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<mods:title id="DECCD7B9EDB713B7C13D7F05B0EA1066">The genera Agathocles and Surenus (Hemiptera: Heteroptera: Pentatomidae) tribal reassessment, redescription, new synonyms, and description of two new species</mods:title>
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<mods:namePart id="D83E976B1ED2B9783C93C4BAC0143851">Kment, Petr</mods:namePart>
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<mods:affiliation id="C573B9E8C42FA7E061589D6766D09389">Department of Entomology, National Museum, Cirkusová 1740, CZ- 193 00 Praha 9 - Horní Počernice, Czech Republic sigara @ post. cz; https: // orcid. org / 0000 - 0002 - 7026 - 5691</mods:affiliation>
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<taxonomicName id="4C2C4D616804FFC64E8D66BCFD8FFE2C" authority="Ahmad &amp; Afzal, 1977" authorityName="Ahmad &amp; Afzal" authorityYear="1977" box="[151,626,402,429]" class="Insecta" family="Pentatomidae" genus="Riazocoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="39" pageNumber="549" phylum="Arthropoda" rank="species" species="niger">
<emphasis id="B958EAF06804FFC64E8D66BCFD8FFE2C" bold="true" box="[151,626,402,429]" pageId="39" pageNumber="549">
<emphasis id="B958EAF06804FFC64E8D66BCFEA3FE2D" bold="true" box="[151,350,403,429]" italics="true" pageId="39" pageNumber="549">Riazocoris niger</emphasis>
<bibRefCitation id="EFBD4B136804FFC64F7966BDFD8FFE2C" author="Ahmad, I. &amp; Afzal, M. &amp; Siddiqui, M. I." box="[355,626,402,428]" pageId="39" pageNumber="549" pagination="161 - 165" refId="ref27925" refString="Ahmad, I., Afzal, M. &amp; Siddiqui, M. I. (1977) Aspects of morphology of Riazocoris nigra Ahmad and Afzal (Pentatominae: Caystrini) with phylogenetic considerations. Pakistan Journal of Zoology, 9 (2), 161 - 165." type="journal article" year="1977">Ahmad &amp; Afzal, 1977</bibRefCitation>
</emphasis>
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</paragraph>
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<subSubSection id="C33665696804FFC64E8D66F7FD6AFC57" pageId="39" pageNumber="549" type="reference_group">
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<treatmentCitationGroup id="AB3C11CC6804FFC64E8D66F7FD5DFD8E" pageId="39" pageNumber="549">
<taxonomicName id="4C2C4D616804FFC64E8D66F7FDCEFE6F" authority="Ahmad &amp; Afzal, 1977" authorityName="Ahmad &amp; Afzal" authorityYear="1977" box="[151,563,471,495]" class="Insecta" family="Pentatomidae" genus="Riazocoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="39" pageNumber="549" phylum="Arthropoda" rank="species" species="nigra">
<emphasis id="B958EAF06804FFC64E8D66F7FEBEFE6F" box="[151,323,472,495]" italics="true" pageId="39" pageNumber="549">Riazocoris nigra</emphasis>
<bibRefCitation id="EFBD4B136804FFC64F5066F8FDCEFE6F" author="Ahmad, I. &amp; Afzal, M. &amp; Siddiqui, M. I." box="[330,563,471,495]" pageId="39" pageNumber="549" pagination="161 - 165" refId="ref27925" refString="Ahmad, I., Afzal, M. &amp; Siddiqui, M. I. (1977) Aspects of morphology of Riazocoris nigra Ahmad and Afzal (Pentatominae: Caystrini) with phylogenetic considerations. Pakistan Journal of Zoology, 9 (2), 161 - 165." type="journal article" year="1977">
Ahmad &amp; Afzal,
<quantity id="4CD49B076804FFC64C1A66F8FDCEFE6F" box="[512,563,471,495]" metricMagnitude="1" metricUnit="m" metricValue="5.02158" pageId="39" pageNumber="549" unit="in" value="1977.0">1977</quantity>
</bibRefCitation>
</taxonomicName>
in
<treatmentCitation id="0A8D10F36804FFC64C4C66F8FC9FFE6F" author="Ahmad, I. &amp; Afzal, M. &amp; Siddiqui, M. I." box="[598,866,471,495]" page="161" pageId="39" pageNumber="549" year="1977">
<bibRefCitation id="EFBD4B136804FFC64C4C66F8FC9FFE6F" author="Ahmad, I. &amp; Afzal, M. &amp; Siddiqui, M. I." box="[598,866,471,495]" pageId="39" pageNumber="549" pagination="161 - 165" refId="ref27925" refString="Ahmad, I., Afzal, M. &amp; Siddiqui, M. I. (1977) Aspects of morphology of Riazocoris nigra Ahmad and Afzal (Pentatominae: Caystrini) with phylogenetic considerations. Pakistan Journal of Zoology, 9 (2), 161 - 165." type="journal article" year="1977">
Ahmad
<emphasis id="B958EAF06804FFC64CB366F7FD23FE6F" box="[681,734,471,495]" italics="true" pageId="39" pageNumber="549">et al.</emphasis>
(1977: 161)
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</treatmentCitation>
.
<typeStatus id="549788406804FFC64D7466F7FC2EFE70" box="[878,979,472,496]" pageId="39" pageNumber="549" type="lectotype">Lectotype</typeStatus>
(designated by
<treatmentCitation id="0A8D10F36804FFC64A6066F8FA84FE6F" author="Ahmad, I. &amp; Afzal, M." box="[1146,1401,471,495]" page="7" pageId="39" pageNumber="549" year="1979">
<bibRefCitation id="EFBD4B136804FFC64A6066F8FA84FE6F" author="Ahmad, I. &amp; Afzal, M." box="[1146,1401,471,495]" pageId="39" pageNumber="549" pagination="1 - 14" refId="ref27835" refString="Ahmad, I. &amp; Afzal, M. (1979) Resurrection of the tribe Caystrini Stal (Heteroptera, Pentatomidae, Pentatominae) with description of two new genera from Oriental region. Annotationes Zoologicae et Botanicae, 133, 1 - 14." type="journal article" year="1979">Ahmad &amp; Afzal 1979: 7</bibRefCitation>
</treatmentCitation>
, as
<typeStatus id="549788406804FFC64EDD66D9FED8FD8E" box="[199,293,502,526]" pageId="39" pageNumber="549" type="holotype">holotype</typeStatus>
):
<specimenCount id="9D2AFD6B6804FFC64F2C66D9FEB6FD8D" box="[310,331,502,525]" pageId="39" pageNumber="549" type="male"></specimenCount>
,
<collectingCountry id="F33B76726804FFC64F4F66D9FE52FD8E" box="[341,431,502,526]" name="Pakistan" pageId="39" pageNumber="549">Pakistan</collectingCountry>
:
<collectingRegion id="49E8F8006804FFC64FA366D9FE16FD8E" box="[441,491,502,526]" country="Pakistan" name="Sindh" pageId="39" pageNumber="549">Sind</collectingRegion>
: Thatta (NHMK).
</treatmentCitationGroup>
</paragraph>
<paragraph id="8B9336E26804FFC64E8D653AFD45FDCB" blockId="39.[151,1437,471,587]" pageId="39" pageNumber="549">
<treatmentCitationGroup id="AB3C11CC6804FFC64E8D653AFD45FDCB" pageId="39" pageNumber="549">
<treatmentCitation id="0A8D10F36804FFC64E8D653AFDA2FDAD" author="Ahmad, I. &amp; Afzal, M." box="[151,607,532,557]" page="7" pageId="39" pageNumber="549" year="1979">
<taxonomicName id="4C2C4D616804FFC64E8D653AFDA2FDAD" authority=": Ahmad &amp; Afzal, 1979: 4 - 7" authorityName=": Ahmad &amp; Afzal" authorityPageNumber="4 - 7" authorityYear="1979" box="[151,607,532,557]" class="Insecta" family="Pentatomidae" genus="Riazocoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="39" pageNumber="549" phylum="Arthropoda" rank="species" species="nigra">
<emphasis id="B958EAF06804FFC64E8D653AFEBCFDAC" box="[151,321,533,556]" italics="true" pageId="39" pageNumber="549">Riazocoris nigra</emphasis>
:
<bibRefCitation id="EFBD4B136804FFC64F56653BFDA2FDAD" author="Ahmad, I. &amp; Afzal, M." box="[332,607,532,557]" pageId="39" pageNumber="549" pagination="1 - 14" refId="ref27835" refString="Ahmad, I. &amp; Afzal, M. (1979) Resurrection of the tribe Caystrini Stal (Heteroptera, Pentatomidae, Pentatominae) with description of two new genera from Oriental region. Annotationes Zoologicae et Botanicae, 133, 1 - 14." type="journal article" year="1979">Ahmad &amp; Afzal, 1979: 47</bibRefCitation>
</taxonomicName>
</treatmentCitation>
(redescription, illustrations, host plant);
<treatmentCitation id="0A8D10F36804FFC64DE3653BFB3DFDAD" author="Ahmad, I." box="[1017,1216,532,557]" page="137" pageId="39" pageNumber="549" year="1980">
<bibRefCitation id="EFBD4B136804FFC64DE3653BFB6CFDAD" author="Ahmad, I." box="[1017,1169,532,557]" pageId="39" pageNumber="549" pagination="115 - 155" refId="ref27587" refString="Ahmad, I. (1980) Insect fauna of Pakistan and Azad Kashmir - some groups within the order Hemiptera. Proceedings of the Pakistan Congress of Zoology, 1, 115 - 155." type="journal article" year="1980">Ahmad (1980)</bibRefCitation>
: 137
</treatmentCitation>
(list);
<treatmentCitation id="0A8D10F36804FFC64B1B653BFF1EFDCB" author="Ahmad, I." page="52" pageId="39" pageNumber="549" year="1981">
<bibRefCitation id="EFBD4B136804FFC64B1B653BFA64FDAD" author="Ahmad, I." box="[1281,1433,532,557]" pageId="39" pageNumber="549" pagination="1 - 113" refId="ref27623" refString="Ahmad, I. (1981) A revision of the superfamilies Coreoidea and Pentatomoidea (Heteroptera: Pentatomomorpha) from Pakistan, Azad Kashmir and Bangladesh. Part 1: Additions and corrections of coreid and pentatomid fauna with phylogenetic considerations. Supplement of the Entomological Society of Karachi, 4 (1), 1 - 113. [1979]" type="journal article" year="1981">Ahmad (1981)</bibRefCitation>
: 52
</treatmentCitation>
, 105, fig. 8 (checklist, host plant, illustration).
</treatmentCitationGroup>
</paragraph>
<paragraph id="8B9336E26804FFC64E8D6556FD6AFC57" blockId="39.[151,1437,633,984]" pageId="39" pageNumber="549">
<emphasis id="B958EAF06804FFC64E8D6556FEBAFD13" bold="true" box="[151,327,633,659]" pageId="39" pageNumber="549">Nomenclature.</emphasis>
The name
<taxonomicName id="4C2C4D616804FFC64FD66555FD76FD13" authorityName=": Ahmad &amp; Afzal" authorityYear="1979" box="[460,651,634,659]" class="Insecta" family="Pentatomidae" genus="Riazocoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="39" pageNumber="549" phylum="Arthropoda" rank="species" species="nigra">
<emphasis id="B958EAF06804FFC64FD66555FD76FD13" box="[460,651,634,659]" italics="true" pageId="39" pageNumber="549">Riazocoris nigra</emphasis>
</taxonomicName>
with explicitly given authorship of Ahmad and Afzal first appeared in morphological paper devoted to description of its internal organs by
<bibRefCitation id="EFBD4B136804FFC64DA165B2FB5FFD37" author="Ahmad, I. &amp; Afzal, M. &amp; Siddiqui, M. I." box="[955,1186,669,695]" pageId="39" pageNumber="549" pagination="161 - 165" refId="ref27925" refString="Ahmad, I., Afzal, M. &amp; Siddiqui, M. I. (1977) Aspects of morphology of Riazocoris nigra Ahmad and Afzal (Pentatominae: Caystrini) with phylogenetic considerations. Pakistan Journal of Zoology, 9 (2), 161 - 165." type="journal article" year="1977">
Ahmad
<emphasis id="B958EAF06804FFC64A0F65B1FBB3FD37" box="[1045,1102,669,695]" italics="true" pageId="39" pageNumber="549">et al.</emphasis>
(1977)
</bibRefCitation>
, which is sufficient to make it available from that publication. The intended original description appeared later in
<bibRefCitation id="EFBD4B136804FFC64A8265EEFF4CFD7F" author="Ahmad, I. &amp; Afzal, M." pageId="39" pageNumber="549" pagination="1 - 14" refId="ref27835" refString="Ahmad, I. &amp; Afzal, M. (1979) Resurrection of the tribe Caystrini Stal (Heteroptera, Pentatomidae, Pentatominae) with description of two new genera from Oriental region. Annotationes Zoologicae et Botanicae, 133, 1 - 14." type="journal article" year="1979">Ahmad &amp; Afzal (1979: 4)</bibRefCitation>
. The fact that
<taxonomicName id="4C2C4D616804FFC64F4265C9FE2DFD7F" authorityName="Ahmad &amp; Afzal" authorityYear="1979" box="[344,464,742,767]" class="Insecta" family="Pentatomidae" genus="Riazocoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="39" pageNumber="549" phylum="Arthropoda" rank="genus">
<emphasis id="B958EAF06804FFC64F4265C9FE2DFD7F" box="[344,464,742,767]" italics="true" pageId="39" pageNumber="549">Riazocoris</emphasis>
</taxonomicName>
has been combined with an unavailable generic name in
<bibRefCitation id="EFBD4B136804FFC64A7B65CAFAB4FD7F" author="Ahmad, I. &amp; Afzal, M. &amp; Siddiqui, M. I." box="[1121,1353,741,767]" pageId="39" pageNumber="549" pagination="161 - 165" refId="ref27925" refString="Ahmad, I., Afzal, M. &amp; Siddiqui, M. I. (1977) Aspects of morphology of Riazocoris nigra Ahmad and Afzal (Pentatominae: Caystrini) with phylogenetic considerations. Pakistan Journal of Zoology, 9 (2), 161 - 165." type="journal article" year="1977">
Ahmad
<emphasis id="B958EAF06804FFC64AA665C9FB0BFD7F" box="[1212,1270,741,767]" italics="true" pageId="39" pageNumber="549">et al.</emphasis>
(1977)
</bibRefCitation>
has no effect on its availability (see
<bibRefCitation id="EFBD4B136804FFC64FCF6426FDA4FCA3" author="ICZN" box="[469,601,777,803]" pageId="39" pageNumber="549" refId="ref31005" refString="ICZN (1999) International Code of Zoological Nomenclature. 4 th Edition. The International Trust for Zoological Nomenclature, London, 306 pp." type="book" year="1999">ICZN 1999</bibRefCitation>
: Art. 11.9.3.1). There is no mention of type material in
<bibRefCitation id="EFBD4B136804FFC64AAF6426FA65FCA3" author="Ahmad, I. &amp; Afzal, M. &amp; Siddiqui, M. I." box="[1205,1432,777,803]" pageId="39" pageNumber="549" pagination="161 - 165" refId="ref27925" refString="Ahmad, I., Afzal, M. &amp; Siddiqui, M. I. (1977) Aspects of morphology of Riazocoris nigra Ahmad and Afzal (Pentatominae: Caystrini) with phylogenetic considerations. Pakistan Journal of Zoology, 9 (2), 161 - 165." type="journal article" year="1977">
Ahmad
<emphasis id="B958EAF06804FFC64B146425FAB8FCA3" box="[1294,1349,777,803]" italics="true" pageId="39" pageNumber="549">et al.</emphasis>
(1977)
</bibRefCitation>
. As
<typeStatus id="549788406804FFC64EDA6402FEDFFCC7" box="[192,290,813,839]" pageId="39" pageNumber="549" type="holotype">holotype</typeStatus>
is defined as the single specimen upon which a new nominal species-group taxon is based in the original publication (
<bibRefCitation id="EFBD4B136804FFC64F90647EFDECFCEB" author="ICZN" box="[394,529,849,875]" pageId="39" pageNumber="549" refId="ref31005" refString="ICZN (1999) International Code of Zoological Nomenclature. 4 th Edition. The International Trust for Zoological Nomenclature, London, 306 pp." type="book" year="1999">ICZN 1999</bibRefCitation>
: Art. 73.1), the subsequently designated holotype by
<bibRefCitation id="EFBD4B136804FFC64A62647EFA61FCEC" author="Ahmad, I. &amp; Afzal, M." box="[1144,1436,849,876]" pageId="39" pageNumber="549" pagination="1 - 14" refId="ref27835" refString="Ahmad, I. &amp; Afzal, M. (1979) Resurrection of the tribe Caystrini Stal (Heteroptera, Pentatomidae, Pentatominae) with description of two new genera from Oriental region. Annotationes Zoologicae et Botanicae, 133, 1 - 14." type="journal article" year="1979">Ahmad &amp; Afzal (1979: 7)</bibRefCitation>
must be considered the
<typeStatus id="549788406804FFC64FB8645AFDF4FC0F" box="[418,521,885,911]" pageId="39" pageNumber="549" type="lectotype">lectotype</typeStatus>
according to
<bibRefCitation id="EFBD4B136804FFC64CB9645AFCCFFC0F" author="ICZN" box="[675,818,885,911]" pageId="39" pageNumber="549" refId="ref31005" refString="ICZN (1999) International Code of Zoological Nomenclature. 4 th Edition. The International Trust for Zoological Nomenclature, London, 306 pp." type="book" year="1999">ICZN (1999</bibRefCitation>
: Art. 74.5). As the generic names ending with -
<emphasis id="B958EAF06804FFC64B5D6459FA82FC0F" box="[1351,1407,886,911]" italics="true" pageId="39" pageNumber="549">coris</emphasis>
(a Latinized Greek noun meaning true bug) are masculines, the gender agreement of the species name is corrected as
<taxonomicName id="4C2C4D616804FFC64E8D6491FF0CFC57" authorityName="Ahmad &amp; Afzal" authorityYear="1977" box="[151,241,958,983]" class="Insecta" family="Pentatomidae" genus="Riazocoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="39" pageNumber="549" phylum="Arthropoda" rank="species" species="niger">
<emphasis id="B958EAF06804FFC64E8D6491FF0CFC57" box="[151,241,958,983]" italics="true" pageId="39" pageNumber="549">R. niger</emphasis>
</taxonomicName>
according to
<bibRefCitation id="EFBD4B136804FFC64F916492FDE5FC57" author="ICZN" box="[395,536,957,983]" pageId="39" pageNumber="549" refId="ref31005" refString="ICZN (1999) International Code of Zoological Nomenclature. 4 th Edition. The International Trust for Zoological Nomenclature, London, 306 pp." type="book" year="1999">ICZN (1999</bibRefCitation>
: Art. 31.2).
</paragraph>
</subSubSection>
</treatment>
</document>