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<document id="91B529503ACC1CD97E4D2777F6AC1C26" ID-DOI="10.5281/zenodo.205995" ID-GBIF-Dataset="556343c6-a289-4d17-841e-ef4666ca7406" ID-ISSN="1175-5326" ID-Zenodo-Dep="205995" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460274268212" checkinUser="plazi" docAuthor="Meyer, Harry A." docDate="2011" docId="0399878AFFD2FF8B58B6C05AFB513C2C" docLanguage="en" docName="zt02812p040.pdf" docOrigin="Zootaxa 2812" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Macrobiotus caymanensis Meyer, 2011, sp. n." docType="treatment" docVersion="9" lastPageNumber="38" masterDocId="FFA0FFF2FFD5FF815821C179FFD2387B" masterDocTitle="Tardigrada of Grand Cayman, West Indies, with descriptions of two new species of eutardigrade, Doryphoribius tessellatus (Hypsibiidae) and Macrobiotus caymanensis (Macrobiotidae)" masterLastPageNumber="40" masterPageNumber="28" pageNumber="35" updateTime="1698247824567" updateUser="plazi">
<mods:mods id="FEABFD1E5B8DE8FA362978D3F5D297A6" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="603EB762575C9C48105DD3C878A64C00">Tardigrada of Grand Cayman, West Indies, with descriptions of two new species of eutardigrade, Doryphoribius tessellatus (Hypsibiidae) and Macrobiotus caymanensis (Macrobiotidae)</mods:title>
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<mods:namePart id="969A265194D9039E58CB3E7F4FA4D4A0">Meyer, Harry A.</mods:namePart>
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<mods:date id="6F32354FB2164DF04F99DCFD418D2C48">2011</mods:date>
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<mods:classification id="3FE7DE15FECFCCEB2D45EAABE0A7A241">journal article</mods:classification>
<mods:identifier id="DD8D0F093EC86F2E5DF9636C72304F37" type="DOI">10.5281/zenodo.205995</mods:identifier>
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<treatment id="0399878AFFD2FF8B58B6C05AFB513C2C" ID-DOI="http://doi.org/10.5281/zenodo.5684275" ID-GBIF-Taxon="119395759" ID-Zenodo-Dep="5684275" LSID="urn:lsid:plazi:treatment:0399878AFFD2FF8B58B6C05AFB513C2C" httpUri="http://treatment.plazi.org/id/0399878AFFD2FF8B58B6C05AFB513C2C" lastPageId="10" lastPageNumber="38" pageId="7" pageNumber="35">
<subSubSection id="C32A6517FFD2FF8658B6C05AFE673924" pageId="7" pageNumber="35" type="nomenclature">
<paragraph id="8B8F369CFFD2FF8658B6C05AFDD83947" blockId="7.[151,522,291,351]" box="[151,522,291,316]" pageId="7" pageNumber="35">
<heading id="D0C781F0FFD2FF8658B6C05AFDD83947" bold="true" box="[151,522,291,316]" fontSize="10" level="2" pageId="7" pageNumber="35" reason="2">
<emphasis id="B944EA8EFFD2FF8658B6C05AFDD83947" bold="true" box="[151,522,291,316]" pageId="7" pageNumber="35">
<taxonomicName id="4C304D1FFFD2FF8658B6C05AFE163947" ID-CoL="563925f8-8758-472b-8e01-83cd686009ab" box="[151,452,291,316]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="7" pageNumber="35" phylum="Tardigrada" rank="species" species="caymanensis" status="sp. nov.">
<emphasis id="B944EA8EFFD2FF8658B6C05AFE163947" bold="true" box="[151,452,291,316]" italics="true" pageId="7" pageNumber="35">Macrobiotus caymanensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27757F5FFD2FF8659EAC05DFDD83947" box="[459,522,292,316]" pageId="7" pageNumber="35" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8B8F369CFFD2FF8658B6C03EFE673924" blockId="7.[151,522,291,351]" box="[151,437,327,351]" pageId="7" pageNumber="35">
(
<figureCitation id="130B2A19FFD2FF8658BEC03EFEF53924" box="[159,295,327,351]" captionStart-0="FIGURES 13 – 16" captionStart-1="FIGURES 17 – 18" captionStartId-0="7.[151,264,1914,1937]" captionStartId-1="9.[151,264,578,601]" captionTargetBox-0="[245,1332,814,1878]" captionTargetBox-1="[406,1181,194,541]" captionTargetId-0="figure@7.[234,1353,787,1893]" captionTargetId-1="figure@9.[403,1185,193,545]" captionTargetPageId-0="7" captionTargetPageId-1="9" captionText-0="FIGURES 13 – 16. Macrobiotus caymanensis sp. n. (13) Cuticular pores (arrows). (14) Buccopharyngeal apparatus. (15) Buccal cavity, with arrows indicating peribuccal lamellae and transverse ridges. (16) Claw, leg II. Scale bars: 13, 15 – 16, 10 μm; 13, 20 μm." captionText-1="FIGURES 17 – 18. Macrobiotus caymanensis sp. n. (17) Egg. (18) Egg shell with processes and hexagonal areolation. Scale bars: 10 μm." httpUri-0="https://zenodo.org/record/206001/files/figure.png" httpUri-1="https://zenodo.org/record/206002/files/figure.png" pageId="7" pageNumber="35">Figs. 13–18</figureCitation>
, Tables 2–3)
</paragraph>
</subSubSection>
<subSubSection id="C32A6517FFD2FF8658B6C0F6FC813A6F" pageId="7" pageNumber="35" type="diagnosis">
<paragraph id="8B8F369CFFD2FF8658B6C0F6FC813A6F" blockId="7.[151,1436,399,749]" pageId="7" pageNumber="35">
<emphasis id="B944EA8EFFD2FF8658B6C0F6FEC339D3" bold="true" box="[151,273,399,424]" pageId="7" pageNumber="35">Diagnosis.</emphasis>
A
<taxonomicName id="4C304D1FFFD2FF865915C0E9FE1039D3" box="[308,450,400,424]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="7" pageNumber="35" phylum="Tardigrada" rank="genus">
<emphasis id="B944EA8EFFD2FF865915C0E9FE1039D3" box="[308,450,400,424]" italics="true" pageId="7" pageNumber="35">Macrobiotus</emphasis>
</taxonomicName>
of the
<taxonomicName id="4C304D1FFFD2FF865A32C0E9FDAA39D3" box="[531,632,400,424]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="7" pageNumber="35" phylum="Tardigrada" rank="species" species="polyopus">
<emphasis id="B944EA8EFFD2FF865A32C0E9FDAA39D3" box="[531,632,400,424]" italics="true" pageId="7" pageNumber="35">polyopus</emphasis>
</taxonomicName>
-group with minute cuticular pores, two macroplacoids and a microplacoid, a relatively short buccal tube and posterior stylet support insertion (pt 77.5–79.3) and eggs, laid free, with ornamented shell of reticular design with hexagonal mesh and a small number (
<quantity id="4CC89B79FFD2FF865C2BC0AEFBB23994" box="[1034,1120,471,495]" metricMagnitude="-1" metricUnit="m" metricValue="2.667" metricValueMax="3.048" metricValueMin="2.286" pageId="7" pageNumber="35" unit="in" value="10.5" valueMax="12.0" valueMin="9.0">9–12 in</quantity>
the circumference and
<quantity id="4CC89B79FFD2FF865D45C0AEFF7F3A6F" metricMagnitude="-1" metricUnit="m" metricValue="2.794" metricValueMax="3.302" metricValueMin="2.286" pageId="7" pageNumber="35" unit="in" value="11.0" valueMax="13.0" valueMin="9.0">9–13 in</quantity>
the hemisphere) of large, conical processes (9–20 μm high).
</paragraph>
</subSubSection>
<subSubSection id="C32A6517FFD2FF8658E4C359FDA03A04" pageId="7" pageNumber="35" type="materials_examined">
<paragraph id="8B8F369CFFD2FF8658E4C359FDA03A04" blockId="7.[151,1436,399,749]" pageId="7" pageNumber="35">
<emphasis id="B944EA8EFFD2FF8658E4C359FE253A43" bold="true" box="[197,503,543,568]" pageId="7" pageNumber="35">
<typeStatus id="548B883EFFD2FF8658E4C359FED23A43" box="[197,256,544,568]" pageId="7" pageNumber="35">Type</typeStatus>
material examined.
</emphasis>
<typeStatus id="548B883EFFD2FF865A23C366FDBF3A4C" box="[514,621,543,567]" pageId="7" pageNumber="35" type="holotype">Holotype</typeStatus>
, seven
<typeStatus id="548B883EFFD2FF865AE8C359FCE53A43" box="[713,823,544,568]" pageId="7" pageNumber="35" type="paratype">paratypes</typeStatus>
, one embryonated egg: sample 4b (
<tableCitation id="C6B20327FFD2FF865CC1C366FAEB3A4C" box="[1248,1337,543,567]" captionStart="TABLE 1" captionStartId="1.[151,235,506,527]" captionTargetBox="[151,1436,592,1919]" captionTargetPageId="1" captionText="TABLE 1. Samples containing terrestrial tardigrades from Grand Cayman, Cayman Islands, West Indies. Letters after site num-" httpUri="http://table.plazi.org/id/DF4F6614FFD4FF8058B6C083FA4E3A6B" pageId="7" pageNumber="35" tableUuid="DF4F6614FFD4FF8058B6C083FA4E3A6B">Table 1</tableCitation>
),
<taxonomicName id="4C304D1FFFD2FF865D77C366FF263A27" authority="Reserve" authorityName="Reserve" class="Entognatha" family="Paronellidae" genus="Salina" kingdom="Animalia" order="Collembola" pageId="7" pageNumber="35" phylum="Arthropoda" rank="genus">Salina Reserve</taxonomicName>
, East
<collectingCountry id="F327760CFFD2FF865918C33DFEB43A27" box="[313,358,580,604]" name="India" pageId="7" pageNumber="35">End</collectingCountry>
District, Grand
<collectingCountry id="F327760CFFD2FF865A00C33DFD533A27" box="[545,641,580,604]" name="Cayman Islands" pageId="7" pageNumber="35">Cayman</collectingCountry>
,
<collectingCountry id="F327760CFFD2FF865AAFC33DFC943A27" box="[654,838,580,604]" name="Cayman Islands" pageId="7" pageNumber="35">Cayman Islands</collectingCountry>
, West Indies, 1921ʹ0 0.3ʺ N and 8105ʹ48.7ʺ W,
<quantity id="4CC89B79FFD2FF865D42C33DFA4B3A27" box="[1379,1433,580,604]" metricMagnitude="0" metricUnit="m" metricValue="9.1" pageId="7" pageNumber="35" unit="m" value="9.1">9.1m</quantity>
asl, dry tropical forest, squamulose lichen.
</paragraph>
</subSubSection>
<subSubSection id="C32A6517FFD2FF8858E4C3F2FDF43D44" lastPageId="9" lastPageNumber="37" pageId="7" pageNumber="35" type="description">
<paragraph id="8B8F369CFFD2FF8658E4C3F2FF223A97" blockId="7.[151,1436,399,749]" pageId="7" pageNumber="35">
<emphasis id="B944EA8EFFD2FF8658E4C3F2FE373ADF" bold="true" box="[197,485,651,676]" pageId="7" pageNumber="35">
Description of
<typeStatus id="548B883EFFD2FF865956C3F5FE323ADF" box="[375,480,652,676]" pageId="7" pageNumber="35" type="holotype">holotype</typeStatus>
.
</emphasis>
Body length 452.7 μm, colourless. Sex undetermined. Cuticle smooth, with numerous, uniformly distributed small, circular pores about 1.0 μm in width (
<figureCitation id="130B2A19FFD2FF865BE2C3D6FBCA3ABC" box="[963,1048,687,711]" captionStart="FIGURES 13 – 16" captionStartId="7.[151,264,1914,1937]" captionTargetBox="[245,1332,814,1878]" captionTargetId="figure@7.[234,1353,787,1893]" captionTargetPageId="7" captionText="FIGURES 13 – 16. Macrobiotus caymanensis sp. n. (13) Cuticular pores (arrows). (14) Buccopharyngeal apparatus. (15) Buccal cavity, with arrows indicating peribuccal lamellae and transverse ridges. (16) Claw, leg II. Scale bars: 13, 15 – 16, 10 μm; 13, 20 μm." httpUri="https://zenodo.org/record/206001/files/figure.png" pageId="7" pageNumber="35">Fig. 13</figureCitation>
). Eyes present. No granulation on any leg.
</paragraph>
<caption id="DF4F6614FFD2FF8658B6C603FF0D3FB6" httpUri="https://zenodo.org/record/206001/files/figure.png" pageId="7" pageNumber="35" targetBox="[245,1332,814,1878]" targetPageId="7">
<paragraph id="8B8F369CFFD2FF8658B6C603FF0D3FB6" blockId="7.[151,1436,1914,1997]" pageId="7" pageNumber="35">
<emphasis id="B944EA8EFFD2FF8658B6C603FE863FEB" bold="true" box="[151,340,1914,1937]" pageId="7" pageNumber="35">FIGURES 13–16.</emphasis>
<taxonomicName id="4C304D1FFFD2FF86597AC603FDB13FEB" box="[347,611,1914,1936]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="7" pageNumber="35" phylum="Tardigrada" rank="species" species="caymanensis" status="sp. nov.">
<emphasis id="B944EA8EFFD2FF86597AC603FDB13FEB" box="[347,611,1914,1936]" italics="true" pageId="7" pageNumber="35">Macrobiotus caymanensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27757F5FFD2FF865A4AC602FD763FEB" box="[619,676,1915,1936]" pageId="7" pageNumber="35" rank="species">
<emphasis id="B944EA8EFFD2FF865A4AC602FD763FEB" bold="true" box="[619,676,1915,1936]" pageId="7" pageNumber="35">sp. n.</emphasis>
</taxonomicNameLabel>
(13) Cuticular pores (arrows). (14) Buccopharyngeal apparatus. (15) Buccal cavity, with arrows indicating peribuccal lamellae and transverse ridges. (16) Claw, leg II. Scale bars: 13, 15–16, 10 μm; 13, 20 μm.
</paragraph>
</caption>
<caption id="DF4F6614FFDCFF8858B6C33BFEC43A0C" httpUri="https://zenodo.org/record/206002/files/figure.png" pageId="9" pageNumber="37" targetBox="[406,1181,194,541]" targetPageId="9">
<paragraph id="8B8F369CFFDCFF8858B6C33BFEC43A0C" blockId="9.[151,1436,578,631]" pageId="9" pageNumber="37">
<emphasis id="B944EA8EFFDCFF8858B6C33BFE873A23" bold="true" box="[151,341,578,601]" pageId="9" pageNumber="37">FIGURES 17–18.</emphasis>
<taxonomicName id="4C304D1FFFDCFF88597CC33AFDB53A22" box="[349,615,579,601]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="9" pageNumber="37" phylum="Tardigrada" rank="species" species="caymanensis" status="sp. nov.">
<emphasis id="B944EA8EFFDCFF88597CC33AFDB53A22" box="[349,615,579,601]" italics="true" pageId="9" pageNumber="37">Macrobiotus caymanensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27757F5FFDCFF885A4EC33AFD783A23" box="[623,682,579,600]" pageId="9" pageNumber="37" rank="species">
<emphasis id="B944EA8EFFDCFF885A4EC33AFD783A23" bold="true" box="[623,682,579,600]" pageId="9" pageNumber="37">sp. n.</emphasis>
</taxonomicNameLabel>
(17) Egg. (18) Egg shell with processes and hexagonal areolation. Scale bars: 10 μm.
</paragraph>
</caption>
<paragraph id="8B8F369CFFDCFF8858E4C3E6FA8A3B58" blockId="9.[151,1437,671,1344]" pageId="9" pageNumber="37">
Mouth opening terminal with ten small peribuccal lamellae (
<figureCitation id="130B2A19FFDCFF885B48C3E6FC343ACC" box="[873,998,671,695]" captionStart="FIGURES 13 – 16" captionStartId="7.[151,264,1914,1937]" captionTargetBox="[245,1332,814,1878]" captionTargetId="figure@7.[234,1353,787,1893]" captionTargetPageId="7" captionText="FIGURES 13 – 16. Macrobiotus caymanensis sp. n. (13) Cuticular pores (arrows). (14) Buccopharyngeal apparatus. (15) Buccal cavity, with arrows indicating peribuccal lamellae and transverse ridges. (16) Claw, leg II. Scale bars: 13, 15 – 16, 10 μm; 13, 20 μm." httpUri="https://zenodo.org/record/206001/files/figure.png" pageId="9" pageNumber="37">Fig. 14, 15</figureCitation>
). Buccal cavity small. Buccal armature very simple and difficult to see with light microscopy, system of 3 very thin transversal ridges present with posterior band of very small teeth (
<figureCitation id="130B2A19FFDCFF8859C6C39EFDE93A84" box="[487,571,743,767]" captionStart="FIGURES 13 – 16" captionStartId="7.[151,264,1914,1937]" captionTargetBox="[245,1332,814,1878]" captionTargetId="figure@7.[234,1353,787,1893]" captionTargetPageId="7" captionText="FIGURES 13 – 16. Macrobiotus caymanensis sp. n. (13) Cuticular pores (arrows). (14) Buccopharyngeal apparatus. (15) Buccal cavity, with arrows indicating peribuccal lamellae and transverse ridges. (16) Claw, leg II. Scale bars: 13, 15 – 16, 10 μm; 13, 20 μm." httpUri="https://zenodo.org/record/206001/files/figure.png" pageId="9" pageNumber="37">Fig. 15</figureCitation>
). Buccal tube 39.2 μm long and 6.9 μm [
<emphasis id="B944EA8EFFDCFF885C2DC39EFBEF3A84" box="[1036,1085,743,767]" italics="true" pageId="9" pageNumber="37">17.6</emphasis>
] wide. Ventral lamina 21.6 μm [
<emphasis id="B944EA8EFFDCFF885881C272FF033B58" box="[160,209,779,803]" italics="true" pageId="9" pageNumber="37">55.1</emphasis>
] long, double the length of the stylet sheaths. Stylet supports inserted on buccal tube at 30.4 μm [
<emphasis id="B944EA8EFFDCFF885D37C272FA953B58" box="[1302,1351,779,803]" italics="true" pageId="9" pageNumber="37">77.6</emphasis>
].
</paragraph>
<paragraph id="8B8F369CFFDCFF8858E4C256FD0E3BC8" blockId="9.[151,1437,671,1344]" pageId="9" pageNumber="37">
Oval pharyngeal bulb with apophyses, two rod-shaped macroplacoids decreasing in length and a small, rodshaped microplacoid (
<figureCitation id="130B2A19FFDCFF8859BAC22AFE273B10" box="[411,501,851,875]" captionStart="FIGURES 13 – 16" captionStartId="7.[151,264,1914,1937]" captionTargetBox="[245,1332,814,1878]" captionTargetId="figure@7.[234,1353,787,1893]" captionTargetPageId="7" captionText="FIGURES 13 – 16. Macrobiotus caymanensis sp. n. (13) Cuticular pores (arrows). (14) Buccopharyngeal apparatus. (15) Buccal cavity, with arrows indicating peribuccal lamellae and transverse ridges. (16) Claw, leg II. Scale bars: 13, 15 – 16, 10 μm; 13, 20 μm." httpUri="https://zenodo.org/record/206001/files/figure.png" pageId="9" pageNumber="37">Fig. 14</figureCitation>
). First macroplacoid 11.0 μm [
<emphasis id="B944EA8EFFDCFF885B51C22AFC733B10" box="[880,929,851,875]" italics="true" pageId="9" pageNumber="37">28.1</emphasis>
] long, with a central constriction. Second macroplacoid 7.0 μm [
<emphasis id="B944EA8EFFDCFF8859BEC20EFE023BF4" box="[415,464,887,911]" italics="true" pageId="9" pageNumber="37">17.9</emphasis>
] long, not constricted. Microplacoid 2.9 μm [
<emphasis id="B944EA8EFFDCFF885BC6C20EFBD83BF4" box="[999,1034,887,911]" italics="true" pageId="9" pageNumber="37">7.4</emphasis>
] long. Macroplacoid row 19.8 μm [
<emphasis id="B944EA8EFFDCFF885881C2E2FF033BC8" box="[160,209,923,947]" italics="true" pageId="9" pageNumber="37">50.5</emphasis>
] long, entire placoid row 23.5 μm [
<emphasis id="B944EA8EFFDCFF885A40C2E2FD403BC8" box="[609,658,923,947]" italics="true" pageId="9" pageNumber="37">59.9</emphasis>
] long.
</paragraph>
<paragraph id="8B8F369CFFDCFF8858E4C2C6FBA03C1C" blockId="9.[151,1437,671,1344]" pageId="9" pageNumber="37">
Claws of
<taxonomicName id="4C304D1FFFDCFF885913C2C6FE4E3BAC" box="[306,412,959,983]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="9" pageNumber="37" phylum="Tardigrada" rank="species" species="hufelandi">
<emphasis id="B944EA8EFFDCFF885913C2C6FE4E3BAC" box="[306,412,959,983]" italics="true" pageId="9" pageNumber="37">hufelandi</emphasis>
</taxonomicName>
-
<typeStatus id="548B883EFFDCFF885985C2B9FE0B3BA3" box="[420,473,960,984]" pageId="9" pageNumber="37">type</typeStatus>
, short (especially the common basal tract) and stout (
<figureCitation id="130B2A19FFDCFF885C17C2C6FB583BAC" box="[1078,1162,959,983]" captionStart="FIGURES 13 – 16" captionStartId="7.[151,264,1914,1937]" captionTargetBox="[245,1332,814,1878]" captionTargetId="figure@7.[234,1353,787,1893]" captionTargetPageId="7" captionText="FIGURES 13 – 16. Macrobiotus caymanensis sp. n. (13) Cuticular pores (arrows). (14) Buccopharyngeal apparatus. (15) Buccal cavity, with arrows indicating peribuccal lamellae and transverse ridges. (16) Claw, leg II. Scale bars: 13, 15 – 16, 10 μm; 13, 20 μm." httpUri="https://zenodo.org/record/206001/files/figure.png" pageId="9" pageNumber="37">Fig. 16</figureCitation>
). External claw lengths: leg I primary branch 10.5 μm [
<emphasis id="B944EA8EFFDCFF8859D7C29AFDF53B80" box="[502,551,995,1019]" italics="true" pageId="9" pageNumber="37">26.8</emphasis>
], secondary branch 8.7 μm [
<emphasis id="B944EA8EFFDCFF885B4CC29AFC4C3B80" box="[877,926,995,1019]" italics="true" pageId="9" pageNumber="37">22.2</emphasis>
]; leg II pb 10.4 μm [
<emphasis id="B944EA8EFFDCFF885CAEC29AFB123B80" box="[1167,1216,995,1019]" italics="true" pageId="9" pageNumber="37">26.5</emphasis>
], sb 8.7 μm [
<emphasis id="B944EA8EFFDCFF885D7BC29AFA593B80" box="[1370,1419,995,1019]" italics="true" pageId="9" pageNumber="37">22.2</emphasis>
]; leg III pb 11.2 μm [
<emphasis id="B944EA8EFFDCFF885950C57EFE703C64" box="[369,418,1031,1055]" italics="true" pageId="9" pageNumber="37">28.6</emphasis>
], sb 9.6 μm [
<emphasis id="B944EA8EFFDCFF885A16C57EFDBA3C64" box="[567,616,1031,1055]" italics="true" pageId="9" pageNumber="37">24.5</emphasis>
]. Posterior claw lengths: leg IV pb 13.3 μm [
<emphasis id="B944EA8EFFDCFF885C42C57EFB463C64" box="[1123,1172,1031,1055]" italics="true" pageId="9" pageNumber="37">33.9</emphasis>
], sb 9.0 μm [
<emphasis id="B944EA8EFFDCFF885D0BC57EFA893C64" box="[1322,1371,1031,1055]" italics="true" pageId="9" pageNumber="37">23.0</emphasis>
]. Primary branches of all claws with very well-developed accessory points. Small, smooth lunules present on all legs (leg II, 3.1 μm [
<emphasis id="B944EA8EFFDCFF885968C536FEBE3C1C" box="[329,364,1103,1127]" italics="true" pageId="9" pageNumber="37">7.9</emphasis>
] wide; leg IV, 4.1 μm [
<emphasis id="B944EA8EFFDCFF885A53C536FD713C1C" box="[626,675,1103,1127]" italics="true" pageId="9" pageNumber="37">10.5</emphasis>
] wide). No cuticular bars visible on legs.
</paragraph>
<paragraph id="8B8F369CFFDCFF8858E4C50AFB8F3CD4" blockId="9.[151,1437,671,1344]" pageId="9" pageNumber="37">
Morphometric data and summary statistics for the
<typeStatus id="548B883EFFDCFF885B2EC50AFCA33CF0" box="[783,881,1139,1163]" pageId="9" pageNumber="37" type="holotype">holotype</typeStatus>
and seven
<typeStatus id="548B883EFFDCFF885BD6C50DFBB03CF7" box="[1015,1122,1140,1164]" pageId="9" pageNumber="37" type="paratype">paratypes</typeStatus>
are given in Table 2. Four
<typeStatus id="548B883EFFDCFF8858B6C5E1FED03CCB" box="[151,258,1176,1200]" pageId="9" pageNumber="37" type="paratype">paratypes</typeStatus>
(50%) lacked discernable eyes when examined immediately after mounting.
</paragraph>
<paragraph id="8B8F369CFFDCFF8858E4C5C5FDF43D44" blockId="9.[151,1437,671,1344]" pageId="9" pageNumber="37">
Eggs laid freely (
<figureCitation id="130B2A19FFDCFF8859A8C5C2FE0E3CA8" box="[393,476,1211,1235]" captionStart="FIGURES 17 – 18" captionStartId="9.[151,264,578,601]" captionTargetBox="[406,1181,194,541]" captionTargetId="figure@9.[403,1185,193,545]" captionTargetPageId="9" captionText="FIGURES 17 – 18. Macrobiotus caymanensis sp. n. (17) Egg. (18) Egg shell with processes and hexagonal areolation. Scale bars: 10 μm." httpUri="https://zenodo.org/record/206002/files/figure.png" pageId="9" pageNumber="37">Fig. 17</figureCitation>
), with small number of large processes (9–12 around the circumference,
<quantity id="4CC89B79FFDCFF885D3DC5C2FAA03CA8" box="[1308,1394,1211,1235]" metricMagnitude="-1" metricUnit="m" metricValue="2.794" metricValueMax="3.302" metricValueMin="2.286" pageId="9" pageNumber="37" unit="in" value="11.0" valueMax="13.0" valueMin="9.0">9–13 in</quantity>
the hemisphere). Egg shell areolated, six polygonal areolae around each process present (
<figureCitation id="130B2A19FFDCFF885C76C5A6FB783C8C" box="[1111,1194,1247,1271]" captionStart="FIGURES 17 – 18" captionStartId="9.[151,264,578,601]" captionTargetBox="[406,1181,194,541]" captionTargetId="figure@9.[403,1185,193,545]" captionTargetPageId="9" captionText="FIGURES 17 – 18. Macrobiotus caymanensis sp. n. (17) Egg. (18) Egg shell with processes and hexagonal areolation. Scale bars: 10 μm." httpUri="https://zenodo.org/record/206002/files/figure.png" pageId="9" pageNumber="37">Fig. 18</figureCitation>
). Egg shell otherwise smooth. Processes conical, tapering to a sharp point with very faint annulations. Measurements of some structures from five eggs are given in Table 3.
</paragraph>
</subSubSection>
<paragraph id="8B8F369CFFDCFF8858B6C416FAC63F67" pageId="9" pageNumber="37">
<table id="F930C43CFFDC007E58B6C416FA4F3F66" box="[151,1437,1391,1821]" gridcols="4" gridrows="5" pageId="9" pageNumber="37">
<tr id="350034DEFFDC007E58B6C416FA4F3DD3" box="[151,1437,1391,1448]" gridrow="0" pageId="9" pageNumber="37">
<th id="76D15DA2FFDC007E58B6C416FD143DD3" box="[151,710,1391,1448]" gridcol="0" gridrow="0" pageId="9" pageNumber="37">
<emphasis id="B944EA8EFFDCFF8858B6C416FED73DFF" bold="true" box="[151,261,1391,1412]" pageId="9" pageNumber="37">TABLE 3.</emphasis>
Measurements (in micrometres) of selected Cayman mounted in polyvinyl lactophenol.
</th>
<th id="76D15DA2FFDC007E5AECC416FCA83DD3" box="[717,890,1391,1448]" gridcol="1" gridrow="0" pageId="9" pageNumber="37">structures of five</th>
<th id="76D15DA2FFDC007E5BA0C416FB0E3DD3" box="[897,1244,1391,1448]" gridcol="2" gridrow="0" pageId="9" pageNumber="37">
eggs of
<taxonomicName id="4C304D1FFFDCFF885BF2C416FB0E3DFE" box="[979,1244,1391,1413]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="9" pageNumber="37" phylum="Tardigrada" rank="species" species="caymanensis">
<emphasis id="B944EA8EFFDCFF885BF2C416FB0E3DFE" box="[979,1244,1391,1413]" italics="true" pageId="9" pageNumber="37">Macrobiotus caymanensis</emphasis>
</taxonomicName>
</th>
<th id="76D15DA2FFDC007E5CC2C416FA4F3DD3" box="[1251,1437,1391,1448]" gridcol="3" gridrow="0" pageId="9" pageNumber="37">
<emphasis id="B944EA8EFFDCFF885CC2C416FACF3DFF" bold="true" box="[1251,1309,1391,1412]" pageId="9" pageNumber="37">sp. n.</emphasis>
from Grand
</th>
</tr>
<tr id="350034DEFFDC007E58B6C4BDFA4F3DA7" box="[151,1437,1476,1500]" gridrow="1" pageId="9" pageNumber="37">
<th id="76D15DA2FFDC007E58B6C4BDFD143DA7" box="[151,710,1476,1500]" gridcol="0" gridrow="1" pageId="9" pageNumber="37">Character 1</th>
<td id="76D15DA2FFDC007E5AECC4BDFCA83DA7" box="[717,890,1476,1500]" gridcol="1" gridrow="1" pageId="9" pageNumber="37">2</td>
<td id="76D15DA2FFDC007E5BA0C4BDFB0E3DA7" box="[897,1244,1476,1500]" gridcol="2" gridrow="1" pageId="9" pageNumber="37">3 4</td>
<td id="76D15DA2FFDC007E5CC2C4BDFA4F3DA7" box="[1251,1437,1476,1500]" gridcol="3" gridrow="1" pageId="9" pageNumber="37">5</td>
</tr>
<tr id="350034DEFFDC007E58B6C489FA4F3ED0" box="[151,1437,1520,1707]" gridrow="2" pageId="9" pageNumber="37">
<th id="76D15DA2FFDC007E58B6C489FD143ED0" box="[151,710,1520,1707]" gridcol="0" gridrow="2" pageId="9" pageNumber="37">Diameter of egg without processes 73.7 Diameter of egg with processes 94.6 Processes height 9.1–13.2 Basal diameter of processes 12.3–18.1 Distance between processes 2.4–6.6</th>
<td id="76D15DA2FFDC007E5AECC489FCA83ED0" box="[717,890,1520,1707]" gridcol="1" gridrow="2" pageId="9" pageNumber="37">72.3 87.6 9.3–13.5 16.1–17.3 2.9–7.7</td>
<td id="76D15DA2FFDC007E5BA0C489FB0E3ED0" box="[897,1244,1520,1707]" gridcol="2" gridrow="2" pageId="9" pageNumber="37">71.8 66.0 104.1 90.9 8.3–20.0 8.9–13.2 15.0–19.2 13.7–18.2 4.9–8.3 3.0–8.3</td>
<td id="76D15DA2FFDC007E5CC2C489FA4F3ED0" box="[1251,1437,1520,1707]" gridcol="3" gridrow="2" pageId="9" pageNumber="37">73.1 98.5 10.4–13.1 14.6–18.6 3.4–6.0</td>
</tr>
<tr id="350034DEFFDC007E58B6C7C6FA4F3E88" box="[151,1437,1727,1779]" gridrow="3" pageId="9" pageNumber="37">
<th id="76D15DA2FFDC007E58B6C7C6FD143E88" box="[151,710,1727,1779]" gridcol="0" gridrow="3" pageId="9" pageNumber="37">Number of processes around the circum- 12 ference</th>
<td id="76D15DA2FFDC007E5AECC7C6FCA83E88" box="[717,890,1727,1779]" gridcol="1" gridrow="3" pageId="9" pageNumber="37">11</td>
<td id="76D15DA2FFDC007E5BA0C7C6FB0E3E88" box="[897,1244,1727,1779]" gridcol="2" gridrow="3" pageId="9" pageNumber="37">10 9</td>
<td id="76D15DA2FFDC007E5CC2C7C6FA4F3E88" box="[1251,1437,1727,1779]" gridcol="3" gridrow="3" pageId="9" pageNumber="37">11</td>
</tr>
<tr id="350034DEFFDC007E58B6C67EFA4F3F66" box="[151,1437,1799,1821]" gridrow="4" pageId="9" pageNumber="37">
<th id="76D15DA2FFDC007E58B6C67EFD143F66" box="[151,710,1799,1821]" gridcol="0" gridrow="4" pageId="9" pageNumber="37">Number of processes in the hemisphere 13</th>
<td id="76D15DA2FFDC007E5AECC67EFCA83F66" box="[717,890,1799,1821]" gridcol="1" gridrow="4" pageId="9" pageNumber="37">10</td>
<td id="76D15DA2FFDC007E5BA0C67EFB0E3F66" box="[897,1244,1799,1821]" gridcol="2" gridrow="4" pageId="9" pageNumber="37">10 9</td>
<td id="76D15DA2FFDC007E5CC2C67EFA4F3F66" box="[1251,1437,1799,1821]" gridcol="3" gridrow="4" pageId="9" pageNumber="37">13</td>
</tr>
</table>
</paragraph>
<subSubSection id="C32A6517FFDCFF8858E4C61CFF2E3FBF" pageId="9" pageNumber="37" type="discussion">
<paragraph id="8B8F369CFFDCFF8858E4C61CFF2E3FBF" blockId="9.[151,1436,1892,1988]" pageId="9" pageNumber="37">
<emphasis id="B944EA8EFFDCFF8858E4C61CFEEB3F06" bold="true" box="[197,313,1893,1917]" pageId="9" pageNumber="37">Remarks.</emphasis>
Eighty-two additional specimens were found in samples 1, 2a, 2b, 3, 4a, 4d,
<specimenCount id="9D36FD15FFDCFF885CBEC61DFB6C3F07" box="[1183,1214,1892,1916]" pageId="9" pageNumber="37" type="generic">4f.</specimenCount>
4h and
<date id="FF8E105CFFDCFF885D39C61DFAFC3F07" box="[1304,1326,1892,1916]" pageId="9" pageNumber="37">4i</date>
(
<tableCitation id="C6B20327FFDCFF885D1CC61DFA423F07" box="[1341,1424,1892,1916]" captionStart="TABLE 1" captionStartId="1.[151,235,506,527]" captionTargetBox="[151,1436,592,1919]" captionTargetPageId="1" captionText="TABLE 1. Samples containing terrestrial tardigrades from Grand Cayman, Cayman Islands, West Indies. Letters after site num-" httpUri="http://table.plazi.org/id/DF4F6614FFD4FF8058B6C083FA4E3A6B" pageId="9" pageNumber="37" tableUuid="DF4F6614FFD4FF8058B6C083FA4E3A6B">Table 1</tableCitation>
) in foliose and squamulose lichen, bark and moss.
<taxonomicName id="4C304D1FFFDCFF885AECC6F1FC203FDB" box="[717,1010,1928,1952]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="9" pageNumber="37" phylum="Tardigrada" rank="species" species="caymanensis" status="sp. nov.">
<emphasis id="B944EA8EFFDCFF885AECC6F1FC203FDB" box="[717,1010,1928,1952]" italics="true" pageId="9" pageNumber="37">Macrobiotus caymanensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27757F5FFDCFF885BDBC6F1FBE93FDB" box="[1018,1083,1928,1952]" pageId="9" pageNumber="37" rank="species">
<emphasis id="B944EA8EFFDCFF885BDBC6F1FBE93FDB" bold="true" box="[1018,1083,1928,1952]" pageId="9" pageNumber="37">sp. n.</emphasis>
</taxonomicNameLabel>
is widely distributed on Grand
<collectingCountry id="F327760CFFDCFF8858B6C6D5FF2A3FBF" box="[151,248,1964,1988]" name="Cayman Islands" pageId="9" pageNumber="37">Cayman</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C32A6517FFDFFF8B58E4C1E1FEBB38AF" pageId="10" pageNumber="38" type="etymology">
<paragraph id="8B8F369CFFDFFF8B58E4C1E1FEBB38AF" blockId="10.[151,1437,151,1112]" pageId="10" pageNumber="38">
<emphasis id="B944EA8EFFDFFF8B58E4C1E1FE9938CB" bold="true" box="[197,331,152,176]" pageId="10" pageNumber="38">Etymology.</emphasis>
The specific name is a masculine adjective commemorating the
<collectingCountry id="F327760CFFDFFF8B5C11C1EEFB3A38D4" box="[1072,1256,151,175]" name="Cayman Islands" pageId="10" pageNumber="38">Cayman Islands</collectingCountry>
, where the species was collected.
</paragraph>
</subSubSection>
<subSubSection id="C32A6517FFDFFF8B58E4C199FB513C2C" pageId="10" pageNumber="38" type="materials_examined">
<paragraph id="8B8F369CFFDFFF8B58E4C199FB873944" blockId="10.[151,1437,151,1112]" pageId="10" pageNumber="38">
<emphasis id="B944EA8EFFDFFF8B58E4C199FE553883" bold="true" box="[197,391,223,248]" pageId="10" pageNumber="38">
<typeStatus id="548B883EFFDFFF8B58E4C199FED23883" box="[197,256,224,248]" pageId="10" pageNumber="38">Type</typeStatus>
repository.
</emphasis>
The
<typeStatus id="548B883EFFDFFF8B59E1C1A6FDF0388C" box="[448,546,223,247]" pageId="10" pageNumber="38" type="holotype">holotype</typeStatus>
and three
<typeStatus id="548B883EFFDFFF8B5AB6C199FCD13883" box="[663,771,224,248]" pageId="10" pageNumber="38" type="paratype">paratypes</typeStatus>
(
<collectionCode id="ED21AE59FFDFFF8B5B33C1A6FCB0388C" box="[786,866,223,247]" pageId="10" pageNumber="38">SMLA</collectionCode>
9401) are deposited in the W.A.K. Seale Museum, Department of Biology and Health Science, McNeese State University in Lake Charles, Louisiana,
<collectingCountry id="F327760CFFDFFF8B5D30C07DFA8C3967" box="[1297,1374,260,284]" name="United States of America" pageId="10" pageNumber="38">U.S.A.</collectingCountry>
Four
<typeStatus id="548B883EFFDFFF8B58B6C051FED0393B" box="[151,258,296,320]" pageId="10" pageNumber="38" type="paratype">paratypes</typeStatus>
(
<collectionCode id="ED21AE59FFDFFF8B5931C05EFE813944" box="[272,339,295,319]" pageId="10" pageNumber="38">NTCI</collectionCode>
9402) are deposited with the National Trust for the
<collectingCountry id="F327760CFFDFFF8B5BBBC05EFB833944" box="[922,1105,295,319]" name="Cayman Islands" pageId="10" pageNumber="38">Cayman Islands</collectingCountry>
.
</paragraph>
<paragraph id="8B8F369CFFDFFF8B58E4C032FECB3A6C" blockId="10.[151,1437,151,1112]" pageId="10" pageNumber="38">
<emphasis id="B944EA8EFFDFFF8B58E4C032FE19391F" bold="true" box="[197,459,331,356]" pageId="10" pageNumber="38">Differential diagnosis.</emphasis>
<taxonomicName id="4C304D1FFFDFFF8B59F3C035FD24391F" box="[466,758,332,356]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="10" pageNumber="38" phylum="Tardigrada" rank="species" species="caymanensis" status="sp. nov.">
<emphasis id="B944EA8EFFDFFF8B59F3C035FD24391F" box="[466,758,332,356]" italics="true" pageId="10" pageNumber="38">Macrobiotus caymanensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27757F5FFDFFF8B5ADCC035FCEF391F" box="[765,829,332,356]" pageId="10" pageNumber="38" rank="species">
<emphasis id="B944EA8EFFDFFF8B5ADCC035FCEF391F" bold="true" box="[765,829,332,356]" pageId="10" pageNumber="38">sp. n.</emphasis>
</taxonomicNameLabel>
belongs to the
<taxonomicName id="4C304D1FFFDFFF8B5BC9C035FB9F391F" box="[1000,1101,332,356]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="10" pageNumber="38" phylum="Tardigrada" rank="species" species="polyopus">
<emphasis id="B944EA8EFFDFFF8B5BC9C035FB9F391F" box="[1000,1101,332,356]" italics="true" pageId="10" pageNumber="38">polyopus</emphasis>
</taxonomicName>
-group of species, which have in common a pharyngeal bulb with two macroplacoids and a small or absent microplacoid, short claws and egg shells with reticular design with hexagonal mesh (
<bibRefCitation id="EFA14B6DFFDFFF8B5AECC0EDFC8739D7" author="Pilato" box="[717,853,404,428]" pageId="10" pageNumber="38" refString="Pilato, G. (2006) Remarks on the Macrobiotus polyopus group, with the description of two new species. Zootaxa, 1298, 37 - 47." type="journal article" year="2006">Pilato 2006</bibRefCitation>
). Five species are currently attributed to this group:
<taxonomicName id="4C304D1FFFDFFF8B58B6C0C0FE1639B4" authority="Marcus, 1928" authorityName="Marcus" authorityYear="1928" box="[151,452,439,464]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="10" pageNumber="38" phylum="Tardigrada" rank="species" species="polyopus">
<emphasis id="B944EA8EFFDFFF8B58B6C0C0FEF039AB" box="[151,290,440,464]" italics="true" pageId="10" pageNumber="38">M. polyopus</emphasis>
<bibRefCitation id="EFA14B6DFFDFFF8B590BC0C1FE1639B4" author="Marcus" box="[298,452,439,463]" pageId="10" pageNumber="38" refString="Marcus, E. (1928) Spinnentiere oder Arachoides. IV. Bartirchen (Tardigrada). In: Dahl, F. (Ed.), Die Tierreich Deutschlands und der angrenzenden Meeresteile Jena. Gustav Fischer, Jena, pp. 1 - 230." type="book chapter" year="1928">
Marcus,
<quantity id="4CC89B79FFDFFF8B59ADC0CEFE1639B4" box="[396,452,439,463]" metricMagnitude="1" metricUnit="m" metricValue="4.897119999999999" pageId="10" pageNumber="38" unit="in" value="1928.0">1928</quantity>
</bibRefCitation>
</taxonomicName>
in Sumatra,
<taxonomicName id="4C304D1FFFDFFF8B5A74C0C0FC5A39B4" authority="Pilato, 1974" authorityName="Pilato" authorityYear="1974" box="[597,904,439,464]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="10" pageNumber="38" phylum="Tardigrada" rank="species" species="mandalaae">
<emphasis id="B944EA8EFFDFFF8B5A74C0C0FD2A39AB" box="[597,760,440,464]" italics="true" pageId="10" pageNumber="38">M. mandalaae</emphasis>
<bibRefCitation id="EFA14B6DFFDFFF8B5B21C0CEFC5A39B4" author="Pilato" box="[768,904,439,463]" pageId="10" pageNumber="38" refString="Pilato, G. (1974) Tre nuove specie di Tardigradi delle isole Andamane. Bolletino del Museo Civico di Storia Naturale di Verona, 7, 281 - 291." type="journal article" year="1974">
Pilato,
<quantity id="4CC89B79FFDFFF8B5B71C0CEFC5A39B4" box="[848,904,439,463]" metricMagnitude="1" metricUnit="m" metricValue="5.01396" pageId="10" pageNumber="38" unit="in" value="1974.0">1974</quantity>
</bibRefCitation>
</taxonomicName>
in southern
<collectingCountry id="F327760CFFDFFF8B5C34C0CEFB8939B4" box="[1045,1115,439,463]" name="China" pageId="10" pageNumber="38">China</collectingCountry>
,
<taxonomicName id="4C304D1FFFDFFF8B5C47C0C0FAAC39B4" authority="Pilato, 2006" authorityName="Pilato" authorityYear="2006" box="[1126,1406,439,464]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="10" pageNumber="38" phylum="Tardigrada" rank="species" species="insularis">
<emphasis id="B944EA8EFFDFFF8B5C47C0C0FB3C39AB" box="[1126,1262,440,464]" italics="true" pageId="10" pageNumber="38">M. insularis</emphasis>
<bibRefCitation id="EFA14B6DFFDFFF8B5CD7C0CEFAAC39B4" author="Pilato" box="[1270,1406,439,463]" pageId="10" pageNumber="38" refString="Pilato, G. (2006) Remarks on the Macrobiotus polyopus group, with the description of two new species. Zootaxa, 1298, 37 - 47." type="journal article" year="2006">
Pilato,
<quantity id="4CC89B79FFDFFF8B5D67C0CEFAAC39B4" box="[1350,1406,439,463]" metricMagnitude="1" metricUnit="m" metricValue="5.0952399999999995" pageId="10" pageNumber="38" unit="in" value="2006.0">2006</quantity>
</bibRefCitation>
</taxonomicName>
in the Andaman Islands,
<taxonomicName id="4C304D1FFFDFFF8B59B4C0A4FD66398F" authority="Pilato, 2006" authorityName="Pilato" authorityYear="2006" box="[405,692,476,500]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="10" pageNumber="38" phylum="Tardigrada" rank="species" species="ocotensis">
<emphasis id="B944EA8EFFDFFF8B59B4C0A4FDF1398F" box="[405,547,477,500]" italics="true" pageId="10" pageNumber="38">M. ocotensis</emphasis>
<bibRefCitation id="EFA14B6DFFDFFF8B5A0DC0A5FD66398F" author="Pilato" box="[556,692,476,500]" pageId="10" pageNumber="38" refString="Pilato, G. (2006) Remarks on the Macrobiotus polyopus group, with the description of two new species. Zootaxa, 1298, 37 - 47." type="journal article" year="2006">
Pilato,
<quantity id="4CC89B79FFDFFF8B5A5DC0A5FD66398F" box="[636,692,476,500]" metricMagnitude="1" metricUnit="m" metricValue="5.0952399999999995" pageId="10" pageNumber="38" unit="in" value="2006.0">2006</quantity>
</bibRefCitation>
</taxonomicName>
in southern
<collectingCountry id="F327760CFFDFFF8B5B62C0A5FC4A398F" box="[835,920,476,500]" name="Mexico" pageId="10" pageNumber="38">Mexico</collectingCountry>
and
<taxonomicName id="4C304D1FFFDFFF8B5BF0C0A4FAAC398F" authority="Pilato &amp; Kaczmarek, 2007" authorityName="Pilato &amp; Kaczmarek" authorityYear="2007" box="[977,1406,476,500]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="10" pageNumber="38" phylum="Tardigrada" rank="species" species="alvaroi">
<emphasis id="B944EA8EFFDFFF8B5BF0C0A4FB9A398F" box="[977,1096,476,500]" italics="true" pageId="10" pageNumber="38">M. alvaroi</emphasis>
<bibRefCitation id="EFA14B6DFFDFFF8B5C71C0A5FAAC398F" author="Pilato" box="[1104,1406,476,500]" pageId="10" pageNumber="38" refString="Pilato, G. &amp; Kaczmarek, L. (2007) Macrobiotus alvaroi, a new species of eutardigrade (Tardigrada, Macrobiotidae) of the polyopus group from Costa Rica (Central America). Zootaxa, 1479, 1 - 7." type="journal article" year="2007">
Pilato &amp; Kaczmarek,
<quantity id="4CC89B79FFDFFF8B5D67C0A5FAAC398F" box="[1350,1406,476,500]" metricMagnitude="1" metricUnit="m" metricValue="5.097779999999999" pageId="10" pageNumber="38" unit="in" value="2007.0">2007</quantity>
</bibRefCitation>
</taxonomicName>
in
<collectingCountry id="F327760CFFDFFF8B58B6C086FEC63A6C" box="[151,276,511,535]" name="Costa Rica" pageId="10" pageNumber="38">Costa Rica</collectingCountry>
.
</paragraph>
<paragraph id="8B8F369CFFDFFF8B58E4C35DFB513C2C" blockId="10.[151,1437,151,1112]" pageId="10" pageNumber="38">
<taxonomicName id="4C304D1FFFDFFF8B58E4C35DFE393A47" box="[197,491,548,572]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="10" pageNumber="38" phylum="Tardigrada" rank="species" species="caymanensis" status="sp. nov.">
<emphasis id="B944EA8EFFDFFF8B58E4C35DFE393A47" box="[197,491,548,572]" italics="true" pageId="10" pageNumber="38">Macrobiotus caymanensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27757F5FFDFFF8B59D5C35DFDE43A47" box="[500,566,548,572]" pageId="10" pageNumber="38" rank="species">
<emphasis id="B944EA8EFFDFFF8B59D5C35DFDE43A47" bold="true" box="[500,566,548,572]" pageId="10" pageNumber="38">sp. n.</emphasis>
</taxonomicNameLabel>
differs from other known species in the
<taxonomicName id="4C304D1FFFDFFF8B5C2DC35DFBA33A47" box="[1036,1137,548,572]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="10" pageNumber="38" phylum="Tardigrada" rank="species" species="polyopus">
<emphasis id="B944EA8EFFDFFF8B5C2DC35DFBA33A47" box="[1036,1137,548,572]" italics="true" pageId="10" pageNumber="38">polyopus</emphasis>
</taxonomicName>
group in having a shorter buccal tube (28.2 μm long in a
<taxonomicName id="4C304D1FFFDFFF8B59D3C330FD743A1B" box="[498,678,585,608]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="10" pageNumber="38" phylum="Tardigrada" rank="species" species="caymanensis" status="sp. nov.">
<emphasis id="B944EA8EFFDFFF8B59D3C330FD743A1B" box="[498,678,585,608]" italics="true" pageId="10" pageNumber="38">M. caymanensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27757F5FFDFFF8B5A8DC331FD393A1B" box="[684,747,584,608]" pageId="10" pageNumber="38" rank="species">
<emphasis id="B944EA8EFFDFFF8B5A8DC331FD393A1B" bold="true" box="[684,747,584,608]" pageId="10" pageNumber="38">sp. n.</emphasis>
</taxonomicNameLabel>
specimen 282.1 μm long, 29.2–38.8 μm in specimens of similar length from the other species) and a more posterior stylet support insertion (
<emphasis id="B944EA8EFFDFFF8B5C32C314FBB33AFF" box="[1043,1121,620,644]" italics="true" pageId="10" pageNumber="38">
pt
<quantity id="4CC89B79FFDFFF8B5C11C315FBB33AFF" box="[1072,1121,620,644]" metricMagnitude="0" metricUnit="m" metricValue="1.98882" pageId="10" pageNumber="38" unit="in" value="78.3">78.3</quantity>
</emphasis>
in the new species,
<emphasis id="B944EA8EFFDFFF8B5D61C314FF1A3AD3" italics="true" pageId="10" pageNumber="38">
pt
<quantity id="4CC89B79FFDFFF8B5D7CC315FF1A3AD3" metricMagnitude="0" metricUnit="m" metricValue="1.83515" metricValueMax="1.86436" metricValueMin="1.80594" pageId="10" pageNumber="38" unit="in" value="72.25" valueMax="73.4" valueMin="71.1">71.1– 73.4</quantity>
</emphasis>
in the other described species).
<taxonomicName id="4C304D1FFFDFFF8B5A15C3E9FC8B3AD3" box="[564,857,656,680]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="10" pageNumber="38" phylum="Tardigrada" rank="species" species="caymanensis" status="sp. nov.">
<emphasis id="B944EA8EFFDFFF8B5A15C3E9FC8B3AD3" box="[564,857,656,680]" italics="true" pageId="10" pageNumber="38">Macrobiotus caymanensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27757F5FFDFFF8B5B40C3E9FC733AD3" box="[865,929,656,680]" pageId="10" pageNumber="38" rank="species">
<emphasis id="B944EA8EFFDFFF8B5B40C3E9FC733AD3" bold="true" box="[865,929,656,680]" pageId="10" pageNumber="38">sp. n.</emphasis>
</taxonomicNameLabel>
further differs from
<taxonomicName id="4C304D1FFFDFFF8B5CAAC3E8FAC43AD3" box="[1163,1302,656,680]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="10" pageNumber="38" phylum="Tardigrada" rank="species" species="polyopus">
<emphasis id="B944EA8EFFDFFF8B5CAAC3E8FAC43AD3" box="[1163,1302,656,680]" italics="true" pageId="10" pageNumber="38">M. polyopus</emphasis>
</taxonomicName>
in having a microplacoid, more robust claws, stronger accessory points and fewer processes on the egg (18 around the circumference and
<quantity id="4CC89B79FFDFFF8B593EC3AEFE853A94" box="[287,343,727,751]" metricMagnitude="-1" metricUnit="m" metricValue="9.906" pageId="10" pageNumber="38" unit="in" value="39.0">39 in</quantity>
the hemisphere in
<taxonomicName id="4C304D1FFFDFFF8B5A0BC3A0FD663A8B" box="[554,692,728,752]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="10" pageNumber="38" phylum="Tardigrada" rank="species" species="polyopus">
<emphasis id="B944EA8EFFDFFF8B5A0BC3A0FD663A8B" box="[554,692,728,752]" italics="true" pageId="10" pageNumber="38">M. polyopus</emphasis>
</taxonomicName>
, 9–12 around the circumference and
<quantity id="4CC89B79FFDFFF8B5C73C3AEFB753A94" box="[1106,1191,727,751]" metricMagnitude="-1" metricUnit="m" metricValue="2.794" metricValueMax="3.302" metricValueMin="2.286" pageId="10" pageNumber="38" unit="in" value="11.0" valueMax="13.0" valueMin="9.0">9–13 in</quantity>
the hemisphere in the new species). It differs from
<taxonomicName id="4C304D1FFFDFFF8B59FDC384FDAD3B6F" box="[476,639,764,788]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="10" pageNumber="38" phylum="Tardigrada" rank="species" species="mandalaae">
<emphasis id="B944EA8EFFDFFF8B59FDC384FDAD3B6F" box="[476,639,764,788]" italics="true" pageId="10" pageNumber="38">M. mandalaae</emphasis>
</taxonomicName>
in having a wider buccal tube (
<emphasis id="B944EA8EFFDFFF8B5BC5C384FBE03B6F" box="[996,1074,764,788]" italics="true" pageId="10" pageNumber="38">
pt
<quantity id="4CC89B79FFDFFF8B5C20C385FBE03B6F" box="[1025,1074,764,788]" metricMagnitude="-1" metricUnit="m" metricValue="3.1496" pageId="10" pageNumber="38" unit="in" value="12.4">12.4</quantity>
</emphasis>
in
<taxonomicName id="4C304D1FFFDFFF8B5C76C384FB2B3B6F" box="[1111,1273,764,788]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="10" pageNumber="38" phylum="Tardigrada" rank="species" species="mandalaae">
<emphasis id="B944EA8EFFDFFF8B5C76C384FB2B3B6F" box="[1111,1273,764,788]" italics="true" pageId="10" pageNumber="38">M. mandalaae</emphasis>
</taxonomicName>
and
<emphasis id="B944EA8EFFDFFF8B5D11C384FAAD3B6F" box="[1328,1407,764,788]" italics="true" pageId="10" pageNumber="38">
pt
<quantity id="4CC89B79FFDFFF8B5D6FC385FAAD3B6F" box="[1358,1407,764,788]" metricMagnitude="-1" metricUnit="m" metricValue="3.9878" pageId="10" pageNumber="38" unit="in" value="15.7">15.7</quantity>
</emphasis>
in the new species), in lacking leg granulation and in having fewer and larger egg process (24–26 around the circumference,
<quantity id="4CC89B79FFDFFF8B58D8C23DFE8F3B27" box="[249,349,836,860]" metricMagnitude="0" metricUnit="m" metricValue="1.8415" metricValueMax="1.905" metricValueMin="1.778" pageId="10" pageNumber="38" unit="in" value="72.5" valueMax="75.0" valueMin="70.0">70–75 in</quantity>
the hemisphere, 3.4–4.7 μm high, 3.2–4.5 μm wide in
<taxonomicName id="4C304D1FFFDFFF8B5BF0C23CFBA63B27" box="[977,1140,836,860]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="10" pageNumber="38" phylum="Tardigrada" rank="species" species="mandalaae">
<emphasis id="B944EA8EFFDFFF8B5BF0C23CFBA63B27" box="[977,1140,836,860]" italics="true" pageId="10" pageNumber="38">M. mandalaae</emphasis>
</taxonomicName>
; 8.3–20.0 μm high, 12.3– 19.2 μm wide in
<taxonomicName id="4C304D1FFFDFFF8B5972C210FDD53BFB" box="[339,519,873,896]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="10" pageNumber="38" phylum="Tardigrada" rank="species" species="caymanensis" status="sp. nov.">
<emphasis id="B944EA8EFFDFFF8B5972C210FDD53BFB" box="[339,519,873,896]" italics="true" pageId="10" pageNumber="38">M. caymanensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27757F5FFDFFF8B5A2FC211FD9F3BFB" box="[526,589,872,896]" pageId="10" pageNumber="38" rank="species">
<emphasis id="B944EA8EFFDFFF8B5A2FC211FD9F3BFB" bold="true" box="[526,589,872,896]" pageId="10" pageNumber="38">sp. n.</emphasis>
</taxonomicNameLabel>
).
<taxonomicName id="4C304D1FFFDFFF8B5A45C211FC553BFB" box="[612,903,872,896]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="10" pageNumber="38" phylum="Tardigrada" rank="species" species="caymanensis" status="sp. nov.">
<emphasis id="B944EA8EFFDFFF8B5A45C211FC553BFB" box="[612,903,872,896]" italics="true" pageId="10" pageNumber="38">Macrobiotus caymanensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27757F5FFDFFF8B5BAFC211FC1F3BFB" box="[910,973,872,896]" pageId="10" pageNumber="38" rank="species">
<emphasis id="B944EA8EFFDFFF8B5BAFC211FC1F3BFB" bold="true" box="[910,973,872,896]" pageId="10" pageNumber="38">sp. n.</emphasis>
</taxonomicNameLabel>
differs from
<taxonomicName id="4C304D1FFFDFFF8B5C7EC210FB343BFB" box="[1119,1254,872,896]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="10" pageNumber="38" phylum="Tardigrada" rank="species" species="insularis">
<emphasis id="B944EA8EFFDFFF8B5C7EC210FB343BFB" box="[1119,1254,872,896]" italics="true" pageId="10" pageNumber="38">M. insularis</emphasis>
</taxonomicName>
in having fewer and larger egg processes (
<quantity id="4CC89B79FFDFFF8B5999C2F5FE203BDF" box="[440,498,908,932]" metricMagnitude="-1" metricUnit="m" metricValue="8.89" pageId="10" pageNumber="38" unit="in" value="35.0">35 in</quantity>
the hemisphere, 7.7–8.8 μm high and 10.9–13.2 μm wide in
<taxonomicName id="4C304D1FFFDFFF8B5CBDC2F4FAF63BDF" box="[1180,1316,908,932]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="10" pageNumber="38" phylum="Tardigrada" rank="species" species="insularis">
<emphasis id="B944EA8EFFDFFF8B5CBDC2F4FAF63BDF" box="[1180,1316,908,932]" italics="true" pageId="10" pageNumber="38">M. insularis</emphasis>
</taxonomicName>
). The new species differs from
<taxonomicName id="4C304D1FFFDFFF8B595CC2C8FDD93BB3" box="[381,523,945,968]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="10" pageNumber="38" phylum="Tardigrada" rank="species" species="ocotensis">
<emphasis id="B944EA8EFFDFFF8B595CC2C8FDD93BB3" box="[381,523,945,968]" italics="true" pageId="10" pageNumber="38">M. ocotensis</emphasis>
</taxonomicName>
in lacking leg granulation, in having a wider buccal tube (
<emphasis id="B944EA8EFFDFFF8B5C81C2C8FB643BB3" box="[1184,1206,945,968]" italics="true" pageId="10" pageNumber="38">pt</emphasis>
<quantity id="4CC89B79FFDFFF8B5C9CC2D6FAEF3BBC" box="[1213,1341,943,967]" metricMagnitude="-1" metricUnit="m" metricValue="2.3876" metricValueMax="2.6924" metricValueMin="2.0827999999999998" pageId="10" pageNumber="38" unit="in" value="9.399999999999999" valueMax="10.6" valueMin="8.2">8.2–10.6 in</quantity>
<taxonomicName id="4C304D1FFFDFFF8B5D65C2C8FF053B97" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="10" pageNumber="38" phylum="Tardigrada" rank="species" species="ocotensis">
<emphasis id="B944EA8EFFDFFF8B5D65C2C8FF053B97" italics="true" pageId="10" pageNumber="38">M. ocotensis</emphasis>
</taxonomicName>
and
<quantity id="4CC89B79FFDFFF8B592FC2ADFE4E3B97" box="[270,412,980,1004]" metricMagnitude="-1" metricUnit="m" metricValue="4.0512999999999995" metricValueMax="4.6228" metricValueMin="3.4798" pageId="10" pageNumber="38" unit="in" value="15.95" valueMax="18.2" valueMin="13.7">13.7–18.2 in</quantity>
the new species) and fewer and larger egg processes (25 around the circumference,
<quantity id="4CC89B79FFDFFF8B5D75C2ADFA4E3B97" box="[1364,1436,980,1004]" metricMagnitude="0" metricUnit="m" metricValue="2.54" pageId="10" pageNumber="38" unit="in" value="100.0">100 in</quantity>
the hemisphere, 4.7–6.0 μm high and 4.9–6.0 μm wide in
<taxonomicName id="4C304D1FFFDFFF8B5B3AC280FC7A3C6B" box="[795,936,1017,1040]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="10" pageNumber="38" phylum="Tardigrada" rank="species" species="ocotensis">
<emphasis id="B944EA8EFFDFFF8B5B3AC280FC7A3C6B" box="[795,936,1017,1040]" italics="true" pageId="10" pageNumber="38">M. ocotensis</emphasis>
</taxonomicName>
). The new species differs from
<taxonomicName id="4C304D1FFFDFFF8B5D2AC280FA523C6B" box="[1291,1408,1016,1040]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="10" pageNumber="38" phylum="Tardigrada" rank="species" species="alvaroi">
<emphasis id="B944EA8EFFDFFF8B5D2AC280FA523C6B" box="[1291,1408,1016,1040]" italics="true" pageId="10" pageNumber="38">M. alvaroi</emphasis>
</taxonomicName>
in having a wider buccal tube (
<emphasis id="B944EA8EFFDFFF8B59F6C564FE3F3C4F" box="[471,493,1053,1076]" italics="true" pageId="10" pageNumber="38">pt</emphasis>
<quantity id="4CC89B79FFDFFF8B59D5C565FD533C4F" box="[500,641,1052,1076]" metricMagnitude="-1" metricUnit="m" metricValue="2.667" metricValueMax="2.7686" metricValueMin="2.5654" pageId="10" pageNumber="38" unit="in" value="10.5" valueMax="10.9" valueMin="10.1">10.1–10.9 in</quantity>
<taxonomicName id="4C304D1FFFDFFF8B5AA9C564FD2C3C4F" box="[648,766,1052,1076]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="10" pageNumber="38" phylum="Tardigrada" rank="species" species="alvaroi">
<emphasis id="B944EA8EFFDFFF8B5AA9C564FD2C3C4F" box="[648,766,1052,1076]" italics="true" pageId="10" pageNumber="38">M. alvaroi</emphasis>
</taxonomicName>
) and fewer and larger egg processes (23–26 around the circumference,
<quantity id="4CC89B79FFDFFF8B5906C546FEBC3C2C" box="[295,366,1087,1111]" metricMagnitude="0" metricUnit="m" metricValue="2.54" pageId="10" pageNumber="38" unit="in" value="100.0">100 in</quantity>
the hemisphere, 3.6–6.1 μm high and 3.3–4.1 μm wide in
<taxonomicName id="4C304D1FFFDFFF8B5BDCC538FBA13C23" box="[1021,1139,1088,1112]" class="Eutardigrada" family="Macrobiotidae" genus="Macrobiotus" kingdom="Animalia" order="Parachela" pageId="10" pageNumber="38" phylum="Tardigrada" rank="species" species="alvaroi">
<emphasis id="B944EA8EFFDFFF8B5BDCC538FBA13C23" box="[1021,1139,1088,1112]" italics="true" pageId="10" pageNumber="38">M. alvaroi</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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