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<document id="BD7FAA5A495857682340E9B025C3FBEB" ID-CLB-Dataset="99508" ID-DOI="10.5281/zenodo.6419781" ID-GBIF-Dataset="37ddff76-af79-4681-a698-fc8f6a58bc6c" ID-ISBN="978-84-16728-19-0" ID-Zenodo-Dep="6419781" IM.metadata_requiresApprovalFor="plazi" IM.taxonomicNames_requiresApprovalFor="plazi" checkinTime="1600874096722" checkinUser="plazi" docAuthor="Don E. Wilson &amp; Russell A. Mittermeier" docDate="2019" docId="03A637439152FFEEE4E4CC8F27C9A2B6" docLanguage="en" docName="hbmw_9_Mormoopidae_424.pdf.imf" docOrigin="Handbook of the Mammals of the World Volume 9 Bats, Barcelona: Lynx Edicions" docTitle="Pleronotus macleayii" docType="treatment" docVersion="14" lastPageNumber="439" masterDocId="FF9F4F3B9157FFEBE472C9232F51A072" masterDocTitle="Mormoopidae" masterLastPageNumber="443" masterPageNumber="424" pageNumber="439" updateTime="1698845891737" updateUser="ExternalLinkService">
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<mods:title id="FB1D629B649E4E12082E237D3E0B9E9D">Mormoopidae</mods:title>
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<mods:namePart id="8EDC283F069C5531EABEAD922322AC07">Don E. Wilson</mods:namePart>
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<mods:namePart id="A5874257626622CC465CFF1C4BE85F2A">Russell A. Mittermeier</mods:namePart>
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<mods:title id="DA8461D24303401E0EBE9F5CBA46FB58">Handbook of the Mammals of the World Volume 9 Bats</mods:title>
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<treatment id="03A637439152FFEEE4E4CC8F27C9A2B6" ID-DOI="http://doi.org/10.5281/zenodo.6606812" ID-GBIF-Taxon="197821935" ID-Zenodo-Dep="6606812" LSID="urn:lsid:plazi:treatment:03A637439152FFEEE4E4CC8F27C9A2B6" httpUri="http://treatment.plazi.org/id/03A637439152FFEEE4E4CC8F27C9A2B6" lastPageNumber="439" pageId="5" pageNumber="439">
<subSubSection id="C315D5DE9152FFEEE4E4CC8F2FE7A5A8" box="[150,182,1452,1498]" pageId="5" pageNumber="439" type="multiple">
<paragraph id="8BB086559152FFEEE4E4CC8F2FE7A5A8" blockId="5.[148,1246,1452,1538]" box="[150,182,1452,1498]" pageId="5" pageNumber="439">
<heading id="D0F831399152FFEEE4E4CC8F2FE7A5A8" box="[150,182,1452,1498]" pageId="5" pageNumber="439">
<figureCitation id="13349AD09152FFEEE4E4CC8F2FE7A5A8" box="[150,182,1452,1498]" captionStart="Plate 34: Mormoopidae" captionStartId="3.[148,178,3292,3317]" captionTargetBox="[14,2785,14,3657]" captionTargetPageId="2" captionText="1. Antllean Ghost-faced Bat (Mormoops blainuviller), 2. Peterss Ghostfaced Bat (Mormoops megalophylla), 3. Davys Naked-backed Bat (Pteronotus davyi), 4. Thomas's Naked-backed Bat (Pteronotus fulvus), 5. Big Naked-backed Bat (Pteronotus gymnonotus), 6. Macleays Mustached Bat (Pleronotus macleayii), 7. Sooty Mustached Bat (Pleronotus quadridens), 8. Wagner's Lesser Mustached Bat (Pteronotus personatus), 9. Dobsons Lesser Mustached Bat (Pteronotus psilotis), 10. Parnells Common Mustached Bat (Pteronotus parnellii), 11. Hispaniolan Common Mustached Bat (Preronotus pusillus), 12. Puerto Rican Common Mustached Bat (Pteronotus portoricensis), 13. Mexican Common Mustached Bat (Pteronotus mexicanus), 14. Mesoamerican Common Mustached Bat (Pteronotus mesoamericanus), 15. Allens Common Mustached Bat (Pleronotus fuscus), 16. Paraguana Common Mustached Bat (Pteronotus paraguanensis), 17. Womurs Common Mustached Bat (Pteronotus rubiginosus), 18. Amazonian Common Mustached Bat (Pteronotus alitonus)" figureDoi="http://doi.org/10.5281/zenodo.6419849" httpUri="https://zenodo.org/record/6419849/files/figure.png" pageId="5" pageNumber="439">6.</figureCitation>
</heading>
</paragraph>
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<subSubSection id="C315D5DE9152FFEEE4B4CC8F2D80A5A8" box="[198,721,1452,1498]" pageId="5" pageNumber="439" type="vernacular_names">
<paragraph id="8BB086559152FFEEE4B4CC8F2D80A5A8" blockId="5.[148,1246,1452,1538]" box="[198,721,1452,1498]" pageId="5" pageNumber="439">
<heading id="D0F831399152FFEEE4B4CC8F2D80A5A8" box="[198,721,1452,1498]" pageId="5" pageNumber="439">
<vernacularName id="050CF67B9152FFEEE4B4CC8F2D80A5A8" box="[198,721,1452,1498]" pageId="5" pageNumber="439">Macleays Mustached Bat</vernacularName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C315D5DE9152FFEEE761CC8F2BD0A5A8" box="[787,1153,1452,1498]" pageId="5" pageNumber="439" type="nomenclature">
<paragraph id="8BB086559152FFEEE761CC8F2BD0A5A8" blockId="5.[148,1246,1452,1538]" box="[787,1153,1452,1498]" pageId="5" pageNumber="439">
<heading id="D0F831399152FFEEE761CC8F2BD0A5A8" box="[787,1153,1452,1498]" pageId="5" pageNumber="439">
<taxonomicName id="4C0FFDD69152FFEEE761CC8F2BD0A5A8" authority="(J. E. Gray, 1839)" baseAuthorityName="J. E. Gray" baseAuthorityYear="1839" box="[787,1153,1452,1498]" class="Gastropoda" family="Muricidae" genus="Pleronotus" kingdom="Animalia" order="Neogastropoda" pageId="5" pageNumber="439" phylum="Mollusca" rank="species" species="macleayii">
<emphasis id="B97B5A479152FFEEE761CC8F2BD0A5A8" box="[787,1153,1452,1498]" italics="true" pageId="5" pageNumber="439">Pleronotus macleayii</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C315D5DE9152FFEEE4E7CCCF2B8FA673" box="[149,1246,1516,1537]" pageId="5" pageNumber="439" type="vernacular_names">
<paragraph id="8BB086559152FFEEE4E7CCCF2B8FA673" blockId="5.[148,1246,1452,1538]" box="[149,1246,1516,1537]" pageId="5" pageNumber="439">
<heading id="D0F831399152FFEEE4E7CCCF2B8FA673" box="[149,1246,1516,1537]" pageId="5" pageNumber="439">
<emphasis id="B97B5A479152FFEEE4E7CCCF2FB1A673" bold="true" box="[149,224,1516,1537]" pageId="5" pageNumber="439">French:</emphasis>
<vernacularName id="050CF67B9152FFEEE498CCCF2E97A673" box="[234,454,1516,1537]" pageId="5" pageNumber="439">Ptéronote de Macleay</vernacularName>
/
<emphasis id="B97B5A479152FFEEE5A9CCCF2D64A673" bold="true" box="[475,565,1516,1537]" pageId="5" pageNumber="439">German:</emphasis>
<vernacularName id="050CF67B9152FFEEE64DCCCF2C2DA673" box="[575,892,1516,1537]" pageId="5" pageNumber="439">Macleay-Schnurrbartfledermaus</vernacularName>
/
<emphasis id="B97B5A479152FFEEE7E3CCCF2CBDA673" bold="true" box="[913,1004,1516,1537]" pageId="5" pageNumber="439">Spanish:</emphasis>
<vernacularName id="050CF67B9152FFEEE785CCCF2B8FA673" box="[1015,1246,1516,1537]" pageId="5" pageNumber="439">Pteronotus de Macleay</vernacularName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C315D5DE9152FFEEE688CF0D2CC4A605" pageId="5" pageNumber="439" type="reference_group">
<paragraph id="8BB086559152FFEEE688CF0D2CC4A605" blockId="5.[761,1354,1582,2010]" pageId="5" pageNumber="439">
<emphasis id="B97B5A479152FFEEE688CF0D2CC4A63D" bold="true" box="[762,917,1582,1615]" pageId="5" pageNumber="439">Taxonomy.</emphasis>
<taxonomicName id="4C0FFDD69152FFEEE7C5CF0D2CC4A605" authority="J. E." class="Gastropoda" family="Muricidae" genus="Chilonycteris" kingdom="Animalia" order="Neogastropoda" pageId="5" pageNumber="439" phylum="Mollusca" rank="species" species="macleayii">Chilonycteris macleayii J. E. Gray, 1839,</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="C315D5DE9152FFEEE7D0CF7D2C87A6B4" pageId="5" pageNumber="439" type="materials_examined">
<paragraph id="8BB086559152FFEEE7D0CF7D2C87A6B4" blockId="5.[761,1354,1582,2010]" pageId="5" pageNumber="439">
<materialsCitation id="3B678C089152FFEEE7D0CF7D2C87A6B4" ID-GBIF-Occurrence="3863927313" pageId="5" pageNumber="439">
<collectingCountry id="F318C6C59152FFEEE7DCCF7D2CADA605" box="[942,1020,1630,1655]" name="Cuba" pageId="5" pageNumber="439">Cuba</collectingCountry>
.” Restricted by G. SilvaTaboada in 1976 to Guanabacoa,
<collectingRegion id="49CB48B79152FFEEE0ADCF5D2C26A6B4" country="Cuba" name="La Habana" pageId="5" pageNumber="439">Havana Province</collectingRegion>
,
<collectingCountry id="F318C6C59152FFEEE7F5CF8E2C83A6B4" box="[903,978,1709,1734]" name="Cuba" pageId="5" pageNumber="439">Cuba</collectingCountry>
.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C315D5DE9152FFEEE689CFEF2EF7A89C" pageId="5" pageNumber="439" type="discussion">
<paragraph id="8BB086559152FFEEE689CFEF2EF7A89C" blockId="5.[761,1354,1582,2010]" lastBlockId="5.[146,1356,2017,3470]" pageId="5" pageNumber="439">
Description of
<taxonomicName id="4C0FFDD69152FFEEE798CFEF2B30A69F" box="[1002,1121,1740,1773]" class="Gastropoda" family="Muricidae" genus="Pleronotus" kingdom="Animalia" order="Neogastropoda" pageId="5" pageNumber="439" phylum="Mollusca" rank="species" species="macleayii">macleayii</taxonomicName>
was associated with three specimens from
<collectingCountry id="F318C6C59152FFEEE0F0CFD72B9DA767" box="[1154,1228,1780,1813]" name="Cuba" pageId="5" pageNumber="439">Cuba</collectingCountry>
presented by W. S. Macleay to J. E. Gray; original description was detailed and accurate and apparently based on only one male among these specimens. It is believed that such material, however, contained both species of Antillean mustached bats,
<taxonomicName id="4C0FFDD69152FFEEE77DCEC22CFFA870" box="[783,942,2017,2050]" class="Gastropoda" family="Muricidae" genus="Pleronotus" kingdom="Animalia" order="Neogastropoda" pageId="5" pageNumber="439" phylum="Mollusca" rank="species" species="macleayii">P. macleayii</taxonomicName>
and
<taxonomicName id="4C0FFDD69152FFEEE78DCEC22BE6A870" baseAuthorityName="Gundlach" baseAuthorityYear="1840" box="[1023,1207,2017,2050]" class="Mammalia" family="Mormoopidae" genus="Pteronotus" kingdom="Animalia" order="Chiroptera" pageId="5" pageNumber="439" phylum="Chordata" rank="species" species="quadridens">P. quadridens</taxonomicName>
, and that during World WarII, the incorrect specimen (an individual of
<taxonomicName id="4C0FFDD69152FFEEE066C12B2B94A85B" baseAuthorityName="Gundlach" baseAuthorityYear="1840" box="[1044,1221,2056,2089]" class="Mammalia" family="Mormoopidae" genus="Pteronotus" kingdom="Animalia" order="Chiroptera" pageId="5" pageNumber="439" phylum="Chordata" rank="species" species="quadridens">P. quadridens</taxonomicName>
) was segregated as the holotype of
<taxonomicName id="4C0FFDD69152FFEEE665C1132DE0A823" box="[535,689,2096,2129]" class="Gastropoda" family="Muricidae" genus="Pleronotus" kingdom="Animalia" order="Neogastropoda" pageId="5" pageNumber="439" phylum="Mollusca" rank="species" species="macleayii">P. macleayii</taxonomicName>
and subsequently labeled. J. D. Smith in 1972 designated another of these specimens as the holotype. Fossil records of P. macleayu: are reported from
<collectingCountry id="F318C6C59152FFEEE5D8C15D2DD2A8ED" box="[426,643,2174,2207]" name="Dominican Republic" pageId="5" pageNumber="439">the Dominican</collectingCountry>
Republic and New Providence Island,
<collectingCountry id="F318C6C59152FFEEE0B2C15D2A13A8ED" box="[1216,1346,2174,2207]" name="Bahamas" pageId="5" pageNumber="439">Bahamas</collectingCountry>
, suggesting a more extensive distribution in the late Quaternary than today. Two subspecies recognized.
</paragraph>
</subSubSection>
<subSubSection id="C315D5DE9152FFEEE4E6C1D62D25A916" pageId="5" pageNumber="439" type="synonymic_list">
<caption id="DF70D6DD9152FFEEE4E6C1D62D25A916" ID-DOI="http://doi.org/10.5281/zenodo.6419813" ID-Zenodo-Dep="6419813" httpUri="https://zenodo.org/record/6419813/files/figure.png" inLine="true" pageId="5" pageNumber="439" targetBox="[147,738,1588,2003]" targetPageId="5">
<paragraph id="8BB086559152FFEEE4E6C1D62D25A916" blockId="5.[146,1356,2017,3470]" pageId="5" pageNumber="439">
<emphasis id="B97B5A479152FFEEE4E6C1D62D74A964" bold="true" box="[148,549,2293,2326]" pageId="5" pageNumber="439">Subspecies and Distribution.</emphasis>
<taxonomicName id="4C0FFDD69152FFEEE4E7C03F2CBAA94F" authority="]. E. Gray, 1839" authorityName="]. E. Gray" authorityYear="1839" box="[149,1003,2332,2365]" class="Gastropoda" family="Muricidae" genus="Pleronotus" kingdom="Animalia" order="Neogastropoda" pageId="5" pageNumber="439" phylum="Mollusca" rank="subSpecies" species="macleayii" subSpecies="macleayii">P.m.macleayii].E.Gray,1839—CubaandIsladelaJuventud.</taxonomicName>
<taxonomicName id="4C0FFDD69152FFEEE4E6C0602E8CA916" authority="Gosse, 1851" authorityName="Gosse" authorityYear="1851" box="[148,477,2371,2404]" class="Gastropoda" family="Muricidae" genus="Pleronotus" kingdom="Animalia" order="Neogastropoda" pageId="5" pageNumber="439" phylum="Mollusca" rank="subSpecies" species="macleayii" subSpecies="griseus">P. m. griseus Gosse, 1851</taxonomicName>
<collectingCountry id="F318C6C59152FFEEE58EC0602D21A916" box="[508,624,2371,2404]" name="Jamaica" pageId="5" pageNumber="439">Jamaica</collectingCountry>
.
</paragraph>
</caption>
</subSubSection>
<subSubSection id="C315D5DE9152FFEEE4E6C0502B14AAB5" pageId="5" pageNumber="439" type="description">
<paragraph id="8BB086559152FFEEE4E6C0502B14AAB5" blockId="5.[146,1356,2017,3470]" pageId="5" pageNumber="439">
<emphasis id="B97B5A479152FFEEE4E6C0502EC0A9FE" bold="true" box="[148,401,2419,2444]" pageId="5" pageNumber="439">Descriptive notes.</emphasis>
Head-body ¢.41-53 mm, tail 20-25 mm, ear 9-10 mm, hindfoot 9-15 mm, forearm 41-45 mm; weight 4-8 g. Pelage is tricolored dorsally, basal and tip bands vary from dark brown or olive-brown to orange, and central band is whitish to grayish. Ventral fur is paler. Tragus is long and spatulated, with prominent secondary fold. Labionasal plate is very distinct: nostrils have squared flaps above them and prominent lateral spikes to either side. Calcar length is c.1-5 times longer than hindfoot. Skull has slightly elevated rostrum, which is longer than one-half total length of skull. Braincase is globular. Condylo-basal lengths are 14-16 mm. Subspecies macleayi: is smaller than griseus based on external and cranial measurements.
</paragraph>
</subSubSection>
<subSubSection id="C315D5DE9152FFEEE4E6C3F12D90AB4C" pageId="5" pageNumber="439" type="biology_ecology">
<paragraph id="8BB086559152FFEEE4E6C3F12D90AB4C" blockId="5.[146,1356,2017,3470]" pageId="5" pageNumber="439">
<emphasis id="B97B5A479152FFEEE4E6C3F12E52AA9D" bold="true" box="[148,259,2770,2799]" pageId="5" pageNumber="439">Habitat.</emphasis>
Semi-open spaces over streams and watercourses for foraging. Macleays Mustached Bat is considered an obligate cave-dweller, preferably roosting in high-domed chambers in deep and wet cave systems.
</paragraph>
</subSubSection>
<subSubSection id="C315D5DE9152FFEEE4E6C2672D16ABC6" pageId="5" pageNumber="439" type="food_feeding">
<paragraph id="8BB086559152FFEEE4E6C2672D16ABC6" blockId="5.[146,1356,2017,3470]" pageId="5" pageNumber="439">
<emphasis id="B97B5A479152FFEEE4E6C2672EF9AB17" bold="true" box="[148,424,2884,2917]" pageId="5" pageNumber="439">Food and Feeding.</emphasis>
Macleays Mustached Bat feeds on a high diversity of arthropods, including lepidopterans, dipterans, coleopterans, orthopterans, hemipterans, ephemeropterans, and spiders.
</paragraph>
</subSubSection>
<subSubSection id="C315D5DE9152FFEEE4E6C2982D08AC59" pageId="5" pageNumber="439" type="breeding">
<paragraph id="8BB086559152FFEEE4E6C2982D08AC59" blockId="5.[146,1356,2017,3470]" pageId="5" pageNumber="439">
<emphasis id="B97B5A479152FFEEE4E6C2982E4BABAE" bold="true" box="[148,282,3003,3036]" pageId="5" pageNumber="439">Breeding.</emphasis>
Macleays Mustached Bat is monoestrous and bears one young each pregnancy. Breeding begins in March, pregnant females are observed in March—July, and lactation occurs in June-August.
</paragraph>
</subSubSection>
<subSubSection id="C315D5DE9152FFEEE4E0C5122B48ADFC" pageId="5" pageNumber="439" type="activity">
<paragraph id="8BB086559152FFEEE4E0C5122B48ADFC" blockId="5.[146,1356,2017,3470]" pageId="5" pageNumber="439">
<emphasis id="B97B5A479152FFEEE4E0C5122ED3AC20" bold="true" box="[146,386,3121,3154]" pageId="5" pageNumber="439">Activity patterns.</emphasis>
Macleays Mustached Bat has an intermediate flight speed and is most active in edge habitats, usually foraging relatively higher than syntopic species of
<taxonomicName id="4C0FFDD69152FFEEE4E7C5A32E48ACD3" box="[149,281,3200,3233]" class="Mammalia" family="Mormoopidae" genus="Pteronotus" kingdom="Animalia" order="Chiroptera" pageId="5" pageNumber="439" phylum="Chordata" rank="genus">Pteronotus</taxonomicName>
. There is temporal displacement in activity patterns among Macleays Mustached Bat, the Antillean Ghostfaced Bat (
<taxonomicName id="4C0FFDD69152FFEEE773C5882B46ACBA" authorityName="Leach" authorityYear="1821" box="[769,1047,3243,3272]" class="Mammalia" family="Mormoopidae" genus="Mormoops" kingdom="Animalia" order="Chiroptera" pageId="5" pageNumber="439" phylum="Chordata" rank="species" species="blainvillei">Mormoops blainvillei</taxonomicName>
), and the Sooty Mustached Bat (
<taxonomicName id="4C0FFDD69152FFEEE533C5EC2EA0AC82" baseAuthorityName="Gundlach" baseAuthorityYear="1840" box="[321,497,3279,3312]" class="Mammalia" family="Mormoopidae" genus="Pteronotus" kingdom="Animalia" order="Chiroptera" pageId="5" pageNumber="439" phylum="Chordata" rank="species" species="quadridens">P. quadridens</taxonomicName>
) when foraging in the same habitat. In
<collectingCountry id="F318C6C59152FFEEE052C5EC2BC5AC82" box="[1056,1172,3279,3312]" name="Jamaica" pageId="5" pageNumber="439">Jamaica</collectingCountry>
, its activity 1s more intense a few hours after sunset. In
<collectingCountry id="F318C6C59152FFEEE6ACC5D92C72AD65" box="[734,803,3322,3351]" name="Cuba" pageId="5" pageNumber="439">Cuba</collectingCountry>
,it exhibits a unimodal peak of activity in the middle of the night. Echolocation calls during search phase consist of short CF-FM signals of ¢.4-3 milliseconds, with up to three harmonics. Second harmonic is usually most intense and starts at ¢.71 kHz and ends at ¢.56 kHz.
</paragraph>
</subSubSection>
<subSubSection id="C315D5DE9152FFEEE1E2C83B2873A1A5" pageId="5" pageNumber="439" type="biology_ecology">
<paragraph id="8BB086559152FFEEE1E2C83B2873A1A5" blockId="5.[1423,2630,280,710]" pageId="5" pageNumber="439">
<emphasis id="B97B5A479152FFEEE1E2C83B2702A14B" bold="true" box="[1424,2131,280,313]" pageId="5" pageNumber="439">Movements, Home range and Social organization.</emphasis>
Macleays Mustached Bat roosts in association with other bat species, particularly mormoopids, but it forms spatially separated clusters. In
<collectingCountry id="F318C6C59152FFEEE2FFC8482989A1FA" box="[1677,1752,363,392]" name="Cuba" pageId="5" pageNumber="439">Cuba</collectingCountry>
, colonies ofseveral thousand are found; in
<collectingCountry id="F318C6C59152FFEEED3DC8482692A1FA" box="[2383,2499,363,392]" name="Jamaica" pageId="5" pageNumber="439">Jamaica</collectingCountry>
, it sometimes congregates in smaller clusters. Males and females might disappear from daily roosts during mating season.
</paragraph>
</subSubSection>
<subSubSection id="C315D5DE9152FFEEE1E3C8FE2AB0A23F" pageId="5" pageNumber="439" type="conservation">
<paragraph id="8BB086559152FFEEE1E3C8FE2AB0A23F" blockId="5.[1423,2630,280,710]" pageId="5" pageNumber="439">
<emphasis id="B97B5A479152FFEEE1E3C8FE29BEA18C" bold="true" box="[1425,1775,477,510]" pageId="5" pageNumber="439">Status and Conservation.</emphasis>
Classified as Least Concern on The IUCN Red List. Macleays Mustached Bat is less abundant and has a more restricted distribution in
<collectingCountry id="F318C6C59152FFEEEDF8CB2726ACA257" box="[2442,2557,516,549]" name="Jamaica" pageId="5" pageNumber="439">Jamaica</collectingCountry>
than
<collectingCountry id="F318C6C59152FFEEE1E3CB0F2A8DA23F" box="[1425,1500,556,589]" name="Cuba" pageId="5" pageNumber="439">Cuba</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C315D5DE9152FFEEE1E3CB4327C9A2B6" pageId="5" pageNumber="439" type="bibRefCitation_list">
<paragraph id="8BB086559152FFEEE1E3CB4327C9A2B6" blockId="5.[1423,2630,280,710]" pageId="5" pageNumber="439">
<emphasis id="B97B5A479152FFEEE1E3CB43297BA207" bold="true" box="[1425,1578,608,629]" pageId="5" pageNumber="439">Bibliography.</emphasis>
Emrich et al. (2014), Genoways et al. (2005), Mancina (2005), Mancina, Echenique-Diaz et al. (2007), Mancina, Garcia-Rivera &amp; Miller (2012), Mora &amp; Macias (2011), Morgan (2001), Silva-Taboada (1976b), Simmons &amp; Conway (2001), Smith (1972), Velazco, O'Neill et al. (2013).
</paragraph>
</subSubSection>
</treatment>
</document>