treatments-xml/data/03/99/87/039987E8FFEE325E04EEFF37FBA7F74C.xml

<document id="4743230CA583CA6B08D09B787A2C1A24" ID-DOI="10.11646/zootaxa.5496.3.7" ID-ISSN="1175-5326" ID-Zenodo-Dep="13346961" ID-ZooBank="C331701F-4EDB-43D5-8C4F-947028239A03" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1724133683584" checkinUser="plazi" docAuthor="Carvalho, Rafael N. &amp; Kury, Adriano B." docDate="2024" docId="039987E8FFEE325E04EEFF37FBA7F74C" docLanguage="en" docName="zootaxa.5496.3.7.pdf" docOrigin="Zootaxa 5496 (3)" docSource="http://dx.doi.org/10.11646/zootaxa.5496.3.7" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Lacronia utaru Carvalho &amp; Kury, 2024, sp. nov." docType="treatment" docUuid="47119D64-A910-438E-8EFD-A211D80F6831" docUuidSource="ZooBank" docVersion="4" lastPageNumber="426" masterDocId="FFA0FF90FFED32570479FFA0FFEBF571" masterDocTitle="Filling a geographic gap: a new species of genus Lacronia Strand, 1942 (Opiliones, Gonyleptidae) from southern Brazil" masterLastPageNumber="428" masterPageNumber="417" pageNumber="420" updateTime="1725045541638" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="E0CBD6A16793136229BDA15F4D4A021C">Filling a geographic gap: a new species of genus Lacronia Strand, 1942 (Opiliones, Gonyleptidae) from southern Brazil</mods:title>
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<mods:namePart id="1B44F53B9DCD9E0D1A2CBFD90F08B98F">Carvalho, Rafael N.</mods:namePart>
<mods:affiliation id="E9BA81900C091FD31136C1772DACF4BC">Departamento de Invertebrados, Museu Nacional, Universidade Federal do Rio de Janeiro, Quinta da Boa Vista, São Cristóvão, 20.940 - 040, Rio de Janeiro - RJ - BRAZIL</mods:affiliation>
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<mods:namePart id="3EC5B58003ECC5F6140C2503C4C00B58">Kury, Adriano B.</mods:namePart>
<mods:affiliation id="05F90EE5D4ABD13E56B360F08FF0D64F">Departamento de Invertebrados, Museu Nacional, Universidade Federal do Rio de Janeiro, Quinta da Boa Vista, São Cristóvão, 20.940 - 040, Rio de Janeiro - RJ - BRAZIL</mods:affiliation>
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<taxonomicName id="4C304D7DFFEE325404EEFF37FEBBF5C0" authorityName="Carvalho &amp; Kury" authorityYear="2024" box="[151,336,151,177]" class="Arachnida" family="Gonyleptidae" genus="Lacronia" kingdom="Animalia" order="Opiliones" pageId="3" pageNumber="420" phylum="Arthropoda" rank="species" species="utaru" status="sp. nov.">
<emphasis id="B944EAECFFEE325404EEFF37FEBBF5C0" bold="true" box="[151,336,151,177]" italics="true" pageId="3" pageNumber="420">Lacronia utaru</emphasis>
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<paragraph id="8B8F36FEFFEE325404EEFF7FFEABF588" blockId="3.[151,955,151,250]" box="[151,320,223,250]" pageId="3" pageNumber="420">
(
<figureCitation id="130B2A7BFFEE325404E6FF7FFEE0F58B" box="[159,267,223,250]" captionStart-0="FIGURE 2" captionStart-1="FIGURE 3" captionStart-2="FIGURE 4" captionStart-3="FIGURE 5" captionStartId-0="4.[151,250,1546,1570]" captionStartId-1="5.[151,250,1184,1208]" captionStartId-2="6.[151,250,1878,1902]" captionStartId-3="8.[151,250,1048,1072]" captionTargetBox-0="[151,1436,181,1522]" captionTargetBox-1="[151,1436,181,1160]" captionTargetBox-2="[151,1436,237,1855]" captionTargetBox-3="[151,1436,181,1024]" captionTargetId-0="figure-19@4.[151,1436,181,1522]" captionTargetId-1="figure-17@5.[151,1436,181,1160]" captionTargetId-2="figure-48@6.[151,1436,237,1855]" captionTargetId-3="figure-19@8.[151,1436,181,1024]" captionTargetPageId-0="4" captionTargetPageId-1="5" captionTargetPageId-2="6" captionTargetPageId-3="8" captionText-0="FIGURE 2. Habitus in vivo of Lacronia utaru sp. nov., ♂ paratype (MNRJ 3697), from Brazil, Paraná, Paranaguá, Estação Ecológica do Guaraguaçu. A: dorsal view. B: fronto-dorsal view. C: posterior view. D: dorso-posterior view. E: ventral view. Photographs by Adriano Kury." captionText-1="FIGURE 3. Lacronia utaru sp. nov., ♂ holotype (MHNCI). A: habitus, dorsal view. B: ventral view. C: lateral view. D: posterior view. E: Fe–Ta IV, prolateral view (left leg) and retrolateral view (right leg). Scale bars = 1 mm." captionText-2="FIGURE 4. Lacronia utaru sp. nov., ♂ holotype (MHNCI). A: dorsal scutum, chelicerae, trochanters and coxae IV, dorsal view. B: dorsal scutum, lateral view. C: ocularium, anterior view. D: right Fe–Ti III, retrolateral view. E: armature of mesotergal area III, posterior view. F: detail of the prodorsal apophysis of coxa IV, posterior view. G–J: right Tr–Ti IV, G: dorsal view. H: prolateral view. I: ventral view. J: retrolateral view. K: right Mt–Ta IV, retrolateral view. Scale bars = 1 mm." captionText-3="FIGURE 5. Lacronia utaru sp. nov., ♂ holotype (MHNCI), genitalia, distal part.A: dorsal view. B: lateral view. Scale bars: 100 μm. Letters A to E represent the AE11 macrosetae scheme (Kury &amp; Villarreal 2015, Kury &amp; Machado 2021)." figureDoi-0="http://doi.org/10.5281/zenodo.13346967" figureDoi-1="http://doi.org/10.5281/zenodo.13346969" figureDoi-2="http://doi.org/10.5281/zenodo.13346971" figureDoi-3="http://doi.org/10.5281/zenodo.13346973" httpUri-0="https://zenodo.org/record/13346967/files/figure.png" httpUri-1="https://zenodo.org/record/13346969/files/figure.png" httpUri-2="https://zenodo.org/record/13346971/files/figure.png" httpUri-3="https://zenodo.org/record/13346973/files/figure.png" pageId="3" pageNumber="420">Figs. 2–5</figureCitation>
, 
<figureCitation id="130B2A7BFFEE3254056EFF7FFED3F588" box="[279,312,223,249]" captionStart="FIGURE 6" captionStartId="8.[151,250,1732,1756]" captionTargetBox="[151,1436,1132,1708]" captionTargetId="figure-62@8.[151,1436,1132,1708]" captionTargetPageId="8" captionText="FIGURE 6. Schematic representation of males’ DS, showing the two main sets of colored tubercles in the ‘not areolated Lacronia’. The red rectangle indicates the area of the central set, and the blue rectangles indicate the AS lateral sets.A: Lacronia serripes Mello-Leitão, ♂ (MZSP 14342). B: Lacronia utaru sp. nov., ♂ holotype (MHNCI). Scale bars = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.13346975" httpUri="https://zenodo.org/record/13346975/files/figure.png" pageId="3" pageNumber="420">6B</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C32A6575FFEE325404EEFE87FB68F4DF" pageId="3" pageNumber="420" type="materials_examined">
<paragraph id="8B8F36FEFFEE325404EEFE87FB21F4F8" blockId="3.[151,1437,295,1006]" pageId="3" pageNumber="420">
<emphasis id="B944EAECFFEE325404EEFE87FEC9F430" bold="true" box="[151,290,295,321]" pageId="3" pageNumber="420">Type series.</emphasis>
<materialsCitation id="3B583CA3FFEE32540551FE87FF23F417" collectingDate="1989-06-10" collectingDateMax="2024-01-16" collectingDateMin="1989-06-10" collectionCode="MHNCI, V, R" collectorName="Segalla. M." country="Brazil" county="Porto de Cima" elevation="16" location="Estacao Ecologica do Guaraguacu" municipality="Paranagua" pageId="3" pageNumber="420" specimenCode="MNRJ 3697" specimenCount="2" specimenCount-male="2" stateProvince="Parana" typeStatus="holotype">
<typeStatus id="548B885CFFEE32540551FE87FE7BF430" box="[296,400,295,321]" pageId="3" pageNumber="420" type="holotype">Holotype</typeStatus>
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(
<collectionCode id="ED21AE3BFFEE325405C0FE87FDF7F430" box="[441,540,295,321]" country="0" httpUri="http://grbio.org/cool/ej4f-8m62" name="Museu de Historia Natural Capao de Imbuia (Brazil)" pageId="3" pageNumber="420">MHNCI</collectionCode>
), from 
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, 
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<collectingRegion id="49F4F81CFFEE325406C5FE88FCA5F430" box="[700,846,295,322]" country="Brazil" name="Parana" pageId="3" pageNumber="420">Paraná State</collectingRegion>
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, 
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, 
<collectingCounty id="62EE4E72FFEE325407B7FE88FB9AF430" box="[974,1137,295,322]" pageId="3" pageNumber="420">
Porto 
<collectorName id="26C55328FFEE32540069FE87FB9AF430" box="[1040,1137,295,321]" pageId="3" pageNumber="420">de Cima</collectorName>
</collectingCounty>
, 
<date id="FF8E103EFFEE32540002FE87FB1EF430" box="[1147,1269,295,321]" pageId="3" pageNumber="420" value="1989-06-10">
<collectingDate id="EFCAE9D6FFEE32540002FE87FB1EF430" box="[1147,1269,295,321]" pageId="3" pageNumber="420" value="1989-06-10">10.vi.1989</collectingDate>
</date>
, 
<collectorName id="26C55328FFEE32540179FE87FA94F433" box="[1280,1407,295,322]" pageId="3" pageNumber="420">Segalla. M.</collectorName>
<collectionCode id="ED21AE3BFFEE325401FCFE88FA72F433" box="[1413,1433,296,322]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="3" pageNumber="420" type="Museum">V</collectionCode>
. 
<emphasis id="B944EAECFFEE325404EEFEEBFF2AF414" box="[151,193,331,357]" italics="true" pageId="3" pageNumber="420">leg.</emphasis>
;
</materialsCitation>
<materialsCitation id="3B583CA3FFEE325404B7FEEBFB21F4F8" collectingDate="2024-01-15" elevation="16.0" pageId="3" pageNumber="420">
<specimenCount id="9D36FD77FFEE325404B7FEEBFF13F417" box="[206,248,331,358]" count="1" pageId="3" pageNumber="420" type="male">1 ♂</specimenCount>
<typeStatus id="548B885CFFEE32540487FEECFEB5F417" box="[254,350,332,358]" pageId="3" pageNumber="420" type="paratype">paratype</typeStatus>
(
<specimenCode id="DB969E85FFEE32540515FEEBFE10F414" box="[364,507,331,357]" collectionCode="MNRJ" country="Brazil" httpUri="http://grbio.org/cool/zi1i-a0b5" name="Museu Nacional/Universidade Federal de Rio de Janeiro" pageId="3" pageNumber="420">MNRJ 3697</specimenCode>
), from 
<collectingCountry id="F327766EFFEE32540633FEEBFD7AF414" box="[586,657,331,357]" name="Brazil" pageId="3" pageNumber="420">Brazil</collectingCountry>
, 
<emphasis id="B944EAECFFEE325406E5FEECFCC4F414" box="[668,815,331,358]" italics="true" pageId="3" pageNumber="420">
<collectingRegion id="49F4F81CFFEE325406E5FEECFCC4F414" box="[668,815,331,358]" country="Brazil" name="Parana" pageId="3" pageNumber="420">Paraná State</collectingRegion>
</emphasis>
, 
<collectingMunicipality id="6BEBAC84FFEE32540745FEEBFC5FF414" box="[828,948,331,357]" pageId="3" pageNumber="420">Paranaguá</collectingMunicipality>
, 
<location id="8EEF6025FFEE325407C7FEECFAD5F414" LSID="urn:lsid:plazi:treatment:039987E8FFEE325E04EEFF37FBA7F74C:8EEF6025FFEE325407C7FEECFAD5F414" box="[958,1342,331,358]" country="Brazil" county="Porto de Cima" municipality="Paranagua" name="Estacao Ecologica do Guaraguacu" pageId="3" pageNumber="420" stateProvince="Parana">Estação Ecológica do Guaraguaçu</location>
, 
<location id="8EEF6025FFEE32540131FEEBFEE5F4FB" LSID="urn:lsid:plazi:treatment:039987E8FFEE325E04EEFF37FBA7F74C:8EEF6025FFEE32540131FEEBFEE5F4FB" country="Brazil" county="Porto de Cima" municipality="Paranagua" name="Estrada do Bosque" pageId="3" pageNumber="420" stateProvince="Parana">Estrada do Bosque</location>
(-25.6193°, -48.52682°, alt.: 
<quantity id="4CC89B1BFFEE32540620FECFFD7DF4F8" box="[601,662,367,393]" metricMagnitude="1" metricUnit="m" metricValue="1.6" pageId="3" pageNumber="420" unit="m" value="16.0">
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); 
<date id="FF8E103EFFEE325406D3FECFFCAAF4FB" box="[682,833,367,394]" pageId="3" pageNumber="420" value="2024-01-15" valueMax="2024-01-16" valueMin="2024-01-15">
<collectingDate id="EFCAE9D6FFEE325406D3FECFFCAAF4FB" box="[682,833,367,394]" pageId="3" pageNumber="420" value="2024-01-15" valueMax="2024-01-16" valueMin="2024-01-15">15–16.i.2024</collectingDate>
</date>
, Carvalho 
<taxonomicNameLabel id="A2775797FFEE325407C3FED0FC1DF4FB" box="[954,1014,368,394]" pageId="3" pageNumber="420" rank="subSpecies">
<collectionCode id="ED21AE3BFFEE325407C3FED0FC3BF4FB" box="[954,976,368,394]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="3" pageNumber="420">R</collectionCode>
. N.
</taxonomicNameLabel>
&amp; 
<collectorName id="26C55328FFEE32540060FED0FB72F4FB" box="[1049,1177,367,394]" pageId="3" pageNumber="420">Kury, A. B.</collectorName>
<emphasis id="B944EAECFFEE325400D9FECFFB21F4F8" box="[1184,1226,367,393]" italics="true" pageId="3" pageNumber="420">leg.</emphasis>
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</paragraph>
<paragraph id="8B8F36FEFFEE325404BEFE33FB68F4DF" blockId="3.[151,1437,295,1006]" box="[199,1155,403,430]" pageId="3" pageNumber="420">
<emphasis id="B944EAECFFEE325404BEFE33FE8DF4DC" bold="true" box="[199,358,403,429]" pageId="3" pageNumber="420">
<typeStatus id="548B885CFFEE325404BEFE33FEE9F4DC" box="[199,258,403,429]" pageId="3" pageNumber="420">Type</typeStatus>
locality.
</emphasis>
<collectingCountry id="F327766EFFEE32540515FE33FE58F4DC" box="[364,435,403,429]" name="Brazil" pageId="3" pageNumber="420">Brazil</collectingCountry>
, 
<emphasis id="B944EAECFFEE325405C6FE34FDB9F4DC" box="[447,594,403,430]" italics="true" pageId="3" pageNumber="420">
<collectingRegion id="49F4F81CFFEE325405C6FE34FDB9F4DC" box="[447,594,403,430]" country="Brazil" name="Parana" pageId="3" pageNumber="420">Paraná State</collectingRegion>
</emphasis>
, Morretes, 
<location id="8EEF6025FFEE325406AAFE34FC9EF4DC" LSID="urn:lsid:plazi:treatment:039987E8FFEE325E04EEFF37FBA7F74C:8EEF6025FFEE325406AAFE34FC9EF4DC" box="[723,885,403,430]" country="Brazil" name="Porto de Cima" pageId="3" pageNumber="420" stateProvince="Parana">
Porto 
<collectorName id="26C55328FFEE3254076FFE33FC9EF4DC" box="[790,885,403,429]" pageId="3" pageNumber="420">de Cima</collectorName>
</location>
(-25.422°, -48.88552°).
</paragraph>
</subSubSection>
<subSubSection id="C32A6575FFEE325404BEFE17FF1BF74C" pageId="3" pageNumber="420" type="etymology">
<paragraph id="8B8F36FEFFEE325404BEFE17FF1BF74C" blockId="3.[151,1437,295,1006]" pageId="3" pageNumber="420">
<emphasis id="B944EAECFFEE325404BEFE17FEA5F4A0" bold="true" box="[199,334,439,465]" pageId="3" pageNumber="420">Etymology.</emphasis>
The specific name is a noun in apposition referring to the Utaru, an agrarian tribe in the universe of the game Horizon Forbidden West developed by Guerrilla Games. The body painting of the Utaru exhibits a color and shape pattern very similar to that found on the tubercles highlighted on the 
<collectingCountry id="F327766EFFEE32540037FE5FFB99F768" box="[1102,1138,511,537]" name="American Samoa" pageId="3" pageNumber="420">AS</collectingCountry>
lateral borders of this new species.
</paragraph>
</subSubSection>
<subSubSection id="C32A6575FFEE325404BEFDE7FDA1F6D4" pageId="3" pageNumber="420" type="diagnosis">
<paragraph id="8B8F36FEFFEE325404BEFDE7FDA1F6D4" blockId="3.[151,1437,295,1006]" pageId="3" pageNumber="420">
<emphasis id="B944EAECFFEE325404BEFDE7FEA9F710" bold="true" box="[199,322,583,609]" pageId="3" pageNumber="420">Diagnosis.</emphasis>
<taxonomicName id="4C304D7DFFEE32540531FDE8FE1AF710" authorityName="Carvalho &amp; Kury" authorityYear="2024" box="[328,497,584,609]" class="Arachnida" family="Gonyleptidae" genus="Lacronia" kingdom="Animalia" order="Opiliones" pageId="3" pageNumber="420" phylum="Arthropoda" rank="species" species="utaru" status="sp. nov.">
<emphasis id="B944EAECFFEE32540531FDE8FE1AF710" box="[328,497,584,609]" italics="true" pageId="3" pageNumber="420">Lacronia utaru</emphasis>
</taxonomicName>
<emphasis id="B944EAECFFEE3254058EFDE7FDBAF710" bold="true" box="[503,593,583,609]" pageId="3" pageNumber="420">
<taxonomicNameLabel id="A2775797FFEE3254058EFDE7FDBAF710" box="[503,593,583,609]" pageId="3" pageNumber="420" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
can be distinguished from congeners by the following combination of male characters: a) 
<collectingCountry id="F327766EFFEE32540541FDCBFEB7F7F4" box="[312,348,619,645]" name="American Samoa" pageId="3" pageNumber="420">AS</collectingCountry>
lateral borders with light-colored tubercles contrasting with its background (without areolated spots surrounding the tubercles) at the height of mesotergum areas II–IV (
<figureCitation id="130B2A7BFFEE32540072FD2FFB7CF7D8" box="[1035,1175,655,681]" captionStart="FIGURE 2" captionStartId="4.[151,250,1546,1570]" captionTargetBox="[151,1436,181,1522]" captionTargetId="figure-19@4.[151,1436,181,1522]" captionTargetPageId="4" captionText="FIGURE 2. Habitus in vivo of Lacronia utaru sp. nov., ♂ paratype (MNRJ 3697), from Brazil, Paraná, Paranaguá, Estação Ecológica do Guaraguaçu. A: dorsal view. B: fronto-dorsal view. C: posterior view. D: dorso-posterior view. E: ventral view. Photographs by Adriano Kury." figureDoi="http://doi.org/10.5281/zenodo.13346967" httpUri="https://zenodo.org/record/13346967/files/figure.png" pageId="3" pageNumber="420">Figs. 2A–D</figureCitation>
, 
<figureCitation id="130B2A7BFFEE325400DEFD2FFAFFF7D8" box="[1191,1300,655,681]" captionStart="FIGURE 3" captionStartId="5.[151,250,1184,1208]" captionTargetBox="[151,1436,181,1160]" captionTargetId="figure-17@5.[151,1436,181,1160]" captionTargetPageId="5" captionText="FIGURE 3. Lacronia utaru sp. nov., ♂ holotype (MHNCI). A: habitus, dorsal view. B: ventral view. C: lateral view. D: posterior view. E: Fe–Ta IV, prolateral view (left leg) and retrolateral view (right leg). Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13346969" httpUri="https://zenodo.org/record/13346969/files/figure.png" pageId="3" pageNumber="420">3A, C–D</figureCitation>
, 
<figureCitation id="130B2A7BFFEE3254015CFD2FFA87F7D8" box="[1317,1388,655,681]" captionStart="FIGURE 4" captionStartId="6.[151,250,1878,1902]" captionTargetBox="[151,1436,237,1855]" captionTargetId="figure-48@6.[151,1436,237,1855]" captionTargetPageId="6" captionText="FIGURE 4. Lacronia utaru sp. nov., ♂ holotype (MHNCI). A: dorsal scutum, chelicerae, trochanters and coxae IV, dorsal view. B: dorsal scutum, lateral view. C: ocularium, anterior view. D: right Fe–Ti III, retrolateral view. E: armature of mesotergal area III, posterior view. F: detail of the prodorsal apophysis of coxa IV, posterior view. G–J: right Tr–Ti IV, G: dorsal view. H: prolateral view. I: ventral view. J: retrolateral view. K: right Mt–Ta IV, retrolateral view. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13346971" httpUri="https://zenodo.org/record/13346971/files/figure.png" pageId="3" pageNumber="420">4A–B</figureCitation>
); b) Ocularium elliptical in dorsal view, almost perpendicular in relation to the carapace (
<figureCitation id="130B2A7BFFEE32540016FD13FB21F7BC" box="[1135,1226,691,717]" captionStart="FIGURE 4" captionStartId="6.[151,250,1878,1902]" captionTargetBox="[151,1436,237,1855]" captionTargetId="figure-48@6.[151,1436,237,1855]" captionTargetPageId="6" captionText="FIGURE 4. Lacronia utaru sp. nov., ♂ holotype (MHNCI). A: dorsal scutum, chelicerae, trochanters and coxae IV, dorsal view. B: dorsal scutum, lateral view. C: ocularium, anterior view. D: right Fe–Ti III, retrolateral view. E: armature of mesotergal area III, posterior view. F: detail of the prodorsal apophysis of coxa IV, posterior view. G–J: right Tr–Ti IV, G: dorsal view. H: prolateral view. I: ventral view. J: retrolateral view. K: right Mt–Ta IV, retrolateral view. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13346971" httpUri="https://zenodo.org/record/13346971/files/figure.png" pageId="3" pageNumber="420">Fig. 4B</figureCitation>
); c) Carapace and mesotergum areas I–IV with ordinary tubercles not contrasting with its background (
<figureCitation id="130B2A7BFFEE3254003EFD77FB25F780" box="[1095,1230,727,753]" captionStart="FIGURE 2" captionStartId="4.[151,250,1546,1570]" captionTargetBox="[151,1436,181,1522]" captionTargetId="figure-19@4.[151,1436,181,1522]" captionTargetPageId="4" captionText="FIGURE 2. Habitus in vivo of Lacronia utaru sp. nov., ♂ paratype (MNRJ 3697), from Brazil, Paraná, Paranaguá, Estação Ecológica do Guaraguaçu. A: dorsal view. B: fronto-dorsal view. C: posterior view. D: dorso-posterior view. E: ventral view. Photographs by Adriano Kury." figureDoi="http://doi.org/10.5281/zenodo.13346967" httpUri="https://zenodo.org/record/13346967/files/figure.png" pageId="3" pageNumber="420">Figs. 2A–D</figureCitation>
, 
<figureCitation id="130B2A7BFFEE325400A1FD77FAABF780" box="[1240,1344,727,753]" captionStart="FIGURE 3" captionStartId="5.[151,250,1184,1208]" captionTargetBox="[151,1436,181,1160]" captionTargetId="figure-17@5.[151,1436,181,1160]" captionTargetPageId="5" captionText="FIGURE 3. Lacronia utaru sp. nov., ♂ holotype (MHNCI). A: habitus, dorsal view. B: ventral view. C: lateral view. D: posterior view. E: Fe–Ta IV, prolateral view (left leg) and retrolateral view (right leg). Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13346969" httpUri="https://zenodo.org/record/13346969/files/figure.png" pageId="3" pageNumber="420">3A, C–D</figureCitation>
, 
<figureCitation id="130B2A7BFFEE32540132FD77FA9AF780" box="[1355,1393,727,753]" captionStart="FIGURE 4" captionStartId="6.[151,250,1878,1902]" captionTargetBox="[151,1436,237,1855]" captionTargetId="figure-48@6.[151,1436,237,1855]" captionTargetPageId="6" captionText="FIGURE 4. Lacronia utaru sp. nov., ♂ holotype (MHNCI). A: dorsal scutum, chelicerae, trochanters and coxae IV, dorsal view. B: dorsal scutum, lateral view. C: ocularium, anterior view. D: right Fe–Ti III, retrolateral view. E: armature of mesotergal area III, posterior view. F: detail of the prodorsal apophysis of coxa IV, posterior view. G–J: right Tr–Ti IV, G: dorsal view. H: prolateral view. I: ventral view. J: retrolateral view. K: right Mt–Ta IV, retrolateral view. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13346971" httpUri="https://zenodo.org/record/13346971/files/figure.png" pageId="3" pageNumber="420">4A</figureCitation>
); d) Ti III proventral face with a comb of three spines (iII) on the distal third, the larger ones not touching each other’s tip (
<figureCitation id="130B2A7BFFEE325404BDFCBFFEF5F648" box="[196,286,799,825]" captionStart="FIGURE 4" captionStartId="6.[151,250,1878,1902]" captionTargetBox="[151,1436,237,1855]" captionTargetId="figure-48@6.[151,1436,237,1855]" captionTargetPageId="6" captionText="FIGURE 4. Lacronia utaru sp. nov., ♂ holotype (MHNCI). A: dorsal scutum, chelicerae, trochanters and coxae IV, dorsal view. B: dorsal scutum, lateral view. C: ocularium, anterior view. D: right Fe–Ti III, retrolateral view. E: armature of mesotergal area III, posterior view. F: detail of the prodorsal apophysis of coxa IV, posterior view. G–J: right Tr–Ti IV, G: dorsal view. H: prolateral view. I: ventral view. J: retrolateral view. K: right Mt–Ta IV, retrolateral view. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13346971" httpUri="https://zenodo.org/record/13346971/files/figure.png" pageId="3" pageNumber="420">Fig. 4D</figureCitation>
); e) Tr IV prolateral proximal/central portion with an almost isosceles triangle-shaped apophysis with a prodorsal basal protuberance (like in 
<taxonomicName id="4C304D7DFFEE32540643FCE4FD59F62C" baseAuthorityName="Mello-Leitao" baseAuthorityYear="1923" box="[570,690,836,861]" class="Arachnida" family="Gonyleptidae" genus="Lacronia" kingdom="Animalia" order="Opiliones" pageId="3" pageNumber="420" phylum="Arthropoda" rank="species" species="serripes">
<emphasis id="B944EAECFFEE32540643FCE4FD59F62C" box="[570,690,836,861]" italics="true" pageId="3" pageNumber="420">L. serripes</emphasis>
</taxonomicName>
) (
<figureCitation id="130B2A7BFFEE325406B2FCE3FCBAF62C" box="[715,849,835,861]" captionStart="FIGURE 3" captionStartId="5.[151,250,1184,1208]" captionTargetBox="[151,1436,181,1160]" captionTargetId="figure-17@5.[151,1436,181,1160]" captionTargetPageId="5" captionText="FIGURE 3. Lacronia utaru sp. nov., ♂ holotype (MHNCI). A: habitus, dorsal view. B: ventral view. C: lateral view. D: posterior view. E: Fe–Ta IV, prolateral view (left leg) and retrolateral view (right leg). Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13346969" httpUri="https://zenodo.org/record/13346969/files/figure.png" pageId="3" pageNumber="420">Figs. 3A–B</figureCitation>
, 
<figureCitation id="130B2A7BFFEE32540725FCE3FC23F62C" box="[860,968,835,861]" captionStart="FIGURE 4" captionStartId="6.[151,250,1878,1902]" captionTargetBox="[151,1436,237,1855]" captionTargetId="figure-48@6.[151,1436,237,1855]" captionTargetPageId="6" captionText="FIGURE 4. Lacronia utaru sp. nov., ♂ holotype (MHNCI). A: dorsal scutum, chelicerae, trochanters and coxae IV, dorsal view. B: dorsal scutum, lateral view. C: ocularium, anterior view. D: right Fe–Ti III, retrolateral view. E: armature of mesotergal area III, posterior view. F: detail of the prodorsal apophysis of coxa IV, posterior view. G–J: right Tr–Ti IV, G: dorsal view. H: prolateral view. I: ventral view. J: retrolateral view. K: right Mt–Ta IV, retrolateral view. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13346971" httpUri="https://zenodo.org/record/13346971/files/figure.png" pageId="3" pageNumber="420">4A, G–H</figureCitation>
); f) dorsal face of Mt IV with a row of 14 conical spines (iIIIIIIIIIiIii) (
<figureCitation id="130B2A7BFFEE325405A2FCC7FDD5F6F0" box="[475,574,871,897]" captionStart="FIGURE 2" captionStartId="4.[151,250,1546,1570]" captionTargetBox="[151,1436,181,1522]" captionTargetId="figure-19@4.[151,1436,181,1522]" captionTargetPageId="4" captionText="FIGURE 2. Habitus in vivo of Lacronia utaru sp. nov., ♂ paratype (MNRJ 3697), from Brazil, Paraná, Paranaguá, Estação Ecológica do Guaraguaçu. A: dorsal view. B: fronto-dorsal view. C: posterior view. D: dorso-posterior view. E: ventral view. Photographs by Adriano Kury." figureDoi="http://doi.org/10.5281/zenodo.13346967" httpUri="https://zenodo.org/record/13346967/files/figure.png" pageId="3" pageNumber="420">Figs. 2C</figureCitation>
, 
<figureCitation id="130B2A7BFFEE32540633FCC7FD87F6F0" box="[586,620,871,897]" captionStart="FIGURE 3" captionStartId="5.[151,250,1184,1208]" captionTargetBox="[151,1436,181,1160]" captionTargetId="figure-17@5.[151,1436,181,1160]" captionTargetPageId="5" captionText="FIGURE 3. Lacronia utaru sp. nov., ♂ holotype (MHNCI). A: habitus, dorsal view. B: ventral view. C: lateral view. D: posterior view. E: Fe–Ta IV, prolateral view (left leg) and retrolateral view (right leg). Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13346969" httpUri="https://zenodo.org/record/13346969/files/figure.png" pageId="3" pageNumber="420">3E</figureCitation>
, 
<figureCitation id="130B2A7BFFEE32540601FCC7FD75F6F0" box="[632,670,871,897]" captionStart="FIGURE 4" captionStartId="6.[151,250,1878,1902]" captionTargetBox="[151,1436,237,1855]" captionTargetId="figure-48@6.[151,1436,237,1855]" captionTargetPageId="6" captionText="FIGURE 4. Lacronia utaru sp. nov., ♂ holotype (MHNCI). A: dorsal scutum, chelicerae, trochanters and coxae IV, dorsal view. B: dorsal scutum, lateral view. C: ocularium, anterior view. D: right Fe–Ti III, retrolateral view. E: armature of mesotergal area III, posterior view. F: detail of the prodorsal apophysis of coxa IV, posterior view. G–J: right Tr–Ti IV, G: dorsal view. H: prolateral view. I: ventral view. J: retrolateral view. K: right Mt–Ta IV, retrolateral view. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13346971" httpUri="https://zenodo.org/record/13346971/files/figure.png" pageId="3" pageNumber="420">4K</figureCitation>
); g) ventral process of stylus flabellum scallop-shaped, slightly bent ventrad (
<figureCitation id="130B2A7BFFEE32540483FC2BFEB8F6D7" box="[250,339,907,934]" captionStart="FIGURE 5" captionStartId="8.[151,250,1048,1072]" captionTargetBox="[151,1436,181,1024]" captionTargetId="figure-19@8.[151,1436,181,1024]" captionTargetPageId="8" captionText="FIGURE 5. Lacronia utaru sp. nov., ♂ holotype (MHNCI), genitalia, distal part.A: dorsal view. B: lateral view. Scale bars: 100 μm. Letters A to E represent the AE11 macrosetae scheme (Kury &amp; Villarreal 2015, Kury &amp; Machado 2021)." figureDoi="http://doi.org/10.5281/zenodo.13346973" httpUri="https://zenodo.org/record/13346973/files/figure.png" pageId="3" pageNumber="420">Fig. 5B</figureCitation>
). Female is unknown.
</paragraph>
</subSubSection>
<subSubSection id="C32A6575FFEE325E04BEFC0FFBA7F74C" lastPageId="9" lastPageNumber="426" pageId="3" pageNumber="420" type="description">
<paragraph id="8B8F36FEFFEE325404BEFC0FFCAEF69C" blockId="3.[151,1437,295,1006]" pageId="3" pageNumber="420">
<emphasis id="B944EAECFFEE325404BEFC0FFEB2F6B8" bold="true" box="[199,345,943,969]" pageId="3" pageNumber="420">Description.</emphasis>
DS measurements of 
<specimenCount id="9D36FD77FFEE3254062CFC10FD81F6BB" box="[597,618,944,970]" count="1" pageId="3" pageNumber="420" type="male">♂</specimenCount>
<typeStatus id="548B885CFFEE3254060BFC0FFD33F6B8" box="[626,728,943,969]" pageId="3" pageNumber="420" type="holotype">holotype</typeStatus>
: CW 2.7, CL 1.8, AW 4.8, AL 3.0; legs I–IV measurements in 
<tableCitation id="C6B20345FFEE325404EEFC73FF00F69C" box="[151,235,979,1005]" captionStart="TABLE 1" captionStartId="3.[152,244,1053,1077]" captionTargetPageId="3" captionText="TABLE 1. Leg measurements (mm) of Lacronia utaru sp. nov., holotype ♂ (MHNCI)." httpUri="http://table.plazi.org/id/DF4F6676FFEE325404E1FBBDFBD4F144" pageId="3" pageNumber="420" tableUuid="DF4F6676FFEE325404E1FBBDFBD4F144">Table 1</tableCitation>
; tarsal counts: 5(3) / 6(3) - 11(3) / 11(3) - 7 / 7 - 7 / 8.
</paragraph>
<caption id="DF4F6676FFEE325404E1FBBDFBD4F144" ID-Table-UUID="DF4F6676FFEE325404E1FBBDFBD4F144" box="[152,1087,1053,1077]" httpUri="http://table.plazi.org/id/DF4F6676FFEE325404E1FBBDFBD4F144" pageId="3" pageNumber="420" startId="3.[152,244,1053,1077]" targetBox="[163,1376,1093,1319]" targetIsTable="true" targetPageId="3" targetType="table">
<paragraph id="8B8F36FEFFEE325404E1FBBDFBD4F144" blockId="3.[152,1087,1053,1077]" box="[152,1087,1053,1077]" pageId="3" pageNumber="420">
<emphasis id="B944EAECFFEE325404E1FBBDFEE4F144" bold="true" box="[152,271,1053,1077]" pageId="3" pageNumber="420">TABLE 1.</emphasis>
Leg measurements (mm) of 
<taxonomicName id="4C304D7DFFEE3254063DFBBEFD0EF144" authorityName="Carvalho &amp; Kury" authorityYear="2024" box="[580,741,1054,1077]" class="Arachnida" family="Gonyleptidae" genus="Lacronia" kingdom="Animalia" order="Opiliones" pageId="3" pageNumber="420" phylum="Arthropoda" rank="species" species="utaru" status="sp. nov.">
<emphasis id="B944EAECFFEE3254063DFBBEFD0EF144" box="[580,741,1054,1077]" italics="true" pageId="3" pageNumber="420">Lacronia utaru</emphasis>
</taxonomicName>
<emphasis id="B944EAECFFEE32540695FBBDFCAAF144" bold="true" box="[748,833,1053,1077]" pageId="3" pageNumber="420">
<taxonomicNameLabel id="A2775797FFEE32540695FBBDFCAAF144" box="[748,833,1053,1077]" pageId="3" pageNumber="420" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, holotype ♂ (MHNCI).
</paragraph>
</caption>
<paragraph id="8B8F36FEFFEE32540538FBE6FA8BF057" pageId="3" pageNumber="420">
<table id="F930C45EFFEECDA804DAFBE5FA8BF056" box="[163,1376,1093,1319]" gridcols="9" gridrows="6" pageId="3" pageNumber="420">
<tr id="350034BCFFEECDA804DAFBE5FA8BF12C" box="[163,1376,1093,1117]" gridrow="0" pageId="3" pageNumber="420" rowspan-0="1">
<th id="76D15DC0FFEECDA80538FBE5FE86F12C" box="[321,365,1093,1117]" gridcol="1" gridrow="0" pageId="3" pageNumber="420">Tr</th>
<th id="76D15DC0FFEECDA805ABFBE5FE15F12C" box="[466,510,1093,1117]" gridcol="2" gridrow="0" pageId="3" pageNumber="420">Fe</th>
<th id="76D15DC0FFEECDA8061BFBE5FD65F12C" box="[610,654,1093,1117]" gridcol="3" gridrow="0" pageId="3" pageNumber="420">Pa</th>
<th id="76D15DC0FFEECDA8068FFBE5FCC9F12C" box="[758,802,1093,1117]" gridcol="4" gridrow="0" pageId="3" pageNumber="420">Ti</th>
<th id="76D15DC0FFEECDA807FBFBE5FC45F12C" box="[898,942,1093,1117]" gridcol="5" gridrow="0" pageId="3" pageNumber="420">Mt</th>
<th id="76D15DC0FFEECDA80069FBE5FBD7F12C" box="[1040,1084,1093,1117]" gridcol="6" gridrow="0" pageId="3" pageNumber="420">Ta</th>
<th id="76D15DC0FFEECDA800E2FBE5FB2CF12C" box="[1179,1223,1093,1117]" gridcol="7" gridrow="0" pageId="3" pageNumber="420">Cl</th>
<th id="76D15DC0FFEECDA8015EFBE5FA8BF12C" box="[1319,1376,1093,1117]" gridcol="8" gridrow="0" pageId="3" pageNumber="420">Total</th>
</tr>
<tr id="350034BCFFEECDA804DAFBCDFA8BF1F4" box="[163,1376,1133,1157]" gridrow="1" pageId="3" pageNumber="420">
<th id="76D15DC0FFEECDA804DAFBCDFF00F1F4" box="[163,235,1133,1157]" gridcol="0" gridrow="1" pageId="3" pageNumber="420">Pp</th>
<td id="76D15DC0FFEECDA80538FBCDFE86F1F4" box="[321,365,1133,1157]" gridcol="1" gridrow="1" pageId="3" pageNumber="420">0.47</td>
<td id="76D15DC0FFEECDA805ABFBCDFE15F1F4" box="[466,510,1133,1157]" gridcol="2" gridrow="1" pageId="3" pageNumber="420">1.15</td>
<td id="76D15DC0FFEECDA8061BFBCDFD65F1F4" box="[610,654,1133,1157]" gridcol="3" gridrow="1" pageId="3" pageNumber="420">0.61</td>
<td id="76D15DC0FFEECDA8068FFBCDFCC9F1F4" box="[758,802,1133,1157]" gridcol="4" gridrow="1" pageId="3" pageNumber="420">0.78</td>
<td id="76D15DC0FFEECDA807FBFBCDFC45F1F4" box="[898,942,1133,1157]" gridcol="5" gridrow="1" pageId="3" pageNumber="420">–</td>
<td id="76D15DC0FFEECDA80069FBCDFBD7F1F4" box="[1040,1084,1133,1157]" gridcol="6" gridrow="1" pageId="3" pageNumber="420">0.68</td>
<td id="76D15DC0FFEECDA800E2FBCDFB2CF1F4" box="[1179,1223,1133,1157]" gridcol="7" gridrow="1" pageId="3" pageNumber="420">0.54</td>
<td id="76D15DC0FFEECDA8015EFBCDFA8BF1F4" box="[1319,1376,1133,1157]" gridcol="8" gridrow="1" pageId="3" pageNumber="420">4.23</td>
</tr>
<tr id="350034BCFFEECDA804DAFB35FA8BF1DF" box="[163,1376,1173,1198]" gridrow="2" pageId="3" pageNumber="420">
<th id="76D15DC0FFEECDA804DAFB35FF00F1DF" box="[163,235,1173,1198]" gridcol="0" gridrow="2" pageId="3" pageNumber="420">Leg I</th>
<td id="76D15DC0FFEECDA80538FB35FE86F1DF" box="[321,365,1173,1198]" gridcol="1" gridrow="2" pageId="3" pageNumber="420">0.50</td>
<td id="76D15DC0FFEECDA805ABFB35FE15F1DF" box="[466,510,1173,1198]" gridcol="2" gridrow="2" pageId="3" pageNumber="420">1.74</td>
<td id="76D15DC0FFEECDA8061BFB35FD65F1DF" box="[610,654,1173,1198]" gridcol="3" gridrow="2" pageId="3" pageNumber="420">0.71</td>
<td id="76D15DC0FFEECDA8068FFB35FCC9F1DF" box="[758,802,1173,1198]" gridcol="4" gridrow="2" pageId="3" pageNumber="420">1.24</td>
<td id="76D15DC0FFEECDA807FBFB35FC45F1DF" box="[898,942,1173,1198]" gridcol="5" gridrow="2" pageId="3" pageNumber="420">1.96</td>
<td id="76D15DC0FFEECDA80069FB35FBD7F1DF" box="[1040,1084,1173,1198]" gridcol="6" gridrow="2" pageId="3" pageNumber="420">1.40</td>
<td id="76D15DC0FFEECDA800E2FB35FB2CF1DF" box="[1179,1223,1173,1198]" gridcol="7" gridrow="2" pageId="3" pageNumber="420">0.14</td>
<td id="76D15DC0FFEECDA8015EFB35FA8BF1DF" box="[1319,1376,1173,1198]" gridcol="8" gridrow="2" pageId="3" pageNumber="420">7.69</td>
</tr>
<tr id="350034BCFFEECDA804DAFB1EFA8BF1A7" box="[163,1376,1214,1238]" gridrow="3" pageId="3" pageNumber="420">
<th id="76D15DC0FFEECDA804DAFB1EFF00F1A7" box="[163,235,1214,1238]" gridcol="0" gridrow="3" pageId="3" pageNumber="420">Leg II</th>
<td id="76D15DC0FFEECDA80538FB1EFE86F1A7" box="[321,365,1214,1238]" gridcol="1" gridrow="3" pageId="3" pageNumber="420">0.65</td>
<td id="76D15DC0FFEECDA805ABFB1EFE15F1A7" box="[466,510,1214,1238]" gridcol="2" gridrow="3" pageId="3" pageNumber="420">3.66</td>
<td id="76D15DC0FFEECDA8061BFB1EFD65F1A7" box="[610,654,1214,1238]" gridcol="3" gridrow="3" pageId="3" pageNumber="420">1.15</td>
<td id="76D15DC0FFEECDA8068FFB1EFCC9F1A7" box="[758,802,1214,1238]" gridcol="4" gridrow="3" pageId="3" pageNumber="420">2.72</td>
<td id="76D15DC0FFEECDA807FBFB1EFC45F1A7" box="[898,942,1214,1238]" gridcol="5" gridrow="3" pageId="3" pageNumber="420">3.86</td>
<td id="76D15DC0FFEECDA80069FB1EFBD7F1A7" box="[1040,1084,1214,1238]" gridcol="6" gridrow="3" pageId="3" pageNumber="420">4.23</td>
<td id="76D15DC0FFEECDA800E2FB1EFB2CF1A7" box="[1179,1223,1214,1238]" gridcol="7" gridrow="3" pageId="3" pageNumber="420">0.24</td>
<td id="76D15DC0FFEECDA8015EFB1EFA8BF1A7" box="[1319,1376,1214,1238]" gridcol="8" gridrow="3" pageId="3" pageNumber="420">16.51</td>
</tr>
<tr id="350034BCFFEECDA804DAFB46FA8BF18E" box="[163,1376,1254,1279]" gridrow="4" pageId="3" pageNumber="420">
<th id="76D15DC0FFEECDA804DAFB46FF00F18E" box="[163,235,1254,1279]" gridcol="0" gridrow="4" pageId="3" pageNumber="420">Leg III</th>
<td id="76D15DC0FFEECDA80538FB46FE86F18E" box="[321,365,1254,1279]" gridcol="1" gridrow="4" pageId="3" pageNumber="420">0.78</td>
<td id="76D15DC0FFEECDA805ABFB46FE15F18E" box="[466,510,1254,1279]" gridcol="2" gridrow="4" pageId="3" pageNumber="420">2.92</td>
<td id="76D15DC0FFEECDA8061BFB46FD65F18E" box="[610,654,1254,1279]" gridcol="3" gridrow="4" pageId="3" pageNumber="420">1.19</td>
<td id="76D15DC0FFEECDA8068FFB46FCC9F18E" box="[758,802,1254,1279]" gridcol="4" gridrow="4" pageId="3" pageNumber="420">1.87</td>
<td id="76D15DC0FFEECDA807FBFB46FC45F18E" box="[898,942,1254,1279]" gridcol="5" gridrow="4" pageId="3" pageNumber="420">2.93</td>
<td id="76D15DC0FFEECDA80069FB46FBD7F18E" box="[1040,1084,1254,1279]" gridcol="6" gridrow="4" pageId="3" pageNumber="420">1.57</td>
<td id="76D15DC0FFEECDA800E2FB46FB2CF18E" box="[1179,1223,1254,1279]" gridcol="7" gridrow="4" pageId="3" pageNumber="420">0.26</td>
<td id="76D15DC0FFEECDA8015EFB46FA8BF18E" box="[1319,1376,1254,1279]" gridcol="8" gridrow="4" pageId="3" pageNumber="420">11.54</td>
</tr>
<tr id="350034BCFFEECDA804DAFAAEFA8BF056" box="[163,1376,1294,1319]" gridrow="5" pageId="3" pageNumber="420">
<th id="76D15DC0FFEECDA804DAFAAEFF00F056" box="[163,235,1294,1319]" gridcol="0" gridrow="5" pageId="3" pageNumber="420">Leg IV</th>
<td id="76D15DC0FFEECDA80538FAAEFE86F056" box="[321,365,1294,1319]" gridcol="1" gridrow="5" pageId="3" pageNumber="420">1.11</td>
<td id="76D15DC0FFEECDA805ABFAAEFE15F056" box="[466,510,1294,1319]" gridcol="2" gridrow="5" pageId="3" pageNumber="420">3.50</td>
<td id="76D15DC0FFEECDA8061BFAAEFD65F056" box="[610,654,1294,1319]" gridcol="3" gridrow="5" pageId="3" pageNumber="420">1.21</td>
<td id="76D15DC0FFEECDA8068FFAAEFCC9F056" box="[758,802,1294,1319]" gridcol="4" gridrow="5" pageId="3" pageNumber="420">2.74</td>
<td id="76D15DC0FFEECDA807FBFAAEFC45F056" box="[898,942,1294,1319]" gridcol="5" gridrow="5" pageId="3" pageNumber="420">4.38</td>
<td id="76D15DC0FFEECDA80069FAAEFBD7F056" box="[1040,1084,1294,1319]" gridcol="6" gridrow="5" pageId="3" pageNumber="420">1.96</td>
<td id="76D15DC0FFEECDA800E2FAAEFB2CF056" box="[1179,1223,1294,1319]" gridcol="7" gridrow="5" pageId="3" pageNumber="420">0.29</td>
<td id="76D15DC0FFEECDA8015EFAAEFA8BF056" box="[1319,1376,1294,1319]" gridcol="8" gridrow="5" pageId="3" pageNumber="420">15.19</td>
</tr>
</table>
</paragraph>
<paragraph id="8B8F36FEFFEE325404BEFADDFDDEF2C3" blockId="3.[151,1437,1404,1970]" pageId="3" pageNumber="420">
<emphasis id="B944EAECFFEE325404BEFADDFEC2F0E7" box="[199,297,1405,1430]" italics="true" pageId="3" pageNumber="420">Dorsum.</emphasis>
DS gamma-pyriform, as long as wide, with lateral borders of the 
<collectingCountry id="F327766EFFEE3254005DFADCFBA3F0E7" box="[1060,1096,1404,1430]" name="American Samoa" pageId="3" pageNumber="420">AS</collectingCountry>
convex, widest at mesotergal area II and higher at mesotergal area III, with posterior border convex (
<figureCitation id="130B2A7BFFEE325407AEFA00FBD6F0CB" box="[983,1085,1440,1466]" captionStart="FIGURE 3" captionStartId="5.[151,250,1184,1208]" captionTargetBox="[151,1436,181,1160]" captionTargetId="figure-17@5.[151,1436,181,1160]" captionTargetPageId="5" captionText="FIGURE 3. Lacronia utaru sp. nov., ♂ holotype (MHNCI). A: habitus, dorsal view. B: ventral view. C: lateral view. D: posterior view. E: Fe–Ta IV, prolateral view (left leg) and retrolateral view (right leg). Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13346969" httpUri="https://zenodo.org/record/13346969/files/figure.png" pageId="3" pageNumber="420">Figs. 3A</figureCitation>
, 
<figureCitation id="130B2A7BFFEE32540035FA00FB98F0CB" box="[1100,1139,1440,1466]" captionStart="FIGURE 4" captionStartId="6.[151,250,1878,1902]" captionTargetBox="[151,1436,237,1855]" captionTargetId="figure-48@6.[151,1436,237,1855]" captionTargetPageId="6" captionText="FIGURE 4. Lacronia utaru sp. nov., ♂ holotype (MHNCI). A: dorsal scutum, chelicerae, trochanters and coxae IV, dorsal view. B: dorsal scutum, lateral view. C: ocularium, anterior view. D: right Fe–Ti III, retrolateral view. E: armature of mesotergal area III, posterior view. F: detail of the prodorsal apophysis of coxa IV, posterior view. G–J: right Tr–Ti IV, G: dorsal view. H: prolateral view. I: ventral view. J: retrolateral view. K: right Mt–Ta IV, retrolateral view. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13346971" httpUri="https://zenodo.org/record/13346971/files/figure.png" pageId="3" pageNumber="420">4A</figureCitation>
). DS anterior border with a pair of shallow cheliceral sockets separated by a small central projection (
<figureCitation id="130B2A7BFFEE32540079FA64FBB6F0AF" box="[1024,1117,1476,1502]" captionStart="FIGURE 4" captionStartId="6.[151,250,1878,1902]" captionTargetBox="[151,1436,237,1855]" captionTargetId="figure-48@6.[151,1436,237,1855]" captionTargetPageId="6" captionText="FIGURE 4. Lacronia utaru sp. nov., ♂ holotype (MHNCI). A: dorsal scutum, chelicerae, trochanters and coxae IV, dorsal view. B: dorsal scutum, lateral view. C: ocularium, anterior view. D: right Fe–Ti III, retrolateral view. E: armature of mesotergal area III, posterior view. F: detail of the prodorsal apophysis of coxa IV, posterior view. G–J: right Tr–Ti IV, G: dorsal view. H: prolateral view. I: ventral view. J: retrolateral view. K: right Mt–Ta IV, retrolateral view. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13346971" httpUri="https://zenodo.org/record/13346971/files/figure.png" pageId="3" pageNumber="420">Fig. 4A</figureCitation>
). Carapace with one pair of prominent tubercles posterior to the ocularium (
<figureCitation id="130B2A7BFFEE325406C1FA48FCFEF373" box="[696,789,1512,1538]" captionStart="FIGURE 4" captionStartId="6.[151,250,1878,1902]" captionTargetBox="[151,1436,237,1855]" captionTargetId="figure-48@6.[151,1436,237,1855]" captionTargetPageId="6" captionText="FIGURE 4. Lacronia utaru sp. nov., ♂ holotype (MHNCI). A: dorsal scutum, chelicerae, trochanters and coxae IV, dorsal view. B: dorsal scutum, lateral view. C: ocularium, anterior view. D: right Fe–Ti III, retrolateral view. E: armature of mesotergal area III, posterior view. F: detail of the prodorsal apophysis of coxa IV, posterior view. G–J: right Tr–Ti IV, G: dorsal view. H: prolateral view. I: ventral view. J: retrolateral view. K: right Mt–Ta IV, retrolateral view. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13346971" httpUri="https://zenodo.org/record/13346971/files/figure.png" pageId="3" pageNumber="420">Fig. 4A</figureCitation>
). Ocularium elliptical (in dorsal view), high (
<emphasis id="B944EAECFFEE32540165FA49FAD5F373" box="[1308,1342,1513,1538]" italics="true" pageId="3" pageNumber="420">ca.</emphasis>
4 × eye diameter), almost forming a 90º angle with DS, placed in the anterior portion of the carapace (
<figureCitation id="130B2A7BFFEE325400A5F9ACFAA9F357" box="[1244,1346,1548,1574]" captionStart="FIGURE 2" captionStartId="4.[151,250,1546,1570]" captionTargetBox="[151,1436,181,1522]" captionTargetId="figure-19@4.[151,1436,181,1522]" captionTargetPageId="4" captionText="FIGURE 2. Habitus in vivo of Lacronia utaru sp. nov., ♂ paratype (MNRJ 3697), from Brazil, Paraná, Paranaguá, Estação Ecológica do Guaraguaçu. A: dorsal view. B: fronto-dorsal view. C: posterior view. D: dorso-posterior view. E: ventral view. Photographs by Adriano Kury." figureDoi="http://doi.org/10.5281/zenodo.13346967" httpUri="https://zenodo.org/record/13346967/files/figure.png" pageId="3" pageNumber="420">Figs. 2A</figureCitation>
, 
<figureCitation id="130B2A7BFFEE32540129F9ACFA73F357" box="[1360,1432,1548,1574]" captionStart="FIGURE 3" captionStartId="5.[151,250,1184,1208]" captionTargetBox="[151,1436,181,1160]" captionTargetId="figure-17@5.[151,1436,181,1160]" captionTargetPageId="5" captionText="FIGURE 3. Lacronia utaru sp. nov., ♂ holotype (MHNCI). A: habitus, dorsal view. B: ventral view. C: lateral view. D: posterior view. E: Fe–Ta IV, prolateral view (left leg) and retrolateral view (right leg). Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13346969" httpUri="https://zenodo.org/record/13346969/files/figure.png" pageId="3" pageNumber="420">3A, C</figureCitation>
, 
<figureCitation id="130B2A7BFFEE325404EEF990FF35F33B" box="[151,222,1584,1610]" captionStart="FIGURE 4" captionStartId="6.[151,250,1878,1902]" captionTargetBox="[151,1436,237,1855]" captionTargetId="figure-48@6.[151,1436,237,1855]" captionTargetPageId="6" captionText="FIGURE 4. Lacronia utaru sp. nov., ♂ holotype (MHNCI). A: dorsal scutum, chelicerae, trochanters and coxae IV, dorsal view. B: dorsal scutum, lateral view. C: ocularium, anterior view. D: right Fe–Ti III, retrolateral view. E: armature of mesotergal area III, posterior view. F: detail of the prodorsal apophysis of coxa IV, posterior view. G–J: right Tr–Ti IV, G: dorsal view. H: prolateral view. I: ventral view. J: retrolateral view. K: right Mt–Ta IV, retrolateral view. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13346971" httpUri="https://zenodo.org/record/13346971/files/figure.png" pageId="3" pageNumber="420">4A–B</figureCitation>
). Ocularium with a pair of almost parallel spines (ca 2 × the eye diameter), slightly inclined frontwards (
<figureCitation id="130B2A7BFFEE325404E6F9F4FF17F31F" box="[159,252,1620,1646]" captionStart="FIGURE 2" captionStartId="4.[151,250,1546,1570]" captionTargetBox="[151,1436,181,1522]" captionTargetId="figure-19@4.[151,1436,181,1522]" captionTargetPageId="4" captionText="FIGURE 2. Habitus in vivo of Lacronia utaru sp. nov., ♂ paratype (MNRJ 3697), from Brazil, Paraná, Paranaguá, Estação Ecológica do Guaraguaçu. A: dorsal view. B: fronto-dorsal view. C: posterior view. D: dorso-posterior view. E: ventral view. Photographs by Adriano Kury." figureDoi="http://doi.org/10.5281/zenodo.13346967" httpUri="https://zenodo.org/record/13346967/files/figure.png" pageId="3" pageNumber="420">Figs 2B</figureCitation>
, 
<figureCitation id="130B2A7BFFEE32540571F9F4FEA4F31F" box="[264,335,1620,1646]" captionStart="FIGURE 4" captionStartId="6.[151,250,1878,1902]" captionTargetBox="[151,1436,237,1855]" captionTargetId="figure-48@6.[151,1436,237,1855]" captionTargetPageId="6" captionText="FIGURE 4. Lacronia utaru sp. nov., ♂ holotype (MHNCI). A: dorsal scutum, chelicerae, trochanters and coxae IV, dorsal view. B: dorsal scutum, lateral view. C: ocularium, anterior view. D: right Fe–Ti III, retrolateral view. E: armature of mesotergal area III, posterior view. F: detail of the prodorsal apophysis of coxa IV, posterior view. G–J: right Tr–Ti IV, G: dorsal view. H: prolateral view. I: ventral view. J: retrolateral view. K: right Mt–Ta IV, retrolateral view. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13346971" httpUri="https://zenodo.org/record/13346971/files/figure.png" pageId="3" pageNumber="420">4A–C</figureCitation>
). Mesotergum divided in four clearly defined areas (
<figureCitation id="130B2A7BFFEE325407E6F9F4FBCBF31E" box="[927,1056,1620,1647]" captionStart="FIGURE 2" captionStartId="4.[151,250,1546,1570]" captionTargetBox="[151,1436,181,1522]" captionTargetId="figure-19@4.[151,1436,181,1522]" captionTargetPageId="4" captionText="FIGURE 2. Habitus in vivo of Lacronia utaru sp. nov., ♂ paratype (MNRJ 3697), from Brazil, Paraná, Paranaguá, Estação Ecológica do Guaraguaçu. A: dorsal view. B: fronto-dorsal view. C: posterior view. D: dorso-posterior view. E: ventral view. Photographs by Adriano Kury." figureDoi="http://doi.org/10.5281/zenodo.13346967" httpUri="https://zenodo.org/record/13346967/files/figure.png" pageId="3" pageNumber="420">Figs 2A, D</figureCitation>
, 
<figureCitation id="130B2A7BFFEE32540055F9F4FBBBF31F" box="[1068,1104,1620,1646]" captionStart="FIGURE 3" captionStartId="5.[151,250,1184,1208]" captionTargetBox="[151,1436,181,1160]" captionTargetId="figure-17@5.[151,1436,181,1160]" captionTargetPageId="5" captionText="FIGURE 3. Lacronia utaru sp. nov., ♂ holotype (MHNCI). A: habitus, dorsal view. B: ventral view. C: lateral view. D: posterior view. E: Fe–Ta IV, prolateral view (left leg) and retrolateral view (right leg). Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13346969" httpUri="https://zenodo.org/record/13346969/files/figure.png" pageId="3" pageNumber="420">3A</figureCitation>
, 
<figureCitation id="130B2A7BFFEE32540024F9F4FB68F31F" box="[1117,1155,1620,1646]" captionStart="FIGURE 4" captionStartId="6.[151,250,1878,1902]" captionTargetBox="[151,1436,237,1855]" captionTargetId="figure-48@6.[151,1436,237,1855]" captionTargetPageId="6" captionText="FIGURE 4. Lacronia utaru sp. nov., ♂ holotype (MHNCI). A: dorsal scutum, chelicerae, trochanters and coxae IV, dorsal view. B: dorsal scutum, lateral view. C: ocularium, anterior view. D: right Fe–Ti III, retrolateral view. E: armature of mesotergal area III, posterior view. F: detail of the prodorsal apophysis of coxa IV, posterior view. G–J: right Tr–Ti IV, G: dorsal view. H: prolateral view. I: ventral view. J: retrolateral view. K: right Mt–Ta IV, retrolateral view. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13346971" httpUri="https://zenodo.org/record/13346971/files/figure.png" pageId="3" pageNumber="420">4A</figureCitation>
). 
<collectingCountry id="F327766EFFEE325400EFF9F4FB51F31F" box="[1174,1210,1620,1646]" name="American Samoa" pageId="3" pageNumber="420">AS</collectingCountry>
lateral borders with 8–12 yellowish prominent tubercles contrasting with the background at level of mesotergum areas II–IV (
<figureCitation id="130B2A7BFFEE3254011DF9D8FF34F3C7" captionStart="FIGURE 2" captionStartId="4.[151,250,1546,1570]" captionTargetBox="[151,1436,181,1522]" captionTargetId="figure-19@4.[151,1436,181,1522]" captionTargetPageId="4" captionText="FIGURE 2. Habitus in vivo of Lacronia utaru sp. nov., ♂ paratype (MNRJ 3697), from Brazil, Paraná, Paranaguá, Estação Ecológica do Guaraguaçu. A: dorsal view. B: fronto-dorsal view. C: posterior view. D: dorso-posterior view. E: ventral view. Photographs by Adriano Kury." figureDoi="http://doi.org/10.5281/zenodo.13346967" httpUri="https://zenodo.org/record/13346967/files/figure.png" pageId="3" pageNumber="420">Figs. 2A–D</figureCitation>
, 
<figureCitation id="130B2A7BFFEE32540493F93CFEDBF3C7" box="[234,304,1692,1718]" captionStart="FIGURE 3" captionStartId="5.[151,250,1184,1208]" captionTargetBox="[151,1436,181,1160]" captionTargetId="figure-17@5.[151,1436,181,1160]" captionTargetPageId="5" captionText="FIGURE 3. Lacronia utaru sp. nov., ♂ holotype (MHNCI). A: habitus, dorsal view. B: ventral view. C: lateral view. D: posterior view. E: Fe–Ta IV, prolateral view (left leg) and retrolateral view (right leg). Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13346969" httpUri="https://zenodo.org/record/13346969/files/figure.png" pageId="3" pageNumber="420">3A, C</figureCitation>
, 
<figureCitation id="130B2A7BFFEE32540545F93CFE68F3C7" box="[316,387,1692,1718]" captionStart="FIGURE 4" captionStartId="6.[151,250,1878,1902]" captionTargetBox="[151,1436,237,1855]" captionTargetId="figure-48@6.[151,1436,237,1855]" captionTargetPageId="6" captionText="FIGURE 4. Lacronia utaru sp. nov., ♂ holotype (MHNCI). A: dorsal scutum, chelicerae, trochanters and coxae IV, dorsal view. B: dorsal scutum, lateral view. C: ocularium, anterior view. D: right Fe–Ti III, retrolateral view. E: armature of mesotergal area III, posterior view. F: detail of the prodorsal apophysis of coxa IV, posterior view. G–J: right Tr–Ti IV, G: dorsal view. H: prolateral view. I: ventral view. J: retrolateral view. K: right Mt–Ta IV, retrolateral view. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13346971" httpUri="https://zenodo.org/record/13346971/files/figure.png" pageId="3" pageNumber="420">4A–B</figureCitation>
). Mesotergum area I divided into left and right halves by a longitudinal median groove (
<figureCitation id="130B2A7BFFEE3254011DF93CFEEBF3AA" captionStart="FIGURE 2" captionStartId="4.[151,250,1546,1570]" captionTargetBox="[151,1436,181,1522]" captionTargetId="figure-19@4.[151,1436,181,1522]" captionTargetPageId="4" captionText="FIGURE 2. Habitus in vivo of Lacronia utaru sp. nov., ♂ paratype (MNRJ 3697), from Brazil, Paraná, Paranaguá, Estação Ecológica do Guaraguaçu. A: dorsal view. B: fronto-dorsal view. C: posterior view. D: dorso-posterior view. E: ventral view. Photographs by Adriano Kury." figureDoi="http://doi.org/10.5281/zenodo.13346967" httpUri="https://zenodo.org/record/13346967/files/figure.png" pageId="3" pageNumber="420">Figs. 2A–B, D</figureCitation>
, 
<figureCitation id="130B2A7BFFEE32540575F960FEDBF3AB" box="[268,304,1728,1754]" captionStart="FIGURE 3" captionStartId="5.[151,250,1184,1208]" captionTargetBox="[151,1436,181,1160]" captionTargetId="figure-17@5.[151,1436,181,1160]" captionTargetPageId="5" captionText="FIGURE 3. Lacronia utaru sp. nov., ♂ holotype (MHNCI). A: habitus, dorsal view. B: ventral view. C: lateral view. D: posterior view. E: Fe–Ta IV, prolateral view (left leg) and retrolateral view (right leg). Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13346969" httpUri="https://zenodo.org/record/13346969/files/figure.png" pageId="3" pageNumber="420">3A</figureCitation>
, 
<figureCitation id="130B2A7BFFEE32540544F960FE88F3AB" box="[317,355,1728,1754]" captionStart="FIGURE 4" captionStartId="6.[151,250,1878,1902]" captionTargetBox="[151,1436,237,1855]" captionTargetId="figure-48@6.[151,1436,237,1855]" captionTargetPageId="6" captionText="FIGURE 4. Lacronia utaru sp. nov., ♂ holotype (MHNCI). A: dorsal scutum, chelicerae, trochanters and coxae IV, dorsal view. B: dorsal scutum, lateral view. C: ocularium, anterior view. D: right Fe–Ti III, retrolateral view. E: armature of mesotergal area III, posterior view. F: detail of the prodorsal apophysis of coxa IV, posterior view. G–J: right Tr–Ti IV, G: dorsal view. H: prolateral view. I: ventral view. J: retrolateral view. K: right Mt–Ta IV, retrolateral view. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13346971" httpUri="https://zenodo.org/record/13346971/files/figure.png" pageId="3" pageNumber="420">4A</figureCitation>
). All areas tuberculate, with tiny tubercles not contrasting with the background (
<figureCitation id="130B2A7BFFEE32540096F960FA73F3AA" box="[1263,1432,1728,1755]" captionStart="FIGURE 2" captionStartId="4.[151,250,1546,1570]" captionTargetBox="[151,1436,181,1522]" captionTargetId="figure-19@4.[151,1436,181,1522]" captionTargetPageId="4" captionText="FIGURE 2. Habitus in vivo of Lacronia utaru sp. nov., ♂ paratype (MNRJ 3697), from Brazil, Paraná, Paranaguá, Estação Ecológica do Guaraguaçu. A: dorsal view. B: fronto-dorsal view. C: posterior view. D: dorso-posterior view. E: ventral view. Photographs by Adriano Kury." figureDoi="http://doi.org/10.5281/zenodo.13346967" httpUri="https://zenodo.org/record/13346967/files/figure.png" pageId="3" pageNumber="420">Figs. 2A–B, D</figureCitation>
, 
<figureCitation id="130B2A7BFFEE325404EEF944FF50F38F" box="[151,187,1764,1790]" captionStart="FIGURE 3" captionStartId="5.[151,250,1184,1208]" captionTargetBox="[151,1436,181,1160]" captionTargetId="figure-17@5.[151,1436,181,1160]" captionTargetPageId="5" captionText="FIGURE 3. Lacronia utaru sp. nov., ♂ holotype (MHNCI). A: habitus, dorsal view. B: ventral view. C: lateral view. D: posterior view. E: Fe–Ta IV, prolateral view (left leg) and retrolateral view (right leg). Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13346969" httpUri="https://zenodo.org/record/13346969/files/figure.png" pageId="3" pageNumber="420">3A</figureCitation>
, 
<figureCitation id="130B2A7BFFEE325404B0F944FF04F38F" box="[201,239,1764,1790]" captionStart="FIGURE 4" captionStartId="6.[151,250,1878,1902]" captionTargetBox="[151,1436,237,1855]" captionTargetId="figure-48@6.[151,1436,237,1855]" captionTargetPageId="6" captionText="FIGURE 4. Lacronia utaru sp. nov., ♂ holotype (MHNCI). A: dorsal scutum, chelicerae, trochanters and coxae IV, dorsal view. B: dorsal scutum, lateral view. C: ocularium, anterior view. D: right Fe–Ti III, retrolateral view. E: armature of mesotergal area III, posterior view. F: detail of the prodorsal apophysis of coxa IV, posterior view. G–J: right Tr–Ti IV, G: dorsal view. H: prolateral view. I: ventral view. J: retrolateral view. K: right Mt–Ta IV, retrolateral view. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13346971" httpUri="https://zenodo.org/record/13346971/files/figure.png" pageId="3" pageNumber="420">4A</figureCitation>
). Mesotergum area I with a pair of prominent tubercles (
<figureCitation id="130B2A7BFFEE3254070CF944FC16F38F" box="[885,1021,1764,1790]" captionStart="FIGURE 4" captionStartId="6.[151,250,1878,1902]" captionTargetBox="[151,1436,237,1855]" captionTargetId="figure-48@6.[151,1436,237,1855]" captionTargetPageId="6" captionText="FIGURE 4. Lacronia utaru sp. nov., ♂ holotype (MHNCI). A: dorsal scutum, chelicerae, trochanters and coxae IV, dorsal view. B: dorsal scutum, lateral view. C: ocularium, anterior view. D: right Fe–Ti III, retrolateral view. E: armature of mesotergal area III, posterior view. F: detail of the prodorsal apophysis of coxa IV, posterior view. G–J: right Tr–Ti IV, G: dorsal view. H: prolateral view. I: ventral view. J: retrolateral view. K: right Mt–Ta IV, retrolateral view. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13346971" httpUri="https://zenodo.org/record/13346971/files/figure.png" pageId="3" pageNumber="420">Figs. 4A–B</figureCitation>
). Mesotergum area III with a pair of outstanding paramedian spines (
<emphasis id="B944EAECFFEE32540587F8A9FDF4F253" box="[510,543,1801,1826]" italics="true" pageId="3" pageNumber="420">ca.</emphasis>
1.5 × the ocularium spines) with subconical apex. (
<figureCitation id="130B2A7BFFEE32540021F8A8FB3CF253" box="[1112,1239,1800,1826]" captionStart="FIGURE 2" captionStartId="4.[151,250,1546,1570]" captionTargetBox="[151,1436,181,1522]" captionTargetId="figure-19@4.[151,1436,181,1522]" captionTargetPageId="4" captionText="FIGURE 2. Habitus in vivo of Lacronia utaru sp. nov., ♂ paratype (MNRJ 3697), from Brazil, Paraná, Paranaguá, Estação Ecológica do Guaraguaçu. A: dorsal view. B: fronto-dorsal view. C: posterior view. D: dorso-posterior view. E: ventral view. Photographs by Adriano Kury." figureDoi="http://doi.org/10.5281/zenodo.13346967" httpUri="https://zenodo.org/record/13346967/files/figure.png" pageId="3" pageNumber="420">Figs 2A–D</figureCitation>
, 
<figureCitation id="130B2A7BFFEE32540098F8A8FAA3F253" box="[1249,1352,1800,1826]" captionStart="FIGURE 3" captionStartId="5.[151,250,1184,1208]" captionTargetBox="[151,1436,181,1160]" captionTargetId="figure-17@5.[151,1436,181,1160]" captionTargetPageId="5" captionText="FIGURE 3. Lacronia utaru sp. nov., ♂ holotype (MHNCI). A: habitus, dorsal view. B: ventral view. C: lateral view. D: posterior view. E: Fe–Ta IV, prolateral view (left leg) and retrolateral view (right leg). Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13346969" httpUri="https://zenodo.org/record/13346969/files/figure.png" pageId="3" pageNumber="420">3A, C–D</figureCitation>
, 
<figureCitation id="130B2A7BFFEE3254012BF8A8FF47F237" captionStart="FIGURE 4" captionStartId="6.[151,250,1878,1902]" captionTargetBox="[151,1436,237,1855]" captionTargetId="figure-48@6.[151,1436,237,1855]" captionTargetPageId="6" captionText="FIGURE 4. Lacronia utaru sp. nov., ♂ holotype (MHNCI). A: dorsal scutum, chelicerae, trochanters and coxae IV, dorsal view. B: dorsal scutum, lateral view. C: ocularium, anterior view. D: right Fe–Ti III, retrolateral view. E: armature of mesotergal area III, posterior view. F: detail of the prodorsal apophysis of coxa IV, posterior view. G–J: right Tr–Ti IV, G: dorsal view. H: prolateral view. I: ventral view. J: retrolateral view. K: right Mt–Ta IV, retrolateral view. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13346971" httpUri="https://zenodo.org/record/13346971/files/figure.png" pageId="3" pageNumber="420">4A–B, E</figureCitation>
). Mesotergum area IV with a transversal row of four to five prominent tubercles (
<figureCitation id="130B2A7BFFEE32540035F88CFB43F237" box="[1100,1192,1836,1862]" captionStart="FIGURE 4" captionStartId="6.[151,250,1878,1902]" captionTargetBox="[151,1436,237,1855]" captionTargetId="figure-48@6.[151,1436,237,1855]" captionTargetPageId="6" captionText="FIGURE 4. Lacronia utaru sp. nov., ♂ holotype (MHNCI). A: dorsal scutum, chelicerae, trochanters and coxae IV, dorsal view. B: dorsal scutum, lateral view. C: ocularium, anterior view. D: right Fe–Ti III, retrolateral view. E: armature of mesotergal area III, posterior view. F: detail of the prodorsal apophysis of coxa IV, posterior view. G–J: right Tr–Ti IV, G: dorsal view. H: prolateral view. I: ventral view. J: retrolateral view. K: right Mt–Ta IV, retrolateral view. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13346971" httpUri="https://zenodo.org/record/13346971/files/figure.png" pageId="3" pageNumber="420">Fig. 4A</figureCitation>
). DS posterior border with a transversal row of prominent tubercles (same color as the background) (
<figureCitation id="130B2A7BFFEE32540065F8F0FB2DF21B" box="[1052,1222,1872,1898]" captionStart="FIGURE 2" captionStartId="4.[151,250,1546,1570]" captionTargetBox="[151,1436,181,1522]" captionTargetId="figure-19@4.[151,1436,181,1522]" captionTargetPageId="4" captionText="FIGURE 2. Habitus in vivo of Lacronia utaru sp. nov., ♂ paratype (MNRJ 3697), from Brazil, Paraná, Paranaguá, Estação Ecológica do Guaraguaçu. A: dorsal view. B: fronto-dorsal view. C: posterior view. D: dorso-posterior view. E: ventral view. Photographs by Adriano Kury." figureDoi="http://doi.org/10.5281/zenodo.13346967" httpUri="https://zenodo.org/record/13346967/files/figure.png" pageId="3" pageNumber="420">Figs. 2A, C–D</figureCitation>
, 
<figureCitation id="130B2A7BFFEE325400AAF8F0FAD7F21B" box="[1235,1340,1872,1898]" captionStart="FIGURE 3" captionStartId="5.[151,250,1184,1208]" captionTargetBox="[151,1436,181,1160]" captionTargetId="figure-17@5.[151,1436,181,1160]" captionTargetPageId="5" captionText="FIGURE 3. Lacronia utaru sp. nov., ♂ holotype (MHNCI). A: habitus, dorsal view. B: ventral view. C: lateral view. D: posterior view. E: Fe–Ta IV, prolateral view (left leg) and retrolateral view (right leg). Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13346969" httpUri="https://zenodo.org/record/13346969/files/figure.png" pageId="3" pageNumber="420">3A, C–D</figureCitation>
, 
<figureCitation id="130B2A7BFFEE32540130F8F0FA7BF21B" box="[1353,1424,1872,1898]" captionStart="FIGURE 4" captionStartId="6.[151,250,1878,1902]" captionTargetBox="[151,1436,237,1855]" captionTargetId="figure-48@6.[151,1436,237,1855]" captionTargetPageId="6" captionText="FIGURE 4. Lacronia utaru sp. nov., ♂ holotype (MHNCI). A: dorsal scutum, chelicerae, trochanters and coxae IV, dorsal view. B: dorsal scutum, lateral view. C: ocularium, anterior view. D: right Fe–Ti III, retrolateral view. E: armature of mesotergal area III, posterior view. F: detail of the prodorsal apophysis of coxa IV, posterior view. G–J: right Tr–Ti IV, G: dorsal view. H: prolateral view. I: ventral view. J: retrolateral view. K: right Mt–Ta IV, retrolateral view. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13346971" httpUri="https://zenodo.org/record/13346971/files/figure.png" pageId="3" pageNumber="420">4A–B</figureCitation>
). Free tergites I–III with a transversal row of ordinary tubercles (
<figureCitation id="130B2A7BFFEE3254071FF8D4FC22F2FF" box="[870,969,1908,1934]" captionStart="FIGURE 2" captionStartId="4.[151,250,1546,1570]" captionTargetBox="[151,1436,181,1522]" captionTargetId="figure-19@4.[151,1436,181,1522]" captionTargetPageId="4" captionText="FIGURE 2. Habitus in vivo of Lacronia utaru sp. nov., ♂ paratype (MNRJ 3697), from Brazil, Paraná, Paranaguá, Estação Ecológica do Guaraguaçu. A: dorsal view. B: fronto-dorsal view. C: posterior view. D: dorso-posterior view. E: ventral view. Photographs by Adriano Kury." figureDoi="http://doi.org/10.5281/zenodo.13346967" httpUri="https://zenodo.org/record/13346967/files/figure.png" pageId="3" pageNumber="420">Figs. 2C</figureCitation>
, 
<figureCitation id="130B2A7BFFEE325407AFF8D4FBD4F2FF" box="[982,1087,1908,1934]" captionStart="FIGURE 3" captionStartId="5.[151,250,1184,1208]" captionTargetBox="[151,1436,181,1160]" captionTargetId="figure-17@5.[151,1436,181,1160]" captionTargetPageId="5" captionText="FIGURE 3. Lacronia utaru sp. nov., ♂ holotype (MHNCI). A: habitus, dorsal view. B: ventral view. C: lateral view. D: posterior view. E: Fe–Ta IV, prolateral view (left leg) and retrolateral view (right leg). Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13346969" httpUri="https://zenodo.org/record/13346969/files/figure.png" pageId="3" pageNumber="420">3A, C–D</figureCitation>
, 
<figureCitation id="130B2A7BFFEE32540035F8D4FB99F2FF" box="[1100,1138,1908,1934]" captionStart="FIGURE 4" captionStartId="6.[151,250,1878,1902]" captionTargetBox="[151,1436,237,1855]" captionTargetId="figure-48@6.[151,1436,237,1855]" captionTargetPageId="6" captionText="FIGURE 4. Lacronia utaru sp. nov., ♂ holotype (MHNCI). A: dorsal scutum, chelicerae, trochanters and coxae IV, dorsal view. B: dorsal scutum, lateral view. C: ocularium, anterior view. D: right Fe–Ti III, retrolateral view. E: armature of mesotergal area III, posterior view. F: detail of the prodorsal apophysis of coxa IV, posterior view. G–J: right Tr–Ti IV, G: dorsal view. H: prolateral view. I: ventral view. J: retrolateral view. K: right Mt–Ta IV, retrolateral view. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13346971" httpUri="https://zenodo.org/record/13346971/files/figure.png" pageId="3" pageNumber="420">4A</figureCitation>
). Anal operculum covered by ordinary tubercles (
<figureCitation id="130B2A7BFFEE325405EDF838FE1CF2C3" box="[404,503,1944,1970]" captionStart="FIGURE 2" captionStartId="4.[151,250,1546,1570]" captionTargetBox="[151,1436,181,1522]" captionTargetId="figure-19@4.[151,1436,181,1522]" captionTargetPageId="4" captionText="FIGURE 2. Habitus in vivo of Lacronia utaru sp. nov., ♂ paratype (MNRJ 3697), from Brazil, Paraná, Paranaguá, Estação Ecológica do Guaraguaçu. A: dorsal view. B: fronto-dorsal view. C: posterior view. D: dorso-posterior view. E: ventral view. Photographs by Adriano Kury." figureDoi="http://doi.org/10.5281/zenodo.13346967" httpUri="https://zenodo.org/record/13346967/files/figure.png" pageId="3" pageNumber="420">Figs. 2C</figureCitation>
, 
<figureCitation id="130B2A7BFFEE3254067BF838FDC3F2C3" box="[514,552,1944,1970]" captionStart="FIGURE 3" captionStartId="5.[151,250,1184,1208]" captionTargetBox="[151,1436,181,1160]" captionTargetId="figure-17@5.[151,1436,181,1160]" captionTargetPageId="5" captionText="FIGURE 3. Lacronia utaru sp. nov., ♂ holotype (MHNCI). A: habitus, dorsal view. B: ventral view. C: lateral view. D: posterior view. E: Fe–Ta IV, prolateral view (left leg) and retrolateral view (right leg). Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13346969" httpUri="https://zenodo.org/record/13346969/files/figure.png" pageId="3" pageNumber="420">3D</figureCitation>
).
</paragraph>
<caption id="DF4F6676FFE9325304EEF9AAFE25F31B" ID-DOI="http://doi.org/10.5281/zenodo.13346967" ID-Zenodo-Dep="13346967" httpUri="https://zenodo.org/record/13346967/files/figure.png" pageId="4" pageNumber="421" startId="4.[151,250,1546,1570]" targetBox="[151,1436,181,1522]" targetPageId="4" targetType="figure">
<paragraph id="8B8F36FEFFE9325304EEF9AAFE25F31B" blockId="4.[151,1437,1546,1642]" pageId="4" pageNumber="421">
<emphasis id="B944EAECFFE9325304EEF9AAFEFEF353" bold="true" box="[151,277,1546,1570]" pageId="4" pageNumber="421">FIGURE 2.</emphasis>
Habitus 
<emphasis id="B944EAECFFE93253050DF9AAFE51F350" box="[372,442,1546,1569]" italics="true" pageId="4" pageNumber="421">in vivo</emphasis>
of 
<taxonomicName id="4C304D7DFFE9325305A6F9AAFD90F353" authorityName="Carvalho &amp; Kury" authorityYear="2024" box="[479,635,1546,1570]" class="Arachnida" family="Gonyleptidae" genus="Lacronia" kingdom="Animalia" order="Opiliones" pageId="4" pageNumber="421" phylum="Arthropoda" rank="species" species="utaru" status="sp. nov.">
<emphasis id="B944EAECFFE9325305A6F9AAFD90F353" box="[479,635,1546,1570]" italics="true" pageId="4" pageNumber="421">Lacronia utaru</emphasis>
</taxonomicName>
<emphasis id="B944EAECFFE9325306FAF9AAFD3EF353" bold="true" box="[643,725,1546,1570]" pageId="4" pageNumber="421">
<taxonomicNameLabel id="A2775797FFE9325306FAF9AAFD3EF353" box="[643,725,1546,1570]" pageId="4" pageNumber="421" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, ♂ paratype (MNRJ 3697), from Brazil, Paraná, Paranaguá, Estação Ecológica do Guaraguaçu. A: dorsal view. B: fronto-dorsal view. C: posterior view. D: dorso-posterior view. E: ventral view. Photographs by Adriano Kury.
</paragraph>
</caption>
<paragraph id="8B8F36FEFFE9325304BEF91DFD0FF2A3" blockId="4.[151,1437,1724,2003]" pageId="4" pageNumber="421">
<emphasis id="B944EAECFFE9325304BEF91DFEF8F3A7" box="[199,275,1725,1750]" italics="true" pageId="4" pageNumber="421">Venter.</emphasis>
Cx I–III sub-parallel to each other, each with ventral longitudinal rows of setiferous tubercles (Cx I rows with higher and sharper tubercles than the others) (
<figureCitation id="130B2A7BFFE9325306A0F940FCDAF38B" box="[729,817,1760,1786]" captionStart="FIGURE 2" captionStartId="4.[151,250,1546,1570]" captionTargetBox="[151,1436,181,1522]" captionTargetId="figure-19@4.[151,1436,181,1522]" captionTargetPageId="4" captionText="FIGURE 2. Habitus in vivo of Lacronia utaru sp. nov., ♂ paratype (MNRJ 3697), from Brazil, Paraná, Paranaguá, Estação Ecológica do Guaraguaçu. A: dorsal view. B: fronto-dorsal view. C: posterior view. D: dorso-posterior view. E: ventral view. Photographs by Adriano Kury." figureDoi="http://doi.org/10.5281/zenodo.13346967" httpUri="https://zenodo.org/record/13346967/files/figure.png" pageId="4" pageNumber="421">Fig. 2E</figureCitation>
). Cx II retro-ventral margin with a longitudinal row of acuminate tubercles on the distal third (
<figureCitation id="130B2A7BFFE932530624F8A4FD5DF26F" box="[605,694,1796,1822]" captionStart="FIGURE 3" captionStartId="5.[151,250,1184,1208]" captionTargetBox="[151,1436,181,1160]" captionTargetId="figure-17@5.[151,1436,181,1160]" captionTargetPageId="5" captionText="FIGURE 3. Lacronia utaru sp. nov., ♂ holotype (MHNCI). A: habitus, dorsal view. B: ventral view. C: lateral view. D: posterior view. E: Fe–Ta IV, prolateral view (left leg) and retrolateral view (right leg). Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13346969" httpUri="https://zenodo.org/record/13346969/files/figure.png" pageId="4" pageNumber="421">Fig. 3B</figureCitation>
). Cx III retro-ventral border with a transversal row of acuminate tubercles on the centro-distal two thirds (
<figureCitation id="130B2A7BFFE93253061AF888FD50F233" box="[611,699,1832,1858]" captionStart="FIGURE 3" captionStartId="5.[151,250,1184,1208]" captionTargetBox="[151,1436,181,1160]" captionTargetId="figure-17@5.[151,1436,181,1160]" captionTargetPageId="5" captionText="FIGURE 3. Lacronia utaru sp. nov., ♂ holotype (MHNCI). A: habitus, dorsal view. B: ventral view. C: lateral view. D: posterior view. E: Fe–Ta IV, prolateral view (left leg) and retrolateral view (right leg). Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13346969" httpUri="https://zenodo.org/record/13346969/files/figure.png" pageId="4" pageNumber="421">Fig. 3B</figureCitation>
). Cx IV much larger than the others, directed obliquely (
<figureCitation id="130B2A7BFFE93253014EF888FA73F233" box="[1335,1432,1832,1858]" captionStart="FIGURE 2" captionStartId="4.[151,250,1546,1570]" captionTargetBox="[151,1436,181,1522]" captionTargetId="figure-19@4.[151,1436,181,1522]" captionTargetPageId="4" captionText="FIGURE 2. Habitus in vivo of Lacronia utaru sp. nov., ♂ paratype (MNRJ 3697), from Brazil, Paraná, Paranaguá, Estação Ecológica do Guaraguaçu. A: dorsal view. B: fronto-dorsal view. C: posterior view. D: dorso-posterior view. E: ventral view. Photographs by Adriano Kury." figureDoi="http://doi.org/10.5281/zenodo.13346967" httpUri="https://zenodo.org/record/13346967/files/figure.png" pageId="4" pageNumber="421">Figs. 2E</figureCitation>
, 
<figureCitation id="130B2A7BFFE9325304EEF8ECFF50F217" box="[151,187,1868,1894]" captionStart="FIGURE 3" captionStartId="5.[151,250,1184,1208]" captionTargetBox="[151,1436,181,1160]" captionTargetId="figure-17@5.[151,1436,181,1160]" captionTargetPageId="5" captionText="FIGURE 3. Lacronia utaru sp. nov., ♂ holotype (MHNCI). A: habitus, dorsal view. B: ventral view. C: lateral view. D: posterior view. E: Fe–Ta IV, prolateral view (left leg) and retrolateral view (right leg). Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13346969" httpUri="https://zenodo.org/record/13346969/files/figure.png" pageId="4" pageNumber="421">3B</figureCitation>
). Intercoxal bridges well-marked (
<figureCitation id="130B2A7BFFE932530646F8ECFD72F217" box="[575,665,1868,1894]" captionStart="FIGURE 2" captionStartId="4.[151,250,1546,1570]" captionTargetBox="[151,1436,181,1522]" captionTargetId="figure-19@4.[151,1436,181,1522]" captionTargetPageId="4" captionText="FIGURE 2. Habitus in vivo of Lacronia utaru sp. nov., ♂ paratype (MNRJ 3697), from Brazil, Paraná, Paranaguá, Estação Ecológica do Guaraguaçu. A: dorsal view. B: fronto-dorsal view. C: posterior view. D: dorso-posterior view. E: ventral view. Photographs by Adriano Kury." figureDoi="http://doi.org/10.5281/zenodo.13346967" httpUri="https://zenodo.org/record/13346967/files/figure.png" pageId="4" pageNumber="421">Figs 2E</figureCitation>
, 
<figureCitation id="130B2A7BFFE9325306DCF8ECFD22F217" box="[677,713,1868,1894]" captionStart="FIGURE 3" captionStartId="5.[151,250,1184,1208]" captionTargetBox="[151,1436,181,1160]" captionTargetId="figure-17@5.[151,1436,181,1160]" captionTargetPageId="5" captionText="FIGURE 3. Lacronia utaru sp. nov., ♂ holotype (MHNCI). A: habitus, dorsal view. B: ventral view. C: lateral view. D: posterior view. E: Fe–Ta IV, prolateral view (left leg) and retrolateral view (right leg). Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13346969" httpUri="https://zenodo.org/record/13346969/files/figure.png" pageId="4" pageNumber="421">3B</figureCitation>
). Stigmatic area inverted-Y-shaped, clearly sunken in relation to Cx IV distal part (
<figureCitation id="130B2A7BFFE93253051EF8D0FE22F2FB" box="[359,457,1904,1930]" captionStart="FIGURE 2" captionStartId="4.[151,250,1546,1570]" captionTargetBox="[151,1436,181,1522]" captionTargetId="figure-19@4.[151,1436,181,1522]" captionTargetPageId="4" captionText="FIGURE 2. Habitus in vivo of Lacronia utaru sp. nov., ♂ paratype (MNRJ 3697), from Brazil, Paraná, Paranaguá, Estação Ecológica do Guaraguaçu. A: dorsal view. B: fronto-dorsal view. C: posterior view. D: dorso-posterior view. E: ventral view. Photographs by Adriano Kury." figureDoi="http://doi.org/10.5281/zenodo.13346967" httpUri="https://zenodo.org/record/13346967/files/figure.png" pageId="4" pageNumber="421">Figs. 2E</figureCitation>
, 
<figureCitation id="130B2A7BFFE9325305AFF8D0FE12F2FB" box="[470,505,1904,1930]" captionStart="FIGURE 3" captionStartId="5.[151,250,1184,1208]" captionTargetBox="[151,1436,181,1160]" captionTargetId="figure-17@5.[151,1436,181,1160]" captionTargetPageId="5" captionText="FIGURE 3. Lacronia utaru sp. nov., ♂ holotype (MHNCI). A: habitus, dorsal view. B: ventral view. C: lateral view. D: posterior view. E: Fe–Ta IV, prolateral view (left leg) and retrolateral view (right leg). Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13346969" httpUri="https://zenodo.org/record/13346969/files/figure.png" pageId="4" pageNumber="421">3B</figureCitation>
). Cx IV covered by ordinary tubercles. (
<figureCitation id="130B2A7BFFE9325307BFF8D0FBC9F2FB" box="[966,1058,1904,1930]" captionStart="FIGURE 2" captionStartId="4.[151,250,1546,1570]" captionTargetBox="[151,1436,181,1522]" captionTargetId="figure-19@4.[151,1436,181,1522]" captionTargetPageId="4" captionText="FIGURE 2. Habitus in vivo of Lacronia utaru sp. nov., ♂ paratype (MNRJ 3697), from Brazil, Paraná, Paranaguá, Estação Ecológica do Guaraguaçu. A: dorsal view. B: fronto-dorsal view. C: posterior view. D: dorso-posterior view. E: ventral view. Photographs by Adriano Kury." figureDoi="http://doi.org/10.5281/zenodo.13346967" httpUri="https://zenodo.org/record/13346967/files/figure.png" pageId="4" pageNumber="421">Figs 2E</figureCitation>
, 
<figureCitation id="130B2A7BFFE932530056F8D0FBB9F2FB" box="[1071,1106,1904,1930]" captionStart="FIGURE 3" captionStartId="5.[151,250,1184,1208]" captionTargetBox="[151,1436,181,1160]" captionTargetId="figure-17@5.[151,1436,181,1160]" captionTargetPageId="5" captionText="FIGURE 3. Lacronia utaru sp. nov., ♂ holotype (MHNCI). A: habitus, dorsal view. B: ventral view. C: lateral view. D: posterior view. E: Fe–Ta IV, prolateral view (left leg) and retrolateral view (right leg). Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13346969" httpUri="https://zenodo.org/record/13346969/files/figure.png" pageId="4" pageNumber="421">3B</figureCitation>
). Cx IV posterior border and stigmatic area each with a transversal row of ordinary tubercles (
<figureCitation id="130B2A7BFFE9325307F0F834FC0FF2DF" box="[905,996,1940,1966]" captionStart="FIGURE 2" captionStartId="4.[151,250,1546,1570]" captionTargetBox="[151,1436,181,1522]" captionTargetId="figure-19@4.[151,1436,181,1522]" captionTargetPageId="4" captionText="FIGURE 2. Habitus in vivo of Lacronia utaru sp. nov., ♂ paratype (MNRJ 3697), from Brazil, Paraná, Paranaguá, Estação Ecológica do Guaraguaçu. A: dorsal view. B: fronto-dorsal view. C: posterior view. D: dorso-posterior view. E: ventral view. Photographs by Adriano Kury." figureDoi="http://doi.org/10.5281/zenodo.13346967" httpUri="https://zenodo.org/record/13346967/files/figure.png" pageId="4" pageNumber="421">Fig. 2E</figureCitation>
). Stigmata visible (
<figureCitation id="130B2A7BFFE9325300BDF834FACAF2DF" box="[1220,1313,1940,1966]" captionStart="FIGURE 2" captionStartId="4.[151,250,1546,1570]" captionTargetBox="[151,1436,181,1522]" captionTargetId="figure-19@4.[151,1436,181,1522]" captionTargetPageId="4" captionText="FIGURE 2. Habitus in vivo of Lacronia utaru sp. nov., ♂ paratype (MNRJ 3697), from Brazil, Paraná, Paranaguá, Estação Ecológica do Guaraguaçu. A: dorsal view. B: fronto-dorsal view. C: posterior view. D: dorso-posterior view. E: ventral view. Photographs by Adriano Kury." figureDoi="http://doi.org/10.5281/zenodo.13346967" httpUri="https://zenodo.org/record/13346967/files/figure.png" pageId="4" pageNumber="421">Figs 2E</figureCitation>
, 
<figureCitation id="130B2A7BFFE932530149F834FAB8F2DF" box="[1328,1363,1940,1966]" captionStart="FIGURE 3" captionStartId="5.[151,250,1184,1208]" captionTargetBox="[151,1436,181,1160]" captionTargetId="figure-17@5.[151,1436,181,1160]" captionTargetPageId="5" captionText="FIGURE 3. Lacronia utaru sp. nov., ♂ holotype (MHNCI). A: habitus, dorsal view. B: ventral view. C: lateral view. D: posterior view. E: Fe–Ta IV, prolateral view (left leg) and retrolateral view (right leg). Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13346969" httpUri="https://zenodo.org/record/13346969/files/figure.png" pageId="4" pageNumber="421">3B</figureCitation>
). Free sternites with a transverse row of ordinary tubercles.
</paragraph>
<caption id="DF4F6676FFE8325204EEFB00FB2DF1AD" ID-DOI="http://doi.org/10.5281/zenodo.13346969" ID-Zenodo-Dep="13346969" httpUri="https://zenodo.org/record/13346969/files/figure.png" pageId="5" pageNumber="422" startId="5.[151,250,1184,1208]" targetBox="[151,1436,181,1160]" targetPageId="5" targetType="figure">
<paragraph id="8B8F36FEFFE8325204EEFB00FB2DF1AD" blockId="5.[151,1437,1184,1244]" pageId="5" pageNumber="422">
<emphasis id="B944EAECFFE8325204EEFB00FEFCF1C9" bold="true" box="[151,279,1184,1208]" pageId="5" pageNumber="422">FIGURE 3.</emphasis>
<taxonomicName id="4C304D7DFFE832520558FB01FE55F1C9" authorityName="Carvalho &amp; Kury" authorityYear="2024" box="[289,446,1185,1208]" class="Arachnida" family="Gonyleptidae" genus="Lacronia" kingdom="Animalia" order="Opiliones" pageId="5" pageNumber="422" phylum="Arthropoda" rank="species" species="utaru" status="sp. nov.">
<emphasis id="B944EAECFFE832520558FB01FE55F1C9" box="[289,446,1185,1208]" italics="true" pageId="5" pageNumber="422">Lacronia utaru</emphasis>
</taxonomicName>
<emphasis id="B944EAECFFE8325205B1FB00FDF7F1C9" bold="true" box="[456,540,1184,1208]" pageId="5" pageNumber="422">
<taxonomicNameLabel id="A2775797FFE8325205B1FB00FDF7F1C9" box="[456,540,1184,1208]" pageId="5" pageNumber="422" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, ♂ holotype (MHNCI). A: habitus, dorsal view. B: ventral view. C: lateral view. D: posterior view. E: Fe–Ta IV, prolateral view (left leg) and retrolateral view (right leg). Scale bars = 1 mm.
</paragraph>
</caption>
<paragraph id="8B8F36FEFFE8325204BEFA8EFC9DF01D" blockId="5.[151,1437,1326,2037]" pageId="5" pageNumber="422">
<emphasis id="B944EAECFFE8325204BEFA8EFEA1F039" box="[199,330,1326,1352]" italics="true" pageId="5" pageNumber="422">Chelicerae.</emphasis>
Basichelicerite elongate, bulla well-marked, with marginal setiferous tubercles (two–three mesal and posterior, two ectal) (
<figureCitation id="130B2A7BFFE8325205CCFAF2FDF3F01D" box="[437,536,1362,1388]" captionStart="FIGURE 2" captionStartId="4.[151,250,1546,1570]" captionTargetBox="[151,1436,181,1522]" captionTargetId="figure-19@4.[151,1436,181,1522]" captionTargetPageId="4" captionText="FIGURE 2. Habitus in vivo of Lacronia utaru sp. nov., ♂ paratype (MNRJ 3697), from Brazil, Paraná, Paranaguá, Estação Ecológica do Guaraguaçu. A: dorsal view. B: fronto-dorsal view. C: posterior view. D: dorso-posterior view. E: ventral view. Photographs by Adriano Kury." figureDoi="http://doi.org/10.5281/zenodo.13346967" httpUri="https://zenodo.org/record/13346967/files/figure.png" pageId="5" pageNumber="422">Figs. 2B</figureCitation>
, 
<figureCitation id="130B2A7BFFE83252065AFAF2FD82F01D" box="[547,617,1362,1388]" captionStart="FIGURE 3" captionStartId="5.[151,250,1184,1208]" captionTargetBox="[151,1436,181,1160]" captionTargetId="figure-17@5.[151,1436,181,1160]" captionTargetPageId="5" captionText="FIGURE 3. Lacronia utaru sp. nov., ♂ holotype (MHNCI). A: habitus, dorsal view. B: ventral view. C: lateral view. D: posterior view. E: Fe–Ta IV, prolateral view (left leg) and retrolateral view (right leg). Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13346969" httpUri="https://zenodo.org/record/13346969/files/figure.png" pageId="5" pageNumber="422">3A, C</figureCitation>
, 
<figureCitation id="130B2A7BFFE83252060DFAF2FD70F01D" box="[628,667,1362,1388]" captionStart="FIGURE 4" captionStartId="6.[151,250,1878,1902]" captionTargetBox="[151,1436,237,1855]" captionTargetId="figure-48@6.[151,1436,237,1855]" captionTargetPageId="6" captionText="FIGURE 4. Lacronia utaru sp. nov., ♂ holotype (MHNCI). A: dorsal scutum, chelicerae, trochanters and coxae IV, dorsal view. B: dorsal scutum, lateral view. C: ocularium, anterior view. D: right Fe–Ti III, retrolateral view. E: armature of mesotergal area III, posterior view. F: detail of the prodorsal apophysis of coxa IV, posterior view. G–J: right Tr–Ti IV, G: dorsal view. H: prolateral view. I: ventral view. J: retrolateral view. K: right Mt–Ta IV, retrolateral view. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13346971" httpUri="https://zenodo.org/record/13346971/files/figure.png" pageId="5" pageNumber="422">4A</figureCitation>
); hand not swollen.
</paragraph>
<paragraph id="8B8F36FEFFE8325204BEFAD6FBD9F0A9" blockId="5.[151,1437,1326,2037]" pageId="5" pageNumber="422">
<emphasis id="B944EAECFFE8325204BEFAD6FED4F0E1" box="[199,319,1398,1424]" italics="true" pageId="5" pageNumber="422">Pedipalps.</emphasis>
Tr with two geminate ventral setiferous tubercles. Fe with a ventral basal and a mesal apical setiferous tubercle. Pa unarmed. Ti with two rows of four spines (IiIi) on ventro-mesal and ventro-ectal faces. Ta with two rows of spines: three (Iii) on ventro-mesal face and four (IiIi) on ventro-ectal face.
</paragraph>
<paragraph id="8B8F36FEFFE8325204BEFA43FBA6F3A5" blockId="5.[151,1437,1326,2037]" pageId="5" pageNumber="422">
<emphasis id="B944EAECFFE8325204BEFA43FEEFF08D" box="[199,260,1507,1532]" italics="true" pageId="5" pageNumber="422">Legs.</emphasis>
All the unmentioned podomeres are unarmed or without relevant armature. Tr I–III each with several ventral tubercles (
<figureCitation id="130B2A7BFFE83252051BF9A6FE02F351" box="[354,489,1542,1568]" captionStart="FIGURE 3" captionStartId="5.[151,250,1184,1208]" captionTargetBox="[151,1436,181,1160]" captionTargetId="figure-17@5.[151,1436,181,1160]" captionTargetPageId="5" captionText="FIGURE 3. Lacronia utaru sp. nov., ♂ holotype (MHNCI). A: habitus, dorsal view. B: ventral view. C: lateral view. D: posterior view. E: Fe–Ta IV, prolateral view (left leg) and retrolateral view (right leg). Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13346969" httpUri="https://zenodo.org/record/13346969/files/figure.png" pageId="5" pageNumber="422">Figs. 3A–B</figureCitation>
). Fe I sub-straight (
<figureCitation id="130B2A7BFFE8325206BFF9A6FCC3F351" box="[710,808,1542,1568]" captionStart="FIGURE 2" captionStartId="4.[151,250,1546,1570]" captionTargetBox="[151,1436,181,1522]" captionTargetId="figure-19@4.[151,1436,181,1522]" captionTargetPageId="4" captionText="FIGURE 2. Habitus in vivo of Lacronia utaru sp. nov., ♂ paratype (MNRJ 3697), from Brazil, Paraná, Paranaguá, Estação Ecológica do Guaraguaçu. A: dorsal view. B: fronto-dorsal view. C: posterior view. D: dorso-posterior view. E: ventral view. Photographs by Adriano Kury." figureDoi="http://doi.org/10.5281/zenodo.13346967" httpUri="https://zenodo.org/record/13346967/files/figure.png" pageId="5" pageNumber="422">Figs. 2E</figureCitation>
, 
<figureCitation id="130B2A7BFFE83252074DF9A6FC97F351" box="[820,892,1542,1568]" captionStart="FIGURE 3" captionStartId="5.[151,250,1184,1208]" captionTargetBox="[151,1436,181,1160]" captionTargetId="figure-17@5.[151,1436,181,1160]" captionTargetPageId="5" captionText="FIGURE 3. Lacronia utaru sp. nov., ♂ holotype (MHNCI). A: habitus, dorsal view. B: ventral view. C: lateral view. D: posterior view. E: Fe–Ta IV, prolateral view (left leg) and retrolateral view (right leg). Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13346969" httpUri="https://zenodo.org/record/13346969/files/figure.png" pageId="5" pageNumber="422">3A–B</figureCitation>
); Fe II straight (
<figureCitation id="130B2A7BFFE83252004AF9A6FB5CF350" box="[1075,1207,1542,1569]" captionStart="FIGURE 2" captionStartId="4.[151,250,1546,1570]" captionTargetBox="[151,1436,181,1522]" captionTargetId="figure-19@4.[151,1436,181,1522]" captionTargetPageId="4" captionText="FIGURE 2. Habitus in vivo of Lacronia utaru sp. nov., ♂ paratype (MNRJ 3697), from Brazil, Paraná, Paranaguá, Estação Ecológica do Guaraguaçu. A: dorsal view. B: fronto-dorsal view. C: posterior view. D: dorso-posterior view. E: ventral view. Photographs by Adriano Kury." figureDoi="http://doi.org/10.5281/zenodo.13346967" httpUri="https://zenodo.org/record/13346967/files/figure.png" pageId="5" pageNumber="422">Figs. 2A, E</figureCitation>
, 
<figureCitation id="130B2A7BFFE8325200BDF9A6FB02F351" box="[1220,1257,1542,1568]" captionStart="FIGURE 3" captionStartId="5.[151,250,1184,1208]" captionTargetBox="[151,1436,181,1160]" captionTargetId="figure-17@5.[151,1436,181,1160]" captionTargetPageId="5" captionText="FIGURE 3. Lacronia utaru sp. nov., ♂ holotype (MHNCI). A: habitus, dorsal view. B: ventral view. C: lateral view. D: posterior view. E: Fe–Ta IV, prolateral view (left leg) and retrolateral view (right leg). Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13346969" httpUri="https://zenodo.org/record/13346969/files/figure.png" pageId="5" pageNumber="422">3A</figureCitation>
); Fe III sinuous (
<figureCitation id="130B2A7BFFE8325204E6F98AFEC3F334" box="[159,296,1578,1605]" captionStart="FIGURE 2" captionStartId="4.[151,250,1546,1570]" captionTargetBox="[151,1436,181,1522]" captionTargetId="figure-19@4.[151,1436,181,1522]" captionTargetPageId="4" captionText="FIGURE 2. Habitus in vivo of Lacronia utaru sp. nov., ♂ paratype (MNRJ 3697), from Brazil, Paraná, Paranaguá, Estação Ecológica do Guaraguaçu. A: dorsal view. B: fronto-dorsal view. C: posterior view. D: dorso-posterior view. E: ventral view. Photographs by Adriano Kury." figureDoi="http://doi.org/10.5281/zenodo.13346967" httpUri="https://zenodo.org/record/13346967/files/figure.png" pageId="5" pageNumber="422">Figs. 2A, E</figureCitation>
, 
<figureCitation id="130B2A7BFFE832520541F98AFE6BF334" box="[312,384,1578,1605]" captionStart="FIGURE 3" captionStartId="5.[151,250,1184,1208]" captionTargetBox="[151,1436,181,1160]" captionTargetId="figure-17@5.[151,1436,181,1160]" captionTargetPageId="5" captionText="FIGURE 3. Lacronia utaru sp. nov., ♂ holotype (MHNCI). A: habitus, dorsal view. B: ventral view. C: lateral view. D: posterior view. E: Fe–Ta IV, prolateral view (left leg) and retrolateral view (right leg). Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13346969" httpUri="https://zenodo.org/record/13346969/files/figure.png" pageId="5" pageNumber="422">3A, B</figureCitation>
, 
<figureCitation id="130B2A7BFFE8325205F6F98AFE5EF335" box="[399,437,1578,1604]" captionStart="FIGURE 4" captionStartId="6.[151,250,1878,1902]" captionTargetBox="[151,1436,237,1855]" captionTargetId="figure-48@6.[151,1436,237,1855]" captionTargetPageId="6" captionText="FIGURE 4. Lacronia utaru sp. nov., ♂ holotype (MHNCI). A: dorsal scutum, chelicerae, trochanters and coxae IV, dorsal view. B: dorsal scutum, lateral view. C: ocularium, anterior view. D: right Fe–Ti III, retrolateral view. E: armature of mesotergal area III, posterior view. F: detail of the prodorsal apophysis of coxa IV, posterior view. G–J: right Tr–Ti IV, G: dorsal view. H: prolateral view. I: ventral view. J: retrolateral view. K: right Mt–Ta IV, retrolateral view. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13346971" httpUri="https://zenodo.org/record/13346971/files/figure.png" pageId="5" pageNumber="422">4D</figureCitation>
). Fe and Ti I–III with all faces covered by longitudinal rows of small tubercles (
<figureCitation id="130B2A7BFFE83252011DF98AFF35F318" captionStart="FIGURE 3" captionStartId="5.[151,250,1184,1208]" captionTargetBox="[151,1436,181,1160]" captionTargetId="figure-17@5.[151,1436,181,1160]" captionTargetPageId="5" captionText="FIGURE 3. Lacronia utaru sp. nov., ♂ holotype (MHNCI). A: habitus, dorsal view. B: ventral view. C: lateral view. D: posterior view. E: Fe–Ta IV, prolateral view (left leg) and retrolateral view (right leg). Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13346969" httpUri="https://zenodo.org/record/13346969/files/figure.png" pageId="5" pageNumber="422">Figs. 3A, B</figureCitation>
, 
<figureCitation id="130B2A7BFFE832520492F9EEFEF9F319" box="[235,274,1614,1640]" captionStart="FIGURE 4" captionStartId="6.[151,250,1878,1902]" captionTargetBox="[151,1436,237,1855]" captionTargetId="figure-48@6.[151,1436,237,1855]" captionTargetPageId="6" captionText="FIGURE 4. Lacronia utaru sp. nov., ♂ holotype (MHNCI). A: dorsal scutum, chelicerae, trochanters and coxae IV, dorsal view. B: dorsal scutum, lateral view. C: ocularium, anterior view. D: right Fe–Ti III, retrolateral view. E: armature of mesotergal area III, posterior view. F: detail of the prodorsal apophysis of coxa IV, posterior view. G–J: right Tr–Ti IV, G: dorsal view. H: prolateral view. I: ventral view. J: retrolateral view. K: right Mt–Ta IV, retrolateral view. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13346971" httpUri="https://zenodo.org/record/13346971/files/figure.png" pageId="5" pageNumber="422">4D</figureCitation>
). Fe II–III with an apical retro-dorsal spur (
<figureCitation id="130B2A7BFFE83252077FF9EEFC67F318" box="[774,908,1614,1641]" captionStart="FIGURE 2" captionStartId="4.[151,250,1546,1570]" captionTargetBox="[151,1436,181,1522]" captionTargetId="figure-19@4.[151,1436,181,1522]" captionTargetPageId="4" captionText="FIGURE 2. Habitus in vivo of Lacronia utaru sp. nov., ♂ paratype (MNRJ 3697), from Brazil, Paraná, Paranaguá, Estação Ecológica do Guaraguaçu. A: dorsal view. B: fronto-dorsal view. C: posterior view. D: dorso-posterior view. E: ventral view. Photographs by Adriano Kury." figureDoi="http://doi.org/10.5281/zenodo.13346967" httpUri="https://zenodo.org/record/13346967/files/figure.png" pageId="5" pageNumber="422">Figs. 2A, E</figureCitation>
, 
<figureCitation id="130B2A7BFFE8325207E3F9EEFC55F319" box="[922,958,1614,1640]" captionStart="FIGURE 3" captionStartId="5.[151,250,1184,1208]" captionTargetBox="[151,1436,181,1160]" captionTargetId="figure-17@5.[151,1436,181,1160]" captionTargetPageId="5" captionText="FIGURE 3. Lacronia utaru sp. nov., ♂ holotype (MHNCI). A: habitus, dorsal view. B: ventral view. C: lateral view. D: posterior view. E: Fe–Ta IV, prolateral view (left leg) and retrolateral view (right leg). Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13346969" httpUri="https://zenodo.org/record/13346969/files/figure.png" pageId="5" pageNumber="422">3A</figureCitation>
, 
<figureCitation id="130B2A7BFFE8325207B5F9EEFC19F319" box="[972,1010,1614,1640]" captionStart="FIGURE 4" captionStartId="6.[151,250,1878,1902]" captionTargetBox="[151,1436,237,1855]" captionTargetId="figure-48@6.[151,1436,237,1855]" captionTargetPageId="6" captionText="FIGURE 4. Lacronia utaru sp. nov., ♂ holotype (MHNCI). A: dorsal scutum, chelicerae, trochanters and coxae IV, dorsal view. B: dorsal scutum, lateral view. C: ocularium, anterior view. D: right Fe–Ti III, retrolateral view. E: armature of mesotergal area III, posterior view. F: detail of the prodorsal apophysis of coxa IV, posterior view. G–J: right Tr–Ti IV, G: dorsal view. H: prolateral view. I: ventral view. J: retrolateral view. K: right Mt–Ta IV, retrolateral view. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13346971" httpUri="https://zenodo.org/record/13346971/files/figure.png" pageId="5" pageNumber="422">4D</figureCitation>
). Fe III with an apical prodorsal spur (much smaller than the retro-dorsal one) (
<figureCitation id="130B2A7BFFE832520612F9D2FD28F3FD" box="[619,707,1650,1676]" captionStart="FIGURE 3" captionStartId="5.[151,250,1184,1208]" captionTargetBox="[151,1436,181,1160]" captionTargetId="figure-17@5.[151,1436,181,1160]" captionTargetPageId="5" captionText="FIGURE 3. Lacronia utaru sp. nov., ♂ holotype (MHNCI). A: habitus, dorsal view. B: ventral view. C: lateral view. D: posterior view. E: Fe–Ta IV, prolateral view (left leg) and retrolateral view (right leg). Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13346969" httpUri="https://zenodo.org/record/13346969/files/figure.png" pageId="5" pageNumber="422">Fig. 3C</figureCitation>
). Fe III and Ti III with two rows (proventral and retro-ventral) of small acuminate tubercles, distally becoming spines (outstanding spines on Ti III) (
<figureCitation id="130B2A7BFFE832520044F936FB4BF3C1" box="[1085,1184,1686,1712]" captionStart="FIGURE 3" captionStartId="5.[151,250,1184,1208]" captionTargetBox="[151,1436,181,1160]" captionTargetId="figure-17@5.[151,1436,181,1160]" captionTargetPageId="5" captionText="FIGURE 3. Lacronia utaru sp. nov., ♂ holotype (MHNCI). A: habitus, dorsal view. B: ventral view. C: lateral view. D: posterior view. E: Fe–Ta IV, prolateral view (left leg) and retrolateral view (right leg). Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13346969" httpUri="https://zenodo.org/record/13346969/files/figure.png" pageId="5" pageNumber="422">Figs. 3B</figureCitation>
, 
<figureCitation id="130B2A7BFFE8325200D5F936FB39F3C1" box="[1196,1234,1686,1712]" captionStart="FIGURE 4" captionStartId="6.[151,250,1878,1902]" captionTargetBox="[151,1436,237,1855]" captionTargetId="figure-48@6.[151,1436,237,1855]" captionTargetPageId="6" captionText="FIGURE 4. Lacronia utaru sp. nov., ♂ holotype (MHNCI). A: dorsal scutum, chelicerae, trochanters and coxae IV, dorsal view. B: dorsal scutum, lateral view. C: ocularium, anterior view. D: right Fe–Ti III, retrolateral view. E: armature of mesotergal area III, posterior view. F: detail of the prodorsal apophysis of coxa IV, posterior view. G–J: right Tr–Ti IV, G: dorsal view. H: prolateral view. I: ventral view. J: retrolateral view. K: right Mt–Ta IV, retrolateral view. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13346971" httpUri="https://zenodo.org/record/13346971/files/figure.png" pageId="5" pageNumber="422">4D</figureCitation>
). Pa I–III covered dorsally by tubercles (
<figureCitation id="130B2A7BFFE8325205E9F91AFE1FF3A5" box="[400,500,1722,1748]" captionStart="FIGURE 2" captionStartId="4.[151,250,1546,1570]" captionTargetBox="[151,1436,181,1522]" captionTargetId="figure-19@4.[151,1436,181,1522]" captionTargetPageId="4" captionText="FIGURE 2. Habitus in vivo of Lacronia utaru sp. nov., ♂ paratype (MNRJ 3697), from Brazil, Paraná, Paranaguá, Estação Ecológica do Guaraguaçu. A: dorsal view. B: fronto-dorsal view. C: posterior view. D: dorso-posterior view. E: ventral view. Photographs by Adriano Kury." figureDoi="http://doi.org/10.5281/zenodo.13346967" httpUri="https://zenodo.org/record/13346967/files/figure.png" pageId="5" pageNumber="422">Figs. 2D</figureCitation>
, 
<figureCitation id="130B2A7BFFE832520586F91AFDCDF3A5" box="[511,550,1722,1748]" captionStart="FIGURE 4" captionStartId="6.[151,250,1878,1902]" captionTargetBox="[151,1436,237,1855]" captionTargetId="figure-48@6.[151,1436,237,1855]" captionTargetPageId="6" captionText="FIGURE 4. Lacronia utaru sp. nov., ♂ holotype (MHNCI). A: dorsal scutum, chelicerae, trochanters and coxae IV, dorsal view. B: dorsal scutum, lateral view. C: ocularium, anterior view. D: right Fe–Ti III, retrolateral view. E: armature of mesotergal area III, posterior view. F: detail of the prodorsal apophysis of coxa IV, posterior view. G–J: right Tr–Ti IV, G: dorsal view. H: prolateral view. I: ventral view. J: retrolateral view. K: right Mt–Ta IV, retrolateral view. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13346971" httpUri="https://zenodo.org/record/13346971/files/figure.png" pageId="5" pageNumber="422">4D</figureCitation>
). Ti III mace-shaped (
<figureCitation id="130B2A7BFFE832520765F91AFC56F3A4" box="[796,957,1722,1749]" captionStart="FIGURE 2" captionStartId="4.[151,250,1546,1570]" captionTargetBox="[151,1436,181,1522]" captionTargetId="figure-19@4.[151,1436,181,1522]" captionTargetPageId="4" captionText="FIGURE 2. Habitus in vivo of Lacronia utaru sp. nov., ♂ paratype (MNRJ 3697), from Brazil, Paraná, Paranaguá, Estação Ecológica do Guaraguaçu. A: dorsal view. B: fronto-dorsal view. C: posterior view. D: dorso-posterior view. E: ventral view. Photographs by Adriano Kury." figureDoi="http://doi.org/10.5281/zenodo.13346967" httpUri="https://zenodo.org/record/13346967/files/figure.png" pageId="5" pageNumber="422">Figs. 2B–C, E</figureCitation>
, 
<figureCitation id="130B2A7BFFE8325207B0F91AFBE4F3A5" box="[969,1039,1722,1748]" captionStart="FIGURE 3" captionStartId="5.[151,250,1184,1208]" captionTargetBox="[151,1436,181,1160]" captionTargetId="figure-17@5.[151,1436,181,1160]" captionTargetPageId="5" captionText="FIGURE 3. Lacronia utaru sp. nov., ♂ holotype (MHNCI). A: habitus, dorsal view. B: ventral view. C: lateral view. D: posterior view. E: Fe–Ta IV, prolateral view (left leg) and retrolateral view (right leg). Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13346969" httpUri="https://zenodo.org/record/13346969/files/figure.png" pageId="5" pageNumber="422">3A–B</figureCitation>
, 
<figureCitation id="130B2A7BFFE832520063F91AFBAAF3A5" box="[1050,1089,1722,1748]" captionStart="FIGURE 4" captionStartId="6.[151,250,1878,1902]" captionTargetBox="[151,1436,237,1855]" captionTargetId="figure-48@6.[151,1436,237,1855]" captionTargetPageId="6" captionText="FIGURE 4. Lacronia utaru sp. nov., ♂ holotype (MHNCI). A: dorsal scutum, chelicerae, trochanters and coxae IV, dorsal view. B: dorsal scutum, lateral view. C: ocularium, anterior view. D: right Fe–Ti III, retrolateral view. E: armature of mesotergal area III, posterior view. F: detail of the prodorsal apophysis of coxa IV, posterior view. G–J: right Tr–Ti IV, G: dorsal view. H: prolateral view. I: ventral view. J: retrolateral view. K: right Mt–Ta IV, retrolateral view. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13346971" httpUri="https://zenodo.org/record/13346971/files/figure.png" pageId="5" pageNumber="422">4D</figureCitation>
).
</paragraph>
<paragraph id="8B8F36FEFFE8325204BEF97EFE5CF2F9" blockId="5.[151,1437,1326,2037]" pageId="5" pageNumber="422">
Coxa IV reaching the level of the mesotergum areas III–IV of DS (
<figureCitation id="130B2A7BFFE8325207CEF97EFBFFF389" box="[951,1044,1758,1784]" captionStart="FIGURE 2" captionStartId="4.[151,250,1546,1570]" captionTargetBox="[151,1436,181,1522]" captionTargetId="figure-19@4.[151,1436,181,1522]" captionTargetPageId="4" captionText="FIGURE 2. Habitus in vivo of Lacronia utaru sp. nov., ♂ paratype (MNRJ 3697), from Brazil, Paraná, Paranaguá, Estação Ecológica do Guaraguaçu. A: dorsal view. B: fronto-dorsal view. C: posterior view. D: dorso-posterior view. E: ventral view. Photographs by Adriano Kury." figureDoi="http://doi.org/10.5281/zenodo.13346967" httpUri="https://zenodo.org/record/13346967/files/figure.png" pageId="5" pageNumber="422">Figs 2A</figureCitation>
, 
<figureCitation id="130B2A7BFFE832520058F97EFBAEF389" box="[1057,1093,1758,1784]" captionStart="FIGURE 3" captionStartId="5.[151,250,1184,1208]" captionTargetBox="[151,1436,181,1160]" captionTargetId="figure-17@5.[151,1436,181,1160]" captionTargetPageId="5" captionText="FIGURE 3. Lacronia utaru sp. nov., ♂ holotype (MHNCI). A: habitus, dorsal view. B: ventral view. C: lateral view. D: posterior view. E: Fe–Ta IV, prolateral view (left leg) and retrolateral view (right leg). Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13346969" httpUri="https://zenodo.org/record/13346969/files/figure.png" pageId="5" pageNumber="422">3A</figureCitation>
, 
<figureCitation id="130B2A7BFFE832520028F97EFB9CF389" box="[1105,1143,1758,1784]" captionStart="FIGURE 4" captionStartId="6.[151,250,1878,1902]" captionTargetBox="[151,1436,237,1855]" captionTargetId="figure-48@6.[151,1436,237,1855]" captionTargetPageId="6" captionText="FIGURE 4. Lacronia utaru sp. nov., ♂ holotype (MHNCI). A: dorsal scutum, chelicerae, trochanters and coxae IV, dorsal view. B: dorsal scutum, lateral view. C: ocularium, anterior view. D: right Fe–Ti III, retrolateral view. E: armature of mesotergal area III, posterior view. F: detail of the prodorsal apophysis of coxa IV, posterior view. G–J: right Tr–Ti IV, G: dorsal view. H: prolateral view. I: ventral view. J: retrolateral view. K: right Mt–Ta IV, retrolateral view. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13346971" httpUri="https://zenodo.org/record/13346971/files/figure.png" pageId="5" pageNumber="422">4A</figureCitation>
); its prodorsal and ventral faces are covered by a mix of ordinary and prominent acuminated tubercles (
<figureCitation id="130B2A7BFFE83252005CF8A2FB44F26C" box="[1061,1199,1794,1821]" captionStart="FIGURE 2" captionStartId="4.[151,250,1546,1570]" captionTargetBox="[151,1436,181,1522]" captionTargetId="figure-19@4.[151,1436,181,1522]" captionTargetPageId="4" captionText="FIGURE 2. Habitus in vivo of Lacronia utaru sp. nov., ♂ paratype (MNRJ 3697), from Brazil, Paraná, Paranaguá, Estação Ecológica do Guaraguaçu. A: dorsal view. B: fronto-dorsal view. C: posterior view. D: dorso-posterior view. E: ventral view. Photographs by Adriano Kury." figureDoi="http://doi.org/10.5281/zenodo.13346967" httpUri="https://zenodo.org/record/13346967/files/figure.png" pageId="5" pageNumber="422">Figs. 2A, E</figureCitation>
, 
<figureCitation id="130B2A7BFFE8325200B9F8A2FAEEF26D" box="[1216,1285,1794,1820]" captionStart="FIGURE 3" captionStartId="5.[151,250,1184,1208]" captionTargetBox="[151,1436,181,1160]" captionTargetId="figure-17@5.[151,1436,181,1160]" captionTargetPageId="5" captionText="FIGURE 3. Lacronia utaru sp. nov., ♂ holotype (MHNCI). A: habitus, dorsal view. B: ventral view. C: lateral view. D: posterior view. E: Fe–Ta IV, prolateral view (left leg) and retrolateral view (right leg). Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13346969" httpUri="https://zenodo.org/record/13346969/files/figure.png" pageId="5" pageNumber="422">3A–C</figureCitation>
, 
<figureCitation id="130B2A7BFFE83252016DF8A2FAD0F26D" box="[1300,1339,1794,1820]" captionStart="FIGURE 4" captionStartId="6.[151,250,1878,1902]" captionTargetBox="[151,1436,237,1855]" captionTargetId="figure-48@6.[151,1436,237,1855]" captionTargetPageId="6" captionText="FIGURE 4. Lacronia utaru sp. nov., ♂ holotype (MHNCI). A: dorsal scutum, chelicerae, trochanters and coxae IV, dorsal view. B: dorsal scutum, lateral view. C: ocularium, anterior view. D: right Fe–Ti III, retrolateral view. E: armature of mesotergal area III, posterior view. F: detail of the prodorsal apophysis of coxa IV, posterior view. G–J: right Tr–Ti IV, G: dorsal view. H: prolateral view. I: ventral view. J: retrolateral view. K: right Mt–Ta IV, retrolateral view. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13346971" httpUri="https://zenodo.org/record/13346971/files/figure.png" pageId="5" pageNumber="422">4A</figureCitation>
); with a prodorsal distal thick cylindrical apophysis (slightly curved backwards in the distal fourth), posteriorly crenated (
<figureCitation id="130B2A7BFFE8325204E6F8EAFEC3F215" box="[159,296,1866,1892]" captionStart="FIGURE 3" captionStartId="5.[151,250,1184,1208]" captionTargetBox="[151,1436,181,1160]" captionTargetId="figure-17@5.[151,1436,181,1160]" captionTargetPageId="5" captionText="FIGURE 3. Lacronia utaru sp. nov., ♂ holotype (MHNCI). A: habitus, dorsal view. B: ventral view. C: lateral view. D: posterior view. E: Fe–Ta IV, prolateral view (left leg) and retrolateral view (right leg). Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13346969" httpUri="https://zenodo.org/record/13346969/files/figure.png" pageId="5" pageNumber="422">Figs. 3A–D</figureCitation>
, 
<figureCitation id="130B2A7BFFE83252054CF8EAFE78F215" box="[309,403,1866,1892]" captionStart="FIGURE 4" captionStartId="6.[151,250,1878,1902]" captionTargetBox="[151,1436,237,1855]" captionTargetId="figure-48@6.[151,1436,237,1855]" captionTargetPageId="6" captionText="FIGURE 4. Lacronia utaru sp. nov., ♂ holotype (MHNCI). A: dorsal scutum, chelicerae, trochanters and coxae IV, dorsal view. B: dorsal scutum, lateral view. C: ocularium, anterior view. D: right Fe–Ti III, retrolateral view. E: armature of mesotergal area III, posterior view. F: detail of the prodorsal apophysis of coxa IV, posterior view. G–J: right Tr–Ti IV, G: dorsal view. H: prolateral view. I: ventral view. J: retrolateral view. K: right Mt–Ta IV, retrolateral view. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13346971" httpUri="https://zenodo.org/record/13346971/files/figure.png" pageId="5" pageNumber="422">4A, F–I</figureCitation>
); a retrolateral distal spiniform apophysis, fused with a small secondary branch (
<figureCitation id="130B2A7BFFE83252014BF8EAFF26F2F8" captionStart="FIGURE 2" captionStartId="4.[151,250,1546,1570]" captionTargetBox="[151,1436,181,1522]" captionTargetId="figure-19@4.[151,1436,181,1522]" captionTargetPageId="4" captionText="FIGURE 2. Habitus in vivo of Lacronia utaru sp. nov., ♂ paratype (MNRJ 3697), from Brazil, Paraná, Paranaguá, Estação Ecológica do Guaraguaçu. A: dorsal view. B: fronto-dorsal view. C: posterior view. D: dorso-posterior view. E: ventral view. Photographs by Adriano Kury." figureDoi="http://doi.org/10.5281/zenodo.13346967" httpUri="https://zenodo.org/record/13346967/files/figure.png" pageId="5" pageNumber="422">Figs. 2A, D–E</figureCitation>
, 
<figureCitation id="130B2A7BFFE8325204A0F8CEFEABF2F8" box="[217,320,1902,1929]" captionStart="FIGURE 3" captionStartId="5.[151,250,1184,1208]" captionTargetBox="[151,1436,181,1160]" captionTargetId="figure-17@5.[151,1436,181,1160]" captionTargetPageId="5" captionText="FIGURE 3. Lacronia utaru sp. nov., ♂ holotype (MHNCI). A: habitus, dorsal view. B: ventral view. C: lateral view. D: posterior view. E: Fe–Ta IV, prolateral view (left leg) and retrolateral view (right leg). Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13346969" httpUri="https://zenodo.org/record/13346969/files/figure.png" pageId="5" pageNumber="422">3A–B, D</figureCitation>
, 
<figureCitation id="130B2A7BFFE832520535F8CEFE42F2F9" box="[332,425,1902,1928]" captionStart="FIGURE 4" captionStartId="6.[151,250,1878,1902]" captionTargetBox="[151,1436,237,1855]" captionTargetId="figure-48@6.[151,1436,237,1855]" captionTargetPageId="6" captionText="FIGURE 4. Lacronia utaru sp. nov., ♂ holotype (MHNCI). A: dorsal scutum, chelicerae, trochanters and coxae IV, dorsal view. B: dorsal scutum, lateral view. C: ocularium, anterior view. D: right Fe–Ti III, retrolateral view. E: armature of mesotergal area III, posterior view. F: detail of the prodorsal apophysis of coxa IV, posterior view. G–J: right Tr–Ti IV, G: dorsal view. H: prolateral view. I: ventral view. J: retrolateral view. K: right Mt–Ta IV, retrolateral view. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13346971" httpUri="https://zenodo.org/record/13346971/files/figure.png" pageId="5" pageNumber="422">4A, G, I</figureCitation>
).
</paragraph>
<paragraph id="8B8F36FEFFE8325104BEF832FC2BF5A4" blockId="5.[151,1437,1326,2037]" lastBlockId="6.[151,1436,151,213]" lastPageId="6" lastPageNumber="423" pageId="5" pageNumber="422">
Trochanter IV square-shaped in dorsal view (
<figureCitation id="130B2A7BFFE8325206CCF832FCFCF2DD" box="[693,791,1938,1964]" captionStart="FIGURE 2" captionStartId="4.[151,250,1546,1570]" captionTargetBox="[151,1436,181,1522]" captionTargetId="figure-19@4.[151,1436,181,1522]" captionTargetPageId="4" captionText="FIGURE 2. Habitus in vivo of Lacronia utaru sp. nov., ♂ paratype (MNRJ 3697), from Brazil, Paraná, Paranaguá, Estação Ecológica do Guaraguaçu. A: dorsal view. B: fronto-dorsal view. C: posterior view. D: dorso-posterior view. E: ventral view. Photographs by Adriano Kury." figureDoi="http://doi.org/10.5281/zenodo.13346967" httpUri="https://zenodo.org/record/13346967/files/figure.png" pageId="5" pageNumber="422">Figs. 2A</figureCitation>
, 
<figureCitation id="130B2A7BFFE832520758F832FCAEF2DD" box="[801,837,1938,1964]" captionStart="FIGURE 3" captionStartId="5.[151,250,1184,1208]" captionTargetBox="[151,1436,181,1160]" captionTargetId="figure-17@5.[151,1436,181,1160]" captionTargetPageId="5" captionText="FIGURE 3. Lacronia utaru sp. nov., ♂ holotype (MHNCI). A: habitus, dorsal view. B: ventral view. C: lateral view. D: posterior view. E: Fe–Ta IV, prolateral view (left leg) and retrolateral view (right leg). Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13346969" httpUri="https://zenodo.org/record/13346969/files/figure.png" pageId="5" pageNumber="422">3A</figureCitation>
, 
<figureCitation id="130B2A7BFFE832520736F832FC7EF2DD" box="[847,917,1938,1964]" captionStart="FIGURE 4" captionStartId="6.[151,250,1878,1902]" captionTargetBox="[151,1436,237,1855]" captionTargetId="figure-48@6.[151,1436,237,1855]" captionTargetPageId="6" captionText="FIGURE 4. Lacronia utaru sp. nov., ♂ holotype (MHNCI). A: dorsal scutum, chelicerae, trochanters and coxae IV, dorsal view. B: dorsal scutum, lateral view. C: ocularium, anterior view. D: right Fe–Ti III, retrolateral view. E: armature of mesotergal area III, posterior view. F: detail of the prodorsal apophysis of coxa IV, posterior view. G–J: right Tr–Ti IV, G: dorsal view. H: prolateral view. I: ventral view. J: retrolateral view. K: right Mt–Ta IV, retrolateral view. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13346971" httpUri="https://zenodo.org/record/13346971/files/figure.png" pageId="5" pageNumber="422">4A, G</figureCitation>
); prodorsal distal face with three-four tubercles (two prolateral most prominent) (
<figureCitation id="130B2A7BFFE832520666F816FD25F2A1" box="[543,718,1974,2000]" captionStart="FIGURE 4" captionStartId="6.[151,250,1878,1902]" captionTargetBox="[151,1436,237,1855]" captionTargetId="figure-48@6.[151,1436,237,1855]" captionTargetPageId="6" captionText="FIGURE 4. Lacronia utaru sp. nov., ♂ holotype (MHNCI). A: dorsal scutum, chelicerae, trochanters and coxae IV, dorsal view. B: dorsal scutum, lateral view. C: ocularium, anterior view. D: right Fe–Ti III, retrolateral view. E: armature of mesotergal area III, posterior view. F: detail of the prodorsal apophysis of coxa IV, posterior view. G–J: right Tr–Ti IV, G: dorsal view. H: prolateral view. I: ventral view. J: retrolateral view. K: right Mt–Ta IV, retrolateral view. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13346971" httpUri="https://zenodo.org/record/13346971/files/figure.png" pageId="5" pageNumber="422">Figs. 4A, F–G</figureCitation>
); prolateral proximal/central portion with an almost isosceles triangle-shaped apophysis, with a prodorsal basal protuberance (
<figureCitation id="130B2A7BFFE83252070DF87AFBE7F285" box="[884,1036,2010,2036]" captionStart="FIGURE 4" captionStartId="6.[151,250,1878,1902]" captionTargetBox="[151,1436,237,1855]" captionTargetId="figure-48@6.[151,1436,237,1855]" captionTargetPageId="6" captionText="FIGURE 4. Lacronia utaru sp. nov., ♂ holotype (MHNCI). A: dorsal scutum, chelicerae, trochanters and coxae IV, dorsal view. B: dorsal scutum, lateral view. C: ocularium, anterior view. D: right Fe–Ti III, retrolateral view. E: armature of mesotergal area III, posterior view. F: detail of the prodorsal apophysis of coxa IV, posterior view. G–J: right Tr–Ti IV, G: dorsal view. H: prolateral view. I: ventral view. J: retrolateral view. K: right Mt–Ta IV, retrolateral view. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13346971" httpUri="https://zenodo.org/record/13346971/files/figure.png" pageId="5" pageNumber="422">Fig. 4A, G–I</figureCitation>
); ventral face tuberculate (
<figureCitation id="130B2A7BFFE832520145F87AFA73F285" box="[1340,1432,2010,2036]" captionStart="FIGURE 2" captionStartId="4.[151,250,1546,1570]" captionTargetBox="[151,1436,181,1522]" captionTargetId="figure-19@4.[151,1436,181,1522]" captionTargetPageId="4" captionText="FIGURE 2. Habitus in vivo of Lacronia utaru sp. nov., ♂ paratype (MNRJ 3697), from Brazil, Paraná, Paranaguá, Estação Ecológica do Guaraguaçu. A: dorsal view. B: fronto-dorsal view. C: posterior view. D: dorso-posterior view. E: ventral view. Photographs by Adriano Kury." figureDoi="http://doi.org/10.5281/zenodo.13346967" httpUri="https://zenodo.org/record/13346967/files/figure.png" pageId="5" pageNumber="422">Figs 2E</figureCitation>
, 
<figureCitation id="130B2A7BFFEB325104EEFF37FF51F5C0" box="[151,186,151,177]" captionStart="FIGURE 3" captionStartId="5.[151,250,1184,1208]" captionTargetBox="[151,1436,181,1160]" captionTargetId="figure-17@5.[151,1436,181,1160]" captionTargetPageId="5" captionText="FIGURE 3. Lacronia utaru sp. nov., ♂ holotype (MHNCI). A: habitus, dorsal view. B: ventral view. C: lateral view. D: posterior view. E: Fe–Ta IV, prolateral view (left leg) and retrolateral view (right leg). Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13346969" httpUri="https://zenodo.org/record/13346969/files/figure.png" pageId="6" pageNumber="423">3B</figureCitation>
, 
<figureCitation id="130B2A7BFFEB325104BEFF37FEEDF5C0" box="[199,262,151,177]" captionStart="FIGURE 4" captionStartId="6.[151,250,1878,1902]" captionTargetBox="[151,1436,237,1855]" captionTargetId="figure-48@6.[151,1436,237,1855]" captionTargetPageId="6" captionText="FIGURE 4. Lacronia utaru sp. nov., ♂ holotype (MHNCI). A: dorsal scutum, chelicerae, trochanters and coxae IV, dorsal view. B: dorsal scutum, lateral view. C: ocularium, anterior view. D: right Fe–Ti III, retrolateral view. E: armature of mesotergal area III, posterior view. F: detail of the prodorsal apophysis of coxa IV, posterior view. G–J: right Tr–Ti IV, G: dorsal view. H: prolateral view. I: ventral view. J: retrolateral view. K: right Mt–Ta IV, retrolateral view. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13346971" httpUri="https://zenodo.org/record/13346971/files/figure.png" pageId="6" pageNumber="423">4H–J</figureCitation>
), retro-lateral face with a proximal conical apophysis (slightly curved dorsad on the distal portion), and with a reduced distal subconical apophysis (
<figureCitation id="130B2A7BFFEB325106FDFF1BFD34F5A4" box="[644,735,187,213]" captionStart="FIGURE 2" captionStartId="4.[151,250,1546,1570]" captionTargetBox="[151,1436,181,1522]" captionTargetId="figure-19@4.[151,1436,181,1522]" captionTargetPageId="4" captionText="FIGURE 2. Habitus in vivo of Lacronia utaru sp. nov., ♂ paratype (MNRJ 3697), from Brazil, Paraná, Paranaguá, Estação Ecológica do Guaraguaçu. A: dorsal view. B: fronto-dorsal view. C: posterior view. D: dorso-posterior view. E: ventral view. Photographs by Adriano Kury." figureDoi="http://doi.org/10.5281/zenodo.13346967" httpUri="https://zenodo.org/record/13346967/files/figure.png" pageId="6" pageNumber="423">Figs 2E</figureCitation>
, 
<figureCitation id="130B2A7BFFEB32510693FF1BFCDBF5A4" box="[746,816,187,213]" captionStart="FIGURE 3" captionStartId="5.[151,250,1184,1208]" captionTargetBox="[151,1436,181,1160]" captionTargetId="figure-17@5.[151,1436,181,1160]" captionTargetPageId="5" captionText="FIGURE 3. Lacronia utaru sp. nov., ♂ holotype (MHNCI). A: habitus, dorsal view. B: ventral view. C: lateral view. D: posterior view. E: Fe–Ta IV, prolateral view (left leg) and retrolateral view (right leg). Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13346969" httpUri="https://zenodo.org/record/13346969/files/figure.png" pageId="6" pageNumber="423">3A–B</figureCitation>
, 
<figureCitation id="130B2A7BFFEB32510742FF1BFC5FF5A4" box="[827,948,187,213]" captionStart="FIGURE 4" captionStartId="6.[151,250,1878,1902]" captionTargetBox="[151,1436,237,1855]" captionTargetId="figure-48@6.[151,1436,237,1855]" captionTargetPageId="6" captionText="FIGURE 4. Lacronia utaru sp. nov., ♂ holotype (MHNCI). A: dorsal scutum, chelicerae, trochanters and coxae IV, dorsal view. B: dorsal scutum, lateral view. C: ocularium, anterior view. D: right Fe–Ti III, retrolateral view. E: armature of mesotergal area III, posterior view. F: detail of the prodorsal apophysis of coxa IV, posterior view. G–J: right Tr–Ti IV, G: dorsal view. H: prolateral view. I: ventral view. J: retrolateral view. K: right Mt–Ta IV, retrolateral view. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13346971" httpUri="https://zenodo.org/record/13346971/files/figure.png" pageId="6" pageNumber="423">4A, G, I–J</figureCitation>
).
</paragraph>
<caption id="DF4F6676FFEB325104EEF8F6FB37F2AB" ID-DOI="http://doi.org/10.5281/zenodo.13346971" ID-Zenodo-Dep="13346971" httpUri="https://zenodo.org/record/13346971/files/figure.png" pageId="6" pageNumber="423" startId="6.[151,250,1878,1902]" targetBox="[151,1436,237,1855]" targetPageId="6" targetType="figure">
<paragraph id="8B8F36FEFFEB325104EEF8F6FB37F2AB" blockId="6.[151,1437,1878,2011]" pageId="6" pageNumber="423">
<emphasis id="B944EAECFFEB325104EEF8F6FEFEF21F" bold="true" box="[151,277,1878,1902]" pageId="6" pageNumber="423">FIGURE 4.</emphasis>
<taxonomicName id="4C304D7DFFEB32510567F8F7FE51F21F" authorityName="Carvalho &amp; Kury" authorityYear="2024" box="[286,442,1879,1902]" class="Arachnida" family="Gonyleptidae" genus="Lacronia" kingdom="Animalia" order="Opiliones" pageId="6" pageNumber="423" phylum="Arthropoda" rank="species" species="utaru" status="sp. nov.">
<emphasis id="B944EAECFFEB32510567F8F7FE51F21F" box="[286,442,1879,1902]" italics="true" pageId="6" pageNumber="423">Lacronia utaru</emphasis>
</taxonomicName>
<emphasis id="B944EAECFFEB325105BAF8F7FDFDF21E" bold="true" box="[451,534,1879,1903]" pageId="6" pageNumber="423">
<taxonomicNameLabel id="A2775797FFEB325105BAF8F7FDFDF21E" box="[451,534,1879,1903]" pageId="6" pageNumber="423" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, ♂ holotype (MHNCI). A: dorsal scutum, chelicerae, trochanters and coxae IV, dorsal view. B: dorsal scutum, lateral view. C: ocularium, anterior view. D: right Fe–Ti III, retrolateral view. E: armature of mesotergal area III, posterior view. F: detail of the prodorsal apophysis of coxa IV, posterior view. G–J: right Tr–Ti IV, G: dorsal view. H: prolateral view. I: ventral view. J: retrolateral view. K: right Mt–Ta IV, retrolateral view. Scale bars = 1 mm.
</paragraph>
</caption>
<paragraph id="8B8F36FEFFEA325004BEFF37FECFF710" blockId="7.[151,1437,151,1869]" pageId="7" pageNumber="424">
Femur IV straight in dorsal view and swollen at the centro-distal thirds (
<figureCitation id="130B2A7BFFEA3250078BFF37FB9EF5C0" box="[1010,1141,151,177]" captionStart="FIGURE 2" captionStartId="4.[151,250,1546,1570]" captionTargetBox="[151,1436,181,1522]" captionTargetId="figure-19@4.[151,1436,181,1522]" captionTargetPageId="4" captionText="FIGURE 2. Habitus in vivo of Lacronia utaru sp. nov., ♂ paratype (MNRJ 3697), from Brazil, Paraná, Paranaguá, Estação Ecológica do Guaraguaçu. A: dorsal view. B: fronto-dorsal view. C: posterior view. D: dorso-posterior view. E: ventral view. Photographs by Adriano Kury." figureDoi="http://doi.org/10.5281/zenodo.13346967" httpUri="https://zenodo.org/record/13346967/files/figure.png" pageId="7" pageNumber="424">Figs. 2A–E</figureCitation>
, 
<figureCitation id="130B2A7BFFEA325000F8FF37FB0EF5C0" box="[1153,1253,151,177]" captionStart="FIGURE 3" captionStartId="5.[151,250,1184,1208]" captionTargetBox="[151,1436,181,1160]" captionTargetId="figure-17@5.[151,1436,181,1160]" captionTargetPageId="5" captionText="FIGURE 3. Lacronia utaru sp. nov., ♂ holotype (MHNCI). A: habitus, dorsal view. B: ventral view. C: lateral view. D: posterior view. E: Fe–Ta IV, prolateral view (left leg) and retrolateral view (right leg). Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13346969" httpUri="https://zenodo.org/record/13346969/files/figure.png" pageId="7" pageNumber="424">3A–C, E</figureCitation>
, 
<figureCitation id="130B2A7BFFEA32500088FF37FADBF5C0" box="[1265,1328,151,177]" captionStart="FIGURE 4" captionStartId="6.[151,250,1878,1902]" captionTargetBox="[151,1436,237,1855]" captionTargetId="figure-48@6.[151,1436,237,1855]" captionTargetPageId="6" captionText="FIGURE 4. Lacronia utaru sp. nov., ♂ holotype (MHNCI). A: dorsal scutum, chelicerae, trochanters and coxae IV, dorsal view. B: dorsal scutum, lateral view. C: ocularium, anterior view. D: right Fe–Ti III, retrolateral view. E: armature of mesotergal area III, posterior view. F: detail of the prodorsal apophysis of coxa IV, posterior view. G–J: right Tr–Ti IV, G: dorsal view. H: prolateral view. I: ventral view. J: retrolateral view. K: right Mt–Ta IV, retrolateral view. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13346971" httpUri="https://zenodo.org/record/13346971/files/figure.png" pageId="7" pageNumber="424">4G–J</figureCitation>
); a dorsal row of nine prominent subconical tubercles and conical spine, with a short apical spur (
<figureCitation id="130B2A7BFFEA32500006FF1BFAF0F5A4" box="[1151,1307,187,213]" captionStart="FIGURE 4" captionStartId="6.[151,250,1878,1902]" captionTargetBox="[151,1436,237,1855]" captionTargetId="figure-48@6.[151,1436,237,1855]" captionTargetPageId="6" captionText="FIGURE 4. Lacronia utaru sp. nov., ♂ holotype (MHNCI). A: dorsal scutum, chelicerae, trochanters and coxae IV, dorsal view. B: dorsal scutum, lateral view. C: ocularium, anterior view. D: right Fe–Ti III, retrolateral view. E: armature of mesotergal area III, posterior view. F: detail of the prodorsal apophysis of coxa IV, posterior view. G–J: right Tr–Ti IV, G: dorsal view. H: prolateral view. I: ventral view. J: retrolateral view. K: right Mt–Ta IV, retrolateral view. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13346971" httpUri="https://zenodo.org/record/13346971/files/figure.png" pageId="7" pageNumber="424">Figs 4G–H, J</figureCitation>
); prodorsal face with a row of 16–17 prominent subconical tubercles (
<figureCitation id="130B2A7BFFEA32500748FF7FFC51F588" box="[817,954,223,249]" captionStart="FIGURE 4" captionStartId="6.[151,250,1878,1902]" captionTargetBox="[151,1436,237,1855]" captionTargetId="figure-48@6.[151,1436,237,1855]" captionTargetPageId="6" captionText="FIGURE 4. Lacronia utaru sp. nov., ♂ holotype (MHNCI). A: dorsal scutum, chelicerae, trochanters and coxae IV, dorsal view. B: dorsal scutum, lateral view. C: ocularium, anterior view. D: right Fe–Ti III, retrolateral view. E: armature of mesotergal area III, posterior view. F: detail of the prodorsal apophysis of coxa IV, posterior view. G–J: right Tr–Ti IV, G: dorsal view. H: prolateral view. I: ventral view. J: retrolateral view. K: right Mt–Ta IV, retrolateral view. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13346971" httpUri="https://zenodo.org/record/13346971/files/figure.png" pageId="7" pageNumber="424">Figs. 4G–H</figureCitation>
); prolateral face with a row of 14–15 subconical prominent tubercles (
<figureCitation id="130B2A7BFFEA3250059AFEA3FD88F46C" box="[483,611,259,285]" captionStart="FIGURE 4" captionStartId="6.[151,250,1878,1902]" captionTargetBox="[151,1436,237,1855]" captionTargetId="figure-48@6.[151,1436,237,1855]" captionTargetPageId="6" captionText="FIGURE 4. Lacronia utaru sp. nov., ♂ holotype (MHNCI). A: dorsal scutum, chelicerae, trochanters and coxae IV, dorsal view. B: dorsal scutum, lateral view. C: ocularium, anterior view. D: right Fe–Ti III, retrolateral view. E: armature of mesotergal area III, posterior view. F: detail of the prodorsal apophysis of coxa IV, posterior view. G–J: right Tr–Ti IV, G: dorsal view. H: prolateral view. I: ventral view. J: retrolateral view. K: right Mt–Ta IV, retrolateral view. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13346971" httpUri="https://zenodo.org/record/13346971/files/figure.png" pageId="7" pageNumber="424">Figs. 4G–I</figureCitation>
); proventral face with a row of seven sub-conical tubercles on proximal half, and eight prominent conical tubercles on distal half (
<figureCitation id="130B2A7BFFEA32500744FE87FC5CF430" box="[829,951,295,321]" captionStart="FIGURE 4" captionStartId="6.[151,250,1878,1902]" captionTargetBox="[151,1436,237,1855]" captionTargetId="figure-48@6.[151,1436,237,1855]" captionTargetPageId="6" captionText="FIGURE 4. Lacronia utaru sp. nov., ♂ holotype (MHNCI). A: dorsal scutum, chelicerae, trochanters and coxae IV, dorsal view. B: dorsal scutum, lateral view. C: ocularium, anterior view. D: right Fe–Ti III, retrolateral view. E: armature of mesotergal area III, posterior view. F: detail of the prodorsal apophysis of coxa IV, posterior view. G–J: right Tr–Ti IV, G: dorsal view. H: prolateral view. I: ventral view. J: retrolateral view. K: right Mt–Ta IV, retrolateral view. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13346971" httpUri="https://zenodo.org/record/13346971/files/figure.png" pageId="7" pageNumber="424">Figs 4H–I</figureCitation>
); ventral face with three–four sub-conical tubercles on proximal fifth (
<figureCitation id="130B2A7BFFEA325005AEFEEBFDC7F414" box="[471,556,331,357]" captionStart="FIGURE 4" captionStartId="6.[151,250,1878,1902]" captionTargetBox="[151,1436,237,1855]" captionTargetId="figure-48@6.[151,1436,237,1855]" captionTargetPageId="6" captionText="FIGURE 4. Lacronia utaru sp. nov., ♂ holotype (MHNCI). A: dorsal scutum, chelicerae, trochanters and coxae IV, dorsal view. B: dorsal scutum, lateral view. C: ocularium, anterior view. D: right Fe–Ti III, retrolateral view. E: armature of mesotergal area III, posterior view. F: detail of the prodorsal apophysis of coxa IV, posterior view. G–J: right Tr–Ti IV, G: dorsal view. H: prolateral view. I: ventral view. J: retrolateral view. K: right Mt–Ta IV, retrolateral view. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13346971" httpUri="https://zenodo.org/record/13346971/files/figure.png" pageId="7" pageNumber="424">Figs 4I</figureCitation>
); retro-ventral face with a row of nine subconical tubercles on proximal half, and six prominent conical tubercles on distal half (the second distal larger than the others, almost spine-shaped) (
<figureCitation id="130B2A7BFFEA325004E6FE33FEFFF4DC" box="[159,276,403,429]" captionStart="FIGURE 4" captionStartId="6.[151,250,1878,1902]" captionTargetBox="[151,1436,237,1855]" captionTargetId="figure-48@6.[151,1436,237,1855]" captionTargetPageId="6" captionText="FIGURE 4. Lacronia utaru sp. nov., ♂ holotype (MHNCI). A: dorsal scutum, chelicerae, trochanters and coxae IV, dorsal view. B: dorsal scutum, lateral view. C: ocularium, anterior view. D: right Fe–Ti III, retrolateral view. E: armature of mesotergal area III, posterior view. F: detail of the prodorsal apophysis of coxa IV, posterior view. G–J: right Tr–Ti IV, G: dorsal view. H: prolateral view. I: ventral view. J: retrolateral view. K: right Mt–Ta IV, retrolateral view. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13346971" httpUri="https://zenodo.org/record/13346971/files/figure.png" pageId="7" pageNumber="424">Figs. 4I–J</figureCitation>
); a retro-lateral row of three sub-conical tubercles on the proximal third, three prominent tubercles and three conical spines interspersed on the centro-distal portion, and five–six ordinary tubercles on the distal fourth (
<figureCitation id="130B2A7BFFEA325004E6FE7BFEC7F484" box="[159,300,475,501]" captionStart="FIGURE 4" captionStartId="6.[151,250,1878,1902]" captionTargetBox="[151,1436,237,1855]" captionTargetId="figure-48@6.[151,1436,237,1855]" captionTargetPageId="6" captionText="FIGURE 4. Lacronia utaru sp. nov., ♂ holotype (MHNCI). A: dorsal scutum, chelicerae, trochanters and coxae IV, dorsal view. B: dorsal scutum, lateral view. C: ocularium, anterior view. D: right Fe–Ti III, retrolateral view. E: armature of mesotergal area III, posterior view. F: detail of the prodorsal apophysis of coxa IV, posterior view. G–J: right Tr–Ti IV, G: dorsal view. H: prolateral view. I: ventral view. J: retrolateral view. K: right Mt–Ta IV, retrolateral view. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13346971" httpUri="https://zenodo.org/record/13346971/files/figure.png" pageId="7" pageNumber="424">Figs 4G, I–J</figureCitation>
); a retro-dorsal row of 15 sub-conical tubercles on the proximal and central thirds, with a conical spine (slightly bent retro-laterad) on the distal third (
<figureCitation id="130B2A7BFFEA325006E5FE5FFCFAF768" box="[668,785,511,537]" captionStart="FIGURE 4" captionStartId="6.[151,250,1878,1902]" captionTargetBox="[151,1436,237,1855]" captionTargetId="figure-48@6.[151,1436,237,1855]" captionTargetPageId="6" captionText="FIGURE 4. Lacronia utaru sp. nov., ♂ holotype (MHNCI). A: dorsal scutum, chelicerae, trochanters and coxae IV, dorsal view. B: dorsal scutum, lateral view. C: ocularium, anterior view. D: right Fe–Ti III, retrolateral view. E: armature of mesotergal area III, posterior view. F: detail of the prodorsal apophysis of coxa IV, posterior view. G–J: right Tr–Ti IV, G: dorsal view. H: prolateral view. I: ventral view. J: retrolateral view. K: right Mt–Ta IV, retrolateral view. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13346971" httpUri="https://zenodo.org/record/13346971/files/figure.png" pageId="7" pageNumber="424">Figs 4G, J</figureCitation>
); a sizeable spur on prodorsal and retro-dorsal apical faces (retro-dorsal larger than prodorsal) (
<figureCitation id="130B2A7BFFEA32500651FD83FD55F74C" box="[552,702,547,573]" captionStart="FIGURE 4" captionStartId="6.[151,250,1878,1902]" captionTargetBox="[151,1436,237,1855]" captionTargetId="figure-48@6.[151,1436,237,1855]" captionTargetPageId="6" captionText="FIGURE 4. Lacronia utaru sp. nov., ♂ holotype (MHNCI). A: dorsal scutum, chelicerae, trochanters and coxae IV, dorsal view. B: dorsal scutum, lateral view. C: ocularium, anterior view. D: right Fe–Ti III, retrolateral view. E: armature of mesotergal area III, posterior view. F: detail of the prodorsal apophysis of coxa IV, posterior view. G–J: right Tr–Ti IV, G: dorsal view. H: prolateral view. I: ventral view. J: retrolateral view. K: right Mt–Ta IV, retrolateral view. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13346971" httpUri="https://zenodo.org/record/13346971/files/figure.png" pageId="7" pageNumber="424">Figs 4G–H, J</figureCitation>
); an outstanding spine on proventral and retro-ventral apical faces (
<figureCitation id="130B2A7BFFEA325004E6FDE7FEFCF710" box="[159,279,583,609]" captionStart="FIGURE 4" captionStartId="6.[151,250,1878,1902]" captionTargetBox="[151,1436,237,1855]" captionTargetId="figure-48@6.[151,1436,237,1855]" captionTargetPageId="6" captionText="FIGURE 4. Lacronia utaru sp. nov., ♂ holotype (MHNCI). A: dorsal scutum, chelicerae, trochanters and coxae IV, dorsal view. B: dorsal scutum, lateral view. C: ocularium, anterior view. D: right Fe–Ti III, retrolateral view. E: armature of mesotergal area III, posterior view. F: detail of the prodorsal apophysis of coxa IV, posterior view. G–J: right Tr–Ti IV, G: dorsal view. H: prolateral view. I: ventral view. J: retrolateral view. K: right Mt–Ta IV, retrolateral view. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13346971" httpUri="https://zenodo.org/record/13346971/files/figure.png" pageId="7" pageNumber="424">Figs 4H–J</figureCitation>
).
</paragraph>
<paragraph id="8B8F36FEFFEA325004BEFDCBFB29F7D8" blockId="7.[151,1437,151,1869]" pageId="7" pageNumber="424">
Patella IV dorsally covered by sub-conical tubercles and conical spines (
<figureCitation id="130B2A7BFFEA32500072FDCBFB5CF7F4" box="[1035,1207,619,645]" captionStart="FIGURE 2" captionStartId="4.[151,250,1546,1570]" captionTargetBox="[151,1436,181,1522]" captionTargetId="figure-19@4.[151,1436,181,1522]" captionTargetPageId="4" captionText="FIGURE 2. Habitus in vivo of Lacronia utaru sp. nov., ♂ paratype (MNRJ 3697), from Brazil, Paraná, Paranaguá, Estação Ecológica do Guaraguaçu. A: dorsal view. B: fronto-dorsal view. C: posterior view. D: dorso-posterior view. E: ventral view. Photographs by Adriano Kury." figureDoi="http://doi.org/10.5281/zenodo.13346967" httpUri="https://zenodo.org/record/13346967/files/figure.png" pageId="7" pageNumber="424">Figs. 2A, C–D</figureCitation>
, 
<figureCitation id="130B2A7BFFEA325000BCFDCBFAE3F7F4" box="[1221,1288,619,645]" captionStart="FIGURE 3" captionStartId="5.[151,250,1184,1208]" captionTargetBox="[151,1436,181,1160]" captionTargetId="figure-17@5.[151,1436,181,1160]" captionTargetPageId="5" captionText="FIGURE 3. Lacronia utaru sp. nov., ♂ holotype (MHNCI). A: habitus, dorsal view. B: ventral view. C: lateral view. D: posterior view. E: Fe–Ta IV, prolateral view (left leg) and retrolateral view (right leg). Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13346969" httpUri="https://zenodo.org/record/13346969/files/figure.png" pageId="7" pageNumber="424">3C–E</figureCitation>
, 
<figureCitation id="130B2A7BFFEA3250016CFDCBFA9CF7F4" box="[1301,1399,619,645]" captionStart="FIGURE 4" captionStartId="6.[151,250,1878,1902]" captionTargetBox="[151,1436,237,1855]" captionTargetId="figure-48@6.[151,1436,237,1855]" captionTargetPageId="6" captionText="FIGURE 4. Lacronia utaru sp. nov., ♂ holotype (MHNCI). A: dorsal scutum, chelicerae, trochanters and coxae IV, dorsal view. B: dorsal scutum, lateral view. C: ocularium, anterior view. D: right Fe–Ti III, retrolateral view. E: armature of mesotergal area III, posterior view. F: detail of the prodorsal apophysis of coxa IV, posterior view. G–J: right Tr–Ti IV, G: dorsal view. H: prolateral view. I: ventral view. J: retrolateral view. K: right Mt–Ta IV, retrolateral view. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13346971" httpUri="https://zenodo.org/record/13346971/files/figure.png" pageId="7" pageNumber="424">4G–H, J</figureCitation>
); a proventral row of three spines (IIi) (
<figureCitation id="130B2A7BFFEA32500653FD2FFD4BF7D8" box="[554,672,655,681]" captionStart="FIGURE 4" captionStartId="6.[151,250,1878,1902]" captionTargetBox="[151,1436,237,1855]" captionTargetId="figure-48@6.[151,1436,237,1855]" captionTargetPageId="6" captionText="FIGURE 4. Lacronia utaru sp. nov., ♂ holotype (MHNCI). A: dorsal scutum, chelicerae, trochanters and coxae IV, dorsal view. B: dorsal scutum, lateral view. C: ocularium, anterior view. D: right Fe–Ti III, retrolateral view. E: armature of mesotergal area III, posterior view. F: detail of the prodorsal apophysis of coxa IV, posterior view. G–J: right Tr–Ti IV, G: dorsal view. H: prolateral view. I: ventral view. J: retrolateral view. K: right Mt–Ta IV, retrolateral view. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13346971" httpUri="https://zenodo.org/record/13346971/files/figure.png" pageId="7" pageNumber="424">Figs 4H–I</figureCitation>
); with retro-ventral three spines (iII) (
<figureCitation id="130B2A7BFFEA32500031FD2FFB5DF7D8" box="[1096,1206,655,681]" captionStart="FIGURE 4" captionStartId="6.[151,250,1878,1902]" captionTargetBox="[151,1436,237,1855]" captionTargetId="figure-48@6.[151,1436,237,1855]" captionTargetPageId="6" captionText="FIGURE 4. Lacronia utaru sp. nov., ♂ holotype (MHNCI). A: dorsal scutum, chelicerae, trochanters and coxae IV, dorsal view. B: dorsal scutum, lateral view. C: ocularium, anterior view. D: right Fe–Ti III, retrolateral view. E: armature of mesotergal area III, posterior view. F: detail of the prodorsal apophysis of coxa IV, posterior view. G–J: right Tr–Ti IV, G: dorsal view. H: prolateral view. I: ventral view. J: retrolateral view. K: right Mt–Ta IV, retrolateral view. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13346971" httpUri="https://zenodo.org/record/13346971/files/figure.png" pageId="7" pageNumber="424">Figs 4I–J</figureCitation>
).
</paragraph>
<paragraph id="8B8F36FEFFEA325004BEFD13FE45F160" blockId="7.[151,1437,151,1869]" pageId="7" pageNumber="424">
Tibia IV dorsal face with a proximal conical tubercle on the proximal fifth, and two ordinary tubercles plus an apical reduced spur on the distal third (
<figureCitation id="130B2A7BFFEA3250063FFD77FD37F780" box="[582,732,727,753]" captionStart="FIGURE 4" captionStartId="6.[151,250,1878,1902]" captionTargetBox="[151,1436,237,1855]" captionTargetId="figure-48@6.[151,1436,237,1855]" captionTargetPageId="6" captionText="FIGURE 4. Lacronia utaru sp. nov., ♂ holotype (MHNCI). A: dorsal scutum, chelicerae, trochanters and coxae IV, dorsal view. B: dorsal scutum, lateral view. C: ocularium, anterior view. D: right Fe–Ti III, retrolateral view. E: armature of mesotergal area III, posterior view. F: detail of the prodorsal apophysis of coxa IV, posterior view. G–J: right Tr–Ti IV, G: dorsal view. H: prolateral view. I: ventral view. J: retrolateral view. K: right Mt–Ta IV, retrolateral view. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13346971" httpUri="https://zenodo.org/record/13346971/files/figure.png" pageId="7" pageNumber="424">Figs 4G–H, J</figureCitation>
); prodorsal face with six prominent subconical tubercles on the proximal 4/5 fifths (
<figureCitation id="130B2A7BFFEA3250050EFD5BFE14F664" box="[375,511,763,789]" captionStart="FIGURE 4" captionStartId="6.[151,250,1878,1902]" captionTargetBox="[151,1436,237,1855]" captionTargetId="figure-48@6.[151,1436,237,1855]" captionTargetPageId="6" captionText="FIGURE 4. Lacronia utaru sp. nov., ♂ holotype (MHNCI). A: dorsal scutum, chelicerae, trochanters and coxae IV, dorsal view. B: dorsal scutum, lateral view. C: ocularium, anterior view. D: right Fe–Ti III, retrolateral view. E: armature of mesotergal area III, posterior view. F: detail of the prodorsal apophysis of coxa IV, posterior view. G–J: right Tr–Ti IV, G: dorsal view. H: prolateral view. I: ventral view. J: retrolateral view. K: right Mt–Ta IV, retrolateral view. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13346971" httpUri="https://zenodo.org/record/13346971/files/figure.png" pageId="7" pageNumber="424">Figs. 4G–H</figureCitation>
); prolateral face with nine prominent subconical tubercles on its entire length (
<figureCitation id="130B2A7BFFEA32500113FD5BFF0BF648" captionStart="FIGURE 4" captionStartId="6.[151,250,1878,1902]" captionTargetBox="[151,1436,237,1855]" captionTargetId="figure-48@6.[151,1436,237,1855]" captionTargetPageId="6" captionText="FIGURE 4. Lacronia utaru sp. nov., ♂ holotype (MHNCI). A: dorsal scutum, chelicerae, trochanters and coxae IV, dorsal view. B: dorsal scutum, lateral view. C: ocularium, anterior view. D: right Fe–Ti III, retrolateral view. E: armature of mesotergal area III, posterior view. F: detail of the prodorsal apophysis of coxa IV, posterior view. G–J: right Tr–Ti IV, G: dorsal view. H: prolateral view. I: ventral view. J: retrolateral view. K: right Mt–Ta IV, retrolateral view. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13346971" httpUri="https://zenodo.org/record/13346971/files/figure.png" pageId="7" pageNumber="424">Figs 4G–H</figureCitation>
); proventral face with three ordinary tubercles on the basal fourth, five subconical tubercles on the central 2/4 fourths, and one subconical tubercle and one spine (I) on the distal fourth (
<figureCitation id="130B2A7BFFEA3250006AFCE3FB60F62C" box="[1043,1163,835,861]" captionStart="FIGURE 4" captionStartId="6.[151,250,1878,1902]" captionTargetBox="[151,1436,237,1855]" captionTargetId="figure-48@6.[151,1436,237,1855]" captionTargetPageId="6" captionText="FIGURE 4. Lacronia utaru sp. nov., ♂ holotype (MHNCI). A: dorsal scutum, chelicerae, trochanters and coxae IV, dorsal view. B: dorsal scutum, lateral view. C: ocularium, anterior view. D: right Fe–Ti III, retrolateral view. E: armature of mesotergal area III, posterior view. F: detail of the prodorsal apophysis of coxa IV, posterior view. G–J: right Tr–Ti IV, G: dorsal view. H: prolateral view. I: ventral view. J: retrolateral view. K: right Mt–Ta IV, retrolateral view. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13346971" httpUri="https://zenodo.org/record/13346971/files/figure.png" pageId="7" pageNumber="424">Figs 4H–I</figureCitation>
); retro-ventral face with five–six prominent subconical tubercles on the proximal 2/3 thirds and three spines (iiI) on the distal third (
<figureCitation id="130B2A7BFFEA32500113FCC7FF27F6D4" captionStart="FIGURE 4" captionStartId="6.[151,250,1878,1902]" captionTargetBox="[151,1436,237,1855]" captionTargetId="figure-48@6.[151,1436,237,1855]" captionTargetPageId="6" captionText="FIGURE 4. Lacronia utaru sp. nov., ♂ holotype (MHNCI). A: dorsal scutum, chelicerae, trochanters and coxae IV, dorsal view. B: dorsal scutum, lateral view. C: ocularium, anterior view. D: right Fe–Ti III, retrolateral view. E: armature of mesotergal area III, posterior view. F: detail of the prodorsal apophysis of coxa IV, posterior view. G–J: right Tr–Ti IV, G: dorsal view. H: prolateral view. I: ventral view. J: retrolateral view. K: right Mt–Ta IV, retrolateral view. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13346971" httpUri="https://zenodo.org/record/13346971/files/figure.png" pageId="7" pageNumber="424">Figs 4I–J</figureCitation>
); retro-lateral face with six–eight subconical spines on its entire length (
<figureCitation id="130B2A7BFFEA32500078FC2BFB78F6D4" box="[1025,1171,907,933]" captionStart="FIGURE 4" captionStartId="6.[151,250,1878,1902]" captionTargetBox="[151,1436,237,1855]" captionTargetId="figure-48@6.[151,1436,237,1855]" captionTargetPageId="6" captionText="FIGURE 4. Lacronia utaru sp. nov., ♂ holotype (MHNCI). A: dorsal scutum, chelicerae, trochanters and coxae IV, dorsal view. B: dorsal scutum, lateral view. C: ocularium, anterior view. D: right Fe–Ti III, retrolateral view. E: armature of mesotergal area III, posterior view. F: detail of the prodorsal apophysis of coxa IV, posterior view. G–J: right Tr–Ti IV, G: dorsal view. H: prolateral view. I: ventral view. J: retrolateral view. K: right Mt–Ta IV, retrolateral view. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13346971" httpUri="https://zenodo.org/record/13346971/files/figure.png" pageId="7" pageNumber="424">Figs 4G, I–J</figureCitation>
); retro-dorsal face with six–eight subconical spines on the proximal 4/5 fifths, with three subconical tubercles on the distal fifth (
<figureCitation id="130B2A7BFFEA32500143FC0FFF4FF69C" captionStart="FIGURE 4" captionStartId="6.[151,250,1878,1902]" captionTargetBox="[151,1436,237,1855]" captionTargetId="figure-48@6.[151,1436,237,1855]" captionTargetPageId="6" captionText="FIGURE 4. Lacronia utaru sp. nov., ♂ holotype (MHNCI). A: dorsal scutum, chelicerae, trochanters and coxae IV, dorsal view. B: dorsal scutum, lateral view. C: ocularium, anterior view. D: right Fe–Ti III, retrolateral view. E: armature of mesotergal area III, posterior view. F: detail of the prodorsal apophysis of coxa IV, posterior view. G–J: right Tr–Ti IV, G: dorsal view. H: prolateral view. I: ventral view. J: retrolateral view. K: right Mt–Ta IV, retrolateral view. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13346971" httpUri="https://zenodo.org/record/13346971/files/figure.png" pageId="7" pageNumber="424">Figs 4G, J</figureCitation>
); proventral and retro-ventral faces with an outstanding spine/spur on apical portion (retro-ventral largest than proventral) (
<figureCitation id="130B2A7BFFEA3250055AFC57FE4AF160" box="[291,417,1015,1041]" captionStart="FIGURE 4" captionStartId="6.[151,250,1878,1902]" captionTargetBox="[151,1436,237,1855]" captionTargetId="figure-48@6.[151,1436,237,1855]" captionTargetPageId="6" captionText="FIGURE 4. Lacronia utaru sp. nov., ♂ holotype (MHNCI). A: dorsal scutum, chelicerae, trochanters and coxae IV, dorsal view. B: dorsal scutum, lateral view. C: ocularium, anterior view. D: right Fe–Ti III, retrolateral view. E: armature of mesotergal area III, posterior view. F: detail of the prodorsal apophysis of coxa IV, posterior view. G–J: right Tr–Ti IV, G: dorsal view. H: prolateral view. I: ventral view. J: retrolateral view. K: right Mt–Ta IV, retrolateral view. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13346971" httpUri="https://zenodo.org/record/13346971/files/figure.png" pageId="7" pageNumber="424">Figs. 4H–J</figureCitation>
).
</paragraph>
<paragraph id="8B8F36FEFFEA325004BEFBBBFA63F10C" blockId="7.[151,1437,151,1869]" pageId="7" pageNumber="424">
Metatarsus IV dorsal face with a row of 14 conical spines (iIIIIIIIIIiIii) (
<figureCitation id="130B2A7BFFEA3250079AFBBBFBAFF144" box="[995,1092,1051,1077]" captionStart="FIGURE 2" captionStartId="4.[151,250,1546,1570]" captionTargetBox="[151,1436,181,1522]" captionTargetId="figure-19@4.[151,1436,181,1522]" captionTargetPageId="4" captionText="FIGURE 2. Habitus in vivo of Lacronia utaru sp. nov., ♂ paratype (MNRJ 3697), from Brazil, Paraná, Paranaguá, Estação Ecológica do Guaraguaçu. A: dorsal view. B: fronto-dorsal view. C: posterior view. D: dorso-posterior view. E: ventral view. Photographs by Adriano Kury." figureDoi="http://doi.org/10.5281/zenodo.13346967" httpUri="https://zenodo.org/record/13346967/files/figure.png" pageId="7" pageNumber="424">Figs. 2C</figureCitation>
, 
<figureCitation id="130B2A7BFFEA32500037FBBBFB9BF144" box="[1102,1136,1051,1077]" captionStart="FIGURE 3" captionStartId="5.[151,250,1184,1208]" captionTargetBox="[151,1436,181,1160]" captionTargetId="figure-17@5.[151,1436,181,1160]" captionTargetPageId="5" captionText="FIGURE 3. Lacronia utaru sp. nov., ♂ holotype (MHNCI). A: habitus, dorsal view. B: ventral view. C: lateral view. D: posterior view. E: Fe–Ta IV, prolateral view (left leg) and retrolateral view (right leg). Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13346969" httpUri="https://zenodo.org/record/13346969/files/figure.png" pageId="7" pageNumber="424">3E</figureCitation>
, 
<figureCitation id="130B2A7BFFEA32500003FBBBFB4AF144" box="[1146,1185,1051,1077]" captionStart="FIGURE 4" captionStartId="6.[151,250,1878,1902]" captionTargetBox="[151,1436,237,1855]" captionTargetId="figure-48@6.[151,1436,237,1855]" captionTargetPageId="6" captionText="FIGURE 4. Lacronia utaru sp. nov., ♂ holotype (MHNCI). A: dorsal scutum, chelicerae, trochanters and coxae IV, dorsal view. B: dorsal scutum, lateral view. C: ocularium, anterior view. D: right Fe–Ti III, retrolateral view. E: armature of mesotergal area III, posterior view. F: detail of the prodorsal apophysis of coxa IV, posterior view. G–J: right Tr–Ti IV, G: dorsal view. H: prolateral view. I: ventral view. J: retrolateral view. K: right Mt–Ta IV, retrolateral view. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13346971" httpUri="https://zenodo.org/record/13346971/files/figure.png" pageId="7" pageNumber="424">4K</figureCitation>
); prodorsal, prolateral, retro-lateral and retrodorsal faces with a row of tiny tubercles (
<figureCitation id="130B2A7BFFEA3250071AFB9FFC54F128" box="[867,959,1087,1113]" captionStart="FIGURE 4" captionStartId="6.[151,250,1878,1902]" captionTargetBox="[151,1436,237,1855]" captionTargetId="figure-48@6.[151,1436,237,1855]" captionTargetPageId="6" captionText="FIGURE 4. Lacronia utaru sp. nov., ♂ holotype (MHNCI). A: dorsal scutum, chelicerae, trochanters and coxae IV, dorsal view. B: dorsal scutum, lateral view. C: ocularium, anterior view. D: right Fe–Ti III, retrolateral view. E: armature of mesotergal area III, posterior view. F: detail of the prodorsal apophysis of coxa IV, posterior view. G–J: right Tr–Ti IV, G: dorsal view. H: prolateral view. I: ventral view. J: retrolateral view. K: right Mt–Ta IV, retrolateral view. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13346971" httpUri="https://zenodo.org/record/13346971/files/figure.png" pageId="7" pageNumber="424">Fig. 4K</figureCitation>
); proventral and retro-ventral faces with a row of sub-conical tubercles (
<figureCitation id="130B2A7BFFEA3250059CFBC3FDD4F10C" box="[485,575,1123,1149]" captionStart="FIGURE 4" captionStartId="6.[151,250,1878,1902]" captionTargetBox="[151,1436,237,1855]" captionTargetId="figure-48@6.[151,1436,237,1855]" captionTargetPageId="6" captionText="FIGURE 4. Lacronia utaru sp. nov., ♂ holotype (MHNCI). A: dorsal scutum, chelicerae, trochanters and coxae IV, dorsal view. B: dorsal scutum, lateral view. C: ocularium, anterior view. D: right Fe–Ti III, retrolateral view. E: armature of mesotergal area III, posterior view. F: detail of the prodorsal apophysis of coxa IV, posterior view. G–J: right Tr–Ti IV, G: dorsal view. H: prolateral view. I: ventral view. J: retrolateral view. K: right Mt–Ta IV, retrolateral view. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13346971" httpUri="https://zenodo.org/record/13346971/files/figure.png" pageId="7" pageNumber="424">Fig. 4K</figureCitation>
); proventral and retro-ventral faces with a spur on apical portion (
<figureCitation id="130B2A7BFFEA32500158FBC3FA90F10C" box="[1313,1403,1123,1149]" captionStart="FIGURE 4" captionStartId="6.[151,250,1878,1902]" captionTargetBox="[151,1436,237,1855]" captionTargetId="figure-48@6.[151,1436,237,1855]" captionTargetPageId="6" captionText="FIGURE 4. Lacronia utaru sp. nov., ♂ holotype (MHNCI). A: dorsal scutum, chelicerae, trochanters and coxae IV, dorsal view. B: dorsal scutum, lateral view. C: ocularium, anterior view. D: right Fe–Ti III, retrolateral view. E: armature of mesotergal area III, posterior view. F: detail of the prodorsal apophysis of coxa IV, posterior view. G–J: right Tr–Ti IV, G: dorsal view. H: prolateral view. I: ventral view. J: retrolateral view. K: right Mt–Ta IV, retrolateral view. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13346971" httpUri="https://zenodo.org/record/13346971/files/figure.png" pageId="7" pageNumber="424">Fig. 4K</figureCitation>
).
</paragraph>
<paragraph id="8B8F36FEFFEA325004BEFB27FC71F304" blockId="7.[151,1437,151,1869]" pageId="7" pageNumber="424">
<emphasis id="B944EAECFFEA325004BEFB27FE85F1D0" box="[199,366,1159,1185]" italics="true" pageId="7" pageNumber="424">Color (in vivo)</emphasis>
(MNRJ 3697, 
<typeStatus id="548B885CFFEA3250066EFB27FD92F1D0" box="[535,633,1159,1185]" pageId="7" pageNumber="424" type="paratype">paratype</typeStatus>
) (
<figureCitation id="130B2A7BFFEA325006E9FB27FCE5F1D0" box="[656,782,1159,1185]" captionStart="FIGURE 2" captionStartId="4.[151,250,1546,1570]" captionTargetBox="[151,1436,181,1522]" captionTargetId="figure-19@4.[151,1436,181,1522]" captionTargetPageId="4" captionText="FIGURE 2. Habitus in vivo of Lacronia utaru sp. nov., ♂ paratype (MNRJ 3697), from Brazil, Paraná, Paranaguá, Estação Ecológica do Guaraguaçu. A: dorsal view. B: fronto-dorsal view. C: posterior view. D: dorso-posterior view. E: ventral view. Photographs by Adriano Kury." figureDoi="http://doi.org/10.5281/zenodo.13346967" httpUri="https://zenodo.org/record/13346967/files/figure.png" pageId="7" pageNumber="424">Figs 2A–E</figureCitation>
). Ocularium, DS background and its regular tubercles in a mixture of 
<emphasis id="B944EAECFFEA3250056DFB0BFE4BF1B5" box="[276,416,1195,1221]" italics="true" pageId="7" pageNumber="424">Light Brown</emphasis>
(57) and 
<emphasis id="B944EAECFFEA32500675FB0BFD21F1B5" box="[524,714,1195,1221]" italics="true" pageId="7" pageNumber="424">Moderate Brown</emphasis>
(58). Pair of spines on ocularium and mesotergum area III 
<emphasis id="B944EAECFFEA3250011AFB0BFEAFF199" italics="true" pageId="7" pageNumber="424">Dark Grayish Brown</emphasis>
(62). DS lateral borders in a mixture of 
<emphasis id="B944EAECFFEA32500768FB6FFC31F198" box="[785,986,1231,1257]" italics="true" pageId="7" pageNumber="424">Brownish Orange</emphasis>
(54) and 
<emphasis id="B944EAECFFEA32500030FB6FFADFF198" box="[1097,1332,1231,1257]" italics="true" pageId="7" pageNumber="424">Light Orange Yellow</emphasis>
(70). 
<collectingCountry id="F327766EFFEA32500101FB6FFA77F198" box="[1400,1436,1231,1257]" name="American Samoa" pageId="7" pageNumber="424">AS</collectingCountry>
posterior border and free tergites I–III 
<emphasis id="B944EAECFFEA3250062AFB53FCA5F07C" box="[595,846,1267,1293]" italics="true" pageId="7" pageNumber="424">Dark Greenish Yellow</emphasis>
(103). Prominent tubercles contrasting with the 
<collectingCountry id="F327766EFFEA32500101FB53FA77F07C" box="[1400,1436,1267,1293]" name="American Samoa" pageId="7" pageNumber="424">AS</collectingCountry>
lateral borders background 
<emphasis id="B944EAECFFEA325005BEFAB7FDA1F040" box="[455,586,1303,1329]" italics="true" pageId="7" pageNumber="424">Pale Yellow</emphasis>
(89). Ch and Pp background in a mixture of 
<emphasis id="B944EAECFFEA3250004EFAB7FAC7F040" box="[1079,1324,1303,1329]" italics="true" pageId="7" pageNumber="424">Dark Greenish Yellow</emphasis>
(103) and 
<emphasis id="B944EAECFFEA325004EEFA9BFEDDF024" box="[151,310,1339,1365]" italics="true" pageId="7" pageNumber="424">Grayish Olive</emphasis>
(110). Cx I–III background in a mixture of 
<emphasis id="B944EAECFFEA3250075EFA9BFC13F025" box="[807,1016,1339,1365]" italics="true" pageId="7" pageNumber="424">Light Olive Brown</emphasis>
(94) and 
<emphasis id="B944EAECFFEA3250001FFA9BFAEEF024" box="[1126,1285,1339,1365]" italics="true" pageId="7" pageNumber="424">Grayish Olive</emphasis>
(110), with a dorsal distal semicircle 
<emphasis id="B944EAECFFEA325005D9FAFFFD72F008" box="[416,665,1375,1401]" italics="true" pageId="7" pageNumber="424">Light Greenish Yellow</emphasis>
(101). Fe–Mt I–II and Fe III proximal and central thirds background in a mixture of 
<emphasis id="B944EAECFFEA32500528FA23FE38F0EC" box="[337,467,1411,1437]" italics="true" pageId="7" pageNumber="424">Light Olive</emphasis>
(106) and 
<emphasis id="B944EAECFFEA3250062FFA23FD3CF0EC" box="[598,727,1411,1437]" italics="true" pageId="7" pageNumber="424">Olive Gray</emphasis>
(113). Fe III distal third and Pa–Mt III almost entirely 
<emphasis id="B944EAECFFEA3250011AFA23FEDEF0B0" italics="true" pageId="7" pageNumber="424">Dark Grayish Olive</emphasis>
(111). Cx–Tr IV background in a mixture of 
<emphasis id="B944EAECFFEA32500753FA07FC5FF0B1" box="[810,948,1447,1473]" italics="true" pageId="7" pageNumber="424">Dark Brown</emphasis>
(59) and 
<emphasis id="B944EAECFFEA32500067FA07FAC3F0B1" box="[1054,1320,1447,1473]" italics="true" pageId="7" pageNumber="424">Strong Yellowish Brown</emphasis>
(74), with a Tr IV dorsal distal semicircle 
<emphasis id="B944EAECFFEA3250058FFA6BFD0AF094" box="[502,737,1483,1509]" italics="true" pageId="7" pageNumber="424">Dark Grayish Yellow</emphasis>
(91). Cx–Tr IV main apophyses 
<emphasis id="B944EAECFFEA32500029FA6BFA76F095" box="[1104,1437,1483,1509]" italics="true" pageId="7" pageNumber="424">Dark Grayish Reddish Brown</emphasis>
(47), with apex 
<emphasis id="B944EAECFFEA32500534FA4FFE1EF378" box="[333,501,1519,1545]" italics="true" pageId="7" pageNumber="424">Strong Orange</emphasis>
(50). Fe–Pa IV and Ti IV proximal 4/5 almost entirely 
<emphasis id="B944EAECFFEA32500009FA4FFAB5F379" box="[1136,1374,1519,1545]" italics="true" pageId="7" pageNumber="424">Dark Reddish Brown</emphasis>
(44), with apex of the main spines 
<emphasis id="B944EAECFFEA3250059AF9B3FD20F35C" box="[483,715,1555,1581]" italics="true" pageId="7" pageNumber="424">Dark Orange Yellow</emphasis>
(72). Ti IV distal fifth background Moderate Olive Brown (95), with proventral and retro-ventral spurs 
<emphasis id="B944EAECFFEA32500633F997FD3EF320" box="[586,725,1591,1617]" italics="true" pageId="7" pageNumber="424">Light Yellow</emphasis>
(86). Mt IV background 
<emphasis id="B944EAECFFEA32500793F997FAFCF321" box="[1002,1303,1591,1617]" italics="true" pageId="7" pageNumber="424">Moderate Yellowish Brown</emphasis>
(77). Mt IV dorsal spines 
<emphasis id="B944EAECFFEA32500549F9FBFE46F304" box="[304,429,1627,1653]" italics="true" pageId="7" pageNumber="424">Olive Gray</emphasis>
(113), with apex 
<emphasis id="B944EAECFFEA3250060BF9FBFCB5F304" box="[626,862,1627,1653]" italics="true" pageId="7" pageNumber="424">Dark Grayish Yellow</emphasis>
(91).
</paragraph>
<paragraph id="8B8F36FEFFEA325004BEF9DFFD2DF23C" blockId="7.[151,1437,151,1869]" pageId="7" pageNumber="424">
<emphasis id="B944EAECFFEA325004BEF9DFFEE3F3E8" box="[199,264,1663,1689]" italics="true" pageId="7" pageNumber="424">Color</emphasis>
(
<emphasis id="B944EAECFFEA32500562F9DFFE64F3E8" box="[283,399,1663,1689]" italics="true" pageId="7" pageNumber="424">in ethanol</emphasis>
) (MHNCI, 
<typeStatus id="548B885CFFEA32500661F9DFFD97F3E8" box="[536,636,1663,1689]" pageId="7" pageNumber="424" type="holotype">holotype</typeStatus>
) (
<figureCitation id="130B2A7BFFEA325006EFF9DFFCF6F3E8" box="[662,797,1663,1689]" captionStart="FIGURE 3" captionStartId="5.[151,250,1184,1208]" captionTargetBox="[151,1436,181,1160]" captionTargetId="figure-17@5.[151,1436,181,1160]" captionTargetPageId="5" captionText="FIGURE 3. Lacronia utaru sp. nov., ♂ holotype (MHNCI). A: habitus, dorsal view. B: ventral view. C: lateral view. D: posterior view. E: Fe–Ta IV, prolateral view (left leg) and retrolateral view (right leg). Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.13346969" httpUri="https://zenodo.org/record/13346969/files/figure.png" pageId="7" pageNumber="424">Figs. 3A–E</figureCitation>
). 
<collectorName id="26C55328FFEA3250074BF9DFFC34F3E8" box="[818,991,1663,1689]" pageId="7" pageNumber="424">Ocularium, DS</collectorName>
background, free tergites I– 
<collectionCode id="ED21AE3BFFEA3250015DF9DFFAABF3E8" box="[1316,1344,1663,1689]" pageId="7" pageNumber="424">III</collectionCode>
and 
<collectorName id="26C55328FFEA32500105F9DFFE30F3CC" pageId="7" pageNumber="424">
Cx IV 
<emphasis id="B944EAECFFEA325004C4F903FE30F3CC" box="[189,475,1699,1725]" italics="true" pageId="7" pageNumber="424">Moderate Orange Yellow</emphasis>
</collectorName>
(71). Spines of the mesotergum area 
<collectionCode id="ED21AE3BFFEA325007F5F903FC43F3CC" box="[908,936,1699,1725]" pageId="7" pageNumber="424">III</collectionCode>
and 
<collectorName id="26C55328FFEA3250079BF903FBC2F3CC" box="[994,1065,1699,1725]" pageId="7" pageNumber="424">Cx IV</collectorName>
prodorsal distal apophysis 
<emphasis id="B944EAECFFEA3250011AF903FEB9F391" italics="true" pageId="7" pageNumber="424">Dark Yellowish Brown</emphasis>
(78), with apex 
<emphasis id="B944EAECFFEA32500671F967FCCAF390" box="[520,801,1735,1761]" italics="true" pageId="7" pageNumber="424">Moderate Orange Yellow</emphasis>
(71). Prominent tubercles contrasting with the 
<collectingCountry id="F327766EFFEA32500155F967FABBF390" box="[1324,1360,1735,1761]" name="American Samoa" pageId="7" pageNumber="424">AS</collectingCountry>
lateral borders background 
<emphasis id="B944EAECFFEA325005FCF94BFDD0F274" box="[389,571,1771,1797]" italics="true" pageId="7" pageNumber="424">Yellowish White</emphasis>
(92). Ch and Leg 
<collectionCode id="ED21AE3BFFEA32500763F94BFCDDF274" box="[794,822,1771,1797]" pageId="7" pageNumber="424">III</collectionCode>
background 
<emphasis id="B944EAECFFEA325007A9F94BFB2BF274" box="[976,1216,1771,1797]" italics="true" pageId="7" pageNumber="424">
<collectorName id="26C55328FFEA325007A9F94BFB2BF274" box="[976,1216,1771,1797]" pageId="7" pageNumber="424">Light Orange Yellow</collectorName>
</emphasis>
(70). 
<collectorName id="26C55328FFEA32500173F94BFAC3F274" box="[1290,1320,1771,1797]" pageId="7" pageNumber="424">Pp</collectorName>
and 
<collectorName id="26C55328FFEA3250011FF94BFF4BF258" pageId="7" pageNumber="424">Legs I</collectorName>
–II background 
<emphasis id="B944EAECFFEA3250052CF8AFFE14F258" box="[341,511,1807,1833]" italics="true" pageId="7" pageNumber="424">
<collectorName id="26C55328FFEA3250052CF8AFFE14F258" box="[341,511,1807,1833]" pageId="7" pageNumber="424">Purplish White</collectorName>
</emphasis>
(231). 
<collectorName id="26C55328FFEA3250062BF8AFFD87F258" box="[594,620,1807,1833]" pageId="7" pageNumber="424">Tr</collectorName>
– 
<collectorName id="26C55328FFEA32500603F8AFFD54F258" box="[634,703,1807,1833]" pageId="7" pageNumber="424">Mt IV</collectorName>
background 
<emphasis id="B944EAECFFEA3250072DF8AFFBA6F258" box="[852,1101,1807,1833]" italics="true" pageId="7" pageNumber="424">
<collectorName id="26C55328FFEA3250072DF8AFFBA6F258" box="[852,1101,1807,1833]" pageId="7" pageNumber="424">Dark Greenish Yellow</collectorName>
</emphasis>
(103). 
<collectorName id="26C55328FFEA325000D9F8AFFB0DF258" box="[1184,1254,1807,1833]" pageId="7" pageNumber="424">Cx IV</collectorName>
and 
<collectorName id="26C55328FFEA32500167F8AFFAB7F258" box="[1310,1372,1807,1833]" pageId="7" pageNumber="424">Tr IV</collectorName>
main apophyses background 
<emphasis id="B944EAECFFEA325005E7F893FD61F23D" box="[414,650,1843,1869]" italics="true" pageId="7" pageNumber="424">
<collectorName id="26C55328FFEA325005E7F893FD61F23D" box="[414,650,1843,1869]" pageId="7" pageNumber="424">Dark Grayish Brown</collectorName>
</emphasis>
(62).
</paragraph>
<caption id="DF4F6676FFE5325F04EEFBB8FB0DF125" ID-DOI="http://doi.org/10.5281/zenodo.13346973" ID-Zenodo-Dep="13346973" httpUri="https://zenodo.org/record/13346973/files/figure.png" pageId="8" pageNumber="425" startId="8.[151,250,1048,1072]" targetBox="[151,1436,181,1024]" targetPageId="8" targetType="figure">
<paragraph id="8B8F36FEFFE5325F04EEFBB8FB0DF125" blockId="8.[151,1436,1048,1109]" pageId="8" pageNumber="425">
<emphasis id="B944EAECFFE5325F04EEFBB8FEF9F141" bold="true" box="[151,274,1048,1072]" pageId="8" pageNumber="425">FIGURE 5.</emphasis>
<taxonomicName id="4C304D7DFFE5325F056EFBB9FE5BF141" authorityName="Carvalho &amp; Kury" authorityYear="2024" box="[279,432,1049,1072]" class="Arachnida" family="Gonyleptidae" genus="Lacronia" kingdom="Animalia" order="Opiliones" pageId="8" pageNumber="425" phylum="Arthropoda" rank="species" species="utaru" status="sp. nov.">
<emphasis id="B944EAECFFE5325F056EFBB9FE5BF141" box="[279,432,1049,1072]" italics="true" pageId="8" pageNumber="425">Lacronia utaru</emphasis>
</taxonomicName>
<emphasis id="B944EAECFFE5325F05CCFBB8FDEEF141" bold="true" box="[437,517,1048,1072]" pageId="8" pageNumber="425">
<taxonomicNameLabel id="A2775797FFE5325F05CCFBB8FDEEF141" box="[437,517,1048,1072]" pageId="8" pageNumber="425" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, ♂ holotype (MHNCI), genitalia, distal part. A: dorsal view. B: lateral view. Scale bars: 100 μm. Letters A to E represent the AE11 macrosetae scheme (
<bibRefCitation id="EFA14B0FFFE5325F0688FB9CFC0AF125" author="Kury, A. B. &amp; Villarreal, O." box="[753,993,1084,1108]" pageId="8" pageNumber="425" pagination="1 - 46" refId="ref8076" refString="Kury, A. B. &amp; Villarreal, O. (2015) The prickly blade mapped: establishing homologies and a chaetotaxy for macrosetae of penis ventral plate in Gonyleptoidea (Arachnida, Opiliones, Laniatores). Zoological Journal of the Linnean Society, 174 (1), 1 - 46. https: // doi. org / 10.1111 / zoj. 12225" type="journal article" year="2015">Kury &amp; Villarreal 2015</bibRefCitation>
, 
<bibRefCitation id="EFA14B0FFFE5325F0792FB9DFB30F125" author="Kury, A. B. &amp; Machado, G." box="[1003,1243,1084,1109]" pageId="8" pageNumber="425" pagination="358 - 370" refId="ref7851" refString="Kury, A. B. &amp; Machado, G. (2021) The genus Eurytromma from Sri Lanka: the homology of penial macrosetae in Podoctidae matches the gonyleptoid AE 11 pattern (Opiliones: Laniatores: Epedanoidea). The Journal of Arachnology, 49, 358 - 370. https: // doi. org / 10.1636 / JoA-S- 20 - 078" type="journal article" year="2021">Kury &amp; Machado 2021</bibRefCitation>
).
</paragraph>
</caption>
<caption id="DF4F6676FFE5325F04EEF964FAE2F252" ID-DOI="http://doi.org/10.5281/zenodo.13346975" ID-Zenodo-Dep="13346975" httpUri="https://zenodo.org/record/13346975/files/figure.png" pageId="8" pageNumber="425" startId="8.[151,250,1732,1756]" targetBox="[151,1436,1132,1708]" targetPageId="8" targetType="figure">
<paragraph id="8B8F36FEFFE5325F04EEF964FAE2F252" blockId="8.[151,1436,1732,1828]" pageId="8" pageNumber="425">
<emphasis id="B944EAECFFE5325F04EEF964FEFDF3AD" bold="true" box="[151,278,1732,1756]" pageId="8" pageNumber="425">FIGURE 6.</emphasis>
Schematic representation of males’ DS, showing the two main sets of colored tubercles in the ‘not areolated 
<taxonomicName id="4C304D7DFFE5325F04EEF949FF18F271" authorityName="Strand" authorityYear="1942" box="[151,243,1769,1792]" class="Arachnida" family="Gonyleptidae" genus="Lacronia" kingdom="Animalia" order="Opiliones" pageId="8" pageNumber="425" phylum="Arthropoda" rank="genus">
<emphasis id="B944EAECFFE5325F04EEF949FF18F271" box="[151,243,1769,1792]" italics="true" pageId="8" pageNumber="425">Lacronia</emphasis>
</taxonomicName>
’. The red rectangle indicates the area of the central set, and the blue rectangles indicate the AS lateral sets. A: 
<taxonomicName id="4C304D7DFFE5325F0139F949FE9CF255" authority="Mello-Leitao" authorityName="Mello-Leitao" class="Arachnida" family="Gonyleptidae" genus="Lacronia" kingdom="Animalia" order="Opiliones" pageId="8" pageNumber="425" phylum="Arthropoda" rank="species" species="serripes">
<emphasis id="B944EAECFFE5325F0139F949FF02F255" italics="true" pageId="8" pageNumber="425">Lacronia serripes</emphasis>
Mello-Leitão
</taxonomicName>
, ♂ (MZSP 14342). B: 
<taxonomicName id="4C304D7DFFE5325F0627F8ADFD1CF255" authorityName="Carvalho &amp; Kury" authorityYear="2024" box="[606,759,1805,1828]" class="Arachnida" family="Gonyleptidae" genus="Lacronia" kingdom="Animalia" order="Opiliones" pageId="8" pageNumber="425" phylum="Arthropoda" rank="species" species="utaru" status="sp. nov.">
<emphasis id="B944EAECFFE5325F0627F8ADFD1CF255" box="[606,759,1805,1828]" italics="true" pageId="8" pageNumber="425">Lacronia utaru</emphasis>
</taxonomicName>
<emphasis id="B944EAECFFE5325F0684F8ACFCA5F255" bold="true" box="[765,846,1804,1828]" pageId="8" pageNumber="425">
<taxonomicNameLabel id="A2775797FFE5325F0684F8ACFCA5F255" box="[765,846,1804,1828]" pageId="8" pageNumber="425" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, ♂ holotype (MHNCI). Scale bars = 2 mm.
</paragraph>
</caption>
<paragraph id="8B8F36FEFFE5325E04BEF8D6FB30F484" blockId="8.[151,1437,1910,2009]" lastBlockId="9.[151,1437,150,573]" lastPageId="9" lastPageNumber="426" pageId="8" pageNumber="425">
<emphasis id="B944EAECFFE5325F04BEF8D6FE9DF2E1" box="[199,374,1910,1936]" italics="true" pageId="8" pageNumber="425">Male genitalia.</emphasis>
VP slightly divided into an apical square-shaped half (with latero-apical flaps), and a basal half elliptical (
<figureCitation id="130B2A7BFFE5325F0545F83AFE7CF2C4" box="[316,407,1946,1973]" captionStart="FIGURE 5" captionStartId="8.[151,250,1048,1072]" captionTargetBox="[151,1436,181,1024]" captionTargetId="figure-19@8.[151,1436,181,1024]" captionTargetPageId="8" captionText="FIGURE 5. Lacronia utaru sp. nov., ♂ holotype (MHNCI), genitalia, distal part.A: dorsal view. B: lateral view. Scale bars: 100 μm. Letters A to E represent the AE11 macrosetae scheme (Kury &amp; Villarreal 2015, Kury &amp; Machado 2021)." figureDoi="http://doi.org/10.5281/zenodo.13346973" httpUri="https://zenodo.org/record/13346973/files/figure.png" pageId="8" pageNumber="425">Fig. 5A</figureCitation>
); ventral surface entirely covered with microsetae of 
<typeStatus id="548B885CFFE5325F078BF83BFBC9F2C4" box="[1010,1058,1947,1973]" pageId="8" pageNumber="425">type</typeStatus>
1. All macrosetae inserted on the ventro-laterals of VP. MS A1–A3 cylindrical, thick, and acuminate, forming a diagonal row in lateral view (MS A1 dorsalmost) (
<figureCitation id="130B2A7BFFE4325E0555FF37FE5EF5C0" box="[300,437,150,177]" captionStart="FIGURE 5" captionStartId="8.[151,250,1048,1072]" captionTargetBox="[151,1436,181,1024]" captionTargetId="figure-19@8.[151,1436,181,1024]" captionTargetPageId="8" captionText="FIGURE 5. Lacronia utaru sp. nov., ♂ holotype (MHNCI), genitalia, distal part.A: dorsal view. B: lateral view. Scale bars: 100 μm. Letters A to E represent the AE11 macrosetae scheme (Kury &amp; Villarreal 2015, Kury &amp; Machado 2021)." figureDoi="http://doi.org/10.5281/zenodo.13346973" httpUri="https://zenodo.org/record/13346973/files/figure.png" pageId="9" pageNumber="426">Figs. 5A–B</figureCitation>
). MS B1 small, inserted ventrally and close to MS A3 (
<figureCitation id="130B2A7BFFE4325E004AFF37FB66F5C0" box="[1075,1165,150,177]" captionStart="FIGURE 5" captionStartId="8.[151,250,1048,1072]" captionTargetBox="[151,1436,181,1024]" captionTargetId="figure-19@8.[151,1436,181,1024]" captionTargetPageId="8" captionText="FIGURE 5. Lacronia utaru sp. nov., ♂ holotype (MHNCI), genitalia, distal part.A: dorsal view. B: lateral view. Scale bars: 100 μm. Letters A to E represent the AE11 macrosetae scheme (Kury &amp; Villarreal 2015, Kury &amp; Machado 2021)." figureDoi="http://doi.org/10.5281/zenodo.13346973" httpUri="https://zenodo.org/record/13346973/files/figure.png" pageId="9" pageNumber="426">Fig. 5B</figureCitation>
). MS C1–C3 forming a longitudinal row (MS C3 dorsalmost) on the apical quarter of VP (
<figureCitation id="130B2A7BFFE4325E0701FF1BFC15F5A4" box="[888,1022,186,213]" captionStart="FIGURE 5" captionStartId="8.[151,250,1048,1072]" captionTargetBox="[151,1436,181,1024]" captionTargetId="figure-19@8.[151,1436,181,1024]" captionTargetPageId="8" captionText="FIGURE 5. Lacronia utaru sp. nov., ♂ holotype (MHNCI), genitalia, distal part.A: dorsal view. B: lateral view. Scale bars: 100 μm. Letters A to E represent the AE11 macrosetae scheme (Kury &amp; Villarreal 2015, Kury &amp; Machado 2021)." figureDoi="http://doi.org/10.5281/zenodo.13346973" httpUri="https://zenodo.org/record/13346973/files/figure.png" pageId="9" pageNumber="426">Figs. 5A–B</figureCitation>
). MS D1 small, closer to MS C3 than the MS A1 (
<figureCitation id="130B2A7BFFE4325E0559FF7FFE4DF588" box="[288,422,222,249]" captionStart="FIGURE 5" captionStartId="8.[151,250,1048,1072]" captionTargetBox="[151,1436,181,1024]" captionTargetId="figure-19@8.[151,1436,181,1024]" captionTargetPageId="8" captionText="FIGURE 5. Lacronia utaru sp. nov., ♂ holotype (MHNCI), genitalia, distal part.A: dorsal view. B: lateral view. Scale bars: 100 μm. Letters A to E represent the AE11 macrosetae scheme (Kury &amp; Villarreal 2015, Kury &amp; Machado 2021)." figureDoi="http://doi.org/10.5281/zenodo.13346973" httpUri="https://zenodo.org/record/13346973/files/figure.png" pageId="9" pageNumber="426">Figs. 5A–B</figureCitation>
). MS E1–E2 small, more ventrally on the laterodistal flange of VP. MS E1 placed between MS C1–C2 level, MS E2 same, below C3 (but closer to MS C3 than MS D1) (
<figureCitation id="130B2A7BFFE4325E0070FEA3FB8AF46C" box="[1033,1121,258,285]" captionStart="FIGURE 5" captionStartId="8.[151,250,1048,1072]" captionTargetBox="[151,1436,181,1024]" captionTargetId="figure-19@8.[151,1436,181,1024]" captionTargetPageId="8" captionText="FIGURE 5. Lacronia utaru sp. nov., ♂ holotype (MHNCI), genitalia, distal part.A: dorsal view. B: lateral view. Scale bars: 100 μm. Letters A to E represent the AE11 macrosetae scheme (Kury &amp; Villarreal 2015, Kury &amp; Machado 2021)." figureDoi="http://doi.org/10.5281/zenodo.13346973" httpUri="https://zenodo.org/record/13346973/files/figure.png" pageId="9" pageNumber="426">Fig. 5B</figureCitation>
). Glans sac arising from the middle bulge on the podium, not extended as a dorsal process (
<figureCitation id="130B2A7BFFE4325E0729FE87FC3BF430" box="[848,976,294,321]" captionStart="FIGURE 5" captionStartId="8.[151,250,1048,1072]" captionTargetBox="[151,1436,181,1024]" captionTargetId="figure-19@8.[151,1436,181,1024]" captionTargetPageId="8" captionText="FIGURE 5. Lacronia utaru sp. nov., ♂ holotype (MHNCI), genitalia, distal part.A: dorsal view. B: lateral view. Scale bars: 100 μm. Letters A to E represent the AE11 macrosetae scheme (Kury &amp; Villarreal 2015, Kury &amp; Machado 2021)." figureDoi="http://doi.org/10.5281/zenodo.13346973" httpUri="https://zenodo.org/record/13346973/files/figure.png" pageId="9" pageNumber="426">Figs 5A–B</figureCitation>
). Stylus and its ventral process axis fused basally, forming a prominent trapezoidal-shaped pedestal (
<figureCitation id="130B2A7BFFE4325E0753FEEBFC68F414" box="[810,899,330,357]" captionStart="FIGURE 5" captionStartId="8.[151,250,1048,1072]" captionTargetBox="[151,1436,181,1024]" captionTargetId="figure-19@8.[151,1436,181,1024]" captionTargetPageId="8" captionText="FIGURE 5. Lacronia utaru sp. nov., ♂ holotype (MHNCI), genitalia, distal part.A: dorsal view. B: lateral view. Scale bars: 100 μm. Letters A to E represent the AE11 macrosetae scheme (Kury &amp; Villarreal 2015, Kury &amp; Machado 2021)." figureDoi="http://doi.org/10.5281/zenodo.13346973" httpUri="https://zenodo.org/record/13346973/files/figure.png" pageId="9" pageNumber="426">Fig. 5B</figureCitation>
). Stylus cylindrical, almost straight, inserted on the pedestal in a 10º angle, without a conspicuous head (
<figureCitation id="130B2A7BFFE4325E0764FECFFC4FF4F8" box="[797,932,366,393]" captionStart="FIGURE 5" captionStartId="8.[151,250,1048,1072]" captionTargetBox="[151,1436,181,1024]" captionTargetId="figure-19@8.[151,1436,181,1024]" captionTargetPageId="8" captionText="FIGURE 5. Lacronia utaru sp. nov., ♂ holotype (MHNCI), genitalia, distal part.A: dorsal view. B: lateral view. Scale bars: 100 μm. Letters A to E represent the AE11 macrosetae scheme (Kury &amp; Villarreal 2015, Kury &amp; Machado 2021)." figureDoi="http://doi.org/10.5281/zenodo.13346973" httpUri="https://zenodo.org/record/13346973/files/figure.png" pageId="9" pageNumber="426">Figs. 5A–B</figureCitation>
); its subapical portion slightly swollen, with subdistal tiny spines on the lateral and ventral faces (
<figureCitation id="130B2A7BFFE4325E077AFE33FC66F4DC" box="[771,909,402,429]" captionStart="FIGURE 5" captionStartId="8.[151,250,1048,1072]" captionTargetBox="[151,1436,181,1024]" captionTargetId="figure-19@8.[151,1436,181,1024]" captionTargetPageId="8" captionText="FIGURE 5. Lacronia utaru sp. nov., ♂ holotype (MHNCI), genitalia, distal part.A: dorsal view. B: lateral view. Scale bars: 100 μm. Letters A to E represent the AE11 macrosetae scheme (Kury &amp; Villarreal 2015, Kury &amp; Machado 2021)." figureDoi="http://doi.org/10.5281/zenodo.13346973" httpUri="https://zenodo.org/record/13346973/files/figure.png" pageId="9" pageNumber="426">Figs. 5A–B</figureCitation>
). Ventral process of stylus cylindrical, almost straight, with ¾ of the stylus length, and a flabellum on the apical half (
<figureCitation id="130B2A7BFFE4325E07BCFE17FBF5F4A0" box="[965,1054,438,465]" captionStart="FIGURE 5" captionStartId="8.[151,250,1048,1072]" captionTargetBox="[151,1436,181,1024]" captionTargetId="figure-19@8.[151,1436,181,1024]" captionTargetPageId="8" captionText="FIGURE 5. Lacronia utaru sp. nov., ♂ holotype (MHNCI), genitalia, distal part.A: dorsal view. B: lateral view. Scale bars: 100 μm. Letters A to E represent the AE11 macrosetae scheme (Kury &amp; Villarreal 2015, Kury &amp; Machado 2021)." figureDoi="http://doi.org/10.5281/zenodo.13346973" httpUri="https://zenodo.org/record/13346973/files/figure.png" pageId="9" pageNumber="426">Fig. 5B</figureCitation>
). Flabellum slightly bent ventrad, scallop-shaped with spines, with approximately 50% of the ventral process stem length (
<figureCitation id="130B2A7BFFE4325E000FFE7BFB25F484" box="[1142,1230,474,501]" captionStart="FIGURE 5" captionStartId="8.[151,250,1048,1072]" captionTargetBox="[151,1436,181,1024]" captionTargetId="figure-19@8.[151,1436,181,1024]" captionTargetPageId="8" captionText="FIGURE 5. Lacronia utaru sp. nov., ♂ holotype (MHNCI), genitalia, distal part.A: dorsal view. B: lateral view. Scale bars: 100 μm. Letters A to E represent the AE11 macrosetae scheme (Kury &amp; Villarreal 2015, Kury &amp; Machado 2021)." figureDoi="http://doi.org/10.5281/zenodo.13346973" httpUri="https://zenodo.org/record/13346973/files/figure.png" pageId="9" pageNumber="426">Fig. 5B</figureCitation>
).
</paragraph>
<paragraph id="8B8F36FEFFE4325E04BEFE5FFA88F768" blockId="9.[151,1437,150,573]" box="[199,1379,511,537]" pageId="9" pageNumber="426">
<emphasis id="B944EAECFFE4325E04BEFE5FFE21F769" box="[199,458,511,537]" italics="true" pageId="9" pageNumber="426">Intraspecific variation.</emphasis>
It was not possible to detect morphologically 
<emphasis id="B944EAECFFE4325E07B6FE5FFBF9F769" box="[975,1042,511,536]" italics="true" pageId="9" pageNumber="426">minor</emphasis>
males in the studied material.
</paragraph>
<paragraph id="8B8F36FEFFE4325E04BEFD82FBA7F74C" blockId="9.[151,1437,150,573]" box="[199,1100,546,573]" pageId="9" pageNumber="426">
<emphasis id="B944EAECFFE4325E04BEFD82FE04F74D" bold="true" box="[199,495,546,572]" pageId="9" pageNumber="426">Geographic distribution.</emphasis>
<collectingCountry id="F327766EFFE4325E058FFD83FDB5F74C" box="[502,606,547,573]" name="Brazil" pageId="9" pageNumber="426">BRAZIL</collectingCountry>
, 
<collectingRegion id="49F4F81CFFE4325E0610FD83FD13F74C" box="[617,760,547,573]" country="Brazil" name="Parana" pageId="9" pageNumber="426">Paraná State</collectingRegion>
: Morretes, Paranaguá (
<figureCitation id="130B2A7BFFE4325E0783FD83FBD0F74C" box="[1018,1083,547,573]" captionStart="FIGURE 1" captionStartId="2.[151,250,1788,1812]" captionTargetBox="[235,1348,921,1760]" captionTargetId="figure-389@2.[235,1351,921,1765]" captionTargetPageId="2" captionText="FIGURE 1. Southeastern and South Brazil, showing distribution of the eight species of Lacronia. The area shaded in dark green displays the Atlantic Province, as defined by Morrone et al. (2022). Inset: Location of the represented sector in South America." figureDoi="http://doi.org/10.5281/zenodo.13346965" httpUri="https://zenodo.org/record/13346965/files/figure.png" pageId="9" pageNumber="426">Fig. 1</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>