,1100,1124]" pageId="7" pageNumber="370">
|
||
<taxonomicNameLabel box="[429,485,1100,1124]" pageId="7" pageNumber="370" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, holotype, ovigerous female (cl 4.8 mm), CBM-ZC 17112. A, close up of ventrolateral margin of ischium of right cheliped; B, carpus of right cheliped, extensor view; C, merus and carpus of right cheliped, ventrolateral view; D, chela of right cheliped, extensor view; E, same, ventrolateral view; F–H, dactyli of pereopods 3–5, lateral view.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph blockId="7.[151,1437,1242,1881]" pageId="7" pageNumber="370">
|
||
A partial fragment of the 16S rRNA gene (542 bp) was sequenced for the
|
||
<typeStatus box="[1029,1127,1242,1268]" pageId="7" pageNumber="370">holotype</typeStatus>
|
||
of the present new species. Sequences of the16S rRNA gene are available for eight species (nine sequences) of
|
||
<taxonomicName authorityName="Leach" authorityYear="1814" box="[1085,1177,1278,1304]" class="Malacostraca" family="Alpheidae" genus="Athanas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="370" phylum="Arthropoda" rank="genus">
|
||
<emphasis box="[1085,1177,1278,1304]" italics="true" pageId="7" pageNumber="370">Athanas</emphasis>
|
||
</taxonomicName>
|
||
in the GenBank (
|
||
<tableCitation captionStart="TABLE 1" captionStartId="1.[151,239,945,969]" captionTargetPageId="1" captionText="TABLE 1. Sequences of the mitochondrial 16S rRNA gene of species of Athanas Leach, 1814 used for genetic analysis." httpUri="http://table.plazi.org/id/DF78661ACC6A9D64EEA7FC5BFA2436BB" pageId="7" pageNumber="370" tableUuid="DF78661ACC6A9D64EEA7FC5BFA2436BB">Table 1</tableCitation>
|
||
), although no sequence data is available for
|
||
<taxonomicName authorityName="Anker & Ahyong" authorityYear="2007" box="[678,831,1314,1341]" class="Malacostraca" family="Alpheidae" genus="Athanas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="370" phylum="Arthropoda" rank="species" species="sydnyensis">
|
||
<emphasis box="[678,831,1314,1341]" italics="true" pageId="7" pageNumber="370">A. sydnyensis</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName authorityName="Holthuis" authorityYear="1951" box="[889,1022,1315,1341]" class="Malacostraca" family="Alpheidae" genus="Athanas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="370" phylum="Arthropoda" rank="species" species="amazone">
|
||
<emphasis box="[889,1022,1315,1341]" italics="true" pageId="7" pageNumber="370">A. amazone</emphasis>
|
||
</taxonomicName>
|
||
, mentioned above. The interspecific K2P genetic divergence among the nine
|
||
<taxonomicName box="[601,780,1350,1376]" pageId="7" pageNumber="370">
|
||
<emphasis box="[601,693,1350,1376]" italics="true" pageId="7" pageNumber="370">Athanas</emphasis>
|
||
species
|
||
</taxonomicName>
|
||
is high, ranging from 14.0 to 40.4% (
|
||
<tableCitation box="[1199,1286,1350,1377]" captionStart="TABLE 2" captionStartId="8.[151,239,151,175]" captionTargetPageId="8" captionText="TABLE 2. Genetic divergences of the mitochondrial 16S rRNA gene among nine species of Athanas (10 sequences) based on Kimura 2-parameter model. Standard error estimate(s) are shown above the diagonal. The rate variation among sites was modeled with a gamma distribution (shape parameter = 1). There were a total of 531 bp in the final dataset." httpUri="http://table.plazi.org/id/DF78661ACC639D6DEEA7FF7DFACE3585" pageId="7" pageNumber="370" tableUuid="DF78661ACC639D6DEEA7FF7DFACE3585">Table 2</tableCitation>
|
||
). The genetic divergence between
|
||
<taxonomicName authority="Komai & Henmi, 2023" authorityName="Komai & Henmi" authorityYear="2023" box="[381,539,1386,1412]" class="Malacostraca" family="Alpheidae" genus="Athanas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="370" phylum="Arthropoda" rank="species" species="exilis" status="n. sp.">
|
||
<emphasis box="[381,539,1386,1412]" italics="true" pageId="7" pageNumber="370">Athanas exilis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis bold="true" box="[546,609,1387,1413]" pageId="7" pageNumber="370">
|
||
<taxonomicNameLabel box="[546,609,1387,1413]" pageId="7" pageNumber="370" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
and the other eight species ranges from 21.1 to 33.4%. It should be noted that the identity of the voucher specimen of the sequence KU312972 with
|
||
<taxonomicName authority="Leach, 1813" authorityName="Leach" authorityYear="1813" box="[978,1256,1422,1449]" class="Malacostraca" family="Alpheidae" genus="Athanas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="370" phylum="Arthropoda" rank="species" species="nitescens">
|
||
<emphasis box="[978,1112,1423,1449]" italics="true" pageId="7" pageNumber="370">A. nitescens</emphasis>
|
||
<bibRefCitation author="Leach, W. E." box="[1118,1256,1422,1449]" pageId="7" pageNumber="370" pagination="383 - 437" refId="ref6948" refString="Leach, W. E. (1813 - 1814) Crustaceology. In: Brewster, D., The Edinburgh Encyclopaedia. A. Balfour, Edinburgh, pp. 383 - 437." type="book chapter" year="1813">Leach, 1813</bibRefCitation>
|
||
</taxonomicName>
|
||
is questionable, because the sequence closely matches MZ661664 referred to
|
||
<taxonomicName authority="Coutiere, 1911" authorityName="Coutiere" authorityYear="1911" box="[844,1152,1458,1485]" class="Malacostraca" family="Alpheidae" genus="Athanas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="370" phylum="Arthropoda" rank="species" species="glimardii">
|
||
<emphasis box="[844,979,1458,1485]" italics="true" pageId="7" pageNumber="370">A. glimardii</emphasis>
|
||
<bibRefCitation author="Coutiere, H." box="[986,1152,1458,1485]" pageId="7" pageNumber="370" pagination="1 - 7" refId="ref5980" refString="Coutiere, H. (1911) Sur les Alpheidae du genre Athanas Leach, provenant des collections de S. A. S. le Prince de Monaco. Bulletin de l'Institut Oceanographique, 197, 1 - 7." type="journal article" year="1911">Coutiére, 1911</bibRefCitation>
|
||
</taxonomicName>
|
||
(K2P genetic divergence 0.8%), rather than to MZ661665 referred to
|
||
<taxonomicName authorityName="Leach" authorityYear="1813" box="[646,780,1495,1521]" class="Malacostraca" family="Alpheidae" genus="Athanas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="370" phylum="Arthropoda" rank="species" species="nitescens">
|
||
<emphasis box="[646,780,1495,1521]" italics="true" pageId="7" pageNumber="370">A. nitescens</emphasis>
|
||
</taxonomicName>
|
||
(K2P genetic divergence 14.8%).
|
||
</paragraph>
|
||
<paragraph blockId="7.[151,1437,1242,1881]" pageId="7" pageNumber="370">
|
||
Of the 13 species known from
|
||
<collectingCountry box="[544,611,1531,1557]" name="Japan" pageId="7" pageNumber="370">Japan</collectingCountry>
|
||
, 10 species are widely distributed in the tropical or subtropical Indo-West Pacific (
|
||
<bibRefCitation author="Miya, Y. & Miyake, S." box="[246,502,1566,1593]" pageId="7" pageNumber="370" pagination="129 - 162" refId="ref7153" refString="Miya, Y. & Miyake, S. (1968) Revision of the genus Athanas of Japan and the Ryukyu Islands, with description of a new species (Crustacea, Decapoda, Alpheidae). Publications from the Amakusa Marine Biological Laboratory, Kyushu University, 1, 129 - 162." type="journal article" year="1968">Miya & Miyake 1968</bibRefCitation>
|
||
;
|
||
<bibRefCitation author="Banner, D. M. & Banner, A. H." box="[517,788,1567,1593]" pageId="7" pageNumber="370" pagination="291 - 382" refId="ref5652" refString="Banner, D. M. & Banner, A. H. (1973) The alpheid shrimp of Australia, part I: the lower genera. Records of the Australian Museum, 28, 291 - 382. https: // doi. org / 10.3853 / j. 0067 - 1975.28.1973.407" type="journal article" year="1973">Banner & Banner 1973</bibRefCitation>
|
||
;
|
||
<bibRefCitation author="Chace, F. A. Jr." box="[802,941,1566,1593]" pageId="7" pageNumber="370" pagination="1 - 99" refId="ref5842" refString="Chace, F. A. Jr. (1988) The caridean shrimps (Crustacea: Decapoda) of the Albatross Philippine Expedition, 1907 - 1910, part 5: Family Alpheidae. Smithsonian Contributions to Zoology, 466, i - v + 1 - 99. https: // doi. org / 10.5479 / si. 19436696.391.1" type="journal article" year="1988">Chace 1988</bibRefCitation>
|
||
;
|
||
<bibRefCitation author="Hayashi, K. - I." box="[955,1129,1566,1593]" pageId="7" pageNumber="370" pagination="2 - 6" refId="ref6490" refString="Hayashi, K. - I. (1995 a) Prawns, shrimps and lobsters from Japan (81), Family Alpheidae - Genus Athanas 2. Aquabiology, 96, 2 - 6. [in Japanese]" type="journal article" year="1995">Hayashi 1995a</bibRefCitation>
|
||
;
|
||
<bibRefCitation author="Sha, Z. & Wang, Y. & Cui, D." box="[1143,1323,1566,1593]" pageId="7" pageNumber="370" refId="ref7384" refString="Sha, Z., Wang, Y. & Cui, D. (2019) The Alpheidae from China Seas. Springer Nature Singapore Pte Ltd., Singapore, 323 pp. https: // doi. org / 10.1007 / 978 - 981 - 15 - 0648 - 2 _ 2" type="book" year="2019">
|
||
Sha
|
||
<emphasis box="[1194,1253,1566,1592]" italics="true" pageId="7" pageNumber="370">et al.</emphasis>
|
||
2019
|
||
</bibRefCitation>
|
||
), whereas three species, including the present new species, are so far restricted to temperate areas in East Asian waters:
|
||
<taxonomicName authorityName="Hayashi" authorityYear="2002" class="Malacostraca" family="Alpheidae" genus="Athanas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="370" phylum="Arthropoda" rank="species" species="squillophilus">
|
||
<emphasis italics="true" pageId="7" pageNumber="370">A. squillophilus</emphasis>
|
||
</taxonomicName>
|
||
was originally described from Ishikari Bay,
|
||
<collectingRegion box="[792,905,1638,1664]" country="Japan" name="Hokkaido" pageId="7" pageNumber="370">Hokkaido</collectingRegion>
|
||
,
|
||
<collectingCountry box="[917,981,1639,1665]" name="Japan" pageId="7" pageNumber="370">Japan</collectingCountry>
|
||
(
|
||
<bibRefCitation author="Hayashi, K. - I." box="[995,1155,1638,1665]" pageId="7" pageNumber="370" pagination="395 - 403" refId="ref6572" refString="Hayashi, K. - I. (2002) A new species of the genus Athanas (Decapoda, Caridea, Alpheidae) living in the burrows of a mantis shrimp. Crustaceana, 75, 395 - 403. https: // doi. org / 10.1163 / 156854002760095462" type="journal article" year="2002">Hayashi 2002</bibRefCitation>
|
||
), and later recorded from the Yellow Sea coast of northern
|
||
<collectingCountry box="[526,593,1674,1700]" name="China" pageId="7" pageNumber="370">China</collectingCountry>
|
||
(
|
||
<bibRefCitation author="Sha, Z. & Wang, Y. & Cui, D." box="[609,785,1674,1701]" pageId="7" pageNumber="370" refId="ref7384" refString="Sha, Z., Wang, Y. & Cui, D. (2019) The Alpheidae from China Seas. Springer Nature Singapore Pte Ltd., Singapore, 323 pp. https: // doi. org / 10.1007 / 978 - 981 - 15 - 0648 - 2 _ 2" type="book" year="2019">
|
||
Sha
|
||
<emphasis box="[660,717,1674,1700]" italics="true" pageId="7" pageNumber="370">et al.</emphasis>
|
||
2019
|
||
</bibRefCitation>
|
||
);
|
||
<taxonomicName authority="Marin, 2017" authorityName="Marin" authorityYear="2017" box="[805,1153,1674,1701]" class="Malacostraca" family="Alpheidae" genus="Athanas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="370" phylum="Arthropoda" rank="species" species="alpheusophilus">
|
||
<emphasis box="[805,1007,1674,1701]" italics="true" pageId="7" pageNumber="370">A. alpheusophilus</emphasis>
|
||
<bibRefCitation author="Marin, I. N." box="[1014,1153,1674,1701]" pageId="7" pageNumber="370" pagination="50 - 62" refId="ref7020" refString="Marin, I. N. (2017) Athanas alpheusophilus sp. nov. (Decapoda: Alpheidae) - a new Alpheus - associated shrimp from the Russian coast of the Sea of Japan. Zootaxa, 4324 (1), 50 - 62. https: // doi. org / 10.11646 / zootaxa. 4324.1.3" type="journal article" year="2017">Marin, 2017</bibRefCitation>
|
||
</taxonomicName>
|
||
was originally described from Posjeta Bay, Far Eastern
|
||
<collectingCountry box="[483,558,1710,1736]" name="Russia" pageId="7" pageNumber="370">Russia</collectingCountry>
|
||
(Sea of
|
||
<collectingCountry box="[645,714,1711,1737]" name="Japan" pageId="7" pageNumber="370">Japan</collectingCountry>
|
||
); and
|
||
<taxonomicName authority="Komai & Henmi, 2023" authorityName="Komai & Henmi" authorityYear="2023" box="[776,864,1710,1737]" class="Malacostraca" family="Alpheidae" genus="Athanas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="370" phylum="Arthropoda" rank="species" species="exilis" status="n. sp.">
|
||
<emphasis box="[776,864,1710,1737]" italics="true" pageId="7" pageNumber="370">A. exilis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis bold="true" box="[869,930,1711,1737]" pageId="7" pageNumber="370">
|
||
<taxonomicNameLabel box="[869,930,1711,1737]" pageId="7" pageNumber="370" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
is so far known only from Wakasa Bay,
|
||
<collectingCountry box="[1365,1432,1711,1737]" name="Japan" pageId="7" pageNumber="370">Japan</collectingCountry>
|
||
, although its real distribution remains clarified. The
|
||
<typeStatus box="[725,823,1746,1772]" pageId="7" pageNumber="370">holotype</typeStatus>
|
||
of
|
||
<taxonomicName authority="Komai & Henmi, 2023" authorityName="Komai & Henmi" authorityYear="2023" box="[859,949,1746,1773]" class="Malacostraca" family="Alpheidae" genus="Athanas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="370" phylum="Arthropoda" rank="species" species="exilis" status="n. sp.">
|
||
<emphasis box="[859,949,1746,1773]" italics="true" pageId="7" pageNumber="370">A. exilis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis bold="true" box="[956,1018,1747,1773]" pageId="7" pageNumber="370">
|
||
<taxonomicNameLabel box="[956,1018,1747,1773]" pageId="7" pageNumber="370" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
was collected from the sea bottom of the temperature 11.8°C (see above), suggesting that the new species is adapted to the temperate environment.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="7" pageNumber="370" type="etymology">
|
||
<paragraph blockId="7.[151,1437,1242,1881]" pageId="7" pageNumber="370">
|
||
<emphasis bold="true" box="[199,334,1819,1845]" pageId="7" pageNumber="370">Etymology.</emphasis>
|
||
From the Latin
|
||
<taxonomicName authorityName="Komai & Henmi" authorityYear="2023" box="[516,575,1818,1844]" class="Malacostraca" family="Alpheidae" genus="Athanas" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="370" phylum="Arthropoda" rank="species" species="exilis">
|
||
<emphasis box="[516,575,1818,1844]" italics="true" pageId="7" pageNumber="370">exilis</emphasis>
|
||
</taxonomicName>
|
||
(= slender) in reference to the slender, elongate dactyli of the pereopods
|
||
<quantity metricMagnitude="-1" metricUnit="m" metricValue="1.016" metricValueMax="1.27" metricValueMin="0.762" pageId="7" pageNumber="370" unit="in" value="4.0" valueMax="5.0" valueMin="3.0">3–5 in</quantity>
|
||
the new species.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |