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<document id="2B7654F634304021729366510E54C270" ID-CLB-Dataset="21904" ID-DOI="10.5852/ejt.2016.199" ID-GBIF-Dataset="d88184f4-ceef-4392-a444-0ac1552778d3" ID-ISSN="2118-9773" ID-Zenodo-Dep="3840484" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1590159499057" checkinUser="valdenar" docAuthor="Pimvichai, Piyatida, Enghoff, Henrik, Panha, Somsak &amp; Backeljau, Thierry" docDate="2016" docId="0385FA5C9600FFB2FD008E36DEA3C2DC" docLanguage="en" docName="ejt-2016-199.pdf.imf" docOrigin="European Journal of Taxonomy 199" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Thyropygus cimi Pimvichai &amp; Enghoff &amp; Panha &amp; Backeljau 2016, sp. nov." docType="treatment" docUuid="0252646C-AD4E-4B8B-8A7C-AEEA80DD4A72" docUuidSource="ZooBank" docVersion="13" lastPageNumber="16" masterDocId="FFBC8224960DFFBDFF848A31DC7FC003" masterDocTitle="A revision of the Thyropygus allevatus group. Part V: Nine new species of the extended opinatus subgroup, based on morphological and DNA sequence data (Diplopoda: Spirostreptida: Harpagophoridae)" masterLastPageNumber="37" masterPageNumber="1" pageNumber="14" updateTime="1701998209473" updateUser="plazi" zenodo-license-document="CC-BY-3.0" zenodo-license-figures="CC-BY-3.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="AAE3660E3CC044309FAF45D650FE3C89">A revision of the Thyropygus allevatus group. Part V: Nine new species of the extended opinatus subgroup, based on morphological and DNA sequence data (Diplopoda: Spirostreptida: Harpagophoridae)</mods:title>
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<mods:name id="5C6F40A2153BB41FE8C956578BE51C6F" type="personal">
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<mods:namePart id="0E03DFFBB65641CA0B38B63B0907B980">Pimvichai, Piyatida</mods:namePart>
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<mods:affiliation id="9FD2CBD8E7B52C6F7E496A79FE057BC8">Department of Biology, Faculty of Science, Mahasarakham University, Kantharawichai District, Maha Sarakham 44150, Thailand. &amp; urn: lsid: zoobank. org: author: 4 D 882 A 89 - E 0 C 1 - 4 F 78 - 93 E 7 - FA 1 B 5 A 2650 F 2 &amp; Corresponding author: piyatida _ pimvichai @ yahoo. com</mods:affiliation>
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<mods:namePart id="D32D70CB90EE54FF34F3CFE90F36D34A">Enghoff, Henrik</mods:namePart>
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<mods:affiliation id="E9792DDBD606A7232A622DEBF1FB3B2E">Natural History Museum of Denmark, University of Copenhagen, Universitetsparken 15, DK- 2100 Copenhagen Ø, Denmark. &amp; Email: henghoff @ snm. ku. dk &amp; urn: lsid: zoobank. org: author: FB 09 A 817 - 000 D- 43 C 3 - BCC 4 - 2 BC 1 E 5373635</mods:affiliation>
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<mods:namePart id="7179D440C61286CE6F613FC5F2FB19F1">Panha, Somsak</mods:namePart>
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<mods:affiliation id="AB22F25DA833C1FFB5F99EB4B3ED8A82">Animal Systematics Research Unit, Department of Biology, Faculty of Science, Chulalongkorn University, Bangkok 10330, Thailand. &amp; Email: somsakp @ sc. chula. ac. th &amp; urn: lsid: zoobank. org: author: AC 935098 - D 901 - 4 F 35 - A 414 - 4 B 0 D 4 FE 44 E 79</mods:affiliation>
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<mods:namePart id="002C886E3B49430CA8329951289AF270">Backeljau, Thierry</mods:namePart>
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<mods:affiliation id="F7B6E15D89889BBC1730CA0AB7F72BD8">University of Antwerp, Evolutionary Ecology Group, Groenenborgerlaan 171, B- 2020 Antwerp, Belgium and Royal Belgian Institute of Natural Sciences, OD Taxonomy and Phylogeny, and JEMU, Vautierstraat 29, B- 1000 Brussels, Belgium. &amp; Email: Thierry. Backeljau @ naturalsciences. be &amp; urn: lsid: zoobank. org: author: E 6 F 42575 - 36 AE- 4 AD 7 - 98 C 6 - D 083 EF 052568</mods:affiliation>
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<mods:date id="ECE725E1FF0DC831CB9BCBCFF4E566A0">2016</mods:date>
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<treatment id="0385FA5C9600FFB2FD008E36DEA3C2DC" ID-DOI="http://doi.org/10.5281/zenodo.3853336" ID-GBIF-Taxon="164249491" ID-Zenodo-Dep="3853336" LSID="urn:lsid:zoobank.org:act:0252646C-AD4E-4B8B-8A7C-AEEA80DD4A72" httpUri="http://treatment.plazi.org/id/0385FA5C9600FFB2FD008E36DEA3C2DC" lastPageId="15" lastPageNumber="16" pageId="13" pageNumber="14">
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<heading id="D0DBFC269600FFB0FD008E36DFCFC421" box="[644,944,1031,1058]" centered="true" fontSize="11" level="2" pageId="13" pageNumber="14" reason="2">
<taxonomicName id="4C2C30C99600FFB0FD008E36DF31C422" ID-CoL="56SZ9" authority="Pimvichai &amp; Enghoff &amp; Panha &amp; Backeljau, 2016" authorityName="Pimvichai &amp; Enghoff &amp; Panha &amp; Backeljau" authorityYear="2016" box="[644,846,1031,1058]" class="Diplopoda" family="Harpagophoridae" genus="Thyropygus" kingdom="Animalia" order="Spirostreptida" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="cimi" status="sp. nov.">
<emphasis id="B95897589600FFB0FD008E36DF31C422" bold="true" box="[644,846,1031,1058]" italics="true" pageId="13" pageNumber="14">Thyropygus cimi</emphasis>
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<taxonomicNameLabel id="A26B2A239600FFB0FCD28E39DFCFC421" box="[854,944,1032,1058]" pageId="13" pageNumber="14" rank="species">sp. nov.</taxonomicNameLabel>
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urn:lsid:zoobank.org:act:
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<paragraph id="8B934B4A9600FFB0FD598E7FDF28C46B" blockId="13.[352,1235,1031,1128]" box="[733,855,1102,1128]" pageId="13" pageNumber="14">
<figureCitation id="131757CF9600FFB0FD598E7FDF5EC46B" box="[733,801,1102,1128]" captionStart="Fig" captionStartId="14.[189,232,1906,1932]" captionTargetBox="[200,1383,662,1870]" captionTargetId="figure@14.[189,1398,654,1878]" captionTargetPageId="14" captionText="Fig. 2. Thyropygus cimi sp. nov., from Namwang Srithammasokrach, holotype (CUMZ-D00086), ³, gonopods. A. Anterior view, left telopodite removed. B. Posterior view, left telopodite removed. C. Lateral view. D. Left telopodite, posterior-mesal view. E. Left telopodite, anterior-lateral view." figureDoi="http://doi.org/10.5281/zenodo.3840488" httpUri="https://zenodo.org/record/3840488/files/figure.png" pageId="13" pageNumber="14">Fig. 2</figureCitation>
AE
</paragraph>
</subSubSection>
<subSubSection id="C33618C19600FFB0FF398EBEDFFDC566" pageId="13" pageNumber="14" type="diagnosis">
<paragraph id="8B934B4A9600FFB0FF398EBEDD49C4AA" blockId="13.[189,310,1167,1193]" box="[189,310,1167,1193]" pageId="13" pageNumber="14">
<heading id="D0DBFC269600FFB0FF398EBEDD49C4AA" bold="true" box="[189,310,1167,1193]" fontSize="11" level="3" pageId="13" pageNumber="14" reason="3">
<emphasis id="B95897589600FFB0FF398EBEDD49C4AA" bold="true" box="[189,310,1167,1193]" pageId="13" pageNumber="14">Diagnosis</emphasis>
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A species of the
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<emphasis id="B95897589600FFB0FEFF8E8EDD9FC4DB" box="[379,480,1215,1240]" italics="true" pageId="13" pageNumber="14">opinatus</emphasis>
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subgroup. Lateral process of anterior coxal fold (
<emphasis id="B95897589600FFB0FBA48E8FD83AC4DB" box="[1056,1093,1214,1240]" italics="true" pageId="13" pageNumber="14">alp</emphasis>
) regularly curved, basally broad, gradually tapering towards end and ending in sharp point, with a lateral crest near tip. Similar in this respect to
<taxonomicName id="4C2C30C99600FFB0FE0B8F34DE71C51D" baseAuthorityName="Demange" baseAuthorityYear="1986" box="[399,526,1284,1310]" class="Diplopoda" family="Harpagophoridae" genus="Thyropygus" kingdom="Animalia" order="Spirostreptida" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="bifurcus">
<emphasis id="B95897589600FFB0FE0B8F34DE71C51D" box="[399,526,1284,1310]" italics="true" pageId="13" pageNumber="14">T. bifurcus</emphasis>
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. Differs from this species by having the mesal process of anterior coxal fold (
<emphasis id="B95897589600FFB0FF7A8F19DD4EC542" box="[254,305,1320,1345]" italics="true" pageId="13" pageNumber="14">amp</emphasis>
) straight, directed distad, by having the mesal process of posterior coxal fold (
<emphasis id="B95897589600FFB0FB538F19D975C542" box="[1239,1290,1320,1345]" italics="true" pageId="13" pageNumber="14">pmp</emphasis>
) slender, directed anteriad, and by having only one femoral spine (
<emphasis id="B95897589600FFB0FCD88F7ADF0EC566" box="[860,881,1355,1381]" italics="true" pageId="13" pageNumber="14">fe</emphasis>
).
</paragraph>
</subSubSection>
<subSubSection id="C33618C19600FFB0FF398FBDDD3EC618" pageId="13" pageNumber="14" type="etymology">
<paragraph id="8B934B4A9600FFB0FF398FBDDD3BC5A5" blockId="13.[189,324,1420,1446]" box="[189,324,1420,1446]" pageId="13" pageNumber="14">
<heading id="D0DBFC269600FFB0FF398FBDDD3BC5A5" bold="true" box="[189,324,1420,1446]" fontSize="11" level="3" pageId="13" pageNumber="14" reason="3">
<emphasis id="B95897589600FFB0FF398FBDDD3BC5A5" bold="true" box="[189,324,1420,1446]" pageId="13" pageNumber="14">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="8B934B4A9600FFB0FF398F8ADD3EC618" blockId="13.[189,1399,1467,1563]" pageId="13" pageNumber="14">
This species is named after the organization “
<collectingRegion id="49E885A89600FFB0FD5A8F8ADF50C5D6" box="[734,815,1467,1493]" country="France" name="Centre" pageId="13" pageNumber="14">Centre</collectingRegion>
International de Myriapodologie CIM” (www. myriapodology.org) in recognition of its immense importance for inspiring and supporting research on myriapods.
</paragraph>
</subSubSection>
<subSubSection id="C33618C19600FFB0FF398C73D908C751" pageId="13" pageNumber="14" type="materials_examined">
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<heading id="D0DBFC269600FFB0FF398C73DDD5C65F" bold="true" box="[189,426,1602,1628]" fontSize="11" level="3" pageId="13" pageNumber="14" reason="3">
<emphasis id="B95897589600FFB0FF398C73DDD5C65F" bold="true" box="[189,426,1602,1628]" pageId="13" pageNumber="14">Material examined</emphasis>
</heading>
</paragraph>
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<typeStatus id="5497F5E89600FFB0FF6E8C40DD23C688" box="[234,348,1649,1675]" pageId="13" pageNumber="14" type="holotype">Holotype</typeStatus>
</emphasis>
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<collectingCountry id="F33B0BDA9600FFB0FF398CA5DD26C6AD" box="[189,345,1684,1710]" name="Thailand" pageId="13" pageNumber="14">THAILAND</collectingCountry>
: ³,
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,
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,
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,
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,
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,
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<collectingDate id="EFD694629600FFB0FD858C86DED3C6D1" box="[513,684,1719,1746]" pageId="13" pageNumber="14" value="2010-05-16">16 May 2010</collectingDate>
</date>
, leg.
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, P. Pimvichai and members of the
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(
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).
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<typeStatus id="5497F5E89600FFB0FF6E8D24DD18C72C" box="[234,359,1813,1839]" pageId="13" pageNumber="14" type="paratype">Paratypes</typeStatus>
</emphasis>
</paragraph>
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<collectingCountry id="F33B0BDA9600FFB0FF398D09DD26C751" box="[189,345,1848,1874]" name="Thailand" pageId="13" pageNumber="14">THAILAND</collectingCountry>
: 2 ³³,
<specimenCount id="9D2A80C39600FFB0FE348D09DDA4C751" box="[432,475,1848,1874]" pageId="13" pageNumber="14" type="female">1 ♀</specimenCount>
, same data as
<typeStatus id="5497F5E89600FFB0FD048D09DE98C751" box="[640,743,1848,1874]" pageId="13" pageNumber="14" type="holotype">holotype</typeStatus>
(
<collectionCode id="ED3DD38F9600FFB0FD708D09DF13C750" box="[756,876,1848,1875]" pageId="13" pageNumber="14">CUMZ-D</collectionCode>
00096); 1 ³, same data as
<typeStatus id="5497F5E89600FFB0FB1E8D09D97EC751" box="[1178,1281,1848,1874]" pageId="13" pageNumber="14" type="holotype">holotype</typeStatus>
(
<collectionCode id="ED3DD38F9600FFB0FA8B8D09D914C751" box="[1295,1387,1848,1874]" country="Denmark" httpUri="http://grbio.org/cool/mci8-ehqk" name="Zoological Museum, University of Copenhagen" pageId="13" pageNumber="14">ZMUC</collectionCode>
).
</paragraph>
</subSubSection>
<subSubSection id="C33618C19600FFB2FF398D48DE26C26C" lastPageId="15" lastPageNumber="16" pageId="13" pageNumber="14" type="description">
<paragraph id="8B934B4A9600FFB0FF398D48DD31C790" blockId="13.[189,334,1913,1939]" box="[189,334,1913,1939]" pageId="13" pageNumber="14">
<heading id="D0DBFC269600FFB0FF398D48DD31C790" bold="true" box="[189,334,1913,1939]" fontSize="11" level="3" pageId="13" pageNumber="14" reason="3">
<emphasis id="B95897589600FFB0FF398D48DD31C790" bold="true" box="[189,334,1913,1939]" pageId="13" pageNumber="14">Description</emphasis>
</heading>
</paragraph>
<paragraph id="8B934B4A9600FFB0FF398D99D909C7E6" blockId="13.[189,1399,1959,2022]" pageId="13" pageNumber="14">
Adult males with 6568 podous rings, no apodous rings. Length
<quantity id="4CD4E6AF9600FFB0FC418D99D841C7C1" box="[965,1086,1960,1986]" metricMagnitude="-1" metricUnit="m" metricValue="1.45" metricValueMax="1.6" metricValueMin="1.3" pageId="13" pageNumber="14" unit="cm" value="14.5" valueMax="16.0" valueMin="13.0">1316 cm</quantity>
, width
<quantity id="4CD4E6AF9600FFB0FB138D99D956C7C1" box="[1175,1321,1960,1986]" metricMagnitude="-3" metricUnit="m" metricValue="7.95" metricValueMax="8.3" metricValueMin="7.6" pageId="13" pageNumber="14" unit="mm" value="7.95" valueMax="8.3" valueMin="7.6">7.68.3 mm</quantity>
. Adult female with 65 podous rings, no apodous rings. Length
<quantity id="4CD4E6AF9600FFB0FCCB8DFADFE4C7E6" box="[847,923,1995,2021]" metricMagnitude="-1" metricUnit="m" metricValue="1.3" pageId="13" pageNumber="14" unit="cm" value="13.0">13 cm</quantity>
, width
<quantity id="4CD4E6AF9600FFB0FC758DFAD830C7E6" box="[1009,1103,1995,2021]" metricMagnitude="-3" metricUnit="m" metricValue="7.4" pageId="13" pageNumber="14" unit="mm" value="7.4">7.4 mm</quantity>
. Colour in life unknown;
</paragraph>
<paragraph id="8B934B4A9603FFB3FF398B3ADE94C14B" blockId="14.[189,1398,267,329]" pageId="14" pageNumber="15">preserved specimens with head, antennae, prozona, middorsal metazona and legs brown; metazona, epiproct, paraprocts and hypoproct dark brown.</paragraph>
<paragraph id="8B934B4A9603FFB2FF398B41DDC4C16F" blockId="14.[189,1399,368,606]" lastBlockId="15.[189,1398,267,364]" lastPageId="15" lastPageNumber="16" pageId="14" pageNumber="15">
GONOPODS (
<figureCitation id="131757CF9603FFB3FEC08B41DDF6C189" box="[324,393,368,394]" captionStart="Fig" captionStartId="14.[189,232,1906,1932]" captionTargetBox="[200,1383,662,1870]" captionTargetId="figure@14.[189,1398,654,1878]" captionTargetPageId="14" captionText="Fig. 2. Thyropygus cimi sp. nov., from Namwang Srithammasokrach, holotype (CUMZ-D00086), ³, gonopods. A. Anterior view, left telopodite removed. B. Posterior view, left telopodite removed. C. Lateral view. D. Left telopodite, posterior-mesal view. E. Left telopodite, anterior-lateral view." figureDoi="http://doi.org/10.5281/zenodo.3840488" httpUri="https://zenodo.org/record/3840488/files/figure.png" pageId="14" pageNumber="15">Fig. 2</figureCitation>
AE). Anterior coxal fold (
<emphasis id="B95897589603FFB3FD428B40DE9DC189" box="[710,738,369,394]" italics="true" pageId="14" pageNumber="15">ac</emphasis>
;
<figureCitation id="131757CF9603FFB3FD758B41DF2FC189" box="[753,848,368,394]" captionStart="Fig" captionStartId="14.[189,232,1906,1932]" captionTargetBox="[200,1383,662,1870]" captionTargetId="figure@14.[189,1398,654,1878]" captionTargetPageId="14" captionText="Fig. 2. Thyropygus cimi sp. nov., from Namwang Srithammasokrach, holotype (CUMZ-D00086), ³, gonopods. A. Anterior view, left telopodite removed. B. Posterior view, left telopodite removed. C. Lateral view. D. Left telopodite, posterior-mesal view. E. Left telopodite, anterior-lateral view." figureDoi="http://doi.org/10.5281/zenodo.3840488" httpUri="https://zenodo.org/record/3840488/files/figure.png" pageId="14" pageNumber="15">Fig. 2A</figureCitation>
): lateral process (
<emphasis id="B95897589603FFB3FB998B41D83DC189" box="[1053,1090,368,394]" italics="true" pageId="14" pageNumber="15">alp</emphasis>
) regularly curved, basally broad, gradually tapering towards end and ending in sharp point, with lateral crest near tip (
<figureCitation id="131757CF9603FFB3FA908BA2D90EC1AE" box="[1300,1393,403,429]" captionStart="Fig" captionStartId="14.[189,232,1906,1932]" captionTargetBox="[200,1383,662,1870]" captionTargetId="figure@14.[189,1398,654,1878]" captionTargetPageId="14" captionText="Fig. 2. Thyropygus cimi sp. nov., from Namwang Srithammasokrach, holotype (CUMZ-D00086), ³, gonopods. A. Anterior view, left telopodite removed. B. Posterior view, left telopodite removed. C. Lateral view. D. Left telopodite, posterior-mesal view. E. Left telopodite, anterior-lateral view." figureDoi="http://doi.org/10.5281/zenodo.3840488" httpUri="https://zenodo.org/record/3840488/files/figure.png" pageId="14" pageNumber="15">Fig. 2C</figureCitation>
, arrow); mesal process (
<emphasis id="B95897589603FFB3FE588B86DE70C1D3" box="[476,527,439,464]" italics="true" pageId="14" pageNumber="15">amp</emphasis>
) slightly shorter than
<emphasis id="B95897589603FFB3FCA58B87DF39C1D3" box="[801,838,438,464]" italics="true" pageId="14" pageNumber="15">alp</emphasis>
, straight, directed distad. Posterior coxal fold (
<emphasis id="B95897589603FFB3FF438BEBDC9CC1F0" box="[199,227,474,499]" italics="true" pageId="14" pageNumber="15">pc</emphasis>
;
<figureCitation id="131757CF9603FFB3FF778BE8DD32C1F7" box="[243,333,473,500]" captionStart="Fig" captionStartId="14.[189,232,1906,1932]" captionTargetBox="[200,1383,662,1870]" captionTargetId="figure@14.[189,1398,654,1878]" captionTargetPageId="14" captionText="Fig. 2. Thyropygus cimi sp. nov., from Namwang Srithammasokrach, holotype (CUMZ-D00086), ³, gonopods. A. Anterior view, left telopodite removed. B. Posterior view, left telopodite removed. C. Lateral view. D. Left telopodite, posterior-mesal view. E. Left telopodite, anterior-lateral view." figureDoi="http://doi.org/10.5281/zenodo.3840488" httpUri="https://zenodo.org/record/3840488/files/figure.png" pageId="14" pageNumber="15">Fig. 2B</figureCitation>
) basally with moderately high lateral paracoxites (
<emphasis id="B95897589603FFB3FC2D8BEBDFBAC1F0" box="[937,965,474,499]" italics="true" pageId="14" pageNumber="15">px</emphasis>
), distally truncate, forming shelf for accommodation of telopodite, mesal process (
<emphasis id="B95897589603FFB3FD658BCFDF6BC214" box="[737,788,510,535]" italics="true" pageId="14" pageNumber="15">pmp</emphasis>
) slender, directed anteriad. Telopodite (
<figureCitation id="131757CF9603FFB3FB748BCCD949C214" box="[1264,1334,509,535]" captionStart="Fig" captionStartId="14.[189,232,1906,1932]" captionTargetBox="[200,1383,662,1870]" captionTargetId="figure@14.[189,1398,654,1878]" captionTargetPageId="14" captionText="Fig. 2. Thyropygus cimi sp. nov., from Namwang Srithammasokrach, holotype (CUMZ-D00086), ³, gonopods. A. Anterior view, left telopodite removed. B. Posterior view, left telopodite removed. C. Lateral view. D. Left telopodite, posterior-mesal view. E. Left telopodite, anterior-lateral view." figureDoi="http://doi.org/10.5281/zenodo.3840488" httpUri="https://zenodo.org/record/3840488/files/figure.png" pageId="14" pageNumber="15">Fig. 2</figureCitation>
DE) leaving coxite over shelf of posterior coxal fold; femoral spine (
<emphasis id="B95897589603FFB3FC128811DFD4C239" box="[918,939,544,570]" italics="true" pageId="14" pageNumber="15">fe</emphasis>
) very long, slender, curving downward,
<emphasis id="B95897589603FFB3FF398875DD78C25E" box="[189,263,580,605]" italics="true" pageId="14" pageNumber="15">in situ</emphasis>
resting between
<emphasis id="B95897589603FFB3FE518872DD85C25E" box="[469,506,579,605]" italics="true" pageId="14" pageNumber="15">alp</emphasis>
and
<emphasis id="B95897589603FFB3FDBC8875DE14C25E" box="[568,619,580,605]" italics="true" pageId="14" pageNumber="15">amp</emphasis>
; tibial spine (
<emphasis id="B95897589603FFB3FC968875DF5DC25E" box="[786,802,580,605]" italics="true" pageId="14" pageNumber="15">ti</emphasis>
) long, slender, curving in horizontal plane, its tip
<emphasis id="B95897589602FFB2FF398B3DDD78C126" box="[189,263,268,293]" italics="true" pageId="15" pageNumber="16">in situ</emphasis>
resting close to base of
<emphasis id="B95897589602FFB2FDAB8B3DDE3BC125" box="[559,580,268,294]" italics="true" pageId="15" pageNumber="16">fe</emphasis>
; apical part: lamellar lobe (
<emphasis id="B95897589602FFB2FC178B3ADFDCC126" box="[915,931,267,293]" italics="true" pageId="15" pageNumber="16">ll</emphasis>
) broadly expanded, bent down; palette (
<emphasis id="B95897589602FFB2FF438B1EDC9BC14B" box="[199,228,303,328]" italics="true" pageId="15" pageNumber="16">pa</emphasis>
) simple, gutter-like, with longitudinal rounded crest (
<emphasis id="B95897589602FFB2FCDC8B1EDF0FC14B" box="[856,880,303,328]" italics="true" pageId="15" pageNumber="16">cr</emphasis>
) at middle; distally with about ten brownish blepharochaetae (
<emphasis id="B95897589602FFB2FE098B63DDD5C16F" box="[397,426,338,364]" italics="true" pageId="15" pageNumber="16">bp</emphasis>
).
</paragraph>
<caption id="DF531BC29603FFB3FF398D43D949C7D0" ID-DOI="http://doi.org/10.5281/zenodo.3840488" ID-Zenodo-Dep="3840488" httpUri="https://zenodo.org/record/3840488/files/figure.png" pageId="14" pageNumber="15" startId="14.[189,232,1906,1932]" targetBox="[200,1383,662,1870]" targetPageId="14">
<paragraph id="8B934B4A9603FFB3FF398D43D949C7D0" blockId="14.[189,1399,1906,2003]" pageId="14" pageNumber="15">
<emphasis id="B95897589603FFB3FF398D43DD6EC78F" bold="true" box="[189,273,1906,1932]" pageId="14" pageNumber="15">Fig. 2.</emphasis>
<taxonomicName id="4C2C30C99603FFB3FE9B8D43DD96C78F" authority="Pimvichai &amp; Enghoff &amp; Panha &amp; Backeljau, 2016" authorityName="Pimvichai &amp; Enghoff &amp; Panha &amp; Backeljau" authorityYear="2016" box="[287,489,1906,1932]" class="Diplopoda" family="Harpagophoridae" genus="Thyropygus" kingdom="Animalia" order="Spirostreptida" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="cimi" status="sp. nov.">
<emphasis id="B95897589603FFB3FE9B8D43DD96C78F" box="[287,489,1906,1932]" italics="true" pageId="14" pageNumber="15">Thyropygus cimi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26B2A239603FFB3FE728D42DE25C78E" box="[502,602,1907,1933]" pageId="14" pageNumber="15" rank="species">sp. nov.</taxonomicNameLabel>
, from Namwang Srithammasokrach, holotype (CUMZ-D00086), ³, gonopods.
<emphasis id="B95897589603FFB3FEEE8DA4DD00C7AC" bold="true" box="[362,383,1941,1967]" pageId="14" pageNumber="15">A</emphasis>
. Anterior view, left telopodite removed.
<emphasis id="B95897589603FFB3FCF68DA7DFF9C7B3" bold="true" box="[882,902,1942,1968]" pageId="14" pageNumber="15">B</emphasis>
. Posterior view, left telopodite removed.
<emphasis id="B95897589603FFB3FF398D88DCADC7D0" bold="true" box="[189,210,1977,2003]" pageId="14" pageNumber="15">C</emphasis>
. Lateral view.
<emphasis id="B95897589603FFB3FE058D88DDE9C7D0" bold="true" box="[385,406,1977,2003]" pageId="14" pageNumber="15">D</emphasis>
. Left telopodite, posterior-mesal view.
<emphasis id="B95897589603FFB3FCE08D88DF07C7D0" bold="true" box="[868,888,1977,2003]" pageId="14" pageNumber="15">E</emphasis>
. Left telopodite, anterior-lateral view.
</paragraph>
</caption>
<paragraph id="8B934B4A9602FFB2FF398BA2DD19C1AE" blockId="15.[189,358,403,429]" box="[189,358,403,429]" pageId="15" pageNumber="16">
<heading id="D0DBFC269602FFB2FF398BA2DD19C1AE" bold="true" box="[189,358,403,429]" fontSize="11" level="3" pageId="15" pageNumber="16" reason="3">
<emphasis id="B95897589602FFB2FF398BA2DD19C1AE" bold="true" box="[189,358,403,429]" pageId="15" pageNumber="16">DNA barcode</emphasis>
</heading>
</paragraph>
<paragraph id="8B934B4A9602FFB2FF398BF3DDF3C203" blockId="15.[189,1398,450,512]" pageId="15" pageNumber="16">
The GenBank accession number of the barcode of the
<typeStatus id="5497F5E89602FFB2FC118BF3DF83C1DF" box="[917,1020,450,476]" pageId="15" pageNumber="16" type="holotype">holotype</typeStatus>
is
<accessionNumber id="947FD6A99602FFB2FBB68BF3D8CBC1DE" box="[1074,1204,450,477]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/KU306519" pageId="15" pageNumber="16">KU306519</accessionNumber>
(voucher code CUMZ-D00086).
</paragraph>
<paragraph id="8B934B4A9602FFB2FF398817DDBBC243" blockId="15.[189,452,550,576]" box="[189,452,550,576]" pageId="15" pageNumber="16">
<emphasis id="B95897589602FFB2FF398817DD29C243" bold="true" box="[189,342,550,576]" pageId="15" pageNumber="16">Distribution</emphasis>
(
<figureCitation id="131757CF9602FFB2FEE28817DDC4C243" box="[358,443,550,576]" captionStart="Fig" captionStartId="30.[189,232,1998,2024]" captionTargetBox="[189,1398,829,1964]" captionTargetId="figure@30.[189,1398,829,1964]" captionTargetPageId="30" captionText="Fig. 12. Known distribution of the species of the T. opinatus subgroup." figureDoi="http://doi.org/10.5281/zenodo.3840510" httpUri="https://zenodo.org/record/3840510/files/figure.png" pageId="15" pageNumber="16">Fig. 12</figureCitation>
)
</paragraph>
<paragraph id="8B934B4A9602FFB2FF398864DE26C26C" blockId="15.[189,601,597,623]" box="[189,601,597,623]" pageId="15" pageNumber="16">
Known only from the
<typeStatus id="5497F5E89602FFB2FE468864DD8AC26C" box="[450,501,597,623]" pageId="15" pageNumber="16">type</typeStatus>
locality.
</paragraph>
</subSubSection>
<subSubSection id="C33618C19602FFB2FF3988A6DEA3C2DC" box="[189,732,663,735]" pageId="15" pageNumber="16" type="discussion">
<paragraph id="8B934B4A9602FFB2FF3988A6DD50C2B2" blockId="15.[189,303,663,689]" box="[189,303,663,689]" pageId="15" pageNumber="16">
<heading id="D0DBFC269602FFB2FF3988A6DD50C2B2" bold="true" box="[189,303,663,689]" fontSize="11" level="3" pageId="15" pageNumber="16" reason="3">
<emphasis id="B95897589602FFB2FF3988A6DD50C2B2" bold="true" box="[189,303,663,689]" pageId="15" pageNumber="16">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="8B934B4A9602FFB2FF3988F4DEA3C2DC" blockId="15.[189,732,709,735]" box="[189,732,709,735]" pageId="15" pageNumber="16">
Coexisting with the smaller
<taxonomicName id="4C2C30C99602FFB2FD8C88F7DE05C2DC" authority="Pimvichai &amp; Enghoff &amp; Panha &amp; Backeljau, 2016" authorityName="Pimvichai &amp; Enghoff &amp; Panha &amp; Backeljau" authorityYear="2016" box="[520,634,709,735]" class="Diplopoda" family="Harpagophoridae" genus="Thyropygus" kingdom="Animalia" order="Spirostreptida" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="forceps" status="sp. nov.">
<emphasis id="B95897589602FFB2FD8C88F7DE05C2DC" box="[520,634,709,735]" italics="true" pageId="15" pageNumber="16">T. forceps</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26B2A239602FFB2FD0588F4DEA3C2DC" box="[641,732,709,735]" pageId="15" pageNumber="16" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
</treatment>
</document>