779 lines
130 KiB
XML
779 lines
130 KiB
XML
<document id="929B932B9F07E58B20BF872D89100834" ID-DOI="10.4202/app.00652.2019" ID-ISSN="1732-2421" ID-Zenodo-Dep="10626640" IM.bibliography_approvedBy="karina" IM.illustrations_approvedBy="karina" IM.taxonomicNames_approvedBy="karina" IM.treatments_approvedBy="karina" checkinTime="1707250169927" checkinUser="felipe" docAuthor="Müller, Rodrigo T., Baczko, M. Belén Von, Desojo, Julia B. & Nesbitt, Sterling J." docDate="2020" docId="038D87A7FFC8B72BFF39FA804852FA89" docLanguage="en" docName="ActaPalaeontolPol.65.1.1-10.pdf" docOrigin="Acta Palaeontologica Polonica 65 (1)" docSource="http://dx.doi.org/10.4202/app.00652.2019" docStyle="DocumentStyle:E4780F6D34569985EDF9CACFB9BC780E.4:ActaPalaeontolPol.2015-.journal_article" docStyleId="E4780F6D34569985EDF9CACFB9BC780E" docStyleName="ActaPalaeontolPol.2015-.journal_article" docStyleVersion="4" docTitle="Dynamosuchus collisensis Müller & Baczko & Desojo & Nesbitt 2020, sp. nov." docType="treatment" docVersion="4" lastPageNumber="7" masterDocId="FFB4FFDFFFCAB72DFFAEFFFC497FFFD0" masterDocTitle="The first ornithosuchid from Brazil and its macroevolutionary and phylogenetic implications for Late Triassic faunas in Gondwana" masterLastPageNumber="10" masterPageNumber="1" pageNumber="3" updateTime="1714422813977" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
|
||
<mods:mods id="4B036D56E0D1395AA2DA4492AF998A7E" xmlns:mods="http://www.loc.gov/mods/v3">
|
||
<mods:titleInfo id="98D0608C4B93B7B1D84A2815639CF6F3">
|
||
<mods:title id="7055A89993EA110A2B4BEC944D1AC7DD">The first ornithosuchid from Brazil and its macroevolutionary and phylogenetic implications for Late Triassic faunas in Gondwana</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:name id="9AA8A6E34A020B1D326E69954A408B0E" type="personal">
|
||
<mods:role id="6D4FD1017F1CE17081E9DB8F78DFF3BE">
|
||
<mods:roleTerm id="2EA2E7F54C24EE999E7E5F25AB6D7A5F">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="4B2500C2591CA34CFFF5C12F5C0E9D87">Müller, Rodrigo T.</mods:namePart>
|
||
</mods:name>
|
||
<mods:name id="0D3A8B45F483AE63322B6D98533C5957" type="personal">
|
||
<mods:role id="7E06709560DE9F642D15B76D82FBF4FF">
|
||
<mods:roleTerm id="73639643BE033B58ED64ECAD0ACED835">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="9436904A5AF231D8307DA8A4F3CEB4DB">Baczko, M. Belén Von</mods:namePart>
|
||
</mods:name>
|
||
<mods:name id="498B0F804C33DEB6F80C9FF1F78CF31A" type="personal">
|
||
<mods:role id="2842A6DDDA154A9C226BB1788FAEF8A5">
|
||
<mods:roleTerm id="5C3AD5E7B956AB88F6835FE19F6C9605">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="0342E31107F03C0FFB03F77043569F07">Desojo, Julia B.</mods:namePart>
|
||
</mods:name>
|
||
<mods:name id="333DF4F4B7245EB2D2018C46B71B75B4" type="personal">
|
||
<mods:role id="9C37058B630DA8C31B6BAD62CC1F197C">
|
||
<mods:roleTerm id="546EBEF997359EA591C920E91F291E63">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="61780D762707A15AAE4C3E9B167FCA67">Nesbitt, Sterling J.</mods:namePart>
|
||
</mods:name>
|
||
<mods:typeOfResource id="FF0C36609026CBB222298575B51F7976">text</mods:typeOfResource>
|
||
<mods:relatedItem id="792F90A653E343905CA1D9D27B00528E" type="host">
|
||
<mods:titleInfo id="ACA48EF0D474D20A265B71AFDD8B4B04">
|
||
<mods:title id="E5F8A84589A36B6C13D7D4D25EA36EF1">Acta Palaeontologica Polonica</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:part id="E492D01B5006AE56322A7EE8EC046846">
|
||
<mods:date id="3C829AFD2D0F4786DA46A1F8FBBE4811">2020</mods:date>
|
||
<mods:detail id="2F14E14C217095FB08793FB3CE1EA7E8" type="pubDate">
|
||
<mods:number id="5751ADAD8AAEE186EEF9506D584A5390">2020-01-31</mods:number>
|
||
</mods:detail>
|
||
<mods:detail id="34F5893D23E3F402BB91F6E79E55CD42" type="volume">
|
||
<mods:number id="B590C39BECF4EBB4B1D0D020AEA7293E">65</mods:number>
|
||
</mods:detail>
|
||
<mods:detail id="58C6294D1ED8AB9FCC5146DB03E8EF75" type="issue">
|
||
<mods:number id="71F5B72FDAA364B910F3728DCDD3EC86">1</mods:number>
|
||
</mods:detail>
|
||
<mods:extent id="56AD3DDA706D5161A52671EB84A91D63" unit="page">
|
||
<mods:start id="FEBD9434F263E27B7895614136162B3C">1</mods:start>
|
||
<mods:end id="B7CCECA447754C060DDD05452DC93F9B">10</mods:end>
|
||
</mods:extent>
|
||
</mods:part>
|
||
</mods:relatedItem>
|
||
<mods:location id="13B71C93188D952582130326F71F52D4">
|
||
<mods:url id="26352A2EC2390DEA754906EF3889791B">http://dx.doi.org/10.4202/app.00652.2019</mods:url>
|
||
</mods:location>
|
||
<mods:classification id="7B6A409EEC28534DDF5F8990949A1CFB">journal article</mods:classification>
|
||
<mods:identifier id="EE7A3F2586F6D82A2DBB001C8FEC87FA" type="DOI">10.4202/app.00652.2019</mods:identifier>
|
||
<mods:identifier id="AC4473D6B81C403E3FC71E1376389496" type="ISSN">1732-2421</mods:identifier>
|
||
<mods:identifier id="3BC9B6D0AB04B85CF94D7CC15C5AE6FC" type="Zenodo-Dep">10626640</mods:identifier>
|
||
<mods:identifier id="5AED073C5596DD6E2B086A14A826941D" type="ZooBank">C3CCF3E0-B3C3-48B7-BCB9-AF493D9655E9</mods:identifier>
|
||
</mods:mods>
|
||
<treatment id="038D87A7FFC8B72BFF39FA804852FA89" ID-DOI="http://doi.org/10.5281/zenodo.11088360" ID-Zenodo-Dep="11088360" LSID="urn:lsid:plazi:treatment:038D87A7FFC8B72BFF39FA804852FA89" httpUri="http://treatment.plazi.org/id/038D87A7FFC8B72BFF39FA804852FA89" lastPageId="6" lastPageNumber="7" pageId="2" pageNumber="3">
|
||
<subSubSection id="C33E653AFFC8B72FFF39FA8049AAFA6E" pageId="2" pageNumber="3" type="nomenclature">
|
||
<paragraph id="8B9B36B1FFC8B72FFF39FA804B2CFA48" blockId="2.[151,595,1404,1432]" box="[151,595,1404,1432]" pageId="2" pageNumber="3">
|
||
<heading id="D0D381DDFFC8B72FFF39FA804B2CFA48" box="[151,595,1404,1432]" fontSize="12" level="2" pageId="2" pageNumber="3" reason="2">
|
||
<taxonomicName id="4C244D32FFC8B72FFF39FA804897FA48" authority="Müller & Baczko & Desojo & Nesbitt, 2020" authorityName="Müller & Baczko & Desojo & Nesbitt" authorityYear="2020" box="[151,488,1404,1432]" class="Reptilia" family="Ornithosuchidae" genus="Dynamosuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Pseudosuchia" pageId="2" pageNumber="3" phylum="Chordata" rank="species" species="collisensis" status="sp. nov.">
|
||
<emphasis id="B950EAA3FFC8B72FFF39FA804897FA48" box="[151,488,1404,1432]" italics="true" pageId="2" pageNumber="3">Dynamosuchus collisensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A26357D8FFC8B72FFE5EFA804B2CFA48" box="[496,595,1404,1432]" pageId="2" pageNumber="3" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8B9B36B1FFC8B72FFF39FA5549AAFA6E" blockId="2.[151,213,1449,1470]" box="[151,213,1449,1470]" pageId="2" pageNumber="3">
|
||
<figureCitation id="131F2A34FFC8B72FFF39FA5549AEFA6E" box="[151,209,1449,1470]" captionStart="→" captionStartId="4.[116,144,1757,1787]" captionTargetId="figure-8@3.[98,1479,161,1951]" captionTargetPageId="3" captionText="→ Fig. 2. Representative skeletal elements of ornithosuchid archosaur Dynamosuchus collisensis gen. et sp. nov. (CAPPA/UFSM 0248) from Janner outcrop, Carnian, Late Triassic. A. Selected skull bones in left lateral view. B. Reconstruction of the skull. C. Skull in ventral view. D. Left quadrate and quadratojugal in posterodorsal view. E. Parabasisphenoid in left lateral view. F. Neural arch of an anterior cervical vertebra in anterior view. G. Centrum of a cervical vertebra in left lateral view. H. Right osteoderm in dorsal view. I. Neural arch of an anterior dorsal vertebra in left lateral view. J. Left ilium in lateral view. L. Right humerus in anterior view. M. Right forearm in medial view. N. Left manus in dorsal view. O. Right (reversed) pubis in lateral view. P. Left femur in anterior view. Q. Left fibula in lateral view. Some unpreserved portions are modified from Baczko et al. in press, for the reconstruction of the skeleton of CAPPA/UFSM 0248 (preserved elements indicated in orange) (K). Scale bars 20 mm." figureDoi="http://doi.org/10.5281/zenodo.11088354" httpUri="https://zenodo.org/record/11088354/files/figure.png" pageId="2" pageNumber="3">Fig. 2</figureCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C33E653AFFC8B72BFF39FA2C4852FA89" lastPageId="6" lastPageNumber="7" pageId="2" pageNumber="3" type="description">
|
||
<paragraph id="8B9B36B1FFC8B72FFF39FA2C484AF9D1" blockId="2.[151,824,1488,1537]" pageId="2" pageNumber="3">
|
||
<emphasis id="B950EAA3FFC8B72FFF39FA2C485AFA35" box="[151,293,1488,1509]" italics="true" pageId="2" pageNumber="3">ZooBank LSID</emphasis>
|
||
:
|
||
<uri id="FFB53AB3FFC8B72FFE9FFA2C484AF9D1" pageId="2" pageNumber="3">
|
||
urn:lsid:zoobank.org:act:
|
||
<uuid id="FF820C64FFC8B72FFDB6FA2C484AF9D1" pageId="2" pageNumber="3">4B1FFAFB-6287-4448-9E34- B7976D307C80</uuid>
|
||
</uri>
|
||
</paragraph>
|
||
<paragraph id="8B9B36B1FFC8B72FFF39F9F24B51F98C" blockId="2.[151,824,1550,1628]" pageId="2" pageNumber="3">
|
||
<emphasis id="B950EAA3FFC8B72FFF39F9F24981F9F3" box="[151,254,1550,1571]" italics="true" pageId="2" pageNumber="3">Etymology</emphasis>
|
||
: From Latin
|
||
<emphasis id="B950EAA3FFC8B72FFED3F9F248CEF9F3" box="[381,433,1550,1571]" italics="true" pageId="2" pageNumber="3">collis</emphasis>
|
||
, hill, reflecting the fact that the
|
||
<typeStatus id="549F8813FFC8B72FFD4AF9F24A47F9F3" box="[740,824,1550,1571]" pageId="2" pageNumber="3">holotype</typeStatus>
|
||
was collected from the Janner site, which is located at the base of the Agudo Hill (Morro Agudo in Portuguese).
|
||
</paragraph>
|
||
<paragraph id="8B9B36B1FFC8B72FFF39F9954B46F8DD" blockId="2.[151,824,1641,1805]" pageId="2" pageNumber="3">
|
||
<materialsCitation id="3B4C3CECFFC8B72FFF39F9954B46F8DD" collectionCode="CAPPA" pageId="2" pageNumber="3" specimenCode="UFSM 0248" specimenCount="1" typeStatus="holotype">
|
||
<emphasis id="B950EAA3FFC8B72FFF39F9954990F9AE" box="[151,239,1641,1662]" italics="true" pageId="2" pageNumber="3">
|
||
<typeStatus id="549F8813FFC8B72FFF39F9954990F9AE" box="[151,239,1641,1662]" pageId="2" pageNumber="3">Holotype</typeStatus>
|
||
</emphasis>
|
||
:
|
||
<collectionCode id="ED35AE74FFC8B72FFF50F9954834F9AE" box="[254,331,1641,1662]" pageId="2" pageNumber="3">CAPPA</collectionCode>
|
||
/
|
||
<specimenCode id="DB829ECAFFC8B72FFEFCF99548B3F9AE" box="[338,460,1641,1662]" collectionCode="UFSM" pageId="2" pageNumber="3">UFSM 0248</specimenCode>
|
||
, a partial skeleton including the premaxillae, left maxilla, frontals, right postfrontal, right parietal, squamosals, quadratojugals, quadrates, parabasisphenoid, both hemimandibles, some cervical, dorsal, sacral and caudal vertebrae, several ribs, gastralia, osteoderms, left scapula, both forelimbs, left ilium, right pubis, left femur, right tibia, and left fibula.
|
||
</materialsCitation>
|
||
</paragraph>
|
||
<paragraph id="8B9B36B1FFC8B72FFF39F8E6489AF89B" blockId="2.[151,824,1818,1867]" pageId="2" pageNumber="3">
|
||
<emphasis id="B950EAA3FFC8B72FFF39F8E6486BF8FF" box="[151,276,1818,1839]" italics="true" pageId="2" pageNumber="3">Type locality</emphasis>
|
||
:
|
||
<materialsCitation id="3B4C3CECFFC8B72FFE8DF8E6489EF89B" country="Brazil" latitude="-29.653025" location="Agudo" longLatPrecision="1" longitude="-53.29283" pageId="2" pageNumber="3" specimenCount="1" stateProvince="Rio Grande do Sul">
|
||
Janner site (
|
||
<geoCoordinate id="EE105076FFC8B72FFE35F8E64B4DF8FF" box="[411,562,1818,1839]" degrees="29" direction="south" minutes="39" orientation="latitude" pageId="2" pageNumber="3" precision="1" seconds="10.89" value="-29.653025">
|
||
29°39
|
||
<emphasis id="B950EAA3FFC8B72FFE7BF8E648A5F8FF" bold="true" box="[469,474,1818,1839]" italics="true" pageId="2" pageNumber="3">′</emphasis>
|
||
10.89
|
||
<emphasis id="B950EAA3FFC8B72FFDBFF8E64B64F8FF" bold="true" box="[529,539,1818,1839]" italics="true" pageId="2" pageNumber="3">″</emphasis>
|
||
S
|
||
</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE105076FFC8B72FFD90F8E64BA0F8FF" box="[574,735,1818,1839]" degrees="53" direction="west" minutes="17" orientation="longitude" pageId="2" pageNumber="3" precision="1" seconds="34.20" value="-53.29283">
|
||
53°17
|
||
<emphasis id="B950EAA3FFC8B72FFDD6F8E64B02F8FF" bold="true" box="[632,637,1818,1839]" italics="true" pageId="2" pageNumber="3">′</emphasis>
|
||
34.20
|
||
<emphasis id="B950EAA3FFC8B72FFD1DF8E64BC2F8FF" bold="true" box="[691,701,1818,1839]" italics="true" pageId="2" pageNumber="3">″</emphasis>
|
||
W
|
||
</geoCoordinate>
|
||
),
|
||
<location id="8EFB606AFFC8B72FFD5FF8E64A4AF8FF" LSID="urn:lsid:plazi:treatment:038D87A7FFC8B72BFF39FA804852FA89:8EFB606AFFC8B72FFD5FF8E64A4AF8FF" box="[753,821,1818,1839]" country="Brazil" latitude="-29.653025" longLatPrecision="1" longitude="-53.29283" name="Agudo" pageId="2" pageNumber="3" stateProvince="Rio Grande do Sul">Agudo</location>
|
||
,
|
||
<collectingRegion id="49E0F853FFC8B72FFF39F8CA4833F89B" box="[151,332,1846,1867]" country="Brazil" name="Rio Grande do Sul" pageId="2" pageNumber="3">Rio Grande do Sul</collectingRegion>
|
||
,
|
||
<collectingCountry id="F3337621FFC8B72FFEF9F8CA48EDF89B" box="[343,402,1846,1867]" name="Brazil" pageId="2" pageNumber="3">Brazil</collectingCountry>
|
||
(
|
||
<figureCitation id="131F2A34FFC8B72FFE30F8CA48A9F89B" box="[414,470,1846,1867]" captionStart="Fig" captionStartId="1.[98,131,1004,1024]" captionTargetBox="[104,1479,167,974]" captionTargetId="figure-385@1.[100,1479,161,975]" captionTargetPageId="1" captionText="Fig. 1. A. Location map of the Janner site and the surface distribution of the geologic units in the area (modified from Müller et al. 2017). B. Stratigraphic column of the Janner site (modified from Da-Rosa 2015) depicting its fossiliferous content: 1, Exaeretodon; 2, Hyperodapedon; 3, Pampadromaeus; 4, Trucidocynodon; 5, Bagualosaurus; and 6, the new ornithosuchid." figureDoi="http://doi.org/10.5281/zenodo.10626642" httpUri="https://zenodo.org/record/10626642/files/figure.png" pageId="2" pageNumber="3">Fig. 1</figureCitation>
|
||
)
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8B9B36B1FFC8B72FFF39F8A44CB3FF3D" blockId="2.[151,824,1880,1958]" lastBlockId="2.[862,1535,160,237]" pageId="2" pageNumber="3">
|
||
<emphasis id="B950EAA3FFC8B72FFF39F8A44869F8BD" box="[151,278,1880,1901]" italics="true" pageId="2" pageNumber="3">Type horizon</emphasis>
|
||
: Santa Maria Formation; Candelária Sequence, Paraná Basin. The presence of the genus
|
||
<taxonomicName id="4C244D32FFC8B72FFE73F8894B08F85A" authorityName="Huxley" authorityYear="1859" box="[477,631,1909,1930]" class="Reptilia" family="Rhynchosauridae" genus="Hyperodapedon" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="2" pageNumber="3" phylum="Chordata" rank="genus">
|
||
<emphasis id="B950EAA3FFC8B72FFE73F8894B08F85A" box="[477,631,1909,1930]" italics="true" pageId="2" pageNumber="3">Hyperodapedon</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C244D32FFC8B72FFD08F8894A5FF85A" authorityName="Cabrera" authorityYear="1943" box="[678,800,1909,1930]" class="Reptilia" family="Traversodontidae" genus="Exaeretodon" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="2" pageNumber="3" phylum="Chordata" rank="genus">
|
||
<emphasis id="B950EAA3FFC8B72FFD08F8894A5FF85A" box="[678,800,1909,1930]" italics="true" pageId="2" pageNumber="3">Exaeretodon</emphasis>
|
||
</taxonomicName>
|
||
at the Janner site (
|
||
<bibRefCitation id="EFB54B40FFC8B72FFE9DF86D4895F876" author="Langer, M. C. & Ribeiro, A. M. & Schultz, C. L. & Ferigolo, J." box="[307,490,1937,1958]" pageId="2" pageNumber="3" pagination="201 - 218" refId="ref8527" refString="Langer, M. C., Ribeiro, A. M., Schultz, C. L., and Ferigolo, J. 2007. The continental tetrapodbearing Triassic of south Brazil. New Mexico Museum of Natural History & Science Bulletin 41: 201 - 218." type="journal article" year="2007">Langer et al. 2007</bibRefCitation>
|
||
) supports a Carnian age based on correlation with radioisotope dating of another Brazilian site (
|
||
<bibRefCitation id="EFB54B40FFC8B72FFA17FF5C4AB8FF01" author="Langer, M. C. & Ramezani, J. & Da-Rosa, A. A. S." pageId="2" pageNumber="3" pagination="133 - 140" refId="ref8486" refString="Langer, M. C., Ramezani, J., and Da-Rosa, A. A. S. 2018. U-Pb age constraints on dinosaur rise from south Brazil. Gondwana Research 57: 133 - 140." type="journal article" year="2018">Langer et al. 2018</bibRefCitation>
|
||
) and
|
||
<collectingCountry id="F3337621FFC8B72FFC50FF404D21FF01" box="[1022,1118,188,209]" name="Argentina" pageId="2" pageNumber="3">Argentine</collectingCountry>
|
||
beds from the La Peña Member of the Ischigualasto Formation (
|
||
<bibRefCitation id="EFB54B40FFC8B72FFBEFFF244C7BFF3D" author="Martinez, R. N. & Sereno, P. C. & Alcober, O. A. & Colombi, C. E. & Renne, P. R. & Montanez, I. P. & Currie, B. S." box="[1089,1284,216,237]" pageId="2" pageNumber="3" pagination="206 - 210" refId="ref8630" refString="Martinez, R. N., Sereno, P. C., Alcober, O. A., Colombi, C. E., Renne, P. R., Montanez, I. P., and Currie, B. S. 2011. A basal dinosaur from the dawn of the dinosaur era in southwestern Pangaea. Science 331: 206 - 210." type="journal article" year="2011">Martínez et al. 2011</bibRefCitation>
|
||
) with the same taxa.
|
||
</paragraph>
|
||
<paragraph id="8B9B36B1FFC8B72FFCF0FEFC4D67FCCE" blockId="2.[862,1536,256,1960]" pageId="2" pageNumber="3">
|
||
<emphasis id="B950EAA3FFC8B72FFCF0FEFC4C50FEC9" box="[862,1327,256,281]" italics="true" pageId="2" pageNumber="3">
|
||
Diagnosis.—
|
||
<taxonomicName id="4C244D32FFC8B72FFC5DFEFC4C50FEC9" authorityName="Müller & Baczko & Desojo & Nesbitt" authorityYear="2020" box="[1011,1327,256,281]" class="Reptilia" family="Ornithosuchidae" genus="Dynamosuchus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="2" pageNumber="3" phylum="Chordata" rank="species" species="collisensis">Dynamosuchus collisensis</taxonomicName>
|
||
</emphasis>
|
||
differs from all other known ornithosuchids based on a unique combination of character states (* autapomorphy): dorsal process of the quadratojugal mostly dorsally oriented and with the lateral surface densely covered by rugosities; additional foramen on the medial wall of the quadrate foramen*; parabasisphenoid component of the basal tubera ventrally separated from the basipterygoid process by a slight notch at the midline; main axis of the parabasisphenoid process posteroventrally directed; surangular with a conspicuous lateral shelf and an anteroposteriorly wide surangular foramen; anterior dorsal vertebrae with anterior and posterior centrodiapophyseal laminae and without prezygodiapophyseal and postzygodiapophyseal laminae; dorsal body osteoderms with an external pattern of dense crests and grooves; and iliofibularis tubercle of the fibula located within the proximal half of the bone.
|
||
</paragraph>
|
||
<paragraph id="8B9B36B1FFC8B72FFC2AFCD94AADFAD2" blockId="2.[862,1536,256,1960]" pageId="2" pageNumber="3">
|
||
<taxonomicName id="4C244D32FFC8B72FFC2AFCD94DCEFCEE" baseAuthorityName="Newton" baseAuthorityYear="1894" box="[900,1201,805,830]" class="Reptilia" family="Ornithosuchidae" genus="Ornithosuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prolacertiformes" pageId="2" pageNumber="3" phylum="Chordata" rank="species" species="woodwardi">
|
||
<emphasis id="B950EAA3FFC8B72FFC2AFCD94DCEFCEE" box="[900,1201,805,830]" italics="true" pageId="2" pageNumber="3">Ornithosuchus woodwardi</emphasis>
|
||
</taxonomicName>
|
||
differs from
|
||
<taxonomicName id="4C244D32FFC8B72FFAFDFCD94AAFFC8E" authorityName="Müller & Baczko & Desojo & Nesbitt" authorityYear="2020" class="Reptilia" family="Ornithosuchidae" genus="Dynamosuchus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="2" pageNumber="3" phylum="Chordata" rank="species" species="collisensis">
|
||
<emphasis id="B950EAA3FFC8B72FFAFDFCD94AAFFC8E" italics="true" pageId="2" pageNumber="3">Dynamosuchus collisensis</emphasis>
|
||
</taxonomicName>
|
||
in that
|
||
<taxonomicName id="4C244D32FFC8B72FFB82FCB94DB3FC8E" baseAuthorityName="Newton" baseAuthorityYear="1894" box="[1068,1228,837,862]" class="Reptilia" family="Ornithosuchidae" genus="Ornithosuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prolacertiformes" pageId="2" pageNumber="3" phylum="Chordata" rank="species" species="woodwardi">
|
||
<emphasis id="B950EAA3FFC8B72FFB82FCB94DB3FC8E" box="[1068,1228,837,862]" italics="true" pageId="2" pageNumber="3">O. woodwardi</emphasis>
|
||
</taxonomicName>
|
||
has (for comparative illustrations see the SOM, Supplementary Online Material available at http://app.pan.pl/SOM/app65-Muller_etal_SOM.pdf): dorsal process of the quadratojugal is more anteriorly projected (
|
||
<taxonomicName id="4C244D32FFC8B72FFC01FC3B4D33FC30" baseAuthorityName="Newton" baseAuthorityYear="1894" box="[943,1100,967,992]" class="Reptilia" family="Ornithosuchidae" genus="Ornithosuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prolacertiformes" pageId="2" pageNumber="3" phylum="Chordata" rank="species" species="woodwardi">
|
||
<emphasis id="B950EAA3FFC8B72FFC01FC3B4D33FC30" box="[943,1100,967,992]" italics="true" pageId="2" pageNumber="3">O. woodwardi</emphasis>
|
||
</taxonomicName>
|
||
, 45°;
|
||
<taxonomicName id="4C244D32FFC8B72FFB20FC3B4C5DFC30" authorityName="Müller & Baczko & Desojo & Nesbitt" authorityYear="2020" box="[1166,1314,967,992]" class="Reptilia" family="Ornithosuchidae" genus="Dynamosuchus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="2" pageNumber="3" phylum="Chordata" rank="species" species="collisensis">
|
||
<emphasis id="B950EAA3FFC8B72FFB20FC3B4C5DFC30" box="[1166,1314,967,992]" italics="true" pageId="2" pageNumber="3">D. collisensis</emphasis>
|
||
</taxonomicName>
|
||
, 90°); no additional foramen on the medial wall of the quadrate foramen; the lateral surface of the angular has ornamentation; less laterally extended surangular shelf; presence of prezygodiapophyseal and postzygodiapophyseal lamina in the anterior dorsal vertebrae; presence of a fossa ventral to the neurocentral suture on the lateral side of the dorsal centra; presence of an accessory process on the anterior margin of the neural arch of the middle caudal vertebrae; and a less expanded distal end of the radius.
|
||
</paragraph>
|
||
<paragraph id="8B9B36B1FFC8B72FFC2AFAF54DBCF9B5" blockId="2.[862,1536,256,1960]" pageId="2" pageNumber="3">
|
||
<taxonomicName id="4C244D32FFC8B72FFC2AFAF54DE0FAF2" baseAuthorityName="Bonaparte" baseAuthorityYear="1970" box="[900,1183,1289,1314]" family="Ornithosuchidae" genus="Venaticosuchus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="2" pageNumber="3" phylum="Chordata" rank="species" species="rusconii">
|
||
<emphasis id="B950EAA3FFC8B72FFC2AFAF54DE0FAF2" box="[900,1183,1289,1314]" italics="true" pageId="2" pageNumber="3">Venaticosuchus rusconii</emphasis>
|
||
</taxonomicName>
|
||
differs from
|
||
<taxonomicName id="4C244D32FFC8B72FFAEFFAF54CA2FAF2" authorityName="Müller & Baczko & Desojo & Nesbitt" authorityYear="2020" box="[1345,1501,1289,1314]" class="Reptilia" family="Ornithosuchidae" genus="Dynamosuchus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="2" pageNumber="3" phylum="Chordata" rank="species" species="collisensis">
|
||
<emphasis id="B950EAA3FFC8B72FFAEFFAF54CA2FAF2" box="[1345,1501,1289,1314]" italics="true" pageId="2" pageNumber="3">D. collisensis</emphasis>
|
||
</taxonomicName>
|
||
in that
|
||
<taxonomicName id="4C244D32FFC8B72FFC3EFAD64D74FA93" baseAuthorityName="Bonaparte" baseAuthorityYear="1970" box="[912,1035,1322,1347]" family="Ornithosuchidae" genus="Venaticosuchus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="2" pageNumber="3" phylum="Chordata" rank="species" species="rusconii">
|
||
<emphasis id="B950EAA3FFC8B72FFC3EFAD64D74FA93" box="[912,1035,1322,1347]" italics="true" pageId="2" pageNumber="3">V. rusconii</emphasis>
|
||
</taxonomicName>
|
||
has: absence of foramina on the lateral surface of the maxilla; no additional foramen on the medial wall of the quadrate foramen; parabasisphenoid component of the basal tubera ventrally separated from the basipterygoid process by a notch that forms an acute angle; basipterygoid processes more transversely separated from each other in comparison to
|
||
<taxonomicName id="4C244D32FFC8B72FFB8DFA174DC5F9D4" authorityName="Müller & Baczko & Desojo & Nesbitt" authorityYear="2020" box="[1059,1210,1515,1540]" class="Reptilia" family="Ornithosuchidae" genus="Dynamosuchus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="2" pageNumber="3" phylum="Chordata" rank="species" species="collisensis">
|
||
<emphasis id="B950EAA3FFC8B72FFB8DFA174DC5F9D4" box="[1059,1210,1515,1540]" italics="true" pageId="2" pageNumber="3">D. collisensis</emphasis>
|
||
</taxonomicName>
|
||
; basipterygoid processes are ventrally directed with a semicircular shape in lateral view; and absence of the surangular foramen (autapomorphy of
|
||
<taxonomicName id="4C244D32FFC8B72FFCF0F9B04AA8F9B5" baseAuthorityName="Bonaparte" baseAuthorityYear="1970" box="[862,983,1612,1637]" family="Ornithosuchidae" genus="Venaticosuchus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="2" pageNumber="3" phylum="Chordata" rank="species" species="rusconii">
|
||
<emphasis id="B950EAA3FFC8B72FFCF0F9B04AA8F9B5" box="[862,983,1612,1637]" italics="true" pageId="2" pageNumber="3">V. rusconii</emphasis>
|
||
</taxonomicName>
|
||
;
|
||
<bibRefCitation id="EFB54B40FFC8B72FFC48F9B04DC8F9B5" author="Baczko, M. B. von & Desojo, J. B. & Pol, D." box="[998,1207,1612,1637]" pageId="2" pageNumber="3" pagination="1342 - 1356" refId="ref7276" refString="Baczko, M. B. von, Desojo, J. B., and Pol, D. 2014. Anatomy and phylogenetic position of Venaticosuchus rusconii Bonaparte, 1970 (Archosauria, Pseudosuchia), from the Ischigualasto Formation (Late Triassic), La Rioja, Argentina. Journal of Vertebrate Paleontology 34: 1342 - 1356." type="journal article" year="2014">Baczko et al. 2014</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B9B36B1FFC8B729FC2AF9904B99FEAC" blockId="2.[862,1536,256,1960]" lastBlockId="4.[151,824,160,380]" lastPageId="4" lastPageNumber="5" pageId="2" pageNumber="3">
|
||
<taxonomicName id="4C244D32FFC8B72FFC2AF9904DECF955" baseAuthorityName="Bonaparte" baseAuthorityYear="1967" box="[900,1171,1644,1669]" class="Reptilia" family="Ornithosuchidae" genus="Riojasuchus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="2" pageNumber="3" phylum="Chordata" rank="species" species="tenuisceps">
|
||
<emphasis id="B950EAA3FFC8B72FFC2AF9904DECF955" box="[900,1171,1644,1669]" italics="true" pageId="2" pageNumber="3">Riojasuchus tenuisceps</emphasis>
|
||
</taxonomicName>
|
||
differs from
|
||
<taxonomicName id="4C244D32FFC8B72FFA93F9904CA3F955" authorityName="Müller & Baczko & Desojo & Nesbitt" authorityYear="2020" box="[1341,1500,1644,1669]" class="Reptilia" family="Ornithosuchidae" genus="Dynamosuchus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="2" pageNumber="3" phylum="Chordata" rank="species" species="collisensis">
|
||
<emphasis id="B950EAA3FFC8B72FFA93F9904CA3F955" box="[1341,1500,1644,1669]" italics="true" pageId="2" pageNumber="3">D. collisensis</emphasis>
|
||
</taxonomicName>
|
||
in that
|
||
<taxonomicName id="4C244D32FFC8B72FFC3DF9714D53F976" baseAuthorityName="Bonaparte" baseAuthorityYear="1967" box="[915,1068,1677,1702]" class="Reptilia" family="Ornithosuchidae" genus="Riojasuchus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="2" pageNumber="3" phylum="Chordata" rank="species" species="tenuisceps">
|
||
<emphasis id="B950EAA3FFC8B72FFC3DF9714D53F976" box="[915,1068,1677,1702]" italics="true" pageId="2" pageNumber="3">R. tenuisceps</emphasis>
|
||
</taxonomicName>
|
||
has: absence of foramina on the lateral surface of the maxilla; anterior and ventral margins of the antorbital fossa almost meet with the anterior and ventral edges of the maxilla; lateral surface of the quadratojugal is smooth; dorsal process of the quadratojugal is more anteriorly projected (c. 45°); no additional foramen on the medial wall of the quadrate foramen; ventral process of the squamosal anteroposteriorly larger proportionally; absence of a semilunar depression on the posterolateral surface of the parabasisphenoid; absence of the anterior and posterior centrodiapophyseal laminae of the anterior dorsal vertebrae; osteoderms with smooth ornamentation; internal tuberosity of the humerus is more pronounced; postacetabular process of the ilium is proportionally anteroposteriorly longer; iliac blade with a concave dorsal margin in lateral view; and iliofibularis tubercle of the fibula more distally located.
|
||
</paragraph>
|
||
<paragraph id="8B9B36B1FFCEB729FF39FE734BC8FB07" blockId="4.[151,826,399,1728]" pageId="4" pageNumber="5">
|
||
<emphasis id="B950EAA3FFCEB729FF39FE734843FE78" box="[151,316,399,424]" italics="true" pageId="4" pageNumber="5">Description.—</emphasis>
|
||
The body of the premaxilla of
|
||
<taxonomicName id="4C244D32FFCEB729FD24FE734871FE18" authorityName="Müller & Baczko & Desojo & Nesbitt" authorityYear="2020" class="Reptilia" family="Ornithosuchidae" genus="Dynamosuchus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="4" pageNumber="5" phylum="Chordata" rank="species" species="collisensis">
|
||
<emphasis id="B950EAA3FFCEB729FD24FE734871FE18" italics="true" pageId="4" pageNumber="5">Dynamosuchus collisensis</emphasis>
|
||
</taxonomicName>
|
||
is ventrally deflected at the anterior portion (= downturned) and bears three teeth, as typical of ornithosuchids (
|
||
<bibRefCitation id="EFB54B40FFCEB729FF52FE0C4B58FDD9" author="Baczko, M. B. von & Ezcurra, M. D." box="[252,551,496,521]" pageId="4" pageNumber="5" pagination="187 - 202" refId="ref7181" refString="Baczko, M. B. von and Ezcurra, M. D. 2013. Ornithosuchidae: a group of Triassic archosaurs with a unique ankle joint. Geological Society, London, Special Publications 379: 187 - 202." type="journal article" year="2013">Baczko and Ezcurra 2013</bibRefCitation>
|
||
), and the anterior tip of the premaxilla extends further anteriorly than the anterior tip of the dentary (
|
||
<figureCitation id="131F2A34FFCEB729FECDFDCE48A2FD9B" box="[355,477,562,587]" captionStart="→" captionStartId="4.[116,144,1757,1787]" captionTargetId="figure-8@3.[98,1479,161,1951]" captionTargetPageId="3" captionText="→ Fig. 2. Representative skeletal elements of ornithosuchid archosaur Dynamosuchus collisensis gen. et sp. nov. (CAPPA/UFSM 0248) from Janner outcrop, Carnian, Late Triassic. A. Selected skull bones in left lateral view. B. Reconstruction of the skull. C. Skull in ventral view. D. Left quadrate and quadratojugal in posterodorsal view. E. Parabasisphenoid in left lateral view. F. Neural arch of an anterior cervical vertebra in anterior view. G. Centrum of a cervical vertebra in left lateral view. H. Right osteoderm in dorsal view. I. Neural arch of an anterior dorsal vertebra in left lateral view. J. Left ilium in lateral view. L. Right humerus in anterior view. M. Right forearm in medial view. N. Left manus in dorsal view. O. Right (reversed) pubis in lateral view. P. Left femur in anterior view. Q. Left fibula in lateral view. Some unpreserved portions are modified from Baczko et al. in press, for the reconstruction of the skeleton of CAPPA/UFSM 0248 (preserved elements indicated in orange) (K). Scale bars 20 mm." figureDoi="http://doi.org/10.5281/zenodo.11088354" httpUri="https://zenodo.org/record/11088354/files/figure.png" pageId="4" pageNumber="5">Fig. 2A–C</figureCitation>
|
||
). The elongated palatal process of the premaxilla of
|
||
<taxonomicName id="4C244D32FFCEB729FED4FDAE4B6DFDBB" authorityName="Müller & Baczko & Desojo & Nesbitt" authorityYear="2020" box="[378,530,594,619]" class="Reptilia" family="Ornithosuchidae" genus="Dynamosuchus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="4" pageNumber="5" phylum="Chordata" rank="species" species="collisensis">
|
||
<emphasis id="B950EAA3FFCEB729FED4FDAE4B6DFDBB" box="[378,530,594,619]" italics="true" pageId="4" pageNumber="5">D. collisensis</emphasis>
|
||
</taxonomicName>
|
||
bears a ridge at its ventral margin, like in
|
||
<taxonomicName id="4C244D32FFCEB729FEECFD8F4B29FD5C" baseAuthorityName="Bonaparte" baseAuthorityYear="1970" box="[322,598,627,652]" family="Ornithosuchidae" genus="Venaticosuchus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="4" pageNumber="5" phylum="Chordata" rank="species" species="rusconii">
|
||
<emphasis id="B950EAA3FFCEB729FEECFD8F4B29FD5C" box="[322,598,627,652]" italics="true" pageId="4" pageNumber="5">Venaticosuchus rusconii</emphasis>
|
||
</taxonomicName>
|
||
(PVL 2578;
|
||
<bibRefCitation id="EFB54B40FFCEB729FD4AFD8F4870FD7C" author="Baczko, M. B. von & Desojo, J. B. & Pol, D." pageId="4" pageNumber="5" pagination="1342 - 1356" refId="ref7276" refString="Baczko, M. B. von, Desojo, J. B., and Pol, D. 2014. Anatomy and phylogenetic position of Venaticosuchus rusconii Bonaparte, 1970 (Archosauria, Pseudosuchia), from the Ischigualasto Formation (Late Triassic), La Rioja, Argentina. Journal of Vertebrate Paleontology 34: 1342 - 1356." type="journal article" year="2014">Baczko et al. 2014</bibRefCitation>
|
||
). There is a lateral edentulous constriction at the posterior process of the premaxilla, where two caniniform teeth from the dentary fit. The lateral surface of the maxilla is pierced by several foramina (see the Supplementary Online Material for additional figures), which are not recognized in
|
||
<taxonomicName id="4C244D32FFCEB729FEB4FCCA48EAFC9F" baseAuthorityName="Bonaparte" baseAuthorityYear="1970" box="[282,405,822,847]" family="Ornithosuchidae" genus="Venaticosuchus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="4" pageNumber="5" phylum="Chordata" rank="species" species="rusconii">
|
||
<emphasis id="B950EAA3FFCEB729FEB4FCCA48EAFC9F" box="[282,405,822,847]" italics="true" pageId="4" pageNumber="5">V. rusconii</emphasis>
|
||
</taxonomicName>
|
||
(PVL 2578;
|
||
<bibRefCitation id="EFB54B40FFCEB729FD85FCCA4B80FC9F" author="Baczko, M. B. von & Desojo, J. B. & Pol, D." box="[555,767,822,847]" pageId="4" pageNumber="5" pagination="1342 - 1356" refId="ref7276" refString="Baczko, M. B. von, Desojo, J. B., and Pol, D. 2014. Anatomy and phylogenetic position of Venaticosuchus rusconii Bonaparte, 1970 (Archosauria, Pseudosuchia), from the Ischigualasto Formation (Late Triassic), La Rioja, Argentina. Journal of Vertebrate Paleontology 34: 1342 - 1356." type="journal article" year="2014">Baczko et al. 2014</bibRefCitation>
|
||
) and
|
||
<taxonomicName id="4C244D32FFCEB729FF39FCAB484DFCA0" baseAuthorityName="Bonaparte" baseAuthorityYear="1967" box="[151,306,855,880]" class="Reptilia" family="Ornithosuchidae" genus="Riojasuchus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="4" pageNumber="5" phylum="Chordata" rank="species" species="tenuisceps">
|
||
<emphasis id="B950EAA3FFCEB729FF39FCAB484DFCA0" box="[151,306,855,880]" italics="true" pageId="4" pageNumber="5">R. tenuisceps</emphasis>
|
||
</taxonomicName>
|
||
(PVL 3827, PVL 3828;
|
||
<bibRefCitation id="EFB54B40FFCEB729FDF8FCAB49AFFC41" author="Baczko, M. B. von & Desojo, J. B." pageId="4" pageNumber="5" pagination="0148575" refId="ref7138" refString="Baczko, M. B. von and Desojo, J. B. 2016. Cranial anatomy and palaeoneurology of the archosaur Riojasuchus tenuisceps from the Los Colorados Formation, La Rioja, Argentina. PloS One 11: e 0148575." type="journal article" year="2016">Baczko and Desojo 2016</bibRefCitation>
|
||
), but present in
|
||
<taxonomicName id="4C244D32FFCEB729FE27FC844BC7FC41" baseAuthorityName="Newton" baseAuthorityYear="1894" box="[393,696,888,913]" class="Reptilia" family="Ornithosuchidae" genus="Ornithosuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prolacertiformes" pageId="4" pageNumber="5" phylum="Chordata" rank="species" species="woodwardi">
|
||
<emphasis id="B950EAA3FFCEB729FE27FC844BC7FC41" box="[393,696,888,913]" italics="true" pageId="4" pageNumber="5">Ornithosuchus woodwardi</emphasis>
|
||
</taxonomicName>
|
||
(NHMUK PV R 3143;
|
||
<bibRefCitation id="EFB54B40FFCEB729FEB2FC6448D0FC61" author="Walker, A. D." box="[284,431,920,945]" pageId="4" pageNumber="5" pagination="53 - 134" refId="ref9347" refString="Walker, A. D. 1964. Triassic reptiles from the Elgin area Ornithosuchus and the origin of Carnosaurs. Philosophical Transactions of the Royal Society of London B, Biological Sciences 744: 53 - 134." type="journal article" year="1964">Walker 1964</bibRefCitation>
|
||
). The anterior and ventral portions of the antorbital fossa do not coincide with the anterior and ventral edges of the maxilla in
|
||
<taxonomicName id="4C244D32FFCEB729FE56FC254BF0FC22" authorityName="Müller & Baczko & Desojo & Nesbitt" authorityYear="2020" box="[504,655,985,1010]" class="Reptilia" family="Ornithosuchidae" genus="Dynamosuchus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="4" pageNumber="5" phylum="Chordata" rank="species" species="collisensis">
|
||
<emphasis id="B950EAA3FFCEB729FE56FC254BF0FC22" box="[504,655,985,1010]" italics="true" pageId="4" pageNumber="5">D. collisensis</emphasis>
|
||
</taxonomicName>
|
||
, whereas in
|
||
<taxonomicName id="4C244D32FFCEB729FCB1FC254871FBC3" baseAuthorityName="Bonaparte" baseAuthorityYear="1967" class="Reptilia" family="Ornithosuchidae" genus="Riojasuchus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="4" pageNumber="5" phylum="Chordata" rank="species" species="tenuisceps">
|
||
<emphasis id="B950EAA3FFCEB729FCB1FC254871FBC3" italics="true" pageId="4" pageNumber="5">R. tenuisceps</emphasis>
|
||
</taxonomicName>
|
||
(PVL 3827, PVL 3828) these margins almost coincide with each other (
|
||
<bibRefCitation id="EFB54B40FFCEB729FE31FBE74BC4FBE4" author="Baczko, M. B. von & Desojo, J. B." box="[415,699,1051,1076]" pageId="4" pageNumber="5" pagination="0148575" refId="ref7138" refString="Baczko, M. B. von and Desojo, J. B. 2016. Cranial anatomy and palaeoneurology of the archosaur Riojasuchus tenuisceps from the Los Colorados Formation, La Rioja, Argentina. PloS One 11: e 0148575." type="journal article" year="2016">Baczko and Desojo 2016</bibRefCitation>
|
||
). There are six tooth positions preserved; however, the total number of maxillary teeth of
|
||
<taxonomicName id="4C244D32FFCEB729FEC1FBA04B76FBA5" authorityName="Müller & Baczko & Desojo & Nesbitt" authorityYear="2020" box="[367,521,1116,1141]" class="Reptilia" family="Ornithosuchidae" genus="Dynamosuchus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="4" pageNumber="5" phylum="Chordata" rank="species" species="collisensis">
|
||
<emphasis id="B950EAA3FFCEB729FEC1FBA04B76FBA5" box="[367,521,1116,1141]" italics="true" pageId="4" pageNumber="5">D. collisensis</emphasis>
|
||
</taxonomicName>
|
||
is uncertain, but probably does not exceeds seven or eight teeth based on the preserved length of the maxilla, the length of the hemimandible, and through comparisons with other ornithosuchids.
|
||
</paragraph>
|
||
<paragraph id="8B9B36B1FFCEB729FF13FB224CDDFD2C" blockId="4.[151,826,399,1728]" lastBlockId="4.[862,1536,160,1728]" pageId="4" pageNumber="5">
|
||
The right frontal, postfrontal, and anterior portion of the parietal were preserved in articulation showing a markedly constricted interorbital area and delimiting the medial half of the supratemporal fenestra (SOM: fig. S2). A depression for the olfactory bulbs occurs on the ventral surface of the anterior half of the frontal. Like in
|
||
<taxonomicName id="4C244D32FFCEB729FDF7FA7D4B9AFA4A" authorityName="Bonaparte" authorityYear="1969" box="[601,741,1409,1434]" class="Reptilia" family="Ornithosuchidae" genus="Riojasuchus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="4" pageNumber="5" phylum="Chordata" rank="species" species="tenuiscep">
|
||
<emphasis id="B950EAA3FFCEB729FDF7FA7D4B9AFA4A" box="[601,741,1409,1434]" italics="true" pageId="4" pageNumber="5">R. tenuiscep</emphasis>
|
||
</taxonomicName>
|
||
s (PVL 3827), the supratemporal fenestra is anteroposteriorly short, approximately half of the orbit, whereas in
|
||
<taxonomicName id="4C244D32FFCEB729FD08FA3F49A2FA2C" authorityName="Reig" authorityYear="1959" class="Reptilia" genus="Saurosuchus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="4" pageNumber="5" phylum="Chordata" rank="species" species="galilei">
|
||
<emphasis id="B950EAA3FFCEB729FD08FA3F49A2FA2C" italics="true" pageId="4" pageNumber="5">Saurosuchus galilei</emphasis>
|
||
</taxonomicName>
|
||
(PVSJ 32;
|
||
<bibRefCitation id="EFB54B40FFCEB729FEC9FA1F4B77FA2C" author="Alcober, O." box="[359,520,1507,1532]" pageId="4" pageNumber="5" pagination="302 - 316" refId="ref7074" refString="Alcober, O. 2000. Redescription of the skull of Saurosuchus galilei (Archosauria: Rauisuchidae). Journal of Vertebrate Paleontology 20: 302 - 316." type="journal article" year="2000">Alcober 2000</bibRefCitation>
|
||
),
|
||
<taxonomicName id="4C244D32FFCEB729FDB1FA1F49CBF9CD" authorityName="Huene" authorityYear="1942" class="Reptilia" genus="Prestosuchus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="4" pageNumber="5" phylum="Chordata" rank="species" species="chiniquensis">
|
||
<emphasis id="B950EAA3FFCEB729FDB1FA1F49CBF9CD" italics="true" pageId="4" pageNumber="5">Prestosuchus chiniquensis</emphasis>
|
||
</taxonomicName>
|
||
(UFRGS-PV-0629-T;
|
||
<bibRefCitation id="EFB54B40FFCEB729FE1FF9F84BBAF9CD" author="Mastrantonio, B. & Baczko, M. B. von & Desojo, J. B. & Schultz, C. L." box="[433,709,1540,1565]" pageId="4" pageNumber="5" pagination="171 - 198" refId="ref8702" refString="Mastrantonio, B., Baczko, M. B. von, Desojo, J. B., and Schultz, C. L. 2019. The skull anatomy and cranial endocast of the pseudosuchid archosaur Prestosuchus chiniquensis from the Triassic of Brazil. Acta Palaeontologia Polonica 64: 171 - 198." type="journal article" year="2019">Mastrantonio et al. 2019</bibRefCitation>
|
||
),
|
||
<taxonomicName id="4C244D32FFCEB729FD76F9F8499FF9ED" authorityName="Arcucci & Marsicano" authorityYear="1999" class="Reptilia" family="Doswelliidae" genus="Tarjadia" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="4" pageNumber="5" phylum="Chordata" rank="species" species="ruthae">
|
||
<emphasis id="B950EAA3FFCEB729FD76F9F8499FF9ED" italics="true" pageId="4" pageNumber="5">Tarjadia ruthae</emphasis>
|
||
</taxonomicName>
|
||
(CRILAR-Pv 495;
|
||
<bibRefCitation id="EFB54B40FFCEB729FE6DF9D84BE2F9ED" author="Ezcurra, M. D. & Fiorelli, L. E. & Martinelli, A. G. & Rocher, S. M. & Baczko, M. B. von & Ezpeleta, M. & Taborda, J. R. A. & Hechenleitner, E. M. & Trotteyn, M. J. & Desojo, J. B." box="[451,669,1572,1597]" pageId="4" pageNumber="5" pagination="1477 - 1483" refId="ref8018" refString="Ezcurra, M. D., Fiorelli, L. E., Martinelli, A. G., Rocher, S. M., Baczko, M. B. von, Ezpeleta, M., Taborda, J. R. A., Hechenleitner, E. M., Trotteyn, M. J., and Desojo, J. B. 2017. Deep faunistic changes preceded the raise of dinosaurs 1 in southwestern Pangaea. Nature Ecology & Evolution 1: 1477 - 1483." type="journal article" year="2017">Ezcurra et al. 2017</bibRefCitation>
|
||
) and in aetosaurs (e.g.
|
||
<taxonomicName id="4C244D32FFCEB729FEB7F9B94B34F98E" box="[281,587,1605,1630]" family="Stagonolepididae" genus="Neoaetosauroides" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="4" pageNumber="5" phylum="Chordata" rank="species" species="engaeus">
|
||
<emphasis id="B950EAA3FFCEB729FEB7F9B94B34F98E" box="[281,587,1605,1630]" italics="true" pageId="4" pageNumber="5">Neoaetosauroides engaeus</emphasis>
|
||
</taxonomicName>
|
||
; PVL 5698;
|
||
<bibRefCitation id="EFB54B40FFCEB729FD44F9B94833F9AF" author="Desojo, J. B. & Baez, A. M." pageId="4" pageNumber="5" pagination="267 - 276" refId="ref7695" refString="Desojo, J. B. and Baez, A. M. 2007. Cranial morphology of the Late Triassic South American archosaur Neoaetosauroides engaeus: evidence for aetosaurian diversity. Palaeontology 50: 267 - 276." type="journal article" year="2007">Desojo and Báez 2007</bibRefCitation>
|
||
), the supratemporal fenestra is equivalent to the length of the orbit. The ventral process of the squamosal of
|
||
<taxonomicName id="4C244D32FFCEB729FEABF95B48E1F910" authorityName="Müller & Baczko & Desojo & Nesbitt" authorityYear="2020" box="[261,414,1703,1728]" class="Reptilia" family="Ornithosuchidae" genus="Dynamosuchus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="4" pageNumber="5" phylum="Chordata" rank="species" species="collisensis">
|
||
<emphasis id="B950EAA3FFCEB729FEABF95B48E1F910" box="[261,414,1703,1728]" italics="true" pageId="4" pageNumber="5">D. collisensis</emphasis>
|
||
</taxonomicName>
|
||
is elongated and slender (
|
||
<figureCitation id="131F2A34FFCEB729FD7FF95B4A53F910" box="[721,812,1703,1728]" captionStart="→" captionStartId="4.[116,144,1757,1787]" captionTargetId="figure-8@3.[98,1479,161,1951]" captionTargetPageId="3" captionText="→ Fig. 2. Representative skeletal elements of ornithosuchid archosaur Dynamosuchus collisensis gen. et sp. nov. (CAPPA/UFSM 0248) from Janner outcrop, Carnian, Late Triassic. A. Selected skull bones in left lateral view. B. Reconstruction of the skull. C. Skull in ventral view. D. Left quadrate and quadratojugal in posterodorsal view. E. Parabasisphenoid in left lateral view. F. Neural arch of an anterior cervical vertebra in anterior view. G. Centrum of a cervical vertebra in left lateral view. H. Right osteoderm in dorsal view. I. Neural arch of an anterior dorsal vertebra in left lateral view. J. Left ilium in lateral view. L. Right humerus in anterior view. M. Right forearm in medial view. N. Left manus in dorsal view. O. Right (reversed) pubis in lateral view. P. Left femur in anterior view. Q. Left fibula in lateral view. Some unpreserved portions are modified from Baczko et al. in press, for the reconstruction of the skeleton of CAPPA/UFSM 0248 (preserved elements indicated in orange) (K). Scale bars 20 mm." figureDoi="http://doi.org/10.5281/zenodo.11088354" httpUri="https://zenodo.org/record/11088354/files/figure.png" pageId="4" pageNumber="5">Fig. 2A</figureCitation>
|
||
), resembling that of
|
||
<taxonomicName id="4C244D32FFCEB729FB91FF5C4D9BFF69" baseAuthorityName="Newton" baseAuthorityYear="1894" box="[1087,1252,160,185]" class="Reptilia" family="Ornithosuchidae" genus="Ornithosuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prolacertiformes" pageId="4" pageNumber="5" phylum="Chordata" rank="species" species="woodwardi">
|
||
<emphasis id="B950EAA3FFCEB729FB91FF5C4D9BFF69" box="[1087,1252,160,185]" italics="true" pageId="4" pageNumber="5">O. woodwardi</emphasis>
|
||
</taxonomicName>
|
||
(NHMUK PV R 2409, NHMUK PV R 3562), whereas in
|
||
<taxonomicName id="4C244D32FFCEB729FB70FF3C4C0CFF09" baseAuthorityName="Bonaparte" baseAuthorityYear="1967" box="[1246,1395,192,217]" class="Reptilia" family="Ornithosuchidae" genus="Riojasuchus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="4" pageNumber="5" phylum="Chordata" rank="species" species="tenuisceps">
|
||
<emphasis id="B950EAA3FFCEB729FB70FF3C4C0CFF09" box="[1246,1395,192,217]" italics="true" pageId="4" pageNumber="5">R. tenuisceps</emphasis>
|
||
</taxonomicName>
|
||
(PVL 3827) it is proportionally wider anteroposteriorly. There is a longitudinally oriented sulcus bounded medially by a sharp crest on the dorsal surface of the squamosal of
|
||
<taxonomicName id="4C244D32FFCEB729FA9EFEDC4CBAFEE9" authorityName="Müller & Baczko & Desojo & Nesbitt" authorityYear="2020" box="[1328,1477,288,313]" class="Reptilia" family="Ornithosuchidae" genus="Dynamosuchus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="4" pageNumber="5" phylum="Chordata" rank="species" species="collisensis">
|
||
<emphasis id="B950EAA3FFCEB729FA9EFEDC4CBAFEE9" box="[1328,1477,288,313]" italics="true" pageId="4" pageNumber="5">D. collisensis</emphasis>
|
||
</taxonomicName>
|
||
. The general morphology of the quadratojugal of
|
||
<taxonomicName id="4C244D32FFCEB729FACAFEBC4C81FE89" authorityName="Müller & Baczko & Desojo & Nesbitt" authorityYear="2020" box="[1380,1534,320,345]" class="Reptilia" family="Ornithosuchidae" genus="Dynamosuchus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="4" pageNumber="5" phylum="Chordata" rank="species" species="collisensis">
|
||
<emphasis id="B950EAA3FFCEB729FACAFEBC4C81FE89" box="[1380,1534,320,345]" italics="true" pageId="4" pageNumber="5">D. collisensis</emphasis>
|
||
</taxonomicName>
|
||
resembles that of
|
||
<taxonomicName id="4C244D32FFCEB729FB87FE9D4C4AFEAA" authority="(PVL 2578)" baseAuthorityName="PVL" baseAuthorityYear="2578" box="[1065,1333,353,378]" family="Ornithosuchidae" genus="Venaticosuchus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="4" pageNumber="5" phylum="Chordata" rank="species" species="rusconii">
|
||
<emphasis id="B950EAA3FFCEB729FB87FE9D4DD9FEAA" box="[1065,1190,353,378]" italics="true" pageId="4" pageNumber="5">V. rusconii</emphasis>
|
||
(PVL 2578)
|
||
</taxonomicName>
|
||
, where the dorsal process is less anteriorly inclined than in
|
||
<taxonomicName id="4C244D32FFCEB729FA88FE7D4CC5FE4A" baseAuthorityName="Bonaparte" baseAuthorityYear="1967" box="[1318,1466,385,410]" class="Reptilia" family="Ornithosuchidae" genus="Riojasuchus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="4" pageNumber="5" phylum="Chordata" rank="species" species="tenuisceps">
|
||
<emphasis id="B950EAA3FFCEB729FA88FE7D4CC5FE4A" box="[1318,1466,385,410]" italics="true" pageId="4" pageNumber="5">R. tenuisceps</emphasis>
|
||
</taxonomicName>
|
||
(PVL 3827). Like in
|
||
<taxonomicName id="4C244D32FFCEB729FBBEFE5D4DCBFE6A" baseAuthorityName="Newton" baseAuthorityYear="1894" box="[1040,1204,417,442]" class="Reptilia" family="Ornithosuchidae" genus="Ornithosuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prolacertiformes" pageId="4" pageNumber="5" phylum="Chordata" rank="species" species="woodwardi">
|
||
<emphasis id="B950EAA3FFCEB729FBBEFE5D4DCBFE6A" box="[1040,1204,417,442]" italics="true" pageId="4" pageNumber="5">O. woodwardi</emphasis>
|
||
</taxonomicName>
|
||
(NHMUK PV R 3142), the lateral surface of the quadratojugal of
|
||
<taxonomicName id="4C244D32FFCEB729FAB1FE3D4CC7FE0A" authorityName="Müller & Baczko & Desojo & Nesbitt" authorityYear="2020" box="[1311,1464,449,474]" class="Reptilia" family="Ornithosuchidae" genus="Dynamosuchus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="4" pageNumber="5" phylum="Chordata" rank="species" species="collisensis">
|
||
<emphasis id="B950EAA3FFCEB729FAB1FE3D4CC7FE0A" box="[1311,1464,449,474]" italics="true" pageId="4" pageNumber="5">D. collisensis</emphasis>
|
||
</taxonomicName>
|
||
bears a strongly rugose surface (
|
||
<figureCitation id="131F2A34FFCEB729FB3EFE1D4D96FE2A" box="[1168,1257,481,506]" captionStart="→" captionStartId="4.[116,144,1757,1787]" captionTargetId="figure-8@3.[98,1479,161,1951]" captionTargetPageId="3" captionText="→ Fig. 2. Representative skeletal elements of ornithosuchid archosaur Dynamosuchus collisensis gen. et sp. nov. (CAPPA/UFSM 0248) from Janner outcrop, Carnian, Late Triassic. A. Selected skull bones in left lateral view. B. Reconstruction of the skull. C. Skull in ventral view. D. Left quadrate and quadratojugal in posterodorsal view. E. Parabasisphenoid in left lateral view. F. Neural arch of an anterior cervical vertebra in anterior view. G. Centrum of a cervical vertebra in left lateral view. H. Right osteoderm in dorsal view. I. Neural arch of an anterior dorsal vertebra in left lateral view. J. Left ilium in lateral view. L. Right humerus in anterior view. M. Right forearm in medial view. N. Left manus in dorsal view. O. Right (reversed) pubis in lateral view. P. Left femur in anterior view. Q. Left fibula in lateral view. Some unpreserved portions are modified from Baczko et al. in press, for the reconstruction of the skeleton of CAPPA/UFSM 0248 (preserved elements indicated in orange) (K). Scale bars 20 mm." figureDoi="http://doi.org/10.5281/zenodo.11088354" httpUri="https://zenodo.org/record/11088354/files/figure.png" pageId="4" pageNumber="5">Fig. 2D</figureCitation>
|
||
), whereas in other ornithosuchids the homologous surface is smooth (
|
||
<bibRefCitation id="EFB54B40FFCEB729FAC5FDFD4AE9FDEB" author="Desojo, J. B. & Langer, M. C. & Da-Rosa, A. A. S. & Ezpeleta, M. & Ezcurra, M. D. & Martinelli, A. G. & Baczko, M. B. von & Fiorelli, L. & Taborda, J. R. A. & Trotteyn, M. & Hechenleitner, M. & Montefeltro, F. & Ramezani, J." pageId="4" pageNumber="5" pagination="46" refId="ref7810" refString="Desojo, J. B., Langer, M. C., Da-Rosa, A. A. S., Ezpeleta, M., Ezcurra, M. D., Martinelli, A. G., Baczko, M. B. von, Fiorelli, L., Taborda, J. R. A., Trotteyn, M., Hechenleitner, M., Montefeltro, F., and Ramezani, J. 2018. Nuevos hallazgos y dataciones en la Formacion Ischigualasto en Cerro Las Lajas (Cuenca Ischigualasto-Villa Union), Provincia de La Rioja, Argentina. Publicacion Electronica de la Asociacion Paleontologica Argentina 19: 46." type="journal article" year="2018">Baczko et al. 2018</bibRefCitation>
|
||
). The quadrate foramen lies between the medial margin of the quadratojugal and the lateral margin of the quadrate. In addition, there is a foramen piercing the wall that forms the medial margin of the quadrate foramen in the quadrate (
|
||
<figureCitation id="131F2A34FFCEB729FCC8FD5E4ABFFD6B" box="[870,960,674,699]" captionStart="→" captionStartId="4.[116,144,1757,1787]" captionTargetId="figure-8@3.[98,1479,161,1951]" captionTargetPageId="3" captionText="→ Fig. 2. Representative skeletal elements of ornithosuchid archosaur Dynamosuchus collisensis gen. et sp. nov. (CAPPA/UFSM 0248) from Janner outcrop, Carnian, Late Triassic. A. Selected skull bones in left lateral view. B. Reconstruction of the skull. C. Skull in ventral view. D. Left quadrate and quadratojugal in posterodorsal view. E. Parabasisphenoid in left lateral view. F. Neural arch of an anterior cervical vertebra in anterior view. G. Centrum of a cervical vertebra in left lateral view. H. Right osteoderm in dorsal view. I. Neural arch of an anterior dorsal vertebra in left lateral view. J. Left ilium in lateral view. L. Right humerus in anterior view. M. Right forearm in medial view. N. Left manus in dorsal view. O. Right (reversed) pubis in lateral view. P. Left femur in anterior view. Q. Left fibula in lateral view. Some unpreserved portions are modified from Baczko et al. in press, for the reconstruction of the skeleton of CAPPA/UFSM 0248 (preserved elements indicated in orange) (K). Scale bars 20 mm." figureDoi="http://doi.org/10.5281/zenodo.11088354" httpUri="https://zenodo.org/record/11088354/files/figure.png" pageId="4" pageNumber="5">Fig. 2D</figureCitation>
|
||
). Although present in some archosauriforms (e.g.,
|
||
<taxonomicName id="4C244D32FFCEB729FCF0FD3E4D0FFD0B" authorityName="Sennikov" authorityYear="1994" box="[862,1136,706,731]" class="Reptilia" genus="Sarmatosuchus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="4" pageNumber="5" phylum="Chordata" rank="species" species="otschevi">
|
||
<emphasis id="B950EAA3FFCEB729FCF0FD3E4D0FFD0B" box="[862,1136,706,731]" italics="true" pageId="4" pageNumber="5">Sarmatosuchus otschevi</emphasis>
|
||
</taxonomicName>
|
||
; PIN 2865/68;
|
||
<bibRefCitation id="EFB54B40FFCEB729FA8AFD3E4CC0FD0B" author="Ezcurra, M. D." box="[1316,1471,706,731]" pageId="4" pageNumber="5" pagination="1778" refId="ref7932" refString="Ezcurra, M. D. 2016. The phylogenetic relationships of basal archosauromorphs, with an emphasis on the systematics of proterosuchian archosauriforms. PeerJ 4: e 1778." type="journal article" year="2016">Ezcurra 2016</bibRefCitation>
|
||
), this trait was not reported previously for ornithosuchids.
|
||
</paragraph>
|
||
<paragraph id="8B9B36B1FFCEB729FC2AFCFF4C80FA6F" blockId="4.[862,1536,160,1728]" pageId="4" pageNumber="5">
|
||
There is a shallow semilunar depression on the posterolateral surface of the parabasisphenoid of
|
||
<taxonomicName id="4C244D32FFCEB729FAC8FCDF4C80FCEC" authorityName="Müller & Baczko & Desojo & Nesbitt" authorityYear="2020" box="[1382,1535,803,828]" class="Reptilia" family="Ornithosuchidae" genus="Dynamosuchus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="4" pageNumber="5" phylum="Chordata" rank="species" species="collisensis">
|
||
<emphasis id="B950EAA3FFCEB729FAC8FCDF4C80FCEC" box="[1382,1535,803,828]" italics="true" pageId="4" pageNumber="5">D. collisensis</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="131F2A34FFCEB729FCC8FCBF4AC4FC8C" box="[870,955,835,860]" captionStart="→" captionStartId="4.[116,144,1757,1787]" captionTargetId="figure-8@3.[98,1479,161,1951]" captionTargetPageId="3" captionText="→ Fig. 2. Representative skeletal elements of ornithosuchid archosaur Dynamosuchus collisensis gen. et sp. nov. (CAPPA/UFSM 0248) from Janner outcrop, Carnian, Late Triassic. A. Selected skull bones in left lateral view. B. Reconstruction of the skull. C. Skull in ventral view. D. Left quadrate and quadratojugal in posterodorsal view. E. Parabasisphenoid in left lateral view. F. Neural arch of an anterior cervical vertebra in anterior view. G. Centrum of a cervical vertebra in left lateral view. H. Right osteoderm in dorsal view. I. Neural arch of an anterior dorsal vertebra in left lateral view. J. Left ilium in lateral view. L. Right humerus in anterior view. M. Right forearm in medial view. N. Left manus in dorsal view. O. Right (reversed) pubis in lateral view. P. Left femur in anterior view. Q. Left fibula in lateral view. Some unpreserved portions are modified from Baczko et al. in press, for the reconstruction of the skeleton of CAPPA/UFSM 0248 (preserved elements indicated in orange) (K). Scale bars 20 mm." figureDoi="http://doi.org/10.5281/zenodo.11088354" httpUri="https://zenodo.org/record/11088354/files/figure.png" pageId="4" pageNumber="5">Fig. 2E</figureCitation>
|
||
), a trait shared with
|
||
<taxonomicName id="4C244D32FFCEB729FB0EFCBF4C65FC8C" baseAuthorityName="PVL" baseAuthorityYear="2578" box="[1184,1306,835,860]" family="Ornithosuchidae" genus="Venaticosuchus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="4" pageNumber="5" phylum="Chordata" rank="species" species="rusconii">
|
||
<emphasis id="B950EAA3FFCEB729FB0EFCBF4C65FC8C" box="[1184,1306,835,860]" italics="true" pageId="4" pageNumber="5">V. rusconii</emphasis>
|
||
</taxonomicName>
|
||
(PVL 2578;
|
||
<bibRefCitation id="EFB54B40FFCEB729FA04FCBF4AA6FCAC" author="Baczko, M. B. von & Desojo, J. B. & Pol, D." pageId="4" pageNumber="5" pagination="1342 - 1356" refId="ref7276" refString="Baczko, M. B. von, Desojo, J. B., and Pol, D. 2014. Anatomy and phylogenetic position of Venaticosuchus rusconii Bonaparte, 1970 (Archosauria, Pseudosuchia), from the Ischigualasto Formation (Late Triassic), La Rioja, Argentina. Journal of Vertebrate Paleontology 34: 1342 - 1356." type="journal article" year="2014">Baczko et al. 2014</bibRefCitation>
|
||
). Like in
|
||
<taxonomicName id="4C244D32FFCEB729FBFDFC9F4D92FCAC" baseAuthorityName="Bonaparte" baseAuthorityYear="1967" box="[1107,1261,867,892]" class="Reptilia" family="Ornithosuchidae" genus="Riojasuchus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="4" pageNumber="5" phylum="Chordata" rank="species" species="tenuisceps">
|
||
<emphasis id="B950EAA3FFCEB729FBFDFC9F4D92FCAC" box="[1107,1261,867,892]" italics="true" pageId="4" pageNumber="5">R. tenuisceps</emphasis>
|
||
</taxonomicName>
|
||
(PVL 3827), in lateral view, the parabasisphenoid component of the basal tubera of
|
||
<taxonomicName id="4C244D32FFCEB729FCF0FC5F4A88FC6C" authorityName="Müller & Baczko & Desojo & Nesbitt" authorityYear="2020" box="[862,1015,931,956]" class="Reptilia" family="Ornithosuchidae" genus="Dynamosuchus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="4" pageNumber="5" phylum="Chordata" rank="species" species="collisensis">
|
||
<emphasis id="B950EAA3FFCEB729FCF0FC5F4A88FC6C" box="[862,1015,931,956]" italics="true" pageId="4" pageNumber="5">D. collisensis</emphasis>
|
||
</taxonomicName>
|
||
is ventrally separated from the basipterygoid process by a gentle notch, as in most of the pseudosuchians (
|
||
<figureCitation id="131F2A34FFCEB729FCC8FC184AC5FC2D" box="[870,954,996,1021]" captionStart="→" captionStartId="4.[116,144,1757,1787]" captionTargetId="figure-8@3.[98,1479,161,1951]" captionTargetPageId="3" captionText="→ Fig. 2. Representative skeletal elements of ornithosuchid archosaur Dynamosuchus collisensis gen. et sp. nov. (CAPPA/UFSM 0248) from Janner outcrop, Carnian, Late Triassic. A. Selected skull bones in left lateral view. B. Reconstruction of the skull. C. Skull in ventral view. D. Left quadrate and quadratojugal in posterodorsal view. E. Parabasisphenoid in left lateral view. F. Neural arch of an anterior cervical vertebra in anterior view. G. Centrum of a cervical vertebra in left lateral view. H. Right osteoderm in dorsal view. I. Neural arch of an anterior dorsal vertebra in left lateral view. J. Left ilium in lateral view. L. Right humerus in anterior view. M. Right forearm in medial view. N. Left manus in dorsal view. O. Right (reversed) pubis in lateral view. P. Left femur in anterior view. Q. Left fibula in lateral view. Some unpreserved portions are modified from Baczko et al. in press, for the reconstruction of the skeleton of CAPPA/UFSM 0248 (preserved elements indicated in orange) (K). Scale bars 20 mm." figureDoi="http://doi.org/10.5281/zenodo.11088354" httpUri="https://zenodo.org/record/11088354/files/figure.png" pageId="4" pageNumber="5">Fig. 2E</figureCitation>
|
||
). On the other hand, in
|
||
<taxonomicName id="4C244D32FFCEB729FB10FC184CBFFC2D" authority="(PVL 2578)" baseAuthorityName="PVL" baseAuthorityYear="2578" box="[1214,1472,996,1021]" family="Ornithosuchidae" genus="Venaticosuchus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="4" pageNumber="5" phylum="Chordata" rank="species" species="rusconii">
|
||
<emphasis id="B950EAA3FFCEB729FB10FC184C45FC2D" box="[1214,1338,996,1021]" italics="true" pageId="4" pageNumber="5">V. rusconii</emphasis>
|
||
(PVL 2578)
|
||
</taxonomicName>
|
||
there is a more acute angle (about to 40°) formed by the main axis of the basal tubera and the basipterygoid process in lateral view, whereas in
|
||
<taxonomicName id="4C244D32FFCEB729FB8BFBB84DC0FB8D" authorityName="Müller & Baczko & Desojo & Nesbitt" authorityYear="2020" box="[1061,1215,1092,1117]" class="Reptilia" family="Ornithosuchidae" genus="Dynamosuchus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="4" pageNumber="5" phylum="Chordata" rank="species" species="collisensis">
|
||
<emphasis id="B950EAA3FFCEB729FB8BFBB84DC0FB8D" box="[1061,1215,1092,1117]" italics="true" pageId="4" pageNumber="5">D. collisensis</emphasis>
|
||
</taxonomicName>
|
||
the angle is wider (about to 60°). The basipterygoid processes of
|
||
<taxonomicName id="4C244D32FFCEB729FB55FB984C81FBAD" authority="(PVL 2578)" baseAuthorityName="PVL" baseAuthorityYear="2578" box="[1275,1534,1124,1149]" family="Ornithosuchidae" genus="Venaticosuchus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="4" pageNumber="5" phylum="Chordata" rank="species" species="rusconii">
|
||
<emphasis id="B950EAA3FFCEB729FB55FB984C0BFBAD" box="[1275,1396,1124,1149]" italics="true" pageId="4" pageNumber="5">V.rusconii</emphasis>
|
||
(PVL 2578)
|
||
</taxonomicName>
|
||
are more transversely separated from each other when compared to
|
||
<taxonomicName id="4C244D32FFCEB729FC6EFB594D29FB6E" authorityName="Müller & Baczko & Desojo & Nesbitt" authorityYear="2020" box="[960,1110,1189,1214]" class="Reptilia" family="Ornithosuchidae" genus="Dynamosuchus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="4" pageNumber="5" phylum="Chordata" rank="species" species="collisensis">
|
||
<emphasis id="B950EAA3FFCEB729FC6EFB594D29FB6E" box="[960,1110,1189,1214]" italics="true" pageId="4" pageNumber="5">D. collisensis</emphasis>
|
||
</taxonomicName>
|
||
. Moreover, in
|
||
<taxonomicName id="4C244D32FFCEB729FB55FB594C81FB6E" authority="(PVL 2578)" baseAuthorityName="PVL" baseAuthorityYear="2578" box="[1275,1534,1189,1214]" family="Ornithosuchidae" genus="Venaticosuchus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="4" pageNumber="5" phylum="Chordata" rank="species" species="rusconii">
|
||
<emphasis id="B950EAA3FFCEB729FB55FB594C08FB6E" box="[1275,1399,1189,1214]" italics="true" pageId="4" pageNumber="5">V. rusconii</emphasis>
|
||
(PVL 2578)
|
||
</taxonomicName>
|
||
the basipterygoid processes are ventrally directed with a semicircular shape in lateral view (
|
||
<bibRefCitation id="EFB54B40FFCEB729FAA5FB194C8CFB2E" author="Baczko, M. B. von & Desojo, J. B. & Pol, D." box="[1291,1523,1253,1278]" pageId="4" pageNumber="5" pagination="1342 - 1356" refId="ref7276" refString="Baczko, M. B. von, Desojo, J. B., and Pol, D. 2014. Anatomy and phylogenetic position of Venaticosuchus rusconii Bonaparte, 1970 (Archosauria, Pseudosuchia), from the Ischigualasto Formation (Late Triassic), La Rioja, Argentina. Journal of Vertebrate Paleontology 34: 1342 - 1356." type="journal article" year="2014">Baczko et al. 2014</bibRefCitation>
|
||
), whereas in
|
||
<taxonomicName id="4C244D32FFCEB729FC70FAF94D0AFACE" authorityName="Müller & Baczko & Desojo & Nesbitt" authorityYear="2020" box="[990,1141,1285,1310]" class="Reptilia" family="Ornithosuchidae" genus="Dynamosuchus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="4" pageNumber="5" phylum="Chordata" rank="species" species="collisensis">
|
||
<emphasis id="B950EAA3FFCEB729FC70FAF94D0AFACE" box="[990,1141,1285,1310]" italics="true" pageId="4" pageNumber="5">D. collisensis</emphasis>
|
||
</taxonomicName>
|
||
, the processes lie posteroventrally, which results in a more elongated ventral margin in comparison. The basipterygoid recess of
|
||
<taxonomicName id="4C244D32FFCEB729FB7DFAB94C15FA8E" authorityName="Müller & Baczko & Desojo & Nesbitt" authorityYear="2020" box="[1235,1386,1349,1374]" class="Reptilia" family="Ornithosuchidae" genus="Dynamosuchus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="4" pageNumber="5" phylum="Chordata" rank="species" species="collisensis">
|
||
<emphasis id="B950EAA3FFCEB729FB7DFAB94C15FA8E" box="[1235,1386,1349,1374]" italics="true" pageId="4" pageNumber="5">D. collisensis</emphasis>
|
||
</taxonomicName>
|
||
is well-excavated differing from the shallower recess seen in
|
||
<taxonomicName id="4C244D32FFCEB729FA3FFA9A4AEEFA4F" baseAuthorityName="Bonaparte" baseAuthorityYear="1967" class="Reptilia" family="Ornithosuchidae" genus="Riojasuchus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="4" pageNumber="5" phylum="Chordata" rank="species" species="tenuisceps">
|
||
<emphasis id="B950EAA3FFCEB729FA3FFA9A4AEEFA4F" italics="true" pageId="4" pageNumber="5">R. tenuisceps</emphasis>
|
||
</taxonomicName>
|
||
(PVL 3827) and in aetosaurs (e.g.,
|
||
<taxonomicName id="4C244D32FFCEB729FA9CFA7A4A88FA6F" authority="PVL" authorityName="PVL" family="Stagonolepididae" genus="Neoaetosauroides" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="4" pageNumber="5" phylum="Chordata" rank="species" species="engaeus">
|
||
<emphasis id="B950EAA3FFCEB729FA9CFA7A4AC4FA6F" italics="true" pageId="4" pageNumber="5">Neoaetosauroides engaeus</emphasis>
|
||
PVL
|
||
</taxonomicName>
|
||
5698,
|
||
<taxonomicName id="4C244D32FFCEB729FBEEFA5A4CCEFA6F" authority="TTUP" authorityName="TTUP" box="[1088,1457,1446,1471]" class="Reptilia" family="Desmatosuchidae" genus="Desmatosuchus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="4" pageNumber="5" phylum="Chordata" rank="species" species="spurensis">
|
||
<emphasis id="B950EAA3FFCEB729FBEEFA5A4C1CFA6F" box="[1088,1379,1446,1471]" italics="true" pageId="4" pageNumber="5">Desmatosuchus spurensis</emphasis>
|
||
TTUP
|
||
</taxonomicName>
|
||
9024).
|
||
</paragraph>
|
||
<paragraph id="8B9B36B1FFCEB728FC2AFA3A483EFD4D" blockId="4.[862,1536,160,1728]" lastBlockId="5.[98,773,160,1960]" lastPageId="5" lastPageNumber="6" pageId="4" pageNumber="5">
|
||
The lower jaws of
|
||
<taxonomicName id="4C244D32FFCEB729FBC6FA3A4C7CFA0F" authorityName="Müller & Baczko & Desojo & Nesbitt" authorityYear="2020" box="[1128,1283,1478,1503]" class="Reptilia" family="Ornithosuchidae" genus="Dynamosuchus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="4" pageNumber="5" phylum="Chordata" rank="species" species="collisensis">
|
||
<emphasis id="B950EAA3FFCEB729FBC6FA3A4C7CFA0F" box="[1128,1283,1478,1503]" italics="true" pageId="4" pageNumber="5">D. collisensis</emphasis>
|
||
</taxonomicName>
|
||
are anteroposteriorly shorter than the skull (
|
||
<figureCitation id="131F2A34FFCEB729FBF6FA1A4DAEFA2F" box="[1112,1233,1510,1535]" captionStart="→" captionStartId="4.[116,144,1757,1787]" captionTargetId="figure-8@3.[98,1479,161,1951]" captionTargetPageId="3" captionText="→ Fig. 2. Representative skeletal elements of ornithosuchid archosaur Dynamosuchus collisensis gen. et sp. nov. (CAPPA/UFSM 0248) from Janner outcrop, Carnian, Late Triassic. A. Selected skull bones in left lateral view. B. Reconstruction of the skull. C. Skull in ventral view. D. Left quadrate and quadratojugal in posterodorsal view. E. Parabasisphenoid in left lateral view. F. Neural arch of an anterior cervical vertebra in anterior view. G. Centrum of a cervical vertebra in left lateral view. H. Right osteoderm in dorsal view. I. Neural arch of an anterior dorsal vertebra in left lateral view. J. Left ilium in lateral view. L. Right humerus in anterior view. M. Right forearm in medial view. N. Left manus in dorsal view. O. Right (reversed) pubis in lateral view. P. Left femur in anterior view. Q. Left fibula in lateral view. Some unpreserved portions are modified from Baczko et al. in press, for the reconstruction of the skeleton of CAPPA/UFSM 0248 (preserved elements indicated in orange) (K). Scale bars 20 mm." figureDoi="http://doi.org/10.5281/zenodo.11088354" httpUri="https://zenodo.org/record/11088354/files/figure.png" pageId="4" pageNumber="5">Fig. 2A–C</figureCitation>
|
||
). The symphysis is approximately 35% of the mandible length (synapomorphic character state of ornithosuchids;
|
||
<bibRefCitation id="EFB54B40FFCEB729FB03F9DB4C48F990" author="Sereno, P. C." box="[1197,1335,1575,1600]" pageId="4" pageNumber="5" pagination="1 - 53" refId="ref9258" refString="Sereno, P. C. 1991. Basal archosaurs: phylogenetic relationships and functional implications. Journal of Vertebrate Paleontology 11: 1 - 53." type="journal article" year="1991">Sereno 1991</bibRefCitation>
|
||
) and the external mandibular fenestra is also anteroposteriorly longer than tall in comparison with other ornithosuchids. The dorsal margin of the anterior portion of the dentary is elevated relative to the more posterior portion. The splenial forms most of the medial portion of the lower jaw and lacks any foramen. The dorsoventrally slender angular is anteroposteriorly elongated, and delimits the posterior ¾ of the ventral margin of the external mandibular fenestra. Posteriorly, the angular meets the surangular, which delimits the posterodorsal and dorsal margin of the fenestra. There is a wide surangular foramen dorsal to the contact with the angular (
|
||
<figureCitation id="131F2A34FFCFB728FDCEFE9D4BC7FEAA" box="[608,696,353,378]" captionStart="→" captionStartId="4.[116,144,1757,1787]" captionTargetId="figure-8@3.[98,1479,161,1951]" captionTargetPageId="3" captionText="→ Fig. 2. Representative skeletal elements of ornithosuchid archosaur Dynamosuchus collisensis gen. et sp. nov. (CAPPA/UFSM 0248) from Janner outcrop, Carnian, Late Triassic. A. Selected skull bones in left lateral view. B. Reconstruction of the skull. C. Skull in ventral view. D. Left quadrate and quadratojugal in posterodorsal view. E. Parabasisphenoid in left lateral view. F. Neural arch of an anterior cervical vertebra in anterior view. G. Centrum of a cervical vertebra in left lateral view. H. Right osteoderm in dorsal view. I. Neural arch of an anterior dorsal vertebra in left lateral view. J. Left ilium in lateral view. L. Right humerus in anterior view. M. Right forearm in medial view. N. Left manus in dorsal view. O. Right (reversed) pubis in lateral view. P. Left femur in anterior view. Q. Left fibula in lateral view. Some unpreserved portions are modified from Baczko et al. in press, for the reconstruction of the skeleton of CAPPA/UFSM 0248 (preserved elements indicated in orange) (K). Scale bars 20 mm." figureDoi="http://doi.org/10.5281/zenodo.11088354" httpUri="https://zenodo.org/record/11088354/files/figure.png" pageId="5" pageNumber="6">Fig. 2A</figureCitation>
|
||
). Such structure occurs in the other ornithosuchids, except in
|
||
<taxonomicName id="4C244D32FFCFB728FD43FE7E49C1FE6B" baseAuthorityName="PVL" baseAuthorityYear="2578" family="Ornithosuchidae" genus="Venaticosuchus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="5" pageNumber="6" phylum="Chordata" rank="species" species="rusconii">
|
||
<emphasis id="B950EAA3FFCFB728FD43FE7E49C1FE6B" italics="true" pageId="5" pageNumber="6">V. rusconii</emphasis>
|
||
</taxonomicName>
|
||
(PVL 2578;
|
||
<bibRefCitation id="EFB54B40FFCFB728FEE7FE5E4B69FE6B" author="Baczko, M. B. von & Desojo, J. B. & Pol, D." box="[329,534,418,443]" pageId="5" pageNumber="6" pagination="1342 - 1356" refId="ref7276" refString="Baczko, M. B. von, Desojo, J. B., and Pol, D. 2014. Anatomy and phylogenetic position of Venaticosuchus rusconii Bonaparte, 1970 (Archosauria, Pseudosuchia), from the Ischigualasto Formation (Late Triassic), La Rioja, Argentina. Journal of Vertebrate Paleontology 34: 1342 - 1356." type="journal article" year="2014">Baczko et al. 2014</bibRefCitation>
|
||
).
|
||
<taxonomicName id="4C244D32FFCFB728FD8BFE5E49CAFE0B" authorityName="Müller & Baczko & Desojo & Nesbitt" authorityYear="2020" class="Reptilia" family="Ornithosuchidae" genus="Dynamosuchus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="5" pageNumber="6" phylum="Chordata" rank="species" species="collisensis">
|
||
<emphasis id="B950EAA3FFCFB728FD8BFE5E49CAFE0B" italics="true" pageId="5" pageNumber="6">Dynamosuchus collisensis</emphasis>
|
||
</taxonomicName>
|
||
bears a well-developed surangular shelf, which resembles the sharp shelf of
|
||
<taxonomicName id="4C244D32FFCFB728FE24FE1E4B60FE2B" baseAuthorityName="Bonaparte" baseAuthorityYear="1967" box="[394,543,482,507]" class="Reptilia" family="Ornithosuchidae" genus="Riojasuchus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="5" pageNumber="6" phylum="Chordata" rank="species" species="tenuisceps">
|
||
<emphasis id="B950EAA3FFCFB728FE24FE1E4B60FE2B" box="[394,543,482,507]" italics="true" pageId="5" pageNumber="6">R. tenuisceps</emphasis>
|
||
</taxonomicName>
|
||
(PVL 3827;
|
||
<bibRefCitation id="EFB54B40FFCFB728FD01FE1E4862FDCC" author="Baczko, M. B. von & Desojo, J. B." pageId="5" pageNumber="6" pagination="0148575" refId="ref7138" refString="Baczko, M. B. von and Desojo, J. B. 2016. Cranial anatomy and palaeoneurology of the archosaur Riojasuchus tenuisceps from the Los Colorados Formation, La Rioja, Argentina. PloS One 11: e 0148575." type="journal article" year="2016">Baczko and Desojo 2016</bibRefCitation>
|
||
) and
|
||
<taxonomicName id="4C244D32FFCFB728FEF5FDFF48ABFDCC" baseAuthorityName="PVL" baseAuthorityYear="2578" box="[347,468,515,540]" family="Ornithosuchidae" genus="Venaticosuchus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="5" pageNumber="6" phylum="Chordata" rank="species" species="rusconii">
|
||
<emphasis id="B950EAA3FFCFB728FEF5FDFF48ABFDCC" box="[347,468,515,540]" italics="true" pageId="5" pageNumber="6">V. rusconii</emphasis>
|
||
</taxonomicName>
|
||
(PVL 2578;
|
||
<bibRefCitation id="EFB54B40FFCFB728FDCBFDFF4B88FDCC" author="Baczko, M. B. von" box="[613,759,515,540]" pageId="5" pageNumber="6" pagination="365 - 380" refId="ref7103" refString="Baczko, M. B. von 2018. Rediscovered cranial material of Venaticosuchus rusconii enables the first jaw biomechanics in Ornithosuchidae (Archosauria: Pseudosuchia). Ameghiniana 55: 365 - 380." type="journal article" year="2018">Baczko 2018</bibRefCitation>
|
||
), whereas in
|
||
<taxonomicName id="4C244D32FFCFB728FF48FDDF4BE7FDEC" authority="(NHMUK PV R 2409)" baseAuthorityName="NHMUK PV R" baseAuthorityYear="2409" box="[230,664,547,572]" class="Reptilia" family="Ornithosuchidae" genus="Ornithosuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prolacertiformes" pageId="5" pageNumber="6" phylum="Chordata" rank="species" species="woodwardi">
|
||
<emphasis id="B950EAA3FFCFB728FF48FDDF48F7FDEC" box="[230,392,547,572]" italics="true" pageId="5" pageNumber="6">O. woodwardi</emphasis>
|
||
(NHMUK PV R 2409)
|
||
</taxonomicName>
|
||
the shelf is less pronounced. This shelf is also present in proterochampsids, erpetosuchids, and gracilisuchids (
|
||
<bibRefCitation id="EFB54B40FFCFB728FDCFFD984B81FDAD" author="Ezcurra, M. D." box="[609,766,612,637]" pageId="5" pageNumber="6" pagination="1778" refId="ref7932" refString="Ezcurra, M. D. 2016. The phylogenetic relationships of basal archosauromorphs, with an emphasis on the systematics of proterosuchian archosauriforms. PeerJ 4: e 1778." type="journal article" year="2016">Ezcurra 2016</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFB54B40FFCFB728FFCCFD784849FD4D" author="Ezcurra, M. D. & Fiorelli, L. E. & Martinelli, A. G. & Rocher, S. M. & Baczko, M. B. von & Ezpeleta, M. & Taborda, J. R. A. & Hechenleitner, E. M. & Trotteyn, M. J. & Desojo, J. B." box="[98,310,644,669]" pageId="5" pageNumber="6" pagination="1477 - 1483" refId="ref8018" refString="Ezcurra, M. D., Fiorelli, L. E., Martinelli, A. G., Rocher, S. M., Baczko, M. B. von, Ezpeleta, M., Taborda, J. R. A., Hechenleitner, E. M., Trotteyn, M. J., and Desojo, J. B. 2017. Deep faunistic changes preceded the raise of dinosaurs 1 in southwestern Pangaea. Nature Ecology & Evolution 1: 1477 - 1483." type="journal article" year="2017">Ezcurra et al. 2017</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
<caption id="DF5B6639FFCEB729FFDAF9214D35F874" ID-DOI="http://doi.org/10.5281/zenodo.11088354" ID-Zenodo-Dep="11088354" httpUri="https://zenodo.org/record/11088354/files/figure.png" pageId="4" pageNumber="5" startId="4.[116,144,1757,1787]" subCaptionStartIDs="4.[151,184,1768,1788]" subCaptionStarts="Fig. 2" targetBox="[104,1478,170,1946]" targetPageId="3">
|
||
<paragraph id="8B9B36B1FFCEB729FFDAF9214D35F874" blockId="4.[116,1537,1757,1956]" pageId="4" pageNumber="5">
|
||
<emphasis id="B950EAA3FFCEB729FFDAF92149EFF92B" bold="true" box="[116,144,1757,1787]" pageId="4" pageNumber="5">→</emphasis>
|
||
Fig. 2. Representative skeletal elements of ornithosuchid archosaur
|
||
<taxonomicName id="4C244D32FFCEB729FD58F9144A9BF92C" authority="Müller & Baczko & Desojo & Nesbitt, 2020" authorityName="Müller & Baczko & Desojo & Nesbitt" authorityYear="2020" box="[758,996,1768,1788]" class="Reptilia" family="Ornithosuchidae" genus="Dynamosuchus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="4" pageNumber="5" phylum="Chordata" rank="species" species="collisensis" status="gen. et sp. nov.">
|
||
<emphasis id="B950EAA3FFCEB729FD58F9144A9BF92C" box="[758,996,1768,1788]" italics="true" pageId="4" pageNumber="5">Dynamosuchus collisensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A26357D8FFCEB729FC47F9144D10F92C" box="[1001,1135,1768,1788]" pageId="4" pageNumber="5" rank="species">gen. et sp. nov.</taxonomicNameLabel>
|
||
(CAPPA/UFSM 0248) from Janner outcrop, Carnian, Late Triassic.
|
||
<emphasis id="B950EAA3FFCEB729FEC5F8FF4804F8C8" bold="true" box="[363,379,1795,1816]" pageId="4" pageNumber="5">A</emphasis>
|
||
. Selected skull bones in left lateral view.
|
||
<emphasis id="B950EAA3FFCEB729FD59F8FF4A79F8C8" bold="true" box="[759,774,1795,1816]" pageId="4" pageNumber="5">B</emphasis>
|
||
. Reconstruction of the skull.
|
||
<emphasis id="B950EAA3FFCEB729FBBDF8FE4D5CF8C8" bold="true" box="[1043,1059,1794,1816]" pageId="4" pageNumber="5">C</emphasis>
|
||
. Skull in ventral view.
|
||
<emphasis id="B950EAA3FFCEB729FB56F8FF4C77F8C8" bold="true" box="[1272,1288,1795,1816]" pageId="4" pageNumber="5">D</emphasis>
|
||
. Left quadrate and quadratojugal in posterodorsal view.
|
||
<emphasis id="B950EAA3FFCEB729FE02F8E348C4F8E4" bold="true" box="[428,443,1823,1844]" pageId="4" pageNumber="5">E</emphasis>
|
||
. Parabasisphenoid in left lateral view.
|
||
<emphasis id="B950EAA3FFCEB729FCB1F8E34A52F8E4" bold="true" box="[799,813,1823,1844]" pageId="4" pageNumber="5">F</emphasis>
|
||
. Neural arch of an anterior cervical vertebra in anterior view.
|
||
<emphasis id="B950EAA3FFCEB729FAC6F8E24C05F8E4" bold="true" box="[1384,1402,1822,1844]" pageId="4" pageNumber="5">G</emphasis>
|
||
.
|
||
<collectingRegion id="49E0F853FFCEB729FA28F8DC4CAAF8E4" box="[1414,1493,1824,1844]" country="France" name="Centre" pageId="4" pageNumber="5">Centrum</collectingRegion>
|
||
of a cervical vertebra in left lateral view.
|
||
<emphasis id="B950EAA3FFCEB729FE49F8C74886F880" bold="true" box="[487,505,1851,1872]" pageId="4" pageNumber="5">H</emphasis>
|
||
. Right osteoderm in dorsal view.
|
||
<emphasis id="B950EAA3FFCEB729FC80F8C74A48F880" bold="true" box="[814,823,1851,1872]" pageId="4" pageNumber="5">I</emphasis>
|
||
. Neural arch of an anterior dorsal vertebra in left lateral view.
|
||
<emphasis id="B950EAA3FFCEB729FADAF8C74C00F880" bold="true" box="[1396,1407,1851,1872]" pageId="4" pageNumber="5">J</emphasis>
|
||
. Left ilium in lateral view.
|
||
<emphasis id="B950EAA3FFCEB729FEA5F8AB4865F8BC" bold="true" box="[267,282,1879,1900]" pageId="4" pageNumber="5">L</emphasis>
|
||
. Right humerus in anterior view.
|
||
<emphasis id="B950EAA3FFCEB729FDE7F8AB4B21F8BC" bold="true" box="[585,606,1879,1900]" pageId="4" pageNumber="5">M</emphasis>
|
||
. Right forearm in medial view.
|
||
<emphasis id="B950EAA3FFCEB729FCD3F8AB4AF2F8BC" bold="true" box="[893,909,1879,1900]" pageId="4" pageNumber="5">N</emphasis>
|
||
. Left manus in dorsal view.
|
||
<emphasis id="B950EAA3FFCEB729FB20F8AA4DDFF8BC" bold="true" box="[1166,1184,1878,1900]" pageId="4" pageNumber="5">O</emphasis>
|
||
. Right (reversed) pubis in lateral view.
|
||
<emphasis id="B950EAA3FFCEB729FF39F88F49DAF858" bold="true" box="[151,165,1907,1928]" pageId="4" pageNumber="5">P</emphasis>
|
||
. Left femur in anterior view.
|
||
<emphasis id="B950EAA3FFCEB729FE1DF88E48BAF858" bold="true" box="[435,453,1906,1928]" pageId="4" pageNumber="5">Q</emphasis>
|
||
. Left fibula in lateral view. Some unpreserved portions are modified from Baczko et al. in press, for the reconstruction of the skeleton of CAPPA/UFSM 0248 (preserved elements indicated in orange) (
|
||
<emphasis id="B950EAA3FFCEB729FCD3F8734AF0F874" bold="true" box="[893,911,1935,1956]" pageId="4" pageNumber="5">K</emphasis>
|
||
). Scale bars
|
||
<quantity id="4CDC9B54FFCEB729FBADF86C4D39F874" box="[1027,1094,1936,1956]" metricMagnitude="-2" metricUnit="m" metricValue="2.0" pageId="4" pageNumber="5" unit="mm" value="20.0">20 mm</quantity>
|
||
.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8B9B36B1FFCFB728FF29FD584818F916" blockId="5.[98,773,160,1960]" pageId="5" pageNumber="6">
|
||
The axial neural spine is anteroposteriorly longer than the neural spine of the other preserved post-axial cervical vertebrae. At the ventral surface of the postzygapophysis of the cervical vertebrae there is a posterolaterally facing fossa (
|
||
<figureCitation id="131F2A34FFCFB728FF00FCD94878FCEE" box="[174,263,805,830]" captionStart="→" captionStartId="4.[116,144,1757,1787]" captionTargetId="figure-8@3.[98,1479,161,1951]" captionTargetPageId="3" captionText="→ Fig. 2. Representative skeletal elements of ornithosuchid archosaur Dynamosuchus collisensis gen. et sp. nov. (CAPPA/UFSM 0248) from Janner outcrop, Carnian, Late Triassic. A. Selected skull bones in left lateral view. B. Reconstruction of the skull. C. Skull in ventral view. D. Left quadrate and quadratojugal in posterodorsal view. E. Parabasisphenoid in left lateral view. F. Neural arch of an anterior cervical vertebra in anterior view. G. Centrum of a cervical vertebra in left lateral view. H. Right osteoderm in dorsal view. I. Neural arch of an anterior dorsal vertebra in left lateral view. J. Left ilium in lateral view. L. Right humerus in anterior view. M. Right forearm in medial view. N. Left manus in dorsal view. O. Right (reversed) pubis in lateral view. P. Left femur in anterior view. Q. Left fibula in lateral view. Some unpreserved portions are modified from Baczko et al. in press, for the reconstruction of the skeleton of CAPPA/UFSM 0248 (preserved elements indicated in orange) (K). Scale bars 20 mm." figureDoi="http://doi.org/10.5281/zenodo.11088354" httpUri="https://zenodo.org/record/11088354/files/figure.png" pageId="5" pageNumber="6">Fig. 2F</figureCitation>
|
||
), a trait unusual in pseudosuchians, except by
|
||
<taxonomicName id="4C244D32FFCFB728FF2AFCB948ABFC8E" box="[132,468,837,862]" class="Reptilia" family="Sphenosuchidae" genus="Gracilisuchus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="5" pageNumber="6" phylum="Chordata" rank="species" species="stipanicicorum">
|
||
<emphasis id="B950EAA3FFCFB728FF2AFCB948ABFC8E" box="[132,468,837,862]" italics="true" pageId="5" pageNumber="6">Gracilisuchus stipanicicorum</emphasis>
|
||
</taxonomicName>
|
||
(PULR 08;
|
||
<bibRefCitation id="EFB54B40FFCFB728FDF0FCB94B87FC8E" author="Ezcurra, M. D." box="[606,760,837,862]" pageId="5" pageNumber="6" pagination="1778" refId="ref7932" refString="Ezcurra, M. D. 2016. The phylogenetic relationships of basal archosauromorphs, with an emphasis on the systematics of proterosuchian archosauriforms. PeerJ 4: e 1778." type="journal article" year="2016">Ezcurra 2016</bibRefCitation>
|
||
). The cervical neural spines are tall relative to the neural arch, straight, and possess transverse expansions at their dorsal end, forming “spine tables”. The cervical centra have a strongly developed ventral keel that extends longitudinally and project ventral to the margin of the centra (
|
||
<figureCitation id="131F2A34FFCFB728FDC9FC1B4BC1FBD0" box="[615,702,999,1024]" captionStart="→" captionStartId="4.[116,144,1757,1787]" captionTargetId="figure-8@3.[98,1479,161,1951]" captionTargetPageId="3" captionText="→ Fig. 2. Representative skeletal elements of ornithosuchid archosaur Dynamosuchus collisensis gen. et sp. nov. (CAPPA/UFSM 0248) from Janner outcrop, Carnian, Late Triassic. A. Selected skull bones in left lateral view. B. Reconstruction of the skull. C. Skull in ventral view. D. Left quadrate and quadratojugal in posterodorsal view. E. Parabasisphenoid in left lateral view. F. Neural arch of an anterior cervical vertebra in anterior view. G. Centrum of a cervical vertebra in left lateral view. H. Right osteoderm in dorsal view. I. Neural arch of an anterior dorsal vertebra in left lateral view. J. Left ilium in lateral view. L. Right humerus in anterior view. M. Right forearm in medial view. N. Left manus in dorsal view. O. Right (reversed) pubis in lateral view. P. Left femur in anterior view. Q. Left fibula in lateral view. Some unpreserved portions are modified from Baczko et al. in press, for the reconstruction of the skeleton of CAPPA/UFSM 0248 (preserved elements indicated in orange) (K). Scale bars 20 mm." figureDoi="http://doi.org/10.5281/zenodo.11088354" httpUri="https://zenodo.org/record/11088354/files/figure.png" pageId="5" pageNumber="6">Fig. 2G</figureCitation>
|
||
). This condition is uncommon within pseudosuchians being present only in
|
||
<taxonomicName id="4C244D32FFCFB728FF4AFBDB4804FB90" baseAuthorityName="Bonaparte" baseAuthorityYear="1967" box="[228,379,1063,1088]" class="Reptilia" family="Ornithosuchidae" genus="Riojasuchus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="5" pageNumber="6" phylum="Chordata" rank="species" species="tenuisceps">
|
||
<emphasis id="B950EAA3FFCFB728FF4AFBDB4804FB90" box="[228,379,1063,1088]" italics="true" pageId="5" pageNumber="6">R. tenuisceps</emphasis>
|
||
</taxonomicName>
|
||
(PVL 3827),
|
||
<taxonomicName id="4C244D32FFCFB728FDB7FBDB487FFBB1" authority="NHMUK PV" authorityName="NHMUK PV" class="Amphibia" family="Erpetosuchidae" genus="Erpetosuchus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="5" pageNumber="6" phylum="Chordata" rank="species" species="granti">
|
||
<emphasis id="B950EAA3FFCFB728FDB7FBDB4A7DFB90" box="[537,770,1063,1088]" italics="true" pageId="5" pageNumber="6">Erpetosuchus granti</emphasis>
|
||
NHMUK PV
|
||
</taxonomicName>
|
||
3139),
|
||
<taxonomicName id="4C244D32FFCFB728FEE1FBB44B22FBB1" authorityName="Agassiz" authorityYear="1844" box="[335,605,1096,1121]" class="Reptilia" family="Stagonolepididae" genus="Stagonolepis" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="5" pageNumber="6" phylum="Chordata" rank="species" species="robertsoni">
|
||
<emphasis id="B950EAA3FFCFB728FEE1FBB44B22FBB1" box="[335,605,1096,1121]" italics="true" pageId="5" pageNumber="6">Stagonolepis robertsoni</emphasis>
|
||
</taxonomicName>
|
||
(NHMUK PV 4784), and
|
||
<taxonomicName id="4C244D32FFCFB728FF4AFB944B3BFB51" authorityName="Gower" authorityYear="1999" box="[228,580,1128,1153]" class="Reptilia" genus="Batrachotomus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="5" pageNumber="6" phylum="Chordata" rank="species" species="kupferzellensis">
|
||
<emphasis id="B950EAA3FFCFB728FF4AFB944B3BFB51" box="[228,580,1128,1153]" italics="true" pageId="5" pageNumber="6">Batrachotomus kupferzellensis</emphasis>
|
||
</taxonomicName>
|
||
(SMNS 80284). The anterior dorsal vertebrae (
|
||
<figureCitation id="131F2A34FFCFB728FE16FB744B7BFB71" box="[440,516,1160,1185]" captionStart="→" captionStartId="4.[116,144,1757,1787]" captionTargetId="figure-8@3.[98,1479,161,1951]" captionTargetPageId="3" captionText="→ Fig. 2. Representative skeletal elements of ornithosuchid archosaur Dynamosuchus collisensis gen. et sp. nov. (CAPPA/UFSM 0248) from Janner outcrop, Carnian, Late Triassic. A. Selected skull bones in left lateral view. B. Reconstruction of the skull. C. Skull in ventral view. D. Left quadrate and quadratojugal in posterodorsal view. E. Parabasisphenoid in left lateral view. F. Neural arch of an anterior cervical vertebra in anterior view. G. Centrum of a cervical vertebra in left lateral view. H. Right osteoderm in dorsal view. I. Neural arch of an anterior dorsal vertebra in left lateral view. J. Left ilium in lateral view. L. Right humerus in anterior view. M. Right forearm in medial view. N. Left manus in dorsal view. O. Right (reversed) pubis in lateral view. P. Left femur in anterior view. Q. Left fibula in lateral view. Some unpreserved portions are modified from Baczko et al. in press, for the reconstruction of the skeleton of CAPPA/UFSM 0248 (preserved elements indicated in orange) (K). Scale bars 20 mm." figureDoi="http://doi.org/10.5281/zenodo.11088354" httpUri="https://zenodo.org/record/11088354/files/figure.png" pageId="5" pageNumber="6">Fig. 2I</figureCitation>
|
||
) of
|
||
<taxonomicName id="4C244D32FFCFB728FD9EFB744BB7FB71" authorityName="Müller & Baczko & Desojo & Nesbitt" authorityYear="2020" box="[560,712,1160,1185]" class="Reptilia" family="Ornithosuchidae" genus="Dynamosuchus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="5" pageNumber="6" phylum="Chordata" rank="species" species="collisensis">
|
||
<emphasis id="B950EAA3FFCFB728FD9EFB744BB7FB71" box="[560,712,1160,1185]" italics="true" pageId="5" pageNumber="6">D. collisensis</emphasis>
|
||
</taxonomicName>
|
||
have an anterior and posterior centrodiapophyseal lamina, absent in
|
||
<taxonomicName id="4C244D32FFCFB728FF2CFB354864FB32" baseAuthorityName="Bonaparte" baseAuthorityYear="1967" box="[130,283,1225,1250]" class="Reptilia" family="Ornithosuchidae" genus="Riojasuchus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="5" pageNumber="6" phylum="Chordata" rank="species" species="tenuisceps">
|
||
<emphasis id="B950EAA3FFCFB728FF2CFB354864FB32" box="[130,283,1225,1250]" italics="true" pageId="5" pageNumber="6">R. tenuisceps</emphasis>
|
||
</taxonomicName>
|
||
(PVL 3827) and unknown in
|
||
<taxonomicName id="4C244D32FFCFB728FDD1FB354B84FB32" baseAuthorityName="PVL" baseAuthorityYear="2578" box="[639,763,1225,1250]" family="Ornithosuchidae" genus="Venaticosuchus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="5" pageNumber="6" phylum="Chordata" rank="species" species="rusconii">
|
||
<emphasis id="B950EAA3FFCFB728FDD1FB354B84FB32" box="[639,763,1225,1250]" italics="true" pageId="5" pageNumber="6">V. rusconii</emphasis>
|
||
</taxonomicName>
|
||
, but lack prezygodiapophyseal and postzygodiapophyseal lamina. The dorsal vertebrae of
|
||
<taxonomicName id="4C244D32FFCFB728FE71FAF54B04FAF2" authorityName="Müller & Baczko & Desojo & Nesbitt" authorityYear="2020" box="[479,635,1289,1314]" class="Reptilia" family="Ornithosuchidae" genus="Dynamosuchus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="5" pageNumber="6" phylum="Chordata" rank="species" species="collisensis">
|
||
<emphasis id="B950EAA3FFCFB728FE71FAF54B04FAF2" box="[479,635,1289,1314]" italics="true" pageId="5" pageNumber="6">D. collisensis</emphasis>
|
||
</taxonomicName>
|
||
lack hyposphene-hypantrum accessory intervertebral articulation similar to
|
||
<taxonomicName id="4C244D32FFCFB728FF74FAB6480CFAB3" baseAuthorityName="Bonaparte" baseAuthorityYear="1967" box="[218,371,1354,1379]" class="Reptilia" family="Ornithosuchidae" genus="Riojasuchus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="5" pageNumber="6" phylum="Chordata" rank="species" species="tenuisceps">
|
||
<emphasis id="B950EAA3FFCFB728FF74FAB6480CFAB3" box="[218,371,1354,1379]" italics="true" pageId="5" pageNumber="6">R. tenuisceps</emphasis>
|
||
</taxonomicName>
|
||
(PVL 3827). The condition is uncertain in other ornithosuchids. The centra of
|
||
<taxonomicName id="4C244D32FFCFB728FDD3FA9649FFFA73" authorityName="Müller & Baczko & Desojo & Nesbitt" authorityYear="2020" class="Reptilia" family="Ornithosuchidae" genus="Dynamosuchus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="5" pageNumber="6" phylum="Chordata" rank="species" species="collisensis">
|
||
<emphasis id="B950EAA3FFCFB728FDD3FA9649FFFA73" italics="true" pageId="5" pageNumber="6">D. collisensis</emphasis>
|
||
</taxonomicName>
|
||
lack a fossa just ventral to the neurocentral suture like in
|
||
<taxonomicName id="4C244D32FFCFB728FF2BFA57485EFA14" baseAuthorityName="Bonaparte" baseAuthorityYear="1967" box="[133,289,1451,1476]" class="Reptilia" family="Ornithosuchidae" genus="Riojasuchus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="5" pageNumber="6" phylum="Chordata" rank="species" species="tenuisceps">
|
||
<emphasis id="B950EAA3FFCFB728FF2BFA57485EFA14" box="[133,289,1451,1476]" italics="true" pageId="5" pageNumber="6">R. tenuisceps</emphasis>
|
||
</taxonomicName>
|
||
(PVL 3827), whereas a poorly rimmed fossa occurs in
|
||
<taxonomicName id="4C244D32FFCFB728FEA3FA3748D2FA34" baseAuthorityName="NHMUK PV R" baseAuthorityYear="2409" box="[269,429,1483,1508]" class="Reptilia" family="Ornithosuchidae" genus="Ornithosuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prolacertiformes" pageId="5" pageNumber="6" phylum="Chordata" rank="species" species="woodwardi">
|
||
<emphasis id="B950EAA3FFCFB728FEA3FA3748D2FA34" box="[269,429,1483,1508]" italics="true" pageId="5" pageNumber="6">O. woodwardi</emphasis>
|
||
</taxonomicName>
|
||
(NHMUK PV R 3916;
|
||
<bibRefCitation id="EFB54B40FFCFB728FD1FFA3749E3F9D4" author="Walker, A. D." pageId="5" pageNumber="6" pagination="53 - 134" refId="ref9347" refString="Walker, A. D. 1964. Triassic reptiles from the Elgin area Ornithosuchus and the origin of Carnosaurs. Philosophical Transactions of the Royal Society of London B, Biological Sciences 744: 53 - 134." type="journal article" year="1964">Walker 1964</bibRefCitation>
|
||
). The preserved dorsal vertebrae
|
||
<taxonomicName id="4C244D32FFCFB728FDB4FA174BCCF9D4" authorityName="Müller & Baczko & Desojo & Nesbitt" authorityYear="2020" box="[538,691,1515,1540]" class="Reptilia" family="Ornithosuchidae" genus="Dynamosuchus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="5" pageNumber="6" phylum="Chordata" rank="species" species="collisensis">
|
||
<emphasis id="B950EAA3FFCFB728FDB4FA174BCCF9D4" box="[538,691,1515,1540]" italics="true" pageId="5" pageNumber="6">D. collisensis</emphasis>
|
||
</taxonomicName>
|
||
do not preserve the distal tip of the neural spine. The neural spine of the caudal vertebra is posterodorsally directed and bears a pointed posterior projection.
|
||
<taxonomicName id="4C244D32FFCFB728FE7CF9B04A7DF9B5" authorityName="Müller & Baczko & Desojo & Nesbitt" authorityYear="2020" box="[466,770,1612,1637]" class="Reptilia" family="Ornithosuchidae" genus="Dynamosuchus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="5" pageNumber="6" phylum="Chordata" rank="species" species="collisensis">
|
||
<emphasis id="B950EAA3FFCFB728FE7CF9B04A7DF9B5" box="[466,770,1612,1637]" italics="true" pageId="5" pageNumber="6">Dynamosuchus collisensis</emphasis>
|
||
</taxonomicName>
|
||
lacks an accessory laminar process on the anterior margin of the caudal vertebrae, differing from
|
||
<taxonomicName id="4C244D32FFCFB728FDF0F9714862F916" authority="NHMUK PV R" authorityName="NHMUK PV R" class="Reptilia" family="Ornithosuchidae" genus="Ornithosuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prolacertiformes" pageId="5" pageNumber="6" phylum="Chordata" rank="species" species="woodwardi">
|
||
<emphasis id="B950EAA3FFCFB728FDF0F9714A7DF976" box="[606,770,1677,1702]" italics="true" pageId="5" pageNumber="6">O. woodwardi</emphasis>
|
||
NHMUK PV R
|
||
</taxonomicName>
|
||
3561).
|
||
</paragraph>
|
||
<paragraph id="8B9B36B1FFCFB728FF29F9314DD1FDAD" blockId="5.[98,773,160,1960]" lastBlockId="5.[808,1481,160,1960]" pageId="5" pageNumber="6">
|
||
The preserved paramedian osteoderms are square in dorsal view and strongly ornamented by radial ridges and elongated grooves (
|
||
<figureCitation id="131F2A34FFCFB728FEBAF8F2480FF8F7" box="[276,368,1806,1831]" captionStart="→" captionStartId="4.[116,144,1757,1787]" captionTargetId="figure-8@3.[98,1479,161,1951]" captionTargetPageId="3" captionText="→ Fig. 2. Representative skeletal elements of ornithosuchid archosaur Dynamosuchus collisensis gen. et sp. nov. (CAPPA/UFSM 0248) from Janner outcrop, Carnian, Late Triassic. A. Selected skull bones in left lateral view. B. Reconstruction of the skull. C. Skull in ventral view. D. Left quadrate and quadratojugal in posterodorsal view. E. Parabasisphenoid in left lateral view. F. Neural arch of an anterior cervical vertebra in anterior view. G. Centrum of a cervical vertebra in left lateral view. H. Right osteoderm in dorsal view. I. Neural arch of an anterior dorsal vertebra in left lateral view. J. Left ilium in lateral view. L. Right humerus in anterior view. M. Right forearm in medial view. N. Left manus in dorsal view. O. Right (reversed) pubis in lateral view. P. Left femur in anterior view. Q. Left fibula in lateral view. Some unpreserved portions are modified from Baczko et al. in press, for the reconstruction of the skeleton of CAPPA/UFSM 0248 (preserved elements indicated in orange) (K). Scale bars 20 mm." figureDoi="http://doi.org/10.5281/zenodo.11088354" httpUri="https://zenodo.org/record/11088354/files/figure.png" pageId="5" pageNumber="6">Fig. 2H</figureCitation>
|
||
), like in
|
||
<taxonomicName id="4C244D32FFCFB728FE73F8F24BFEF8F7" box="[477,641,1806,1831]" class="Reptilia" family="Ornithosuchidae" genus="Ornithosuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prolacertiformes" pageId="5" pageNumber="6" phylum="Chordata" rank="species" species="woodwardi">
|
||
<emphasis id="B950EAA3FFCFB728FE73F8F24BFEF8F7" box="[477,641,1806,1831]" italics="true" pageId="5" pageNumber="6">O. woodwardi</emphasis>
|
||
</taxonomicName>
|
||
(NHMUK PV R 3916;
|
||
<bibRefCitation id="EFB54B40FFCFB728FF43F8D248FFF897" author="Walker, A. D." box="[237,384,1838,1863]" pageId="5" pageNumber="6" pagination="53 - 134" refId="ref9347" refString="Walker, A. D. 1964. Triassic reptiles from the Elgin area Ornithosuchus and the origin of Carnosaurs. Philosophical Transactions of the Royal Society of London B, Biological Sciences 744: 53 - 134." type="journal article" year="1964">Walker 1964</bibRefCitation>
|
||
) whereas the same surface of the osteoderms of
|
||
<taxonomicName id="4C244D32FFCFB728FEA0F8B248D8F8B7" baseAuthorityName="Bonaparte" baseAuthorityYear="1967" box="[270,423,1870,1895]" class="Reptilia" family="Ornithosuchidae" genus="Riojasuchus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="5" pageNumber="6" phylum="Chordata" rank="species" species="tenuisceps">
|
||
<emphasis id="B950EAA3FFCFB728FEA0F8B248D8F8B7" box="[270,423,1870,1895]" italics="true" pageId="5" pageNumber="6">R. tenuisceps</emphasis>
|
||
</taxonomicName>
|
||
(PVL 3827) bear a gentle radial pattern (
|
||
<bibRefCitation id="EFB54B40FFCFB728FF5DF89348B3F858" author="Baczko, M. B. von & Desojo, J. B. & Ponce, D." box="[243,460,1903,1928]" pageId="5" pageNumber="6" pagination="1693396" refId="ref7338" refString="Baczko, M. B. von, Desojo, J. B., and Ponce, D. 2020. Postcranial anatomy and osteoderm histology of Riojasuchus tenuisceps and a phylogenetic update on Ornithosuchidae (Archosauria, Pseudosuchia). Journal of Vertebrate Paleontology 39 (5): e 1693396." type="journal article" year="2020">Baczko et al. 2020</bibRefCitation>
|
||
). Nevertheless,
|
||
<taxonomicName id="4C244D32FFCFB728FDD1F89349FFF878" authorityName="Müller & Baczko & Desojo & Nesbitt" authorityYear="2020" class="Reptilia" family="Ornithosuchidae" genus="Dynamosuchus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="5" pageNumber="6" phylum="Chordata" rank="species" species="collisensis">
|
||
<emphasis id="B950EAA3FFCFB728FDD1F89349FFF878" italics="true" pageId="5" pageNumber="6">D. collisensis</emphasis>
|
||
</taxonomicName>
|
||
shares with the other ornithosuchids the presence of a prominence on the dorsolateral surface of the osteoderms. In
|
||
<taxonomicName id="4C244D32FFCFB728FCE2FF3C4A96FF09" authorityName="Müller & Baczko & Desojo & Nesbitt" authorityYear="2020" box="[844,1001,192,217]" class="Reptilia" family="Ornithosuchidae" genus="Dynamosuchus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="5" pageNumber="6" phylum="Chordata" rank="species" species="collisensis">
|
||
<emphasis id="B950EAA3FFCFB728FCE2FF3C4A96FF09" box="[844,1001,192,217]" italics="true" pageId="5" pageNumber="6">D. collisensis</emphasis>
|
||
</taxonomicName>
|
||
, this dorsolateral prominence is marked by a rugose surface and, in some elements, it has a pointed dorsal tip. The dorsolateral prominence becomes less developed progressively in posterior osteoderms of the cervical and dorsal series. A longitudinally oriented ridge is absent on the dorsal surface of the osteoderms, such as in other ornithosuchids. The anterior margin of the osteoderms is straight and lacks an anterior process or an unornamented anterior articular lamina. The posterior margin is concave in dorsal view (
|
||
<figureCitation id="131F2A34FFCFB728FC77FE1E4D4CFE2B" box="[985,1075,482,507]" captionStart="→" captionStartId="4.[116,144,1757,1787]" captionTargetId="figure-8@3.[98,1479,161,1951]" captionTargetPageId="3" captionText="→ Fig. 2. Representative skeletal elements of ornithosuchid archosaur Dynamosuchus collisensis gen. et sp. nov. (CAPPA/UFSM 0248) from Janner outcrop, Carnian, Late Triassic. A. Selected skull bones in left lateral view. B. Reconstruction of the skull. C. Skull in ventral view. D. Left quadrate and quadratojugal in posterodorsal view. E. Parabasisphenoid in left lateral view. F. Neural arch of an anterior cervical vertebra in anterior view. G. Centrum of a cervical vertebra in left lateral view. H. Right osteoderm in dorsal view. I. Neural arch of an anterior dorsal vertebra in left lateral view. J. Left ilium in lateral view. L. Right humerus in anterior view. M. Right forearm in medial view. N. Left manus in dorsal view. O. Right (reversed) pubis in lateral view. P. Left femur in anterior view. Q. Left fibula in lateral view. Some unpreserved portions are modified from Baczko et al. in press, for the reconstruction of the skeleton of CAPPA/UFSM 0248 (preserved elements indicated in orange) (K). Scale bars 20 mm." figureDoi="http://doi.org/10.5281/zenodo.11088354" httpUri="https://zenodo.org/record/11088354/files/figure.png" pageId="5" pageNumber="6">Fig. 2H</figureCitation>
|
||
). The ventral surface is smooth and ventrally convex in anterior view. The length of the osteoderms indicate that there is a pair of osteoderms per vertebra, forming two paramedial rows along the dorsal surface of the body (unknown for the tail).
|
||
</paragraph>
|
||
<paragraph id="8B9B36B1FFCFB728FCE0FD784DE4FA93" blockId="5.[808,1481,160,1960]" pageId="5" pageNumber="6">
|
||
The acromion process of the scapula gently rises from the anterior margin of the scapular blade at an angle of approximately 120°, resembling the condition of
|
||
<taxonomicName id="4C244D32FFCFB728FAF0FD384A15FD2E" class="Reptilia" family="Ornithosuchidae" genus="Ornithosuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prolacertiformes" pageId="5" pageNumber="6" phylum="Chordata" rank="species" species="woodwardi">
|
||
<emphasis id="B950EAA3FFCFB728FAF0FD384A15FD2E" italics="true" pageId="5" pageNumber="6">O. woodwardi</emphasis>
|
||
</taxonomicName>
|
||
(NHMUK PV R 3916). In contrast,
|
||
<taxonomicName id="4C244D32FFCFB728FAAAFD194CE2FD2E" authorityName="Müller & Baczko & Desojo & Nesbitt" authorityYear="2020" box="[1284,1437,741,766]" class="Reptilia" family="Ornithosuchidae" genus="Dynamosuchus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="5" pageNumber="6" phylum="Chordata" rank="species" species="collisensis">
|
||
<emphasis id="B950EAA3FFCFB728FAAAFD194CE2FD2E" box="[1284,1437,741,766]" italics="true" pageId="5" pageNumber="6">D. collisensis</emphasis>
|
||
</taxonomicName>
|
||
has a well-developed acromion process, which is nearly flat in
|
||
<taxonomicName id="4C244D32FFCFB728FC86FCD94AB3FCEE" box="[808,972,805,830]" class="Reptilia" family="Ornithosuchidae" genus="Ornithosuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prolacertiformes" pageId="5" pageNumber="6" phylum="Chordata" rank="species" species="woodwardi">
|
||
<emphasis id="B950EAA3FFCFB728FC86FCD94AB3FCEE" box="[808,972,805,830]" italics="true" pageId="5" pageNumber="6">O. woodwardi</emphasis>
|
||
</taxonomicName>
|
||
(NHMUK PV R 3916); the condition of
|
||
<taxonomicName id="4C244D32FFCFB728FA00FCD94AE2FC8F" authorityName="Müller & Baczko & Desojo & Nesbitt" authorityYear="2020" class="Reptilia" family="Ornithosuchidae" genus="Dynamosuchus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="5" pageNumber="6" phylum="Chordata" rank="species" species="collisensis">
|
||
<emphasis id="B950EAA3FFCFB728FA00FCD94AE2FC8F" italics="true" pageId="5" pageNumber="6">D. collisensis</emphasis>
|
||
</taxonomicName>
|
||
resembles that of
|
||
<taxonomicName id="4C244D32FFCFB728FBC2FCBA4C7DFC8F" baseAuthorityName="Bonaparte" baseAuthorityYear="1967" box="[1132,1282,838,863]" class="Reptilia" family="Ornithosuchidae" genus="Riojasuchus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="5" pageNumber="6" phylum="Chordata" rank="species" species="tenuisceps">
|
||
<emphasis id="B950EAA3FFCFB728FBC2FCBA4C7DFC8F" box="[1132,1282,838,863]" italics="true" pageId="5" pageNumber="6">R. tenuisceps</emphasis>
|
||
</taxonomicName>
|
||
(PVL 3828). The humerus (
|
||
<figureCitation id="131F2A34FFCFB728FC34FC9A4A92FCAF" box="[922,1005,870,895]" captionStart="→" captionStartId="4.[116,144,1757,1787]" captionTargetId="figure-8@3.[98,1479,161,1951]" captionTargetPageId="3" captionText="→ Fig. 2. Representative skeletal elements of ornithosuchid archosaur Dynamosuchus collisensis gen. et sp. nov. (CAPPA/UFSM 0248) from Janner outcrop, Carnian, Late Triassic. A. Selected skull bones in left lateral view. B. Reconstruction of the skull. C. Skull in ventral view. D. Left quadrate and quadratojugal in posterodorsal view. E. Parabasisphenoid in left lateral view. F. Neural arch of an anterior cervical vertebra in anterior view. G. Centrum of a cervical vertebra in left lateral view. H. Right osteoderm in dorsal view. I. Neural arch of an anterior dorsal vertebra in left lateral view. J. Left ilium in lateral view. L. Right humerus in anterior view. M. Right forearm in medial view. N. Left manus in dorsal view. O. Right (reversed) pubis in lateral view. P. Left femur in anterior view. Q. Left fibula in lateral view. Some unpreserved portions are modified from Baczko et al. in press, for the reconstruction of the skeleton of CAPPA/UFSM 0248 (preserved elements indicated in orange) (K). Scale bars 20 mm." figureDoi="http://doi.org/10.5281/zenodo.11088354" httpUri="https://zenodo.org/record/11088354/files/figure.png" pageId="5" pageNumber="6">Fig. 2L</figureCitation>
|
||
) is robust, about 0.6 times the total length of the femur (0.65–0.75 in
|
||
<taxonomicName id="4C244D32FFCFB728FBDEFC7A4C75FC4F" baseAuthorityName="Bonaparte" baseAuthorityYear="1967" box="[1136,1290,902,927]" class="Reptilia" family="Ornithosuchidae" genus="Riojasuchus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="5" pageNumber="6" phylum="Chordata" rank="species" species="tenuisceps">
|
||
<emphasis id="B950EAA3FFCFB728FBDEFC7A4C75FC4F" box="[1136,1290,902,927]" italics="true" pageId="5" pageNumber="6">R. tenuisceps</emphasis>
|
||
</taxonomicName>
|
||
), similar to the ration in some aetosaurs and
|
||
<taxonomicName id="4C244D32FFCFB728FB2DFC5A4A16FC30" authority="(PIMUZ 2817)" baseAuthorityName="PIMUZ" baseAuthorityYear="2817" class="Reptilia" family="Rauisuchidae" genus="Ticinosuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="5" pageNumber="6" phylum="Chordata" rank="species" species="ferox">
|
||
<emphasis id="B950EAA3FFCFB728FB2DFC5A4C22FC6F" box="[1155,1373,934,959]" italics="true" pageId="5" pageNumber="6">Ticinosuchus ferox</emphasis>
|
||
(PIMUZ 2817)
|
||
</taxonomicName>
|
||
, among pseudosuchians (
|
||
<bibRefCitation id="EFB54B40FFCFB728FB2AFC3B4C5FFC30" author="Ezcurra, M. D." box="[1156,1312,967,992]" pageId="5" pageNumber="6" pagination="1778" refId="ref7932" refString="Ezcurra, M. D. 2016. The phylogenetic relationships of basal archosauromorphs, with an emphasis on the systematics of proterosuchian archosauriforms. PeerJ 4: e 1778." type="journal article" year="2016">Ezcurra 2016</bibRefCitation>
|
||
). The proximal end of the humerus is widely expanded with a moderately developed internal tuberosity, which is less pronounced in
|
||
<taxonomicName id="4C244D32FFCFB728FC86FBDB4ABEFB90" authorityName="Müller & Baczko & Desojo & Nesbitt" authorityYear="2020" box="[808,961,1063,1088]" class="Reptilia" family="Ornithosuchidae" genus="Dynamosuchus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="5" pageNumber="6" phylum="Chordata" rank="species" species="collisensis">
|
||
<emphasis id="B950EAA3FFCFB728FC86FBDB4ABEFB90" box="[808,961,1063,1088]" italics="true" pageId="5" pageNumber="6">D. collisensis</emphasis>
|
||
</taxonomicName>
|
||
than in
|
||
<taxonomicName id="4C244D32FFCFB728FB8EFBDB4DC9FB90" baseAuthorityName="Bonaparte" baseAuthorityYear="1967" box="[1056,1206,1063,1088]" class="Reptilia" family="Ornithosuchidae" genus="Riojasuchus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="5" pageNumber="6" phylum="Chordata" rank="species" species="tenuisceps">
|
||
<emphasis id="B950EAA3FFCFB728FB8EFBDB4DC9FB90" box="[1056,1206,1063,1088]" italics="true" pageId="5" pageNumber="6">R. tenuisceps</emphasis>
|
||
</taxonomicName>
|
||
(PVL 3826). The deltopectoral crest of
|
||
<taxonomicName id="4C244D32FFCFB728FC5AFBB44DEDFBB1" authorityName="Müller & Baczko & Desojo & Nesbitt" authorityYear="2020" box="[1012,1170,1096,1121]" class="Reptilia" family="Ornithosuchidae" genus="Dynamosuchus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="5" pageNumber="6" phylum="Chordata" rank="species" species="collisensis">
|
||
<emphasis id="B950EAA3FFCFB728FC5AFBB44DEDFBB1" box="[1012,1170,1096,1121]" italics="true" pageId="5" pageNumber="6">D. collisensis</emphasis>
|
||
</taxonomicName>
|
||
is well-developed and restricted to the proximal third of the bone.
|
||
<taxonomicName id="4C244D32FFCFB728FAB5FB944AE1FB71" authorityName="Müller & Baczko & Desojo & Nesbitt" authorityYear="2020" class="Reptilia" family="Ornithosuchidae" genus="Dynamosuchus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="5" pageNumber="6" phylum="Chordata" rank="species" species="collisensis">
|
||
<emphasis id="B950EAA3FFCFB728FAB5FB944AE1FB71" italics="true" pageId="5" pageNumber="6">Dynamosuchus collisensis</emphasis>
|
||
</taxonomicName>
|
||
lacks an ectepicondylar flange or it is poorly developed. The ulna (
|
||
<figureCitation id="131F2A34FFCFB728FB86FB544DF6FB11" box="[1064,1161,1192,1217]" captionStart="→" captionStartId="4.[116,144,1757,1787]" captionTargetId="figure-8@3.[98,1479,161,1951]" captionTargetPageId="3" captionText="→ Fig. 2. Representative skeletal elements of ornithosuchid archosaur Dynamosuchus collisensis gen. et sp. nov. (CAPPA/UFSM 0248) from Janner outcrop, Carnian, Late Triassic. A. Selected skull bones in left lateral view. B. Reconstruction of the skull. C. Skull in ventral view. D. Left quadrate and quadratojugal in posterodorsal view. E. Parabasisphenoid in left lateral view. F. Neural arch of an anterior cervical vertebra in anterior view. G. Centrum of a cervical vertebra in left lateral view. H. Right osteoderm in dorsal view. I. Neural arch of an anterior dorsal vertebra in left lateral view. J. Left ilium in lateral view. L. Right humerus in anterior view. M. Right forearm in medial view. N. Left manus in dorsal view. O. Right (reversed) pubis in lateral view. P. Left femur in anterior view. Q. Left fibula in lateral view. Some unpreserved portions are modified from Baczko et al. in press, for the reconstruction of the skeleton of CAPPA/UFSM 0248 (preserved elements indicated in orange) (K). Scale bars 20 mm." figureDoi="http://doi.org/10.5281/zenodo.11088354" httpUri="https://zenodo.org/record/11088354/files/figure.png" pageId="5" pageNumber="6">Fig. 2M</figureCitation>
|
||
) bears a moderately developed olecranon process and tappers towards its distal end, whereas the radius expands distally. Metacarpal I is the widest, whereas the metacarpal III is the longest of the metacarpals preserved (
|
||
<figureCitation id="131F2A34FFCFB728FB9BFAD64DF0FA93" box="[1077,1167,1322,1347]" captionStart="→" captionStartId="4.[116,144,1757,1787]" captionTargetId="figure-8@3.[98,1479,161,1951]" captionTargetPageId="3" captionText="→ Fig. 2. Representative skeletal elements of ornithosuchid archosaur Dynamosuchus collisensis gen. et sp. nov. (CAPPA/UFSM 0248) from Janner outcrop, Carnian, Late Triassic. A. Selected skull bones in left lateral view. B. Reconstruction of the skull. C. Skull in ventral view. D. Left quadrate and quadratojugal in posterodorsal view. E. Parabasisphenoid in left lateral view. F. Neural arch of an anterior cervical vertebra in anterior view. G. Centrum of a cervical vertebra in left lateral view. H. Right osteoderm in dorsal view. I. Neural arch of an anterior dorsal vertebra in left lateral view. J. Left ilium in lateral view. L. Right humerus in anterior view. M. Right forearm in medial view. N. Left manus in dorsal view. O. Right (reversed) pubis in lateral view. P. Left femur in anterior view. Q. Left fibula in lateral view. Some unpreserved portions are modified from Baczko et al. in press, for the reconstruction of the skeleton of CAPPA/UFSM 0248 (preserved elements indicated in orange) (K). Scale bars 20 mm." figureDoi="http://doi.org/10.5281/zenodo.11088354" httpUri="https://zenodo.org/record/11088354/files/figure.png" pageId="5" pageNumber="6">Fig. 2N</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B9B36B1FFCFB72BFCE0FAB64B62FEAC" blockId="5.[808,1481,160,1960]" lastBlockId="6.[151,825,160,1293]" lastPageId="6" lastPageNumber="7" pageId="5" pageNumber="6">
|
||
Most of the preacetabular process of the ilium is not preserved (
|
||
<figureCitation id="131F2A34FFCFB728FC0BFA964A8AFA53" box="[933,1013,1386,1411]" captionStart="→" captionStartId="4.[116,144,1757,1787]" captionTargetId="figure-8@3.[98,1479,161,1951]" captionTargetPageId="3" captionText="→ Fig. 2. Representative skeletal elements of ornithosuchid archosaur Dynamosuchus collisensis gen. et sp. nov. (CAPPA/UFSM 0248) from Janner outcrop, Carnian, Late Triassic. A. Selected skull bones in left lateral view. B. Reconstruction of the skull. C. Skull in ventral view. D. Left quadrate and quadratojugal in posterodorsal view. E. Parabasisphenoid in left lateral view. F. Neural arch of an anterior cervical vertebra in anterior view. G. Centrum of a cervical vertebra in left lateral view. H. Right osteoderm in dorsal view. I. Neural arch of an anterior dorsal vertebra in left lateral view. J. Left ilium in lateral view. L. Right humerus in anterior view. M. Right forearm in medial view. N. Left manus in dorsal view. O. Right (reversed) pubis in lateral view. P. Left femur in anterior view. Q. Left fibula in lateral view. Some unpreserved portions are modified from Baczko et al. in press, for the reconstruction of the skeleton of CAPPA/UFSM 0248 (preserved elements indicated in orange) (K). Scale bars 20 mm." figureDoi="http://doi.org/10.5281/zenodo.11088354" httpUri="https://zenodo.org/record/11088354/files/figure.png" pageId="5" pageNumber="6">Fig. 2J</figureCitation>
|
||
). Nevertheless, it is evident that the ilium of
|
||
<taxonomicName id="4C244D32FFCFB728FCE6FA764A9EFA73" authorityName="Müller & Baczko & Desojo & Nesbitt" authorityYear="2020" box="[840,993,1418,1443]" class="Reptilia" family="Ornithosuchidae" genus="Dynamosuchus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="5" pageNumber="6" phylum="Chordata" rank="species" species="collisensis">
|
||
<emphasis id="B950EAA3FFCFB728FCE6FA764A9EFA73" box="[840,993,1418,1443]" italics="true" pageId="5" pageNumber="6">D. collisensis</emphasis>
|
||
</taxonomicName>
|
||
lacks a lateral crest dorsal to the supraacetabular crest, as in other ornithosuchids and many pseudosuchians (e.g.,
|
||
<taxonomicName id="4C244D32FFCFB728FC5AFA374C77FA34" authority="PVL" authorityName="PVL" box="[1012,1288,1483,1508]" class="Reptilia" family="Sphenosuchidae" genus="Gracilisuchus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="5" pageNumber="6" phylum="Chordata" rank="species" species="stipanicicorum">
|
||
<emphasis id="B950EAA3FFCFB728FC5AFA374DB9FA34" box="[1012,1222,1483,1508]" italics="true" pageId="5" pageNumber="6">G. stipanicicorum</emphasis>
|
||
PVL
|
||
</taxonomicName>
|
||
4597;
|
||
<taxonomicName id="4C244D32FFCFB728FAF3FA374CB7FA34" authorityName="Arcucci & Marsicano" authorityYear="1999" box="[1373,1480,1483,1508]" class="Reptilia" family="Doswelliidae" genus="Tarjadia" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="5" pageNumber="6" phylum="Chordata" rank="species" species="ruthae">
|
||
<emphasis id="B950EAA3FFCFB728FAF3FA374CB7FA34" box="[1373,1480,1483,1508]" italics="true" pageId="5" pageNumber="6">T. ruthae</emphasis>
|
||
</taxonomicName>
|
||
CRILAR-Pv 478;
|
||
<bibRefCitation id="EFB54B40FFCFB728FC58FA174DB0F9D4" author="Ezcurra, M. D. & Fiorelli, L. E. & Martinelli, A. G. & Rocher, S. M. & Baczko, M. B. von & Ezpeleta, M. & Taborda, J. R. A. & Hechenleitner, E. M. & Trotteyn, M. J. & Desojo, J. B." box="[1014,1231,1515,1540]" pageId="5" pageNumber="6" pagination="1477 - 1483" refId="ref8018" refString="Ezcurra, M. D., Fiorelli, L. E., Martinelli, A. G., Rocher, S. M., Baczko, M. B. von, Ezpeleta, M., Taborda, J. R. A., Hechenleitner, E. M., Trotteyn, M. J., and Desojo, J. B. 2017. Deep faunistic changes preceded the raise of dinosaurs 1 in southwestern Pangaea. Nature Ecology & Evolution 1: 1477 - 1483." type="journal article" year="2017">Ezcurra et al. 2017</bibRefCitation>
|
||
). As in
|
||
<taxonomicName id="4C244D32FFCFB728FA89FA174CB6F9D4" box="[1319,1481,1515,1540]" class="Reptilia" family="Ornithosuchidae" genus="Ornithosuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prolacertiformes" pageId="5" pageNumber="6" phylum="Chordata" rank="species" species="woodwardi">
|
||
<emphasis id="B950EAA3FFCFB728FA89FA174CB6F9D4" box="[1319,1481,1515,1540]" italics="true" pageId="5" pageNumber="6">O. woodwardi</emphasis>
|
||
</taxonomicName>
|
||
(NHMUK PV R 3561), the dorsal margin of the iliac blade of
|
||
<taxonomicName id="4C244D32FFCFB728FC86F9D04AC0F995" authorityName="Müller & Baczko & Desojo & Nesbitt" authorityYear="2020" box="[808,959,1580,1605]" class="Reptilia" family="Ornithosuchidae" genus="Dynamosuchus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="5" pageNumber="6" phylum="Chordata" rank="species" species="collisensis">
|
||
<emphasis id="B950EAA3FFCFB728FC86F9D04AC0F995" box="[808,959,1580,1605]" italics="true" pageId="5" pageNumber="6">D. collisensis</emphasis>
|
||
</taxonomicName>
|
||
is straight to convex, whereas the homologous area in
|
||
<taxonomicName id="4C244D32FFCFB728FCD2F9B04D6CF9B5" baseAuthorityName="Bonaparte" baseAuthorityYear="1967" box="[892,1043,1612,1637]" class="Reptilia" family="Ornithosuchidae" genus="Riojasuchus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="5" pageNumber="6" phylum="Chordata" rank="species" species="tenuisceps">
|
||
<emphasis id="B950EAA3FFCFB728FCD2F9B04D6CF9B5" box="[892,1043,1612,1637]" italics="true" pageId="5" pageNumber="6">R. tenuisceps</emphasis>
|
||
</taxonomicName>
|
||
(PVL 3828) is concave. The supraacetabular crest merges into the main body of the bone approximately at the mid-length of the acetabulum in
|
||
<taxonomicName id="4C244D32FFCFB728FA82F9714CBDF976" authorityName="Müller & Baczko & Desojo & Nesbitt" authorityYear="2020" box="[1324,1474,1677,1702]" class="Reptilia" family="Ornithosuchidae" genus="Dynamosuchus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="5" pageNumber="6" phylum="Chordata" rank="species" species="collisensis">
|
||
<emphasis id="B950EAA3FFCFB728FA82F9714CBDF976" box="[1324,1474,1677,1702]" italics="true" pageId="5" pageNumber="6">D. collisensis</emphasis>
|
||
</taxonomicName>
|
||
. The postacetabular process is proportionally shorter than in
|
||
<taxonomicName id="4C244D32FFCFB728FC86F9314AC1F936" baseAuthorityName="Bonaparte" baseAuthorityYear="1967" box="[808,958,1741,1766]" class="Reptilia" family="Ornithosuchidae" genus="Riojasuchus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="5" pageNumber="6" phylum="Chordata" rank="species" species="tenuisceps">
|
||
<emphasis id="B950EAA3FFCFB728FC86F9314AC1F936" box="[808,958,1741,1766]" italics="true" pageId="5" pageNumber="6">R. tenuisceps</emphasis>
|
||
</taxonomicName>
|
||
(PVL 3828) and lacks any evidence of a brevis fossa on the ventral surface. There are three articular surfaces for the sacral ribs on the medial surface of the ilium, as in other ornithosuchids and poposauroids, differing from the rest of pseudosuchians, which typically possess two sacral vertebrae. Although the distal end of the pubis is not preserved, the length of the pubis is estimated to be markedly elongated (~0.68 times the total length of the femur) compared to other pseudosuchians (
|
||
<figureCitation id="131F2A34FFCCB72BFD96FF3C4BECFF09" box="[568,659,192,217]" captionStart="→" captionStartId="4.[116,144,1757,1787]" captionTargetId="figure-8@3.[98,1479,161,1951]" captionTargetPageId="3" captionText="→ Fig. 2. Representative skeletal elements of ornithosuchid archosaur Dynamosuchus collisensis gen. et sp. nov. (CAPPA/UFSM 0248) from Janner outcrop, Carnian, Late Triassic. A. Selected skull bones in left lateral view. B. Reconstruction of the skull. C. Skull in ventral view. D. Left quadrate and quadratojugal in posterodorsal view. E. Parabasisphenoid in left lateral view. F. Neural arch of an anterior cervical vertebra in anterior view. G. Centrum of a cervical vertebra in left lateral view. H. Right osteoderm in dorsal view. I. Neural arch of an anterior dorsal vertebra in left lateral view. J. Left ilium in lateral view. L. Right humerus in anterior view. M. Right forearm in medial view. N. Left manus in dorsal view. O. Right (reversed) pubis in lateral view. P. Left femur in anterior view. Q. Left fibula in lateral view. Some unpreserved portions are modified from Baczko et al. in press, for the reconstruction of the skeleton of CAPPA/UFSM 0248 (preserved elements indicated in orange) (K). Scale bars 20 mm." figureDoi="http://doi.org/10.5281/zenodo.11088354" httpUri="https://zenodo.org/record/11088354/files/figure.png" pageId="6" pageNumber="7">Fig. 2O</figureCitation>
|
||
). The pubis is anteroventrally projected, but gently arched ventrally along its length. It has a large obturator foramen and a well-developed pubic apron present along the shaft. The distal end is not preserved and it is not possible to determine whether it has a pubic boot or any expansion.
|
||
</paragraph>
|
||
<paragraph id="8B9B36B1FFCCB72BFF13FE784A46FADD" blockId="6.[151,825,160,1293]" pageId="6" pageNumber="7">
|
||
The hindlimb is estimated to be about to 1.6 times longer than the forelimb. The femur (
|
||
<figureCitation id="131F2A34FFCCB72BFE4BFE594B4BFE6E" box="[485,564,421,446]" captionStart="→" captionStartId="4.[116,144,1757,1787]" captionTargetId="figure-8@3.[98,1479,161,1951]" captionTargetPageId="3" captionText="→ Fig. 2. Representative skeletal elements of ornithosuchid archosaur Dynamosuchus collisensis gen. et sp. nov. (CAPPA/UFSM 0248) from Janner outcrop, Carnian, Late Triassic. A. Selected skull bones in left lateral view. B. Reconstruction of the skull. C. Skull in ventral view. D. Left quadrate and quadratojugal in posterodorsal view. E. Parabasisphenoid in left lateral view. F. Neural arch of an anterior cervical vertebra in anterior view. G. Centrum of a cervical vertebra in left lateral view. H. Right osteoderm in dorsal view. I. Neural arch of an anterior dorsal vertebra in left lateral view. J. Left ilium in lateral view. L. Right humerus in anterior view. M. Right forearm in medial view. N. Left manus in dorsal view. O. Right (reversed) pubis in lateral view. P. Left femur in anterior view. Q. Left fibula in lateral view. Some unpreserved portions are modified from Baczko et al. in press, for the reconstruction of the skeleton of CAPPA/UFSM 0248 (preserved elements indicated in orange) (K). Scale bars 20 mm." figureDoi="http://doi.org/10.5281/zenodo.11088354" httpUri="https://zenodo.org/record/11088354/files/figure.png" pageId="6" pageNumber="7">Fig. 2P</figureCitation>
|
||
) is sigmoidal with both ends expanded relative to the shaft. The proximal end preserves a well-developed posteromedial tuber, however, the anterolateral and anteromedial tubers as well as the greater trochanter have been eroded. There is a proximodistally oriented anterior trochanter at the proximal portion of the bone and a mound-like fourth trochanter more ventrally located at the opposite surface of the shaft. The presence of an anterior trochanter is a condition only registered in the ornithosuchids
|
||
<taxonomicName id="4C244D32FFCCB72BFE81FD3648B9FD33" baseAuthorityName="Bonaparte" baseAuthorityYear="1967" box="[303,454,714,739]" class="Reptilia" family="Ornithosuchidae" genus="Riojasuchus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="6" pageNumber="7" phylum="Chordata" rank="species" species="tenuisceps">
|
||
<emphasis id="B950EAA3FFCCB72BFE81FD3648B9FD33" box="[303,454,714,739]" italics="true" pageId="6" pageNumber="7">R. tenuisceps</emphasis>
|
||
</taxonomicName>
|
||
(PVL 3828) and
|
||
<taxonomicName id="4C244D32FFCCB72BFD3BFD364A47FD33" box="[661,824,714,739]" class="Reptilia" family="Ornithosuchidae" genus="Ornithosuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prolacertiformes" pageId="6" pageNumber="7" phylum="Chordata" rank="species" species="woodwardi">
|
||
<emphasis id="B950EAA3FFCCB72BFD3BFD364A47FD33" box="[661,824,714,739]" italics="true" pageId="6" pageNumber="7">O. woodwardi</emphasis>
|
||
</taxonomicName>
|
||
(NHMUK PV R 2410, NHMUK PV R 3561, NHMUK PV R 3916) within pseudosuchians (unknown in
|
||
<taxonomicName id="4C244D32FFCCB72BFDD5FCF74B8AFCF4" baseAuthorityName="PVL" baseAuthorityYear="2578" box="[635,757,779,804]" family="Ornithosuchidae" genus="Venaticosuchus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="6" pageNumber="7" phylum="Chordata" rank="species" species="rusconii">
|
||
<emphasis id="B950EAA3FFCCB72BFDD5FCF74B8AFCF4" box="[635,757,779,804]" italics="true" pageId="6" pageNumber="7">V. rusconii</emphasis>
|
||
</taxonomicName>
|
||
). The fourth trochanter of
|
||
<taxonomicName id="4C244D32FFCCB72BFE2AFCD04B62FC95" authorityName="Müller & Baczko & Desojo & Nesbitt" authorityYear="2020" box="[388,541,812,837]" class="Reptilia" family="Ornithosuchidae" genus="Dynamosuchus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="6" pageNumber="7" phylum="Chordata" rank="species" species="collisensis">
|
||
<emphasis id="B950EAA3FFCCB72BFE2AFCD04B62FC95" box="[388,541,812,837]" italics="true" pageId="6" pageNumber="7">D. collisensis</emphasis>
|
||
</taxonomicName>
|
||
is less pronounced than in
|
||
<taxonomicName id="4C244D32FFCCB72BFF17FCB1482DFCB6" baseAuthorityName="Bonaparte" baseAuthorityYear="1967" box="[185,338,845,870]" class="Reptilia" family="Ornithosuchidae" genus="Riojasuchus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="6" pageNumber="7" phylum="Chordata" rank="species" species="tenuisceps">
|
||
<emphasis id="B950EAA3FFCCB72BFF17FCB1482DFCB6" box="[185,338,845,870]" italics="true" pageId="6" pageNumber="7">R. tenuisceps</emphasis>
|
||
</taxonomicName>
|
||
(PVL 3828), resembling the condition of
|
||
<taxonomicName id="4C244D32FFCCB72BFF39FC914B39FC56" authority="(NHMUK PV R 2410)" baseAuthorityName="NHMUK PV R" baseAuthorityYear="2410" box="[151,582,877,902]" class="Reptilia" family="Ornithosuchidae" genus="Ornithosuchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prolacertiformes" pageId="6" pageNumber="7" phylum="Chordata" rank="species" species="woodwardi">
|
||
<emphasis id="B950EAA3FFCCB72BFF39FC914847FC56" box="[151,312,877,902]" italics="true" pageId="6" pageNumber="7">O. woodwardi</emphasis>
|
||
(NHMUK PV R 2410)
|
||
</taxonomicName>
|
||
. The anterior surface of the distal end of
|
||
<taxonomicName id="4C244D32FFCCB72BFED6FC724B6FFC77" authorityName="Müller & Baczko & Desojo & Nesbitt" authorityYear="2020" box="[376,528,910,935]" class="Reptilia" family="Ornithosuchidae" genus="Dynamosuchus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="6" pageNumber="7" phylum="Chordata" rank="species" species="collisensis">
|
||
<emphasis id="B950EAA3FFCCB72BFED6FC724B6FFC77" box="[376,528,910,935]" italics="true" pageId="6" pageNumber="7">D. collisensis</emphasis>
|
||
</taxonomicName>
|
||
is gently concave and the popliteal fossa, at the opposite surface, is proximodistally short. The distal condyles were eroded. The tibia is about 0.7 times of the length of the femur. The cnemial crest is poorly expanded from the shaft and has a straight anterior surface. The posterior edge of the proximal end is strongly projected. The fibula (
|
||
<figureCitation id="131F2A34FFCCB72BFE89FBAD48FCFBBA" box="[295,387,1105,1130]" captionStart="→" captionStartId="4.[116,144,1757,1787]" captionTargetId="figure-8@3.[98,1479,161,1951]" captionTargetPageId="3" captionText="→ Fig. 2. Representative skeletal elements of ornithosuchid archosaur Dynamosuchus collisensis gen. et sp. nov. (CAPPA/UFSM 0248) from Janner outcrop, Carnian, Late Triassic. A. Selected skull bones in left lateral view. B. Reconstruction of the skull. C. Skull in ventral view. D. Left quadrate and quadratojugal in posterodorsal view. E. Parabasisphenoid in left lateral view. F. Neural arch of an anterior cervical vertebra in anterior view. G. Centrum of a cervical vertebra in left lateral view. H. Right osteoderm in dorsal view. I. Neural arch of an anterior dorsal vertebra in left lateral view. J. Left ilium in lateral view. L. Right humerus in anterior view. M. Right forearm in medial view. N. Left manus in dorsal view. O. Right (reversed) pubis in lateral view. P. Left femur in anterior view. Q. Left fibula in lateral view. Some unpreserved portions are modified from Baczko et al. in press, for the reconstruction of the skeleton of CAPPA/UFSM 0248 (preserved elements indicated in orange) (K). Scale bars 20 mm." figureDoi="http://doi.org/10.5281/zenodo.11088354" httpUri="https://zenodo.org/record/11088354/files/figure.png" pageId="6" pageNumber="7">Fig. 2Q</figureCitation>
|
||
) has an elliptical proximal end with an expanded posterior portion. There is a well-developed tubercle for the attachment of the iliofibularis muscle at the proximal half of the bone. In
|
||
<taxonomicName id="4C244D32FFCCB72BFE4EFB4F4B0AFB1C" baseAuthorityName="Bonaparte" baseAuthorityYear="1967" box="[480,629,1203,1228]" class="Reptilia" family="Ornithosuchidae" genus="Riojasuchus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="6" pageNumber="7" phylum="Chordata" rank="species" species="tenuisceps">
|
||
<emphasis id="B950EAA3FFCCB72BFE4EFB4F4B0AFB1C" box="[480,629,1203,1228]" italics="true" pageId="6" pageNumber="7">R. tenuisceps</emphasis>
|
||
</taxonomicName>
|
||
(PVL 3827), this tubercle is more distally located (
|
||
<bibRefCitation id="EFB54B40FFCCB72BFDA4FB284A52FB3D" author="Baczko, M. B. von & Ezcurra, M. D." box="[522,813,1236,1261]" pageId="6" pageNumber="7" pagination="187 - 202" refId="ref7181" refString="Baczko, M. B. von and Ezcurra, M. D. 2013. Ornithosuchidae: a group of Triassic archosaurs with a unique ankle joint. Geological Society, London, Special Publications 379: 187 - 202." type="journal article" year="2013">Baczko and Ezcurra 2013</bibRefCitation>
|
||
). The specimen preserves slender gastralia (diameter 1.7 mm).
|
||
</paragraph>
|
||
<paragraph id="8B9B36B1FFCCB72BFF39FADC4852FA89" blockId="6.[151,824,1312,1369]" pageId="6" pageNumber="7">
|
||
<emphasis id="B950EAA3FFCCB72BFF39FADC4B3EFAE9" box="[151,577,1312,1337]" italics="true" pageId="6" pageNumber="7">Geographic and strtigraphic range</emphasis>
|
||
.—
|
||
<typeStatus id="549F8813FFCCB72BFDCFFADC4BE4FAE9" box="[609,667,1312,1337]" pageId="6" pageNumber="7">Type</typeStatus>
|
||
locality and horizon only.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |